Stratigraphic Data of the Middle – Late Permian on Russian Platform Données Stratigraphiques Sur Le Permien Moyen Et Supérieur De La Plate-Forme Russe

Geobios 36 (2003) 533–558 www.elsevier.com/locate/geobio

Stratigraphic data of the Middle – Late Permian on Russian Platform Données stratigraphiques sur le Permien moyen et supérieur de la Plate-forme russe

Vladimir P. Gorsky a, Ekaterina A. Gusseva a,†, Sylvie Crasquin-Soleau b,*, Jean Broutin c

a All-Russian Geological Research Institute (VSEGEI), Sredny pr. 74, St. Petersburg, 199106, Russia b CNRS, FRE2400, université Pierre-et-Marie-Curie, département de géologie sédimentaire, T.15–25, E.4, case 104, 75252 Paris cedex 05, France c Université Pierre-et-Marie-Curie, laboratoire de paléobotanique et paléoécologie, IFR101–CNRS, 12, rue Cuvier, 75005 Paris, France

Received 12 November 2001; accepted 2 December 2002

Abstract

This paper presents the litho– and biostratigraphic data and correlations of the Middle and Late Permian (Uﬁmian, Kazanian and Tatarian) on the Russian Platform. The lithological descriptions and the paleontological content (foraminifera, bivalves, ostracods, brachiopods, vertebrates, plants and acritarchs) of the different units are exposed from the Barents Sea up to the Caspian Sea. © 2003 E´ ditions scientiﬁques et médicales Elsevier SAS. All rights reserved.

Résumé

Cet article présente les descriptions et les corrélations litho– et biostratigraphiques du Permien moyen et supérieur (Uﬁmien, Kazanien, Tatarien) de la Plate-forme russe depuis la mer de Barents jusqu’à la mer Caspienne. Les descriptions lithologiques et le contenu paléontologique (foraminifères, bivalves, ostracodes, brachiopodes, vertébrés, plantes et acritarches) des différentes unités sont exposés. © 2003 E´ ditions scientiﬁques et médicales Elsevier SAS. All rights reserved.

Keywords: Stratigraphic data; Correlations; Middle and Late Permian; Russian Platform; Ostracods; Plants

Mots clés : Données stratigraphiques ; Corrélations ; Permien moyen et supérieur ; Plate-forme russe ; Ostracodes ; Plantes

1. Introduction terial meeting on the regional stratigraphy concerning the Middle and Late Paleozoic of the Russian Platform: the During the International Peri-Tethys Programme (1995– Permian” (Gorsky and Gusseva, 1990). This report is here 2000) and for the realisation of the paleogeographic Atlas completed by the studies realized during the Programme. (Dercourt et al., 2000), very efﬁcient collaborations were This paper presents the stratigraphy of the Middle and Late undertaken between the French and Russian scientists, and Permian. The Early Permian part, included in the Gorsky and particularly between the VSEGEI of St. Petersburg and the Gusseva report (1990), will be published later. Pierre et Marie Curie University in Paris. Several proposals The ostracod fauna is updated by the local works in South- of the Peri-Tethys Programme were focussed on the Permian ern Cis-Urals and Precaspian Depression (E. Gusseva† and S. of the Russian Platform. During the last years, both teams Crasquin-Soleau in this paper; Molostovskaya, 1997; Molos- worked together on the project of publishing in English the tovsky et al., 1997; Kukhtinov and Crasquin-Soleau, 1999; VSEGEI internal report of the “Decisions of the interminis- Crasquin-Soleau, 2003). The paleobotanical data are thor- oughly vetted by without systematic revision (i.e., synonymy * Corresponding author. is not keeping up to date).All the other fossil data are listed as E-mail address: [email protected] (S. Crasquin-Soleau). they were quoted in the original descriptions of sections. † Dr. Ekaterina A. Gusseva passed away at the end of 2000. These checklists have historical value.

It is not possible to cite here all the bibliographic refer- ginal conditions (Gaetani, 2000). The ﬁnal stages of the con- ences (all in Russian) used to draw up theVSEGEI report. We vergence between Laurussia, Kazakhstania and Angara re- refer to this report (Gorsky and Gusseva, 1990). sult in the Ural Orogenic belt. Between 20°E (Baltic Sea) and During the Middle – Late Permian time, almost all the 60°E (the Urals), the Russian Platform lies from the Barents Northern Peri-Tethyan platform is under continental or mar- Sea to the North up to the Caspian Sea to the South (Fig. 1).

Fig. 1. Location of the geographical reference points Fig. 1. Localisation des repères géographiques V.P. Gorsky et al. / Geobios 36 (2003) 533–558 535

2. Middle – Late Permian stratigraphical correlations scale (IUGS 2000) and the Russian one are summarized on Table 1. The time slice considered here covered the Ufimian, Ka- Two main domains of sedimentation are distinguished zanian and Tatarian. The precise correlations with the inter- during the Middle and Late Permian: the Eastern European national scale (IUGS 2000) are not easy. Indeed, quite all the Basin (noted A on Fig. 2) and the Western European one Russian series are under continental environments and North (noted B on Fig. 2). They are subdivided in different facies American series (for the Middle Permian: Guadalupian) and belts presented on Figure 2. These two domains are heritage the Chinese series (for the Late Permian: Lopingian) are of the anterior paleogeographic configuration. The Baltic under marine environments. The main useful tool for the Shield extended eastwards with two major positive struc- correlations between the different scales could be the paleo- tures, namely the Voronezh High and the Ukrainian High. magnetism with the reversal between the Kiaman Permo- These highs separated the Eastern European Basin (Western Triassic superchron and the Illawara mixed polarity super- Urals, Precaspian and Dniepr-Donetz basins connected to- chron. The base of the Illawara mixed zone is recognized in gether) from the Western one which is related to the Polish the Tethyan domain (Haag and Heller, 1991; Baud et al., Basin (Gaetani et al., 2000). 1996). The K/I boundary is also recognized on the Russian The correlations are presented by a series of Tables Platform in the northernmost part of western Urals, the west- (Tables 2 to 11) from North to South, belt by belt, for the ern and central part of the Syktyokar area, in Perm – Viatka Eastern European basin (Tables 2 to 10 p.p.) and after for the area and in Samara area (Gorsky and Gusseva, 1990). Gi- Western European Basin (Tables 10 p.p. and 11). The litho- alanella et al. (1997) published developed results obtained in logical descriptions (when the room was not sufficient in the Tataria (Kazan area) and presented the correlations with the Table an asterisk refers to the Table explanation) and the Tethyan domain. The limit is located at the boundary be- paleontological content (the list is always in the Table expla- tween the Urzhumian and the Severodvinian series i.e. at the nation and is refers by a number in the Table) are given for all boundary between early and late Tatarian (Table 2)asitwas the intervals. The member “aleurolite” is often used in the presented by Lozovsky and Yaroshenko (1994) for the Mos- facies description. “Aleurolite” is equivalent to siltstone in cow syncline. The correlations between the international Russian literature.

Table 1 Correlations of International, Tethyan and Russian Middle and Late Permian scales. No vertical scale. 1.After the International stratigraphic Chart (IUGS 2000). 2. After Baud et al. (1993) and Davidov (1996) Corrélations des échelles internationales, téthysienne et russe pour le Permien moyen et supérieur. Pas d’échelle verticale. 1.D’après la Charte stratigraphique internationale (IUGS 2000). 2.D’après Baud et al. (1993) et Davidov (1996) 536 V.P. Gorsky et al. / Geobios 36 (2003) 533–558

Fig. 2. Regional distribution of Middle and Late Permian deposits on the Russian Platform (on a present day geographical map, identical to Fig. 1. For clarity of the map, a lot of geographic names, present on Figure 1 are not reported here). Fig. 2. Répartition régionale des dépôts du Permien moyen et supérieur de la Plate-forme russe (sur un fond géographique actuel, identique à celui de la Fig. 1). Par souci de clarté, bon nombre de noms géographiques présents sur la Figure 1 ne sont pas reportés ici. V.P. Gorsky et al. / Geobios 36 (2003) 533–558 537

• A. Eastern European sedimentation basin. I. Arkhangelsk Belt (Arkhangelsk – Narian-Mar – Adz’Va – Pechora – Ukhta) (Tables 3 and 4); I.1. North of Arkhangelsk (lower courses of Mezen and Pinega rivers, Kouloi river); Kanin Peninsula; I.2. North East of Mezen area (Pesa river area; Beluse – Safonovo area); I.3. West of Pechora depression (Indiga – Izma river – Ukhta area); I.4. Upper course of Pechora river; south-eastern part of Pechora depression; I.5. North East of Pechora depression (Narian Mar – Adz’Va – Usinsk area); NE of Pechora syneclise; I.6. Tchernishova chain (Adak – Usinsk – Pechora area; Adz’va river). II. Syktyokar Belt (South of Pechora and Mezen syneclises and Northern part of Volga and Urals anteclises) (Table 5). II.1.a. South of Arkhangelsk area and West of Vologda area, North East of Moscow syneclise, lower course of the Dvina river; II.1.b. South East of Arkhangelsk area and East of Vologda area (Sukona river, upper courses of Pinega and Dvina rivers); South of axial part of Mezen syneclise and NE of Moscow syneclise; II.2. South of Komi Republic, North of Viatka (ex Kirov) area, upper course of Vycegda river; II.3. South East of Komi Republic, North of Perm area, West of Ural chain, South of Pechora syneclise. III. Perm belt (South East of Moscow syneclise and North of Volgaand Ural syneclises) (Table 6); III.1. Volgaarea between Nijni Novgorod (ex Gorky) and Iaroslav (between Volga and Oka rivers, lower course of Kostroma river); South West and central Moscow syneclise; III.2. East of Unja river, North of Nijni Novgorod – Tcheboksary area (lower and middle course of Unja river and Vetlonga river area), East of Moscow syneclise; III.3. Area of Viatka (upper and middle course of Viatka river and Kobra river), structural shoulder of Viatka; III.4. Oudmourtia, Perm area; Ufa – Solimansk – Glasov area; Kama river, North West of Tataria shoulder, shoulders of Perm and Bashkiria (with structural shoulder of Ufa), Birsk – Kungur depressions. IV. Kazan belt (middle part of Volga and Ural syneclises) (Table 7); IV.1. South of Nijni-Novgorod – Tcheboksary area, Chuvachia (area between Oka and Sviaga rivers); IV.2. South East of Mari Republic and East of Chuvachia, West of Tataria, Sviaga river area, West of Kazan; IV.3. Tataria (lower course of the Kama and Viatka rivers), shoulder of Tataria, North of Kuybychev Lake; IV.4. Bashkiria, Sterlitamak – Ufa – Birsk area, lower course of Belaya river. V. Samara (ex Kuybychev) Belt (South of Volga and Ural anticlise) (Table 8). Southern part of Kuybychev Lake, Samara (ex Kuybychev) – Saratov – Orenburg area, Samara river area. VI. Volgograd belt (Precaspian syneclise) (Tables 8 and 9); VI.1. Volgograd – Ural’sk – Orenburg Arc, Obchyi syrt; external part of western and northern Precaspian depression; VI.2. Volgograd – Ural’sk – Orenburg Arc, Obchyi syrt; internal part of western and northern Precaspian depression; VI.3. Volga area near Astrakhan, inner zone of southern part of Precaspian depression; VI.4. West of Volga in Astrakhan area, external zone of southern part of Precaspian depression; VI.5. Western Kazakhstan area (lower course of Ural river), central part of Precaspian depression; VI.6. West of Mugodzar Mounts, eastern part of Precaspian depression; VI.7. West of Ural chain, near Aktubynsk, NE part of eastern part of Precaspian depression. VII. Mazyr area (lower course of Pripyat river), Prypiat depression (Table 10). • B. Western European sedimentation basin. VIII. Brest area, Brest depression (eastern part) (Table 11); IX. Kaliningrad area (Polish – Lithuanian depression, southern part) (Table 11).

• A. Bassin de sédimentation Est Européen.I. Ceinture d’Arkhangelsk (Arkhangelsk – Narian-Mar – Adz’Va – Pechora – Ukhta) (Tableaux 3 et 4) ; I.1. Nord d’Arkhangelsk (cours inférieur des rivières Mezen et Pinega, rivière Kouloi), presqu’île de Kanin ; I.2. Nord Est de Mezen (valléedelaPesa;région de Beluse – Safonovo) ; I.3. Ouest de la dépression de Pechora (région de Indiga – rivière Izma – Ukhta) ; I.4. Cours supérieur de la rivière Pechora ; partie SudEstdeladépression de Pechora ; I.5.NordEstdeladépression de Pechora (région de Narian Mar – Adz’Va – Usinsk) ; I.6. Chaîne Tchernishova (région d’Adak – Usinsk – Pechora; rivière Adz’Va). II. Ceinture de Syktyokar (Sud des dépressions de Pechora et de Mezen et partie nord des antiformes de la Volga et de l’Oural) (Tableau 5) ; II.1.a.Régions au Sud d’Arkhangelsk et à l’Ouest de Vologda,Nord Est de la dépression de Moscou, cours inférieur de la Dvina ; II.1.b.Régions Sud Est d’Arkhangelsk et Est de Vologda (rivière Sukhona, cours supérieur des rivières Pinega et Dvina) ; Sud de la partie axiale de la dépression de Mezen et Nord Est de la dépression de Moscou ; II.2. Sud de la République Komi, Nord de la région de Viatka (ex Kirov), cours supérieur de la rivière Vycegda ; II.3.SudEstdelaRépublique Komi, Nord de la région de Perm, Ouest de la chaîne Oural, Sud de la dépression de Pechora ; III. Ceinture de Perm (Sud Est de la dépression de Moscou et Nord des dépressions des rivières Volga et Oural) (Tableau 6) ; III.1.Région de la Volga entre Nijni Novgorod (ex Gorky) et Iaroslav (entre les rivières Volga et Oka, cours inférieur de la Kostroma) ; Sud Ouest et partie centrale de la dépression de Moscou ; III.2. Est de la rivière Unja, Nord de la région de Nijni Novgorod – Tcheboksary (cours inférieur et moyen de la rivière Unja et régiondelarivière Vetlonga), Est de la dépression de Moscou ; III.3.Région de Viatka (rivière Koba et cours supérieur et moyen de la Viatka), épaulement structural de la Viatka ; III.4. Oudmourtie, région de Perm ; région de Ufa – Solimansk – Glasov ; rivière Kama, Nord Ouet de l’épaulement de Tatarie, épaulements de Perm et de la Bashkirie (avec épaulement structural de Ufa), dépressions de Birsk – Kungur. IV. Ceinture de Kazan (cours moyen de la Volgaetdépression de la rivière Oural) (Tableau 7) ; IV.1. Sud de la région de Nijni Novgorod – Tcheboksary, Chouvachie (région entre les rivières Oka et Sviaga) ; IV.2.SudEstdelaRépublique Mari et Est de la Chouvachie, Ouest de la Tatarie, région de la rivière Sviaga, Ouest de Kazan ; IV.3. Tatarie (cours inférieur des rivières Kama et Viatka), épaulement de Tatarie, Nord du lac Kuybychev ; IV.4. Bachkirie, région de Sterlitamak – Ufa – Birsk, cours inférieur de la Belaya. V. Ceinture de Samara (ex Kuybychev) (sud de l’antiforme des rivières Volga et Oural) (Tableau 7). Partie Sud du lac de Kuybychev, région de Samara (ex Kuybychev) – Saratov – Orenburg, région de la rivière Samara. VI. Ceinture de Volgograd (dépression Pré Caspienne) (Tableaux 8 et 9) ; VI.1. Arc de Volgograd – Ural’sk – Orenburg, Obchyi syrt ; partie externe des limites ouest et nord de la dépression Pré Caspienne ; VI.2. Arc de Volgograd – Ural’sk – Orenburg, Obchyi syrt, partie interne des limites Ouest et Nord de la dépression Pré Caspienne ; VI.3.Région de la Volga prèsd’Astrakhan, zone interne de la partie Sud de la dépression Pré Caspienne ; VI.4. Ouest de la Volga dans la région d’Astrakhan, partie externe de la région Sud de la dépression Pré Caspienne ; VI.5. Ouest du Kazakhstan (cours inférieur de la rivière Oural), partie centrale dépression Pré Caspienne ; VI.6. Ouest des Monts Mugodzar, partie Est de la dépression Pré Caspienne ; VI.7. Ouest de la chaîne de l’Oural, prèsd’Aktubynsk, partie Nord Est de l’Estdela dépression Pré Caspienne ; VII.Région de Mazyr (cours inférieur de la rivière Pripyat), dépression de Prypiat (Tableau 10). • B. Bassin de sédimentation Ouest Européen.VIII.Région de Brest, dépression de Brest (partie orientale) (Tableau 11) ; IX.Région de Kaliningrad (dépression Lithuano-Polonaise, partie sud) (Tableau 11). 538 V.P. Gorsky et al. / Geobios 36 (2003) 533Ð558

Table 2 Regional stratigraphic subdivisions of the Middle – Late Permian on the Russian Platform Subdivisions stratigraphiques régionales du Permien moyen et supérieur de la Plate-forme russe

• Foraminifera: 1. Digitina rara UCH., Cornuspira megasphaerica GERKE, Nodosaria elabugae TSCHERD., N. suchonensis K.M. MACL, Pseudono- dosaria lata K.M. MACL., Ps. pygmaeaformis K.M. MACL., Rectoglandulina borealis GERKE. 2. Saccammina rodundata K.M. MACL., Nodosaria hexagona TSCHERD., N. noinskyi TSCHERD., N. pseudoconcinna K.M. MACL., Lingulonodosaria kamensis K.M. MACL., Pseudonodosaria nodosariaeformis K.M. MACL., Ps. lata K.M. MACL., Ps. pygnaeaformis K.M. MACL., Spandelina longissima (K.M. MACL.), S. tzaregradskyi (K.M. MACL.), S. fallax (K.M. MACL.) • Bivalves: 3. Palaeomutela curiosa Zone. Palaeomutela curiosa AMAL., P. abunda NETSCH., P. parva AMAL., P. oleniana GUS., P. plana AMAL., P. keyserlingi AMAL., P. ovalis AMAL., P. inostranzevi AMAL., Oligodontella geinitzi (AMAL.), O. tetraedroides (PLOT.), O.? obliqua (AMAL.), Anthraconaia solemyaeformis (NETSCH.), Palaeanodonta parallela AMAL., P. okensis AMAL., P. ﬁscheri (AMAL.), Opokiella tschernyschevi PLOT. 4. Palaeomutela wohrmani Zone. Palaeomutela wohrmani NETSCH., P. ulemensis GUS., P. numerosa GUS., P. marposadica GUS., P. orthodorsala GUS., P. subparallela AMAL., P. verneuili AMAL., P. semilunulata AMAL., P. orthodonta AMAL., P. keyserlingi AMAL., P. ovalis AMAL., Oligodontella geinitzi (AMAL.), O. tetraedroides (PLOT.), Anthraconaia verneuili (AMAL.), A. tschuvashica GUS., A. concavocarinata (NETSCH.), A. trapezoidalis (AMAL.), A. sambulacovi KUL., Palaeanodonta parallela AMAL., P. okensis AMAL., P. ﬁscheri AMAL., Opokiella tschernyachevi PLOT., O. ignatjevi GUS. 5. Palaeomutela vjatkensis Zone. Palaeomutela vjatkensis GUS., P. extensiva GUS., P. visenda GUS., P. corpulenta GUS., P. monstruosa GUS., P. celebrata GUS., P. krotowi NETSCH., P. doratioformis GUS., P. olgae GUS., P. ulemensis GUS., P. verneuili AMAL., P. semilunulata AMAL., Prilukiella lata (NETSCH.), P. nitida GUS., P. mirabilis (GUS.), P. pugnatoria GUS., P. rukhini PLOT., P. subovata (JONES), Anthraconaia tschuvashica GUS., A. aktubensis GUS., A. castor (EICHW.), A. longissima (NETSCH.), Anthraconauta volgensis GUS., A. uslonensis GUS., A. tscherdynzevi GUS., A. iljinskiensis FED., Sinomya gracilenta GUS. 6. Nuculavus trivialis (EICHW.), Schizodus rossicus VERN., Siphogramyssia kasanensis (GEIN.), Alula kutorgi (VERN.), Streblochondria sericea (VERN.), Pseudobakewellia ceratophagaeformis NOIN., Pseudomonotis permianus MASL., Netschajewia tschernyshewi LICH., Lithophaga consobrina (EICHW.). 7. Polidevcia kasanensis (VERN.), Schizodus rossicus VERN., Pseudobakewellia ceratophagaeformis NOIN., Pseudomonotis speluncaria SCHLOTH., P. garforthensis (KING.), Netachajewia globosa (NETSCH.), Wilkingia komiensis (MASL.). 8. Palaeomutela pseudoumbonata Zone. Palaeomutela umbonata FISCH., P. pseudoumbonata GUS., P. attenuata GUS., P. olgae GUS., P. stegocephalum NETSCH., P. sintaensis GUS., P. celebrata GUS., P. visenda GUS., P. krotowi NETSCH., P. doratiformis GUS., P. quadriangularis (NETSCH.), Concinnella angulata POG., Anthraconaia castor (EICHW.), A. rhomboidea (NETSCH.), A. longissima (NETSCH.), A. kamae GUS. 9. Promytilus sicaricus (LUTK. and LOB.), Cyrtorostra laticostata NETSCH., Aviculopecten talis (LUT. and LOB.), Astartella raricostata DEMB. 10. Palaeomutela stegocephalum NETSCH., P. attenuata GUS., P. sintanensis GUS., P. ovatiformis GUS., Prilukiella alta (POG.), Concinnella concinnaeformis POG., C. angulata POG., Anthraconaia castor (EICHW.), A. rhomboidea (NETSCH.), A. longissima (NETSCH.), Anthraconauta subacuta POG., Mrassiella trapezoidalis (KROT.). • Ostracods: 11. Suchonellina trapezoidea Zone. Suchonellina trapezoidea (SHARAP.), S. parvaeformis (KASH.), S. fragilis (SHARAP.), S. fragilina (BEL.), S. ignatjevi ZEK. and JAN., S. fragiloides ZEK., S. parallela SPIZH., S.? praelongata ZEK., Suchonella cyrta ZEK., S. auriculata (SHARAP.), S. typica SPIZH., Darwinuloides svijazicus (SHARAP.), D. tataricus (POSN.). 12. Suchonella futschiki Zone. Suchonellina futschiki (KASH.), S. parallela SPIZH., S. cultella MISH., S. daedala MISH., S. digitalis MISH., Prasuchonella stelmachovi (SPISH.), P. ovalis (KOTSCH.), P. sulacensis V.P. Gorsky et al. / Geobios 36 (2003) 533Ð558 539

KOTSCH., Darwinuloides buguruslanicus KASH., Tscherdynzeviena pulchra (BEL.), T. busulucensis KASH. 13. Paleodarwinula fragiliformis Zone. Paleodarwinula fragiliformis (KASH.), P. elongata (LUN.), P. chramovi (GLEB.), P. fragilis var. angusta (KASH.), P. torensis (KOTSCH.), P. kassini (BEL.), P. faba (MISH.), P. defluxa (MISH.), P. imposter (MISH.), P. arida (MOLOST.), P.? gariainovi (MOLOST.), P.? tuba (MISH.), P. teodorovichi (BEL.), Prasuchonella nasalis (SHARAP.), P. calantarae (BEL.), Darwinuloides tscherdynzevi KASH., Permiana obloga (POSN.), Tscherdynzeviana aquamosa KOTSCH. 14. Paleodarwinula fainae Zone. Paleodarwinula alexandrinae (BEL.), Paleodarwinula fainae (BEL.), P. irinae (BEL.), P. aronovae (BEL.), P. tichonovichi (BEL.), P. inornatinaeformis (BEL.), P. belousovae (KASH.), P. jatskovae (KASH.), P. actubinensis (STAROZH.), P. cuneata (KOTSCH.), P. tuimazensis (KOTSCH.), P. pigmaea (STAROZH.), Prasuchonella onega (BEL.), P. vinocurovi BEL., P. tichvinskaja BEL., P. belebeica BEL., Darwinuloides sentjakensis (SHARAP.), D. edmistonae BEL., Vymella dobrynini (KASH.), Schneideria kazanica KOTSCH. 14bis. Amphissites tscherdynzevi Zone. Pseudoparaparchites furmidabilis SOHN, Amphissites tscherdynzevi POSN., Kirkbya ingeniosa SOHN, Fascianella notabilis KOTSCH., F. spinosa KOTSCH., Graphiodactylus uchtensis MART., Cavellina grandis SOHN, C. edmistonae HAR. and L., Healdia oblonga KOTSCH., H. postcornuta SOHN, H. pseudosimplex KOTSCH., H. subtriangula KOTSCH., Healdianella vulgaris KOTSCH., Bairdia beedei ULR. and B., B. sobolevensis KHIV., Acratia baschkirica KOTSCH., Fabalicypris crespidabilis KOTSCH., Actuaria diffusa KOTSCH., Bairdianella semilunulata KOTSCH., Moorea facilis SOHN, Monoceratina fascigiata KOTSCH. 15. Paleodarwinula angusta Zone. Paleodarwinula angusta (MAND.), P. abunda (MAND.), P. parphenovae (BEL.), P. ufimica (BEL.), P. inerta (KASH.), P. buraevoensis (PAL.), P. biriensis (PAL.), P. ovataeformis (KASH.), Prasuchonella stelmarta KASH., Sinusuella pergrafica MAND., Bairdia kamaensis GUSS., B. dobrjankaensis GUSS., B. plebeia REUSS, B. cultellata GUSS., B. cuneiformis GUSS., B. keltmanensis MAND. • Bryozoans: 16. Fistulipora dybowskii GOR., Cyclotripa multiformis GOR., Ulrichotripella oculata MOR., Stenopora thuliformis MOR., S. timanensis MOR., Tabulipora ordinata MOR., Rhombotripella superanguata MOR., Dyscritella spinigeriformis MOR., Pinegopora delicata SHISHOVA, Streblascopora fasciculata (BASSLER), Neorhombopora polita SHISHOVA, Fabifenestella shurae (MOR.), Rectifenestella microretiformis (MOR.), R. permulta (MOR.), Parafenestralia arborencens (NETSCH.), Triznella vjatkensis (NIKIF.), Wjatkella wjatkensis (NETSCH.), W. permiana MOR., Polyporella magnicava (MOR.), P. soyanensis (MOR.), Polypora baytuganenais SOSK., P. sakanensis MOR., Reteporidra soloduchoi MOR., R. quasimicropora MOR., Kingspora parvifenestrata MOR., Timanodictya nikiforovae MOR., T. borealis MOR., Girtyoporina applicata MOR. • Brachiopods: 17. Orbiculoidea conincki (GEIN.), Aulosteges fragilis (NETSCH.), A. wangenheimi (VERN.), Cancrinella cancrini (VERN.), Od- ontospirifer subcristatus (NETSCH.), Cleiothyridina pectinifera (SOW.), Beecheria angusta (NETSCH.). 18. Lingula orientalis GOL., Aulosteges horrescens (VERN.), A. gigas NETSCH., Cancrinella cancrini (VERN.), Globiella hemisphaerium (KUT.), Licharewia rugulata (KUT.), L. stuckenbergi (NETSCH.), Tumarinia latiareata (NETSCH.), Pinegathris royssiana (KEYS.), Cleiothyridina pectinifera (SOW.), Beecheria netschajewi GRIG., Licharewia grewingki (NETSCH.), Permospirifer keyserlingi (NETSCH.), Blasispirifer blasii (VERN.). • Vertebrates: 19. Chroniosuchus uralensis TVERD., Ch. paradoxus VJUSCH., Jgosuchus mirabilis (VJUSCH.), J. licharevi RIAB., Dvinosaurus primus AMAL., D. purlensis SHISH., Kotlasnia prima AMAL., Karpinskiosaurus secundus (AMAL.), Raphanodon ultimus (TCHUD. and VJUSCH.), Chthonosaurus velocidens VJUSCH., Pravoslavlevia parva (PRAVOSL.), Inostrancevia alexandri AMAL., I. uralensis TAT., I. latifrons PRAVOSL., Hexacynodon purlensis TAT., Annatherapsidus petri (AMAL.), Nanocynodon seductus TAT., Dvinia prima AMAL., Scutosaurus karpinski (AMAL.), Dicynodon trautscholdi AMAL., D. annae (AMAL.), D. amalitzkii SUSH. 20. Chroniosaurus dongusensis TVERD., Raphanodon tverdochlebovae IVACH., Microphon exiguus IVACH., Jugosuchus boreus IVACH., J. hartmanni IVACH., Bystrowiana permira VJUSCH., Dvinosaurus primus AMAL., D. egregius SHISH., Proburnetia viatkensis TAT., Niukenitia sukhonensis TAT., Sauroctonus progressus (HART.–WEIN.), Noschowhaitsia vjuschkovi TAT., Scutosaurus permianus (HART. -WEIN), S. rossicus (HART.–WEIN.), S. utilensis IVACH. and LEB., Deltavjatia vjatkensis (HART.–WEIN.), Archosaurus rossicus TAT., Dicynodon scholdi AMAL. 21. Platyops rickardi TWELV., P. stuckenbergi TRAUT., P. watsoni EFR., Melosaurus kinelensis VJUSCH., M. paucidens (KONZH.), M. uralensis MEYER, M. vetustus KONZH., Zygosaurus lucius EICHW., Chalcosaurus rossicus MEYER, Permotriturus herrei TAT., Brithopus bashkyricus EFR., B. priscus KUT., B. ponderus EFR., Syodon biarmicus KUT., Titanophoneus potens EFR., Dolionauriscus adamanteus ORL., D. janshinovi ORL., Notosvodon gusevi TCHUD., Mnemeiosaurus jubilaevi NOP., Admetophoneus kargalensis EFR., Deuterosaurus biarmicus EICH., D. gigas EFR., Rhapalodon wangenheimi FISCH., Molybdopygus arcanus TCHUD., Moschops svijagensis (RIAB.), Venjukovia prima AMAL., Phreatosaurus bazhovi EFR., Ph. menneri EFR., Phthinoeuchus discors EFR., Porosteognathus efremovi VJUSCH., Ennatosaurus tecton EFR., Gecatogomphius kavejevi VJUSCH. and TCHUD., Lanthanosuchus lukjanovae JVACH., L. watsoni EFR., Lanthaniscus efremovi JVACH., Macroleter poesicus TVERD. and JVACH., Nycteroleter inaptus EFR., N. kassini TCHUD., Discoaauricus netschajeri RIJAB., Nyctibeotus liteus TCHUD., Enosuchus breviceps KONZH., Nyctiphruretus asudens EFR., Biarmica tchudinovi JVACH., Leptoropha talanophora (TCHUD.), Mesenosaurus romeri EFR. 22. Platyops stuckenbergi TRAUT., Phthinosaurus borisiaki EFR., Nycteroleter bashkyricus EFR., Rhipaeo- saurus tricuspidens EFR., Tokosaurus perforatus TVERD. and IVACH., Belebey vegrandis IVACH., B. maxomovi IVACH. and TVERD., Gecatogomph- ius kavejevi VJUSCH. and TVERD., Brithopus priscus KUT., Archaeosydon praeventor TCHUD., Biarmosuchus tener TCHUD., Eotitanosuchus olsoni TCHUD., Ivantosaurus ensifer TCHUD., Chthomaloporus lenocinator TCHUD., Estemmenosuchus mirabilis TCHUD., B. uralensis TCHUD., Anoplonuchus tenuirostris TCHUD., Parabradysaurus udmurticus EFR., Zopherosuchus luceus TCHUD., Otsheria netzvetaevi TCHUD. 23. Phreatophasma aenigmatum EFR., Phreatosuchus qualeni EFR. • Flora: 24. Pelliothallites tataricus MEYEN, Protosphagum nervatum NEUB., Rhizinigerites neuburgae MEYEN, Fasciostomia delicata GOM., Fefilopteris pilosa GOM., Peltaspermopsis buevichiae GOM. and MEYEN, Tatarina olferievii MEYEN, T. conspiqua MEYEN, T. pinnata MEYEN and GOM, T. mira GOM., Stiphorus biceriatus MEYEN, Glossophyllum permiense MEYEN, Salpingocarpus bicornutus MEYEN, S. variabilis MEYEN, Phylladoderma aequalis MEYEN, P. anullata MEYEN, P. rastorguevii MEYEN, P. trichophora MEYEN, P. tatarica MEYEN, Doliostomia krassilovii MEYEN, Rhaphidopteris kiuntzeliae MEYEN, R. antiqua MEYEN, Permotheca sardykense ZAL., P. striatifera MEYEN and GOM., P. vesicasporoides MEYEN, ESAUL. and GOM., P. vittatinifera MEYEN and GOM., Pursongia amalitzkii ZAL., P. angustifolia ZAL., P. beloussovae (RADCZ.) GOM. and MEYEN, Cordaites clercii ZAL., Crassinervia subpentegonata KONYKOV, Samaropsis irregularis ZAL., Sashinia aristovensis MEYEN, S. borealis MEYEN, Dvinostrobus sagittalis GOM. and MEYEN, Quadrocladus dvinensis MEYEN, Q. borealis MEYEN, Q. schweitzerii MEYEN. 25. Sphenophyllum stoukenbergii SCHMALH., Phylladoderma muravievii BOGOV, Nucicarpus piniformis NEUB. 26. Signacularia noinskii ZAL., Sphenophyllum stoukenbergii SCHMALH., Paracalamites striatus (SCHMALH.) ZAL., Pecopteris helenaeana ZAL., Odontopteris rossica ZAL., Syniopteris nesteronkoi ZAL., Rhaphidopteris praecursoria MEYEN, Cordaites candalepensis (ZAL.) RADCZ., Phylladoderma meridionalis MEYEN, P. fefiloviana BOGOV, Permotheca sardykensis ZAL., Nucicarpus piniformis NEUB. 27. Paracalamites striatus (SCHMALH.) ZAL., Callipteris erosa (MORRIS) ZAL., Psygmophyllum (Syniopteris) expansum BRONGN., P. cuneifolium KUTORGA, Odontopteris rossica ZAL., Phylladoderma spinosa BOGOV, Permotheca sardykensis ZAL. 28. Intia variabilis NEUB., Viatcheslavia vorcutensis ZAL., Sylvella alata ZAL., Samaropsis vorcutana TSCHIRK. • Miospores and Acritarchs: 29. Vitreisporites pallidus Zone (J2). Calamospora hartungiana BALME, Punctatisporites sp., Apiculatisporites hispidus ANDR., Granizonospora granifera LUBER, Osmundacidites sp., Nevesisporites sp., Lycospora papullata (SMIRN.) WAR., Cirratriradites sp., 540 V.P. Gorsky et al. / Geobios 36 (2003) 533Ð558

Azonaletes sp., Ginkgocycadophytus sp., Vittatina cincinneta LUBER, V. costabilis WILS., Cladaitina sp., Vesicaspora schemeli KL., Platysaccus major (LUBER) WAR., Lueckisporites virkkiae POT. and KL., Vitreisporites pallidus (REISSINGER) NILSSON, Vitreisporites sp., Gnetaceaepollenites sp., Limitisporites ambiguus (ZAUER) WAR., Striatopodocarpites tojmensis SED., S. rarus BHARD. and SAL., Striatopiceites suchonensis SAD., Striatoabieites sp., Striatolebacheites sp., Vesicaspora ex gr. magnalis ANDREYEVA, Protohaploxypinus limpidus BALME. 30. Taxodiacites permianus

Zone (J1). Calamospora sp., Trachytriletes sp., Cyclogranisporites sp., Apiculatisporites hispidus ANDR., Cirratriradites sp., Lycospora papullata (SMIRN.) WAR., Azonomonoletes vulgaris IBR. f. minor LOOSE, Densoisporites sp., Osmundacidites sp., Raistrickia sp., Azonaletes sp., Vittatina costabilis WILS., V. elegans ZAUER, V. striata f. major LUBER, V. lata WILS., V. crassa ZAUER, Florinites sp., Cordaitina sp., Falcisporites sp., Caytoniapollenites pallidus LESCHIK, Vesicaspora sp., Taxodiacites permianus ZAUER, Striatopodocarpites tojmensis SED., S. rarus (BHARD. and

SAL.), Striatopiceites suchonensis SED., Taeniasporites sp., Hamiapollenites sp. 31. Lueckisporites virkkiae Zone (I2). Calamospora sp., Apicu- latisporites sp., Trachytriletes sp., Granulatisporites graniferus (K.-M.) WAR., Osmundacidites sp., Raistrickia cardioformis (K.-M.) WAR., Neoraist- rickia selaginelliformis SAMOIL., Leiotriletes inderjensis (MAL.) WAR., Kraeuselisporites sp., Azonaletes sp., Ginkgocycadophytus sp., Vittatina cincinnata LUBER, V. persecta ZAUER, Florinites sp., Cordaitina sp., Lueckisporites virkkiae POT. and KL., Taxodiacites sp., Vesicaspora ovata BALME and HEN., Vestigisporites sp., Falcisporites zapfei (POT. and KL.), Limitisporites sp., Striatopodocarpites tojmensis SEd., Striatopinites dvinensis SED., S. suchonensis SED., Striatohaplopinites perfectus NAUM., Striatodiplopinites lata SCHATK., Taeniasporites hexagonalis JANS. 32.

Striatopodocarpites tojmensis Zone (I1). Trachytriletes punctalatus NAUM., Granulatisporites microgranifer IBRAH., Cyclogranisporites osmundae SAMOIL., Spinosisporites spinosus WALTZ, Raistrickia cardioformis (K.-M.) WAR., R. cornuta (ANDR.) DR., Granulatisporites parviverrucosus WALTZ, Azonaletes sp., Vittatina striata LUBER, V. vittifer LUBER, V. persecta ZAUER, Cordaitina sp., Platysaccus sp., Vesicaspora schemeli KL., V. ovata BALME and HEN., Falcisporites zapfei POT. and KL., Alisporites sp., Platysaccus major LUBER, Limitisporites sp., Chordasporites sp., Striatopodocarpites tojmensis SED., Striatopinites dvinensis SED., Striatolebacheites sp., Lueckisporites sp., Taxodiacidites sp., Hamiapollenites sp.,

Vitreisporites sp., Taeniasporites hexagonalis JANS. 33. Granizonospora vulgaris Zone (H2). Calamospora sp., Spinosisporites parvispinus LUBER, Granulatisporites parviverrucosus WALTZ, Granizonospora vulgaris NAUM., Verrucosisporites sp., Cirratriradites procumbens LUBER, C. ornatus LUBER, Vittatina sp., Cordaitina sp., Vesicaspora sp., Platysaccus sp., Florinites luberae SAMOIL., Crusisaccites sp., Hamiapollenites tractiferinus

SAMOIL., Striatodiplopinites elongatus LUBER, Taeniasporites sp. 34. Cirratriradites ornatus Zone (H1). Calamospora sp., Cyclogranisporites gibberuliformis K.-M., C. polypyrenus LUBER, Granulatisporites gibbosus IBR., G. parviverrucosus WALTZ, Granizonospora vulgaris NAUM., G. granifera LUBER, Spinosisporites rectispinus LUB., Acanthotriletes osmundae SAMOIL., Cirratriradites ornatus LUBER, C. procumbens LUBER, Lycospora variabilis JANS., Remysporites psilopterus LUBER, Ginkgocycadophytus sp., Vittatina striata LUBER, V. vittifer LUBER, Florinites luberae SAMOIL., Crucisaccites ornatus SAMOIL., Alisporites sp., Limitisporites monstruosus LUBER, Striatohaplopinites latissimus LUBER, Hamiapolle- nites sp., Striatolebachiites sp.

Table 3 Eastern European basin, Arkhangelsk belt, part 1 Bassin Est-Européen, ceinture d’Arkhangelsk, première partie V.P. Gorsky et al. / Geobios 36 (2003) 533Ð558 541

35. Bivalves: Oligodontella geinitzi (AMAL.), Anthraconaia(?) verneuili AMAL.; Ostracods: Paleodarwinula chramovi (GLEB.), P. elongata (LUN.), Suchonellina futschiki (KASH.), S. inornata SPIZH., S. inornata var. macra (LUN.), S. parallela SPIZH., S. impostor (MISH.), Prasuchonella nasalis (SHAR.), P. stelmachovi (SPIZH.); Flora s.l.: Ginkgocycadophytus sp., Vittatina costabilis WILS., V. cincinnata LUBER, V. striata LUBER, Cordaitina sp., Podocarpites major (LUBER) WAR., Limitisporites monstruosus (LUBER) HART., Piceites sublevis (LUBER) HART., Vesicaspora schemeli KL., Vitreisporites sp., Striatopodocarpites nudus (LUBER) ZAUER, S. tojmensis SED., S. rarus (BHARD. and SAL.) WAR., Striatolebachiites sp. 36. Aleurolites, red marls – Aleurolites et marnes rouges: Ostracods: Paleodarwinula spizharskyi (POSNER), P. teodorovichi (BEL.), P. fragiliformis (KASH.). 37. Aleurolites, marls, dolomitised marls – Aleurolites, marnes, marnes dolomitisées; aleurolites, marnes, marnes dolomitisées. Ostracods: Paleodar- winula aff. perlonga (SHAR.), Permiana elongata (POSNER), P. oblonga (POSNER); Bivalves: Prilukiella(?) mirabilis GUS. 38. Aleurolites, sandstones, red marls – Aleurolites, grès marnes rouges: Ostracods: Amphissites tscherdynzevi POSNER, Moorea facilis SOHN, Cavellina edminstonae (HAR. and LALL.). 39. Western part: red and thin aleurolites with gypsum; eastern part: grey thin aleurolites – Partie occidentale: aleurolites rouges et peu épaisses avec du gypse; partie orientale: aleurolites grises et peu épaisses. Vertebrates: Nycteroletes inertus EFR., Nyctiphuretus asudens EFR., Mesenosaurus romeri EFR., Microphon poesicus TVERD. and IVACH., Lanthaniscus efremovi IVACH. 40. Flora s.l.: Calamospora microrugosa (IBR.) SCHOPF, WILS. and BENT., Trachytriletes sp., Granulatisporites sp., Cyclogranisporites sp., Acanthot- riletes sp., Cirratriradites ornatus (LUBER) HART., Azonomonoletes vulgaris IBR., Azonaletes microdyctius LUBER, Vittatina striata LUBER, V. cincinnata LUBER, V. costabilis WILS., V. lata WILS., V. crassa ZAUER, Alisporites ovatus JANS., Protopodocarpus alatus (LUBER) SAM., Protohaploxypinus sp., Striatopodocarpites nudus (LUBER) ZAUER, S. tojmensis SED., S. rarus BHARD. and SAL., Striatopinites trivialis (SCHATK.) SIV., Taeniaesporites tenor (ZAUER) WAR. 41. Aleurolites, calcareous sandstones, marls, argillites, limestones – Aleurolites, grès calcaires, marnes, argilites, calcaires. Bivalves: Netschajewia pallasi VERN., Schizodus sp., Plurophoria simplex REIS., Nuculana wimensis KUL., Myonia komiensis (MASL.), Oriocrasatella plana (GOLOV.); Ostracods: Fascianella plotnikovi MAND., Pseudobythocypris pediformis KNIGHT, Ps. cf. osagensis KELLETT, Moorea sp., Cavellina sp. 42. Foraminifera: Pseudonodosaria lata K.M. MACL., Cornuspira microsphaerica GERKE, Memigordius planispiralis K.M. MACL.; Bivalves: Pseudomonotis permianus MASL., Pseudobakewellia antiquaeformis NOIN., Nuculavus wymensis (KEYS.); Bryozoans: Dyscritella spinigeriformis MOR., Rhombotrypella summa MOR.; Ostracods: Healdia pseudosimplex KOTSCH., Cavellina grandis SCHNEID., Actuaria seconda KOTSCH.; Brachiopods: Lingula orientalis GOL., Rhynchopora geinitziana VERN., Pinegathyris royssiana (KEYS.), Dielasma elongeta (SCHLOTH.), Amblypterus tuberculatus (EICHW.); Flora s.l.: Calamospora rotunda (NAUM.) WAR., Trachytriletes punctatus (WALTZ) LUBER, Granulatisporites granifer (K.-M.) WAR., Cyclogranisporites osmundae (SAMOIL.) WAR., Raistrickia cardioformis (LUBER) WAR., Azonaletes plicatus WALTZ, Ginkgocycadophytus sp., Vittatina vittifer LUBER, V. persecta ZAUER, Cordaitina sp. 43. Upper part: marls, limestones, aleurolites – Partie supérieure : marnes, calcaires, aleurolites. Foraminifera: Hyperammina delicatula GERKE, Nodosaria suchonensis K.M. MACL., Pseudonodosaria nodosariaeformis K.M. MACL.; Ostracods: Moorea facilis SCHNEID., Healdia subtriangula KOTSCH., Healdianella vulgaris KOTSCH. Flora s.l.: Aulosteges horrescens (VERN.), Rhynchopora variabilis STUCK., Cleiothyridina pectinifera (SOW.). Lower part: argillaceous biogenic limestones, aleurolites, marls, argillites; in western part, biohermes – Partie inférieure : calcaires biogéniques argileux, aleurolites, marnes, argilites ; biohermes dans la partie occidentale. Foraminifera: Verneuilloides andrei UCH., Nodosaria hexagona TSCHERD., N. pseudoconcinna K.M. MACL., Pseudonodosaria lata K.M. MACL.; Ostracods: Kirkbya ingeniosa SCHN., Amphissites tscherdynzevi POSNER, Healdia postcornuta SCHN. Brachiopods: Licharewia stuckenbergi (NETSCH.), L. latiareata (NETSCH.), Cancrinella cancrini VERN. 44. Foraminifera: Saccamina rotunda K.M. MACL., Nodosaria hexagona TSCHERD., N. elabugae TSCHERD., Lingulonodosaria kamensis K.M. MACL., Frondicularia longissima (K.M. MACL.). Ostracods: Healdia postcornuta SOHN, H. pseudosimplex KOSTCH. Brachiopods: Lingula orientalis GOL., Globiella tschernyschevi (NETSCH.), Schisodus rossicus VERN. Flora s.l.: Phylladoderma arberi ZAL., Pecopteris synica ZAL., Meristophyllum sojamaeanum ZAL. 45. Flora s.l.: Calamospora microrugosa (IBR.) SCHOPF, WILS. and BENT., Cyclogranisporites osmundae (WALTZ) WAR., Spinosisporites spinosus (WALTZ), Raistrickia cardioformis K.M., Lycospora variabilis JANS., Entylissa capereta LUBER, Vittatina cincinnata LUBER, V. striata LUBER, Cordaitina rugulifera (LUBER) SAMOIL. 46a. Flora s.l.: Netschajewia tschernyschewi LICH., Moorea facilis SOHN, Paleodarwinula jatskovae (KASH.), P. fainae (BEL.), P. trita (PAL.), P. irenae (BEL.), Vymella dobryinini (KASH.), Trachytriletes punctulatus NAUM., Cyclogranisporites osmundae (SAMOIL.) WAR., C. gibberuliformis (K.-M.) WAR. 46b. Ostracods: Paleodarwinula procera (MAND.), P. Angusta (MAND.), P. Abunda (MAND.), P. inerta (KASH.), P. sobela (KASH.), P. sinedorensis (KASH.), P. pergusta (KASH.). Brachiopods; Lioestheria simplex MOL., Pseudoestheria mezeniana MOL., Ps. borealis MOL., Notocrypta timanica MOL. Flora s.l.: Calamospora microrugosa (IBR.) SCHOPF, WILS. and BENT., Trachytriletes punctalatus NAUM., Cyclogranisporites gibberuliformis K.-M., Granulatisporites gibbosus IBR., Spinosisporites rectispinus LUBER, Remysporites psilopterus LUBER, Cirratriradites ornatus (LUBER) HART., C. procumbens (LUBER) SIV., Lycospora variabilis JANS., Vittatina vittifer LUBER, V. striata LUBER, Florinites luberae SAMOIL., Alisporites ovatus JANS. 46c. Flora s.l.: Calamospora microrugosa (IBR.) SCHOPF, WILS. and BENT., Trachytriletes asperatus (LUBER) ISCH., Cyclogranisporites gibberu- liferus K.-M., Raistrickia cardioformis (K.-M.) WAR., Azonaletes levis LUBER, Vittatina vittifer LUBER, V. striata LUBER, Florinites luberae SAMOIL., Cordaitina vera (SADK.) SIV., C. subrotata (LUBER) SAMOIL., Protopodocarpites alatus (LUBER) SAMOIL., Striatodiploxypinites striatus (LUBER) WAR., Striatopinites trivialis (SCHATK.) SIV. 47. Ostracods: Paleodarwinula perlonga (SHARAP.), Suchonellina inornata (SPIZH.), S. parallela (SPIZH.) 48a. Brachiopods: Cancrinalia cancrini (VERN.). Bivalves: Promytilus retusus CHRON., Permophorus angustus MUROM., Stutchburia pallasi (VERN.), Myonia komiensis (MASL.), Praeunduloyva leonidi MUROM. 48b. Bivalves: Aviculopecten wymensis LICH., Anisotrypella borealis MOR., Discritella spinigeriformis MOR., Paralioclema multispinosum MOR., Wyatkella permiana MOR.; Brachiopods: Aulosteges longus NETSCH., Cancrinella cancrini (VERN.), Ulugkemia leonidi NOV., U. petri NOV., Limnadia kastiatum NOV. 542 V.P. Gorsky et al. / Geobios 36 (2003) 533Ð558

49. Foraminifera: Nodosaria elabugae TSCHERD., N. suchonensis K.M. MACL. Brachiopods: Discritella spinigeriformis MOR., Anisotrypella borealis MOR., Fistulipora dybowskyi GOR., Cancrinella cancrini (VERN.), Permospirifer keyserlingi (NETSCH.), Licharewia sp., Rhynchopora geinitziana (VERN.), Beecheria netschajewi GRIG.; Flora s.l.: Stepanovites meyeni KOZ. and MOV. 50. Bivalves: Prilukiella mirabilis (GUS.), Maturipina kholunicensis GUS. Ostracods: Paleodarwinula fragiliformis KASH., Sinusuella ignota SPIZH., Placidea lutkevichi (SPIZH.), Suchonellina futschiki (KASH.). 51. Bivalves: Prilukiella mirabilis (GUS.), P. nitida GUS., Palaeanodonta subcastor AMAL. Ostracods: Paleodarwinula belousovae (KASH.), Vymella dobrynini (KASH.), Placidea lutkevichi (SPIZH.). Flora s.l.: Callipteris (Feonia) aequabilis MEYEN, Phylladoderma meridionalis MEYEN, Rhaphidopteris sp., Slivkovia sp., Aequistomia sp.; Miospore complex I2 (see Table 2). 52. Green to grey argillites, marls, aleurolites with coal levels; eastern part: red aleurolites – Argilites vertes à grises, marnes, aleurolites avec des niveaux de charbon; dans la partie orientale: aleurolites rouges. Foraminifera: Pseudonodosaria ventrosa SCHL., Frondicularia pseudotriangularis GERKE. Bivalves: Abiella subovata (JON.), Anthraconauta pseudophillipsi FED., A. iljinnklensis FED., A. globosa FED., Syniella (Pseudosinomya) bella GUS.. Ostracods: Paleodarwinula jatskovae (KASH.), P. irinae (BEL.), P. pergusta (KASH.), P. belousovae (KASH.), P. borealis (KASH.), Prasuchonella tichwinzkajae (BEL.), Placidea petchorica (SCHN.), Volganella laevigata SCHN. 53. Red argillites with intercalations of aleurolites and sandstones – Argilites rouges avec intercalations d’aleurolites et grès. Ostr Paleodarwinula abunda (MAND.), P. angusta (MAND.), P. sobela (KASH.), Prasuchonella scolia (MAND.), Sinusuella pergraphica MAND. 54. Red argillites, marls, aleurolites – Argilites rouges, marnes et aleurolites. Ostracods: Paleodarwinula angusta (MAND.), P. inerta (KASH.), P. procera (MAND.), P. burajevoensis (PAL.) 55. Argillites and sandstones with intercalations of limestones – Argilites et grès à intercalations de calcaires. Ostracods: Paleodarwinula sobela (KASH.), P. lancetiformis (KASH.), Praesuchonella stelmarta KASH. 56. Ostracods: Paleodarwinula belousovae (KASH.), Vymella dobrinini KACH., Placidea lutkovichi (SPIZH.). Brachiopods: Lingula orientalis GOL.;

Flora s.l.: Bardocarpus superus NEUB., Miospore complex I2 (see Table 2). 57. Bivalves: Astartella permocarbonica TSCH.. Brachiopods: Leaia cargalensis NETSCH., Lingula orientalis GOL., L. hyperborea IFAN., L. credneri GEIN.. Ostracods: Paleodarwinula pergusta (KASH.), P. jatskovae (KASH.), Placidea mesotuberculata SCHN., Volganella laevigata SCHN., Cavellina edmistonae HARRIS and LALICKER, C. grandis KOTSCH.; Miospore complex I1 (see Table 2). 58. Foraminifera: Frondicularia tsaregradski K.M. MACL., F. dilema GERKW, F. geinitziniformis (K.M. MACL.), F. longissima (K.M. MACL.), Lingulonodosaria cf. farcimeniformis (K.M.–MACL.), Anthraconaia cf. castor (EICHW.), Nuculavus cf. wymensis KEYS.. Bivalves: Astartella sp. Brachio- pods: Lingula orientalis GOL., L. credneri GOIN., Cancrinella cancriniformis TSCHERN.. Ostracods: Healdia postcornuta SCHN., H. enervis MART., H. kozhimica GUSS., Cavellina permiana KOTSCH., Graphiodactylus uchtensis MART., Monoceratina cf. kungurica (MART.), Bairdia sp., Placidea lutkevichi

(SPIZH.); in the lower part, miospore complex H 2 and H1 (see Table 2).

Table 4 Eastern European basin, Arkhangelsk belt, part 2 Bassin Est-Européen, ceinture d’Arkhangelsk, deuxième partie V.P. Gorsky et al. / Geobios 36 (2003) 533Ð558 543

59. Bivalves: Anthraconauta moltschanovi BETEKHT., A. degeniaensis BETEKHT., Concinnella gravis PAP, C. cf. concinna (JONS.). Flora s.l.: Protosphagum nervatum NEUB., Crassinervia ovifolia NEUB., Rhipidopsis ginkgoides SCHMALH., Phylladoderma arberi ZAL., Bardocarpus superus

NEUB., Nucicarpus piniformis NEUB.; Miospore complex I2 (see Table 2) 60. Bivalves: Palaeomutela visenda GUS., P. monstruosa GUS., P. krotowi NETSCH., Anthraconauta volgensis GUS., A. gibbosa BETEKHT., Peraborella fedotovi (KAN.), Syniella bella (GUS.), S. angusta KAN. Ostracods: Paleodarwinula irinae (BEL.), P. chramovi (GLEB.), P. fainae (BEL.), Placidea lutkevichi (SPIZH.), P. petschorica (SCHN.). Flora s.l.: Intia vermicularia NEUB., Odontopteris rossica ZAL., Callipteris adzvensis ZAL., Cordaites clercii (ZAL.) RADCZ., C. candalepensis ZAL., Ruﬂoria synensis (NEUB.) MEYEN, Rhaphidopteris praecursoria MEYEN, Phylladoderma arberi ZAL., Samaropsis extensa NEUB., S. postfrigida DOMBR., Bardocarpus superus NEUB., Permotheca sardykensis ZAL., Nucicarpus piniformis NEUB.; Miospore complex I2. (see Table 2) 61. Bivalves: Anthraconaia rhomboida (NETSCH.), A. oblonga (BEL.), Mrassiella komiensis KAN., Concinnella angulata POG., Abiella kolvae KAN., Palaeomutela cf. alta POG. Ostracods: Paleodarwinula cf. martjevi (PAL.), P. cf. lubimovae (KASH.), Vymella timanica (KASH.), Sinusuella pergraphica MAHD. Flora s.l.: Intia variabilis NEUB., Kosjunia polyedra FEF., Ruﬂoria synensis (NEUB.) MEYEN, Phylladoderma sp., Zamiopteris neuburgiana

MEYEN, Samaropsis niamdensis NEUB., Bardocarpus superus NEUB.; Miospore complex I1 – H2 (see Table 2) 62. In the upper part – Dans la partie supérieure. Bivalves: Wilkingia komiensis MASL., Nuculavus wymenzis (KSYS). Brachiopods: Lingula ex gr. orientalis GOL. Ostracods: Vymella timanica (KASH.), V. petschorica (KASH.), Placidea cf. mesotuberculata SCHN. In the lower part – Dans la partie inférieure. Foraminifera: Protonodosaria praecursor RAUS., Nodosaria lata SOSSIP., Pseudonodosaria nodosariaeformis K.M. MACL., Frondicularia planilata GERKE, F. bella GERKE. Brachiopods: Dentalina praenuntia GERKE, Rhynchopora cf. nikitini TSCH., Lissochonetes sp. Flora s.l.: Streblopteris ex gr. pusilla SCHLF., S. sericea VERN., Laevidentalium sp. In all the series – Dans toute la série. Bivalves: Astartella permocarbonica TACHERN., Nuculopsis ellipticus GUS. and POG., Intia variabilis NEUB., Kosjunia retusa FEF., Junjagia glottophylla NEUB. Flora s.l.: Vorcutannularia plicata (POG.)

NEUB., Sphenophyllum comiense ZAL., Ruﬂoria recta (NEUB.) MEYEN, Miospore complex H1 (see Table 2) 63. Ostracods: Paleodarwinula borealis (KASH.), P. procera (MAND.), P. pseudopetschorica (KASH.), Sinusuella pergraphica MAND. Foraminifera: Palaeomutela sp., Palaeanodonta vorcutica POG., P.exgr.castor (EICHW.), Concinnella ex gr. concinna JONES. Flora s.l.: Cordaites clercii (ZAL.) RAD., Rhipidopsis sp., Phylladoderma arberi ZAL., Bardocarpus superus NEUB., B. synensis DOMB.; in the upper part: Miospore complex I; in the middle part:

Miospore complex I2 (see Table 2) 64. In the upper and middle parts, dans les parties supérieure et moyenne: Bivalves: Anthraconaia longissima (NETSCH.), A. pseudocastor (BETEKHT.), Abiella subovata (JON.), Mransiellina sp. Foraminifera: Concinnella solita PAP. Brachiopods: Lingula sp., Aulosteges sp., Cancrinella ex gr. cantrini VERN., Waagenoconcha cf. purdoni DAV., Lissochonetes aff. rotundatus (TOULA). Flora s.l.: Intia sp., Comia peraborensis ZAL., Lepeophyllum trigonum NEUB.,

Cordaites clercii (ZAL.) RADCZ., Ruﬂoria sp., Miospore complex I1 (see Table 2) 65. In the lower part – Dans la partie inférieure. Bivalves: Anthraconaia oblonga (BEL.), Mrassiella trapezoidalis (KROT.). Ostracods: Graphiodactylus uchtensis Mart.; Brachiopods – brachiopodes: Swalbardia capitolina TOULA, Rhynchopora geinitziana VERN., Lissochonetes sulaensis V. BARCH., Megousia kulikii FRED. Flora s.l.: Jakovlevia mammata KEYS., Viatscheslavia sp., Viatscheslaviophyllum vorcutense NEUB., Sphenophyllum comiense

TSCHIRK., Samaropsis vorcutana TSCHIRK., S. striata NEUB.; Miospore complex H2 (see Table 2) 66. Foraminifera: Protonodosaria proceraformis GERKE, Nododosaria lata SOSSIP., N. cassiaformis IGON., Rectoglandulina pseudoborealis SOSSIP., Frondicularia sectorialis VOR., F. subtilistriata GORKEL, F. bajcurica SOSSIP. Bivalves: Polidevcia cf. formosa DEMB., Nuculopsis wymensis (KEYS.), Nuculana sinuata (DEMB.). Ostracods: Healdia ovataeformis SCHN. Brachiopods: Megousia kulikii FRED. Flora s.l.: Vorcutannularia plicata (POG.)

NEUB., Sphenophyllum thonii MAHR., Samaropsis subelegans NEUB., Sylvella aleta ZAL., Miospore complex H1 (see Table 2) 67. Flora s.l.: Protosphagum nervatum NEUB., Crassinervia ovifolia NEUB., Rhipidopsis ginkgoides SCHMALH., Phylladoderma arberi ZAL., Bardocarpus superus NEUB. 68. Foraminifera: Palaeomutela visenda GUS., P. petschorica KAN. Bivalves: Syniella bella (GUS.), Anthraconauta volgensis GUS. Ostracods: Paleodarvinula irenae (BEL.), P. fainae (BEL.), P. chramovi (GLEB.), Placidea lutkevichi (SPIZH.), P. petchorica (SCHN.). Flora s.l.: Intia vermicularis NEUB., Comia pereborensis ZAL., Callipteris adzvensis ZAL., Cordaites clercii (ZAL.) RADCZ., Ruﬂoria synensis (NEUB.) MEYEN, Rhaphidopteris praecursoria MEYEN, Phylladoderma arberi ZAL., Samaropsis extensa NEUB., S. postfrigida DOMB., S. pajchoica DOMB., Bardocarpus superus NEUB.;

Miospore complex I2 (see Table 2) 69. In the lower part: Bivalves: Khosedaella alta POG., Concinnella concinnaeformis POG., C.exgr.concinna (JONES), Palaeomutela cf. alata POG., Oligodontella ex gr. geinitzi AMAL. Ostracods: Paleodarwinula cf. martjevi (PAL.), P. cf. lubimovae (KASH.), Vymella timanica (KASH.), Sinusuella pergraphica MAND. Flora s.l.: Intia variabilis NEUB., Kosjunia polyedra FEF., Todites sp., Oligocarpia permiana FET., Ruﬂoria synensis (NEUB.) MEYEN,

Phylladoderma sp., Zamiopteris neuburgiana MEYEN, Samaropteris niamdensis NEUB., Bardocarpus superus NEUB.; Miospore complex H2–I (see Table 2) 70. In the upper part; dans la partie supérieure: Bivalves: Wilkingia komiensis MASL., Nuculavus wymensis (KEYS.). Ostracods: Vymella timanica KASH., Sinusuella pergraphica MAND. In the middle part – Dans la partie moyenne. Foraminifera: Protonodosaria praecursor RAUS., Nodosaria lata SOSS., Frondicularia planilata GERKE. Bryozoans: Rhinchopora nikitini TSCH. In the lower part – Dans la partie inférieure: FLORA s.l.: Intia variabilis NEUB., Junjagia glottophylla NEUB., Vorcutannularia plicata (POG.) NEUB., Ruﬂoria recta (NEUB.) MEYEN. 71. Foraminifera: Palaeomutela talbeica POG., Anthraconauta moltachanovi BETCKHT., Adzvaella tatarica (KAN.), Concinnella buredanica POG. Brachiopods: Lioestheria raslovensis NOVOJ., Pseudoestheria dvinaensis MOLIN., Ps. nordvikensis (NOVOJ.), Ps. shujaensis NOVOJ., Eulimnadia lima NOVOJ., E. pausa NOVOJ. Ostracods: Paleodarwinula chramovi (GLEB.), P. fragiliformis (KASH.), P. theodorovichi (BEL.), Prasuchonella stelmachovi (SPIZH.). Flora s.l.: Polyssaievia deﬂexa NEUB., Viatscheslavia chejagensis NEUB., Cladophlebis aff. nystroromii HALLE, Pecopteris anthriscifolia (GOEPP.) ZAL. f. adzvensis FET., Callipteris elegans (SCHMALH) MEYEN, Phylladoderma arberi ZAL., Sphenobaiera sp., Glossophyllum sp., Sporophyl- lites petschorensis (SCHMALH.) Meyen; in the upper part, Miospore complex J; in the lower part Miospore complex I (see Table 2). In the lower part; dans la partie inférieure: Bivalves: Palaeomutela krotowi NETSCH., P. visenda GUS., P. petschorica KAN., Pereborella fedotovi (KAN.), Anthraconauta gibbosa BETEKHT., Abiella subovata (JON.) 72. Bivalves: Palaeomutela celebrata GUS., Concinnella pajchoica POG. Ostracods: Paleodarwinula belousovae (KASH.). In the upper part; dans la partie supérieure. Flora s.l.: Paikhoia tschernovii ZAL., Asterotheca? pluriseriata FEF., Pecopteris obtusa RADCZ., Syniopteris nesterenkoi ZAL., Phylladoderma arbori ZAL., Pereborites rarinervis ZAL., Bardocarpus superus NEUB., Nucicarpus piniformis NEUB.; Miospore complex I (see Table 2). In 544 V.P. Gorsky et al. / Geobios 36 (2003) 533Ð558 the lower part; dans la partie inférieure. Bivalves: Anthraconauta uralica KAN., Abiella kolvae KAN., Concinnella anomala SIB. and PAP. Flora s.l.: Viatscheslavia vorcutensis ZAL., Sphenophyllum comiense TSCHIRK., Pecopteris compta RADCZ., Compsopteris adzvensis ZAL., Callipteris adzvensis

ZAL., Ruﬂoria synensis (ZAL.) MEYEN, Bardocarpus superus NEUB., Samaropsis vorcutana TSCHIRK.; Miospore complex H2 (see Table 2) 73. Bivalves: Anthraconauta subovalis POG., Concinnella angulata POG., Mranciella trapezoidalis (KROT.). Ostracods: Paleodarwinula angusta (MAND.), Sinusuella pergraphica MAND. In the upper part – Dans la partie supérieure: Compsopteris adzvensis ZAL., Callipteris sp., Samaropsis postfrigida DOMBR.; Miospore complex H2 (see Table 2). In lower part – Dans la partie inférieure: Lingula hyperborea IFAN., L.? miloradovichi IFAN., Astartella raricostata DEMB., Hemicycloleaia elata (ZASP.), Intia variabilis NEUB., Viatscheslavia vorcutensis ZAL., Samaropsis extensa NEUB.; Miospore complex H1 (see Table 2) 74. Foraminifera: Nodosaria netschajewi TSCHERD. Bivalves: Aviculopecten frederixi LUTK. and LOB., Astartella permocarbonica TSCHERN., Schizodus subobscurus LICH.; Brachiopods: Lingula? miloradovichi IFAN., Megousia kuliki (FRCKS.), Waagenowoncha irginae (STUCK.)

Table 5 Eastern European basin, Syktyokar belt Bassin Est-Européen, Ceinture de Syktyokar

75. Poldarskaia formation: aleurolites, sandstones with intercalations of limestones and marls; in the lower part, sands with intercalations of gravels and conglomerates – Formation de Poldarskaia: aleurolites, grès à intercalations de calcaires et de marnes ; dans la partie inférieure, sables à intercalations de graviers et de conglomérats. Ostracods: Prasuchonella stelmachovi (SPIZH.), Suchonellina inornata SPIZH., S. fragilis (SCHN.). Conchostraca: Palaeorthothemos lochtaensis NOVOJ., Sphaerorthothemos cellelatus (LUTK.)

76. Nuksenitskaia member: aleurolites, argillites, marls – Membre Nuksenitskaia: aleurolites, argilites, marnes. Ostracods: Suchonellina inornata SPIZH., S. parallela SPIZH., Prasuchonella stelmachovi (SPIZH.), Sinusuella ignota SPIZH. Vertebrates s.l.: Acrolepis macroderma EICHW. Dmitrievskaia member: dolomites, dolomitic marls, sometimes siliciﬁed – Membre Dmitrievskaia: dolomites, marnes dolomitiques, quelquefois siliciﬁées. Verkhnetozmenskaia member: marls, aleurolites and argillites, sandstones often gypseous with lenses of white sands – Membre Verkhnetozmenskaia: marnes, aleurolites et argilites, grès souvent gypseux à lentilles de sables blancs.

77. Nijneonstinskaia formation; Formation de Nijneonstinskaia. Ostracods: Paleodarwinula sp., Permiana sp. Shardinskaia member: grey marls gypseous with intercalations of limestones and dolomites – Membre Shardinskaia: marnes grises gypseuses avec des intercalations de calcaires et de dolomies. Karpogorskaia member: brown red aleurolites, sandstones strongly gypseous with intercalations of argillites – Membre Karpogorskaia: aleurolites rouge-brun, grès fortement gypseux avec des intercalations d’argilites. Maregorskaia member: brown aleurolites and argillites with gypsum – Membre Maregorskaia: aleurolites brunes et argilites avec gypse. V.P. Gorsky et al. / Geobios 36 (2003) 533Ð558 545

78. Foraminifera: Tetrataxis lata SPAND., Geinitzina postcarbonica SPAND., Glomospira pusilla GEIN., Netschajewia striata SCHLOTH. Bivalves: Pseudomonotis permianus MASL., Aviculopecten mutabilis LICH., Streblochondria sericea VERN., S. pusilla SCHLOTH. Gasteropods: Straparollus rossicus LICH. 79. Foraminifera: Hemigordius discoides K.M. MACL., H. schlumbergeri (HOWCH.), Nodosaria hexagona TSCHERD., N. noinskii TSCHERD., N. farcimeniformis K.M. MACL., N. krotowi TSCHERD., N. cushmani PAALZ., Pseudonodosaria lata K.M. MACL., Lingulonodosaria kamensis K.M. MACL., Cornuspira megasphaerica GERKE, Nuculavus trivialis EICHW., Pleurophorus subocostatus BROWN., Polidevcia kazanensis (VERN.), Schizodus rossicus VERN. Ostracods: Pseudoparaparchites furmidabilis SCHN., Cavellina grandis SCHN., Healdianella vulgaris KOTSCH., Amphissites tscherdynzevi POSN., Kirkbya ingeniosa SCHN. Brachiopods: Neochonetes pinegenis KUL., Globiella tschernyschewi (NETSCH.), Licharewia rugulata (KUT.), Aulosteges horrescens (VERN.), Dielasma elongata SCHLOTH., Cancrinella cancrinoides (KUT.), Rhynchopora geinitziana (VERN.), Stepanoviella hemisphaerium (KUT.), Stepanovites meyeni KOZ. and MOV. 80. Komaritskaia member: red aleurolites and sandstones with carbonated nodules and lenses of sandstone and sand at the bottom – Membre Komaritskaia: aleurolites et grès rouges avec des nodules de carbonates et des lentilles de grès et sables à la base. Foraminifera: Palaeomutela curiosa AMAL., P. keyserlingi AMAL. Bivalves: Opokiella tschernysohewi PLOTN. Ostracods: Suchonellina parallela SPIZH., S. trapezoida (SHAR.), S. fragiloides (ZEK.), S. fragilina (BEL.), Suchonella typica SPIZH., S. cyrta ZEK. Flora s.l.: Tatarina conspicua MEYEN, Peltaspermopsis sp., Permotheca sp. Tsherevkovskaia member: red massive aleurolites, marls with intercalations of argillites, grey marls and limestones with lenses of sandstone and sand – Membre Tsherevkovskaia: aleurolites rouges massives, marnes avec intercalations d’argilites, marnes et calcaires gris avec des lentilles de grès et de sables. Ostracods: Suchonellina inornata SPIZH., S. parallela SPIZH., S. fragilina (BEL.), S. vladimirinae (BEL.), S. ignatjevi ZEK., Suchonella cyrta ZEK., S. typica SPIZH., S. auriculata (SHAR.), Darwinuloides svijazhicus (SHAR.) 81. Vertebrates s.l.: Acrolepis macroderma EICHW., Dvinosaurus primus AMAL., Kotlassia prima (AMAL.), Karpinskiosaurus secundus (AMAL.), Jugosuchus licharevi RIAB., Inostrancevia alexandri AMAL., In. latifrons PRAVOSL., Anatherapsidus petri (AMAL.), Dvinia prima AMAL., Scutosaurus karpinzkii AMAL., Pravoslavlevia parva (PRAVOSL.), Dicynodon amalitzkii SUSH., D. trautscholdi AMAL., D. annae (AMAL.) 82. Poldarskaia formation: marls, dolomitic limestones, argillites, aleurolites, some conglomerates, sandstones, lenses of sandstone and sand with bones – Formation Poldarskaia: marnes, calcaires dolomitiques, argilites, aleurolites, quelques conglomérats, grès, lentilles de grès et de sables avec des fragments d’os. Bivalves – bivalves: Palaeomutela ovalis AMAL., P. oleniana GUS., Prilukiella janischewskia PLOT. Ostracods: Suchonellina inornata SPIZH., S. parallela SPIZH., Prasuchonella stelmachovi (SPIZH.). Vertebrates s.l.: Isadia suchonensis A. MIN., Platysomus biarmicus EICHW., Kotlassia prima (AMAL.), Dvinosaurus primus AMAL., Jugosuchus boreus IVACH., Niuksenitia sukhonensis TAT., Stinamara abrasa SAID. Flora s.l.: Tatarina sp. In the upper part – Dans la partie supérieure: Ostracods: Suchonellina vladimirinae (BEL.), S. fragilina (BEL.), S. fragilis SCHN., Suchonella cyrta ZEK., S. auriculata (SHAR.), Permiana tuberculata KASH. 83. Nuksenitskaia member: alternation of brown, grey dolomites, marls and argillites with rare intercalations of white sands – Membre Nuksenitskaia: alternance de dolomies brunes, grises, de marnes et d’argilites avec de rares intercalations de sables blancs. Ostracods: Prasuchonella stelmachovi (SPIZH.), Suchonellina inornata SPIZH., S. futschiki KASH. 84. Dmitrievskaia member: light grey and white dolomites with silex concretions – Membre Dmitrievskaia: dolomies gris clair et blanches avec des concrétions de silex. Ostracods: Prasuchonella stelmachovi (SPIZH.), Suchonellina inornata SPIZH., Permiana oblonga POSN. Verkhnetoz’menskaia member: dolomitic grey aleurolites and marls, grey and brown dolomites and argillites – Membre Verkhnetoz’menskaia: aleurolites et marnes grises dolomitiques, dolomies et argilites brunes et grises. Ostracods: Suchonellina inornata SPIZH., S. futschiki (KASH.). Vertebrates s.l.: Platysomus biarmicus EICHW., Amblypterus cf. tuberculatus EICHW., Paraamblypterus sp., Acrolepis cf. rhombifera EICHW. 85. Aleurolites, argillites, marls, limestones and dolomites red and grey, with intercalations and lenses of sands and sandstone – Aleurolites, argilites, marnes, calcaires et dolomies rouges et grises, avec des intercalations et des lentilles de sables et de grès. Ostracods: Suchonellina inornata SPIZH., S. parallela SPIZH., S. futschiki (KASH.), S. undulata (MISH.), Prasuchonella stelmachovi (SPIZH.), Permiana elongata (POSN.), Sinusuella ignota SPIZH. Flora s.l.: Calamospora sp., Vittatina vittifer LUBER, Cordaitina sp., Podocarpites alatus LUBER, Piceites sublevis LUBER, Lebachiites sp., Striatohaplo- pinites elongatus LUBER, S. perfectum NAUM., Hamiapollenites tractiferinus (SAMOIL.) JANS, Vitreisporites sp. In upper part – Dans la partie supérieure: Ostracods: Suchonellina fragilis (SCHN.), S. ignatjevi ZEK., Suchonella cyrta ZEK., S. auriculata SHAR., S. typica SPIZH. 86. Sandstones, sands, aleurolites, marls with thin levels of dolomitic limestones and argillites – Grès, sables, aleurolites, marnes avec des lentilles peu épaisses de calcaires dolomitiques et d’argilites. Ostracods: Paleodarwinula sp., Prasuchonella sp. Vertebrates s.l.: Palaeoniscidae gen. ind., Slivkovia sp.; Flora s.l.: Leiotriletes rotundum NAUM., Azonaletes levis LUBER, Ginkgocycadophytus tunguskensis (ANDR.) LUBER, Vittatina striata LUBER, Cordaitina sp., Podocarpites sp., Protodiploxypinus silvestritypus SAMOIL. 87. Sand and sandstones with dolomitic cement with intercalations of argillites, marls, dolomites and gypsum – Sables et grès avec ciment dolomitique et avec des intercalations d’argilites, de marnes de dolomies et de gypse. Vertebrates s.l.: Ennatosaurus tecton VJUSCH. Mar’egorskaia member: argillites and gypseous aleurolites with intercalations of marls, dolomites and sandstones – Membre Mar’egorskaia: argilites et aleurolites gypseuses avec des intercalations de marnes, dolomies et grès. 88. Upper part: anhydrites, in alternation with gypsum, dolomites, marls – Partie supérieure: anhydrites en alternance avec du gypse, des dolomies et des marnes. Middle part: dolomites, dolomitic marls, argillites with intercalations of gypsum – Partie moyenne: dolomies, marnes dolomitiques, argilites ave des intercalations de gypse. Lower part: dolomitic limestones, silicified, with gypsum – Partie inférieure: calcaires dolomitiques, silicifiés avec gypse. Flora s.l.: Trachytriletes sp., Raistrickia cardioformis (K.M.) WAR., Acanthotriletes gracilis (K.M.) WAR., Leiotriletes inderjensis (MAL.) WAR., Azonaletes tenuis LUBER, Vittatina sp., Cordaitina stenolimbata SAMOIL., C. rugulifera LUBER, Striatopodocarpites tojmensis SED., Striatodiplopinites lata (SCHATK.) WAR., Vitreisporites sp. 89. argillaceous and biogenic limestones with intercalations of argillites and marls – Calcaires biogéniques et argileux avec des intercalations d’argilites et de marnes. Foraminifera: Nodosaria suchonensis K.M. MACL., N. hexagona TSCHERD., N. krotowi TSCHERD., Cornuspira microsphaerica K.M. MACL.; Ostracods: Healdia pseudosimplex KOTSCH., Moorea facilis SCHN., Amphissites tscherdynzevi POSN.; Brachiopods – brachiopodes: Lingula orientalis GOL., Stepanoviella tachernyschewi LICH., Licharewia rugulata (KUT.), L. keyserlingi (NESCH.), Aulosteges horrescens (VERN.), Spiriferella blasii (VERN.). Vertebrates s.l.: Anodontacanthus ruthenorum CHAB., Ctenacanthus sp.; Flora s.l.–flore s.l.: Stepanovites meyeni KOZ. and MOV. 90. Vagorskaia formation: argillaceous calcareous sandstones, in alternation with aleurolites and argillites, with intercalations of marls and biogenic limestones with inclusions of gypsum – Formation d’Ivagorskaia: grès calcaires argileux, en alternance avec des aleurolites et argilites, intercalation de marnes et de calcaires biogéniques à inclusions de gypse. Brachiopods: Lingula credneri GEIN., Licharewia stuckenbergi (NETSCH), Cleiothyridina semiconcava (WAAG.). Vertebrates s.l.: Janassa bituminosa (SCHLOTH.) 546 V.P. Gorsky et al. / Geobios 36 (2003) 533Ð558

91. Flora s.l.: Trachytrilites punctatus (WALTZ), T. typicus NAUM., Acanthotriletes spinosellus WALTZ., A. gracilis (K.M.) WAR., Raistrickia cardioformis (K.M.) WAR., Azonaletes sp., Vittatina sp., Podocarpites major (LUBER) WAR., Protohaploxypinites tecturatus (LUBER) SAMOIL., Striatopodocarpites tojmensis SED., Striatodiplopinites elongatus (LUBER) ZAUER. 92. Red marls and argillites with levels of sandstones and nodules of limestones and dolomites; marnes et argilites rouges avec niveaux de grès et nodules de calcaires et dolomies. Ostracods: Paleodarwinula elongata (LUN.), P. fragiliformis (KASH.), P. perlonga (SHARAP.), P. chramovi (GLEB.), Darwinuloides? tscherdynzevi KASH., Permiana elongata (POSN.) 93. Elvinsky levels: marls and limestones with intercalations of argillites and dark grey limestones – Niveaux Elvinsky: marnes et calcaires avec intercalations d’argilites et de calcaires gris foncé. Glotovsky levels: limestones and marls light grey, sandy, gypsum, intercalations of limestones and aleurolites – Niveaux Glotovsky: calcaires et marnes gris clair, sableux, gypseux, intercalations de calcaires et d’aleurolites. Bivalves: Prilukiella? lata

NETSCH. Ostracods: Paleodarwinula elongata (LUN.), P. fragiliformis (KASH.), P. chramovi (GLEB.), P. kassini (BEL.); Miospore complex J1 (see Table 2) + Striatohaplopinites amplus BALME and HEN., S. ellipticus ZAUER, Vittatina subsaccata SAMOIL., f. connectivalis ZAUER. 94. Foraminifera: Nodosaria elabugae TSCHERD. Bivalves: Anthraconaia gapeevi (FED.), Palaeomutela attenuata GUS., Pseudomonotis permianus MASL., Promytilus retusus CHRON., Pteronites timanicus (MASL.), Pseudobakewellia cerathophagaeformis NOIN. Ostracods: Cavellina grandis SCHN., Healdia pseudosimplex KOTSCH. Brachiopods: Aulosteges fragilis (NETSCH.), Cancrinella cancrini VERN. Conchostraca: Ulugkemia borisi NOVOJ.,

Hemicycloleaia raymondi MOL. Flora s.l.: Viatscheslaviophyllum expansum BRONGN., Walchia sp.; Miospore complex I2 (see Table 2)+Leiotriletes sp., Caytonipollenites sp., Acanthotriletes sp., Libumella rugulifera LUBER, Striatohaplopinites perfectus NAUM., Striatolebachiites varius ZAUER, Hamiapol- lenites tractiferinus SAMOIL., Alisporites tenuicorpus KL., Vesicaspora schemeli KL., Vittatina striata LUBER, Stepanovites meyeni KOZ. and MOV. 95. Ostracods: Paleodarwinula borealis (KASH.), P. fainae (BEL.), Vyaella dobrynini KASH., Permiana posneri KASH. 96. Sandy limestones and argillites with intercalations of marls and limestones – Calcaires sableux et argilites avec intercalations de marnes et de calcaires. Foraminifera: Nodosaria vjatkensis TSCHERD., N. pseudoconcinna K.M. MACL., Lingulonodosaria kamensis K.L. MACL. Bivalves: Nuculopsis wymensis (KEYS.), Cyrtorostra laticostata (NETSCH.), Pseudomonotis permianus MACL., Schizodus netschajewi LICH., Oriocrassatella plana (GOL.). Ostracods: Cavellina unica KOTSCH., Amphissites tscherdynzevi POSN., Healdia pseudosimplex KOTSCH. Brachiopods: Licharewia stuckenbergi (NETSCH.), Cancrinella cancrini VERN.; Flora s.l.: Callipteris adzvensis ZAL., Syniopteris nesterenkoi ZAL., Ruﬂoria synensis (ZAL.); Miospore complex

I1 (see Table 2)+Granizonospora vulgaris (NAUM.), Laevigatosporites vulgaris IBR., f. gigantea LUBER, Florinites luberae SAMOIL., Lebachiacites pulcherrimus (ZAUER), Striatohaplopinites productus ABR., S. perfectus NAUM. 97. Ostracods: Paleodarwinula ironae (BEL.), P. belousovae (KASH.), P. jatskovae (KASH.), Iniella oilmensis KASH.

98. Ostracods: Paleodarwinula lubimovae (KASH.), P. biriensis (PAL.), Sinusuella ingloria KASH.; Miospore complex H2 (see Table 2)+Remysporites psilopterus LUBER, Laevigatosporites vulgaris IBR. f. gigantea LUBER, Lebachiacites pulcherrimus ZAUER, Vesicaspora sp. 99. Vitshegodskaia formation: reddish brown aleurolites, argillites, gypseous marls, with intercalations of limestones – Formation de Vitshegodskaia: aleurolites brun-rougâtre, argilites, marnes gypseuses; intercalations de calcaires. Bivalves: Notocrypta timanica MOL., Limnadea rossica MOL.; Miospore complex H1 (see Table 2)+Cirratriradites procumbens LUBER, Granizonospora vulgaris NAUM., Florinites luberae SAMOIL., Lebachiacites pulcherrimus ZAUER, Striatohaplopinites perfectus NAUM., Vittatina striata LUBER. 100. Ziminskaia formation: brown sandstones, aleurolites and marls in alternation with argillites and anhydrites – Formation de Ziminskaia: grès bruns, aleurolites et marnes en alternance avec des argilites et des anhydrites. Miospore complex H1 (see Table 2)+Cordaitina uralensis LUBER, C. rugulifera LUBER, Klausipollenites sp., Scheuringipollenites ovatus BALME and HEN., Striatohaplopinites perfectus NAUM., Striatolebachiites varius ZAUER, Hamiapollenites tractiforinus SAMOIL., Vittatina subsaccata SAMOIL., Azonaletes irregulariplicatus SAMOIL. 101. Oust’koulomsky levels: red argillites with intercalations of limestones and sandstones – Niveaux Oust’koulomsky: argilites rouges avec des intercalations de calcaires et de grès. Ostracods: Paleodarwinula abunda (MAND.), P. angusta (MAND.), Prasuchonella scolia (MAND.), Vymeolla timanica (KASH). 102. Limestones and argillites; calcaires et argilites. Ostracods: Paleodarwinula angusta (MAND.), Prasuchonella scolia (MAND.), Sinusuella pergraphica MAND. 103. Kirdorsky levels: argillites with intercalations of limestones and sandstones – Niveaux Kirdorsky: argilites avec intercalations de calcaires et de grès. Ostracods: Paleodarwinula angusta (MAND.), P. abunda (MAND.). 104. Sindorsky levels: limestones, dolomites, argillites and sandstones – Niveaux Sindorsky: calcaires, dolomies, argilites et grès. Bivalves: Anthraconala castor (EICHW.), Nuculopsis celebrosus (DEMB.). Ostracods: Bairdia plebeia REUS, Paleodarwinula parphenovae (BEL.), P. angusta (MAND.), P. abunda

(MAND.), Volganella spizharskii MAND.; Vertebrates s.l.: Palaeolimnadiopsis eichwaldi (NETSCH.); Miospore complex H1 (see Table 2)+Granu- latisporites larvatus LUBER. 105. Vertebrates s.l.: Clatysomus biarmicus EICHW., Acrolepis cf. rhombifera EICHW., A. cf. murchisonia EICHW., Atherstonia sp., Amblypterus sp. Flora s.l.: Callipteris adzvensis ZAL., Rhipidopsis ginkgoides SCHMALH., Phylladoderma arberi ZAL., Glostophyllus sp.; Miospore complex J (see Table 2). 106. Ostracods: Paleodarwinula abunda (MAND.), Sinusuella pergraphica MAND. Bivalves: Anthraconauta tschernyschewi KHALF., A. gibbosa BETEKHT., Abiella subovata (JONES), Syniella bella (GUS.). 107. Bivalves: Concinnella angulata POG., Anthraconaia(?) rhomboides (NETSCH.), Mrassiella(?) trapezoidalis (KROT.). Ostracods: Volganella objecta KASH., Iniella intaensis KASH., Placidea cf. petschorica SCHN., Sinusuella pergraphica MAND. Flora s.l.: Intia vermicularis NEUB., Viatscheslavia vorkoutensis ZAL., Knorriopsis uralica ZAL.; Miospore complex H (see Table 2) 108. Alternation of grey argillites, aleurolites and marls – Alternance d’argilites grises, aleurolites et marnes. Bivalves: Concinnella concinnaeformis POG., Lithophaga consobrina EICHW., Netschajewia globosa (NETSCH.), Pseudobakewellia ceratophagaeformis NOIN. Flora s.l.: Bardocarpus aliger

ZAL., Samaropsis triquetra ZAL.; Miospore complex H1 (see Table 2). V.P. Gorsky et al. / Geobios 36 (2003) 533Ð558 547

Table 6 Eastern European Basin, Perm belt Bassin Est-Européen, Ceinture de Perm

109. Molomskaia member – Membre Molomskaia. Ostracods: Darwinuloides tataricus POSN., D. swijazhicus (SHARAP.). Loutugskaia member – Membre Loutugskaia. Bivalves: Palaeanodonta okensis AMAL., P. segmentata GUS., P. solemvaetormis (NETSCH.), Maturipupa tolstichinae (REV.). Ostracods: Suchonella typica SPIZH., S. cornuta KASH. Conchostraca: Pseudestheria olferievi NOVOJ. Flora s.l.: Cyclotungusites vokhmaensis NOVOJ., Tatarina olferievi MEYEN, Nahocynodon seductus TAT.; Miospore complex J2 (see Table 2). Zamochnikovskaia member. Bivalves: Palaeanodonta segmentata GUS., P. solemyaeformis (NETSCH.), Surella blomi GUS., Vetlugaia schmidti (REV.). Ostracods: Suchonella typica SPIZH., S. cornuta KASH. Conchostraca: Leptestheria convexa NOVOJ. 110. Bivalves: Palaeomutela orthodonta AMAL., P. keyserlingi (KEYS.), P. ovalis KEYS., P. inostranzevi AMAL., P. curiosa AMAL., Palaeanodonta okensis AMAL., P. ﬁscheri AMAL. Ostracods: Suchonellina parallela SPIZH., S. inornata var. macra LUN., S. spizharskyi POSNER, S. futschiki KASH. Conchostraca: Limnadia jeskinoica NOV.Vertebrates s.l.: Dvinosaurus egregius SHISHK., Bystrowiana permira VJUSCH., Jugosuchus mirabilis (VJUSCH.). 111. Ostracods: Suchonellina parallela SPIZH., S. inornata SPIZH., S. daedala MISH., Prasuchonella stelmachovi (SPIZH.). Conchostraca: Limnadia kastiana NOV.; Miospore complex J2 (see Table 2). 112. Ostracods: Paleodarwinula perlonga (SHARAP.), P. chramovi (GLEB.), P. kassini (BEL.), P. elongata (LUN.), P. impostor (MISH.), Darwinuloides sentjakensis SHARAP., Prasuchonella nasalis SHARAP.

113. Member 2: Miospore complex I2 (see Table 2). Member 1: Miospore complex I1 (see Table 2). 114. Member 2: alternation of dolomites and dolomitised limestones, grey with nodules of gypsum, with organic remains – Alternance de dolomies et de calcaires dolomitiques gris à nodules de gypse et restes organiques. Member 1: light grey biogenic limestones sometimes with gypsum, sands, shales, with concretions of flint, organic remains; in the lower part, gravels and pieces of dolomitic limestones – Calcaires biogéniques gris clair, quelquefois avec du gypse, des sables, des shales, à concrétions siliceuses et débris organiques; dans la partie inférieure, graviers et fragments de calcaires dolomitiques. Bivalves: Pseudomonotis permianus MASL., Netschajewia globosa (NETSCH.), Streblochonaria sericea (VERN.), Aviculopecten cf. russiensis NETZCH. Brachio- pods: Cancrinella cancrini (VERN.). 115. Bivalves: Streblochonaria sericea (VERN.). Brachiopods: Stenoscisma superstes (VERN.), Cleiothyridinna pectinifera (SOW.), Pinegathyris royssiana (KEYS.), Cancrinella cancrini (VERN.), Globiella hemisphaerium (KUT.), Licharewia rugulata (KUT.), Blasispirifer blasii (VERN.). Ostracods: Amphissites tscherdynzevi POSNER, Healdia oblonga KOTSCH., H. simplex ROUNDY, H. reniformis GUSS., H. postcornuta SCHN., Healdianella vulgaris KOTSCH. 116. Nefedovskaia member: yellow-brown sandstones, oblique stratifications, with conglomerates with pebbles coming from local and metamorphic rocks (up to 20 m), in the upper part brownish red shales, limestones, with limestone concretions – Grès brun-jaunâtre, stratifications obliques, avec des conglomérats à galets provenant des roches locales et métamorphiques (jusqu’à 20 m), dans la partie supérieure des shales rouge-brunâtre, des calcaires et des concrétions calcaires. Ostracods: Suchonella inornata SPIZH.; Vertebrates s.l.: Nanocynodon seductus TAT. In western part: brown and reddish brown 548 V.P. Gorsky et al. / Geobios 36 (2003) 533Ð558 shales, friable, greenish grey – Dans la partie occidentale: shales brun-rougeâtre et bruns, friables, gris-verdâtre. Ostracods: Darwinuloides tataricus (POSNER), D. svijazhicus (SHARAP.). Bikovskaia member: brownish grey sandstones, large, oblique stratifications, with conglomerates and gravels of local rocks and metamorphic rocks from Ural; higher, shales and aleurolites reddish brown, limestones with intercalations of variegated and dark grey limestones – Membre Bikovskaia: grès gris-brunâtre, stratifications larges et obliques, avec conglomérats et graviers d’origine locale et métamorphique provenant de l’Oural; plus haut, shales et aleurolites brun-rougeâtre, calcaires avec intercalations de calcaires gris foncé et bariolé. Ostracods: Suchonella typica SPIZH. Westwards, reddish brown shales, with intercalations of friable aleurolites and coarse sandstones, at the bottom gravels – Vers l’Ouest, shales brun-rougeâtre, avec des intercalations d’aleurolites friables et de grès grossiers, graviers à la base. Ostracods: Suchonella parvaeformis (KASH.), S. typica SPIZH., Darwinuloides tataricus (POSNER), D. svijazhicus (SHARAP.). Kalininskaia member: shales and brown aleurolites, limestones, porous with roots – Membre Kalininskaia: shale et aleurolites brunes, calcaires poreux avec des racines. 117. Poutaninskaia member: shales and aleurolites brownish grey, limestones, variegated marls, dark grey shaly limestones with gastropods – Membre Poutaninskaia: shales et aleurolites gris-brun, calcaires, marnes bariolées, calcaires détritiques gris foncéàgastéropodes. Ostracods: Suchonellina inornata SPIZH., Darwinuloides avijazhicus (SHAR.), Permiana oblonga (POSNER). Bivalves: Palaeomutela verneuili AMAL., P. subparallela AMAL. 118. Urpalovkaia member: at the bottom coarse sandstones with oblique stratifications; higher, red aleurolites and shales with concretions of limestones – Membre Urpalovkaia: à la base, grès grossiers avec stratifications entrecroissées; plus haut, aleurolites et shales rouges à concrétions de calcaires. Rare Paleaomutela trapezoidalis AMAL. (Bivalves – bivalves) and ostracods. 119. Vertebrates s.l.: Scutosaurus rossicus (HART.–WEIN.), Deltovjatia vjatkensis (HART.–WEIN.). 120. Filinskaia member: grey limestones, marls pink, grey, green, calcareous grey marls – Membre Filinskaia: calcaires gris, marnes roses, grises, vertes, marnes calcaires grises. Ostracods: Suchonellina parallela SPIZH. Slobodskaia member: in the lower part, sandstones and conglomerates with oblique stratifications; in the upper part shales and red aleurolites – Membre Slobodskaia: partie inférieure, conglomérats et grès à stratifications obliques; partie supérieure, shales et aleurolites rouges. Bivalves: Palaeomutela ulemaensis GUS. Ostracods: Suchonellina paralella SPIZH. 121. Siramskaia member – Membre Siramskaia: Bivalves: Prilukiella nitida GUS., Palaeomutela vjatkensis GUS., P. ovalis AMAL. Ostracods: Paleodarwinula fragiliformis (KASH.), Permiana oblinga (POSNER). 122. Belokholomnizkaia member: white fine grain sandstones, thin bedded, with quartz and limestones, shales, grey yellowish aleurolites, variegated marls with intercalations of limestones – Membre Belokholomnizkaia: grès blancs à grains fins, finement lités, avec quartz et calcaires, argillites, aleurolites gris-jaunâtre, marnes bariolées avec intercalations de calcaires. Ostracods: Permiana tuberculata KASH. 123. Arthropods: Platyops stuckenbergi TRAUT., P. watsoni EFR. Vertebrates s.l.: Melosourus uralensis MEYER, Nycteroleter kassini TCHUD., Nyctiboetus liteus TCHUD., Leptoropha talanophora (TCHUD.), Gecatogomphius kavejevi VJUSCH., Molybdopygus arcanus TCHUD. 124. Ieinskaia member: brownish red calcareous shales and aleurolites, with grains of quartz and limestones, intercalations of poorly sandy marls and limestones – Membre Ieinskaia: aleurolites et shales calcaires rouge-brunâtre, avec des grains de quartz et de calcaires, intercalations de marnes sableuses et de calcaires. Rare Permiana tuberculata KASH. (Ostracods). Makisinovskaia member: alternation of feldspatho-quartzic sandstones, thin grains, with gypsum, aleurolites, shales, variegated marls and limestones – Membre Makisinovskaia: alternance de grès feldspathoÐ quartzeux, à grain fin avec du gypse, shales, aleurolites, marnes et calcaires bariolés. Ostracods: Darwinuloides edmistonus (BEL.) 125. Member 3: in the lower part, greenish grey sandstones mainly quartzitic; higher, alternation of sandstones with gypsum, aleurolites, shales, marls, dolomites, limestones with pelecypods – Membre3:partie inférieure, grès gris-verdâtre principalement quartizitiques ; puis, alternance de grès avec gypse, aleurolites, shales, marnes, dolomites, calcaires avec pélécypodes. Arthropods: Platyops stuceubergi TRAUT. Member 2: in the lower part, green-grey sandstones with quartz, locally gypsum, in alternation with shales and grey aleurolites, higher, limestones, dolomites, grey marls with foraminifers, brachiopods and bivalves – Membre2:partie inférieure, grès gris-vert à quartz, localement du gypse, en alternance avec des shales et des aleurolites grises ; plus haut, calcaires, dolomites, marnes grises à foraminifères, brachiopodes et bivalves. Member 1: in the lower part, green-grey and brown sandstones, with gypsum nodules; higher, grey shales, dolomites, thinly bedded, limestones sometimes oolithic, bryozoans, marine ostracods – Membre1:partie inférieure, grès bruns et gris-vert, avec nodules de gypse ; plus haut, shales gris, dolomies, finement litées, calcaires quelquefois oolithiques, bryozoaires, ostracodes marins. Brachiopods: Cancrinella cancrini (VERN.) 126. Calcareous member: biogenic limestones, reefal with brachiopods and crinoids, algae and bryozoans, oolithes, with intercalations of marls, shales, sandstones. Membre calcaire : calcaire biogénique, récifal avec brachiopodes et crinoïdes, algues et bryozoaires, avec intercalations de marnes, shales, grès. Foraminifera: Nodosaria elabugae TSCHERD., N. pugioidea ZOL. and IGON., N. netschajewi TSCHERD., N. permiana SPAND., N. krotowi TSCHERD., Frondicularia sp. Brachiopods: Cleiothyridina pectinifera (SOW.), Cancrinella sp. Marly calcareous member: biogenic limestones with stratifications, light marls, dark grey shales. Membre marno-calcaire : calcaires biogéniques stratifiés, marnes claires, shales gris foncé. Foraminifera: Nodosaria elabugae TSCHERD., N. netachajewi TSCHERD., N. krotowi TSCHERD. Bivalves: Dentalina sp. Brachiopods: Cancrinella cancrini (VERN.), Globiella hemisphaerium (KUT.). Sandy calcareous member: greenish grey sandstones, glauconitic limestones, dark grey shales, limestones; eastwards, terrigenous rocks with few intercalations of limestones. Membre calcaro-sableux: grès gris-verdâtre, calcaires glauconitiques, shales gris foncé, calcaires; vers l’Est, roches terrigènes avec quelques intercalations de calcaires. Foraminifera: Cornuspira kamae TSCHERD., Nodosaria netachajewi TSCHERD., Prondicularia longissima sintasica IGON. Brachiopods: Cleiothyridina pectinitera (SOW.), Rhynchopora sp. 127. Foraminifera: Palaeomutela oleniana GUS., P. plana AMAL. Ostracods: Suchonellina parallela SPIZH., S. futschiki (KASH.). Vertebrates s.l.: Pareiasauria sp., Gorgonopsia sp., Inostranzevia sp. 128. Ostracods: Suchonellina parallela SPIZH. 129. Upper part: alternation of conglomerate with pebbles of flint and quartz, sandstones, brown shales, grey marls, algae limestones. Partie supérieure: alternance de conglomérats à galets de silice et de quartz, grès, shales bruns, marnes grises, calcaires algaires. Bivalves: Prilukiella nitida GUS., Anthraconauta volgensis GUS., A. uslonensis GUS. Ostracods: Paleodarwinula fragiliformis (KASH.), P. elongata (LUN.). Vertebrates s.l.: Biarmica tchudinovi IVACH., Brithopus priscus KUT. Lower part: alternation of conglomerate, gravels with flint and quartz, with fossil lignite, grey sandstones with oblique stratifications, shales, calcareous brown red aleurolites, variegated marls, dark grey limestones. Partie inférieure : alternance de conglomérats, graviers avec silice et quartz, lignite fossile, grès gris à stratifications obliques, shales, aleurolites calcaires rouge-brunâtre, marnes bariolées, calcaires gris sombre. Bivalves: Prilukiella lata (NETSCH.), Palaeomutela doratiformis GUS., P. celebrata GUS., Prilukiella fastigata GUS., P. pugnatoria GUS. Ostracods: Paleodarwinula kassini (BEL.) V.P. Gorsky et al. / Geobios 36 (2003) 533Ð558 549

130. Ostracods: Prasuchonella serpula BEL., Paleodarwinula varsanoﬁevae (BEL.), P. alexandrinae (BEL.), Darwinuloides edmistonus (BEL.), D. sentjakensis (SHARAP.). Vertebrates s.l.: Parabradysaurus udmurticus EFR., Archaesyodon praeventor TCHUD., Eotitanosuchus olsoni TCHUD., Ivanto- saurus ensifer TCHUD., Chthomaloporus lenocinator TCHUD., Estemmenosuchus mirabilis TCHUD., Est. uralensis TCHUD., Anoplosuchus tenuirostris TCHUD., Zopherosuchus luceus TCHUD., Otsheria netzvetaevi TCHUD., Brithopus priscus KUT. 131. Ostracods: Paleodarwinula parphonovae (BEL.), P. abunda (MAND.), Vymella timanica (KASH.), V. petschorica (KASH.). Flora s.l.: Comia biarmica (ZAL.) VLAD., Sphenopteris biﬁda SCHMALH., Syniopteris cuneifolia KUTORGA, Callipteris erosa (MORRIS) ZAL., Paracalamites striatus SCHMALH., Odontopteris rossica (ZAL.) VLAD., Syniopteris cuneifolia KUTORGA, Permotheca sardykensis ZAL. 132. To the East, limestones, dolomites, grey shales, marls, in the lower part levels with bivalves and brachiopods. Vers l’Est : calcaires, dolomies, shales gris, marnes; dans la partie inférieure, niveaux à bivalves et brachiopodes. Ostracods: Paleodarwinula monitoria (MAND.), Kemeroviana angulata MAND.; in the lower part Brachiopods: Cancrinella cancrini (VERN.), C. koninekiana (KEY.). Flora s.l.: Intia variabilis NEUB., Viatscheslavia vorcutensis ZAL. To the West, variegated shales with gypsum, marls, limestones, dolomites, with rare ostracods. Vers l’Ouest, shales bariolés à gypse, marnes, calcaires, dolomies, rares ostracodes. Flora s.l.: Odontopteris artipinnata ZAL., Entsovia rarisulcata MEYEN, Samaropsis frigida NEUB., S. vorcutana TSCHIRK.

Table 7 Eastern European basin, Kazan belt Bassin Est-Européen, Ceinture de Kazan

133. Vertebrates s.l.: Chroniosuchus sp. Bivalves: Palaeomutela oleniana GUS., P. plana AMAL., Palaeanodonta okensis AMAL., P. abunda NETSCH. Conchostraca: Pseudoestheria suchonensis (NOVOJ.), Ps. nordwikensis (NOVOJ.) 134. Conchostraca: Pseudoestheria suchonensis (NOV.), Ps. nordwikensis (NOV.), Ps. lineata (LUTK.), Asmussia petasa (NOVOJ.), Concherisma trapezoidalis (NETSCH.), Brachyestheria ovalis MOL., Leptestheria uralensis MOL. Bivalves: Palaeomutela keyserlingi AMAL. Ostracods: Suchonellina parallela SPIZH., S. futechiki KASH. 135. Brown red argillites and aleurolites, dolomitised with intercalations of pink and grey marls, argillites with palygorskite. In the lower part, level of calcareous argillites. Argilites et aleurolites rouge-brun, dolimitisées avec intercalations de marnes roses et grises à palygorskite ; dans la partie inférieure, niveau d’argilites calcaires. 136. Dolomites and dolomitised limestones, grey, siliciﬁed with bivalve, brachiopod, gastropod debris. Dolomies et calcaires dolomitisés, gris, siliciﬁés avec débris de bivalves, brachiopodes, gastéropodes. 137. Yellow grey clastic dolomitised limestones with Licharewia rugulata (KUT.) (Brachiopods). Calcaires dolomitisés clastiques grisÐjaune avec Licharewia rugulata (KUT.). 138. Light grey dolomitised limestones, oolitic with Pseudomonototis garforthensis (KING.) (Bivalves). Calcaires dolomitisés gris clair, oolithiques avec Pseudomonototis garforthensis (KING.). 550 V.P. Gorsky et al. / Geobios 36 (2003) 533Ð558

139. Dolomitised biogenic limestones with Lingula orientalis GOL. (Brachiopods). Calcaires biogéniques dolomitisés avec Lingula orientalis GOL. 140. Vertebrates s.l.: Platysomus biarmicus EICHW., Jugosuchus hartmanni IVACH., Sauroctonus progressus (HAR.–WEIN.), Scutosaurus permianus (HART.–WEIN.), Sc. itilenis IVACH. and LAB., Dicynodon trautsholdi ANAL. 141. In the lower part, brown-grey sandstones and conglomerates with pebbles of metamorphic rocks; in the upper part red-brown argillites and aleurolites, variegated marls, algal stratified limestones. Dans la partie inférieure, grès gris-brun et conglomérats à galets de roches métamorphiques ; dans la partie supérieure, argilites et aleurolites brun-rouge, marnes bariolées, calcaires algaires stratifiés. Bivalves: Palaeomutela doratifomis GUS., Antraconauta volgensis GUS., Palaeanodonta (Kuthreconia) castor (EICHW.). Ostracods: Paleodarwinula theodorovichi (BEL.), Prasuchonella nasalis SHARAP. Vertebrates s.l.: Xenosynechodus egloni GLÙK., Melosaurus psucidens (KONZH.), Lanthanosuchus watsoni EFR., Enosuchus breviceps KONZH., Titanophoneus potens EFR., Syodon biarmicum KUT., Doliosauriscus janshinovi ORL. 142. In the lower part, brown sandstones, polymictites with conglomerates (pebbles of chert and marly limestones); in the upper part, brown argillite and aleurolites, variegated marls, stratified platy and algal limestones. Dans la partie inférieure, grès bruns, polymictites avec conglomérats (galets de cherts et de calcaires marneux) ; dans la partie supérieure, argilites et aleurolites brunes, marnes bariolées, calcaires algaires et en plaquettes bien stratifiés. Bivalves: Anthraconauta uslonensis GUS. 143. In the lower part, greenish grey sandstones, in places, very coarse pebbles mainly quartzose; in the upper part variegated marls and argillites. Dans la partie inférieure, grès gris-verdâtre, localement, galets très grossiers principalement quartzeux ; dans la partie supérieure, marnes et argilites, bariolées. Bivalves: Nuculavus trivialis (EICHW.), Schizodus rossicus VERN., Pseudomonotis permianus MASL., Streblochondria sericea (VERN.), Pseudobakewellia ceratophagaeformis NOIN., Lithophaga consobrina (EICHW.); Brachiopods: Cancrinella cancrini (VERN.). Arthropods : Platyops stuckenbergi TRAUT. Vertebrates s.l.: Nyateroleter barshkyricus EFR., Rhipaeosaurus tricuspidens EFR., Phthinosaurus borissiaki EFR., Belebey vegrandis IVACH. Flora s.l.: Signacularia noinskyi ZAL., Sphenophyllum stueckenbergii SCHMALH., Phyllododerma volgensis ESAUL., Permotheca sardykensis ZAL. 144. Light grey cavernous oolithic limestones and sandstones, thin stratifications with in some places gypsum; at the base, brown greyish sandstones, grey yellowish argillites. Grès et calcaires oolithiques caverneux gris clair, fines stratifications avec quelques niveaux de gypse ; à la base, grès brun-grisâtre, argilites gris-jaunâtre. Bivalves: Alula kutorgai (VERN.), Pseudobakewellia ceratophagaeformis NOIN., Pseudomonotis pemianus MASL. Bryozoans: Permonantis cornutus (GOL.), Paralioclemeta multispinosum MOROZ., Fenestella schurae MOROZ., Polypora kasanensis MOROZ., Dyscritella incrustata MOROZ. Brachiopods: Cancrinella cancrini (VERN.), Odontospirifer subcostatus (NETSCH.). 145. Greyish brown limestones, compact and oolithic, dolomies with gypsum. Calcaires brun-grisâtre, compacts et oolithiques, dolomies avec gypse. Foraminifera: Nodosaria krotowi TSCHERD. Cnidaria: Conularia hollebeni GEIN. Bryozoans: Fistulipora dybowskii GOR., Tabulipora ordinata MOROZ., Discritella incrustata MOROZ. Brachiopods: Aulosteges wangenheimi (VERN.), Stenoscisma superates (VERN.), Orbiculoidea konincki (GEIN.) 146. In the lower part, grey argillites and sandstones with gypseous cement; in the upper part, stratified and oolithic limestones and dolomites locally with gypsum. Dans la partie inférieure, argilites grises et grès à ciment gypseux ; dans la partie supérieure, dolomies et calcaires oolithiques et stratifiés, gypse de place en place. Foraminifera: Nodosaria elabugae TSCHERD., N. hexagona TSCHERD., N. suchonensis K.M. MACL.; Brachiopods: Aulosteges fragilis (NETSCH.), A. horrescens (VERN.). Bryozoans: Pseudobatostomella decora MOROZ., Fenestella permulta MOROZ. Bivalves: Pseudomonotis permianus MASL., Pseudobakewellia certophagaefomis NOIN. 147. Bioclastic, reefal limestones with brachiopods and crinoids, with intercalations of argillites, marls and sandstones. Calcaires récifaux, bioclastiques à brachiopodes et crinoïdes, avec intercalations d’argilites, marnes et grès. Foraminifera: Nodosaria elabugae TSCHERD., N. pugioides ZOL. and IGON., N. netschajewi TSCHERD., N. permiana SPAND., N. krotowi TSCHERD., Fronticularia sp. Brachiopods: Cleiothyridina pectinifera (SOW.), Cancrinella sp. 148. Bioclastic limestones with medium and thin foliaceous stratifications, calcareous marls, dark grey argillites. Calcaires bioclastiques avec stratifications en feuillets fins ou moyens, marnes calcaires, argilites gris foncé. Foraminifera: Nodosaria elabugae TSCHERD., N. netschajewi TSCHERD., N. krotowi TSCHERD., Dentalina sp. Brachiopods: Cancrinella cancrini (VERN.), Globiella hemisphaerium (KUT.). 149. Greenish grey sandstones, limestones with glauconite, mainly quartzose, dark grey argillites sometimes calcareous, dolomites. Grès gris-verdâtre, calcaires à glauconite, principalement quartzeux, argilites gris foncé, quelquefois calcaires, dolomies. Foraminifera: Cornuspira kamae TSCHERD., Nodosaria netschajewi TSCHERD., Frondicularia longissima sintasica IGON. Brachiopods: Cleiothyridina pectinifera (SOW.), Rhynchopora sp. 150. Westwards, sandstones, argillites, grey aleurolites and limestones with brachiopods; eastwards, variegated argillites, aleurolites. Vers l’Ouest, grès, argilites, calcaires et aleurolites grises à brachiopodes ; vers l’Est, argilites bariolées, aleurolites. Gasteropods: Goniasma subangulata (GOL.), Polidevcia kazanensis (VERN.). Bivalves: Pseudobakewellia ceratophagaeformis NOIN., Netschajewia globosa (NSTACH.), Lithophaga consobrina (EICHW.). Brachiopods: Cancrinella cancrini (VERN.), Licharewia rugulata (KUT.), Cleiothyridina pectinifera (SOW.). Ostracods: Amphissites tscherdynzevi POSNER, Healdia subtriangula KOTSCH., Cavellina unica KOTSCH. 151. Sandstones, aleurolites, grey argillites with intercalations of marls and limestones. Grès, aleurolites, argilites vertes avec intercalations de marnes et calcaires. Foraminifera: Nodosaria hexagona (TSCHERD.), N. urmarensis K.M. MACL. Brachiopods: Lingulina semivelata TSCHERD, Lingula orientalis GOL., Globiella hemisphaerium (KUT.), Licharewia stuckenbergi (NETSCH.), Pinegathyris royssiana (KEYS.), Beecheria netschajewi GRIG. Bivalves: Pseudobakewellia ceratophagaeformis NOIN. Bryozoans: Wjatkella wjatkensis (NESCH.). Ostracods: Amphissites tscherdynzevi POSN., Healdia subtriangula KOTSCH. Flora s.l.: Signacularia noinskii ZAL., Odontopteris rossica ZAL., Phylladoderma meridionalis NEUB., Nucicarpus tarlovskiensis ESAUL. 152. Argillites with lignite, limestones. Argilites avec lignite, calcaires. Foraminifera: Nodosaria hexagona (TSCHERD.), N. urmarensis K.M. MACL. Tabulates: Tabulata ordinata MOROZ. Bryozoans: Fenestella permulta MOROZ. Brachiopods: Cancrinella cancrini (VERN.), Globiella hemisphaerium (KUT.), Licharewia rugulata (KUT.), Pinegathyris royssiana (KEYS.), Cleiothyridina pectinifera (SOW.), Baitugania netschajewi GRUNT, Beecheria netschajewi GRIG. Ostracods: Amphissites tscherdynzevi POSN., Cavellina unica KOTSCH. Flora s.l.: Phylladoderma fefiloviana BOGOV, Sylvella alata ZAL. 153. Grey sandstones with charred vegetal, with intercalations of argillites. Grès gris avec végétaux carbonisés, intercalations d’argilites. Brachiopods: Lingula orientalis GOL. 154. Ostracods: Paleodarwinula ufimica (PAL.), P. biriensis (PAL.), P. burajavoensis (PAL.). Flora s.l.: Paracalamites striatus ZAL., Odontopteris rossica ZAL., Phylladoderma septata BOGOV. 155. Ostracods: Suchonella inornata (SPIZH.), Suchonella typica (SPIZH.), Tscherdynzeviana buslukensis KASH. 156. Ostracods: Suchonella stelmachovi SPIZH., S. nasalis SHARAP., Paleodarwinula fragiliformis (KASH.), P. polenovae (BEL.). 157. Ostracods: Paleodarwinula fragiliformis (KASH.), Darwinuloides tscherdynzevi KASH., D. dobrjankaensis KASH. 158. Variegated argillites, aleurolites, sandstones, grey marls, limestones, dolomites with undetermined fresh water ostracods. Argilites bariolées, aleurolites, marnes grises, calcaires, dolomites avec des ostracodes d’eau douce indéterminés. 159. Red conglomerates, sandstones, light grey limestones, marls, dolomites. Conglomérats rouges, grès, calcaires gris clair, marnes et dolomies. Ostracods: Prasuchonella nasalis (SHARAP.), P. belebeica BEL., D. fainae BEL., Darwinuloides edmistonus BEL., D. sentjakensis (SHARAP.) V.P. Gorsky et al. / Geobios 36 (2003) 533Ð558 551

160. Light grey limestones foliated and gritty, dolomites, red conglomerates, sandstones, aleurolites, argillites. Calcaires gris clair graveleux et en plaquettes, dolomies, conglomérats rouges, grès, aleurolites et argilites. Ostracods: Schneideria kazanica KOTSCH., Paleodarwinula alexandrinae (BEL.), P. fainae (BEL.), P. tuimazensis (KOTSCH.). Flora s.l.: Signacularia noinskii ZAL., Odontopteris rossica ZAL. 161. Greenish grey sandstones, conglomerates, red aleurolites, dolomites cavernous and porous. Grès gris-verdâtre, conglomérats, aleurolites rouges, dolomies caverneuses et poreuses. Ostracods: Darwinuloides sentjakensis (SHARAP.), D. edmistonus BEL. 162. Foliated limestones, light and dark grey dolomites with intercalations of grey argillites, marls, lignite, gypsum. Calcaires feuilletés, dolomies gris clair à foncé avec intercalations d’argilites, marnes, lignite et gypse. Ostracods: Cavellina unica KOTSCH., Healdia simplex ROUNDY, Schneideria kazanica KOTSCH.; Bivalves: Liebea hausmanni (GOLDFUSS), Pseudomonotis garforthenis KING. 163. Light grey and greenish sandstones with lenses of conglomerates. Grès gris clair et verdâtres avec lentilles de conglomérats. Foraminifera: Cornuspira megasphaerica GERKE. Ostracods: Cavellina grandis SCHN. 164. Argillites, marls, bioclastic limestones, greenish grey sandstones. Argilites, marnes, calcaires bioclastiques, grès gris-verdâtre. Foraminifera: Nodosaria hexagona (TSCHERD.), Frondicularia longissima (K.M. MACL.), Tetrataxis sp. Ostracods: Cavellina unica KOTSCH., Healdia simplex ROUNDY, Bairdia beedei ULR. and B. Brachiopods: Globiella hemisphaerium (KUT.), Licharewia regulata (KUT.), Dielasma elongata SCHLOTH. 165. Greenish grey argillites, grey platy marls. Argilites gris-verdâtre, marnes grises en plaquettes. Foraminifera: Cornuspira megasphaerica GERKE, C. microsphaerica K.M. MACL., Calcivertella sp., Frondicularia longissima (K.M. MACL.). Ostracods: Graphiodactylus sp., Moorea facilis SCHN., Healdia pseudosimplex KOTSCH., H. postcornuta SCHN., H. subtriangula KOTSCH., Bairdia beedei ULR. and B., Acratia baschkirica KOTSCH. Brachiopods: Licharewia rugulata (KUT.), Cleiothyridina pectinifera SOW., Globiella hemisphaerium (KUT.) 166. Grey and red tarry sandstones with charred vegetal, with intercalations of argillites. Grès gris et rouges bitumineux à végétaux fusinisés, avec intercalations d’argilites. 167. Conglomerates, sandstones, argillites, aleurolites gypseous; in the upper part, limestones and cavernous dolomites. Conglomérats, grès, argilites, aleurolites gypseuses ; dans la partie supérieure, calcaires et dolomies caverneuses. Ostracods: Paleodarwinula martjevi (PAL.), P. sobela (KASH.), Prasuchonella scolia (MAND.), P. stelmarta (KASH.) 168. Limestones, dolomites, marls, argillites, sandstones, conglomerates with gypsum lenses. Calcaires, dolomies, marnes, argilites, grès, conglomérats avec lentilles de gypse. Ostracods: Paleodarwinula burajevoensis (PAL.), P. trita (PAL.), Prasuchonella nasuta KOTSCH., Darwinuloides djurtjuliensis PAL., Sinusuella pergraphica MAND. 169. Conglomerates, sandstones, argillites, aleurolites, dolomites, limestones. Conglomérats, grès, argilites, aleurolites, dolomies et calcaires. Ostracods: Paleodarwinula biriensis (PAL.), P. parphenovi (BEL.), P. forschi (PAL.), Prasuchonella kamyschincaensis PAL. 170. Dolomites, limestones, marls, argillites, sandstones with gypseous cement, aleurolites. Dolomies, calcaires, marnes, argilites, grès à ciment gypseux, aleurolites. Ostracods: Paleodarwinula angusta (MAND.), P. biriensis (PAL.), P. forschi (PAL.)

Table 8 Eastern European basin, Samara and Volgograd (ﬁrst part) belts Bassin Est-Européen, Ceintures de Samara et de Volgograd (première partie) 552 V.P. Gorsky et al. / Geobios 36 (2003) 533Ð558

171. Bivalves: Palaeomutela ovalis AMAL., P. curiosa AMAL., P. plana AMAL., P. obloqua (AMAL.), Palaeanodonta ﬁscheri AMAL.; Ostracods: Suchonellina trapezoida (SHARAP.), S. inornata SPIZH., S. fragilis (SCHNEID.), S.(?) praelongata (ZEK.), S. vladimirinae (BEL.), Suchonella auriculata (SHARAP.), S. grandis (BEL.), Darwinuloidea svijazhicus (SHARAP.), Sinusuella vjatkensis POSNER; Conchostraca: Pseudestheria cf. cicatriosa (NOV.), Ps. cf. elongata (NETSCH.), Ps. cf. lineata (LUTK.), Limnadia cf. maitchatica NOV. Vertebrates s.l.: Chroniosuchus uralensis TVERD., Ch. paradoxus VJUSCH., Dvinosaurus sp., Inostrancevia uralensis TAT., Scutosaurus sp., Chthonosaurus velocidaens VJUSCH., Rhaphanodon ultimus (TCHUD. and VJUSCH.), Rh. tverdochlebovae IVACH., Kotlassia prima AMAL. 172. Bivalves: Palaeomutella obliqua AMAL., P. keyserlingi AMAL., P. ovalis AMAL. Ostracods: Suchonellina inornata SPIZH., S. parallela SPIZH., S. parallela var. typica (LUN.), S. undulata (MISH.), S. spizharskyi (POSNER), S. cultella (MISH.), Prasuchonella stelmachovi (SPIZH.), P. sulacensis (STAROZH.), Suchonella grandis BEL., Darwinuloides buguruslanicus KASH., Permiana oblonga (POSNER). Conchostraca: Limnadia turaica NOV., L. uralensis NOV., Pseudestheria nordvikensis (NOV.), Ps. lineata (LUTK.). Arthropods: Polygrapta aff. chatangensis NOV. Vertebrates s.l.: Chroniosaurus dongusensis TVERD., Dvinosaurus sp., Scutosaurus sp., Microphon exigus IVACH. 173. Amanak formation: red or variegated aleurolites, clays, sandstones, limestones and marls. Formation d’Amanak: aleurolites, argiles, grès, calcaires et marnes rouges ou bariolés. Ostracods: Paleodarwinula elongata (LUN.), P. chramovi (GLEB.), P. torensis (KOTSCH.), Prasuchonella nasalis (SHAR.). 174. Paleodarwinula fragiliformis (KASH.), Placidea lutkevichi (SPIZH.). 175. Bivalves: Palaeomutela extensiva GUS., P. krotovi NETESCH. Conchostraca: Ulugkemia petri NOV., Rossoestheria acutangulata NOV. Vertebrates s.l.: Platyops stuckenbergi TRAUT., Melosaurus kinelensis VJUSCH. 176. Ostracods: Paleodarwinula torensis (KOTSCH)., P. kassini (BEL.), P. perlonga (SHAR.), P. fragilis var. angusta (LUN.), P. chramovi (GLEB.), P. elongata (LUN.), P. fragiliformis (KASH.), P. faba (MISH.), P. imposter (MISH.), Volganella magna SPIZH., Prasuchonella nasalis (SHAR.). Vertebrates s.l.: Platysomus biarmicus EICHW., Kargalichthis efremovi MINICH, Melosaurus vetustus KONZH., Platyops stuckenbergi TRAUT., Doliosauriscus admanteum ORL., Syodon biarmicum KUT., Titanophoneus potens EFR., Zygosaurus lucius EICHW., Discosauricus netscha jewi RJAB. 177. Soksk formation: aleurolites with gypsum, sandstones. Formation de Soksk: aleurolites avec gypse, grès. Ostracods: Paleodarwinula elongata

(LUN.), P. tichonovichi (BEL.), Darwinuloides tscherdynzevi KASH.; Miospore complex I2 (see Table 2). 178. Sosnovsk formation: anhydrite, dolomites, limestones, thin beds of clays, marls, aleurolites; inclusions of salt, gypsum. Formation de Sosnovsk: anhydrite, dolomies, calcaires, lits ﬁns d’argiles, marnes, aleurolites, inclusions de sel et de gypse. Ostracods: Paleodarwinula fainae (BEL.), P. alexandrinae (BEL.), P. varsanoﬁevae (BEL.), Prasuchonella onega (BEL.), P. belebeica (BEL.), P. tichwinskaja (BEL.), Darwinuloides sentjakensis (SHARAP.) 179. “hydrochemistry” formation: anhydrites with rare beds of clayed dolomites, salts, aleurolites, sandstones. Formation “ hydrochimique ”: anhydrites avec rares lits de dolomies argileuses, sels, aleurolites, grès. 180. Ostracods: Paleodarwinula fainae (BEL.), P. alexandrae (BEL.), P. justa (SPIR. and MOLOST.), P. cuneata (KOTSCH.), P. lira (SPIR. and MOLOST.), Prasuchonella belebeica (BEL.), P. tichwinskaja (BEL.). Vertebrates s.l.: Platyops stuckenbergi TRAUT., Tokosaurus perforatus TYERD. and

IVACH., Belebey maximovi TVERD., B. vergrandis IVACH.; Miospore complex I2 (see Table 2)+Granizonospora vulgaris LUBER f. elegans STAN., Vittatina elegans ZAUER, V. scutata STAN., Cordaitina rugulifera (LUBER), C. stenolimbata WAR., Protohaploxypinus perfectus (NAUM.) SAMOIL. 181. Barbashinsk unit: light grey pelitic dolomites; dark grey clays, green-grey sandstones, lenses of gravelles. Unité Barbashinsk : dolomies pélitiques gris clair, argiles gris foncé,grès gris-vert, lentilles de gravelles. Foraminifera: Hyperammina borealis GERKE, Nodosaria noinskyi TSCHERD. Ostracods: Healdia simplex ROUNDY, H. subtriangula KOTSCH. 182. Kamyshlinsk unit: light grey organodetritic limestones; dark grey to black clays; thin bedded marls, rare thin beds of aleurolites and sandstones; some inclusions of anhydrite and salt. Unité Kamyshlinsk : calcaires organodétritiques gris clair, argiles noires à gris foncé, marnes finement litées, rares bancs fins d’aleurolites et de grès, quelques inclusions d’anhydrite et de sel. Foraminifera: Saccammina parvula GERKE, Nodosaria pseudoconcinna K.M. MACL. Brachiopods: Lingula semivelata TSCHERD. Ostracods: Pseudoparaparchites furmidabilis SCHN., Amphissites tscherdyncevi POSNER, Healdia oblonga KOTSCH., Bairdia sobolevensis KHIV. 183. Baytugansk unit: light grey thin grained limestones, clays, marls, dolomites; calcaires à grain fin, argiles, marnes et dolomies gris clair. Foraminifera: Cornuspira microsphaerica GERKE. Ostracods: Amphissites tscherdyncevi POSNER, Actuaria diffusa SCHN., Monoceratina fasgiata KOSCH.

184. Miospore complex I1 (see Table 2)+Remysporites psilopterus LUBER, Azonaletes levis LUBER, Vittatina persecta ZAUER, Cordaitina rotata (LUBER) MEDV., Striatolebachites varius (ZAUER) WAR., Striatohaplopinites prolixus LUBER, Protohaploxypinus tecturatus LUBER 185. Kalinovsk formation: interbedded of light grey limestones, marls, clays; rare beds of aleurolites, sandstones. Formation Kalinovsk : calcaires, marnes et argiles gris clair interstratifiées ; rares lits d’aleurolites et de grès. Brachiopods: Lingula orientalis GOL., Dielasma clongata SCHLOTH. Ostracods: Cavellina unica KOTSCH., Healdia simplex ROUNDY, H. postcornuta SCHN., H. subtriangula KOTSCH., Bairdia beedei ULRICH and BASSLER, Acratia stricta SCHN. Vertebrates s.l.: Phreatophasma aenigmatum EFR., Phreatosuchus qualeni EFR. 186. Ostracods: Paleodarwinula procera (MAND.), P. parphenovae (BEL.), P. abunda (MAND.), P. angusta (MAND.), P. pyriformis (KASH.), Prasuchonella stelmarta (KASH.), Sinusuella pergraphica (MAND.); Miospore complex H (see Table 2)+Acanthotriletes osmundae SAMOIL., Granizono- spora vulgaris WAR., Cirratriradites procumbens (LUBER) WAR. 187. Ostracods: Suchonellina trapezoidea (SHAR.), S. parallela SPIZH., S. undulata MISHINA, Wjatkellina fragilina (BEL.), W. fragilis (SCHN.), W. vladimirina (BEL.), Dvinella ex gr. cyrta (ZEKINA), Suchonella typica SPIZH., Tatariella angulata MISHINA, Placidea lutkevichi (SPIZH.), Tscherdynze- viana pulchra (BEL.); in eastern zone, endemic sculptured cytherides, Tscherdynzeviana pennata KOTSCH., Netschajewiana costata KOTSCH., Kashevarovi- ana delicata MOLOST., Pamilio primaris delicata MOLOST. Bivalves: Palaeomutela inostranzevi AMAL., P. orthodonta AMAL., P. ovalis AMAL., P. plana AMAL., Palaeanodonta fisheri AMAL., P.okensis AMAL., Opokiella tschernyschevi PLOTN. Conchostraca: Limnadia angulidorsa NOVOJ., L. maitschatica NOVOJ., L. (Falsisca) furaica NOVOJ., Megasitum kaljugense NOVOJ., M. vanum NOVOJ. Vertebrates s.l.: Chroniosuchus uralensis TVERD., Kotlassia sp., Raphanodon sp., Scutosaurus sp., Inostrancevia uralensis TAT., Scylacosuchus orenburgensis TAT., Dicynodon cf. amalitzkii SUSCH.; Isadia aristoviensis A. MINICH, Wodnika invicta A. MINICH. 188. Ostracods: Suchonellina inornata SPIZH., S. parallela SPIZH, S. undulata (MISH.), S. daedala (MISH.), S. cultella (MISH.), S. degitalis (MISH.), S. futschiki (KASH.), S. sizharskyi (POSNER), Prasuchonella stelmachovi (SPIZH), P. sulacensis (STAROZ.), Darwinuloides buguruslanicus KASH., Permiana elongata POSNER, P. oblonga POSNER, Tscherdynzeviana pylchra BEL., Tsch. buzulukensis KASH.; endemic cytherides (ostracods) in the eastern zone – Tscherdynzeviana busulucensis KASH., Tsch. pennata KOTSCH., Tsch. abscondita KOTSCH., Netschajewiana costata KOTSCH., Nuguschia sinizyni KOTSCH., Kaschevaroviana delicata MOLOST., K. pectunata MOLOST., Jungumulus mirus MOLOST., Permianella inilaterata MOLOST., Pamilio primaris MOLOST. Bivalves: Palaeomutela inostranzevi AMAL., P. keyserlingi AMAL., Palaenodonta fischeri AMAL., Opokiella ignatjevi GUSEV. V.P. Gorsky et al. / Geobios 36 (2003) 533Ð558 553

Conchostraca: Siberioleaia oblonga (MITSCH.), Hemicycloleaia intermediata (MITSCH.), Pseudostheria cicatricosa (NOJ.). Vertebrates s.l.: Chroniosaurus dongusensis TVERD., Rophanodon tverdochlebovae IVACH., Microphon exiquus IVACH.; Isadia aristoviensis A. MINICH, I. suchonensis A. MINICH, Toyemia tverdochlebovi MINICH; Flora s.l.: Pursongia amalitzkii ZAL., Cordaites clercii ZAL.; Limitisporites sp., Jugasporites sp., Lueckisporites virkkiae POT. and KREMP, Taeniaesporites albertae JANS.

189. Western part: lower half, obliquely layered red sandstones, aleurolites, clays and fine-pebble conglomerates; upper half, red aleurolites, clays, fine and more seldom coarse-grained sandstones; central part: high thickness, aleurolite-clayey composition, conglomerate lens occurrence and widely spread nodular limestones; eastern part: lower half, conglomerate/sandstone sequence with rare interlayers of red-brown clays and marls; upper half, marl-clay-sandstone sequence with interlayers of grey clayey limestones and aleurolites. Partie occidentale : moitié inférieure, grès à stratifications obliques, aleurolites, argiles et conglomérats à galets fins ; dans la moitié supérieure, aleurolites rouges, argiles, grès à grain fin et plus rarement à grain grossier ; partie centrale : grande épaisseur d’aleurolites argileuses, lentilles de conglomérats et calcaires noduleux largement répandus ; partie orientale : moitié inférieure, séquence de conglomérats Ð grès avec rares niveaux de marnes et argiles rouge-brun ; moitié supérieure, séquence, marnes Ð argiles Ð grès avec niveaux de calcaires argileux gris et aleurolites. Ostracods: Paleodarwinula fragiliformis (KASCH.), P. elongata (LUNJAK), P. fragilis var. angusta (SCHN.), P. inornatinae (LUNJAK), P. chramovi (GLEB.), P. torensis (KOTSCH.), P. faba (MISCH.), P. impostor (MISCH.), P. defluxa (MISCH.), P. arida (MOLOST.), P. obvia (MOLOST.), P. gariainovi (MOLOST.), Prasuchonella nasalis (SHARAP.), Darwinuloides buguruslanicus KASCH., Sinusuella ignota SPIZH., Placidea lutkevichi (SPIZH.). Bivalves: Abiella subovata (JONES), Microdantella microdonta (KHALFIN), M. plotnikovskiensis (FED.), Palaemutela doratiformis GUSEV, P. parallelogramma KULEVA, P. gusevi KULEVA, Anthraconaia verneuili (AMAL.), A. aktubensis GUSEV, Anthraconauta (Prokopievskia) pseudophillipsi FEDOTOV. Conchostraca: Pseudostheria angulata (LUTK.), Sphaerestheria belorussica NOVOJ., Triedrolophus tverdochlebovi MOLIN, Pseudostheria otchevi MOLIN, Megasitum jaroslavlense NOVOJ. Vertebrates s.l.: Mnemeiosaurus jubilaevi NOVI., Melosaurus sp., Deuterosaurus sp., Platyops sp., Notosyodon sp. Flora s.l. : Lebachiacites pulcherrimus SAUER, Striatolebachites, Jugasporites sp., Faunipollenites sp., Taeniaesporites albertae JANS., Vittatina striata LUBER f. angusticostata SAUER, Platysaccus?, Alisporites striatohaplopinites. The upper Urzhum horizon is characterized in some localities by endemic Cytherides (ostracods). La partie supérieure de l’horizon Urzhum est caractérisé par des Cythéridés (ostracodes) endémiques : Tscherdynzeviana squamosa KOTSCH., Wetluginella (?) tatarica MOLOST., Kaschevaroviana delicata MOLOST., K. pectunata MOLOST.

190. Western part: lagoonal-continental formations; red, less frequently grey clays, aleurolites, sandstones, and, to a smaller extent, limestones, marls, dolomites; central part: clays, aleurolites and sandstones with rare interlayers of limestones and marls. Eastern part : carbonate-terrigenous sequence composed of interlaid brown, lilac and grey clays, aleurolites, marls, limestones, sandstones / sandstones, gravelstones and conglomerates / monotonous interlayers of concretionary and bedded brown limestones. Partie occidentale : formations lagunaires Ð continentales ; argiles, aleurolites, grès rouges ou plus rarement gris, plus rarement calcaires, marnes et dolomies ; partie centrale : argiles, aleurolites et grès avec rares niveaux de calcaires et de marnes ; partie orientale : séquence carbonatée Ð terrigène, composéedeséries interstratifiées grises, brunes et lilas, d’aleurolites Ð marnes Ð calcaires Ð grès/grès Ð graviers Ð conglomérats / séries monotones de calcaires bruns lités ou en concrétions. Ostracods: Paleodarwinula fainae (BEL.), P. alexandrinae (BEL.), P. irenae (BEL.), P. cuneata (KOTSCH.), P. tichonovichi (BEL.), P. tuimazensis (KOTSCH.), P. varsanofievae (BEL.), P. inornatinaeformis (BEL.), P. chramovella (BEL.), P. aronovae (BEL.), P. persimplex (KOTSCH.), P. procurva (KOTSCH.), Garjainowula lija (SPIR. and MOLOST.), G. ex. gr. gracilis (SPIR. and MOLOST.), Prasuchonella tichwinskaja (BEL.), P. belebeica (BEL.), Darwinuloides sentjakensis (SCHAR.), Placidea ex gr. lutkevichi (SPIZH.), Schneideria kazanica KOTSCH. Bivalves: Palaeanodonta rhomboidea (NETSCH.), Palaemutela celebrata GUSEV, P. quadrata KULEVA, P. novalis NETSCH., P. umbonata (FISCHER), P. krotovi NETSCH, P. olgae GUSEV, P. doratiformis GUSEV. Conchostraca: Palaeolimnadiopsis eichwaldi (NETSCH.), Clyptoasmussia exiqua (EICHW.), Cyclotungusites kazanensis MOL., Ulugkemia borisi NOVOJ., U. orengurgiana MOLIN, U. petri NOVOJ., Sphaerestheria ikensis MOLIN, Pseudestheria kaslanica MOLIN. Vertebrates s.l.: Platyops sp., Deuterosaurus sp., Tapinocephalidae. Flora s.l.: Cordaitina sp., Limitisporites leschikii KLAUS, Gigantosporites hallstathensis KLAUS, Striatites marginalis KLAUS, Striatohaplopinites latissimus LUBER, Striatopi- ceites suchinensis SED., Vittatina striata LUBER, Cyclogranisporites, Acanthotriletes, Azonaletes; Carpolithes sp., Phylladoderma sp., Noeggerathiopsis sp., Lepeophyllum sp., Samaropsis sp., Paracalamites sp., Odontopteris sp., Equisetina sp., Rufloria sp., Taeniopteris sp., Pecopteris sp., Voltzia mamadyschensis ZAL., Samaropsis irregularis NEUB., Psygmophyllum expansum BRONGN., Odontopteris cf. rossica ZAL., Ullmannia biarmica EICHW.

191. Western part: grey sandstones, aleurolites, clays and limestones. Partie occidentale: grès gris, aleurolites, argiles et calcaires. Ostracods: Pseudoparaparchites formidabilis SCHN., Aecuminella bella KHIV., Cornigella valosa KHIV., Kirkbya rara KHIV., Amphissites tscherdynzevi POSNER, Moorea facilis SCHN., Cavellina grandis SCHN., Cavellina unica KOTSCH., Healdia postcornuta SCHN., H. oblonga KOTSCH., Healdia simplex ROUNDY, H. pseudosimplex KOTSCH., H. subtriangula KOTSCH., H. reniformis SCHN., Healdianella vulgata KOTSCH., Cribroconcha urschakensis KOTSCH., Bairdia beedei ULRICH and BASSLER, B. garrisonensis UPSON, B. pompiliodes HARLTON, Acratia baschkirica KOTSCH., Fabalicypris crepidalis KOTSCH., Actuaria diffusa SCHNEIDER, A. secunda KOTSCH., Monoceratina fastigiata KOTSCH., M. faveolata KOTSCH., M. parvula KOTSCH., M. exilis SCHNEIDER, Fascianella notabilis SCHN. Brachiopods: Glomospira sp., Ammodiscus cf. bradynus SPANDEL, Verneuilinoides kasanica UCHARS. Foraminifera: Cornuspira megasphaerica GERKE, C. microsphaerica K.M. MACLAY, Calcivertella kasanica K.M. MACLAY, Nodosaria cushmani PAALZOV, N. forcimeniformis K.M.–MACLAY, N. hexagona (TSCHERD.), N.aff.noinskyi TSCHERD., N. krotovi TSCHERD., Pseudonodosaria lata K.M. MACLAY, Fronticularia longissima (K.M. MACLAY), Fr. geinitzinaeformis (K.M. MACLAY), Geinitzina spandeli TSCHERD., Tetrataxis sp., Tristis permiana GERKE. Brachiopods: Cleiothyridina pectinifera SOW, Cl. semiconcava WAAGEN, Cl. reyssiana KEYS., Stepanoviella hemisphaerium KUTOR., Pr. (Cancrinella) cancrini VERNEUIL, Pr. koninckianus KEYS., Licharewia rugulatia KUTOR., L. curvirostris VERNEUIL, Dielasma elongata SCHLOT., D. elliptica NETSCH., D. angusta NETSCH., Lingula orientalis GOLOW., Aulosteges horrescens VERNEUIL. Bivalves: Leibea hausmanni GOLDF., Allorisma elegans KING, Pseudobakewellia sulcata (GEIN.), Ps. ceratophragaeformis NOINSKI, S. tutchburia tschernyschevi LICH. Central and eastern parts: lagoonal-deltaic and littoral-marine grey-coloured sandstones, aleurolites, clays, marls, limestones and dolomites. Parties centrale et orientale : série grise, laguno-deltaïque et littorale composéedegrès, aleurolites, argiles, marnes, calcaires et dolomies. Ostracods: Paleodarwinula cuneata (KOTSCH.), P. fainae (BEL.), P. irinae (BEL.), Prasuchonella belebeica (BEL.), Darwinuloides cf. sentjakensis (SHARAP.). Bivalves: Paleomutela sintasensis GUSEV, P. ovataeformis GUSEV. Flora s.l.: Pecopteris anthriscifolia ZAL., Sphenophyllum sp., Asterodiscus; Cordaitina uralensis (LUBER) SAMOIL., C. cubrotata (LUBER) SAMOIL.; Ginkgocycadophytus erosus (LUBER) SAMOIL.; Protodiploxypinus elongatus SAMOIL., Protopodocarpus alatus SAMOIL.; Azonalites (Rugosina) tenuis LUBER, A. (Tenuella) levis LUBER; Azonotriletes cf. resistens LUBER, A. osmundes SAMOIL. 554 V.P. Gorsky et al. / Geobios 36 (2003) 533Ð558

192. Western part: red terrigenous rocks: clays, aleurolites, sandstones, rare marls, limestones and dolomites interlayers; intensive gypsification in the lower part. Partie occidentale : roches terrigènes rouges : argiles, aleurolites, grès, rares marnes, calcaires et dolomies interstratifiées ; gypsification intense dans la partie inférieure. Ostracods: Paleodarwinula abunda (MAND.), P. angusta (MAND.), P. parphenovae (BEL.), P. procera (MAND.), P. lancetiformis (KASH.), P. trita (PALANT), P. burjevoensis (PALANT), P. lubimovae (KASH.), Garjainovula lija (SPIR. and MOLOST.), G. kulevae (SPIR. and MOLOST.), G. gracilis (SPIR. and MOLOST.), Prasuchonella stelmarta (KASH.), Pr. kargalensis (KOTSCH.), Pr. kamyschencaensis (PALANT), Darwinuloides djurtjuliensis

PALANT, Sinusuella pergraphica MAND. Bivalves: Paleomutela attenuata GUSEV, P. ovataeformis GUSEV, P. sintasensis GUSEV. Miospores complex H1 +

H2 (see Table 2). Central part: red clays, aleurolites and sandstones; eastern part: red conglomerates with ﬂint pebbles, sandstones, aleurolites and marls with rare thin limestone interlayers; reddish-brown argillites with interlayers of brownish-grey sandstones and grey limestones. Partie centrale : argiles, aleurolites et grès rouges ; partie orientale : conglomérats rouges avec galets de silice, grès, aleurolites et marnes avec rare niveaux ﬁns de calcaires ; argilites brun-rougeâtre avec niveauxdegrès gris-brunâtre et de calcaires gris. Ostracods: Paleodarwinula abunda (MAND.), P. angusta (MAND.), P. lancetiformis (KASH.).

193. Ostracods: Suchonellina inornata SPIZH., Suchonellina parallela SPIZH., S. cf. futschiki (KASH.), S. spizharskyi (POSNER), Prasuchonella stelmachovi (SPIZH.), Darwinuloides svijaazhicus SHARAP., Paleodarwinula elongata (LUN.), Paleodarwinula fragiliformis (KASH.), Suchonella nasalis SHAR.,

194. Ostracods: Amphissites tscherdynzevi POSNER, Bairdia beedei ULRICH and BASSLER, Bairdianella semilunulata KOTSCH., Actuaria diffusa

SCHN.; Miospore complex I1 (see Table 2)+Calamospora sp., Cyclogranisporites osmundae SAMOIL., Apiculatisporites tenuispinosus (WALTZ.) SIV., Cirratriradites procumbens LUBER, Azonaletes tenuis LUBER, Vittatina cicinnata LUBER, V. striata LUBER, Florinites luberae SAMOIL., Cordaitina sp., Platysaccus latus LUBER, Vesicaspora schemeli KL., Falcisporites zapfei POT. and KL., Caytonispollenites sp., Striatohaplopinites perfectus NAUM., Striatopodocarpites tojmensis SED., Lueckisporites sp., Taeniaesporites sp.

195. Miospore complex H (see Table 2)+Punctatostorites sp., Cyclogranisporites polypyranus (LUBER) SIV., Spinosisporites cornutus K.-M., Verrucosisporites grumosus IBR., Cirratriradites procumbens LUBER, Granulatisporites sp., Azonaletes irregulariplicatus SAMOIL., Ginkgocycadophytus sp., Vittatina crassa ZAUER, V. striata LUBER, V. laxa ZAUER, Cordaitina sp., Vesicaspora sp., Alisporites sp., Striatohaplopinites glomus HART., Protohaploxypinus latissimus (LUBER) SAMOIL., Hamiapollenites bullaeformis (SAMOIL.) JANS.

196. Stomochara abrasa SAID., Stm. cybaea KIS., Stm. diserta SAID., Horniella concinna KIS.; Miospore complex J2 (see Table 2)+Caytonipollenites sp., Calamospora sp. Cyclogranisporites polypyrens LUBER, Apiculatisporites sphaericus VIRB., Granulatisporites dubia PRESL., Falcisporites sp., Fusacolpites permicus DIUP., Platysaccus sp., Striatopiceites sp., Lueckisporites virkkiae POT. and KR., Taeniaesporites sp., Vittatina vittifer LUBER.

197. Ostracods: Suchonellina inornata (SPIZH.), S. parallela (SPIZH.), S. stelmachovi (SPIZH.), Darwinuloides tataricus POSNER, D. svijazhicus SHAR.

198. Ostracods: Paleodarwinula fragiliformis (KASH.), P. elongata (LUN.), Suchonellina nasalis (SCHAR.).

199. Ostracods: Amphissites tscherdynzevi POSNER, Healdia simplex ROUNDY, Healdia subtriangula KOTSCH., Healdianella ex gr. osagensis KELLETT, Healdianella ex gr. panda KOTSCH., Healdianella parallela KNIGHT, Healdianella vulgata KOTSCH., Cavellina? cf. unica KOTSCH., Cavellina ex gr. edmistonae (HAR. and LAI.). Brachiopods: Cancrinella cancrini VERN., Beecheria netschaevi GRIG., Cleiothyridina reussiana (KEIS.), C. pectinifera (SOW.). Bivalves: Pseudomonotis elengatula NETSCH., Parallelodon kingi VERN., P. cf. striatus (SCHLOTH.), Edmondia elongata HOUSE, Bakevellia (Bakevellia) ceratophaga (SCHLOTH.). Bryozoans: Dyscritella incrustata MOR. Foraminifera: Palaeonubecularia uniserialis REITL., P. ﬂuxa REITL., Geinitzina pusilla GROZD., G. spandeli TSCHERD., G. cf. postcarbonica SPAND., Nodosarina elabugae TSCHERD., N. netschevi TSCHERD., Spandellina ex gr. cordiformis GERCE; Miospore complex I (see Table 2)+Leiotriletes osmundae SAMOIL., Azonamonoletes marattiformis SAMOIL., Azonaletes tenuis LUBER, Vittatina vittifer LUBER, Ginkgocycadophytus sp., Vitreisporites sp., Florinites sp., Striatopinites dvinensis SED., Protohaplopin- ites tecturatus LUBER, Protodiplopinites silvestritypus SAMOIL., Protocedrus parviextensisaccus SAMOIL.

200. Miospore complex J (see Table 2)+Granulatisporites sp., Punctatisporites sp., Verrucosisporites vetlugencis (MAL.) WAR., Densoisporites nejburgii (SCHULTZ) BALME, Spinosisporites (MAL.) WAR., Vittatina striata LUBER, V. costabilis WILS., Cordaitina sp., Podocarpites alatus LUBER, Gardena- sporites heisseli KL., Striatopodocarpus sp., Hamiapollenites sp., Striatolebacheites sp.

201. Miospore complex J (see Table 2)+Vittatina costabilis WILS., V. vittifer LUBER, V. striata LUBER, Vesicaspora schemeli KL., Vestigisporites minutus KL., Gardenasporites heisseli KL., G. leonardii KL., Falcisporites sp., Podocarpites alatus (LUBER), Striatopodocarpites cancellatus KL., Striatolebacheites sp., Lueckisporites virkkiae POT. and KL., Vitreisporites sp.

202. Miospore complex I2 (see Table 2)+Punctatisporites sp., Raistrickia cardioformis (LUBER), Entylissa sp., Vittatina sp., Lueckisporites virkkiae POT. and KL., Vitreisporites sp., Striatolebacheites sp., Striatopodocarpites ellipticus ZAUER, Acritarches.

203. Ostracods: Paleodarwinula sp., Moorea cf. facilis SCHN.; Miospore complex I1 (see Table 2)+Vittatina costabilis WILS., Entylissa sp., Cordaitina sp., Gardenasporites sp., Platysaccus papilionis POT. and KL., Alisporites sp., Vitreisporites sp., Lueckisporites virkkiae POT. and KL., Striatolebacheites sp., Taeniaesporites noviaulensis LESCHIK, Striatopodocarpites ellipticus ZAUER, S. cancellatus KL., Acritarches

204. Miospore complex H (see Table 2)+Leiotriletes sp., Lycospora sp., Zonomonotriletes granulatus (SAMOIL.) DIBN., Vittatina cincinnata LUBER, V. vittifer LUBER, V. striata LUBER, V. costabilis WILS., Platysaccus sp., Limitisporites sp., Striatohaplopinites micros HART., S. verus EFR., Striatites minor KL., Striatodiplopinites elongatus (LUBER) SAMOIL., Acritarches. V.P. Gorsky et al. / Geobios 36 (2003) 533Ð558 555

Table 9 Eastern European basin, Volgograd (second part) belt Bassin Est-Européen, Ceinture de Volgograd (deuxième partie)

205. Ostracods: Suchonella futschiki (KASH.), Darwinuloides tataricus POSNER, D. swijazhicus SHARAP. 206. Ostracods: Suchonellina futschiki (KASH.) 207. Ostracods: Darwinuloides edmistnus (BEL.) 208. Brachiopods: Cleiothyridina royssiana (KEYS.), Beecheria netschajewi GRIG. Bryozoans: Dyscritella incustata MOROZ. 209. Ostracods: Darwinuloides edmistonae BEL., Darwinuloides sentjakensis (SHAR.), Darwinuloides triangula BEL., Paleodarwinula accomodata (STAR.), Paleodarwinula alexandrinae (BEL.), Paleodarwinula inornatina (BEL.), Paleodarwinula varsanofievae (BEL.), Paleodarwinula vinocurovi (BEL.), Sinusuella ignota (SPIZH.), Suchonella belebeica BEL., Prasuchonella onega (BEL.), Suchonella serpula BEL., Palaeanodonta rhomboidea (NECH.), Palaeomutela cf. umbonata (FISCH.), Palaeomutela cf. vjatkensis GUS., Palaeomutela ex gr. doratioformis GUS., Palaeomutela ex gr. krotovi NECH. 210. Paleodarwinula irinae (BEL.), P. varsanofievae (BEL.), Paleodarwinula accomodata (STAR.), Paleodarwinula ex gr. tichwinskaje (BEL.), Paleodarwinula isetica (STAR.), Paleodarwinula varsanofievae (BEL.), Paleodarwinula. irinae (BEL.), Prasuchonella onega (BEL.), Darwinuloides sentjakensis (SHARAP.), Darwinuloides edmistonae BEL., Darwinuloides triangula BEL., Sinusuella cf. ignota SPRZH., Suchonella belebeica BEL., Suchonella sacmarensis STAR., pelecypods, miospores 211. Ostracods: Paleodarwinula angusta (MAND.), P. procera (MAND.), P. abunda (MAND.), P. sobela (KASH.) 212. Ostracods: Paleodarwinula cf. angusta (MAND.), P. abunda (MAND.), Prasuchonella ex gr. scolia (MAND.) 211 – 212. At Lugovskaya 1 well: Ostracods: Paleodarwinula angusta (MAND.), Paleodarwinula burajevoensis (PAL.), Paleodarwinula cf. pyriformis (KASH.), Paleodarwinula cf. trita (PAL.), Paleodarwinula fainae (BEL.), Paleodarwinula lanzetiformis (KASH.), Paleodarwinula lubimovae (KASH.). Bivalves: Palaeomutela cf. stegocephalum NECH., miospores and conchostraca. 213. Bivalves: Palaeomutela verneuili (AMAL.), P. ovalis AMAL. Ostracods: Suchonellina parallela SPIZH. Flora s.l.: Cladophlebis kamiensis VLAD. 214. Ostracods: Paleodarwinula fragiliformis (KASH.). Flora s.l.: Cladophlebis kamiensis VLAD., Voltzia mamadyschensis ZAL., Cordaicarpus czakaensis ZAL., C. burensis ZAL. 215. Bivalves: Antraconauta uslonensis GUS. Ostracods: Darwinuloides elegantella BEL. 216. Bivalves: Palaeomutela umbonata (FISCHER), P. attenuata GUS., P. pseudoumbonota GUS. Ostracods: Paleodarwinula inornatinaeformis (BEL.), Prasuchonella onega (BEL.), Darwinuloides sentjakensis SHARAP. Flora s.l.: Cladophlebis kamiensis VLAD., Phylladoderma sp. 217. Bivalves: Palaeomutela stegocephalum NETSCH., F. sintasensis GUS., P. ovatiformis GUS., Palaeanodonta (Antraconaia) rhomdoidea (NETSCH.). Ostracods: Paleodarwinula abunda (MAND.), P. sobela (KASH.), P. pergusta (KASH.). Flora s.l.: Knorriopsis sp., Viatcheslavia sp. 556 V.P. Gorsky et al. / Geobios 36 (2003) 533Ð558

Table 10 Eastern European Basin, Pripyat depression. Characteristic fossils of Western European Basin Bassin Est-Européen, dépression de Pripyat. Fossiles caractéristiques du Bassin Ouest-Européen 218. Foraminifera: Nodosaria sp.

219. Foraminifera: Haplophragmoides probata SCHERP., H. lithuanicus UCHAR., Ammobaculites rectus UCHAR., Trochammina aff. pulvilla CRESP. and PARR., Verneuilinoides andrei UCHAR., V. distinctus UCHAR. 220. Foraminifera: Ammodicus bradynius (SPAND.), Haplophragmoides probata SCHERP., H. lithuanicus UCHAR., Trochammina aff. pulvilla CRESP. and PARR., Nodosaria permiana (SPAND.), N. conicodensestriata PAALZ., N. scharpi K.M. MACL., Dentalina permiana JONES, Lunucammina hastata SCHERP., Spanaelina delicatula (JURK.). 221. Bivalves: Liebea squamosa (SOW.), L. Septifer (KING), Schizodus obscurus SOW., Sch. truncatus KING, Sch. schlotheimi (GEIN.) 222. Bivalves: Pseudobakewellia ceratophagaeformis NOIN., Ps. antiquaeformis NOIN., Schizodus obscurus SOW., Sch. schlotheimi (GEIN.), Liebea squamosa (SOW.), Astartella vallisneriana (KING.), Permophorus costatus (BROWN). 223. Bivalves: Pseudobakewellina ceratophagaeformis NOIN. 224. Ostracods: Dorsoobliquella pulchra KNUEPF., Cavellina permiana KOTSCH., Healdia dahlgrueni KROEM. 225. Ostracods: Dorsoobliquella pulchra KNUEPF., Cavellina permiana KOTSCH., Healdia dahlgrueni KROEM., Bairdia plebeia REUSS., B. hisingeri (MUENST.). 226. Ostracods: Polycope perminuta KELLETT, Kirkbya permiana (JONES), Cavellina permiana KOTSCH., Healdia dahlgrueni KROEM., Micro- cheilinella artiensis GUSS., Bairdia plebeia REUSS, Basslerella regularis (RICHTER), Monoceratina parvula KOTSCH. 227. Bryozoans: Acanthocladia anceps (SCHLOTH.), Fenestella retiformis (SCHLOTH.), Kingopora ehrenbergi (GEIN.). 228. Brachiopods: Lingula credneri GEIN., Horridonia horrida (SOW.), Strophalosia morrisiana KING, Pterospirifer alatus (SCHLOTH.), Dielasma elongata (SCHLOTH.). 229. Brachiopods: Lingula credneri GEIN. 230. Flora s.l.: Klausipollenites schaubergeri (POT. and KLAUS.) JANS., Gardenasporites tonsus INOS., KRUZ. and SCHWARZ., Lueckisporites virkkiae POT. and KLAUS., Striatopodocarpites tojmensis SED. 231. Flora s.l.: Lueckisporites virkkiae POT. and KLAUS., Kausipollenites schaubergeri (POT. and KLAUS.) JANS. V.P. Gorsky et al. / Geobios 36 (2003) 533Ð558 557

Table 11 Western European Basin, Brest and Kaliningrad belts Bassin Ouest-Européen, Ceintures de Brest et Kaliningrad

232. Foraminifera: Nodosaria permiana (SPANDEL), N. noinskyi TSCHERD., N. geinitzi REUSS, N.exgr.netschajewi TSCHERD., Geinitzina spandeli TSCHORD., Dentalina ex gr. bradyi SPANDEL, Ammodiscus ex gr. semiconstrictus WATH. Bivalves: Schizodus ex gr. obscurus SOW., Pseudobakewellia ex gr. ceratophagaeformis NOIN. Bryozoans: Fenestella sp. Ostracods: Bairdia sp., Acratia sp. Brachiopods: Horridonia ex gr. horrida (SOW.), Licharevia ex gr. rugulata (KUT.)

233. Foraminifera: Nodosaria sp.

234. Foraminifera: Nodosaria sp. Bivalves: Liebea squamosa (SOW.). Ostracods: Dorsoobliquella pulchra KNUEPF., Cavellina permiana KOTSCH., Healdia dahilgrueni KROEM.

235. Foraminifera: Haplophragmoides probata SCHAR., H. lithuanicus UCHAR., Ammobaculites rectus UCHAR., Trochaminna aff. pulvilla CRESPIN and PARR., Verneuilinoides andrei UCHAR., V. distinctus UCHAR. Bivalves: Liebea squamosa (SOW.), L. septifer (KING.), Schizodus truncatus KING., Sch. obscurus SOW., Sch. schlotheimi (GEIN.). Ostracods: Dorsoobliquella pulchra KNUEP., Cavellina permiana KOTSCH., Healdia dahlgrueni KROEM., Bairdia plebeia REUSS, B. hisingeri (MUNST.)

236. Foraminifera: Ammodiscus bradyinus (SPAND.), Haplophragmoides probata SCHERP., H. lithuanicus UCH., Trochammina aff. pulvilla CRESPIN and PARR., Nodosaria permiana (SPANDEL), N. conicodensestriata PAAL., N. scharpi K.M.–MACL., Dentalina permiana JONES, Lunucammina hastata SCHERP., Spandolina delicatula (JURK.). Bivalves: Pseudobakewallia ceratophagaeformis NOIN., Ps. antiquaeformis NOIN., Schizodus obscurus SOW., Sch. schlotheimi (GOIN.), Lisbea squamosa (SOW.), Astartella vallisneriana (KING.), Permophorus costatus (BROWN.). Bryozoans: Acanthocladia anceps (SCHLOTH.), Fenestella retiformis (SCHLOTH.), Kingopora ehrenbergi (GEIN.). Brachiopods: Lingula credneri GEIN., Horridonia horrida (SOW.), Strophalosia morrisiana KING, Pterospirifer alatus (SCHLOTH.), Dielasma elongata (SCHLOTH.). Ostracods: Polycope perminuta KELLETT, Kirkbya permiana (JONES), Cavellina permiana KOTSCH., Healdia dahlgrueni KROEM., Microchellinella artiensis GUSS., Bairdia plebeia REUSS, Basslerella regularis (RICHTER), Monoceratina parvula KOTSCH. Flora s.l.: Klausipollenites schaubergeri (POT. and KLAUS.) JANS., Gardenasporites tonsus INOS., KRUZ. and SCHWARZ., Lueckisporites virkkiae POT. and KL., Striatopodocarpites tojmensis SOD., Stepanovites inﬂatus (BEND. and STOP.)

237. Bivalves: Pseudobakewellia ceratophagaeformis NOIN. Brachiopods: Lingula credneri GOIN. Flora s.l.: Lueckisporites virkkiae POT. and KL., Klausipollenites schaubergeri (POT. and KL.) JANS. 558 V.P. Gorsky et al. / Geobios 36 (2003) 533Ð558

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