Reptile Family Tree - Peters 2015 Distribution of Scales, Scutes, Hair and Feathers Fish scales 100 Ichthyostega Eldeceeon 1990.7.1 Pederpes 91 Eldeceeon holotype Gephyrostegus watsoni Eryops 67 Solenodonsaurus 87 Proterogyrinus 85 100 Chroniosaurus Eoherpeton 94 72 Chroniosaurus PIN3585/124 98 Seymouria Chroniosuchus Kotlassia 58 94 Westlothiana Casineria Utegenia 84 Brouffia 95 78 Amphibamus 71 93 77 Coelostegus Cacops Paleothyris Adelospondylus 91 78 82 99 Hylonomus 100 Brachydectes Protorothyris MCZ1532 Eocaecilia 95 91 Protorothyris CM 8617 77 95 Doleserpeton 98 Gerobatrachus Protorothyris MCZ 2149 Rana 86 52 Microbrachis 92 Elliotsmithia Pantylus 93 Apsisaurus 83 92 Anthracodromeus 84 85 Aerosaurus 95 85 Utaherpeton 82 Varanodon 95 Tuditanus 91 98 61 90 Eoserpeton Varanops Diplocaulus Varanosaurus FMNH PR 1760 88 100 Sauropleura Varanosaurus BSPHM 1901 XV20 78 Ptyonius 98 89 Archaeothyris Scincosaurus 77 84 Ophiacodon 95 Micraroter 79 98 Batropetes Rhynchonkos Cutleria 59 Nikkasaurus 95 54 Biarmosuchus Silvanerpeton 72 Titanophoneus Gephyrostegeus bohemicus 96 Procynosuchus 68 100 Megazostrodon Mammal 88 Homo sapiens 100 66 Stenocybus hair 91 94 IVPP V18117 69 Galechirus 69 97 62 Suminia Niaftasuchus 65 Microurania 98 Urumqia 91 Bruktererpeton 65 IVPP V 18120 85 Venjukovia 98 100 Thuringothyris MNG 7729 Thuringothyris MNG 10183 100 Eodicynodon Dicynodon

91 Cephalerpeton 54 Reiszorhinus Haptodus 62 Concordia KUVP 8702a 95 59 Ianthasaurus 87 87 Concordia KUVP 96/95 85 Edaphosaurus Romeria primus 87 Glaucosaurus Romeria texana Secodontosaurus 86 Protocaptorhinus 75 72 98 Dimetrodon 72 Paracaptorhinus Eocaptorhinus Sphenacodon 100 97 Captorhinus 91 Labidosaurus

Mycterosaurus FMNH UC169 Saurorictus 71 Heleosaurus 90 77 Limnoscelis Mesenosaurus SGU 104V/1558 87 100 Tseajaia Mesenosaurus PIN 3706/15 Tetraceratops 94 53 72 Mesenosaurus PIN 158/1 Orobates Mesenosaurus PIN 3586/8a 60 92 Oradectes 53 97 Archaeovenator 95 Silvadectes 55 Diadectes 97 Orovenator Colobomycter 82 Pyozia 84 Procolophon 100 Broomia 86 69 96 Scolaparia NSM 99 GF82.1 82 Milleropsis BPI-720 90 Leptopleuron 85 Erpetonyx 99 Pentaedrusaurus Hypsognathus Eudibamus 100 Milleretta RC14 100 Aphelosaurus Milleretta RC70 92 Petrolacosaurus 65 Oedaleops Araeoscelis 77 95 Eothyris Ennatosaurus Spinoaequalis Youngina? 93 Casea 87 Galesphyrus 98 Cotylorhynchus 63 94 Youngina BPI 3859 81 Feeserpeton 85 Acerosodontosaurus dorsal scutes 95 Australothyris 72 100 Thadeosaurus Eunotosaurus 98 100 Tangasaurus Acleistorhinus Hovasaurus 95 83 Delorhynchus Claudiosaurus 87 Adelosaurus 100 Atopodentatus 96 Bolosaurus 83 100 Largocephalosaurus Belebey Sinosaurosphargis Stephanospondylus 96 82 81 Arganaceras Anarosaurus Anthodon 61 90 Sclerosaurus Majiashanosaurus 81 59 Elginia Turtle 99 Palatodonta Meiolania Paraplacodus 100 Proganochelys scales/scutes 87 100 Placodus inexpectatus 79 79 Odontochelys 96 88 Placodus gigas Marine taxa Trionyx 72 99 100 99 Henodus Kayentachelys Phonodus 90 Chelonia 88 dorsal scutes 88 Placochelys 58 55 Macrochelys temminckii 85 Cyamodus 75 Terrapene carolina 100 Pelomedusa Pachypleurosaurus Foxemys 75 Cartorhynchus 66 Qianxisaurus 87 Macroleter 81 Keichousaurus 95 Hanosaurus Emeroleter 65 97 Romeriscus Lariosaurus 100 93 Corosaurus Lanthanosuchus 53 Nothosaurus 83 78 75 Simosaurus 99 75 Nyctiphruretus holotype Anningsaura Nyctiphruretus PIN 4659/1 Pistosaurus 68 97 Owenetta rubidgei 100 Plesiosaurus 96 95 Owenetta kitchingorum 100 Trinacromerum Candelaria 75 100 100 Brachauchenius Sauropareion Kronosaurus 93 Barasaurus 83 Kitchingnathus Psilotrachelosaurus 69 Coletta Paliguana Serpianosaurus 87 Sophineta 100 Concavispina 79 Santaisaurus 100 94 Xinpusaurus kohi 76 77 Xinpusaurus suni 79 Thalattosaurus 79 Endennasaurus 97 Saurosternon 100 Clarazia 94 90 Palaegama 85 Coelurosauravus Askeptosaurus 99 90 Lanthanolania 94 Anshunsaurus 54 Kuehneosaurus 100 Miodentosaurus Vancleavea 62 100 Eusaurosphargis 100 94 Icarosaurus 100 74 Xianglong Helveticosaurus dermal scutes Vancleavea

91 Megachirella PIMUZ AIII 0192 99 Pleurosaurus SMF R 4710 100 Marmoretta 94 Stereosternum 87 100 Gephyrosaurus Lepidosaur 100 Brazilosaurus holotype 88 100 Ankylosphenodon 94 Mesosaurus 90 Paleopleurosaurus Wumengosaurus Heleosuchus scales 94 82 Eohupehsuchus Planocephalosaurus 100 Hupehsuchus 77 Kallimodon 94 Parahupehsuchus Sphenodon Grippia 88 100 Brachyrhinodon 100 Utatsusaurus 96 88 Clevosaurus Chaohusaurus Sphenotitan 64 Qianichthyosaurus 53 80 71 Leptosaurus 100 Ichthyosaurus Sapheosaurus 92 85 Ophthalmosaurus 61 Noteosuchus Phalarodon Trilophosaurus 53 96 Mixosaurus Mesosuchus 85 95 96 Cymbospondylus Priosphenodon 97 100 Rhynchosaurus articeps Youngina SAM K7710 98 Bentonyx Youngina BPI 375 Hyperodapedon 86 85 AMNH 9520 9502 Prolacerta 90 80 Protorosaurus Tijubina Jaxtasuchus 77 Homoeosaurus macrodactylus 82 97 Boreopricea Huehuecuetzpalli 74 74 Azendohsaurus 95 Jesairosaurus Pamelaria Hypuronector 83 Vallesaurus 84 60 Megalancosaurus Youngina TM 3603 90 64 Macrocnemus BES SC 111 Youngopsis rubidgei RC90 Macrocnemus T2472 73 Youngopsis kitchingi TM1490 71 Dinocephalosaurus Youngoides RC91 91 64 Langobardisaurus 80 Youngina AMNH 5561 99 Tanytrachelos Youngoides UC1528 76 52 Tanystropheus k and q 99 Cosesaurus Proterosuchus BPI 1 4016 99 Kyrgyzsaurus Fenestrasaur 72 93 Sharovipteryx 54 Proterosuchus NMQR 1484 Proterosuchus RC96 94 Longisquama extradermal membranes MPUM6009 pterosaur 74 59 100 Chasmatosaurus QR 880C.500 Chasmatosaurus SAM PK 11208 54 Chasmatosaurus TM 201 95 Bavarisaurus Elaphrosuchus RC59 Lacertulus 52 Youngina BPI 2871 68 80 Daohugou lizard IVPP V13747 Proterosuchus SAMPKK10603 73 96 75 Meyasaurus 56 Doswellia Carusia 89 Ikechosaurus 60 Hoyalacerta 97 63 90 Simoedosaurus 99 Homoeosaurus solnhofensis 100 Champsosaurus IVPP V14386 Pararchosaur 100 98 Cteniogenys Dalinghosaurus Lazarussuchus 61 99 Hyphalosaurus dorsal scutes 77 100 MFSN 19235 100 Monjurosuchus GMV21678 Monjurosuchus holotype 100 Scandensia 92 99 Diandongosuchus 96 Teraterpeton 95 Mesorhinosuchus 100 Paleorhinus 98 Euposaurus 100 52 Parasuchus Iguana Chanaresuchus 65 Draco volans 83 82 Proterochampsa PVSJ 77 Trioceros 100 83 Proterochampsa PVSJ606 Phrynosoma 87 Pseudochampsa PVSJ 567 84 Gualosuchus 92 Cerritosaurus 70 Calanguban 92 Tropidosuchus PVL 4601 100 Macrocephalosaurus 92 Tropidosuchus PVL 4602 84 Tianyusaurus Lagerpeton 75 100 Eolacerta Tupinambis Acanthodactylus Nundasuchus 61 Liushusaurus 60 Euparkeria Osmolskina Tasmaniosaurus 98 Revueltosaurus Euarchosaur Tchingisaurus 95 92 87 Fugusuchus Gekko smithii 94 100 Garjainia expanded armor 58 Lialis 99 95 100 Erythrosuchus 78 97 Gekko gecko 99 Luperosuchus Eublepharis Shansisuchus

70 100 Riojasuchus Ardeosaurus Ornithosuchus 77 Eichstaettisaurus schroederi 81 Euarchosaur 78 Eichstaettisaurus gouldi 80 Jucaraseps 73 Vjushkovia dorsal scutes 79 Prestosuchus Aphanizocnemus 73 80 Adriosaurus 84 Batrachotomus 96 72 Pontosaurus 90 Saurosuchus 83 Smok 97 70 Dinilysia 92 Teratosaurus 97 88 Pachyrhachis 73 Sikannisuchus Boa Postosuchus 83 Loxocemus 82 92 Xenopeltis Anilius 98 91 Arizonasaurus 96 Cylindrophis 99 56 Yarasuchus 81 Uropeltis Qianosuchus 95 Anomochilus 96 99 Ticinosuchus Leptotyphlops 87 Aeotosauroides Euarchosaur 98 Aetosaurus Stagonolepis expanded armor

Yabeinosaurus IVPP V13284 Decuriasuchus Bahndwivici FMNH PR2260 88 Aigialosaurus 62 100 75 Turfanosuchus 94 Tethysaurus Poposaurus 72 52 Tylosaurus Sacisaurus TA1045 Silesaurus 94 Estesia 100 96 Lotosaurus Varanus komodensis 99 98 99 99 Shuvosaurus Varanus griseus 52 Effigia Shinisaurus 92 52 91 Ophisaurus Gracilisuchus Cryptolacerta 97 Saltopus 60 Scleromochlus 91 Heloderma 51 Lanthanotus 72 Terrestrisuchus 93 62 SMNS12591skull 83 Anniella 83 SMNS 12352 Gobiderma 89 Litargosuchus 79 Chalcides guentheri Erpetosuchus Chalcides ocellatus 63 90 100 Gymnophthalmus 89 60 Hesperosuchus Vanzosaura Pedeticosaurus 86 56 100 Yonghesuchus Crocodilian Sirenoscincus 71 88 Dromicosuchus Sineoamphisbaena 76 92 Sichuanosuchus Crythiosaurus Protosuchus expanded armor 96 Spathorhynchus 93 Dibamus 96 Sphenosuchus Tamaulipasaurus 81 67 Dibothrosuchus 82 Bipes 100 92 66 Teleosaurus Amphisbaena 71 Metriorhynchus 97 Isisfordia 91 Crocodylus niloticus 98 Simosuchus 58 Araripesuchus 100 Caipirasuchus Baurusuchus

Lewisuchus 51 Saltoposuchus Euarchosaur 77 87 Junggarsuchus Pseudhesperosuchus 54 PVL 4597 dorsal scutes Trialestes 54 Herrerasaurus

91 Tawa 95 98 Segisaurus 96 95 Marasuchus 98 Procompsognathus 95 Coelophysis Sinocalliopteryx 97 91 100 Aurornis Archaeopteryx Feathers 99 100 Cathayornis 100 Gallus Struthio

Eoraptor Pampadromaeus 94 Leyesaurus

60 Saturnalia 89 97 90 Pantydraco 94 92 Efraasia Massospondylus kaalae 97 Daemonosaurus Jeholosaurus 93 Haya 99 Pisanosaurus Dinosaur 98 Scelidosaurus 67 Emausaurus Heterodontosaurus expanded armor 84 86 Dryosaurus 81 Hexinlusaurus 98 Yinlong 82 Lesothosaurus 98 Agilisaurus 97 Scutellosaurus