Coloquios de Palcontologia, 2002,53: 33-68 ISSN: 1132-1660 An annotated check-list of genera and species of carpoids Lista anotada de los géneros y especies de carpoideos
Patricio DoMíNcuEz ALONSO1’2, Richard RS. JEFItERIES1 & Dolores GIL CID2
Ábstract: Tisis ciseek-list includes liso al presoní valid names of genera and species of liso carpoids. It is a provisional atíempí lo stabili- ze tse names of MI fornially described carpeids up te date and includes mosí of Ihe combinations of genera and species that bayo been used in tSe seicotifie literatsjre, It sisould be especially useful re tisose roading oidor papers in wbicis a wide variety of genera and spe- cies Sayo beco used, and te tisose iriterested in biodiversity, evelutionaiy studies or isistory of the palaeontology. Tise ciseclc-list is divi- ded mío five sections: 1. índex ofgenera; II. índex of species; Hl. índex of valid speciflc epithets; IV. Tables of stratigrapisical occurrence and y. Tiso mosí complete and up-to-date lisí of references en Ihe taxonomy of carpoids. Key words: ciseck-lisl, index nonrioorrrnr, Seluta, Cornuta, Mitrata, Ctenocystoidea, Cineta, calciciserdates, isomalezoans, stylopisorans.
Resumen: Esta lista recoge los génerc>s y especies actualmente válidos de carpoideos. La lista incluye la práctica totalidad de las cera- binaciones de género y especie que San sido empleadas en la literatura científica. No es una lista exhaustiva de sinonimias pero se ofre- ceo referencias clave para elaborarlas. Este trabajo debería sordo utilidad para aquellos lectores interesados en obras antiguas donde una confusa variedad de formas son descritas o referenciadas y para aquellos interesados en paleodiversidad, procesos evolutivos o isistoria de la paleontología. La lista se encuentra dividida en cinco secciones principales: 1. Índice de géneros; II. índice de especies; III. Indice de epítetos específicos válidos; IV. Tablas de distribución estratigráfica y V. la más completa y actssaalizada relación de referencias biblio- gráficas sobre taxomía de carpoideos. Pajabras clave: indice, índex norniooru,n, Soluta, Comuta, Mitrata, Ctenocystoidea, Ciocta, calcicordados, Homalozna, Stylopisom.
INTRODUCTION In Ihis chock-list a total of 179 species of carpoids aro alphabetically rocorded by Classes (including 22 The carpoids are an informal grouping of Palaeo- solutos, 52 cornutos, 75 mitrales, 24 cinctans and 6 zoic fossils wisich hayo a skolelon ofecisinoder¡n-like ctenocystoids). It is a provisional attempt lo stabilize calcito buí Iack radial symmetry based 00 tSe water be names of alí forrnally described carpoids up lo vascular system. date and includes mosí of the combinatioras of gene- They aro isighly controvorsial being regarded by raand species lhat hayo beon usod in bibliography. It sorne as aberrant echinoderms and by others as mem- should be especially useful for tisose reading oidor bers of liso stem groups of many of Ihe major sub- papers in which a wide variety of genera and spocies groups of deuterostomes (craniales, tunicates, acra- Sayo been usod, and lo thoso interested in biodiver- niales, chordates and conceivably of tSe sity and evolutionary sludies. When placing species echinodem~s, isemichordates, ambulacraria and por- in particular gonera we nocessarily oxpress our own haps of Iho deuterostomes lhemselves). personal opinions. No attempt at pisylogenetic orde- TSe aims of Ihe prosoní work are bibliographical ring has been made. TSe sequence is puroly alpisabe- so we do nol consider the stem-group position of tSe tic. TSe check-list comprises five main sections: various subgroups of carpoids. Instead, we use as groups liso Jaokolian orders plus liso ctonocystoids 1. índex of gonera and within tiseso groups wo arrange tSe genera and II. índex of species species in alphabetical ondeE III. índex of valid specific names IV. Tables of stratigraphical occurrence
¡ Departmení of Palaoontology, TSe Natural History Museuni, London SW7 SBD, 1.3K. [email protected]; rpsj@?nhmacuk Departamento de Paleontología & Instituto de Geología Económica (UCM-CSIC), Fac. CC. Geológicas. Universidad Complu- tense de Madrid, 28040 Madrid, Spain. [email protected]; [email protected]
33 /‘esíiioi’s í)or.5551555c: Alsossto & ¿5:[]. Xis sistroraled check-lisl of gersera asid species of carpoids
100% 50 90 ‘e, 5 80 ‘5, 40 70 a 60 30 50 a 40 20 5 30 2 10 20 ~‘II 10 o 31 0000 0 0000 0000
505 55 90 50 909209 9409 09c01975519990515520® Ficure 2.- Valid described species of carpoids per decado (iSick Fietsre 1- Cusnulaíi»e nuortrbcr of al proseo] das salid described lino) and cutnulati»e percenlage sf »alid described species por sísec¡es ssf carpoids por sean. decade (lisio I¡ae). /-eQisres 1,— :Vsose’ro eseíonu!¿it/ss dc copes ‘es así ro/mcta le ió!ie/css figura 2.- Espee:ics válidas dc earpoideos descritas por décadas s/’ eosj’oielc:os s/sas 54<55 usi ¿sises. (en liesea gruesa) y isoreeniaie acumulado de especies válidas ele’scritas por clúcades (línea tina).
TSe ciseck-list is based típon valid iaxenoniic lite- dala. Tho syrnboí sc >» associated with a namo emp- roture necords ond is arranged taxonomicaíly by Class, isasizes tSe eross-reference íisat musí be followed. ISeo alpisobetically by Gonus and Spoeios witbin oacb Tubbs oJsírodgraphieol occíctrence (Sectien IV) list Class. Entiles o liso hijeo ofGenero (Section 1) inclu- te spocies ordeíing tisom by suratigropisical oceurrence de: Genus; A utisor(s) wiso desenibed liso genus; dato of and ISeo, witisin caeS suage, o alpisabetical orden Dala [br tSe stages aro based en Fo.ssil Record 2 BENTON, liso original pníblication; type species (using uSe origi- nal spelling, ciseck So Índex of .spec:icfl: ouínber of i993 in orden lo relato oursuratigrapisical dala witb equi- specios: included specios (¡So syrnbel e”’» denotes liso valeor doto of otisor groups witiseut preposing a new lype species); stratigrapisical occurrenco; goegrapisical worldwide siraligrapisical correlation. Re nomos of liso distnibiruion; kev relerenees. Entries in the Índex oJ síagos are also reforrod lo in Soctiens 1 and II. Recently doscribod spocios are usually not diffrcult te assign lo a Speeics (Seclion II) includo: combination of genusond specicsúscientific binomir.a);Autisor(s} wisodescnibed particular siage. Olderspocieshíwcvcr~ especiahlytho- se based 00 Sistorical colleclions with only a general riso species (in pa¡entiseses if liso specros rs new ossre- ned lo a clifforetñ genus tisan wison desenibed bv aul- lecation in tSe bibliegrapisy or in Iho labels accompan- ying tSe specimens, are diffrcull lo assign stratigraphi- Sor): date r>f Ibe original pubíication; autisor(s) of tSe cally. lo tiseso casos liso species oppoar in liso labIo with new combí o
Gsñrsc¡sócss lIs’ /‘os)esnílolrsgíss 34 REFERENCES: THORAL, 1935b; UBACHS & CASTER in CASTER, Example 1, a genus nanie: 1967 Lagynocystis JAnKEL, 1918 THoI¿AL, 1935 described a new Soluta as Den- TYI’e 5PECiES: Ano,nalocystites pyrarnidalis (BARRANDE, 1887) drocystites 1/idali, a combination no longer in use NUtrSABER OF srecies: 1 (and Iherefore quoted in square brackels «[1») since t (BAima~oe, INCLUDEI) seecios: Lagynocysíis pyrarnidalis UBÁGES & CASTER, 1967 ira Ihe Treatise on Inverte- 1887) brate Palaeontology (CASTER, [967) proposed Iho STRATiGRAPHicPJ. OCCURRENCE: Lower-Middle Ordovician (Llanviru) genus Minervaecyshs for this soluto and so Iho new GEocítAprircAL DiSIRiBUTiON: Europe (Bohemia, France, combination Minervaecystis vidalí (THORAL, 1935) Spain) became available. The symbol «=» before Minerva- REFERENCES: BAi&4y0e, 1887; CHAUvEL, 1941; CHAUvEL & ecystis vidalí thus indicales Iho currently used NíeN, 1977; JAEKEL, 1918; JEFFI5RrFS, 1973; PAR5LEY, 2000; synonym arad acts as a cross roference lo the cisock- UBAcris, 1991 lisí where moro information is available. The symbol «-~» emphasizes thaI cross roferonce. Note also dio JAEKEL, 1918 proposedthe genus Lagynocystis for capitalised «Vidali» as originally describod. Anomalocystites pyramidalis (BARRANDE, 1887) (which incidentally became type specios by monotypy). Lagynocystis JAEKEL, 1918 is in bold CHECK-LIST OF CARPOIDS type, showing a valid genus natne; thore is only ono valid species doscribed (we inusí search in Ihe Check- 1. íNDEX OF GENERA lisí ofthe species naenes); te specios includod is Lagy- nocystis pyramidalis (BARRANDE, 1887), te symbol * Cineta JAEKEL, 1918 indicates tal Ihis is tho typo species of te gonus; te slratigraphical occurrence and liso geographieal distri- Asturicysds SDZUY, 1993 buhen are given. Finally, somo roferonces are usted. TYPE sPECrEs: Asturicystisjaekeli Sdzuy, [993 NUMBER OF 5PECiE5~ 2 Example 2, a salid species name: INCLUDED spEctEs: Asturicystisjaekeli* SO/UY, 1993; A. /íavliceki FATRA & KORDULE, 2001 Sucocystis melendezi (SCHROEDER, 1973); STRATIGRAPOICAL OCCURRENCE: Middle Cambrian (St. Davids) FRIEDRIcE, 1993 GEOGRAPHíCAL DiSTRIBUTiON: Europe (Spain: Asturias; [=Gyrocystis?,nelendezi SCOROEDER, 19731 Boisemia) l=Troclíocinctus pardail/sanicus TERMíER & TErr=drER,1973] REFERENCES: SDZUY, 1993; FATKA & KORDULE, 2001 STRATicIRAPHrcAL OCcURRENCE: Middle Cambrian (St. David’s) [Cepí’íalocínctusTERMIER & TonseíioR, 1973] GEooRAPurcAL DisTRinurrioN: Europe (Spain. France) =Elliptocinctus TERMíER & TERMíER, [973 > REitRENCES: ScFrRoEnere, [973;F~rorwrcu, 1993 REFERENCES: TERMIER & TERMIER, 1973; FíuroDr<íCH, [993 SCHROEDER, 1973 described Gyrocys/is? melendezi; but lator FRIEDRICH, 1993 proposed te new combina- [Cyclocinc/usTERMíER & TERMIER, 1973] tion Sucocystis melendezi. Troehocinetus pardailhani- parima =Elliptocinctus TERMíER & TERMiER, 1973 cus TERMIER & TERMIER, 1973 is a junior synonym. lElliprocincrus barrandei (MIsNíER-CHALMAs & BERGE- RON, 1889)J Since Sucocystis melendezi (SCHROEDER, 1973) is considerod valid, it is lo bold type in tho check-Iisl; partimn =7 Sucocystís CABraEL, TERMIER & TER>AíER, 1959 [Sucocystis?enaroccana (TERIIíER & TERMíER, however bolh synonyms appear in squaro brackets 1973)1 -9< «[ ]». The symbol «=» previous te Iho names indicates REPERENCES: TERMíER & TERMIER, 1973; FRrEDiaicH, te synonym as well as a cross in te check-Iist. 1993 Example 3, a species name no Ionger used: Davidocinctus FRIEDRICE, 1993 Tv~o 5PECiE5: Davidocinctr¿s pembrokensis FrucoRieR, 1993 [DendrnqstitesVldnIi NeRss, 19351 NUMBER OF spoci~5: 1 =Minervaecys/is vidail (TIIORAL, 1935); UnAcris & CASFER, INCLUDED svrserrss: Davidocinetus pe,nbrokensis* FRirsoRien, 1967 —> [993 35 Coloquios de Paleontología 2002, 53: 33-68 Patricio Domínguez Alonso & ci al, An asisrosated chcck-lisí of genera and species of carpoids STRATic,rcAi’urcÁr occi:RRENce: Middle Cambrian (St. Sotociuctus Síztsy, 1993 David s) Tyr’u sr’ociss: Sotocioctas aboghsi SDZUY, l993 NUMBER OF .Sí’ECiE.Y 1 Gí¿:ocrowrsícAt, DIsTRiBUTiON: Europe (UK: Wales) INcLUOLO sr>ecItis: Sotociucías ubaghsi9< Sozev, 1993: Rer:ertoscms: FRreoRrcH, l 993 STRATiIRAFHiCAi OCGURRENCE: Midd[e Cambrian (St. [DecacestisGíSLÉN, 1927] Davids) GeoGRAPuicÁl. DiSTRiBUTiON: Europe (Spain: Asturias) =Gvrocystis JAF.KíuI., 1918 ReFeRísNí:rzs: FrsieríRicrr. 993; GíSLÉN, 1927 Ru¡r:íLNeLs, Suar’», 1993 Sucocystis CABIBEL, TERMIER & TERMIER, 1959 Elliptocinetus TERMtER & TtiRMlriu, 973 Toc srEciF~: Sí¡co Gyrocystis JAEKEL, 1918 [Troehocysti.snorra. van.] Typn spízcíes: Ge roeystis platessa JAEREL, 1918 — Troehocvstiíes BARRANDE, 1887 NuMiseR OF 51’FCib5: 6 INCLUDID 5PECi[5: Gírocysris badalesiensis Fuirmnuico, 1993: G. eruzae GIL & DoMÍrse,Uez, 1995; G. erecta FRíE- Trochocystites BARRANDE, 1859 TYPE SPEeií’s: Tros:hocvsri/es bohemicus BARRANDE, l887 bRiOl, 1993; G. hispan/ea (GisrÉN, 1927), [dubiousspe- NUMrsER OF 5PECiii5: 1 cies]; G. plales.set* JAEKEL, 1918; G. íestudíformis FRrE- INcLsSOED sPeciEs: Troclrocystiícs bohemicus * BARRANDE, i)RitAi, l993 STrtsric.wwoicAí. O(CURRLNCE: Middle Cambrian (St. 1887 STRATíGRAPHíCAL OGCURRENGE: Middle Cambrian (St. David’s) flavids3 GÉdcít&14fi¿ÁL bétRiñÚTriYp¿ EúfoÑ (S¡SaiitFraoce) GEoGRAFHicM. Oi5TRiBUTiON: Europe (Boisemia) REFERENGES: JAGKtSL. 1918; FRieonJcii, 1993; Go. Cro & NOTE: Mosí early descriplions of Ciocta were given as Tro- DOMÍNGUEZ, [994 elíocy.stitcs or Troclrocystis RrserssENcts: BARIG\NOE, 1859, [887;JAEREL, 1918; TErzMir¿u Ludwigicinctus FRIEDRIcO, 1993 & TERMíER. 1973; FRiFDRico, [993;DOMíNGUEZ, 1999 tít 5PECiE5: Ludwigiei,rctas trancatos FRíeDRíco, 1993 NísMnízR 01’ 5FEcrE5: 1 TrochocystoUes JAEKEL, 1918 lNctuJrsFo sítcííes: Ludwig/cinctus lruncatus* FrirEoRreo, Twr. SPEctr5S: Trochocys¡oides parvas JAEKEL, l91 8 1993 NUMBER OF 5PEGitS: 2 STresriuRAurriesí. OGCISRRENCE: Middle Cambrian (St. INCLUDED SitúEs: Trochocysíoides parias> JAEKEL, 1918; T David s) planas SDZUY, 1993 GEoGiZÁPHrCAL OiSTRiOiiTiON: Europe (Germany) STRxriGrtAprircAr, OcCURRENCE: Middlc Cambrian (St. RereRrNcu.s.FurLoRicH, 1993 Da»ids) GEOGRÁrtiCAL DiSTRiBUTiON: Europe (Boisemia, Spain) Progyrocystis FRILDRtCH, [993 ReFaReNeas: JAEREL, 1918; Fuíeourco, ¡993; Soso», l993 Typiy smzcies: Progvrocvsíi.s disjaneía FRreríí Coloquios de Pe¿/eontología 36 2tM)2, 53: 33-68 Patricio Domí,rgrsez Alonso & 01 al. An arinotated check-iist of gorrera and species of caspoids NUMBER OF sPEciEs: 4 Coleicarpus DALEY, 1996 INCLUDED sPEciF.s: Cienocystis eolo don UBAOIIS & RouísoN, TYPE 5PECIES: Castericystis? sprinklei UBAGHS & RoarsoN, 1988; C. jagol JELL, BURRET & BANK5, 1985; C. smithi 1988 UDAGOs, [987; C. utalíensis> RoísrsoN & SPr¿iNiwe, 1969 NUMBER OF 5PEIZiEs: 1 STRATíGRAPHIIIAL OCCURRENCE: Middle Cambrian (St. INcLUDEY sPECIE5: Coleicarpus sprinklei> (UBAGHS & Rení- David’s) sON, 1988) GEOcrtár’uicsL OiSTRiBUTiI)N: Nertis Amorica = Dendrocystites BARRANDE, 1887 -4 Jugoszovia L»íK & OaLowsKi, 1995 TYPE 5PECiE5: .Iugoszosna usehueoeyathoides D~ii< & OruowsKi, 1995 Dendrocystites BARRANDE, 1887 NUMBER OF 5PECiB5: 1 TYPE 5PECiE5: Cystidea sedgwicki BARRANDE, 1867 INCLUDED 5PECiE5: Jugoszovio arehaeocvarhoides> D~iK & NUMBER OF SEECiES: 2 Caxowsra, [995 INCLUDED 5PECiFS: Dendrocystites barrandei (BATHER, 1913); D. sedgwicki> (BARRANDE, 1867) STRATiGrosr’oicAL OCCURRENCE: N4iddle Cambrian (St. David’s) STRATrorcAí’tsrcAL Ot?cURRENcE: Europe (Bohemia) GEOGRAPOWAL DiSTRiBUTiON: Europe (Poland) GEOGRAPHICAL DiSTRiBUTiON: Midd[e Ordovician (Llandeilo) Raí’ErwNcEs: BARRANDE, 1887; BATHER, 1913; Jsrva, 1918 REFERENCES - D2iK & DRaowsKr, [995;DOMÍNGUEZ, 1999 Dendrocysto ¡des JAEKEL, 1918 TYPF 5PECiES: Dendrocystis scotica BATHER, 1913 Soluta JAEKEL, [901 NUMBER oc sPFCirtS: 1 INCLUDED SPECIES: Dendrocysíoides scoiicus* (BATHER, Belemnocyslites MILLER & GrsRLr±y,1894 1913) TYPE 5PEI?iE5: Bclemnocysíites weíherhyi MILLER & GURLEY, STRAnGRAPoIcÁí. oCCURRENCE: Upper Ordovician (Asisgilí) 1894 GEOORAPHíCAL DisTRiBUriON: Europe (UK: Scotland) NUMBER OF 5P~cics: 1 RerosaNcEs: BATHER, [913; JAEKEL, 1918 JEPFERIES, 1990 INCLUDED sPísciEs: Relemnocv.síi¿es weeherbyi* MiLLER & GURLEY, 1894 Girvanicyslis CASTER, 1967 [1968] STRATiGruxi’í]icÁL OCcURRENCE: Uppor Ordovician (Caradoc) Ts~e SPEIíE5: Girvanicyslis baiherí CASTER, 1967 [19681 GEoGRÁPHicA¡. DISTRiBUTiON: Nortis America (USA) NUMBER OF 5PECiE5: 2 REFERENCF5: MILLER & GURLEV, [894; PAR5LEY, [972; INCLUDED SPECiE5: Girvanicystis bat/ter? CASTos, 1968; G. KOLxíA cl al,, l977 casterí DALEY, [992 STRATIGRAPIIICAL OCCURPENCE: Upper Ordovician (Ashgill) Castericystis UBACES & RoBísoN, 1985 GeooRAPíircAL DiSTRiBUTiON: Europe (UK: Scotland; South TvPE 5PECiE5: Gasiericystis vali UBAGH5 & Ronrsors, 1985 Wales) NUMBER risc SFECiE5: 1 REFERENCES: CASTER, 1967; DALEY, 1992 INCLUDED 5FECiE5: Castericysíis valí> UBAGHS & Ronrsors, 1985 lleckericystis GiL & CASTER, 1960 STRósiGasporcAL OCCURRENCI]: Middle Cambrian (St. TYPE SPECiE5: Dendrocysti/es kackersiana HECKER, 1940 David’s) NUMBER OF SPECiE5~ 1 GEOGRAPHíCAL DiSTRiBUIrON: Nortis America (USA) INCLUDED Sí’ECiES: Heckericvstis kackersiana* (HECKER, REFERENciES: UnAuHs & RonísoN, 1985: DALEY, 1996 1940) 37 Coloquios de Paleonlología 2002, 53: 33-68 Patricio Dssnsíííguez A/o,ísss & O al Ata annotated check-iist of genera and apecies of carpoids STítxTíc;aÁs’criíÁi, oeetsRRisNr:E: Middle Ordovician (Llandeilo) Scalenocystites KOLATA, 1973 GEoGR’rííícesi, DrsTRinrsTioN: Ecropo (Russia: St. Pelersburg TYPE SPECiE5: Scalenocystites striníplei K0ATA, 1973 area) NUMBER OF 5PECiES~ 1 RísnízaisNci.s, HíseKísa. 194(1; Gui, & CAs-res, 1961) INCLUDED SPECiES: Scalcnocysthes strimplei> KOLATA, 1973 STRATíGRArHíCAL OcCURRENcE: Upper Ordovician (Caradee) Jowacyslis THOMAS & LADD, 1926 GEOGRAPHíCAL Oi5TRiBtJTiON: Nortis America (USA: Minne- Ts~o suecias: lesrs’aeystis sagitiaria THOMAS & L>rno, [926 sola) NrssrrsoR ob sPtsr:sn.s: 1 RUFERENCES: KOLATA, [973a, 19735; ROLATA el al., [977 lNeLr:rSEL) 5PFtT7iE5: Ioss’aeystis sagittaria * TuoMÁs & LADO, ¡926 Syringocrinus ErLi.tNús. 1859 STaxrií;RsF’elrt:Ás, (Sec:r;RRENI:F:: tipper Ordovician (Caradoe- Ter’r¿ sueCias: Syringocrinus paradoxicas BíLLíNOS, 1859 Asisgilí) NtsMrnsR or sPF.crtis: 1 GcOGRÁPí]rcst. t}r5TRiBUTrON: Nortis Amorica (USA: Iowa) INCIUDISD sPíwrF.S: Syringocrinus paradoxicus> Brí.LrNGS, REFERENCES: THoMAs & LADD. 1926; PxRsrísy & CASTER, [859;5?. sinclairi PARSLEY & CASTraR, 1965 [965;RelATA es al., [977 STRsxTIGRÁroícAL oCI?URRENcE: Upper Ordovician (Caradoc) GF.OORAI’HíCAL Di5TRiBSsTrON: Nortis America (Canada) Maennilia RozílNov & JEFFERIES, [996 REresErsí:es: BiLLiNGs, 1859; PAR5LEY & CASTOS, 1965; CAs- fyros srrzerr:s: Maevínilia eslotiica Roztsr*ov & ifappesies, tEle, [967 996 rÑsJMBER 1SF SFEciE5: 1 Comuta JAEKEL, 1901 [NCLitOiiOSPttiES: fiel oen,iilia cstooica * ROZHNOs & JH±E- Ríes, l996 Acuticarpus SLsINIRALL, SPRINKLE & GUENSBURG, 1997 STrovrrc.ÑesPr riCAL OCCISRRíZNCU: Upper Ordovician (Caradoc) Tvua SuralEs: Acaticarpus delticas SUMRALL, SPRiNELE & Geooíosrtuicxt, oisTRiuuxros: Europe (Estonia) GUEN5BI5RG, [997 RereRísr=cLs:RoztiNov & JEFFERiE5, 1 996 NUMBER 1SF SPBCiBS: 2 INCLUDEO suecias: Aculicarpus delticus> SUMRALL, SPRiNKLE Minervaecystis UBAGHS & CÁSTISR, 1967 & GUEN5OURG, 1997; A.? repablicensis SUMRALL, SPRireK¿.E TYI’E 5Fi&iLS: Dendrocystiíes Vidalí TíroRÁs., 1935 & GUFNSBIsRG, 997 NUMBER OF SFECiE5: 1 SrRÁTrc;RAPOíGAL OcCURRENGE: Upper Cambrian (Merionetis) INCLSJOLi) 5PEeiE5: Mincrvaeevslis vidali> (THORAL, l935) GEoCRAPricAL DiSTRiBUTiON: Nortis America (USA) Srí=ÁriovsAi’Hícxr.sísIcIsRRLNcE: Lower Ordovician (Trema- REFERENCES: SUMRALL etat, 1997 doc) CieOGrtxruicAi, DiStRiisUTiON: Europe (France) Amygdaiotlrrra IIRA Coloquios cíe Paleoís:o/ogkí 38 2002. 53: 33-68 Patricio Domínguez Alonso & el al. Ata annorated check-iist nl genera atad species ol carpoids STRATIGRAPHICAL OCCURRENCE: Lower Ordovician (Arenig) gutnentaiy platos, liso length of Ihe anterior appendicos and GEoORArrrrcAL oIsTRiBUrIoN: Europe (France) liso ventral spikos of liso marginal trame are veiy difrerení in REFERENCES: LEPEUsRE & VizcAíNo, 1999 liso two genera. STRATíGRAPIUCAL OCtIURRENCE: Middle-Upper Ordovícran Archaeocothurnus SUMRALL, SPRINKLE & OUENS- (Llanvim-Caradoc) BURG, 1997 GEOGRAPHICAL DiSTRIBUTiON: Europe (Bohemia, Spain, Fran- TYPE SPECiES: Co//taníocystis? b 39 Coloquios de Paleontología 2002, 53: 33-68 I’assie(ss I)essníssg’sív: Alssn.ss, & ci al. An arsnotascd chcck-list ol gencra aud spccics ol carpoids Milonicystis CHAUVEL, [986 INGitSDED SuElES: Dí-epaoocarpos auzcí culis> SMITH & JELL, [999 Tvun surcrrs: Milooicvstis kerfornei CHAUvEI., [986 Srre.xrrcRAuhrrcÁs ocnt7rsRRFNriz: Upper Caníbrian (Merionelis) Ns>sroue OF SPLtIIES: 1 Giness:,roxpsriexL Di5TRSBISTiON’: Asísíralia lsci í DPi) SRFCIES: ¡í4ilooisvsiis kccJt cocí> CwsUvizv, 1986 STle’sTIGR’pcsIc.sl OtIslISRREN(U: Middlo Ordovician (Llaodei- RSSIERIiN(r:1S: Svrrrs & liii.. [999 lo) Flabellicystis Mxíeri MUs, 2002 GEoeauÁs’ctscxL DisrRIBuIrioN: Europe (France) Rrvrarsers: (‘russsvní - 1986: Caruus & Dsr.ss, [994 lyi’L Si’ LUí LS. Hsibellicvstis rushtoni MARTÍ MliS. 2002 Ns’Mlt:rs OF 5t’F(iFS: 1 Nanocarpus UBAGHS [991 ¡NUI AS 1)111) 5 Pce ir:s~ Elote/líe: vstize sushtooi* MARTÍ Mlis.20(12 Ssssxrsz;seÁ¡’r-sscxr, OCtURiCLNULi Lower Ordovician (Tronra- TYPE SuElES: Naooezrr-pus dolavobíi tirs,scrrs, l 991 dsse) Nr SM RtiR 5SF 5 upicsses: (3szsss;re,ss’r=ss:,xr,r:sssTs{rBs:rsssN: Esísope Esiglandí 1 NSTIl nEc) surcrrs: Neioosac~su.s eloleimbu 9< ti BAGHs, 1 991 S-FICATs;ÑssuHrcÁs esvcutI Rsí-nur’cr s Re r is. 1 999b Ponluulocarpus SUMRALL & SERIORLE. 1999 Tvss SPECIES: Pssoiis:oloearpus cobí.soni SuSMRALL & Susís- Lobocarpus tií}AGHS, 998 Txs’r: sr’ssr rs /.ssbosesspír sv/zcssiooíL’tsÁesus, ¡998 ELE. 999 Nusstrsse OF 55’LCiLS: ¡ Nsevícs OF sl’I:crrzs: ¡ INS:LS]DED suE(:sEs: Po,síículocarpus resbíso,rí Ss:MRss 1. & IxeruoFIs sr’rxir:s: Lesbsssesryrus sizeoiooi> L’Fs,scniss. 1998 tpper Cambri sss (Mcrismnelis) Srrsíwmc. [999 Sise (esloqshsss sIc Psils’O,s teslss5’ísi 40 9l~fl 53 15-68 Patricio Domínguez Alonso & el al. An anriorated check-iist ot genera and specÍes of carpoids GEOGRAPHIcAL DIsTRIBUTION: Europa (UK: England) GEOGRAPHICAL DISTRmUrioN: Europe (Bohemia, Franco) REFERENCieS: DxtEe, [992 REFEREN5IE5: JEFFERIIJS & PROROr’, [972; JEFFERIE5, 1986 Procothurnocystis WooDs & JEFEEmES, [992 Scotiaecystis CASTER & UBACHS, 1967 TYPE sí>btIIiss: Procotlrarnocysti.s owe,ísi WoODS & JEEFERIES, Tyurs SPECiE5: Coí/turnoc-istis carta/a BATHER, l913 1992 NíssrBER OF SPECIES: l INCLUDED SPECiES: Scoíiaecystis collapsa CRIPr’s, 1988; £ NUsrímísR (SET SPEIiES: 1 9< (BATHER, l913) curva/a INCLUDED SPEcIES: Procoíbunrocystís oveensi> Weons & jiep- STRATIGRAPHICAL OCCURRENCE: Upper Ordovician (Asisgilí) FERIES, 1992 Geograpisical distribution: Europe (UK: Scotland, Nortiseos STRATIGRAIHICAL OtICIJRRENCE: Lower Ordovician (Arenig) Ireland) GEOGRAPHICAL DISTRIBUTION: Europe (UK: Wa[os) REFERENCES: BATHER, 1913; CASTER & UBAGOS in UBAOHS, Rí±FERISNcEs:Wooos & JEFETERIES. [992 [969; JEPFEFSIES, 1967, [968, [986; CRíuPS, 1988 Progalliacystis CRIPPS, 1989 Thoralicystis CHAUX’EL, 1971 TYPE SPIJCIES: Progallíasysti.s ubaghsi CRIuus, 989 TePE 5PF.CIFS: Scotiaecisss/is grrffei UBAGFS5, 1969 NUMBER 01’ SuElES: 1 NUMBER OF SuECiaS: 4 INCLIsDED SFECiI5s: Progalliacvstí.s abag/tsi> Cssír’r’s, 1989 INCLUDED SuElES: Thoralicystis grrffeí* (UBAGHS, 1969); 7? STRATJGRAPHILAL OCCURRENCE: Lower Ordovician (Arenig) ubaghsi LEETEBVRE &VIzcsiNo, 1999; 7? zagoraensis CHAU- GEOGRAPHICAL DISTRInIJTIoN: Europe (France) VFL. [971 STRATIGRAI’HICAL OcCURRF.NCE: Lower Ordovician (Arenig) REFERENUES: UBAGHS, 1969: Cnsi’r’s, [989 GEOGRAPHICAL DISTRIBUTION: Europe (France), Nortisero Prokopicystis CRIPPS, 1989 Africa (Morocco) REFERENCES: UBAGHS, 1969; CHAUvEI., 1971 Tepie srieciES: Prokopicysíi.s mes~g/i CRíuPs, [989 NUMBER OF 51’EtiIES: 1 Trigonocarpus UBAGES, 1994 INcLUDEI> SPECIES: Prokopicvstis voergíi”’ Crapr’s, [989 TePE suuinS: Trígonocarpus singularis UBAGHS, 1994 STRxrioRxrHICAi, OCCURRENUL: Middle Ordovician (Llandel- NUMBER OF SPECIE5~ 1 lo) INCLUDED suEciEs: Trigonocarpus singularís* Uaxous, 1994 GEOGRAFHICAL DiSrRiBISTIoN: Europe (Boisemia) STRATIGRAPOICAL osXURRENCE: Lower Ordovician (Arenig) REFERENCUS: CRIPPS, [989 GEOGRAPIII5?AL DISTRIBUTION: Europe (France) REETERENCES: UBAGH5, [994 Proscotiaecystis UBAGIIS, 1994 TePE suiesnsies: Cot/turnsscvsli.s ,nelchiori UBAOHS, 1983 NCMBER OF SPETUIES: 1 Mitrata JAEKEL, 1901 [rsctUDr’.DSPECiETS: Proscotiaec:vslis orelchiori> (UBAOHS, 1983) STRAT5GRAPHICAL oCCURRENcI:: Lower Ordovician (Arenigí Adoketocarpus RUTA & JELL, 1999 GeOGíuAr’íIIcAs, DiSTRSBUTION: Europe (France) Tear’ sPEcíE.s: Adoketocarpus oc/reron/icus RUTA & JELL, 1999 REETERLN5iIJS: UFSAGES, 1994 NUMBER or’ sp~cries: 2 INCLUDED SPECIES: Adoketocarpus acheronticas> RUTA & Protocystites HICKS, 1872 JELL, 1999; A. janeae RUTA & JELL. 1999 TePE sue 41 Coloquios de Paleontología 21102, 53: 33-68 Patricio Domínguez A/miso & el al. An annorared checle-lisí of gesrera and species of caspoids [NCi.UDFDSFECIES: Ana/sfopsis acato BARRANDE, [872; A. S1rsxriGí*xuHicu OCCURRENCE: Middle-Upper Ordovician ancora DOsrÍNGos-iz & GLTIÉRREZ-MARCO. 1 990; A. arProri- (Llandeil[o-Caradoc) tana (CHAUVEL, ¡941); A. ba/claíclsíensí.s RitEn, 1940; A. GEOGRAPHICAL DISTRiBUTiOiÑ , Nortis America (USA) borrondci (Círsinvar.. 1941); A. bohemíca> BARNANDL /872; REFERENCES: HAECKEL, 1896; CASTER in UBAGHS, 1967c; A. cocaban KouxvxsHi, 1960; A. escandei escandei THORAL, KOLATA & JOLLiE, 1982 ¡935; A. escande/Y t/rorali (CHAP/EL, [966);A. gaicheííensis (CuxisvEi., 1941): Aootí¡ópsis longa BARRANDE, [872; A. Australocysñs CASTER, 1954 minuto (CI]xísvcL, [941): A. popillata (BASSLER, 943); A. lira SPECIES: Ausíralocystis laogeí CASTER, 1954 panctata (BAsSIFR, >950): A. spínosa IJBAGHS, 1979; A. Ira- NUMBER OF SuEcIES: 1 pezisfor,nis Triouxr , 1 935; A. travea.so/ensis (CrsAhsvcr., INCLUDED 55’iSCiES: Austraíocysiis langei> CASTER, [954 [941);A. lrsorcoto Kowxs’xsrsí, 1960 STRxsIGRAPIIWÁI. OCCURRENCE: Lower Devonian (Emsian) SnArIGRArIiíc::sí. IXSCURRENCE: Middle-tipper Ordovician GFoGIssptssCÁr, DISTRiBUTiON: Australia (L¡anvinm-AsSgill) RFFERF\CES (ASTER, 1954a Guoc;axuHIcxt. DIsTRIBsJTION: Europe (Boisemia, France, Spain, Sweden, 11K), NorIS Amorica (tISA), Asia (Korea), Balanocystites BARRANDE, 1887 Nertisorn Africa (Morecco) L=Sagitlac~yatis PARSLEY, PROROr & DERSrLER, 2000] REFERENCES: BARRANDF-., 1872; JrsI’rwwíEs, [986; DoMir=oL+z TYPE sFE(isis: Balonocysíites lagenala BARRANDE, 1887 & GUTIÉRREZ-MARCO, ¡990; PARSLIIY & PRoKou, 1999 NUMBER 01 SiECIES: 2 INcIIsoF.D SPECIES: Balanocyslííes lagenala> BARRANDE, [AnatiferocystisCHAI5vETL, l94¡ 1 887: B. primas (BARRANDE, [872) =Anaofrpsís BARRANDF, 1872 -‘- S,íaxrioísxuHiexL otiCL’RRENCE: Middle Ordovician (Lían- REFERUNCES: CHAUVEL, ¡941; UBrserís, 1979: DOMÍNGUEz & viro) GL’TIÉRRFY-MARCo, 1991) GeocRAuuícAL Di5TRiBItTION: Europe (Beisemia) RUrERENCES: BARRANDE. [887: Jocs’crsírs, [986; PARSLE’I & [KirkocvstísBA5SLLR, 1951)] PRoKou, 1999 =Ana/ífrpsis BARRANDE, [872 TYFE SuadEs: Enoploaro? papilla/a BASsI,aR 1943 Barrandeocarpus UBAGOS, 1979 REpERENCES: BÁsSs,tiu, 1950 TYFET SPECIEN: Barrandeocarpas jaekoli UBAGHS, l979 NUMBER OF SPECIES: 2 Ánomalocystites HALL, 1859 [NCLIsDI:DSirCIES: Barrandeocarpus jaekeli* UBAOHS [979; [=BasslesocystisCASTre. [952(junior synonym)] B. oon’egisa.s CRA5KE & JISFrERIES 1989 TYPE SPEcIES Aoo,nalocz’stites dispar/lis HALL, [859 STRATiGRxrHiCáL OCCURRENCE: Upper Ordevician (Caradee- NLMIsER OF SPiSCiIi5: 2 Ashgill) INCIsIDIzD SFECIES: Anovoalocvstiíes corros/as HAll., 1859; A. GEoGRxuIIIc¡sL DISTRIBUTION: Europe (Bohemia) dispar/lis> HALL, [859 REFERENCES: tJnxc,u,s, l979; CRASRE & JEPIERiES, [989; SNí=xTioÑxrriíext.OCCURRENCE: Lower Devonían RUTA, 1997d GETXíRAPHiCAi, DISTRIBItTION: Nortis America (USA) REFETRENCES (‘ASTER, [952, Kos NrA & Jor trE. [982 rBasslerosvs/is CASTre, 19521 =Anomalor-s’s//tes; HALL 1859 —~ Aspidocarpus UBAGUS, 1979 REFERENtES: CASTER, 1952; KOLATA & JOLLIE, [982 TYFE SPEIiiES: Aspidocarpas ho/reInicas UOAGOS, l979 NUMBER OF SutiCIr¡S: 2 Bokkeveldia RUTA & TFIERON, 1997 tmjr’ ITrST¡rscI,r:c-TLC A ,,,J,,-,,.,,, kt.,,.,i,,,9< Itn,.-’ns, iO’7O LXITlEflá5JN,1fl5 A. riadaneuísis (Crrrss¡vrr., 1937) NFsMBER OF SPECIES: 1 STRATIGRAPIIICAL OCCURRFNCI:: Upper Ordovician (Caradoc) INCIUDEI) SPLCiES: Boklcevcldia oos/haízení> RísTss & THE- Gaocuxuííwzxi, DISTRIRI:r ION: Europe (Bohemia) RON, [997 RErFRETNCFS, UssÁt3tIs, 1979 SrssÁTiGRAPIiiCxL OCCLtRRENCE: Lower Devonian (Emsian) GrOoRAs’r-sií:Ai DISTRIBurTION: Soutisom Africa (Republie of Ateleocystites BILLINGS, 1858 SoutS Africa) zAtelcocvsíis LINDSTROM, 1888 (noin. van.)] REFERENCES: RUTA & THERON, [997 [ Coloquios e/e Paleontología 42 2002, 53: 33-68 Patricio DomínguezAlonso & el al. Aa annorated checí-lisí of genera and speeies of carpoids GEOGRAPHíCAL DISTRIBUTION: Nortiseos Africa (Morocco) Guichenocarpos CHAUVEL, 1981 REFERENCES: Cruí’ps, 1990; LEFEvURE, 2000b TirE SPECiES: Guiehenocarpos nioni CHAtO/EL, 1981 NUMBER OF SPECIES: 1 Chinianocarpos UBAGES, 1961 INCLUISED SPEciEs: Gule/tenocarpos nioni* CHAUVEL, 1981 Tvpn SPECIES: C/tiniatrocarpos lsíroroli iJnxcus, 1961 STRATIGRAPInCAL (XncURRENCE: Middle Ordovician (LI andeilo) NUMBER OF SuEciEs: 1 GEoORAPHICsa. DISTRiBUTiON: Europe (France) INCLUDED SPECIES: Chinianocarpos t/íorali> UHAGHS, 1961 REFERENCES: CHAUVEL, [982 STRATICRAPEICAL OCCURRENCE: Lewer Ordovician (Trema- doc) Jaekelocarpus KOLATA, FREST & MAPaS, 1991 GEOORAPH¡CAL DrsrRiBUTioN: Europe (France) TYPE sPECiES: Jaekelocarpas okla/romensis Kou.xrx, FREST & REFERENCES: UBAGHS, 1961, 1967c, 1969; JEFrERIES, 1986 MAPES, 1991 NUMBER OF SPEciES~ 1 Dalejocysús PROKOP, 1963 INCLUDEIS SuElES: Jaekelocarpus okla/tomensis* KDLATA, TvPE SPECIES: Dalejocystis casteri PaoKou, 1963 FREsT & MAPaS, 1991 NuMMER OP SPECi~S: 1 STLATIGRAPBICAL OCCURRENCE: Upper Carbonifereus (Mes- INCLUDED SPEiES: Dalejocystis coste rl PROKOP, 1963 covian) STRATiGRAPHIcxL OCCURRENCE: Middle Devonian (Eifelian) GEOGRAPHíCAL DISTRIBUTION: North America (USA: Oklaho- GEOGR.APHICAL DiSTRiBUTiON: Europe (Bohemia) ma) REFERENCES: PRoKoP, 1963 REFERENCES: KOLATA el aL, 1991; DOMíNGUEZ el al,, 2002 Diwnphñliocystis KOLATA & GUENSBURG, 1979 Kierocystis PARSLEY, 1991 TYPE SPECiES: Diamphidiocyslis drepanon KOLATA & GUENS- TYPE sPECíEs: Kieroeystis inserta PARSLEY, 1991 EURO, [979 NUMBER OF 5PECiES~ 1 NIR,IBER OF sPEcrES: 1 INCLUDED SuECIaS: Kierocystis inserta* PARSLEY, 1991 INCLUDED SuEciaS: Diamphidiosyslis drepanon* KOLATA & STRATiGRAPHICAL OCCURRENCE: Upper Ordovician (Caradoc) GUENSEURO, l979 GEOGRAPHICAL Di5TRIBISTiON: North America (USA: Ken- STRATíGRAPHíCAL OCcURRENCE: Upper Ordovician (Ashgill) tucky) GEOGRAPHICAL DiSTRIBOrION: Nortis America (USA, Franco) REFERENCieS: PARSLEY, 1991; RUTA, 1999a REFERENCES: KOLATA & GUENSBURO, [979; CHAUN’EL, 1982; LEFEvBRE, 2000a Kopflcystis PARSLEY, 1991 TypE SPECIES: Kopfieystis kir4fieldi PÁIIsLEY,1991 Enoplonra WETHERBY, 1879 NuMBER OF si’~Cia.s: 1 Tira SPECiES: Anomalocyslites (Ateleocystiees?) balanoides INCLUDED SuEllEs: Kopficystis kirkfieldi* PARSLEV, 1991 MEEK, 1872 SNATIGRAPFECxí. OCCURRENCE: Upper Ordovician (Caradoc) NUMBER OP sPEcrES: 5? GE(XSRAPHICAL DíSTRIBUTION: Nortia Aznerica (Canada: Ontario) INcLUDED SI’ECiES: Enoploara balanoides WETHERBY, [879 REFERENCES: PARSLEY, 1991; RUTA, 1999a [MEEK,EB. 18721; Enoploura craslacea (Hxrcia, 1896); Enoploara meeki CASTER, 1952; Enoploura popei CASTER, Liigynocystis JAEKEL, 1918 1952; Enoploura wet/terbyi CASTER, 1952 TYPE SuECIaS: Anomalocystites pyramidalis BARPANDE, 1887 Nora: TSe genus Enoploara needs revision. Mosí of tSe des- NUMBER OF 5PEc1ES~ 1 t (BARRANDE, INCLUDED SuECIaS: Lagynocystis pyramidalis cribed species musí be restricted te ihe types because lSey l887) lack diagnostic characters. STRATiGRAPHICAL oCCURRENcE: Middle Ordovician (tlanvirn) Srí 43 Coloquios de Paleontología 2002,53:33-68 Patricio Doosíoguez Alonso & es al. An annosased check-iiss of genera and species of carpoids INCi,511)FO SUEllES: Mivrocssíclla barrandel JAEKEL, 19<)!; 41. INCL5JDED SPETCiETS: Ovocarpas? circuías-ls UBAcrís, ¡994; 0. inc/picor> (BÁRRxNros, [887);41. incip/ens osiloni (BARRAN- ,nons:ereíi> UBAGOS, 1994 DC, ¡887; CírAu:vEr., ¡941) STRAOGRAPIOCAL OCCURRENCE: Lower Ordovician (Arenig) STRATIGRAPH5CAL OCCiSRRFNCF.: Middle Ordosician (Lían- GLoGRAPIIICAI. DISTRIBUTiON: Europe (Franco) vira-Llandoilo) RIFERENCES: UBAGHS. [994 GETOt;RAulrrUAs. DISTRiaUrioN: Europe (BoSemia, France, Spain> Paranacystis CASTER, 1954 RF.Fr:RISNI:rs: BANRANDE. [887; JArRo,, [91)1; UEesc;us, Tyi’r-~ sueelEs: Parouracvsris pctrii CASTOS, 1954 1967c: Jszrrsieies, [967, [968. [986: PAR5I.EY. [994 NL:MBETR OF SFECIFS: 2 lNUIi~DFt) SUrCiES: Paranacyslis petrii> CASrER, 1954; E Mitrocystites BARRANDF~, ¡887 .s,moneae RIsTA, 1997 Tvssz Si’tiUiLS: Mi/rotrw/res mi/ra BARRANOr. 1887 STRPrsc;RAuuIcs.s. 0t?CURRENCE: Lower Devonian NI’Mnízsz 0v Sus7Cies: 1 (and one doubtfolly altributed te ISis GFOGII.áíticAL DrsTRinUTII)N: SoutS America (Brazil: Paraná) scíeous) R5?FFRPNInETS: CASrER, ¡954b: UnAUrIS, 1967c 1 NCtiT>EIs Sl’ECiETS: Mitrocvslites mitro> Ii ARR ANDE, 1887: «,14/tsosí’stiie.s» stvlo/detss DEHM, ¡934 Peltocystis THORAL. 1935 Srvsxní;RApizicxi, OCCiIRRiiNI?F: Middle Ordovician (Lían- TirE SImUlES: Pcltocvst/s cornuto IÁioRÁL, 935 “ini) Nt;xioszu OF SUECiES: 1 9< THORAL, 1935 GEoGIIxiHSUÁL DISTRIBITION: Europe (BoSeíoia) INCJ1;OUD SETEUrES: Peiiocvstis coníala SrRArsssÑAr’rrICAv, OCcí¡RRENcET: Lower Ordovician (Troma- Reí:rRíz:Ncrs: BARRANDE, [887: iNFREL, [918; UBAGIIS, doc) 1967c; ]ETI:FETRIvSS, ¡967, [968, 1986; PNRSI.ssx, [994 GBOGRAPHíCAL DIsTRrBI:TIoN: Europe (France) Mirroívsli.s BÁíRáíiiE, ¡887; CrssssvFs, 1941. incorrecí spe- RFYLRLNCES: TíloRAr. ¡ 935b: UBAc;íís, [969:JlzrrssRsrzs, [986 lIingj Placooystella RENNIE. 1936 =¡Viitroessritcs BARRANDE, [887 TYPET SuEllES: Placocvsí/s africana RaED, 1925 RPFERRNUF:S: BARRANDE, ¡887: ChrAUva. 1941 NUMBER OF SueCias: 1 INCLIsOFO SusICIES: Placoc-vsrcllo africana> (REED, [925) Mongolocarpos RozHNov, 199<) SraxíiGRxt’oIt:Ai, OCCURRENCE: Lower Devonian (Ernsiaim) TYPET SPECIb:s. Moogolocaipos minzhiíri RozFrleov, 1990 Niza nr-a or SFEUiLS: GEOGRÁPFiICÁL DISsRiBs)TJON: SoutSern Africa (Ropublic of Soutb Africa) INULL’DLD SFEUIFS: Mongolocas-pos oíioz/rmni> Rozusov, 1990 REFERENCES RENNIE, ¡936 STÑATIGRÁFII cAí 5 sCsiIJRRENCF: Si jurian (Ludlow) (ieocssÁuríícÁí. 0sSrRiuuTioN: Asia (Mongolia) Rtzr’ekLrsccs: Rtszrassov, 1990 IP/otee ‘,stss O arr sri 1 SOGí 1 noris van par/ros Pías ocx sri/es oc Korssst:ie, 1869 Notocarpos I’uríuíp, 1981 partan Pías oc ;wcila RENNiE, 1936 Tvus: SPFictcS: A’orccaspos gas-raId Ponto. [981 Placocysfltes DE KoNrNcí<, Nisswra ot~ SItúES: 2 1869 INCI,U 1)5715 St’LCSES: Noeoceu-pcs garratti> Pr isíru, ¡981; A’. ~>ra— [=PiacocystisHAECREL, 1896 (floro, van.)] TYPE SUElleS: Ploctwvsv/tes forbesianus DE KoNswa, 1869 oor (O tAiMAN, [91)4> SraÁTrcmAurrícAr. CsCCLJi 44 Coloquios dv Palessssssslogíer 21)02, 53: 33-68 Patricio Domínguez Alonso & el al, Aa annotated cbeck-iisr of genera auíd species of carpoids GEOGRAPHICAL DISTRIBUTION: Australia INCLUDED SPECIES: Vizcainocarpus dentiger> RUTA, 1997; 1K REFERENCES: RUTA & JELL, 1999b rutai LEFEBvRE, 2000 STRATIGRAPHICAL OCCURRENCE: Lower Ordovician (Trema- Pseudovictoriacystis RUTA & JELL, 1999 doc-Arenig) TYPE 51’ECIES: Pseadovictoriacvstis problematica RLrrA & GEoGRAr’HicmL DISTRiBLrI1oN: Europe (France, England) JELL, [999 REFERENCES: RUTA, 19975; LEFESBRE 20005 NUMBER OF suECIas: 1 INCLUDED SPECIES: Pseadovictoriacvslis problematica* RUTA Wilinzanocystis KOLATA & JOLLIE, 1982 &JELL, 1999 TYPE SPECIES: Willunanocystis denticulatas KOLATX & JOLUIE, SrvsricRAuuicxL OCVURPENCE: lxwer Devonian (Ixchkeviarí) 1982 GEOGRAPHICAL DISTRIBUTION: Australia (Victoria) NUMBER OP SPECiES~ 1 REFERENCES: RUTA & JELL, 1999d INCLUDED SuEcias: Willtna,rocystis denliculala* KOLATA & JOLLiE, 1982 Rhenocystis DEHM, 1932 STRAooRxumcxt. OCCURRENCE: lJpper Ordovician (Caradec) TYPE SPECIES: Rhenocystis latipedanculala DEaIM, 1932 GEOORAPHICAL DISTRIBUTION: Nortis America (USA: Wiscon- NIJMBER OP SPECIES~ 1 sin) INCLUDED SPECIES: Rlrenocystis iatipeduncr¿lata* DEKM, 1932 REFERENCES: KOLATA & JoLLIE, 1982 STRATIGRAI’HICAL OCCURPENCE: Lewer Devonian (Emsian) GEoGRxPHIcAL DiSTRiBIYIioN: Europe (Germany) Yachalicystis HAIJDE, 1995 Teue SPECIES: Yac/talicys/is triangularis HAUDE, 1995 REFERENCIeS: DEHM, [932; RuITA & BARrEES, 1998; SUTcLIr’- NUMBER OF sunC~S: 1 FE et al., 2000 INCLUDED SPECIES: Yaclaalicystis triangularis* HAUDE, 1995 STRATIGP.APHíCAL OCCURPENcE: Lower Devonian (Emsian) [Sagittacystis PARSLEY, PROKOP & DERSTLER, 20001 GEOC,RAPH1CAL DISTRIBErHON: South America (Argentina: San =Balanocystites BARRANDE, 1887 4 Juan) REFERENCES: PARSLEY, PROKOP, & DERSTLER, 2000 REFERENCES: HAUDE, 1995; LEFEvERE, 2000a Spern¡acystis UBAGHS, 1967 [=AnomocystisJAEKEL, 1918] II. íNDEX OF SPECIES T’íPc SPECiES: Anomalocys/i¡es ens~fer BARRANDE, 1887 NUMBER OF SuElES: 1 Cincta JAEKEL, 1918 INcLUDED SPECES: Sperníacystis ensífer* (BARRAr-iDE, 1887) STRsTiGRAr’urCsL OCCURRENCE: Middle Ordovician Asturicystisjackeli SDZUY, 1993 GEOGRAPUICAL DISTRIBUTION: Europe (Bohemia) STRATIGRAPHICAL OCCURRENCE: Middle Cambrian (St. REFERENCES: BARRANDE, 1887; UBAGLIS, 1967c Daviris) GEOIIiRxrKicxL oiSTRiBUTiON: Europe (Spain) Tasnuanicytidium CASTER, 1983 REI’ERENCES: SDZUY, 1993 TirE SPECIES: Tasmanicytidiam barre/ti CAsTER, [983 NUMBER OF SuPCIES: 1 Asturicystis havliceki FArRA & KORDULE, 2001 INCLUDED SPECIES: Tasmanicytidiam barre/ti> CASTER, 1983 STRATIORAPIIICAL OCCURRENCE: Middle Cambrian (St. STRATIORAPHICAL OCCURRENCE: Lower Silurian (Llandovery) David’s) GEOGRAPUiCAL DISTRIBUTiON: Australia (Tasmania) GEOCRAI’KICAL DISTIuBUTiON: Europe (Bohemia) REFERENCES: CASTER, 1983 REFERENCES: FATRA & KORDULE, 2001 Victoriacystis GILL & CASTER, 1960 [Cephalocinctuslabiatus TERMIER & TERMIER 1973] Tirc suEcíEs: Victoriacystis willcinsi GILL & CASTER, 1960 =Efliptocinctas barrandei (MUNIER-CHALMAS & BEROERON, NUMBER OP SPECiES~ 2 1889 INCLUDED SPECiES: Viclorracystzs /roltnensoran, RUTA & JELL, REFERENCES: TERMIER & TERMIER, 1973; FRIEDRICH, 1993 1999; 1? wilkins? GILL & CASTER, 1960 STRATIC,RAuHíCAL OCCURRENCE: Upper Silurian (Ludlow)- [Cyclocinctussabeircalaris TErwraíí & TERMIER [973] Lower Devonian (Lochkovian) =Ellip/ocinctas barrandei (MUNIER-CHALMAS & BERGERON, GEOGRAPHICAL DiSTRIBUTION: Australia (Victoria) 1889) REFERENCES: GILL & CASTER, 1960 REFERENCES: TERMIER & TERMíER, 1973; FRiuosICH, [993 Vizcainocarpus Rris’rA, 1997 LCyclocinctas nraroecanus TERMIER & TEIO.4íER, 1973] TirE SPECiES: Vizeainocarpas den/iger RUTA, 1997 =Sueocystis? maroccana (TERssiER & TERMIER, 1973) —> NUMBER OF SuElES: 2 REFERENCES: TERMIER & TERMIER, 1973; FRIEDRICH, 1993 45 Coloquios de Paleontola gla 2002,53:33-68 Patricio [)ombigssezAlonso & el al. An anírorared clsecle-lisí of gesrera and species of carpoids Davidocinetus pe,nbrokensis FRtEDRICH, 1993 NOTE: Formally Decacvslis hispanica is restricted lo tSe STRATrGRxpírscAí. c>ccLSRISETNcE: Middle Cambrian (St. isolotype. FaíEDarcu 1993 considered it as a probable senior David‘s) synorxym of Gyrocysíis platessa JAEREL, 1918. However, alt- GFOGRAPHICAI, DISTRIBUTION: Europe (UK: Wa¡es) Songis tSe altribulion lo Gyrocystis is obvious, liso atíribution RFFERFNCES FRrrrorcH, ¡993 lo Gvrocv.stis pla/essa JAENEI., l918 is deubíful. (Seo note en (iyroty.sl/.s plates.sa JÁFREI,, 1918) [Oec:acvstis/r/spanic:a GísLÉs, 1927: dubious species] STRAT5GRAPHICAI. OCCISRRENu7E: Middle Cambrian (St. = Gs¿ro e vs//.s /rispon/ca (GalÉs. [927);dubieus spocies David’s) —‘ Gvrocvs/i.s piascsso JAEKEL. [918--> GacscRxrnuiC=wDiSTRIBusTIC)N: Europe (Spain: Castilla-Loán) STRÁTsGRÁPIIIcÁr OCCLiRRENCET: Midd[e Cambrian (St. REPETRENCES: JAEKEI., 1918; GIsLÉN, 1927; FRIEDRIcH, 1993; David s) Gil. Cii) & DoMÍNcsusEz, [994 GrsOcaÁuírIcÁL DiSTRIISUTIoN: Europe (Spain) REFERENCES: JxEKETI., 1918; GísrÉN, 1927: FarEaRícíl, 1993; Gyrocyslisplatessa JAEKEL, 1918 GIL Cío & DosrNcustzz, 1994 [?=Decacystishispan/ca GIsLÉN 1927; dubious species] Decacvs/is praeellip/icas TERMIER & TERMIER, [973] [Dec:acysí/spraeeiiiptic:us TE5zMIER & TERMíER, l 973] NOTE: In cur opinion Gyroczstis platersa JAEREL, l91 8, as = Gyrocx’stis plares.sa JAEREL. [918 revised by FRIEDRIcH, 1993, is a complex of fossils, [listen- REFERENCI;S: FI>IEoIIIcH, [993 cally, any Ciocta witS ten marginal platos was described oit- Seras Gyrocysíis platessa JAEREL, [918oras Decacysl/s his- Elliptocinetus barnandei (MUNIER-CHALMAS & ¡mao/ca GísÉÉrs 1927 ( >). A revision of liso material Bí¿RGr-zrzOÑ, 1889); TeaMuza & TERMIER, 1973 detemiined by FRiEDRrCH, 1993 allowed swo of tSe presení Socxrrcacxs’Híc:xs. tti?t:tJRRFNCE: Middle Cambriarí (St. autisors lo propose Gzrocystis cruzoe GIL & DoMÍNc;oEz, David s) 1995 ( O. Seo Sowevor FRIEDRiCO, 1995 GecxsrcxI’HrexL DisrRiBlsTit)N: Europe (France) Srrsxricsitxi’uicxr, OCIIURRENIIE: Middle Cambrian (St. RSFLRLNCEs: MLNíER-CHALMA5 & BeRc;eRors iii BEROFRON David’s) [889; CaRMeN & IERMIER, [973;FRíEDRICH, 1993 GizoGreAuriicAi. DISTRiBiJTION: Europe (France, Spain) REFERENCrS: JAEKETI, 1918; FaíEDaícH, 1993: GIL CID & Elliplocinetus vizcainol EniEDRicH. 1995 Dosrissjuszz, [994 STRArIGRAIHICAL OCcISRRENCE: Midd[e Cambrian (St. Davids) Gyrocystis testudiformis FRIEDRIcH, 1993 GEoCRxumcxt. DiSTRiBI;TION: NortSern Africa (Morocco) SircÁríorcái’HicXL OCCURRENCE: Middle Cambrian (St. REFETRENCE.S: FRIETiDrsICr-s, 1 995 David s) GFOGRAPHICAI. DiSTRIDUTION: Europe (Spain: Aragón) Gyrocystis badulesiensis FízíísDRieH, 1993 RBr-FR11NU11S: FRrtzDrzrCH, [993:Grí. Cro & DOMiNGUEZ, [994 SmxrscuAuríIcÁI OCCURItENCE: Midd[o Cambrian (St. David si Ludwigicinctus truncatus FR¡EDRtCH, 1993 GEOGRAPHICAL DISTRÍBUTION: Europe (Spain: Aragón) STRÁTIGRÁPHIcsí, OtXiURRENCE: Middle Cambrian (St. Reecaeseas: FareDRice, 1993; Go, Cro & DOMÍNGUW, 1994 David s) GEoGRxPHicM. DiSTRiBLTiON: Europe Coloquios de Paleouísslogia 46 21)02, 53: 33-(sX Patricio Domínguez Alonso & el al, An anriolated check-iist of genera and species of carpoids GEOGRAPHICAL DiSTRIOUTIoN: Europe (France) Trochocystites bohemicus BARRANDE, 1887 REFERENCES: FRIEDRICO, 1993 STRATIORAPHICAL OCtIURRENCE: Middle Cambrian (St. David’s) Sucocystis bretoni FRIEDRICH, [993 GEOGRAPHICAL DISTRIBUTION: Europe (Boisemia) STRATIGRAPHICAL OCCURRENCE: Middle Cambrian (St. REFETRENCES: BARRANDE, 1887; JAEREL, 1918; FRIEDIIiCH, David’s) 1993; DoMÍNGUEZ, 1999 GEOGRAPHICAL DISTRIBUTION: Europe (France) REFERENCES: FRIEDRICH, 1993 [Troc/rocysíitesIon gífossatus JxEKEL, 1918] =Troc/toq’stites bo/temicus BARRANDE, 1887 —~ Sucocystis? maroccana (TERMIER & TERMIER, 1973) REFERENcES: JAEKEL, 1918; GISLÉN, 1930; FRIEDRicFt, 1993 [=Cyelocincíusnraroccanas TERMIER & TERSIIER, [973] STRATIGRAPHICAL OCCURRENCE: Middlo Cambrian (St. [Trigonocystisovalis HAECREL, 1896] David’s) =Troe/rocystites boheinicus BARRANDE, 1887 GEOCRAuHICAL DISTRIBUTION: Nortisoro Africa (Morocco) REFERENCES: HsWCKEL, 1896; FRIEDRICH, l993 REFERENCES: TERMIER & TERMIIsR, [973; FRIEDRICO, 1993 LTrigonocystis ¡rigona HAECKEL, 1896] Sucocyslis melendezi (SCHROEDER, 1973) =Troc/tocystites bo/temicas BARRANDE, 1887 —> [=Trochocinctuspardalíhanicas TETRMIER & TERMiER, 1973] REFERENCES: I-[AEcKEL, 1896; FRIEDRICH, 1993 STRATIGRAPHICAL OCCURRENCE: Middle Cambrian (Sí. David’s) Trochocystoides parvus JAEKEL, 1918 GEOORAPHICAL DISTRIBUTION: Europe (Spain, France) STRATIGRAPHICAL OCCURRENCE: Middlo Cambrian (St. REFERENCES: SCHROEDER, 1973; FRIEDRICH, 1993 David’s) GEOGRAPHICAL i)iSrRIBUTIoN: Europe (Boisemia) Sucocystis quadricornuta FRIEDRICO, 1993 REFERENCES: JAEKEL, 1918 UBAOIIS, 1967a; FíxaEDííícu, STRATIGRAIHICAL OCCURRENCE: Middle Cambrian (St. Davids) 1993; SDZUY, ¡993 GEOGRAFHICAL DISTRIBUTION: Nortisem Africa (Morocco) Trochocystoides phanus SDZUY, REFERENCES: FRIEDRIcH, 1993 1993 STRATIGRAPHICAL OCCURRENCE: Middlo Cambrian (St. David’s) Sucocystis theronensis CABIBEL, TERMIER & TER- GEOGRAPHICAL DISTRIBUTION: Europe (Spain) MIER, 1959 REFERENCES: Sr,zrsy, 1993 STRATIGRAPHICAL OSICURRENCF: Middle Cambrian (St. David’s) GEOORAPHICAL DISTRIBUTION: Europe (France) Ctenocystoidea ROBISON & SPRINKLE, 1969 REFERENCES: CABIBEL el al., 1959; FRIEDRICH, 1993 Ctenocystis cotodon UBACHS RoBísoN, 1988 Sucocystis undata FRIEDRtCH, 1993 & STRATIC,RAPHICAL OCCURRENCE: Midd[e Cambrian STRATIORAFHICAL OCCURRENCE: Middle Cambrian (St. (Sí. David’s) David s) GEOGRAPI4ICAL DISTRIBUTION: NortS America (USA: Utah) GEOORAPHICAL DISTRIBUTION: NertSern Africa (Niorocco) REFERENCES: UBAGHS & RoBísoN, 1988; DOMíNGUEZ, 1999 REFERENCIeS: FRIEDRICR, [993 Ctenocystisjagoi JELL, BURRET BANKS, [Trochocinctaspardail/ranicas TERMIER & TERMIER, 1973] & 1985 STRATIGRAuHICAL OCCURRENCE: Middle Cambrian (St. =Sacocystis melendezi (SCHROEDER, 1973) —9< Davids) REFERENCES: SCHROEDER, 1973; TERMIER & TERMIER, 1973; FRIEDRICH, [993 GEOGRAPOICAL DISTRIBUTiON: Australia REFERENCES: JELL ci al., l985; DOMÍNGUEZ, ¡999 [Troc/tocystitesBarrandei MUNIER-CHALMAS & BERCERON, [889] Ctenocystis smithi UBAGHS, 1987 =Elliptoeinctas barrandel (Mu’NiISR-CHALMAS & BERGERON, STRATIORAFHICAL OCCURRENCE: Middle Cambrian (St. [889) David ‘s) REFERENCES: MUNIER-CHALMAS & BERCERON in BERGERON, GEOGRAPHJCAL DISTRIBUTiON: Europe (France) 1889; FRIEDRICH, 1993 REFERENCES: UBAGHS, 1987; DoMÍNGuEz, 1999 [Troc/tocystisbohevnica] Ctenocystis utahensis RoBísoN & SPR¡NKLE, 1969 =Trochocystites boheenicus BARRANDE, 1887 —> STRATIORAPHICAL OCCURRENCE: Middle Cambrian (St. REFERENCES: HAECKEL, 1896 FRrEDíuícH, 1993 David’s) 47 Coloquios de Paleontología 2002, 53: 33-68 Patricies l3sssuíngs¡ez Alonso & es al. An ansmotated check-Iiss ol genera and spoc’es ol csrrpoids GEcXRAPIIICAI, DISOSIIIUTION: Nortis Amorica (USA: Utah, Dendrocyslites barnandel (BATHER, 1913) IdaSo) [=liScndcos:ystisbaícavrdei BATIIER, 19134 nom.van.] REETI?RF.NCETS: ROsSISON & SuRíNarE, [969;UBAGH5 & ROBS- STRATIGRAPHICAL, OCCURRENCE: Middle Ordovician (Llandoi- SON, [988a;DOMÍNGUEz, [999 lo) GEOGRAPHICAL DISTRIBUTION: Europe (Boisemia) Etoctenocystis bohemica FATKA & KoRDUIri, [985 REFERETNCES: BATIIER, 1913 STrsxsrc.rvxurrrCAs. OCCIIRRENCE: Middle Cambrian (St. i)avids> Dendrocystites sedgwicki (BARRANDE, 1867); GEcscRxuI iICAL DiSTRIi)IJTION: Europe (Boisemia) L3ARRANDE, 1887 REFERENCES: FATKA & KORDr’LE, 1985; DsMÍNGS.s,z, [999 I=Cystidea sedgwicki BARRANLSE, 18671 STresTIGRxuurCxí. oCCI;RRENCE: Middle Ordovician (L[andei- Jugoszor’ua archaeocyathoides IWiK & ORaewsKt, [o) 1995 GF.oGRAPHICAs, DISTRIBUTION: Europe (Bohemia) STRATIoRAuHICAr. OCCs)RRENCE: IVliddle Cambrian (St. REFERENCES: BARRANDE, [887 D;rvids) (ieOc;rsAuHIuc DisrRirsUrioN: Europe (Poland) fDendrocysliteze V,dali TIJORAL, ¡935] REFERENCES D~rie & ORtOwsKr, ¡995; DOMÍNGUEZ, [999 —Mioe,vaecystis vidaíi (TíroRxs., l935); UBAGLIS & CASTER ¡967 -~ REFERENCES: TíroRAr.. 1935b Unxc;íis & CASTER in CASTER, SoRata JAEKEL, 1901 [967 Betemnocystites wetherbyi MtLLER & GIJRI±Y, Dendrocystoides scoticus (RAnIER, 1913); JAEKEL, 1894 1918 Srax-IIGRAPHIcAI. OCCURRENCE: 1Jpper Ordovician (Cara- I=Dendrocvsíis scotica BATHER. 19 l31 doc) STRATIoaxvniICxL OCCURRENCE: Upper Ordovician (AsSgill) GETíKRArrIIeái, DISTRrIIUTION: Nortis Amorica (USA) GEOGRAPHICAL DiSTRiBUTION: Europe (UK: Scotland) REFERENCES: Mír.siyR & GIsRIEY, 1894: PARSIEY, [972; REFERENCES: BATHETR, 1913 JAEREL, ¡9l8 KoIArss et aL, 1977 «Dendrocystites» rossicus JAEKEL 1901 Castericystis vail UBAGOS & ReisísoN, ¡985 [=Dendrocysíitesross/ca SraArIoaAioicxr OCCI¡i Dehmicystis globulus (DEHM [934); CASTER, [967 Heckenicystis kuckersiana (HECKER, 1940); Gui. & L=Dcndrocysliles gb/ralas Disus> 19341 CASTER. 1960 STsexrreRAr’urcx¿ OCCURRENCE: Lowcr Devonian (Emsian) =Dendrocvst/tes /eackersianus HECRER, 1940] GIIOGRAr’HiCsL DISTRiBOrION: Europe (Gornrany) SrRxsIGRArriícxi. OCCIJRRENCF.: Ordovician (Llandeilo) REI±acNceS:DeriM, [934; Grír & CASTER, [960; CASTCR, GEOGRAPH5CAL DISTRiisUTION: Europe (Russia) 1967: Ks’Tsí:ursa, 1975: BARTETLS er al - 1998; HAisDis, REETERENCES: HECKER, 1940; GILE & CASrER, 960; RozriNov 191>3 & IEETFERIES. [996 Coloquios de Palers,srssbo~ía 48 2002, 53: 33-68 Patricio Domínguez Alonso & es al. Arr anísmotaled check-iist of genera sud specÍes of carpoida SrRATIGRArHicxL OCCURRENCE: Lower Devonian Iowacysds sagittaria THOMAS & LADD, 1926 GEOGRAPHICAL DISTRIBUTiON: Australia (Victoria) STRATIGRAPHICAL OCCURRENCE: Upper Ordevician (Caradoc- REFERENCES: GILL & CASTER, 1960; JELL & HOLLDWAY, 1982 Asisgilí) GEOGRAPHICAL DISTRIBUTION: Nortis Ameíica (USA: Iowa) Scalenocystites strimplei KOLATA, 1973 ReraleaNCes: KoLsTsetaí., [977; PARSLETY, 1972; THOMAS & STRATIGRAPFIICAL OCCURRENCE: Upper Ordovician (Caradoc) LADi), 1926 GEoGRAPHICAL DISTRiBUTION: Nortis Aanerica (USA: Minnesota) REFERENCES: KOLATA, 1973a, 19735; KOLATA cí al?, 1975: MaennUia estonica Rozonov & JEEFERtES, 1996 Kolissrx e/al,, 1977 STRATIORAFHICAL OCCURRENCE: Upper Ordovician (Caradoc) GEOORAPHII:AL DISTRiISUTiON: Europe (Estonia) Syringocrinus panadoxicus BtLLINGS, 1859 REFERENCES: RozHNov & IEPFERíES. 1996 STRATIORAuHICAL OCCURRENCE: Upper Ordovician (Caradoc) GEOORAFHICAL DisTRmtríoN: North America (Canada: Que- Minervaecystis vidaui (THORAL, ¡935); UBAOHS & bec, Ontario) CASTER 1967 REFERENCES: B[LLINGS, 1859; PARSLEY & CASTER, 1965; Cxs- [=DendrocystilesVidaíi THORAL, l935] TER, 1967 STRArIGRAPHICAL OCCURRENCE: Lower Ordevician (Trema- doc-Areoig) Syringocninus sinchaini PARSLEY & CASTER, 1965 GEOGRAPHICAL DISTRIBUTION: Europe (France) STRATIGRAPHICAL OCCURRENCE: Midd[e-Upper? Ordovician REFERENCES: THORAL, 1935b; UBAc;r-rs & CASTER in CASTER, GeooraAuHícxL DISTRiBUTiON: NorIl, America (Quebec) 1967 REFERENCES: PARSLEY & CASTER, 1965; CASTER, 1967 Myeinocystites crossmani KOLATA, STRtMPLE & LEVORSON, 1977 Comuta JAEREL, 1901 Myeinocystiles natas STRIMF5F,, 1953 (pars); in KOLATA, 2 figs. 810] 1975: p. 15; pl. SUMRALL, SPRtNKLE & GUENS- STRATIORAPHICAL OCCURRENCE: Upper Ordovician (Caradoc) Acuticarpus delticus GEOGRAPHICAL DiSTRIISUTION: NoriS America (USA) BURO, 1997 REFERENCES: KOLATA, l97$ KOLATA et al., 1977 STRATIGRAPHICAL OCCURRENCE: Upper Cambrian (Merionelis) GEOGRAuHICAL DISTRIBUTION: North America (USA) Myeinocystites natus STRIMPI±,[953 REFEPENcES: SUMRALL eíal., 1997 STRATIcRxuuícAL OCCURRENCr’: tlppor Ordovician (Caradoc) Acuticarpus? nepublicensis SUMRALL, SPR1NKLE GEOGRXPHICAL DISTRIBUTiON: Nertis A,neiiea (USA: Oklahoma) & RFPERENCETS: STRIMPLE, 1953; PARSLEY, ¡972; KOLATA, 1975; GUENSBURG, 1997 KoLATAetO/., 1977 STRxríGRAuorCM. OCCURRENCE: Upper Cambrian (Merionetis) GEOGRAuHICAL DISTRIBUTION: Nortl, America (USA) Rutroclypeusjunori WIWERs, 1933 REPERENCES: SUMRALL e/al., ¡997 NOTE: Described in WITHERs, 1933 as a xiphosuran Dr king craS Amygdalotheca gnffei UBAGHS, 1969 STRAriGaxuviICxL OCCURRENCE: Lower Dovonian STRATIGRAPHICAL OCCURRENCE: Lewer Ordovician (Arenig) GEOGRAuHICAL DISTRIBUTiON: Australia GEOORAPHICAL DISTRIBUTION: Europe (France) REFERENCES: WITHETRS, 1933; JEIí. & HOLLOWAY, 1982 REFERENCES: UBAGHS, 1969; Cr¿íuus, 1989 Rutroclypeus victoruae GILI. & CASTER, 1960 Anupelocarpus handeyranensis LEFEBVRE & VIzCAÍ- NOTE: GiLL & CASTER, 1960 doscribed tSe firsí carpoids NO, 1999 recorded from Australia, inc[uding Vicloriacystis wilkinsi, STRATIGRAPHICAL OCCURRENCE: Lower Ordovician (Arenig) Ratroclypcas victoriae and Ratroctípeas withersi, They alio GEOGRAPHICAL DISTRIBUTION: Europe (France) proposed a new classifrcation for tSe Carpoidea invo[ving REFERENCES: LEFEBvRE & VIzCAÍNO, 1999 íwe new subclasses and íwo new superorders. STRATIGRAPHICAL OCt?LtRRENCE: Lower Devonian Anauricystis pñmaeva (THOiáL, 1935); LEFEBVRE & Geograpisical distribution: Ausiralia (Victoria) VízcAiNo, 1999 REFERENCES: GILL & CASTER. 1960 JELL & HOLLOWAY, 1982 [=Coíharnocystispe/inaeva THORAL, 1935] STRATIGRAPHICAL OCCURRENCE: Lower Ordovician (Arenig) Rutroclypeus withersi Gui & CASTER, 1960 GEOGRAPHICAL DISTRIBUTION: Europe (France) NOTE: seo note for Ratroclypeas victoriae GILL & CASTER, REFERE.NCES: THORAL, [935a; UBAGHS, 1969; LEFEBVRE & 1960 VIZCAiNO, 1999 49 Coloquios de Paleontología 2002, 53: 33-68 Patricio Desnsíng’uez A Ososo & cl al, As anírosated check-liss of gesrera and species of caepoids GEOGRAPHICAL D5STRiBUIiON: Europe (Spain: Castilla-La Arauricystis occilana (UBAGHS, 994>; [±FEBVRE & Mancha) VIZCMNo, 1999 RETFs¡RETN(sS, Gsi. Crí, ce al., [996; LETFEON’RE & VizeXÍNo, 1 =Cot/suroescssíis occilano Usxuy i~, 1 994] 1999 STRMIc;wxr’srrcAt OceuRRENCE: Lowor Ordovician (Areoig) (jevsv¡RsrtliU¡sL OrSrRsBUtroN: Europe (France) Bohemiaecystis guilloal (LEI:UBVRE & VIZCAÍNO, RevewizNces: Usssí}ts, 1994: t.sIs45x11r- & Vrzcsi~o, ¡999 1999); new combinalion Serein [=Sc:oliaecvslisgai/íoai LEFEONRE & VIZCAÍNO, 1999] Archacocothurnus bifidus (UBAGLIS & ROBISON, NoTE: LEETEBVRE & VIZcAÍNo, 1999 described Ibis species SísMaÁri., SPRINaIrs & GUENSBURG, ¡997 [988); under Ssot/oeev.stis CrssTri & URACIIS, [967( 9. Seo note o VAsz hoecscoí/ss,rssus bifle/a (UIsNGirS & RoarsoN, 1 981=): liso ontry br Bo/rernioec-rsíis CASTCR, [967under liso section Su:ssaArL. St’aiNÑs.e & Gtsesssr:rac, 997 (incorrecí gonder)] «Check-list of Genera” ( 9 Coí/rsrrnoevsíis Y 01//e/a UISAGHS & Roersoa. ¡988] SrssxrrGkss’uresL ocUURRETNCET: Midd¡e Ordovician (Llandeile) SrasTss;saAurlrzÁr OUUURRI:NUiL: Middle Casnisrian St. GEoGRsursscAr. DISTRtuLsrrísN: Europe (France) Dsrvid sí RISFERETNCES: LEETEBVRa & VszCMNo, 1999 GEToovsAuitsc:sL DISTRIEI:Tr(sN: Norih America (USA) Rssr’s:tsscr’s: 1 Jnxe;srs & Rrsssrscss. [988: Suwirexs± et aL, ¡95>7 Ceratocystis pernerí JAEKEL, 1901 STreArscsssurrsí:AI. csct’u:RRE\cF: Miádlo Cambrian (Sr. 1)avids> Archaeocothurnus goshutensis SIJNIRAI.t,, SÍ’iz[Ní E IYtSRU:NCUS: .1=ti M RAIL er al, - 1 997 Ceratocysti?~ vizcainol UHAGHS. [987 Ssaxriciesrttrexú tXxtiRRETNct: Middle CamSrian (St. Beryllia miranda CRIPES & DALiIY. [994 Davids) Srrs.Níss;rs,xpssscxL t)cUs:RRs:NUF:: Middle Ordosician (Llandol— GIzoGRAvIsscxr DiSTRtBUTION: Europe (France) Retrzkrtsees: UBAGES. 1987 GvocaAs’wcsr. uSSTSSiEL’TSON: Europe (France) Rt¡rersETszcES: Ciesuus & DAr Ev [994: RL’EX, 19995: LEFEN- Chau velicystis spinosa UBAGIIS, 1969 Bise, 20(>t>a pos-lico =Chaavclis:vstis s’izcoiooi i?1.xí rtv, 1992 STíxrrGRAuHrexL OCVURReNCET: Lower Ordevician (Arenig) Babinocystis dilabida RohfiNON, 1990 Gwoe;RsrwrIcss. i)sSTRIRI:TIoN: Europe (Franco) - St R\rsGRáuI5sCAi (5CUURRETNS?E:Lower Ordovician (Arenia> RFFRRFNUFS tli)NU,HS, 1969, 1983:0MEv. 992 (¡FOCRNFIttt’Ai DíSiRiissznroN: Etíropa Rsrssia: Sainí Peters- bourgt RETI-Crss-Ncrs Ro’rssov, ¡990 Chau velicystis ubaghsi (CHAUVEL,. 1966); UBAGHS, [969 Bohemiaecystis houceki CASTER. [967 [=Coíhurnocs’stisabaghsi CírsovEs., 1966] SrRxrrGRAuHsexi. OCCURRENCE: Lower Ordovician (Arenig) [4/ro ses/ics’sli.s bossce/zi CASTER, [967;LrozvsvRE & VrzcM- NO, [999] GLOGiSsPtaseAL DiSíRsuUrroN: Norlhem Africa (Merocco) Nota: See note i o tSe ersrtry of Bohc’vnisaeevsris Cxsrers. 1 967 RtzFessisrecls:CIom’vu,. 1966a, [971;UBAGHS, 969; DALeY, ooder liso secliosr «Check-list of Generas’ ( e [992 SnrsArst¡ÑÁr’iaseAI, tsCUs:’RRJ¡NUs:: Middle Ordoviciarí (lían- s’im-t.Iarndei lo?’> Chauvelicystis vizcainoi DALEY. 1992 Gt¡o;íAuísscAí ursrierBt:Ti(sN: Europe (Bohemia) STssxrrc,RÁumíIcXt, OCcURRS>Nciz: Lower Ordovician (Areoig) Rerarzse=crs:Cssruslo UaAcrts, 1 967c: GIL Cruel al,, 1996; GE5GRAI’NíCAL OiSrRiOtflION: Eursmpe (France> LEFras-vssr: & Vr/cANo, 1999 RLSLRENCLS: UBAGrsS, 983; Dss.n’, [992 Bohemiaecvstisjefferiesi GIL Cíu. DoMÍN;íw:z. SIL- [=Cothartrosvs/i.sY americano URAc,HS, ¡963] Nevae/aetyscis americana (URAGHS, [963);UBAGHS, 1967 > VAN & EscRluxNo. [996 REFERETNCET5: UBActís, 1963, 1967c; Jvr’r’EssrrzS, 1986; Jeara- [=Seociaesysri,s¡etjériesi (Gr r Ciu, DtsMiNc;UEZ., SsLvAS & RíES e> al., 1987 Esciss}ssrscs. 1996>: LEeizrsvko & VizcAÍNo. [999] Nontz: See note rs tSe entra of Bohemiaecsstis CASTER, [967 sínsier tSe section ss(’Seek-list of Cerrera” [Cotlrus-noc:zs/isYbrflda Urn’sc,us & Ronssors, [988] tpper Ordevician (Caradoc) —Arc/raeocothus-,ras bífidas (UBAGOS & ReniSoN, 1988); STPsrse;Rsrr 55UAL O(cs’RRRNCs:: L SUMRALL, SPRINKLE & GísENsnrsRc, [997 9< STRATIC,RAPHICAL OCCURRENCE: Middle Ordovician (Llaodei- REFERENCES: UBAGHS & RotrísoN, 1988; SUMRALL el al., lo) 1997 GEOGRAuHICAL DISTRIBUrION: Eurepe (France) REFERENCES: CHAUVEL & NION, 1977; CRIPPS & DALEY, Cothurnocystis courtessolei UBAGHS, 1969 1994; LEFEvBRE, 2000a STRATIGRAPHICAL OcCURRENCE: Lower Ordovician (Arenig) GEOGRAPIIICAL DISTRIBUTION: Europe (France) Drepanocarpos australis SMITH & JELL, 1995 REFERENCES: UBAOHS, 1969 STRATIGRAPHICAL OCCURRENCE: Upper Cambrian (Merienetis) GEOURAFHICAL DISTRIBUTION: Australia [Cot/turnocvstiscurvata BATIIER, [9131 REFERENCES: SMITH & JELL, 1998 =Scoíiaecystis cas-vala (BATHER, [913); CASTER & UBAOHS, 1967 ‘ Fkzbellicystis rushtoni MARTÍ MUS, 2002 REFERENCES: BATHER, [913; CASrER & UBAGHS in UBACTHS, STRATIGRAPHICAL OCCURRENCE: Lower Ordovician (Trema- [969 doc) GEOGRAPHICAL DISTRIBUTION: Europe (England) Cothurnocystis elizae BArBER, 1913 REFERENCES: MARTÍ Mos, 2002 SrRArícRssruícAL OCCURRENCE: Upper Ordovician (Asisgilí) GeocRAutucAL DiSTRii)LtTIoN: Europe (UK: Scoiíand) Gafliaecystis lignieresi UBAGES, 1969 REFERENCES: BATHER, 1913, [926: UDAGOS, l963; JEEFERIES, STRATIGRAPHICAL OCCURRENCE: Lower Ordovician (Are- ¡986; JEEFERIES etal., 1987; WooDs & JEFEERIES, 1992 nig) GEOGRAPHICAL i)ISTRIOUTIoN: Europe (France) Cothurnocyslis feilinensis tJBAC~HS, 1969 REFERENCES: Uuwris, l969; CRípus, 1989 STRATIGRAPHICAL oCCIsRRENc:E: Lower Ordovician (Arersig) GEOGRAPHíCAL DISTRIBUTION: Europe (France) Hanusia obtusa CRtPPS, 1989 REFERENIES: Uuxcus, 1969; JrzrpssierEs el al,, 1987 STRATIGRAPHICAL OCCURRENCE: Middle Ordovician (Lían- viro) LCoí/racnocystis melc/riori UISAUSHS, [983] GEOGRAI’HICAL DISTRIBUTION: Europe (Bohemia) =Ps-osí:otiaes:vslis melchiori (Uuscsrís, 1983>; UBAGHS, 1994 REFERENCES: CRípus, 1989 [=T/ros-alysy.stisurelch/ori (UBAGII>,1983)] REPERENCETS: UBAGHS, 1983, [994; CRIPPS, 1991; GIL Cro et Hanusia prilepensis CRIPPS, 1989 al., 1996 STRATIGRAPHICAL OCCURRENCE: Middle Ordevician (Llandei- lo) [Col/rurnocystis occitana UBAGHS, 994] GEOGRAPHICAL DISTRIBUTION: Europe (Bohemia) = Araaricystis occitana (Uusc;íís, l994), LEPEBVRE & Víz- REFERENCES: CRIPPS, 1989 CAtNO, 1999 ‘ REFERENCES: UBAcJHS, 1994; LEFERvRE & ViZCMNo, 1999 Hanusia sarkensis CRIPPS, 1989 STRATIGRAPHICAI. cXYCURRENCE: Middle Ordovician (Lían- (Cot/turnocystis owensi (Wooos & JEPEERIEs, 1992), LEFEBNRE & viro) VIZCAÍNO, [999] GEOORAPHICAL DISTRIBUTiON: Europe (Bohemia) =Procotlras-nocystis owensi Wooos & JEí’rEaIas, 1992 —> REFERENCES: CRIPPS, 1989 REFERENCES: WOODS & JEEFERíES, [992;LEFEBvRE & VIZCM- NO, 1999 Juliaecarpus milnerorum RUTA, [999 [Cot/rurnocyslisprirnaeva TIIoRAL, [9351 SrrexrIcRAuHICsxI, OCCItRRENCE: Upper Ordevician (AsSgill) =Arauricvslis prinaaeva (THORAS., 1935), LEPEBvRE & VIZ- GEOGRAPHICAL DISTRiRUTION: Nortisero Africa (Morocco) REFERENCES: RUTA, 1 999b CAÍNO, 999 9< REFERENCES: THORAL, [935a: UBACHS, 1969; LEFEBNRE & VIZcAÍNo, [999 Lobocarpus vizcainoi UBAGHS, 998 STRATICRAPHICAL OCCURRENCE: Upper Cambrian (Merionetis) [Cotharnocvstisabag/tsi CHAIsvEL, l 966] GEOGRAFHICAL DiSTRIIsUTION: Europe (France) =C/raavelicvstis abag/rsi (CrtsísvEL, 1966) —> REFERENCES: UBAGHS, 1998 RPFERENCES: CHAUX’EI., 1966a; CHAUNEL, 1971; DALEY, l992 Lyricocarpus courtessolei UBAGES, 1994 Domfrontia pissotensis (CHAUVEL & NIoN, 1977); &rR*TíuRAí’uícAL OCCURRENCE: Lower Ordovician (Arenig) CRtPPS & DALEY, 1994 GEOGRAI’HICAL DISTRIBUTION: Europe (France) [=Rel/calocarpos pissoíensis CHAISvEL & NioN, 1977] REFERENCES: UBAOHS, 1994 51 Coloquios de Paleontología 2002, 53: 33-68 Patr(s:io Dsss,síngucz.ilonso & es al, An annotated checic-liss of gesrera and specics of carpoids Milonicystis kerfornei CHAUVEL, [986 Procothurnocystis owensi WooDs & JEEFERIES, SisaiiorvXrHIc:ÁL OCCISRRENCE: Middle Ordovician (Llandei- [992 lo) [=Coehas-oocv.sei.sosve,asi (Wooos & JEFFERIES. 992>, GtzoorssuríreAL i)IsTRInIJTIoN: Europe (France) LuTaNRE & VIzCAíNo, ¡999] Res LRLNCES: Críss’vr¡,, 986; CRur’s & DXs.EY, 1994 SiRXsitsRXI’uic:sr, oCci;RRENcE: Lower Ordovician (Arenig) GEOGRAPH5CAL OISTRSSSLsIIoN: Europe (12K: SoutS Wales) Nanocarpus dohambii UDAGHS, 1991 RETFFRETNCETS: Wooos & Jer-r-euri-s, 1992. IFETEUNRE & VszcM- STEsslcRwítscXL OCeURReNUL: Lower Ordovician (Aronig) Nt), 1999 GEOGRAPISICAL DiSiRiELTION: Europe (France) REFCRLN Es: [BytEs. 1991 Progau¡iecystis ubaghsi CRíuPs. 1989 STÑsrroRsurIICxI. OCCURStETNUET: Lower Ordovician (Aronigí Nevadaecystis americana (UBAGHS, 1963); UBAGHS, GETOGRAPHICAL i)ISTRiOLT ION: Europe 1967 RCFeRENUES UBAUrIS, [969: CRIiI’s, 1989 [=Coe/rarnoe:s’sei.sY amecicaoo UEXc.us, [9631 STRTIGRXuuIeAs, OCCURRENíSE: Llpper Cambrian (Meno- Prokopicystis mergil CRIPES, 1989 noIS) STRATIORAPFIICAI. OcCURRENCE: Midd[e Ordovician (Llandeilo) GaosjRAi’ííseXt DiStRIBUTsttN: NoríS America (USA) GeocavXPoic:rsL i)ISTRIBIJTIoN: Europe (Bohemia) R>’raasz~cr:s: UBAGES, 1963, 1 967c; JEFFERIETs, 1986; JEFFE- Rvrvrsieces: CRípus, 1989 RíES ev al? - 1987 Proscotiaecystis me/chían (UBAGIIS, 1983); UBAGHS, Phytlocystis blayaci Tíiortxí., 1935 1994 tusc;rís, 1983] SrR’srrí;rtus]ress. OCCL’RRETNCE: Lowor Ordovician (Trema- [=Co¡óus-uocvs/ismc/chioci L doc-Arenig) =Thoralicvsíis me/ch/oh (Ursscsís, 1983)1 GEocRAFHsCsí. i)ISTRiBUISON: Europe (France) STwsTscu-NursrcAs, OCCURRENcC: Lower Ordovician (Arenigí REFCR11NUF.S: THORAL, 1935a; Grc;oí:T, [954; UnAGHS, [963, GEooRXuriiCXL i)IsTRiuonO~: Europe (France) 1969 REFETRENCES: UHAGES. 1983; LtB.NGIIs, 1994; CierrEN, 1991: GIL Cso ev al?, 1996 Phyllocystis crassimarginata (TnoRÁu.. 1935) Protocystites menevensis HícKs, [872 PhY’líotx’sC~s Ría :ssú y aricIé írassima vgioa va Tooieex 19351 SrícsTrore’rtsrc-ss. oCCs’RRrINCF: Midéle Canbriao (Si. SrrsírrorvXuvrrCXú Oc:CURRiZNUrY: Lower Grdovici an l15ernsr- Davids) doc-Arenig) GEOGRAPrI5UAL Oi5TRI}31’TION: Europe (¡1K: Wa¡os) RFFERENUES: EiCKs hí HICaS & JONES__[172 JEFFERíES el al,, GLoGRAIosiU¡sL DISTRIBUTION: Europe (Frailee) REFETRENCES: TUORAL, 193.Sa: UBAUSIIS. [969 [987 Phyllocystis? salairica DuíBxrOeovA, [985 Reticulocarpos hanusí JEEFERIES & PROI 52 2>1>1223:33-68Coloquios sil Pesle-esoeología Patricio Donr/ssguez Alonso & el al, An aísrnotated check-iist of genera mrd species of caspoids GEOGRAPHICAL i)ISTRIultzrIoN: Europa (UK: Scotland) Mitrata JAEKEL, ]901 REFERENCES: BATOER, [913; CASTER & UBAGHS in UBAGH5, [969; Jnrraieíns, 1967, 1968,1986; CRírus, 1988 Adoketocarpus acherontkus RUTA & JELL, 1999 SrrtATIC,RsxPHICAL OCCURRENCE: Upper Silurian (PIidoli) I=Scoíiaecystis griffei UBAGHS, 19691 GEoGRAPIiICAL OiSTRiBUTION: Australia (Victoria) =TIros-alicvstis gríffei (UBAGI iS, 1969); CHxusvEí., 197l REFERENCES: RUTA & JELE, 1999a REFERENCES: UBAGHS, 1969; CrsAs’vEs., 1971 Adoketocarpusjaneae RuTA & JELL, 1999 [Scotiaecysíisgailloui LEFEBNRIS & VíZCAiNo, 1999] STRsTIorráríiicsL OCCURRENCE: Lower Devonian (Locisko- =Bo/re,n/aes:vstis gailíoai (LeversvaE & VIzCAÍNO, 1999), vian) new combination Serein —> GEOGRAPHICAL DISTRIBUTION: Australia REFERENI:ES: LEFETIWRE & VIZcAÍNO, 1999 REFERENCES: RUTA & JETLL, 1999a [=Scotiaeczsrisjcfferiesi (GIL CID, DoMÍNGusiz, SILvAN & Escier- Atlanicytidium flemingí CASTER BANO, [996>;Leaenvae & Vizc:XiNo. 19991 & GILL, 1967 Bohem/aecystis jefferiesi Grí Cío. DOMÍNGUEZ, SílváN & STRATIGRAPHICAL oCCIsRRPNCE: Lower Devonian (Emsian) GEOGRAPHICAL i)iSTRIBLrDoN: New Zealand (South Island) EscRIBANo, [996 REFERENCES: CASTER & GILL in UBAGHS, 1967c; FLEMINO, REFERENCES: GIL Cii) el al., [996; LEFEi)NRE & VIZCAÍNO, 1999 l970; RUTA & JELL, 1999a [=T/ros-alicvs/isboacelci (CASrER, 1967), LEFEBvRE & VIZCAiNo, Anatzfopsis acuta BARRANDE, ]872 1999] STRATIGRAPHICAL OCCURRENCE: Upper Ordovician (Caradoc) GEOGRAPHICAL DiSTRiBUTION: Europe (Bohemia) Boherniaecss/is boaceki CASTER, l967 —> REPERENCES: CASTER la UBAISFis, 1 967c; GIL Cii) ev al., 1996; REPERENCES: BARRANDE, 1872 LEFEBNRE & VIZcAÍNo, [999 Anatzfopsis ancona DOMINGUEZ & GUTIÉRREZ-MAR- ThoTauicystis gnffei (UBAGHS, 1969); CHAUVEL, ce, 1990 97] STRATíGRAPHICAL OCCURRENCE: Middlo Ordovician (Llandei- [=Scotiaesvstis gciffei UBA;írs, 1969] [o) STRATICRAPHICAL OCCURRENCE: Lower Ordovician (Arenig) GEOGRAPHICAI. i)isTRIBI:TIoN: Europe (Spain: Castilla-la GEeGRAuííícsL i)ISTRIBL’TION: Europe (France) MancSa) REFERENCLS: UBAGHS, [969; CnXr’vEI., 197l REPERENCES: DOMÍNGUEZ & GUTIÉRRr-z-MXRCo, 1990 [Thos-alicvstismelc/rios-i (UBAcurs, [983)] Anat¡fopsis armoricana (CHAUVEL, 1941); DoMÍN- [=Col/run,ocys/ismelc/riori Une« lIs, 1983] GUEZ & GUTIÉRREZ-MARCO, 1990 =Proscoviaecvs/is ,nelchiori (UBAOHS, [983);UBAGES, 1994 [—4naíiferocystisarmoricana CHAUNEL, 1941] REFERENCES: UBAUSHS, 983, [994: CRIPPS, 1991; GiL Cío ev STRATIRAPIIicAL OCCURRENCE: Middle Ordovician (Llandeilo) al?, 1996 GEOGRAIHICAL DISTRIBUTION: Europe (France) REFERENCES: CHAUvEL, 1941; DOMÍNGUEZ & GUTIÉRREZ- Thonauicystis ubaghsi LEFEBVRE &VIZCAINO, ]999 MARCO, 1990 STRATIGRAPHICAL OCCURRENCE: Lower Ordovician (Areoig) GEOGRAiHIcAL DISTRIBIsTION: Europe (France) Anatzfopsis balclatchiensis REED, [940 REFERENCES: LEFEBNRE & VIzcAÍNo, 1999 [4 Anatífopsis barrandei (CHAUNEL, 1941); JEFFERIES & DALEY, 1996] Thonauicystis zagonaensis CIJALIvEL, 1971 STRATIGRAPHICAL OCCURRENCE: Upper Ordovician (Asisgilí) STrvsviískxvHiCxL OCCttRRENCE: Lower Ordovician (Arenig) GEoGRxuHícxL DISTRiBUTION: Europe (UK: Scotland) GEtx;Rxuaic:s¿ DisTIIiBUTioN: Norrisero Africa (Merocce) REFERENCES: REED, 1940; JEr-PERIES & DALEY, 1996 REETERENCCS: CHAUvEL, 1971 Analifopsis barnandel (CEAUVEL, 1941); DoMÍN- Tnigonocarpus singuhanis UHAGHS, 1994 GUEZ & GUTIÉRREZ-MARCO, 1990 STRATIGRAuHICAI, OCCISRRENCE: Lower Ordovician (Aronig) [=Analiferocystisbarrandei CHAIsvEL, 19413 GEOGRAuHICÁL i)IsTRIBIsTIc)N: Europe (France) STRATIGRAPHICAL tXCURRENCE: Middle Ordovician (Lían- REFERENCES: UBAGHS, 1994 viro) GEOCRAIHICAL DISTRIBUTION: Europe (Bohemia) REFERENCES: CHAUVEL, 1941; DOMÍNGUEZ & GUTIÉRREZ- MARCO, 1990 53 Coloquios de Paleontología 2002, 53: 33-68 Patricio Domí,sgucz Alonso & el al, An annotated check-Iist of genera aud Anatifopsis bohemica BARRANDE, 1872 GEOGRAPHICAL DiSTRIBUTION: Europe (Franco) STsvíxrrowxruieXL OC> URI*ETNCET. Upper Ordovician (Caradoc- REFERENCES: CHAUvETL, 1941; DOMÍNGUeZ & GUTIÉRReZ- AslsrgiilY) MARCO, ¡990 GeoGrvXr¡rrCsL 0151 Ritís 5 l0N~ Europe (Bobomia) Rí ETERIN> es: BXRRANDE. 1872 Anatífopsis longa BARRANDE, ]872 STRATIGRAFHICAL OCCURRtiNCL: Upper Ordovician (Cara- Anatifopsis cocaban KOBAYASHI, [960 doc) STRXrss;RXartseXs OCCIJRsZI:NCE: Lowcr Ordovician GEOGRAPHICAL DiSTRSBUTION: Europe (Bohemia) GETR;RAFHICAL i)sSrRiBs,’rsON: Korea REFERENCES: BARRANDET, 1872 RssFERIZNES: KorssxASHI. [960 DoMÍseistEz & GUTIÉRREz- MARCO, 1990 ,4natifopsis minuta (CHAusvEl., ¡94!); LEFEBVRE, 2000 (A ,sovifispsis cactus II ADi)INe;, ¡913] f=Asrat~/bpsis bacrandei minuto (CríXrsvar., 1941): Dor 5rÍN- =YSoíe,ísscasis7 comer (Hsouíso, 1913) GLEZ & GUTIÉRREZ-MARCO ¡990] Nora: originally described as cirripeds trinavifops/s was Ilion [=Ana/mjitrocvstisbarcandei mina/a CHAUVEL, 1941] considered crastacearr rernains) buí probably are isolalod pia- STRATIGRAPHICAL occsSRRENc?E: Middle Ordoviciarsr (Llandei- les of a polyplacopSoran irsolluse, Seo also Anaíífopsis e/em- ¡o) galas (II ‘xi)i)INII. 1 913)> >) and A.rcstifopsi.s romes- 1 Hsssoisc, Gt¿oGRAPHiCXu i)ISTRiBUTIoN: Europe (France) ¡9l3>( >) RFFFRFNCF CHsísvEs., 1 941 - [982; DoMÍNGIsEz & GIsTÉ- Ssaár1s3aXrrdse:NI. 0CCrJrss =Balttnoevst/ve.s escandti (TísoRAs., 1935): ÑIZCACNo & [Anattfopsisps-/ma BAR/ANDE. 1872] Leaersv~re, [998] [=Anatrfops/sprimas BAR/ANDE, 1872 (incorrecí gender)] STRArrcRexuí rUAr, OCCsSRRETNcET: Lower Ordovician [=Sagillaeystis prima BARRANDE, [872:PARSLEY, PROKoP, & GEOGRAPHICAI. orsrarrs~~roa: Europe (France) DERSTLER, 2000] REs’IzRIsNCsss: TuoRsí., 1935a; CNAs;ser,, 1941; JEFFERiES, =Balanoc-ystives pr/mus BARRANDE, 1887 [987; DoMÍNcrsEz & GLTIÉRRETZ-MARCO, [990; VIzCMNo & REFERENCES: BARRANDE, 1872, l887; JaFaErírEs, [986;PARS- LtzrtvBssE, [998 LEY, PROKOP, & DERSTLER, 2000 Anatifopsis escandei? thorali (CHAUVEL, ]966); Anatifopsis spinosa UBAGHS, [979 DOMÍNGUEZ & GLsríEnivez-MARdo, ¡990 STRATIORAPHICAL OCCURRENCE: Upper Ordovician (Caradee) (=Analiferescyslis esca,ídeiY 1/ros-ah CHileNa,, 194 GeoGrs,uuIexL DISTRIBOTION: Europe (Bohemia) STRXriGRAPHICse (XiCURRENCE: Lower Ordosiciao REFERENCES: 1JBAGIIS, [979;PETR, ¡984 (‘sraocsRssuorcss. tsISTISIBUTION: Norlisoro AÍrica (Morocco) Reruen=ices:CtixísveL. 941, 1966b; DOMÍNGUeZ & GuSTÉ- Ánatifopsis tnapezijiormis THORAL, [935 RRETZ-MARIKS, l990 [=A triapezi]formis THORAL, [935 o DOMíNGUez & Gutie- rrez. 1990, p. 135 (incorrecí spol[ingfl Anahfopsis guichenensis (CHAUVEL, ]941); DoMÍN- STRATIGRAPHICAL OCCURRENCE: Lower Ordovician (SUEZ & CUTIERREZ-MARCO. 1990 [=Anavifes-oevsv/sgaichenensis CHAusvEr., l 9413 GEOGRAPHICAL DiSTRiBUTiON: Europe (France) REFETRENCES: TFIoRAI., 1935a; DoMÍNCs’Pz & GIjireRRez- STrasTIcRsuurcsr. ocCIsRRFsNcF: Middle Ordovician (Llandeilo) MARCO, [990 (?csloquios de Pe¿/contsslesg/o 54 2002, 53: 33-68 Patricio Domínguez Alonso & el al, Aa asrmnotated clieck-list of gonera and species of carpoids Anatifopsis traveusotensis (CHAIJVEL, 194]); Aspidocarpus bohemicus UBAGHS, 1979 DOMÍNGUEZ & GuTIÉRREZ-MARCo, 1990 STRATIGRAPHICAL OCCURRENCE: Upper Ordovician (Caradoc) [=Anatífes-ocysíisirar’eusolensis CIIAUNEL, 1941] GEOGRAIHICAL DISTRIBUTION: Europe (Bohemia) STRAOGRAPHICAL OCCUBRENCE: Middlo Ordevician (Llanviro) REFERENCES: UBAC,HS, 1979 GEoORAFHIcAL oISTRIBUTION: Europe (France) REFERENCES: CHAUvEL, 1941; DOMíNGUEZ & GUTIÉRREZ- Aspidocarpus riadanensis (CHAUVEL, 1937); MARCO, [990 LEFEBVRE, 2000 hMitrocystites riadanensis CHAUX’EL, 1937] Ánatifopsis truncata KoBAYÁshíí, 1960 STRATIGRAPHICAL OCCURRENCE: Upper Ordovician (Caradoc) [=Anat~fopsistrancature KorsrArtuHI, 1960, incorrecí gender] GEOGRAPHICAL DiSTRIBUTION: Europe (France) STRsrIcRAuíIIcsrL OCCURRENCE: Lower Ordevician REPERENCES: CHAUVEI., 1937, l94[; LEFE5JBRE, 2000b; GEOGRAIHICAL i)iSTRiBUTiON: Asia (Korea) LEFENRRE, 2000a REFERENCES: Konsyssrii, [960; DOMÍNGUEZ & GUTIÉRREZ- MARCO, 1990 [Aspidocas-pusdiscoidalis (CREuS, 1990); LEFEBVRE 2000] =Chaar’elia discoidalis CRIPFS, [990 —~ LAnatifopsis comer (Hsi)DING, 1913)] REFERENCES: CRrí’us, 1990; LEFEvERE, 2000b =Solenocaris? vomer Hsi)i)rNo, [913; DOMíNGUEZ & GUTIÉ- RREZ-MARCO Marco, 1990 Ateleocystites huxleyi BtLLrNGS, 1858 Noru: originally described as Aoatifopsis tisese are iselated NOTE: This is the firsí carpoid described, platos of a polyplacophorao mollusc, Seo alse Anaíifopsis Srr-ATicRAPHiCAL OCcURRENCE: Middle Ordovician canas (HADDING, 1913) (—>) and Anatífopsis elongatas (Riso- GEOGRAIHICAL i)iSTRiBUTION: North America (Canada) OrNO, [913) ( Q REFERENCES: BILLINOS, 1858; CsxSTER in KOLATA & JOLLIE, STRATIGRAPHICAL OCCURRENCE: Upper Ordovician 1982 GEooíur’HIcAL DISTRIBUTION: Europe (Sweden) RutRENcrsS: HADOINO, 1913, l915; DOMÍNGUEZ & GUTIÉ- Ate/eocystites guttenbergensis KOLATA & JOLLIE, RREZ-MARCO, [990 1982 STRATIGRAPHICAL OCCURRENCE: Upper Ordovician (Caradoc) Anomalocystites cornutus (MILLER 1889; Subse- GEOGRAIHICAL i)ISTRIOUTION: North Amorica (USA: Wiscon- quení designation) HALL, 1859 sin) STRATIGRAPHICAL OCCURRENCE: Lewer Devonian (Locisko- REFERENCES: KOLATA & JOLLIE, 1982 viso) GEOGRAPH]CAL i)ISTRIRUTIoN: NorIS America (USA: New York) Austnatocystis ¡angel CASTER, 1954 [1956] REFERENCES: HALL, 1859; CASTER lo Uniscus, [967c STRATIGRAPHICA!. OCCURRENCE: Lower Devenian (Emsian) GEOORAI’HICAL DISTRIBUTION: South America (Brasil: Paraná) 4a, 1954b Anomalocystites disparilis His¡.í., 1859 REFERENCES: CASTER, 195 [=Bassles-ocysíisdisparilis (HALl, [859)] STRAOGRAPHICAL OCCURRENCE: Lower Devenian [Ralanocyslitesescandei (THORAL, 1935); VIZCMNo & LEFEBV- GEoGItAPHICAI, DISTRIBUTiON: Nortis America (USA) RE, 1998] REFERENCES: HALL, 1859; SCHI5CHERT, l904; KIRK, 1911; =Anat~fopsis escandel escanclei THORAL, 1935 CASTER, 1952 REFERENCES: THORAL, l935a; CHAUVEL, 1941; JEFEERIES, 1987; DOMíNGUEZ & GUTIÉRREZ-MARCO, 1990; VIZCAÍNO & [Anomocyslisensífer(BsRRXNi)E,1887); JAEKEL, 1918] LEEEvBRE, 1998 —Spermacys/is enstfer UBAGHS 1967 [=Anomaiocyslilesensjfer BARRANDE, 1887] Bahanocystites ¡agenula BARRANDE, [887 STRATIGRAPHICAL OCCURRENCE: Ordovician STRATIGRAI’HICAL OCCURRENCE: Middle Ordovician (Lían- GnoGRAuHrcisL DISTRIBUTION: Europe (Bohemia) viro) REFERENCES: BARRANDE, 1887 JAEKEI,, [900, 1918; CHAU- GEoCRXPHIcAL i)ISTRiOUTioN: Europe (Bohemia) vEL, 1941: UBAOHS, 1967c REFERENCES: BARRANDE, l887 CHAP/EL, 1941 [=Anonaalocysiitesincipiens BARRANDE, 1887] Balanocystitesprimnus (BARRANDE, 1872); JEEFERJES, Miíroeystella incipiens (BARRANDE, 1887); JAEREL, l901 1986 REPERENCES: BARRANDE, 1887; JAEKEL, 1901 [=AnatrfopsisprOna BARRANDE, 18721 [=Anaílfopsisprimas BARRANDE, 1872 (incorrect gender)] [Anoenalocystlles pyramidalis BARRANDE, [887] [=Sagitlacystisprima BARRANDE, 1872; PARSLEY, PROKDP, & =Lagynocysíispyramnidalis (BARRANOE, [887); JAEKEL, 19l8 —> DERSTLER, 2000] REFERENCES: BARRANDE, 1887; JAFNEL, l918 55 Coloquios de Paltonlologia 2002,53:33-68 [‘anidoDcsmíngaez Alonso & es al, Art annosared check-liss of gesrera and species of carpnids Strsxríc;Rss±tCXi.OCCIJRRFNCE: Middle Ordovician (Lían- STRATIGRAPHICAL OCCURRENCE: Upper Ordovician virn) GeooRXr’tsrcsL i)ISsRiBUTIoN: NoríS Arnerica (USA: Ohio) GazoGaoxuhIiCXL i)ISTRIBsJTIt)N: Europe (Bohemia) REFERENCES: METER, 1872: ‘,Vtr¡reísíy [879; WOoowAÑo, RETI-EstrzNCas: BARRANDE, 1872; JISFFERIBS, [986: PARSLETY, [880 PaoEctv, & Dr[RSTSrza, 201)0 Enop/oura crustacea (HAECKEL. [896>; CASTER, [Bcroleross.sti.sdisparilis (lIsta, 1859)] [952 =Arscsorolovseiees dis 1,nc/lis Hssr., 1859 [=PlacocvstisCrasloUea IteCací., 18961 RÚi’erseNCres: HA>>, 1859; SUISUCIIIIas, [904: KIRE. 1911; PErsoplossra beilemoides WI;rHIiRBy, [879(Mssr:R, 1 872) CAvFIz, [952 SrrcXTsGRsFrrIoNs. OCCI;RRENCiB: Upper Ordovician GLOGRAPHICAL DIsTRIBUTION: North Anierica (USA: lísdiasra) Ilarrandeocarpusjaekeli UBAGI rs 1979 Ri:simtzes:es: HAETUNEL, [896:CXsreí< 1952 Sstsrsí [RAu>ItUAS, oCCI!RRENCE: Upper Ordotician (Caradoc) Gessort~s’rs (‘NS It [Sta ¡ IIL’]soN: Europe (Bohonria) Enop/oura mee/ej CASíLR. 1952 Rs ir so vr s, tíissorrS, ¡979: Rr;s.x, 1997d STRÁTsGRsFiiIcAL OCCURRENCE: Upper Ordovician GeOorssrEssCxr, Disí Rs» isroN: Nssrth Anscrica (USA: Ohio) Barrandeocarpus norvegicus CíQexsau & JLFFERIIúS REETERENCES: tiAsríR. [952 [989 STÑXrronsr’rasAL 0CCURt Sud do Plarcaus Central. A neíales Sc-/coces géologiejues - 22: 1 — Upper Ordovician of Norway, asid a new’ approach Itt suhdi- 361. víding a plesion. Paloeonvologsv, 32(1): 69-99. BILuNOS, E. 1858. On rSe Cvslideae of tSe Lower Siluriarí rocks CRruus. AP. [988.Anew species of slem-groupchordalo from Ihe of Canada Figures and descriptions. Conaditín Osgasí/c uppor Ordovician of Northorn Irolanid. Palaeontologv, 31>4): Rano/os, 3: 9-74. lOS 3-1077 — [859.On tSe Crinr:iideae nf tSe loxver Siluriasí rocks of Cana- — 1989 A new genus of ssern chordalo (Cornuta) froro tSe da, Ccsnadiao Organic Remains, 4: v-~’i±7-66. Lower and Middlo Ordovician of Czechoslevakia and oSe CAESHEL, J., TeaMrrsv, H. & TERNIIER, G. 959. Les ochinoderínes sarigin nf hilatora¡ symínetrs o tSe ehordatos, Géobios, 22(2): rnesocansbrierís de la Monmagne Noiro (sud do la France). 215-2 46. Annales de Paléoí>vologie, 44: 281-294. — 1990. A new craniaso trono tSe Ordoviciarí of Morocco and CASTETR. K.E. ¡952 Concorning Eíroploara of mho Uppor Ordovi- mho search for tSe sisler group of tSe craniata, Zoo/ogicol cian asíd los relalictn lo olher carpoid Echinodersnata. Bulletios lotecnt>! ufclic L/czr,eara Sc~ lev’> [00: 27—71 t~Aníerican Paleositologv, 34(141): 5-47 + 120-127. — 1991 A cladislie ‘>nalvsrs of lSc cornules (stern chordales). — 1954a, Brazilian Devonian carpsaids. Jossssíerl of Palen,sco- Zeso/ogie’al ionores! of tIre 1 iocieon Scscict 15’, ¡02: 333—366 uxs?PS; AP.&IIALFWPL 1 1994 f o cornutos fronsihe Middle — 19545. Anew carpoid echinoderín froin Iho Parana Dovonian. Ordovician (Llanderlo) nl Nerniandy, France, and a reinterpre- 4síemis e/o Aceademia Brasileira de Ciencias, 26(1): [23-47. lation of Milonicvs vn ke4oí=té5 Palaeoíríogcaphica. Abteilung — 1967. Honroinstelea, ¡ci: Ti-entise oso lsís’escebrave Pa/tonto- A: Pcs/aeozoo/ogie St> atsgraphne 23214-6): 99-1 32. logv. (5) Echinodernasa [(2). RC. MsoRE, Ed. S58l-S623. DALCY, P.EJ. [992. he new cornutos frníst tSe Lnwor Ordovi- Geological Socioíy of America (and Univ. Kansas Press). ctan of Shrepshire asid seolheco France, Palaeontologz, New York. 35(1): 127-148. — 983. A Iseo> Silurian carpoid echinodoms trono Tasmania and — [995.Anatomy. lnccmmorion and eníogeny of dio soluto Cas- a revisic$I$ of Ihe A¡lanicytidiidao. AlcIrecinga, 7(3-41: 321- ceíicvscis cali from tSe Middle Cambrian of Utah. Céobios, 335 28(5): 585-615. CASrER, K.E. & EATON, J.T.H. ¡956. Microstrucroro of tSe platos — [996.TIme flrst solote w’hieh o ajíached as an adulí: a Mié- tSe carpoicí echinoderno Foraoac: 195>15 Jocesnol of Paleo,>— Cambrian fossi¡ from Utah with echiooderm and chordale vologo 30>3): 611-614. alfinilies. Zoologicol Josacnal of che Linneon Societs’, ¡17: CHAí~MAN, E. 191)4. New nr little known Victorian Íbssils in tSe 405-440. National Museum, Melbourne, Parí 4- Sonto Siluriasí Ostia- Deusí, R. [932 Cystoideeo aus dom rheinischon Unterdeveos. coda and Phxd[ocarida. Prcsceedicrgs cf che Royol Socias oJ Acues Johcbuch fui- Minera/ogie. Ceo/ng/e acid Paiñontolo- Victoria, 17: 298-3 19. gre, B69: 63-93. CvrAs:v¡-s, J. 1937. les Helernstélées aromoricainos, Com~aee Ren- — [934. Unlersoehungon an Cystoidoen des rheinischori Unler- do sommcsire cíe ío ,S’c,e’iévé Céolcagiqae ev Mi,récaltsgique de dex’oos, Sitzangsberiehee des- lSoverisc’heu 4k-odetnie — ¡ 941, Recherehes sur ¡os eystoides elles carpofdcs armori- DOM>NCUFZ, P. 1999. Tho oarly ovolortion nf ochinodorms TSe calos. Société (Áéologique e> Minéraiogiqcee de £celogne. 5: c[ass Clcnocystoidea revisited, lcr: Echmoodcrco Research pi-o- ¡-286. c’eee/iírgs of/he vire Europeos> Confei-eís GLrrtÉRRuz-MnCo, J.C. & MELÉNDEZ, B. 1987. Nuevos hallazgos C,xmsEvALeI & E BONASORO, Eds, págs. 263-268, AA. Bal- do Estilóforos (iIoma]ozoos) en los materiales ordovícicos de la kema Rotterdam. zona centnaibécica. Coloquios de Paleonvología, 41:41-50. DOMÍNGUEZ, P. & GoTiÉreRcz-M~aCO, J.C. 1990. Primeros repre- HADDING, A. 1913. Undre Dicellograptusskeffern i Skane jánite sentantes ibéricos del género Anaíifopsis Barrande, 1872. nagra dlirrned ekvivalenta bildoingan Lunelo Uníversitet4rss- (Homa[ozoa Stylophora; Ordovícico) y su posición sistemáti- knjt nf 2,9(15): 1-91. ca. Acta Salinanticensia, 68(1991)): 121-131. — 1915. Der mitulere Dicellograptrisschiefer aus Bomholm. DOMINGUEZ, P., JACOuSON, KG. & JEFIOERIFS, RES. 2002. Paired Lunds UniverisitctÁs-ssknfl nf 2, 1¡(4): [-39, gil) slits in afossil with a calcite skoloton, Najare, 417:841-844. HAECKEL, E. 1896. Dic Amphoridíen und fúystoideen. Beitrúge DziK & Oretowsr — 1930. Alfinitios between dio Ecliinoelerinata, Enteropneusta — [986. TIre ancestry of tIre vertelwaces. 376 págs. British mrd Chordonia. Zoologiska Bid s-agfrdn Uppsala, 12:199-304. Mrjseum, Natural History. London. 65 Coloquios de Paleontología 21102,53:33-68 Psi> nciss I3iscísísm~srics; A [ — ¡987. 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