Floristics of the Upper Walker River, California

Volume 43 Number 1 Article 8

1-31-1983

Floristics of the upper Walker River, California

Matt Lavin University of Nevada, Reno

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Recommended Citation Lavin, Matt (1983) "Floristics of the upper Walker River, California," Great Basin Naturalist: Vol. 43 : No. 1 , Article 8. Available at: https://scholarsarchive.byu.edu/gbn/vol43/iss1/8

This Article is brought to you for free and open access by the Western North American Naturalist Publications at BYU ScholarsArchive. It has been accepted for inclusion in Great Basin Naturalist by an authorized editor of BYU ScholarsArchive. For more information, please contact [email protected], [email protected]. FLORISTICS OF THE UPPER WALKER RIVER, CALIFORNIA AND NEVADA

Matt Lavin'

Abstract.— A checklist of the vascular flora of the upper Walker River is presented. Listed are 1078 taxa from this 4000 km^ area. The upper Walker River encompasses a portion of the boundary between the Intermountain and Sierra Nevada floristic regions, and hence displays much floristic diversity within a relatively small area. Due to its location along the east slope of the Sierra Nevada, the Walker River drainage is unique in that it contains much elevational variation eastward into the Intermountain Region. This elevational extension is due to the presence of large mountain ranges including the Sweetwater Mountains, the Bodie Hills, and the Wassuk Range. As a result of this elevational variation, there is much overlapping of the two floristic regions. Additionally, the easternmost exposures of the Sierran granodiorites occur within the Walker River basin and may enhance the eastward migration of Sierran plants. The 90 percent floristic similarity (Sorenson's) between the Sweetwater Mountains, lying to the east of the Sierra, and the east slope of the Sierra Nevada (within the Walker River drainage) indicates the Sweetwaters to be more affiliated with the Sierran flora instead of the Intermountain flora.

The upper Walker River drainage includes dominates. This vegetation has also been an area of approximately 4000 km 2, located briefly described by Billings (1951). The at the north end of Mono Coimty, California, Sweetwater Mountains, situated between the the southern ends of Douglas and Lyon coun- Intermountain and Sierran floristic regions, ties, Nevada, and the western edge of Miner- display characteristics of both. al County, Nevada. The western boundary of The vascular flora of the upper Walker this drainage generally follows the crest of River is modestly represented in literature. the Sierra Nevada from the Conway Summit- Works such as Hinton (1975), Reveal (1968), Virginia Lakes area, northward to the Topaz Cox (1972), Reveal and Ertter (1980), Hard- Lake-Monitor Pass area. The eastern bound- ham and True (1972), Strother (1974), Munz ary is delimited by the Bodie, Masonic, and (1968), Bameby (1964), Dempster and Ehren- Pine Grove hills and the southwest slopes of dorfer (1965), Halse (1981), and others cite the Pine Nut Range. The Sweetwater Moim- specific collections made within this area. tains and the Wellington hills lie between Davis (1979) compiled a plant list and keys to these hills and the Sierra Nevada. the plants occurring on the east central East slope Sierran vegetation present here Sierra (Owens Lake to Lake Tahoe) and asso- has been generally described by Billings ciated desert ranges. This work was based (1951) and Rundel et al. (1977). The area is solely on distributions given in existing floras. dominated by mixed conifer forests composed Sharsmith (1940) includes the Walker Riv- of Pinus jeffreyi and Abies concolor at the er portion of the Sierra Nevada as the north- low elevations, and Abies magnifica, Tsuga ernmost boundary of the Sierra alpine floris- mertensiana, Pinus monticola, and P. albi- tic region. This area includes Leavitt Peak caulis at the higher elevations. The alpine south to Dunderberg Peak. Thome (1982) de- vegetation of the Sierra is uniquely adapted fines the upper Walker River basin as the for extreme svmimer drought (Chabot and northern boundary of the transmontane Cali- Billings 1972). fornian floristic region. To the east of the Sierra, Intermountain Major and Taylor (1977) conducted a vege- vegetation (high elevation sagebrush steppe tation study of the alpine zone of the Sweet- situated above woodlands composed of Pinus water Mountains. They list 43 species from monophylla and Juniperus osteosperma) pre- the area. Taylor (1977) indicates a 40-50

'Department of Biology, University of Nevada, Reno, Nevada 89557. Present address: P.O. Box 13494, Reno, Nevada 89507.

93 .

94 Great Basin Naturalist Vol. 43, No. 1 percent floristic similarity (Sorenson's) be- Methods tween the Carson Pass area of the Sierra Ne- vada and the Sweetwater Mountains. Since a The checklist (Table 1) was developed 50 percent turnover rate in plant species oc- from collections made during the course of curs every 650 km along the Cascade-Sierran this study: July 1979 to August 1982. Addi- recorded from the axis (Taylor 1977), the Walker River portion tional collections were of Nevada, of the Sierra must have a very high degree of herbaria at the University Reno New York Botanical Garden similarity with Carson Pass, 60 km to the (RENO), The (NY), and the California Academy of Sciences north. An interpretation of this would mean (CAS). Collections from the upper Walker the Walker River portion of the Sierra has River made by Arnold Tiehm and Margaret approximately a 50 percent floristic sim- Williams, Reno, Nevada; Steve Wharff, ilarity to the Sweetwaters. Tonopah, Nevada; Frank Smith, Smithfield, Bell (1980) described the alpine flora of the Utah; Dennis Breedlove, CAS; Joe Robertson, Wassuk Range, located in the lower Walker E. F. Kleiner, Tom Lugaski, Pat and Ham River drainage just west of Walker Lake (50 Vreeland, H. N. Mozingo, and Fred Ryser, all of the Sweetwater Mountains). She km east from the University of Nevada, Reno, were lists 70 vascular plants from the alpine flora also recorded in this checklist. Approximately of this region. Bell has also recently com- 3850 numbers were recorded. Distributional pleted a study of the alpine flora of the information contained within the checklist Sweetwater Mountains, but this information was continually refined while in the field. is unavailable. Various documentary works, mentioned in Messick (1982) completed a flora of the the introduction, were used to determine Bodie Hills, and the results of this work are those plants that have a probable distribution incorporated in Table 1 within the upper Walker River drainage, but

The upper Walker River drainage is the were not observed during this study. region of several type collections. The Sweet- Almost all of the taxa listed can be found water Mountains are the type locale for on deposit at the University of Nevada, Reno, Cordylanthus ramosus ssp. setosus, Senecio herbarium and the Toiyabe National Forest pattersonensis and Draba lemmonii var. in- Supervisor's Office, Reno, Nevada. crassata. In the Bodie Hills, we have the type The purposes of the checklist (Table 1) are locale for Draba quadricostata, Pinus mon- to both document the flora present in the up- ophylla, Arabis bodiensis (see Rollins 1982), per Walker River and to document the geo- Streptanthus oliganthus, and Phacelia mon- graphical and altitudinal distribution of each taxon within this area. Nomenclature gener- oensis. A. bodiensis, S. pattersonensis, D. lem- ally follows Kartesz Kartesz monii var. incrassata, and D. quadricostata and (1980). are endemic to the Walker River drainage. The Wellington Hills are the type locale Results and Discussion for Astragalus oophorus var. lavinii, and So- nora Pass for Raillardella argentea, Cymop- Table 1 lists 1078 taxa of vascular plants terus cinerarius, and Wyethia mollis. "A dry from the upper Walker River. The upper near Sonora Pass" is the type rocky mountain Walker River drainage is unique in that it ex- location for Astragalus platytropis, A. lentigi- tends much elevational variation of the east nosus var. ineptus, and A. whitneyi. Bameby slope of the Sierra Nevada eastward into the suggests that the type locale for these three Intermountain Region, due to the presence Astragali may be the Sweetwater Moujitains. and close proximity of such large mountain However, during the course of this in- ranges as the Sweetwater Mountains, the vestigation, all three, including A. platy- Bodie Hills, and the Wassuk Range. Along

tropis, were found on Leavitt and Emma any floristic boundary there is bound to be Peak of the Sierra Nevada. Therefore, the some overlap of unique plant species or char- type locale given by Gray could have possi- acteristic vegetation of one flora into anoth- bly referred to the Sierra instead of the er. However, many plants having their center Sweetwaters. of distribution in the Intermountain Region January 1983 Lavin: Walker River Floristics 95 also have outlying populations in the mon- Penstemon bridgesii Phoenicaulis cheiranthoides tane environments of the Sierra Nevada. This Phacelia humilis is discussed Taylor for phenomenon by (1976) Phlox covillei the Carson Pass area of the Sierra. He attri- Plagiobothrys hispidus butes the occurrence of many Intermountain P. kingii var. harknessii Prunus andersonii plants on the east slope of the Sierra to Xero- Purshia tridentata thermic invasion. A list of these plants at the Pinus monophylla headwaters of the Walker River follow: Ribes velutinum Rosa woodsii var. ultramontana Agoseris glauca var. monticola Scrophularia desertorum Allium biceptrum Senecio canus

Allium parvum S. pattersonensis

Arabis bodiensis S. spartioides A. pulchra var. pulchra Sisyrinchium halophilum Artemisia arbuscula Sphaeromeria cana Aster ascendens Stephanomeria spinosa Astragalus platytropis Streptanthus oliganthus A. iodanthus Tetradymia canescens Amelanchier utahensis Thelypodium crispum Antennaria dimorpha Zigadenus paniculatus Balsamorhiza sagittata Calyptridium roseum These 77 plant taxa represent approx- Cercocarpus ledifolius imately 10 percent of the flora present on the Cirsium utahense east slope of the Sierra within the Walker Cheilanthes gracillima River drainage. This is below Taylor's Chenopodium overi (1976) Chorizanthe brevicomu var. spathulata estimate of 20 percent for the east slope Chrysothamnus nauseosus ssp. albicaulis within the Carson River drainage just to the C. viscidiflorus north. However, when distribution records Crepis acuminata become more complete, it would not be sur- C. modocensis ssp. subacaulis Castilleja linariifolia prising to find 20 percent of the Sierran flora Cordylanthus ramosus ssp. setosus within the Walker River basin being com- Cryptantha circumscissa posed of Intermountain elements. Cryptantha echinella The Xerothermic climate may also have Erigeron aphanactis been responsible for the northern migration E. nevadincola E. breweri var. porphyreticus of Amelancheir pallida var. covillei, Cea- E. eatonii ssp. plantagineus nothus greggii var. vestitus, Cercocarpus ledi- Eriogonum elatum folius var. intricatus, Ivesia purpurascens ssp. E. microthecum var. ambiguum congdonis, Cryptantha confertiflora, Phacelia E. ovalifolium var. nevadense jonesii. E. ivrightii var. subscaposum peirsoniana, and Plagiobothrys The Galium multiflorum Walker River drainage may be the north- Gilia leptantha ssp. salticola ernmost location for these plants. {Cryp- Grayia spinosa tantha confertiflora has been fovmd just to Haplopappus acaulis the north in the Carson River drainage.) Heuchera duranii Hydrophyllum capitatum var. alpinum It is documented that the eastward migra- Leptodactylon pungens tion of Sierran plant species into the Inter- Linanthus nuttallii mountain Region is small relative to the Lupinus caudatus ssp. caudatus westward migration of Rocky Mountain L. nevadensis et al. 1978). Lomatium nevadense var. nevadense plants into this region (Harper Lomatium nevadense var. parishii However, the Walker River drainage has Melica striata many Sierra or Pacific cordilleran plant spe- Mentzelia congesta cies occurring well into the Intermountain Mimulus densus Region, some of which occur as far east as M. rubellus Navarretia breweri Masonic Mountain or the Wassuk Range. A Opuntia polyacantha var. rufispina list of these plants follows: Poa nevadensis var. juncifolia Agropyron pringlei P. nevadensis var. nevadensis Allium campanulatum Paeonia brownii 96 Great Basin Naturalist Vol. 43, No. 1

Allophyllum gilioides their ranges far into the Intermountain Re- A. violaceum gion. These plants, also found extending east- Amelanchier pallida within the Walker River basin, include Anelsonia eurycarpa ward Angelica lineariloba Artemisia rothrockii, Carex helleri, Silene sar- Arabis inyoensis gentii, Astragalus whitneyi (see Reveal 1979) A. platysperma var. howellii and Astragalus purshii var. tinctus (see Bam- Arnica nevadensis eby 1964). Silene sargentii has been listed as Astragalus kentrophyta var. danaus endemic to the Sierra Nevada. However, the A. lentiginosus var. ineptus A. purshii var. lectulus type collection, as given by Hitchcock and Calochortus leichtlinii Maguire (1947), is from Table Mountain of Carex tahoensis the Monitor Range in central Nevada. nevadensis Chaenactis The eastward distribution of these Sierran Chrysothamnus parryi ssp. monocephalus plants may be enhanced by the very eastern Claytonia nevadensis Cryptantha glomeriflora exposures of the Sierran granodiorites. These Cryptantha nubigena granites can be found as far east as the Was- Cymopteris cinerarius suk Range (Bateman 1967). However, the Draba lemmonii (var. incrassata) most likely factor enhancing the migration of D. oligosperma var. subsessilis these Sierran plants is the abundance of mon- D. stenoloba var. ramosa habitats eastward into Erigeron petiolaris tane and alpine found E. pygmaeus the Intermountain Region. The floristic com- Eriogonum rosense ponents of the Sweetwater Moujitains point Gentianopsis hohpetala to this eastward migration. Gentiana newberryi The Sweetwaters, situated between the Geum canescens Sierra and Intermountain floristic regions, are Galium hypotrichium ssp. hypotrichium Haplopappus apargioides regarded as belonging to the Intermountain Hieracium hcrridum flora (Cronquist et al. 1972). Sierra conifer Ivesia lycopodioides forests are extensive on the Sweetwaters, es- I. purpurascens ssp. congdonis pecially toward the southern end. The tim- Juniperus occidentalis ssp. australis vegetation is dominated solely by Kalmia microphylla berline Ledum glandulosum var. californica Pinus albicaulis, a Sierran characteristic. Leucophysalis nana Using the information provided in Table 1, Lupinus andersonii Sorenson's index of similarity (Billings 1978) L. caudatus ssp. montigenus between the Sierra and Sweetwaters can be L. confertus determined with regard to floristic elements L. hypolasius L. meionanthus in both the alpine and the montane conifer L. caudatus ssp. montigenus forests. These are calculated to be 90 and 93 L. sellulus var. lobbii percent, respectively. This indicates the L. tegeticulatus (breweri bryoides) Sweetwater Mountains to be more affiliated, Luzula divaricate floristically, with the Sierra than previously Mimulus coccineus Penstemon davidsonii var. davidsonii thought. Pinus jeffreyi However, it might be considered that the P. monticola east slope of the Sierra, within the Walker Polygonum douglasii var. latifolia River drainage, being influenced by Inter- Raillardella argentea mountain vegetation, is bound to yield a high Rhamnus rubra ssp. rubra index of similarity to the Sweetwaters. In Scirpus dementis Senecio fremontii var. occidentalis other words, the whole of the upper Walker

S. scorzonella could be considered "Intermountain" with Sisyrinchium idahoense var. occidentale regard to vegetational composition. This is Symphoricarpos parishii reinforced by the fact that the Walker River Trifolium andersonii ssp. andersonii portion of the Sierra does lack typical "un-

Additionally, work done by Goodrich derstory" brush vegetation that is found just (1981) in central Nevada has shown that sev- to the north in the Carson River drainage. eral other Pacific cordilleran plants extend Arctostaphylos patula, Ceanothus cordulatus January 1983 Lavin: Walker River Floristics 97

and C. prostratus have not been found here sharing with me their intimate knowledge of (or are at least not abundant) and A. neva- the Intermountain and Sierran floras, and to densis was foujid only twice in very small the curators at NY and CAS for allowing ac- populations. Instead, the dominant brush cess to their herbaria. cover consists of Purshia tridentata, Cea- nothus velutinus, Artemisia tridentata ssp. Literature Cited vaseyana, and Symphoricarpos oreophilus. Barneby, R. In comparing all the upper Walker River C. 1964. Atlas of North American Astra- galus. New York Botanical Gardens. 2 Vols. flora with the flora of central Nevada (Good- Bateman, p. C. 1967. The Sierra Nevada Batholith. Sci- rich 1981), both floras being just about equal ence 158: 1407-1417. in number of taxa, approximately a 50 per- Bell, K., and R. Johnson. 1980. Alpine flora of the cent similarity can be determined. The Wassuk Range, Mineral County, Nevada. Madroiio 27: 25-35. Sweetwaters could, therefore, be easily con- Billings, W. D. 1951. Vegetation zonation in the Great sidered as part of the east slope Sierran flora. Basin of western North America. Pages 101-122 Aside from the high indices of similarity, the in Les Bases e regeneration do la des zones Sweetwaters are dominated by many Sierra arides. International Colloquium. Intemat. Union or Pacific cordillera plant species; Lupinus of Bio. Sci. Ser. B.9. 1978. Alpine phytogeography across the Great hypolasius, Chrysothamnus parryi ssp. mon- Basin. Great Basin Nat. Mem. 2: 105-117. ocephalus, and var. Draba oligosperma sub- Chabot, B. F., and W. D. Billings. 1972. Origins and sessilis at the highest elevations, and Pinus ecology of the Sierran alpine flora and vegeta- jeffreyi and P. contorta var. murrayana at the tion. Ecol. Monogr. 42: 163-199. Cox, B. 1972. Biosystematics of Lupinus lepidus-L. lower elevations. J. caespitosus complex. Unpublished dissertation. Univ. of Missouri, Columbia. 443 pp. Cronquist, A., A. N. Holmgren, Holmgren, and J. Acknowledgments Reveal. 1972. Intermountain flora. Vol. 1. Haf- ner. New York. 584 pp. This study was funded by the U.S. Forest Davis, B. 1979. Draft: Plant list and keys. East Side Cen- tral Sierra and associated desert Ranges. U.S. Service, Toiyabe National Forest. I am very Forest Service, Toiyabe National Forest, Reno, grateful to them for this and for their cooper- Nevada. 124 pp. ation. Thanks to Sue Sullivan, U.S. Forest Dempster, L. T., and F. Ehrendorfer. 1965. Evolution Service, the checklist (Table 1) and its trans- of the Galium multiflorum complex in western fer to Brigham Young University Press was North America. II. Critical taxonomic revision. Brittonia 17: 289-334. made possible. For their help with identi- Goodrich, S. 1981. A floristic study of central Nevada. fication of plant specimens, I am indebted to Unpublished thesis, Brigham Young Univ. 400 Warren Wagner (Botrychium), Lincoln Con- pp. stance (Apiaceae), Ted Barkley (Senecio), Al- Halse, R. R. 1981. Taxonomy of Phacelia Sect. Miltitzia mut Jones (Aster), Guy Nesom (Erigeron), (Hydrophyllaceae) Madrono 28: 121-132. Hardham, C. B., and G. H. True. 1972. Malacothrix tor- Gerald Ownbey (Cirsium and Argemone), reyi (Compositae) in California. Madrono 21: 535. Reed Rollins (Brassicaceae), Arthur Cronquist Harper, K. T., D. C. Freeman, W. K. Ostler, and L. (miscellaneous), Alfred Schuyler (aquatic C. Klikoff. 1978. The flora of the Great Basin plants), Gary Wallace (Pyrola), Rupert Barn- mountain ranges: diversity, sources and dispersal ecology. Great Basin Nat. Mem. 2: 81-103. eby (Fabaceae), Richard Halse (Phacelia), HiNTON, W. F. 1975. Systematics of the Calyptridium Duane Atwood (Phacelia), Douglass Hender- umbellatum complex (Portulacaceae). Brittonia son (Sisyrinchium), Paul Fryxell (Malvaceae), 27: 197-208. David Boufford (Circaea), Alva Day (Polemo- Hitchcock, C. L., and B. Maguire. 1947. A revision of niaceae), Lauramae Dempster (Galium), Har- the North American species of Silene. Univ. of Washington Pub. Biol. 13: 1-73. lan Lewis (Gayophytum), Lawrence Heckard R. Kartesz, J., AND Kartesz. 1980. A synonymized (Castilleja, Cordylanthus, Orthocarpus), Fred- checklist of the vascular flora of the United rick Meyer (Valeriana), John T. Howell States, Canada and Greenland. Univ. of North (Carex and miscellaneous), Margaret Wil- Carolina Press. Chapel Hill. 500 pp. Lavin, M. 1981. Floristics of the headwaters of the liams (miscellaneous), Arnold Tiehm (mis- Walker River, California and Nevada. Unpub- cellaneous), and Ken Genz (miscellaneous). I lished thesis. Univ. of Nevada, Reno. 141 pp. am especially grateful to Thomas How- 1977. Alpine. Pages John Major, J., and D. W. Taylor. Major, eds.. ell, Margaret Williams and Arnold Tiehm for 601-675 in M. G. Barbour and J. 98 Great Basin Naturalist Vol. 43, No. 1

Terrestrial vegetation of California. John Wiley RuNDEL, P. W., D. J. Parsons, and D. T. Gordon. 1977. Interscience, New York. Montane and subalpine vegetation of the Sierra Messick, T. 1982. The flora and phytogeography of the Nevada and Cascade ranges. Pages 559-599 in M. Barbour and Major, eds., Terrestrial Bodie Hills of Mono County, CA and Mineral G. J. vegeta- County, NV. Unpublished thesis, Humboldt State tion of California. John Wiley Interscience, New York. Univ. Sharsmith, C. W. 1940. A contribution to the history of MuNZ, P. A., AND D. Keck. 1968. A California flora. the alpine flora of the Sierra Nevada. Unpub- Univ. of California Press. Berkeley. 1681 pp. and lished dissertation. Univ. of California, Berkeley. supplement. 274 pp. Reveal, L. 1968. Notes of Eriogonum— IV. A revision J. L. 1974. of Strother, J. Taxonomy Tetradymia (Com- of the Eriogonum deflexum complex. Brittonia positae: Senecioeae). Brittonia 26: 177-202. 20: 13-33. Taylor, D. W. 1976. Disjunction of Great Basin plants 1979. Biogeography of the Intermountain Re- in the northern Sierra Nevada. Madrono 23: gion. Mentzelia 4: 1-92. 301-310. L., B. 1980. Noteworthy collec- Reveal, J. and Ertter. 1977. Floristic relationships along the Cascade- tions. Madrono 27: 142. Sierran axis. Amer. Midi. Natur. 97: 333-349. Rollins, R. C. 1982. Studies on Arahis (Cruciferae) of Thorne, R. F. 1982. The desert and other transmontane Western North America II. Contr. Gray Her- plant communities of southern California. Aliso barium of Harvard. No. 212:111-114. 10:219-257.

Table 1. Checklist of the vascular flora of the upper Walker River. For the columns under the heading of "AREA":N = Sierra Nevada; S= Sweetwater Mountains; W = Wellington Hills; M = Masonic Hills, Bodie Hills, Pine Grove Hills, and the southwest slopes of the Pine Nut Range. For the columns under the heading of "HABITAT": 1 = riparian; 2= low elevation sagebrush-grass zone; 3=pinyon-juniper woodland; 4 = high elevation sagebrush -grass zone; 5 = Jeffrey pine-white fir forests; 6 = red fir forests; 7 = lodgepole pine forests; 8 = whitebark pine forests; 9= alpine zone (see Billings 1951, Lavin 1981, for a general description of these). An X indicates the plant has been observed in the field or by herbarium specimen; an O indicates the plant has a probable distribution within the area due to information obtained in the literature. AREAS HABITATS N S W M 123456789

Adiantaceae - pteridophyta Aspidotis densa (Brack.) Lellinger X X Cheilanthes gracillima D.C. Eat. X X Cryptogramma crispa (L.) R. Br. ex Hook. ssp. acrostichoides (R. Br.) Hulten Pellaea breweri D.C. Eat. X X bridgesii Hook. X X Aspleniaceae Athyrium distentifolium Tausch ex Opiz var. americanum (Butters) Boivin Cystopteris fragilis (L.) Bemh. Woodsia oregana D.C. Eat. scopulina D.C. Eat. Dennstaedtiaceae Pteridium aquilinum (L.) Kuhn var. pubescens Underwood Equisetaceae Equisetum arvense L. X X hyemale L. var. affine (Engelm.) A. A. Eat. laevigatum A. Braun January 1983 Lavin: Walker River Floristics 99

Table 1 continued. AREAS HABITATS

N S W M 12 3 4 5 7 8 9 ISOETACEAE Isoetes bolanderi Engelm. var. bolanderi XXX Marsileaceae Marsilea vestita Hook. & Grev. Ophioglossaceae Botrychium lunaria (L.) Sw. var. minganese (Victorin) Dole simplex E. Hitchcock

Selaginellaceae Selaginella watsonii Underwood X X XXX

CUPRESSACEAE - PINOPHYTA Juniperus communis L. X X occidentalis Hook. ssp. australis Vasek X osteosperma (Torr.) Little Ephedraceae Ephedra

nevadensis S. Wats. viridis Coville

Pinaceae Abies concoloT (Gord. & Glend.) Hildebr. magnifica A. Murr. Pinus albicaulis Engelm. X X contorta Dougl. ex Loud. var. murrayana (Grev. & Balf.) Engelm. XX o flexilis James XX o jeffreyi Grev. & Balf. X X X X monophylla Torr. & Frem. X X X X monticola Dougl. ex D. Don X X Tsuga mertensiana (Bong.) Carr. X

AcERACEAE - ANTHOPHYTA Acer glabrum Torr. var. torreyi (Greene) Smiley Alismataceae Sagittaria cuneata Sheldon Amaranthaceae Amaranthus alhus L. X X blitoides S. Wats. X X X X retroflexus L. X X Apiaceae Angelica breweri Gray X lineariloba Gray XX X 100 Great Basin Naturalist Vol. 43, No. 1

Table 1 continued. AREAS HABITATS

N S W M 123456789

Berula erecta (Huds.) Coville X Cicuta douglasii (DC.) Coult. & Rose X X Conium maculatum L. X X Cymopterus cinerarius Gray X

globosus (S. Wats.) S. Wats. panamintensis Coult. & Rose var. panamintensis Heracleum lanatum Michx. Ligusticum grayii Coult. & Rose Lonuitiiim dissectum (Nutt.) M. & C. var. multifidum (Nutt.) M. & C. foeniculaceum (Nutt.) Coult. & Rose ssp. macdougalii (Coult. & Rose) Theobald

nevadense (S. Wats.) Coult. & Rose var. nevadense var. parishii (Coult. & Rose) Jepson plummerae (Coult. & Rose) Coult. & Rose var. sonnei (Coult. & Rose) Jepson Osmorhiza chilensis Hook. & Am. occidentalis (Torr. & Gray) Torr. Perideridia

bolanderi (Gray) A. Nels. & J.F. Macbride ssp. bolanderi lemmonii (Coult. & Rose) Chuang & Const. parishii (Coult. & Rose) A. Nels. ssp. latifolia (Gray) Chuang & Const. Podistera

nevadensis (Gray) S. Wats. Pteryxia terebinthina (Hook.) Coult. & Rose var. californica (Coult. & Rose) Mathias Sphenosciadium capitellatum Gray Apocynaceae Apocynum androsaemifolium L. ssp. pumilum (Gray) Boivin X medium Greene ASCLEPIADACEAE Asclepias cryptoceras S. Wats. ssp. cryptoceras ssp. davisii (Woods.) Woods. fascicularis Dene. speciosa Torr. ASTERACEAE Achillea millefolium L. var. alpicola (Rydb.) Garrett var. lanulosa (Nutt.) Piper January 1983 Lavin: Walker River Floristics 101

Table 1 continued. AREAS HABITATS

N S 102 Great Basin Naturalist Vol. 43, No. 1

Table 1 continued. AREAS habitats

N S W M 12 3 4 5 6 7

Bahamorhiza hookeri (Hook.) Nutt. XXX XX X sagittata (Pursh) Nutt. X X X X XX X Bidens cemua L. Brickellia grandiflora (Hook.) Nutt. var. petiolaris Gray microphylla (Nutt.) Gray X X X oblongifolia Nutt. var. linifolia (D.C. Eat.) B.L. Robins. XXX X X Chaenactis alpigerm C.W. Sharsmith X X douglasii (Hook.) Hook. & Am. var. rubricaulis (Rydb.) Ferris X X nevadensis (Kellogg) Gray xantiana Gray Chrysothamnus nanseosus (Pall.) Britton ssp. consimilis (Greene) Hall & Clem, ssp. albicaulis (Nutt.) Hall & Clem, ssp. hololeucus (Gray) Hall & Clem. parryi (Gray) Greene ssp. monocephalus (Nels. & Kenn.) Hall & Clem, ssp. nevadensis (Gray) Hall & Clem. viscidiflorus (Hook.) Nutt. ssp. puberulus (D.C. Eat.) Hall & Clem, ssp. viscidiflorus Cichorium intybus L. Cirsium andersonii (Gray) Petrak congdonii Moore & Frankton eatonii (Gray) B.L. Robins.

pastoris J.T. Howell tioganum (Congd.) Petrak utahense Petrak vulgare (Savi) Tenore Conyza canadensis (L.) Cronquist var. glabrata (Gray) Cronquist Crepis acuminata Nutt. ssp. acuminata intermedia Gray modocensis Greene ssp. subacaulis (Kell.) Babcock & Stebbins nana Richards. ssp. ramosa Babcock & Stebbins ssp. nana occidentalis Nutt. ssp. pumila (Rydb.) Babcock & Stebbins ssp. occidentalis ssp. conjuncta (Jepson) Babcock & Stebbins runcinata (James) Torr. & Gray ssp. hallii Babcock & Stebbins Dugaldia hoopesii (Gray) Rydb. January 1983 Lavin: Walker River Floristics 103

Table 1 continued. AREAS HABITATS NSWM 123456789

Eatonella nivea (D.C. Eat.) Gray X X X X Erigeron aphanactis (Gray) Greene var. aphanactis X X barbellulatus Greene bloomeri Gray var. bhameri breweri Gray var. breweri var. porphyretictis (M.E. Jones) Cronquist clokeyi Cronquist compositus Pursh var. glabratus Macoun coulteri Porter divergens Torr. & Gray eatonii Gray ssp. plantagineus (Greene) Cronquist lonchophyllus Hook. nevadincola Blake peregrinus (Pursh) Greene

ssp. callianthemus (Greene) Cronquist var. angustifolius (Gray) Cronquist

ssp. callianthemus (Greene) Cronquist var. hirsutus Cronquist petiolaris Greene pygmaeus (Gray) Greene tener (Gray) Gray vagus Payson Eriophyllum lanatum (Pursh) Forbes var. integrifolium (Hook.) Smiley Eupatorium occidentale Hook. Glyptopleura marginata D.C. Eat. Gnaphalium palustre Nutt. microcephalum Nutt. var. thermale (E. Nels.) Cronquist Grindelia squarrosa (Pursh) Ehinal var. squarrosa Gutierrezia sarothrae (Pursh) Britt. & Rusby Haplopappus acaulis (Nutt.) Gray apargioides Gray bloomeri (Hook.) Gray lanceolatus (Hook.) Torr. & Gray var. lanceolatus macronema Gray racemosus (Nutt.) Torr.

ssp. glomeratus (Nutt.) Hall suffruticosus (Nutt.) Gray uniflorus (Hook.) Torr. & Gray ssp. uniflorus 104 Great Basin Naturalist Vol. 43, No. 1

Table 1 continued. AREAS habitats NSW January 1983 Lavin: Walker River Floristics 105

Table 1 continued. AREAS HABITATS N 106 Great Basin Naturalist Vol. 43, No. 1

Table 1 continued. AREAS habitats

N S W M 123456789

jamesii (Torr.) Payson var. abortiva (Greene) Payson nuhigena (Greene) Payson X pterocarya (Torr.) Greene var. pterocarya torreyana (Gray) Greene var. torreyana var. pumila (Heller) Johnston watsonii (Gray) Greene

sp. nov. Hackelia floribunda (Lehm.) Johnston micrantha (Eastw.) J.L. Gentry Lappula redowskii (Homem.) Greene var. redowskii Mertensia oblongifolia (Nutt.) G. Don var. nevadensis (A. Nels.) L.O. Williams Pectocarya setosa Gray Plagiobothrys hispidus Gray jonesii Gray

kingii (S. Wats.) Gray var. harknessii (Greene) Jepson var. kingii scouleri (Hook. & Am.) Johnston var. scouleri Tiquilia nuttallii (Benth. ex Hook.) Richards.

Brassicaceae Anelsonia eurycarpa (Gray) Macbr. & Payson Arabis bodiensis Rollins cobrensis M.E. Jones davidsonii Greene divaricarpa A. Nels. drummondii Gray fernaldiana Rollins var. stylosa (S. Wats.) Rollins glabra (L.) Bemh. hirsuta (L.) Scop. var. glabrata Torr. & Gray holboellii Homem. var. penduhcarpa (A. Nels.) Rollins var. pinetorum (Tidestrom) Rollins var. retrofracta (Grahm.) Rydb. var. holboellii inyoensis Rollins

lemmonii S. Wats. var. lemmonii var. depauperata (A. Nels. & Kenn.) Rollins lyallii S. Wats.

var. lyallii platysperma Gray var. platysperma var. howellii (S. Wats.) Jepson January 1983 Lavin: Walker River Floristics 107

Table 1 continued. AREAS

puberula Nutt.

pulchra M.E. Jones ex S. Wats, var. pulchra var. gracilis M.E. Jones sparsiflora Nutt.

var. subvillosa (S. Wats.) Rollins var. sparsiflora Barbarea orthoceras Ledeb. var. dolichocarpa Fern, var. orthoceras Capsella bursa-pastoris (L.) Medic. Cardamine

breweri S. Wats, var. breweri Cardaria pubescens (C.A. Mey) Jarmolenko Caulanthus

pilosus S. Wats. Descurainia califomica (Gray) O.K. Schulz pinnata (Walt.) Britt. ssp. filipes (Gray) Detling ssp. halictorum (Cockerell) Detling ssp. menziesii (DC.) Detling richardsonii (Sweet) O.E. Schulz

ssp. incisa (Engelm.) Detling ssp. viscosa (Rydb.) Detling sophia (L.) Webb ex Prantl Draba albertina Greene

breweri S. Wats. densifolia Nutt. douglasii Gray var. crockeri (Lemmon) C.L. Hitchc.

lemmonii S. Wats, var. lemmonii var. incrassata Rollins oligosperma Hook.

var. subsessilis (S. Wats.) O.E. Schulz var. oligosperma

paysonii J.F. Macbride quadricostata Rollins stenoloba Ledeb. var. ramosa C.L. Hitchcock Erysimum argillosum (Greene) Rydb. capitatum (Dougl.) Greene perenne (S.Wats, ex Coville) Abrams repandum L. Hymenolobus procumbens (L.) Nutt. ex Torr. & Gray Lepidium lasiocarpum Nutt. perfoliatum L. virginicum L. var. pubescens (Greene) C.L. Hitchc. 108 Great Basin Naturalist Vol. 43, No. 1

Table 1 continued. AREAS habitats

N S W M 123456789

Lesquerella

kingii S. Wats, var. kingii X X Nasturtium officinale R. Br. Phoenicaulis cheiranthoides Nutt. Polyctenium

fremontii (S. Wats.) Greene Rorippa curvisiliqua (Hook.) Bess, ex Britton teres (Michx.) R. Stuckey Sisymbrium altissimum L. Stanleya pinnata (Pursh) Britton var. pinnata Streptanthus cordatus Nutt. ex Torr. & Gray oliganthus Rollins tortuosus Kellogg var. orbiculatus (Greene) Hall Thelypodium crispum Greene ex Payson integrifolium (Nutt.) Endl. ssp. complanatum Al-Shehbaz laciniatum (Hook.) Endl. Cactaceae Opuntia polyacantha Haw. var. rufispina (Engelm. & Bigelow) Benson pulchella Engelm.

Callitrichaceae CalUtriche heterophylla Pursh emend. Darby var. bolanderi (Hegelm.) Fassett. verna L. emend. Kuetz. Campanulaceae Nemacladus rigidus Curran Porterella carnosula (Hook. & Am.) Torr. Capparidaceae Cleomella hillmanii A. Nels. parviflora Gray

Caprifoliaceae Lonicera involucrata (Rich.) Banks ex Spreng. Sambucus caerulea Raf. racemosa L. ssp. pubens (Michx.) House var. microbotrys (Rydb.) Kearney & Peebles Symphoricarpos longifloTus Gray January 1983 Lavin: Walker River Floristics 109

Table 1 continued. AREAS

oreophilus Gray var. oreophilus parishii Rydb. Caryophyllaceae Arenaria

aculeate S. Wats. Cerastium alpinum L. vulgatum L. Minuartia nuttallii (Pax) Briq. ssp. fragilis (Maguire & Holmgren) McNeill ssp. gracilis (B.L. Robins.) McNeill obtusiloba (Rydb.) House

rossii (R. Br.) Graebn. rubella (Wahlenb.) Hiem Pseudostellaria jamesiana (Torr.) Weber & Hartman Sagina saginoides (L.) Karst. Saponaria officinalis L. Silene

bemardina S. Wats, ssp. maguirei Bocquet var. maguirei ssp. bemardina menziesii Hook. ssp. dorrii (Kellogg) C.L. Hitchc.& Maguire

nuda (S. Wats.) C.L. Hitchc. & Maguire ssp. insectivora (Henders.) C.L. Hitchc. & Maguire sargentii S. Wats. Stellaria crispa Cham. & Schlecht. longipes Goldie umbellata Turcz. ex Kar. & Kir. Chenopodiaceae Atriplex argentea Nutt. ssp. argentea canescens (Pursh) Nutt. ssp. canescens confertifolia (Torr. & Frem.) S. Wats. heterosperma Bunge patula L. ssp. hastata (L.) H. & S. rosea L. Bassia hyssopifolia (Pallas) Kuntze Ceratoides

lanata (Pursh) J.T. Howell var. lanata Chenopodium album L. atrovirens Rydb. botrys L. dessiccatum A. Nels. var. leptophylloides H.A. Wahl. (J. Murr) var. dessiccatum 110 Great Basin Naturalist Vol. 43, No. 1

Table 1 continued. AREAS habitats N

fremontii S. Wats. leptophyllum (Moq.) Nutt. ex S. Wats. overi Aellen Grayia spinosa (Hook.) Moq. Halogeton glomeratus (Stephan ex Bieb) C.A. Mey. Kochia scoparia (L.) Schrad. Monolepis nuttalliana (Roem. & Schult.) Greene spathulata Gray Sakola iberica Sennen & Pau Sarcobatus vermiculatus (Hook.) Torr. var. vermiculatus Sueda

occidentalis S. Wats.

Clusiaceae Hypericum formosum H.B.K. ssp. scouleri (Hook.) C.L. Hitchc. CONVOLVULACEAE Calystegia polymorpha (Greene) Munz Convolvulus arvensis L. CORNACEAE Cornus sericea L. ssp. sericea

Crassulaceae Sedum integrifolium Coult. & A. Nels. lanceolatum (Nutt.) Britton & Rose obtusatum Gray Crossosomataceae Forsellesia nevadensis (Gray) Greene CUSCUTACEAE Cuscuta suksdorfii Yunker var. subpedicellata Yunker Cyperaceae Carex abrupta Mackenzie aquatilis Wahlenb. athrostachya Olney aurea Nutt. brevipes W. Boott. canescens L. capitata L. congdonii Bailey disperma Dewey douglasii Boott January 1983 Lavin: Walker River Floristics 111

Table 1 continued. AREAS

exserta Mackenzie festivelhi Mackenzie fissuricola Mackenzie haydeniana Olney helleri Mackenzie heteroneura W. Boott

var. epapillosa (Mackenzie) F.J. Herm. var. heteroneura hoodii Boott

jepsonii J.T. Howell jonesii Bailey lanuginosa Michx. leporinella Mackenzie luzulifolia W. Boott microptera Mackenzie nebraskensis Dewey nervina Bailey nigricans C.A. Mey. pachystachya Cham, ex Steudel. phaeocephala Piper praegracilis W. Boott

rossii Boott ex Hook. rostrata Stokes ex With. scopulorum T. H. Holm.

var. bracteosa (Bailey) F.J. Herm. var. scopulorum simulate Mackenzie specifica Bailey spectabilis Dewey straminiformis Bailey subnigricans Stacey tahoensis Smiley vallicola Dewey vemacula Bailey vesicaria L. Eleocharis engelmannii Steud. palustris (L.) Roemer & Schultes pauciflora (Lightf.) Link Eriophorum crinigerum (Gray) Beetle Scirpus acutus Muhl. ex Bigelow americanus Pers. dementis M.E. Jones microcarpus Presl.

nevadensis S. Wats. pungens Vahl. Elaeagnaceae Elaeagnus angustifolia L. Shepherdia argentea (Pursh) Nutt.

Elatinaceae Elatine rubella Rydb. 112 Great Basin Naturalist Vol. 43, No. 1

Table 1 continued. AREAS habitats NSWM 12 3 456789 Ericaceae Arctostaphylos tievadensis Gray Cassiape mertensiana (Bong.) D. Don X X Kalmia microphylla (Hook.) Heller var. microphylla O O X X X Ledum glandulosum Nutt. var. californicum (Kellogg) C.L. Hitchc. X X Orthilia secunda (L.) House ssp. secunda Phyllodoce breweri (Gray) Heller Pterospora andromedea Nutt. Pyrola californica Krisa dentata Sm. var. dentata minor L. Sarcodes sanguinea Torr. Vaccinium caespitosum Michx. var. paludicola (Camp) Hulten X uliginosum L. ssp. occidentale (Gray) Hulten X EUPHORBIACEAE Chamaesyce serpyllifolia (Pers.) Small X X X X Fabaceae Astragalus andersonii Gray X X bolanderi Gray

calycosus Torr. ex S. Wats. var. calycosus canadensis L. var. brevidens (Gand.) Bameby casei Gray

curvicarpus (Heller) J.F. Macbr. var. curvicarpus o gibbsii Kellogg

iodanthus S. Wats. var. iodanthus johnnis-howellii Bameby kentrophyta Gray var. danaus Bameby X X lentiginosus Dougl. ex Hook. var. ineptus (Gray) M.E. Jones XX X malacus Gray XXX obscurus S. Wats. X

oophorus S. Wats. var. lavinii Bameby in ed. X X platytropis Gray January 1983 Lavin: Walker River Floristics 113

Table 1 continued. AREAS HABITATS N S W M 123456789 purshii Dougl. ex Hook. var. lectulus (S. Wats.) M.E. Jones X X X XXX XXX var. tinctus M.E. Jones X X X X XXX whitneyi Gray var. whitneyi X X X XXX XXX Dalea omata (Dougl. ex Hook.) Eat. & Wright X Lupinus andersonii S. Wats. X X X X X X X X X arbustus Dougl. ex Lindl. ssp. calcaratus (Kellogg) Dunn X X X XX XX brevicaulis S. Wats. X X X X X caudatus Kellogg ssp. caudatus X X X X XXX ssp. montigenus (Heller) Hess & Dunn X X X X confertus Kellogg X X X X X hypolasius Greene X X XXX lyallii Gray

var. danatis (Gray) S. Wats, X var. lyallii meionanthus Gray nevadensis Heller polyphyllus Lindl. ssp. superbus (Heller) Munz sellulus Kellogg

var. lobbii (S. Wats.) Cox var. sellulus tegeticulatus Eastw X inyoensis Heller Medicago lupulina L. sativa L. Melilotus albus Medic. officinalis (L.) Pallas Oxytropis parryi Gray Robinia pseudoacacia L. Trifolium andersonii Gray ssp. beatleyae Gillett ssp. andersonii cyathiferum Lindl. hybridum L. longipes Nutt. ssp. longipes monanthum Gray var. monanthum pratense L. productum Greene repens L. wormskjoldii Lehm. Vicia americana Muhl. ex Willd. ssp. americana Fagaceae Castanopsis sempervirens (Kellogg) Dudley 114 Great Basin Naturalist Vol. 43, No. 1

Table 1 continued. AREAS HABITATS

N S W M 123456789 Gentianaceae Frasera speciosa Dougl. ex Griseb. X X Gentiana calycosa Griseb. newberryi Gray Gentianopsis

holopetala (Gray) litis Geraniaceae Erodium cicutarium (L.) L'Her. Geranium richardsonii Fisch. & Trautv. Haloragidaceae Hippuris vulgaris L. Hydrocharitaceae Elodea canadensis Michx. Hydrophyllaceae Hesperochiron califomicus (Benth.) S. Wats. Hydrophyllum capitatum Dougl. ex Benth. var. alpinum S. Wats. Noma aretioides (Hook. & Am.) Brand densum Lemmon rothrockii Gray Nemophila spatulata Coville Phacelia

bicolor Torr. ex S. Wats.

curvipes Torr. ex S. Wats. frigida Greene hastata Dougl. ex Lehm. ssp. compacta (Brand) Heckard heterophylla Pursh ssp. virgata (Greene) Heckard humilis Torr. & Gray var. humilis hydrophylloides Torr. ex Gray linearis (Pursh) Holz. monoensis Halse

peirsoniana J.T. Howell ramosissima Dougl. ex Lehm. var. ramosissima

tetramera J.T. Howell Tricardia

watsonii Torr. ex S. Wats.

Iridaceae Iris missouriensis Nutt. Sisyrinchium fialophilum Greene idahoense Bickn. var. occidentale (Bickn.) Henderson January 1983 Lavin: Walker River Floristics 115

Table 1 continued. AREAS HABITATS N S W M 123456789 JUNCACEAE Juncus balticus Willd. X X

bryoides F.J. Herm. bufonius L.

var. occidentalis F.J. Herm. var. bufonius

capillaris F.J. Herm. chlorocephalus Engelm. drumniondii E. Mey. ensifolius Wikstr. var. montanus (Engelm.) C.L. Hitchcock

hemiendytus F.J. Herm. kello^ii Engelm. longistylis Torr. macrandrus Coville

megaspermus F.J. Herm. mertensianus Bong. mexicanus Willd.

nevadensis S. Wats. orthophyllus Coville

parry i Engelm. saximontanus A. Nels. Luzula

ditxiricata S. Wats. multiflora (Retz.) Lej. ssp. comosa (E. Mey.) Hulten spicata (L.) DC. subcongesta (S. Wats.) Jepson JUNCAGINACEAE Triglochin concinna Burtt-Davy var. debilis (M.E. Jones) J.T. Howell maritima L. Lamiaceae Agastache urticifolia (Benth.) Kuntze Marrubium vulgare L. Mentha arvensis L. ssp. haplocalyx Briq. Monardella odoratissima Benth. ssp. glauca (Greene) Epling Salvia dorrii (Kellogg) Abrams var. dorrii Trichostema austromontana Lewis Lemnaceae Lemna gibba L. minuta H.B.K. trisulca L.

Lentibulariaceae Utricularia vulgaris L. 116 Great Basin Naturalist Vol. 43, No. 1

Table 1 continued. AREAS HABITATS

N S W M 123456789

LiLIACEAE Allium amplectens Torr. anceps Kellogg

atroTubens S. Wats, var. atrorubens

bisceptrum S. Wats. campanulatum S. Wats. lemmonii S. Wats. parvum Kellogg validum S. Wats. Cahchortus bruneaunis A. Nels. & J.F. Macbride

leichtlinii Hook. f. Camassia

leichtlinii (Baker) S. Wats. Fritillaria atropurpurea Nutt. pinetorum A. Davids. Lilium parvum Kellogg Muilla transmontana Greene Smilacina stellate (L.) Desf. var. stellata Triteleia

gracilis (S. Wats.) Greene

ixioides (Ait. f.) Greene ssp. analina (Greene) Lenz Veratrum califomicum Durand var. californicum Zigadenus

paniculatus (Nutt.) S. Wats. venetiosus S. Wats, var. venenosus LiMNANTHACEAE Floerkea proserpinacoides Willd. LiNACEAE Linum lewisii Pursh var. lewisii LOASACEAE Mentzelia albicaulis (Hook.) Torr. & Gray congesta (Nutt.) Torr. & Gray dispersa S. Wats. laevicaulis (Hook.) Torr. & Gray montana (A. Davids.) A. Davids. torreyi Gray var. torreyi veatchiana Kellogg LORANTHACEAE Arceuthobium divaricatum Engelm. Phoradendron juniperinum Engelm. ex Gray ssp. juniperinum January 1983 Lavin: Walker River Floristics 117

Table 1 continued. AREAS HABITATS NSWM 123456789 Malvaceae Malva neglecta Wallr. X X X X Sidalcea glaucescens Greene o o o o o multifida Greene X X X X oregana (Torr. & Gray) Gray ssp. spicata (Kegel) C.L. Hitchc. XX X X X Sphaeralcea atnbigua Gray ssp. monticola Kearney X X X X grossulariifolia (Hook. & Am.) Rydb. X X X X MORACEAE Humulus lupulus L. var. lupuhides E. Small o o Nyctag inaceae Abronia

turbinata Torr. ex S. Wats. o Mirahilis bigelovii Gray var. retroTsa (Heller) Munz X X X X Nymphaeaceae Nuphar luteum (L.) Sibthorp. & Sm. ssp. polysepalum (Engelm.) E.O. Real Oleaceae Menodora spinescens Gray Onagraceae Boisduvalia

densiflora (Lindl.) S. Wats. Camissonia claviformis (Torr. & Frem.) Raven ssp. integrior (Raven) Raven nevadensis (Kellogg) Raven

pubens (S. Wats.) Raven pusilla Raven subacaulis (Pursh) Raven tanacetifolia (Torr. & Gray) Raven ssp. tanacetifolia Circaea alpina L. ssp. pacifica (Aschers. & Magnus) Raven Epilobium anagallidifolium Lam. angustifolium L. brachycarpum Presl. ciliatum Raf. ssp. ciliatum glaberrimum Barbey lactiflorum Hausskn. latifolium L. obcordatum Gray ssp. obcordatum oregonese Hausskn. 118 Great Basin Naturalist Vol. 43, No. 1

Table 1 continued. AREAS habitats N January 1983 Lavin: Walker River Floristics 119

Table 1 continued. AREAS HABITATS

N S 120 Great Basin Naturalist Vol. 43, No. 1

Table 1 continued. AREAS habitats

N S January 1983 Lavin: Walker River Floristics 121

Table 1 continued. AREAS HABITATS

N S 122 Great Basin Naturalist Vol. 43, No. 1

Table 1 continued. AREAS habitats

N S W M 123456789

Leptodactylon pungens (Torr.) Nutt. ex Rydb. ssp. pulchriflonim (Brand) Mason X X X X X X X X XXX Linanthus ciliatus (Benth.) Greene var. neglectus (Greene) Jepson X X nuttallii (Gray) Greene ex Milliken

ssp. nuttallii X X septentrionalis Mason MicTosteris gracilis (Hook.) Greene

ssp. gracilis ssp. humilfs (Greene) V. Grant Navarretia breweri (Gray) Greene divaricata (Torr. ex Gray) Greene var. divaricata Phlox caespitosa Nutt.

ssp. pulvinata Wherry covillei E. Nels. diffusa Benth. ssp. subcarinata Wherry ssp. diffusa hoodii Richards ssp. canescens (Torr. & Gray) Wherry longifolia Nutt. Polemonium caeruleum L. ssp. amygdalinum (Wherry) Munz califomicum Eastw. chartaceum Mason eximium Greene pulcherrimum Hook. POLYGALACEAE Polygala intermontana Wendt

subspinosa S. Wats. POLYGONACEAE Chorizanthe brevicomu Torr. var. spathulata (Rydb.) C.L. Hitchc. watsonii Torr. & Gray Eriogonum

baileyi S. Wats. var. baileyi beatlleyae Reveal brachyantherum Coville caespitosum Nutt. var. cemuum elatum Dougl. ex Benth. var. elatum

esmeraldense S. Wats. heermannii Dur. & Hilg. var. humilius (Stokes) Reveal

hookeri S. Wats. incanum Torr. & Gray January 1983 Lavin: Walker River Floristics 123

Table 1 continued. 124 Great Basin Naturalist Vol. 43, No. 1

Table 1 continued. AREAS habitats

N S W M 123456789 PORTULACACEAE Calyptridium

roseum S. Wats. X X umbellatum (Torr.) Greene Claytonia nevadense S. Wats.

rubra (T.J. Howell) Tidestrom umbellata S. Wats. Leivisia nevadensis (Gray) B.L. Robins. pygmaea (Gray) B.L. Robins, ssp. pygmaea rediviva Pursh

ssp. rediviva

ssp. minor (Rydb.) A. Holmgren sierrae Ferris

triphylla (S. Wats.) B.L. Robins. Montia chamissoi (Ledeb. ex Spreng.) Greene POTAMOGETONACEAE Potamogeton gramineus L. richardsonii (Benn.) Rydb.

Primulaceae Androsace septentrionalis L. var. subumbellata A. Nels. Dodecatheon alpinum (Gray) Greene ssp. alpinum ssp. majtis H.J. Thompson jeffreyi Van Houtte Primula suffrutescens Gray Ranunculaceae Aconitum columbianum Nutt. Actaea rubra (Ait.) WiUd. ssp. arguta (Nutt.) Hulten Anemone drummondii S. Wats. Aquilegia formosa Fisch. var. paucifhra (Greene) Boothman var. formosa pubescens Goville Caltha leptosepala DC. ssp. howellii (Huth) P.G. Smith Clematis ligusticifolius Nutt. var. brevifolia Nutt. Delphinium andersonii Gray

ssp. andersonii glaucum S. Wats. January 1983 Lavin: Walker River Floristics 125

Table 1 continued. AREAS HABITATS NSWM 123456789

parishii Gray

ssp. parishii X X polycladon Eastw. Myosurus aristatus Benth. ex Hook. minimus L.

ssp. montanus Campbell Ranunculus alismifolius Geyer ex Benth. var. alismellus Gray andersonii Gray aquatilis L. var. capillaceus (Thuill.) DC. cymbalaria Pursh var. saximontanus Fern. eschscholtzii Schlecht. var. eschscholtzii var. oxynotis (Gray) Jepson glaberrimus Hook. var. ellipticus Hook. occidentalis Torr. & Gray var. ultramx)ntantis Greene testiculatus Crantz Thalictrum fendleri Gray var. fendleri sparsiflorum Turcz. ex Fisch. & Mey. var. saximontanum Boivin Rhamnaceae Ceanothus greggii Gray var. vestitus (Greene) McMinn X X velutinus Dougl. ex Hook. XX X Rham.nus rubra Greene ssp. rubra o ROSACEAE Amelanchier pallida Greene X pumila Torr. & Gray utahensis Koehne var. covillei (Standi.) Clokey var. utahensis Cercocarpus ledifolius Nutt. ex Torr. & Gray var. ledifolius

var. intricatus (S. Wats.) M.E. Jones Chamaebatiaria millefolium (Torr.) Maxim. Fragaria virginiana Duchesne ssp. platypetala (Rydb.) Staudt Geum canescens (Greene) Munz macrophyllum Willd. 126 Great Basin Naturalist Vol. 43, No. 1

Table 1 continued. AREAS habitats

N S W M 123456789

Holodiscus dumosa (Nutt.) Heller var. glabrescens (Greenm.) C.L. Hitchc. X X X X XXX XX Horkelia fusca Lindl. ssp. pseudocapitata (Rydb.) Keck X X Ivesia gordonii (Hook.) Torr. & Gray lycopodioides (Hook.) Torr. & Gray

ssp. lycopodioides ssp. megalopetala (Rydb.) Keck muirii Graj^

purpurascens (S. Wats.) Keck ssp. congdonis (Rydb.) Keck

shockleyi S. Wats. Potentillc anserina L.

ssp. anserina biennis Greene

breweri S. Wats. diversifolia Lehm. drummondii Lehm.

ssp. bruceae (Rydb.) Keck ssp. drummondii fhbellifolia Hook, ex Torr. & Gray var. fhbellifolia fruticosa L. ssp. floribunda (Pursh) Elkington glandulosa Lindl.

ssp. nevadensis (S. Wats.) Keck gracilis Dougl. ex Hook. var. glabrata (Lehm.) C.L. Hitchc. var. flabelliformis (Lehm.) Torr. & Gray newberryi Gray pectinisecta Rydb. pseudosericea Rydb. Prunus andersonii Gray emarginata (Dougl. ex Hook.) Walp. virginiana L. var. demissa (Nutt.) Sarg. var. melanocarpa (A. Nels.) Sarg. Purshia tridentata (Pursh) DC. Rosa woodsii Lindl.

var. ultrarrumtana (S. Wats.) Jepson Rubus parviflorus Nutt. var. bifarius Fern. Sibbaldia procumbens L. Sorbus califomica Greene Spiraea densiflora Nutt. ex Torr. & Gray ssp. densiflora January 1983 Lavin: Walker River Floristics 127

Table 1 continued. AREAS HABITATS

N S W M 123456789 RUBIACEAE Galium aparine L. X X bifolium S. Wats. X X hypotrichium Gray ssp. ebbettsense Dempster & Ehrend. ssp. hypotrichium multiflorum Kellogg trifidum L. ssp. pacificum Wieg. ssp. subbiflorum (Wieg.) Puff trifhrum Michx. Kelloggia galioides Torr.

Salicaceae Populus balsamifera L. ssp. trichocarpa (Torr. & Gray) Brayshaw

fremontii S. Wats. tremuloides Michx. Salix arctica Pallas drummondiana Barratt ex Hook. eastwoodiae Heller exigtia Nutt. geyeriana Anderss. var. geyeriana var. argentea (Bebb) Schneid. lasiandra Benth. var. lasiandra var. caudata (Nutt.) Sudworth lasiolepis Benth. lemmonii Bebb ligulifolia (Ball) Ball ex Schneid. lutea Nutt. melanopsis Nutt. myrtillifolia Anderss. var. myrtillifolia orestera Schneid. planifolia Pursh var. monica (Bebb) Schneid. reticulata L. ssp. nivalis (Hook.) Love, Love & Kapoor scouleriana Barratt ex Hook.

Saxifragaceae Heuchera duranii Bacig. rubescens Torr. var. alpicola Jepson Lithophraffna glabrum Nutt. Mitella breweri Gray pentandra Hook. Pamassia palustris L. var. califomica Gray 128 Great Basin Naturalist Vol. 43, No. 1

Table 1 continued. AREAS habitats

N S W M 123456789

Ribes aureutn Pursh var. aureum X X cereum Dougl. inebrians Lindl. inerme Rydb. var. inerme montigenum McClatchie velutinum Greene var. velutinum var. glanduliferum (Heller) Jepson viscosissimum, Pursh var. viscosissim,um. Saxifraga aprica Greene bryophora Gray nidifica Greene odontoUmui Piper

oregana T.J. Howell var. sierrae Coville tolmiei Torr. & Gray var. ledifolia (Greene) Engl. & Irmsch.

SCROPHULARIACEAE Antirrhinum

kingii S. Wats. Castilleja applegatei Fern. var. fragilis (Zeile) N. Homgren var. pallida (Eastwood) N. Holmgren chromosa A. Nels. exilis A. Nels. lemmonii Gray linariifolia Benth. ex DC. miniata Dougl. ex Benth. nana Eastwood peirsonii Eastwood

pilosa (S. Wats.) Rydb. Collinsia parviflora Dougl. ex Lindl. Cordylanthus

helleri (Ferris) J.F. Macbride ramosus Nutt. ex Benth. ssp. setosus Pennell Keckiella breviflora (Lindl.) Straw ssp. glabrisepala (Keck) Straw Limosella aquatica L. Mimetanthe pilosa (Benth.) Greene Mimulus breweri (Greene) Coville coccineus Congd. densus A.L. Grant floribundus Dougl. ex Lindl. guttatus Fisch. ex DC. lewisii Pursh mephiticus Greene January 1983 Lavin: Walker River Floristics 129

Table 1 continued. AREAS HABITATS NSWM 1234567 moschatus Dougl. X X X X primuloides Benth. var. primuloides X X O X X X X rubellus Gray XX X X X X suksdorfii Gray XXXX XXXX XXX tilingii Kegel XXXX X X X XXX Orthocarpus copelandii Eastwood var. cryptanthus (Piper) Keck XXX XXX hispidus Benth. XX O X O X X luteus Nutt. o o o o Pedicularis attollens Gray ssp. attollens X X groenlandica Retz.

ssp. groenlandica semibarbata Gray var. semibarbata Penstemon bridgesii Gray dovidsonii Greene var. davidsonii deustus Dougl. ex Lindl. ssp. deustus heterodoxus Gray var. heterodoxus humilis Nutt. ex Gray newberryi Gray ssp. newberryi procerus Dougl. ex Grahm. ssp. formosus (A. Nels.) Keck rubicundis Keck rydbergii A. Nels. var. varians (A. Nels.) Cronq. speciosus Dougl. ex Lindl. Scrophularia

desertorum (Munz) R.J. Shaw Verbascum thapsus L. Veronica americana (Raf.) Schwein. ex Benth. anagallis-aquatica L. beccabunga L. peregrina L. ssp. xalapensis (H.B.K.) Pennell serpyllifolia L. ssp. humifusa (Dickson) Syme ssp. serpyllifolia wormskjoldii Roemer & Schultes ssp. alterniflora (Fern.) Pennell SOLANACEAE Leucophysalis nana (Gray) Averett Nicotiana

attenuata Torr. ex S. Wats. Solanum triflorum Nutt. 130 Great Basin Naturalist Vol. 43, No. 1

Table 1 continued. AREAS habitats NSWM 123456789 Typhaceae Typha latifolia L. X X X X Urticaceae Urtica dioica L. var. holosericea (Nutt.) C.L. Hitchc. X X X X Valebianaceae Plectritis macrocera Torr. & Gray ssp. grayit (Suksdorf) Morey X X Valeriana californica Heller XX o ox XXX Verbenaceae Verbena bracteata Lag. & Rodr. X X ViOLACEAE Viola adunca Sm. var. adunca o bakeri Greene ssp. bakeri beckwithii Torr. & Gray ssp. beckwithii macloskeyi Lloyd ssp. macloskeyi nephrophylla Greene var. nephrophylla purpurea Kellogg ssp. atriplicifolia (Greene) Baker 6f Clausen ssp. aurea (Kellogg) Clausen Zygophyllaceae Tribulus terrestris L.