Reptile Family Tree - Peters 2017 1112 Taxa, 231 Characters
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A New Mid-Permian Burnetiamorph Therapsid from the Main Karoo Basin of South Africa and a Phylogenetic Review of Burnetiamorpha
Editors' choice A new mid-Permian burnetiamorph therapsid from the Main Karoo Basin of South Africa and a phylogenetic review of Burnetiamorpha MICHAEL O. DAY, BRUCE S. RUBIDGE, and FERNANDO ABDALA Day, M.O., Rubidge, B.S., and Abdala, F. 2016. A new mid-Permian burnetiamorph therapsid from the Main Karoo Basin of South Africa and a phylogenetic review of Burnetiamorpha. Acta Palaeontologica Polonica 61 (4): 701–719. Discoveries of burnetiamorph therapsids in the last decade and a half have increased their known diversity but they remain a minor constituent of middle–late Permian tetrapod faunas. In the Main Karoo Basin of South Africa, from where the clade is traditionally best known, specimens have been reported from all of the Permian biozones except the Eodicynodon and Pristerognathus assemblage zones. Although the addition of new taxa has provided more evidence for burnetiamorph synapomorphies, phylogenetic hypotheses for the clade remain incongruent with their appearances in the stratigraphic column. Here we describe a new burnetiamorph specimen (BP/1/7098) from the Pristerognathus Assemblage Zone and review the phylogeny of the Burnetiamorpha through a comprehensive comparison of known material. Phylogenetic analysis suggests that BP/1/7098 is closely related to the Russian species Niuksenitia sukhonensis. Remarkably, the supposed mid-Permian burnetiids Bullacephalus and Pachydectes are not recovered as burnetiids and in most cases are not burnetiamorphs at all, instead representing an earlier-diverging clade of biarmosuchians that are characterised by their large size, dentigerous transverse process of the pterygoid and exclusion of the jugal from the lat- eral temporal fenestra. The evolution of pachyostosis therefore appears to have occurred independently in these genera. -
Ontogenetic Change in the Temporal Region of the Early Permian Parareptile Delorhynchus Cifellii and the Implications for Closure of the Temporal Fenestra in Amniotes
RESEARCH ARTICLE Ontogenetic Change in the Temporal Region of the Early Permian Parareptile Delorhynchus cifellii and the Implications for Closure of the Temporal Fenestra in Amniotes Yara Haridy*, Mark J. Macdougall, Diane Scott, Robert R. Reisz Department of Biology, University of Toronto Mississauga, Mississauga, Ontario, Canada * [email protected] a11111 Abstract A juvenile specimen of Delorhynchus cifellii, collected from the Early Permian fissure-fill deposits of Richards Spur, Oklahoma, permits the first detailed study of cranial ontogeny in this parareptile. The specimen, consisting of a partially articulated skull and mandible, exhib- OPEN ACCESS its several features that identify it as juvenile. The dermal tuberosities that ornament the dor- Citation: Haridy Y, Macdougall MJ, Scott D, Reisz sal side and lateral edges of the largest skull of D. cifellii specimens, are less prominent in RR (2016) Ontogenetic Change in the Temporal the intermediate sized holotype, and are absent in the new specimen. This indicates that the Region of the Early Permian Parareptile new specimen represents an earlier ontogenetic stage than all previously described mem- Delorhynchus cifellii and the Implications for bers of this species. In addition, the incomplete interdigitation of the sutures, most notably Closure of the Temporal Fenestra in Amniotes. PLoS ONE 11(12): e0166819. doi:10.1371/journal. along the fronto-nasal contact, plus the proportionally larger sizes of the orbit and temporal pone.0166819 fenestrae further support an early ontogenetic stage for this specimen. Comparisons Editor: Thierry Smith, Royal Belgian Institute of between this juvenile and previously described specimens reveal that the size and shape of Natural Sciences, BELGIUM the temporal fenestra in Delorhynchus appear to vary through ontogeny, due to changes in Received: July 18, 2016 the shape and size of the bordering cranial elements. -
Reptile Family Tree
Reptile Family Tree - Peters 2015 Distribution of Scales, Scutes, Hair and Feathers Fish scales 100 Ichthyostega Eldeceeon 1990.7.1 Pederpes 91 Eldeceeon holotype Gephyrostegus watsoni Eryops 67 Solenodonsaurus 87 Proterogyrinus 85 100 Chroniosaurus Eoherpeton 94 72 Chroniosaurus PIN3585/124 98 Seymouria Chroniosuchus Kotlassia 58 94 Westlothiana Casineria Utegenia 84 Brouffia 95 78 Amphibamus 71 93 77 Coelostegus Cacops Paleothyris Adelospondylus 91 78 82 99 Hylonomus 100 Brachydectes Protorothyris MCZ1532 Eocaecilia 95 91 Protorothyris CM 8617 77 95 Doleserpeton 98 Gerobatrachus Protorothyris MCZ 2149 Rana 86 52 Microbrachis 92 Elliotsmithia Pantylus 93 Apsisaurus 83 92 Anthracodromeus 84 85 Aerosaurus 95 85 Utaherpeton 82 Varanodon 95 Tuditanus 91 98 61 90 Eoserpeton Varanops Diplocaulus Varanosaurus FMNH PR 1760 88 100 Sauropleura Varanosaurus BSPHM 1901 XV20 78 Ptyonius 98 89 Archaeothyris Scincosaurus 77 84 Ophiacodon 95 Micraroter 79 98 Batropetes Rhynchonkos Cutleria 59 Nikkasaurus 95 54 Biarmosuchus Silvanerpeton 72 Titanophoneus Gephyrostegeus bohemicus 96 Procynosuchus 68 100 Megazostrodon Mammal 88 Homo sapiens 100 66 Stenocybus hair 91 94 IVPP V18117 69 Galechirus 69 97 62 Suminia Niaftasuchus 65 Microurania 98 Urumqia 91 Bruktererpeton 65 IVPP V 18120 85 Venjukovia 98 100 Thuringothyris MNG 7729 Thuringothyris MNG 10183 100 Eodicynodon Dicynodon 91 Cephalerpeton 54 Reiszorhinus Haptodus 62 Concordia KUVP 8702a 95 59 Ianthasaurus 87 87 Concordia KUVP 96/95 85 Edaphosaurus Romeria primus 87 Glaucosaurus Romeria texana Secodontosaurus -
The Princeton Field Guide to Dinosaurs, Second Edition
MASS ESTIMATES - DINOSAURS ETC (largely based on models) taxon k model femur length* model volume ml x specific gravity = model mass g specimen (modeled 1st):kilograms:femur(or other long bone length)usually in decameters kg = femur(or other long bone)length(usually in decameters)3 x k k = model volume in ml x specific gravity(usually for whole model) then divided/model femur(or other long bone)length3 (in most models femur in decameters is 0.5253 = 0.145) In sauropods the neck is assigned a distinct specific gravity; in dinosaurs with large feathers their mass is added separately; in dinosaurs with flight ablity the mass of the fight muscles is calculated separately as a range of possiblities SAUROPODS k femur trunk neck tail total neck x 0.6 rest x0.9 & legs & head super titanosaur femur:~55000-60000:~25:00 Argentinosaurus ~4 PVPH-1:~55000:~24.00 Futalognkosaurus ~3.5-4 MUCPv-323:~25000:19.80 (note:downsize correction since 2nd edition) Dreadnoughtus ~3.8 “ ~520 ~75 50 ~645 0.45+.513=.558 MPM-PV 1156:~26000:19.10 Giraffatitan 3.45 .525 480 75 25 580 .045+.455=.500 HMN MB.R.2181:31500(neck 2800):~20.90 “XV2”:~45000:~23.50 Brachiosaurus ~4.15 " ~590 ~75 ~25 ~700 " +.554=~.600 FMNH P25107:~35000:20.30 Europasaurus ~3.2 “ ~465 ~39 ~23 ~527 .023+.440=~.463 composite:~760:~6.20 Camarasaurus 4.0 " 542 51 55 648 .041+.537=.578 CMNH 11393:14200(neck 1000):15.25 AMNH 5761:~23000:18.00 juv 3.5 " 486 40 55 581 .024+.487=.511 CMNH 11338:640:5.67 Chuanjiesaurus ~4.1 “ ~550 ~105 ~38 ~693 .063+.530=.593 Lfch 1001:~10700:13.75 2 M. -
HOVASAURUS BOULEI, an AQUATIC EOSUCHIAN from the UPPER PERMIAN of MADAGASCAR by P.J
99 Palaeont. afr., 24 (1981) HOVASAURUS BOULEI, AN AQUATIC EOSUCHIAN FROM THE UPPER PERMIAN OF MADAGASCAR by P.J. Currie Provincial Museum ofAlberta, Edmonton, Alberta, T5N OM6, Canada ABSTRACT HovasauTUs is the most specialized of four known genera of tangasaurid eosuchians, and is the most common vertebrate recovered from the Lower Sakamena Formation (Upper Per mian, Dzulfia n Standard Stage) of Madagascar. The tail is more than double the snout-vent length, and would have been used as a powerful swimming appendage. Ribs are pachyostotic in large animals. The pectoral girdle is low, but massively developed ventrally. The front limb would have been used for swimming and for direction control when swimming. Copious amounts of pebbles were swallowed for ballast. The hind limbs would have been efficient for terrestrial locomotion at maturity. The presence of long growth series for Ho vasaurus and the more terrestrial tan~saurid ThadeosauTUs presents a unique opportunity to study differences in growth strategies in two closely related Permian genera. At birth, the limbs were relatively much shorter in Ho vasaurus, but because of differences in growth rates, the limbs of Thadeosau rus are relatively shorter at maturity. It is suggested that immature specimens of Ho vasauTUs spent most of their time in the water, whereas adults spent more time on land for mating, lay ing eggs and/or range dispersal. Specilizations in the vertebrae and carpus indicate close re lationship between Youngina and the tangasaurids, but eliminate tangasaurids from consider ation as ancestors of other aquatic eosuchians, archosaurs or sauropterygians. CONTENTS Page ABREVIATIONS . ..... ... ......... .......... ... ......... ..... ... ..... .. .... 101 INTRODUCTION . -
The First Possible Choristoderan Trackway from the Lower
www.nature.com/scientificreports OPEN The frst possible choristoderan trackway from the Lower Cretaceous Daegu Formation of South Korea and its implications on choristoderan locomotion Yuong‑Nam Lee1*, Dal‑Yong Kong2 & Seung‑Ho Jung2 Here we report a new quadrupedal trackway found in the Lower Cretaceous Daegu Formation (Albian) in the vicinity of Ulsan Metropolitan City, South Korea, in 2018. A total of nine manus‑pes imprints show a strong heteropodous quadrupedal trackway (length ratio is 1:3.36). Both manus and pes tracks are pentadactyl with claw marks. The manus prints rotate distinctly outward while the pes prints are nearly parallel to the direction of travel. The functional axis in manus and pes imprints suggests that the trackmaker moved along the medial side during the stroke progressions (entaxonic), indicating weight support on the inner side of the limbs. There is an indication of webbing between the pedal digits. These new tracks are assigned to Novapes ulsanensis, n. ichnogen., n. ichnosp., which are well‑matched not only with foot skeletons and body size of Monjurosuchus but also the fossil record of choristoderes in East Asia, thereby N. ulsanensis could be made by a monjurosuchid‑like choristoderan and represent the frst possible choristoderan trackway from Asia. N. ulsanensis also suggests that semi‑aquatic choristoderans were capable of walking semi‑erect when moving on the ground with a similar locomotion pattern to that of crocodilians on land. South Korea has become globally famous for various tetrapod footprints from Cretaceous strata1, among which some clades such as frogs2, birds3 and mammals4 have been proved for their existences only with ichnological evidence. -
University of Birmingham the Earliest Bird-Line Archosaurs and The
University of Birmingham The earliest bird-line archosaurs and the assembly of the dinosaur body plan Nesbitt, Sterling; Butler, Richard; Ezcurra, Martin; Barrett, Paul; Stocker, Michelle; Angielczyk, Kenneth; Smith, Roger; Sidor, Christian; Niedzwiedzki, Grzegorz; Sennikov, Andrey; Charig, Alan DOI: 10.1038/nature22037 License: None: All rights reserved Document Version Peer reviewed version Citation for published version (Harvard): Nesbitt, S, Butler, R, Ezcurra, M, Barrett, P, Stocker, M, Angielczyk, K, Smith, R, Sidor, C, Niedzwiedzki, G, Sennikov, A & Charig, A 2017, 'The earliest bird-line archosaurs and the assembly of the dinosaur body plan', Nature, vol. 544, no. 7651, pp. 484-487. https://doi.org/10.1038/nature22037 Link to publication on Research at Birmingham portal Publisher Rights Statement: Checked for eligibility: 03/03/2017. General rights Unless a licence is specified above, all rights (including copyright and moral rights) in this document are retained by the authors and/or the copyright holders. The express permission of the copyright holder must be obtained for any use of this material other than for purposes permitted by law. •Users may freely distribute the URL that is used to identify this publication. •Users may download and/or print one copy of the publication from the University of Birmingham research portal for the purpose of private study or non-commercial research. •User may use extracts from the document in line with the concept of ‘fair dealing’ under the Copyright, Designs and Patents Act 1988 (?) •Users may not further distribute the material nor use it for the purposes of commercial gain. Where a licence is displayed above, please note the terms and conditions of the licence govern your use of this document. -
Mesozoic Marine Reptile Palaeobiogeography in Response to Drifting Plates
ÔØ ÅÒÙ×Ö ÔØ Mesozoic marine reptile palaeobiogeography in response to drifting plates N. Bardet, J. Falconnet, V. Fischer, A. Houssaye, S. Jouve, X. Pereda Suberbiola, A. P´erez-Garc´ıa, J.-C. Rage, P. Vincent PII: S1342-937X(14)00183-X DOI: doi: 10.1016/j.gr.2014.05.005 Reference: GR 1267 To appear in: Gondwana Research Received date: 19 November 2013 Revised date: 6 May 2014 Accepted date: 14 May 2014 Please cite this article as: Bardet, N., Falconnet, J., Fischer, V., Houssaye, A., Jouve, S., Pereda Suberbiola, X., P´erez-Garc´ıa, A., Rage, J.-C., Vincent, P., Mesozoic marine reptile palaeobiogeography in response to drifting plates, Gondwana Research (2014), doi: 10.1016/j.gr.2014.05.005 This is a PDF file of an unedited manuscript that has been accepted for publication. As a service to our customers we are providing this early version of the manuscript. The manuscript will undergo copyediting, typesetting, and review of the resulting proof before it is published in its final form. Please note that during the production process errors may be discovered which could affect the content, and all legal disclaimers that apply to the journal pertain. ACCEPTED MANUSCRIPT Mesozoic marine reptile palaeobiogeography in response to drifting plates To Alfred Wegener (1880-1930) Bardet N.a*, Falconnet J. a, Fischer V.b, Houssaye A.c, Jouve S.d, Pereda Suberbiola X.e, Pérez-García A.f, Rage J.-C.a and Vincent P.a,g a Sorbonne Universités CR2P, CNRS-MNHN-UPMC, Département Histoire de la Terre, Muséum National d’Histoire Naturelle, CP 38, 57 rue Cuvier, -
Live Birth in an Archosauromorph Reptile
ARTICLE Received 8 Sep 2016 | Accepted 30 Dec 2016 | Published 14 Feb 2017 DOI: 10.1038/ncomms14445 OPEN Live birth in an archosauromorph reptile Jun Liu1,2,3, Chris L. Organ4, Michael J. Benton5, Matthew C. Brandley6 & Jonathan C. Aitchison7 Live birth has evolved many times independently in vertebrates, such as mammals and diverse groups of lizards and snakes. However, live birth is unknown in the major clade Archosauromorpha, a group that first evolved some 260 million years ago and is represented today by birds and crocodilians. Here we report the discovery of a pregnant long-necked marine reptile (Dinocephalosaurus) from the Middle Triassic (B245 million years ago) of southwest China showing live birth in archosauromorphs. Our discovery pushes back evidence of reproductive biology in the clade by roughly 50 million years, and shows that there is no fundamental reason that archosauromorphs could not achieve live birth. Our phylogenetic models indicate that Dinocephalosaurus determined the sex of their offspring by sex chromosomes rather than by environmental temperature like crocodilians. Our results provide crucial evidence for genotypic sex determination facilitating land-water transitions in amniotes. 1 School of Resources and Environmental Engineering, Hefei University of Technology, Hefei 230009, China. 2 Chengdu Center, China Geological Survey, Chengdu 610081, China. 3 State Key Laboratory of Palaeobiology and Stratigraphy, Nanjing Institute of Geology and Palaeontology, CAS, Nanjing 210008, China. 4 Department of Earth Sciences, Montana State University, Bozeman, Montana 59717, USA. 5 School of Earth Sciences, University of Bristol, Bristol BS8 1RJ, UK. 6 School of Life and Environmental Sciences, The University of Sydney, Sydney, New South Wales 2006, Australia. -
Difficulties with the Origin of Dinosaurs: a Short Comment on the Current Debate
REVIEW ARTICLE Difficulties with the origin of dinosaurs: a short comment on the current debate MATTHEW G. BARON BPP University, 144 Uxbridge Road, London W12 8AA, UK E-mail: [email protected] Abstract: The origin and early evolutionary history of the dinosaurs is a topic that has recently gone through a period of renewed interest and academic debate. For 130 years, one way of classifying the various dinosaur subgroups persisted as the accepted model, with increasing levels of research in the past quarter-century also providing evidence for the hypothesis that dinosaur origination occurred in the Southern Hemisphere, particularly in South America. It is, after all, from within the Late Triassic strata of countries like Argentina and Brazil that we get some of the very best early dinosaur specimens; many of these specimens are the earliest known representatives of some of the major dinosaur subgroups, such as the theropods and sauropodomorphs. However, some recent analyses have brought about a shift in terms of what is currently accepted and what is now disputed regarding the origin of dinosaurs – the Southern Hemisphere origination hypothesis was questioned (although this was based upon observations and not with quantitative analysis techniques), as has the shape of the dinosaur tree. Responses to the new hypothesis were numerous and robust, and new analyses further supported a Southern Hemisphere point of origin. Whilst the interrelationships between the major dinosaur clades remains to be fully resolved, the current data does seem to comprehensively answer the question of where the dinosaurs first originated. However, it is arguable whether or not the current data that is being used in such palaeobiogeographical analyses is sufficient to provide a meaningful answer to the question of where specifically the dinosaur clade first appeared. -
Final Copy 2019 10 01 Herrera
This electronic thesis or dissertation has been downloaded from Explore Bristol Research, http://research-information.bristol.ac.uk Author: Herrera Flores, Jorge Alfredo A Title: The macroevolution and macroecology of Mesozoic lepidosaurs General rights Access to the thesis is subject to the Creative Commons Attribution - NonCommercial-No Derivatives 4.0 International Public License. A copy of this may be found at https://creativecommons.org/licenses/by-nc-nd/4.0/legalcode This license sets out your rights and the restrictions that apply to your access to the thesis so it is important you read this before proceeding. Take down policy Some pages of this thesis may have been removed for copyright restrictions prior to having it been deposited in Explore Bristol Research. However, if you have discovered material within the thesis that you consider to be unlawful e.g. breaches of copyright (either yours or that of a third party) or any other law, including but not limited to those relating to patent, trademark, confidentiality, data protection, obscenity, defamation, libel, then please contact [email protected] and include the following information in your message: •Your contact details •Bibliographic details for the item, including a URL •An outline nature of the complaint Your claim will be investigated and, where appropriate, the item in question will be removed from public view as soon as possible. This electronic thesis or dissertation has been downloaded from Explore Bristol Research, http://research-information.bristol.ac.uk Author: Herrera Flores, Jorge Alfredo A Title: The macroevolution and macroecology of Mesozoic lepidosaurs General rights Access to the thesis is subject to the Creative Commons Attribution - NonCommercial-No Derivatives 4.0 International Public License. -
First Amphibian Ichthyosaur Fossil Found
First amphibian ichthyosaur fossil found An ancient marine reptile with seal-like flippers may have been adapted to life on the land as well as in the sea, scientists believe. The 250-million-year-old creature is the first amphibious ichthyosaur known. Its relatives were dolphin-like creatures that swam in the oceans at the time of the dinosaurs. They are thought to have had terrestrial ancestors, but previously no fossils had come to light marking the transition of ichthyosaurs from land to sea. “Now we have this fossil showing the transition,” said lead scientist Professor Ryosuke Motani, from the University of California at Davis who reported the discovery in the journal Nature. At 1.5 feet long, Cartorhynchus lenticarpus was also the smallest known ichthyosaur. Its fossil remains found in Anhui Province, China, date from the start of the Triassic period about 248 million years ago. As well as big flippers, Cartorhynchus had flexible wrists which would have been essential for movement on the ground. While most ichthyosaurs have long beak-like snouts, the new specimen possessed a short nose that may have been adapted to suction feeding. Its body also contained thicker bones than other ichthyosaurs. This supports the theory that most marine reptiles that left the land first grew heavier to help them swim through rough coastal waves. The animal lived about 4 million years after the worst mass extinction in history, shedding light on how long it took for life on Earth to recover. The Permian-Triassic extinction, known as the “Great Dying”, wiped out 96% of all species and may have been linked to global warming.