Aliso: A Journal of Systematic and Evolutionary Botany

Volume 37 Issue 1 Issue 1–2 Article 2

2019

Plantae Coulterianae: Thomas Coulter’s Californian Exsiccata

Gary D. Wallace California Botanic Garden, Claremont, CA

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Recommended Citation Wallace, Gary D. (2020) "Plantae Coulterianae: Thomas Coulter’s Californian Exsiccata," Aliso: A Journal of Systematic and Evolutionary Botany: Vol. 37: Iss. 1, Article 2. Available at: https://scholarship.claremont.edu/aliso/vol37/iss1/2 Aliso, 37(1–2), pp. 1–73 ISSN: 0065-6275 (print), 2327-2929 (online)

PLANTAE COULTERIANAE: THOMAS COULTER’S CALIFORNIAN EXSICCATA

Gary D. Wallace California Botanic Garden [formerly Rancho Santa Ana Botanic Garden], 1500 North College Avenue, Claremont, California 91711 ([email protected])

abstract An account of the extent, diversity, and importance of the Californian collections of Thomas Coulter in the herbarium (TCD) of Trinity College, Dublin, Ireland, is presented here. It is based on examination of collections in TCD, several other collections available online, and referenced literature. Additional infor- mation on historical context, content of herbarium labels and annotations is included. Coulter’s collections in TCD are less well known than partial duplicate sets at other herbaria. He was the first botanist to cross the desert of southern California to the Colorado River. Coulter’s collections in TCD include not only 60 vascular plant specimens previously unidentified as type material but also among the first moss andmarine algae specimens known to be collected in California. A list of taxa named for Thomas Coulter is included. Images of the map of his route are presented here as well as comments on some locality errors. Mention is made of works cited in annotations that were, in fact, never published, leading to persistent author citation errors. Coulter’s expansive interests in collecting and natural history included languages of the indigenous tribes he encountered in California. Key words: bryophytes, California, Coulter, exsiccata, flora, herbarium, marine algae, Native Americans, Native American language, TCD.

“This is truly the kingdom of Desolation.” “I fear the whole propriate, correct incorrect determinations, and assign infraspe- flora of California Alta will fall far short of a thousand cific taxa not recognized when the specimens were first identi- species. Indeed, I do not expect above 600 and of these fied. I also hope curators and monographers look for more of not above 200 of much interest.” (Coulter 1832) Coulter’s specimens in other collections and image those on hand. I have attempted to locate as many of Coulter’s Californian preface exsiccata as possible in TCD. There are likely more specimens there under older names or merely misfiled. I have included ref- The title of this paper includes “Plantae Coulterianae” partly erences to specimens at other herbaria historically more likely in recognition of—and homage to—the similarly titled treat- to have been consulted than those at TCD. The account of Coul- ments by Asa Gray and his co-authors about the expeditions ter’s Californian plants (phanerogams and bryophytes) is in- and collections of other explorers of the era, including Plan- cluded (Appendix I) to demonstrate the taxonomic breadth of tae Fendlerianae Novi-Mexicanae (Gray 1849), Plantae Lind- his collecting in California and the distribution of his exsiccata. heimerianae (Engelmann and Gray 1845), Plantae Wrightianae There is also a separate account of Coulter’s marine algae speci- Texano–Neo-Mexicanae (Gray 1852), Plantae Novae Thurberi- mens (Appendix II), taxa for which his collections are type ma- anae (Gray 1854) and Plantas Hartwegianas (Bentham 1839– terial (Appendix III), a list of plant and algae taxa named for 1857) and in part in response to Nelson and Probert’s (1994: 79) him (Appendix IV), a list of taxa claimed as first collected by comment, “No-one has published a Plantae Coulterianae.” Coulter but not typified by his specimens (Appendix V), and a My primary purpose is to present a list of—and guide to— list of obscure generic names and their disposition in this paper the hundreds of Coulter’s Californian specimens in the Trin- (Appendix VI). This contribution is not to be taken as taxonom- ity College Dublin Herbarium (TCD), Dublin, Ireland. His are ically definitive because most determinations have not been ver- among the earliest collections of Californian specimens. It is ified. It was not feasible to attempt to verify or update identifica- beyond the scope and purpose of this contribution to docu- tions of the hundreds of exsiccata on site at TCD due to limited ment the identifiers or provenance of each of Coulter’s Cali- time and onsite resources. There may be other herbaria contain- fornian specimens. The initial identifications of Coulter’s spec- ing significant numbers of duplicates of Coulter’s Californian imens were made over 170 years ago. It is likely that many are exsiccata and I welcome additions and corrections to this effort. misidentified or under-identified (not differentiated to infraspe- An applicable comment on this contribution is: “A measure of cific taxa). I hope the listing here and description of discoveries the usefulness of this study will be the number of additions and (see Findings below) entices systematists to examine the Coulter corrections it elicits” (Renkema and Ardagh 1930). exsiccata at TCD and elsewhere to verify their type status if ap-

introduction © 2019, The Author(s), CC-BY. This open access article is distributed under a Creative Commons Attribution License, which allows unrestricted Thomas Coulter (1793–1843) (Fig. 1) is best known, in Cal- use, distribution, and reproduction in any medium, provided that the orig- ifornia, as an Irish botanist for whom are named Pinus coul- inal author(s) and source are credited. Articles can be downloaded at teri (Coulter pine), which bears the heaviest cones of any http://scholarship.claremont.edu/aliso/. pine species, and the large white-flowered Romneya coulteri 2 Wallace ALISO

Fig. 1. Portrait of Thomas Coulter, artist unknown, oil on canvas. Collection of Trinity College Dublin, The University of Dublin. Reproduced by permission of the Board of Trinity College Dublin, The University of Dublin, Ireland. VOLUME 37(1–2) Plantae Coulterianae 3

(Coulter’s matilija poppy). David Douglas, engaged by the Hor- Probert 1994: 98) but soon quit and went south to Guaymas. ticultural Society of London to collect plants in North Amer- Coulter collected plant specimens in the area, noted by Hem- ica, met Coulter in Monterey in 1831 and later described him as sley (1879–1888) as collected in “Sonora Alta” (the intended “ …a good fellow, most zealous in all he undertakes and anx- scope of this geographic designation is uncertain). A set of these ious to collect a vast collection …” (letter from David Douglas is in the herbarium at Kew (K). Thomas Coulter’s collecting of to William Hooker in London dated 15 April 1833, transcribed “California Alta” plant specimens (he used “California Alta” to by Nelson and Probert 1994: 111). Alphonse de Candolle de- distinguish the region from Baja California; see clarification be- scribed Coulter as an amiable eccentric who kept live lizards low) may have begun, for reasons explained later, with his stay and snakes in his pockets and whistled tunes to them to keep in Sonora, Mexico, in December 1829. Coulter had wanted to them calm (de Candolle’s 1862 memoirs: 332 translated in Nel- travel to California for some time and planned to go to Califor- son and Probert 1994: 26). Coulter, an accomplished all-around nia overland but instead sailed for California Alta from Guay- naturalist, and lexicographer, was also an avid fisherman and a mas on 21 August 1831. Prior to leaving Mexico, Coulter had good shot. Coulter’s contribution to our knowledge of the diver- left instructions that all of his earlier Mexican collections be sent sity of the California flora is largely overlooked and underappre- to London. ciated because narratives of his explorations were never written and his collections were not well known for several years after Coulter’s Mexican Woods his death. Coulter wrote only one short article about his travels in California for the Journal of the Royal Geographical Society Coulter reportedly collected 784 wood samples from Mex- of London (Coulter 1835). ico and California, of which most, if not all, were accompanied Thomas Coulter was among the earliest floristic collectors to by herbarium vouchers (Ball 1846: 3). Other references have visit California. He traveled extensively along the coast and inte- higher estimates of the number of wood samples (e.g., Nelson rior desert and collected intensively wherever he went, vascular and Probert 1994: 92) but no others suggest that any wood sam- and non-vascular plants as well as marine algae. He collected ples were from California. It has been assumed that Coulter’s showy specimens and plain specimens, annuals and perennials Mexican wood specimens were included with the long-term loan alike. He seems to have had a good eye for differences, collect- of Dixon’s specimens of tropical woods from TCD to the Uni- ing many different species of Lupinus and Eschscholzia. Coul- versity of Limerick, Ireland. However, none of Coulter’s wood ter’s specimens are not scanty and he often collected enough ma- specimens has been located with the Dixon specimens. I have terial for duplicates. His Californian plant collections, amassed seen some of the herbarium vouchers for the wood samples at in just over two years, were assigned at least 841 exsiccata num- the National Botanic Garden (DBN) in Dublin (see also Nelson bers. Coulter’s Californian collections are unusual in that spec- and Probert 1994: 92) but many are also at TCD.Theywereall imens in the partial sets distributed from TCD to other herbaria Mexican specimens, so Ball’s suggestion that some were from or individuals after Coulter’s death in 1843 are better known and California is a likely error. A comparison of a few herbarium studied than the first set retained at TCD. No list of Coulter’s vouchers for the wood specimens indicates that the affixed num- Californian exsiccata in TCD has previously been prepared and bers on these wood specimen vouchers are a separate sequence there are few reports of Coulter specimens at other herbaria. The from Coulter’s Mexican exsiccata series. focus of this contribution is to bring attention to the early Cali- Nelson and Probert (1994) present an extensive account of fornian collections represented by Coulter’s specimens in TCD. Coulter’s life and exploits in Mexico and California. Bancroft It is also of interest to note the disparity or lack of current anno- (1885), Coville (1895), McVaugh (1943), McKelvey (1955) and tations among the institutions holding Coulter specimens. The Nelson (1983, 1988) include (occasionally conflicting) descrip- majority of the Coulter’s Californian specimens in TCD have tions of Coulter’s itineraries in Mexico and California as well not been annotated since their original determinations over 170 as aspects of his life and collections. Coulter’s itinerary in Cal- years ago. ifornia is especially difficult to follow because there is very Thomas Coulter obtained a degree in medicine from the Uni- little documentation, scant information on the herbarium la- versity of Dublin. He completed his studies in botany in Geneva bels, and he retraced his routes at times. He collected along under Augustin-Pyramus de Candolle, preparing a monograph the central coast of California north perhaps as far as San Fran- on the Dipsacaceae. When he met Alexander von Humboldt in cisco and south as far as San Diego and inland from the coast Paris in 1823, they probably discussed Coulter’s desire to cross north of San Diego through the desert to the Colorado River South America between Buenos Aires, Argentina, and Lima, near present-day Yuma, Arizona. He included a map of Upper Peru, and then to explore Mexico and California (Nelson and California (Fig. 2–5) in the only paper he wrote about his time in Probert 1994: 30). While seeking the means to travel to South California (Coulter 1835). America, Coulter was offered and accepted the position of med- ical doctor to the Real del Monte Mining Company in the state Alta California Upon Coulter’s Arrival of Hidalgo, Mexico. It was ideal for him because the company also wanted a naturalist on staff (Nelson and Probert 1994: 33). Based on correspondence accounts, Coulter arrived in Mon- Coulter went to Mexico in 1825 where his contract allowed time terey, California, a village of about 700 inhabitants, in late to collect plants, insects, reptiles, and bird skins. There he col- September or early October 1831 (Nelson 1983: 68, 102). For lected hundreds of herbarium specimens as well as wood sam- context, Mexico had won its independence from Spain in 1821. ples and associated herbarium vouchers (Nelson and Probert Alta California, as a Province of Mexico, included all of what 1994: 92). He ultimately resigned from the mining company in is now California, Nevada, Utah, and most of Arizona. It was May 1829 and went to Pitic (Hermosillo) in Sonora, Mexico, in a state of tension when Coulter arrived. The governor of to establish a mint for the Mexican government (Nelson and Alta California issued a Proclamation of Emancipation in 1826 4 Wallace ALISO

Fig. 2. Map of Upper California from Coulter (1835): Thomas Coulter, Notes on Upper California. Proc. Roy. Geogr. Soc. London 5: 59–70 (1835) map, RB 7310, The Huntington Library, San Marino, California. Reproduced by permission. Enlarged versions of the upper, middle and lower parts of this map are shown in Fig. 3–5. VOLUME 37(1–2) Plantae Coulterianae 5

Fig. 3. Enlarged upper portion of Fig. 2: Map of Upper California from Coulter (1835): Thomas Coulter, Notes on Upper California. Proc. Roy. Geogr. Soc. London 5: 59–70 (1835) map, RB 7310, The Huntington Library, San Marino, California. Reproduced by permission. For details, see description in section entitled Coulter’s (1835) map. freeing all Indians in the military districts of San Diego, Santa Coulter and the Native Americans Barbara and Monterey from missionary rule and made them el- igible for Mexican citizenship. The Mexican government de- Coulter, although viewing the Native Americans by the cided to secularize the Franciscan missions, partly because of stereotypical norms of the period, expressed his opinion of the the Franciscan’s continued loyalty to the Catholic Church in condition of the “aboriginal inhabitants”: “ …I believe that a Spain and partly to deprive the friars of their control over the na- great deal of both force and fraud were used in congregating tive Americans who lived and worked around the missions. “The them together in missions; and the moment that force shall Franciscans were at times oppressive and ruthless and the Mex- be altogether withdrawn, I have no doubt that the majority of ican government wanted to remove them from their powerful them will return to the woods. Now that the seaboard is pretty position in the province” (Nelson and Probert 1994: 100). Sec- much occupied by whites, the Indians will probably retire ularization was in progress but formalized with passage of An to their relations still living in free in the interior.” (Coulter Act for the Secularization of the Missions of California in 1833. 1835: 67). Coulter exhibited a talent and interest in languages, Other sources of discord were the competing factions seeking gathering oral vocabularies which he wrote down, of several to replace the former Spanish governor of California. Thomas different dialects spoken by the various Native Americans he Coulter and fellow botanist David Douglas signed on as mem- encountered in California (Nelson and Probert 1994: 109–110). bers of Compañía Extranjera (Company of foreigners) under He shared his lists of words with John Scouler who included Zamorano, a friend of the former governor, to defend Mon- them in a paper on the languages of tribes of northwest Amer- terey, the capital of Alta California (Bancroft 1885: 221; Nelson ica (Scouler 1841: 246–251). Except for the Pima word list and Probert 1994: 104). The company apparently served only as that Coulter likely recorded near the Colorado River, Scouler guards. The extent of Douglas’ and Coulter’s service to the com- included the word lists under the names of the missions where pany is unknown. This is the environment under which Coulter Coulter presumably made his notes: San Diego, San Juan Capis- traveled and collected in Alta California. trano, San Gabriel, Santa Barbara, San Luis Obispo, and San 6 Wallace ALISO

Fig. 4. Enlarged middle portion of Fig. 2: Map of Upper California from Coulter (1835): Thomas Coulter, Notes on Upper California. Proc. Roy. Geogr. Soc. London 5: 59–70 (1835) map, RB 7310, The Huntington Library, San Marino, California. Reproduced by permission.

DEDICATION: I would like to dedicate this section of the paper to Jessica M. Orozco, (MS, Botany, 2016, Claremont Graduate University and Rancho Santa Ana Botanic Garden) whose enthusiasm for botany and all things related to her Native American heritage were tragically cut short. One will seldom meet such a kind and giving parent, talented botanist, eager learner, enthusiastic teacher and Native American advocate. There is likely untold Native American context to many of our studies of the California flora. GDW

Antonio. This is further evidence that Coulter likely trav- His record of comparable vocabularies of several different eled along El Camino Real and routes that connected the languages of the Native Americans reflects the broad interests missions. I have seen no specimen labels that refer to San Juan and talents of Coulter, affirms the likely route of his travels, and Capistrano although Coulter undoubtedly passed through the partially describes the social context of his journeys. area. In addition to drawing attention to this reference and as- pect of Coulter’s exploits, Nelson and Probert (1994: 109) credit Coulter’s Travels in Alta California A.P. Grant for providing dialect names for the word lists Coulter gathered based on his locations, San Diego (Diegueño), San Correlating Coulter’s collection localities and travel itinerary Juan Capistrano (Juaneño-Ajachme), San Gabriel (Gabrielino- in California is difficult. In part because all of his notes were Tongva), Santa Barbara (Barbareño Chumash), San Luis Obispo apparently lost between London and Dublin during his trip (Obispeño), and San Antonio (Antoniaño Salinan). Coulter’s home and subsequently scant if any locality is ascribed to his work was cited as recently as 1970 (Beeler 1970) as an impor- specimens. Coulter (1835) included a map (Fig. 2 –5) to illus- tant contribution to the study of native American languages. trate some land features and his routes in California. A copy of VOLUME 37(1–2) Plantae Coulterianae 7

Fig. 5. Enlarged lower portion of Fig. 2: Map of Upper California from Coulter (1835): Thomas Coulter, Notes on Upper California. Proc. Roy. Geogr. Soc. London 5: 59–70 (1835) map, RB 7310, The Huntington Library, San Marino, California. Reproduced by permission. 8 Wallace ALISO

Coulter’s map was reproduced in Dawson (1951) and Nelson Coulter left Mexico and returned to London in mid-November and Probert (1994). San Luis Obispo, La Purisima, Santa Ynes, 1834 (Nelson and Probert 1994: 117) where he remained for Santa Barbara, San Buenaventura, San Fernando, San Gabriel, a time with at least some of his collections. News of his ex- San Pedro, Pala, San Felipe, Bahia, Carinal, Agua Soda, San Pe- ploits and return spread quickly and fellow botanists were eager dro (desert) and San Pablo as well as San Louis [sic] Rey and San to view his collections. Sorting and labeling of the specimens Diego are shown on Coulter’s map (Fig. 2–5). More discussion began in January 1835 (Nelson and Probert 1994: 118). Dur- of Coulter’s map is below at Notes. There are no dates provided ing this period Coulter loaned several pine cones he had col- to indicate the dates or direction of his travels along the segments lected to David Don, secretary and librarian to Aylmer Bourke of his routes. The best accounting of his California itinerary with Lambert who was continuing Lambert’s monographic work on dates and locations are those by Nelson (1983) based, in part, pines. Don (1836) described five new species of conifers based on Coulter’s correspondence, and Nelson and Probert (1994). on these cones. Coulter had returned to Ireland by August 1835 McKelvey (1955: 428–442) provides similar details of Coul- (Nelson and Probert 1994: 122). ter’s travels in California. These references provide the basis of this section although they do not mention where plant specimens Coulter’s Californian Specimens were collected. Coulter joined a party traveling south to San Gabriel along El Coulter made extensive collections of flowering plants (Fig. Camino Real soon after he arrived in Monterey. They apparently 6), ferns, conifers, bryophytes, and marine algae. He may have left Monterey on 18 October 1831 traveling, likely on horseback collected lichens as well. There is no record of when Coul- with pack mules for supplies and equipment, along the valley of ter’s Californian plant specimens were sent to London or when the Rio San Buenaventura (Salinas River) east of the Santa Lu- they arrived. Unfortunately, Coulter’s journals never arrived and cia Mountains to San Luis Obispo. From there the trail went were presumed lost. In a report on the Dublin University Mu- south to La Purisima and eastward to Santa Ynez then south seum, director Robert Ball (1846: 5) stated that Coulter pro- through to the coast and eastward to Santa Barbara where they cured during his stay in Mexico and California “all together arrived on 20 October 1831. The party stayed for a few days, (probably) 3,000 species.” Robinson (1843: 548), reporting on then continued south through San Buenaventura (Ventura) and the passing of Dr. Coulter to the 16 March 1844 meeting of San Fernando, arriving in San Gabriel on 3 November 1831. the Royal Irish Academy, stated that Coulter had collected up- Coulter did not venture further south on this trip and retraced wards of 50,000 specimens of 10,000 to 11,000 species. That his route to Monterey, arriving on 23 November 1831. That is would include collections from Mexico as well as California. when he met the Scottish plant collector David Douglas who Ball’s and Robinson’s numbers could not have been based on had already been to Santa Barbara and back collecting plants. actual counts of specimens and may have included estimates Coulter and Douglas apparently collected together around of the number of duplicates of collections. Nelson and Probert Monterey during the winter of 1831/1832 (see FINDINGS (1994: 177) consider these figures greatly exaggerated because below). they were able to account for only 1737 exsiccata numbers of Coulter and Douglas parted company when Coulter left Mon- phanerogams and cryptogams from Mexico and 841 exsiccata terey in late March 1832, traveling south along El Camino Real numbers of phanerogams and cryptogams from California. Nel- likely collecting plants along the way. These included the spec- son and Probert (1994: 177) estimate that Coulter collected not imen cone that would be designated the type specimen of Pinus more than 4000 exsiccata numbers in two series, Mexico and coulteri collected from the Santa Lucia Mountains near the mis- California, and that even if he had collected five specimens sion of San Antonio. He stopped in Santa Barbara on 6 April of each number the total number of specimens would only be 1832 and then went on to San Gabriel Mission, arriving by 23 20,000. As was the custom at the time, duplicates of Coulter’s April 1832. His party continued south along the east side of Californian specimens would likely have been sent to individu- the Santa Ana Mountains past a lake shown on his map (Lake als for their private collections. Subsequently these private col- Elsinore) arriving at Pala 30 April 1832. From there they con- lections were transferred to institutions. Coulter’s Californian tinued southeast possibly along San Felipe Creek and Carrizo exsiccata at Brown University, Providence, Rhode Island (BRU) Creek toward the Colorado River. Coulter and his party arrived came from S. Olney, those at the Gray Herbarium, Harvard Uni- at the ford of the Colorado River about 8 miles south of the junc- versity, Cambridge, Massachusetts (GH) came from A. Gray, tion with the Gila River on 8 May 1832. On 17 May 1832 they many at the Royal Botanic Gardens, Kew, England (K) came set out retracing their route to return to Monterey. They took from W. J. Hooker, and those at the New York Botanical Garden longer on the return, presumably collecting plants. By 27 May Bronx, New York (NY) from J. Torrey. All of these appear to 1832 they were in Pala. They arrived back at San Gabriel on be partial sets, so estimates of possibly five duplicates of each 15 June 1832, apparently making a side trip to the Pueblo de exsiccata number are probably high. I have not seen a num- Nuestra Señora Reina de los Ángeles de Porciúncula (Los An- ber higher than 841 associated with Coulter’s Californian plant geles) and San Pedro. On 5–7 July 1832 they were in Santa specimens. However, there are marine algae specimens bearing Barbara and finally reached Monterey 19 July 1832 (Nelson nine previously unreported exsiccata numbers scattered between 1983). numbers 842 and 872. Coulter left Monterey for the third and last time in late 1832, There is no record of where or when duplicates of Coulter’s heading south through San Luis Obispo, Santa Barbara, San Californian exsiccata were sent nor an indication of the extent Buenaventura, San Fernando to San Gabriel. From San Gabriel and content of those sets of duplicates. There are Coulter Cali- he kept to the coast to San Luis Rey and to San Diego where fornian specimens at BM, BRU, E, FH, GH, K, L, NY, UC and he hoped to secure passage to Mexico. I have seen no record of US. There are reportedly specimens at CGE and G but I have when he arrived at or departed from San Diego. not seen any representative images or database entries. As more VOLUME 37(1–2) Plantae Coulterianae 9

Fig. 6. Vascular plant specimen, Coulter 237, Gaultheria shallon (TCD). 10 Wallace ALISO herbaria and specimens are digitized or otherwise databased and herbaria. Duplicates of most of the marine algae collections are made accessible online, many more specimens of Coulter’s Cal- in other herbaria. Because generally no exsiccata numbers were ifornian exsiccata will likely come to light. Lightner (2014: 30– assigned, each specimen is treated here as a separate collec- 37) lists several dozen Coulter specimens from Kew as represen- tion (Fig. 10). The marine algae specimens have disparate and tatives of plant taxa known to occur in San Diego County. How- occasionally dubious identifications among institutions holding ever, I have found only two specimens, both labeled Isomeris them. Coulter’s marine algae at TCD are listed in Appendix II arborea with St. [sic] Diego, California, on the label, Coulter with comments kindly provided by Kathy Miller, Curator of Al- 65 at K noted by Lightner (2014: 33) and Coulter s.n. at TCD, gae, University of California (UC) at Berkeley. Additional Coul- that were collected in San Diego. ter specimens were located on the websites of various herbaria as well as at the Macroalgal Herbarium Portal. Nomenclature for Vascular Plants and Bryophytes the marine algae follows that in the online resource, AlgaeBase. Coulter’s plant specimens were apparently first identified, then sorted into families and genera. Names applied to the Collection Localities specimens may have come from de Candolle’s Prodromus (de Candolle 1823–1873) and Steudel’s Nomenclator Botanicus For most of Coulter’s specimens no more detailed a locality (Steudel 1840–1841) (Webb 1991: 301). This ordering is evi- than California is indicated on the labels. All of the printed la- dent from the general closeness of the exsiccata numbers within bels affixed to Coulter’s Californian collections have “Califor- the families in Appendix I (e.g., Ranunculaceae, Apiaceae) and nia” on the label (Fig. 7), as opposed to “Mexico” which is on even within genera (e.g., Trifolium in Fabaceae). It is likely that the printed labels for his separately numbered exsiccata from W. H. Harvey wrote the scientific names and sequential exsic- Mexico. At the time Coulter was in California it was still part cata numbers on the labels of the vascular plant and bryophyte of Mexico and as noted above there were no specific state specimens. Other individuals may have identified and labeled boundaries (current states of Arizona and Nevada had yet to be the specimens as well. Duplicates were probably numbered at founded). Some labels have additional locality information as the same time. Some exsiccata numbers on the labels are fol- listed below. There are handwritten labels on some specimens lowed by a notation *, bis, x, xx or xxx. These are treated here that bear abbreviations for California as well as other localities. as representing distinct collections from the original exsiccata- Coulter seems to have traveled along the El Camino Real that numbered specimens because I am uncertain of the meaning of connected the California missions. Beyond “California”, any ad- the notations. It is not clear how or when duplicates were la- ditional locality information was most frequently a description beled and numbered. Specimens bearing exsiccata numbers out of the portion of his route where the specimen was collected. of sequence with those of related taxa were likely identified Less frequently there is a single location cited. I have tried to cite and numbered later. Most of Coulter’s Californian collections the localities exactly as they appear on the labels. Although all bear printed labels (Fig. 7) while others have handwritten labels of the labels I have seen refer to “San Barbara”, this abbreviation (Fig. 8), or identifications and exsiccata numbers written directly was either used by Coulter on his bundle labels or interpreted as on the herbarium sheet. such by Harvey when applying labels to Coulter’s specimens be- Coulter collected over a dozen bryophytes that are among cause Coulter’s only published paper on his travels consistently the earliest known specimens from California (Fig. 9). They uses “Santa Barbara” (Coulter 1835; Dawson 1951). The only were assigned exsiccata numbers sequentially following those localities noted on any of the labels beyond “California” are: of the vascular plants and consequently follow them in Appendix I. I am fortunate that a moss expert, Dr. Daniel Kelly, Between Monterey and El Sur is at TCD. Dr. Kelly was kind enough and interested enough Between Monterey and San Francisco to look over the Coulter specimens and identify them. This is Between Monterey and San Gabriel greatly appreciated as is his forwarding some difficult material Between Monterey and San Miguel to Dr. Len Ellis at the British Museum of Natural History for Between San Miguel and San Barbara (not written out as a second opinion on a few collections and to Dr. D. G. Long Santa Barbara) at Edinburgh for confirmation of the identity of the liverwort. El Sur (a Mexican land grant southwest of Monterey on the As with many of the vascular plant collections of Coulter’s, the coast near Big Sur) specimen of Orthotrichum coulteri Mitt. in the Farlow Herbar- Monterey (also abbreviated Mty., Mont’y, and Monty.) ium (FH) at Harvard University is the lectotype, not the one at Monterey to El Sur TCD. Monterey to San Francisco Monterey to San Gabriel Monterey to San Miguel Marine Algae Santa Barbara (always written on labels as San Barbara) Coulter collected marine algae while he was in Monterey, San Diego (written as St. Diego) California. These were likely of interest to Dr. Harvey, who San Francisco (also abbreviated S. Francisco, S. Fco.) was Coulter’s curatorial successor at TCD. Harvey described San Gabriel four new taxa based on Coulter’s Californian collections. Har- San Miguel vey (1852, 1853, 1858) cited Coulter collections for 28 marine San Miguel to El Sur algae taxa. Five that Harvey cited are not included in Appendix San Miguel and San Barbara (not written out as Santa II because no specimens were located. Specimens for eighteen Barbara) taxa for which a Coulter collection was cited by Harvey (1852, San Miguel to San Barbara (not written out as Santa Barbara) 1853, 1858) were not found in TCD but were located in other San Miguel to San Francisco VOLUME 37(1–2) Plantae Coulterianae 11

Fig. 7. Examples of printed labels affixed to most of Coulter’s California exsiccata. Right: Coulter 252, Malacothrix coulteri (TCD).Left: annotation label referencing “Torr. & Gray, Fl.N.Amer. suppl.,” a work that was never published (see text).

The Herbarium at Trinity College, Dublin (TCD) collections but never published them. I have been unable to lo- cate any copy of a manuscript at Harvard University or Trin- Upon his return to Dublin, Coulter was given rooms at Trin- ity College, Dublin, containing these descriptions. As discussed ity College in 1835, but a change of Provost in 1837 brought an below, Gray apparently used those descriptions in subsequent end to this arrangement (Webb 1991: 298). While having to va- publications of new taxa referring to the Harvey and Gray work cate most of the rooms he occupied he retained one room and as “Harv. & Gray MS” or “Harv. & A. Gray, Pl. Coult. ined.” this is almost certain to have held his herbarium and other col- The latter citation was used for several taxa Gray described in lections (J. Parnell, pers. comm. to GDW 2019). By June 1840 Plantae Fendlerianae (Gray 1849). he and the new Provost of Trinity College Dublin were recon- Harvey distributed some of Coulter’s duplicates to Asa Gray ciled and Coulter obtained new rooms in Staircase 40 (Fig. 11), (now incorporated in GH), William Hooker (now incorpo- and he was made the first keeper of the TCD herbarium (Nel- rated in K), John Ball (some now in TCD) and Stephen Ol- son and Probert 1994: 128). During his tenure as curator Coulter ney (now incorporated in BRU). Referenced correspondence in- purchased several other collections for the herbarium while ar- dicates that Harvey had sent some of Coulter’s specimens of ranging the herbarium according to de Candolle’s Prodromus Leguminosae (Fabaceae) to George Bentham for review (Fig. system (Wright 1896: 4). Robinson (1843: 547) asserted that at 12), and when Bentham returned them Harvey sent back du- the time of his death on 28 November 1843, Coulter had com- plicates in August 1848 (Carter 1982: 334). These are prob- pleted arrangement of the European part of the herbarium and ably the specimens of Fabaceae (Leguminosae) of Coulter’s about of 8000 American specimens [likely an exaggeration],” that Bentham cites for taxa in that family in Plantas Hartwe- …and the remaining packages are well furnished with memo- gianas (1839–1857) published in 1848–1849. Asa Gray visited randa so that for them also the arrangement is practicable.” This Harvey in Dublin in December 1850 (J. Gray 1893: 376). It might explain how, although Coulter’s journals had been lost, it is probable that during this time he looked at some of Coul- was still possible to affix localities, although scant, to many of ter’s specimens. There are several specimens at TCD bear- Coulter’s specimens. ing the annotation label “Fl. N. Amer., Torr. & Gray.” When William Henry Harvey was appointed Keeper of the Trinity Harvey died in May 1866, Alexander Dickson was appointed College Dublin Herbarium on 30 March 1844. During his tenure chair of botany but resigned after just two years (Webb 1991: Harvey identified, or caused to be identified, and labeled Coul- 300). ter’s Californian exsiccata (letter from A. Gray to J. Torrey, 1 Edward Perceval Wright who had assisted Harvey was ap- October 1844, referenced in J. Gray 1893 vol. 1: 327–328). This pointed Professor of Botany in 1869 and petitioned to use the was a remarkable accomplishment because there were no pub- herbarium rooms. He effectively became Curator of the Herbar- lished floras of the western United States at that timeand TCD ium in 1870. The cabinets (called presses) had not been opened likely contained very few if any herbarium specimens from the in the three years since Harvey’s death. Wright supervised a region for comparison. From among Coulter’s collections Har- mandatory move of the herbarium in 1882 from Staircase 40 vey and W. J. Hooker described four new genera and five new where it had been for 50 years to new quarters on the campus species (Harvey 1845a,b, 1846). Harvey and Asa Gray prepared above the Great Entrance Hall (Wright 1896: 9). Many speci- descriptions of many new taxa based on Coulter’s Californian mens not in dust-free cabinets during this time suffered insect 12 Wallace ALISO

mains in these quarters today and the arrangement of most of the specimen folders remains based on Durand’s Index. David Webb was appointed to succeed Dixon in 1949. Webb conducted a systematic sampling of the folders in the herbar- ium and published the results (Webb 1991) summarizing the geographic diversity and concentration of the specimens (e.g., North America, 14,000 specimens). Webb also provided a list of collectors represented in the collection and the geographical ranges of their collecting efforts. Thomas Coulter is listed as one of the larger collections (Mexico, California, France, Italy, Ireland, England, Madeira) represented in the herbarium. Webb was followed by John Parnell in 1983 as TCD Curator. Parnell (1982) lists Coulter as one of the collectors of lichens in TCD but does not mention whether the collections were from Europe, Mexico, or California. Dr. Parnell’s assistance and encourage- ment during this long study has been remarkable and greatly appreciated. The herbarium at Trinity College, Dublin (TCD) is currently largely housed in the aforementioned annex to the Botany Build- ing (Fig. 13). The synoptic Ireland collection and the gen- eral collection of dicotyledons, the monocotyledons (except Poaceae) and the bryophytes are in the main room of the herbar- ium. The Poaceae, gymnosperms, ferns, algae and some of the lichens are together down the hall in another area. The speci- mens are kept in tiers of latching wooden cabinets, called presses by staff (Fig. 14). The banks of cabinets are two tiers high ex- cept in one end of the main room of the herbarium where they are three tiers high but ladders are readily available. The specimen folders within families and genera of di- cotyledons arranged and numbered according to Durand’s Index Fig. 8. Example of a handwritten label affixed to Coulter’s Califor- (Durand 1888) are sequentially followed by those of the mono- nia exsiccatum, Coulter s.n. Oligomeris subulata (TCD). cotyledons and gymnosperms. The ferns, bryophytes, algae and lichens are not included in Durand’s Index and conse- quently lack coincident numbers. Family designations and gen- damage (Wright 1896: 10). It is possible that because of dam- era recognized since Durand’s Index was published are hand- age, some of Coulter’s Californian specimens were discarded. written in a reference copy of the Index kept in the TCD During the relocation of the herbarium and accompanying jum- herbarium. bled mixing of folders, Wright discovered that Harvey, with no Each genus folder bears a reference number assigned from assistant available and to save time, had put newly named (iden- Durand’s Index. Within genera in the general collection there tified) and unnamed specimens in the lower shelves of the cab- are separate numbered and labeled folders for each of 15 geo- inets below the genera to which he considered they belonged. graphical areas of the world (Fig. 15). Californian plant speci- This practice may explain why there are taxa from some families mens are found in the “North America” blue folders which are that bear Coulter’s exsiccata numbers that are out of consecutive marked “11 North America” (Fig. 16). Specimens from Mexico order from the majority of the numbered exsiccata for that fam- are in folders labeled “12 Central America”. The area folders ily. Wright adopted Durand’s (1888) Index Generum Phanero- are arranged in numerical order within each genus. All of the gamorum (Index) as the basis for arrangement and numbering specimens I have seen at TCD are in good condition despite of the families and included genera in the herbarium. Coulter’s their age, their storage in wooden cabinets and their relocation Californian specimens were filed according to the sequence in history noted above. Durand’s Index (Durand 1888). The assigned exsiccata numbers generally follow a sequence within a plant family but there is some deviation among families. This system is still in use in methods TCD. Wright retained his post as curator of the herbarium until his death in 1910. In the TCD herbarium, I looked in most of the North Amer- Henry H. Dixon who had been Wright’s assistant for some ican folders of genera likely to occur in California. Following years took over the curatorship in 1910 upon Wright’s death. the arrangement of families and genera according to Durand’s His contribution to the status of the herbarium, and consequently Index (Durand 1888) was not difficult. Many specimens are Coulter’s collections, was to secure funding in 1912 to construct filed under genera not currently recognized or as having species an annex to the recently built School of Botany to house the from California. I started with the more obvious genera and pro- herbarium (Webb 1991: 301). Dixon transferred the herbarium ceeded to look under genera known from the central coast and to its new quarters and marked each folder with its number as Colorado Desert areas of California. Beyond genera listed in dictated by Durand’s Index. The majority of the collection re- current treatments of the Californian flora, detecting which gen- VOLUME 37(1–2) Plantae Coulterianae 13

Fig. 9. Bryophyte specimen. Above: Coulter 835, Polytrichum juniperinum (moss) herbarium packet (TCD).Below:Coulter 835, Polytrichum juniperinum label and annotation (TCD). 14 Wallace ALISO

Fig. 10. Coulter s.n., Gigartina canaliculata (marine alga) (TCD). era to look in for Coulter’s Californian collections was a chal- examined at TCD. There may yet be some Coulter specimens in lenge. I also checked folders of genera noted in Durand’s Index some of the other genus folders of taxa listed in Watson (1889) (Durand 1888) to occur in California. Over 250 of the genera I that I did not check. I took images of the TCD specimens. considered likely to contain Coulter specimens from California I have relied on data on the specimen labels, annotations, either did not contain Coulter specimens or there were no North and information written directly on the herbarium sheets. On- American folders in TCD. Many of these are old names not now line databases were used to locate Coulter’s Californian exsic- recognized. In some instances, specimens of a particular genus cata at institutions other than TCD. The electronic databases of may be found under a more recently recognized name as well BM, E, GH, K, NY and US were accessed to retrieve data on as an older name. Occasionally nomenclature has changed such Coulter specimens in those herbaria. Bundling databases were that specimens are under an old name that is current again (e.g., also consulted (e.g., Consortium of California Herbaria, Consor- Hosackia). As of this writing I have located 726 of Coulter’s tium of Intermountain Herbaria, Macroalgal Herbarium Portal 841 numbered Californian vascular and nonvascular plant exsic- and Integrated Digitized Biocollections). The online database cata. All but 47 of these are represented in TCD. One hundred- of the Consortium of Intermountain Herbaria provided access fifteen exsiccata numbers are unaccounted for. In an effort to to the Coulter collections at Brown University (BRU). The locate these I considered that many might be misfiled at TCD. BRU specimens were all part of the herbarium of Stephen Ol- Several taxa described below bearing “California” labels were ney bequeathed to Brown University in 1878. These electronic probably collected in Sonora, Mexico, near Guaymas like oth- databases are listed at the end of the References Cited. I relied ers noted in the section: Locality errors California. Knowing this on images of specimens wherever possible to note original la- I considered that additional specimens bearing “California” la- bel text, determinations and annotations. Sometimes database bels might be filed in “North America” folders of genera not entries differed from information on the specimens. Differences known to occur in California. In March 2019, I selected likely included exsiccata numbers, collector, and specimen determi- genera from a listing of plants collected by Edward Palmer near nation. Frequently database entries provide determinations used Guaymas, Sonora, Mexico (Watson 1889). I checked for Coulter for filing of specimens under a current name even though the specimens in several of the folders of genera listed that occurred specimens may not have been specifically annotated as such. in the Guaymas area as well as states in the United States other For database entries lacking an associated image it was not than California. I found no Coulter specimens in these folders possible to be sure of exactly what was on the original label and VOLUME 37(1–2) Plantae Coulterianae 15

Fig. 11. Staircase 40, Trinity College Dublin, Ireland, where Coulter resided as curator and where the TCD herbarium was housed until 1882. in some cases the sequence and content of annotations were un- resenting about 600 minimum-rank taxa (species, subspecies, clear. In the list of exsiccata (Appendix I) this is indicated by varieties). The last figure is more tentative because of likely “[no image]”. In some cases, the name associated with the spec- misidentification of specimens. Coulter’s non-vascular plant imen may have been assigned by the institution or an annotator collections include 10 families, 11 genera, and 13 species of and may or may not have appeared on the original label. Occa- mosses and liverworts. His collections of marine algae comprise sionally incorrect attributions were given to Coulter collections 16 families, 23 genera and 29 species. Among Coulter’s Cal- if they were mounted on the same sheet with a different spec- ifornian collections at TCD are specimens that represent type imen that was entered into the database. Unexplained changes material of 60 vascular plants, one bryophyte, and several ma- made in databases with no associated explanation on the speci- rine algae. Most of these were previously unknown among the men can misinform researchers. Coulter’s Californian specimen TCD collections. Coulter collected specimens of 28 taxa now of Krameria 71* [sic] did not appear in the K database or have considered to be introduced to California. This does not include an accession number. It is mounted on the same sheet as a Bar- those listed below under: Locality Errors California. I found clay collection from Mexico with an accession number that is 892 specimens of Coulter’s Californian collections at TCD.On- listed in the K database. line data for Coulter’s Californian collections accounted for 250 specimens at K, 194 at GH, 133 at BRU,65atUC,36at NY,25atBM and fewer than 10 at each of the herbaria E, F, findings FH, L, MEL, PC, PH,andUS for a total of just over 1600 specimens. Coulter’s Californian Collections Thomas Coulter’s Californian collections represent a floristic Coulter’s Exsiccata Numbers coverage of a portion of California made about 30 years prior to that of the Botanical Survey of California under J. D. Whitney Coulter’s Californian collections of phanerogams and cryp- (Brewer et al. 1876–1880). Coulter’s vascular plant collections togams were assigned exsiccata numbers up to 841. I was able include specimens from about 92 families, 330 genera and rep- to locate 726 numbered exsiccata, 47 of which are only known 16 Wallace ALISO

Fig. 12. Hosackia rigida, sp. nov. Type material likely annotated by George Bentham, Coulter s.n. (TCD). VOLUME 37(1–2) Plantae Coulterianae 17

also possible that some of Coulter’s specimens were damaged by insects and discarded after the move of the herbarium de- scribed by Wright (1896: 10). I located about 300 specimens lacking exsiccata numbers, here assigned “s.n.” (sine numero) designations. Forty-five of these are the only vouchers for taxa in the exsiccata list. About 130 of the 300 Coulter s.n. specimens are at TCD. I do not know the extent to which exsiccata num- bers are on Coulter’s other marine algae specimens. No num- bered marine algae specimens were found at TCD.Acursory search was made of the TCD lichen collections but no Coul- ter Californian specimens were located. If any were assigned exsiccata numbers, they would have been beyond the 841 in series.

Specimen Labels Coulter’s specimens usually bear either printed labels (Fig. 7) or handwritten labels (Fig. 8). I was curious to see if there was a pattern to the disposition of these two label types at TCD and the other institutions holding significant numbers of Coulter speci- mens. I saw all of the labels on the TCD specimens as noted in the exsiccata list. Seventy-one percent of the TCD labels are the Fig. 13. Botany Building, Trinity College Dublin, Ireland. printed labels. For the other collections I relied on online images and databases. Database entries lacking an associated image of the specimen may not provide consistent unambiguous data on from herbaria other than TCD. I found nine exsiccata numbers the nature or content of the original label. Some database entries higher than 841 on some marine algae specimens, none of which indicated that the label was one of the printed labels but when were found at TCD. There are several possible explanations such a notation was missing it was not possible to know if that of the missing exsiccata numbers. Probably more of Coulter’s was an accidental omission or whether the label was a handwrit- Californian specimens are at TCD and as yet unlocated. It is ten label. Consequently, I counted only labels I had seen either

Fig. 14. TCD herbarium with banks of latching wooden cabinets (presses). 18 Wallace ALISO

Fig. 15. TCD specimen folders with reference number 3544 for the genus Aster following Durand (1888) and numbered references to geo- graphical regions of the world.

Fig. 16. TCD specimen folders for Area 11, North America. VOLUME 37(1–2) Plantae Coulterianae 19 in person or by an online image. Ninety percent of the Coulter Locality Errors Californian exsiccata at BRU are imaged and of those 99% have printed labels. Comparable figures for those at K are 98% im- aged and 19% have printed labels. For GH 29% are imaged and Monterey 78% have printed labels. Except for the fact that nearly all of the There are several of Coulter’s Californian specimens with Coulter Californian exsiccata sent to Olney bore printed labels “Monterey” written on the label as the collection locality. How- no pattern of interest was apparent. ever, assuming they are correctly identified, these taxa are not The taxonomic distribution of handwritten labels is inter- known to occur in or near Monterey County and furthermore esting. At TCD all or nearly all of the labels for specimens are taxa generally confined to the interior deserts of southern in Caryophyllaceae, Fabaceae, Onagraceae, Papaveraceae and California. These include (by current names): Ranunculaceae are handwritten labels, while nearly all of the Acmispon rigidus (Benth.) Brouillet Asteraceae specimens at K are handwritten labels. Also, at K Astragalus crotalariae (Benth.) Torr. there are fifteen taxa each represented by two Coulter speci- Astragalus lentiginosus var. borreganus M.E. Jones mens, one with a printed label occasionally with more infor- Dalea mollis Benth. mation and the other a brief handwritten label. This supports in- Dithyrea californica Harv. formation provided by Carter (1982: 334) that Harvey sent the Oligomeris linifolia (Hornem.) J.F. Macbr. Coulter Californian collections of Fabaceae (Leguminosae) to Sphaeralcea coulteri (S. Watson) A. Gray Bentham for review and that Harvey sent a set of duplicates to Bentham when Bentham had returned the reviewed set. I have not found references to the other handwritten labeled specimens California from other families at TCD, having been sent to specialists for Several of Coulter’s specimens that bear printed “California” review. labels represent taxa not known to occur in California. Most or all of these specimens were likely collected about August 1831 near Guaymas, Sonora, Mexico. Coulter collected in that area Annotations before sailing for Monterey, California, possibly after he had sent his Mexican collections to London. Coulter probably took Various labels, considered here annotations, are affixed to the specimens with him to California from where they would Coulter’s specimens. Some are merely property labels or labels have eventually been sent back to London with the bulk of his identifying the specimen as a donation from a private herbar- Californian collections. Coulter’s earlier Mexican collections ium to an institution, e.g., “Herbarium Careyanum” or “Herbar- and the Californian collections arrived in Europe at different ium Olneyanum”. Other labels indicate that the specimen was times. The bundles of Coulter’s specimens were accompanied examined for a project, e.g., “Torr. & Gray, Fl. N. Amer.” or by memoranda which reportedly contained some level of infor- “Syn. Fl. N. Amer.” Often plant names are written on the Herbar- mation. The oversight was likely Coulter’s. “California” labels ium Olneyanum labels at BRU. It is unclear when these labels would have been affixed to them along with the other collec- were affixed to the specimens or who added names tothem. tions received at the time as the basic location of collection. It Almost 80 of Coulter’s Californian specimens at TCD bear is possible that specimens of some taxa in Appendix I known to “Torr. & Gray, Fl. N. Amer.” labels, referring to Torrey and occur in California as well as Mexico, were actually collected Gray (1838–1843). None of the TCD specimens bear Gray’s near Guaymas, Sonora, Mexico (e.g., Lyrocarpa coulteri). Gray labels, “Syn. Fl. N. Amer.,” but there are 21 Coulter specimens (1884: 321, 322, 348, and 361) specifically questioned the lo- at GH that bear these labels. This would seem to indicate that cality designation of California on Coulter’s specimens of Per- Gray was in possession of some of Coulter’s duplicates at that ityle plumigera A. Gray, Perityle leptoglossa A. Gray, Helenium time. thurberi A. Gray, Pectis thurberi A. Gray, and Pectis coulteri A. As stated earlier, Coulter’s collections at TCD remained rel- Gray. Likewise, Keil (1975: 190) brought this issue to light for atively unknown while duplicates at GH and K were better Pectis taxa. Coulter’s specimens of the following taxa (listed by known and frequently served as type material. Coulter’s Cali- current name) bearing “California” labels are known to occur fornian specimens in TCD are still infrequently examined. In near Guaymas, Sonora, Mexico, and were probably collected in an effort to provide a clear picture of the limited numbers of that area: Coulter’s specimens that have been afforded recent examina- tion I assessed the collections known to me from TCD, K, Amsinckia inepta J.F. Macbr. GH,andBRU. For the assessment I only included annota- Brickellia coulteri A. Gray tions bearing an associated date which allows more reliable Chromolaena sagittata (A. Gray) R.M. King & H. Rob. comparisons. Prior to my annotations in 2016 and 2017 re- Cyperus articulatus Benth. garding type status of Coulter’s specimens in TCD, only 4% Helenium laciniatum A. Gray of his collections in TCD had been annotated since 1900 and Helenium puberulum DC. those were annotated between 1982 and 2015. Including my Helenium thurberi A. Gray recent annotations this figure is 9.5%. By comparison, since Holographis virgata (Benth. & Hook.) T.F. Daniel 1900, 2% of those at BRU have been annotated, 36% of those Ibervillea sonorae (S. Watson) Greene at K, and 56% of those at GH. Annotations dated in the past Mimosa polyantha Benth. 20 years are 2% for TCD (excluding my annotations of type Pectis coulteri Harv. & A. Gray ex A. Gray status in 2016 and 2017), 2% for BRU,4%forK and 9% Pectis filipes Harv. & A. Gray ex A. Gray for GH. Perityle californica Benth. 20 Wallace ALISO

Perityle leptoglossa Harv. & A. Gray ex A. Gray Fendlerianae (Gray 1849) while others were never validly pub- Perityle microglossa Benth. lished. Ruellia occidentalis (A. Gray) Tharp & F.A. Barkley

References to Publications That Were Never Issued Coulter’s Map of Upper California (Coulter 1835)

Harv. & A. Gray or Harv. & A. Gray, Pl. Coult. Thomas Coulter was the first botanist to cross the California desert and collect plants (Brewer et al. 1876–1880, vol. 2: Throughout this paper are references to scientific names with 555; McKelvey 1955: 434). According to the latter reference the citation “Harv. & A. Gray” or some abbreviated version of he “ …set foot—we must believe—in [present day] Arizona”. this. This is found associated with names on Coulter’s exsic- The original map is the most informative and reliable but the cata labels as well as with published accounts currently recog- most difficult to consult. Electronic versions are often not sharp nized here (e.g., Baileya pleniradiata and B. pauciradiata both enough to see the fine details. I have referred to four versions of cited in Turner 2006: 446). William H. Harvey (Trinity College, Coulter’s map and believe they warrant mention here. Dublin) and A. Gray (Harvard) were apparently collaborating on naming and describing new taxa based on Coulter’s speci- Coulter’s (1835) Map mens from California. I have found no indication that these de- Figure 2 is an image of the complete map. It is an 8 × 10- scriptions were ever assembled as a separate manuscript or that inch map in the original article. Figures 3–5 are detail sections such a manuscript was ever published. Torrey recognized that of the map. Coulter’s routes are indicated by narrow lines. There this work had not been published. He published the description are notations of a small “x” at the sites of the missions where he of the genus Baileya as “Baileya n. gen. Harv. and Gr., ined.” presumably stopped as well as some villages (e.g., San Felipe). (Torrey 1848: 145). Gray also recognized that these names and The “x” marks at unlabeled sites seem to coincide with the lo- descriptions were not yet published when he subsequently pub- cations of Mission Nuestra Señora de la Soledad, Mission San lished them for some species in his Plantae Fendlerianae (Gray Antonio de Padua, Mission San Miguel Arcángel, Mission San 1849) citing as reference “Harv. & Gray, in Pl. Coult. ined.” di- Juan Capistrano and Santa Ysabel Asistencia. There are small rectly or by reference with each of the descriptions. It is clear “x” marks on the west side of the Colorado River at Ford and that Harvey made no direct contribution to Plantae Fendlerianae at San Pedro as well as San Pablo [misspelled as Parblo on the and that Gray was acknowledging their previously prepared de- map] east of the junction of the Gila River with the Colorado scriptions. All “Harv. & A. Gray” references related to descrip- River. Taken in context, the notations on his map seem to indi- tions in Gray (1849) should read “Harv. & A. Gray ex A. Gray” cate that it is possible Coulter crossed the Colorado/Gila rivers or just “A. Gray”. I have chosen to use the longer version to into present-day Arizona. The lines depicting Coulter’s travel show the historical context of the nomenclatural combinations. routes do not extend north from the Ford on the Colorado River This applies to several taxa in the exsiccata list (Appendix I) to San Pedro and San Pablo although there are “x” marks co- (e.g., Acamptopappus sphaerocephalus, Baileya pauciradiata, incident with these locations. Coulter’s 1835 map also shows B. pleniradiata, Chaenactis artemisiifolia, Cirsium occidentale multiple routes between San Buenaventura and San Fernando, var. coulteri, Malacothrix coulteri, Pectis coulteri, P. filipes, P. side trip routes to the Pueblo of Los Angeles and nearby San papposa and Perityle leptoglossa. Pedro, and either a side trip or a trial route between San Luis Obispo and the upper Guadalupe River ( = Santa Maria River). Torr. & Gray, Fl. N. Amer., Suppl. This map does not show any route to San Francisco even though Several of Coulter’s Californian exsiccata specimens in TCD some of his specimens are labeled San Francisco. As noted by and GH bear labels indicating that the specimen represents a Nelson and Probert (1994: 159), Coulter commented that he had new taxon attributed to Harv. & Gray and published in “Torr. not been to the north of San Francisco. However, that does not & Gray, Fl. N. Amer. Suppl.” (Fig. 7). All of these taxa are in mean that he did not get as far as San Francisco. Asteraceae. No such publication has been found. The Torrey and Gray Flora of North America came out in two volumes. Volume Dawson’s (1951) Reprinting of Coulter’s 1835 Map I published in parts between 1838 and 1840 does include a “Sup- Dawson includes a narrow but complete facsimile of Coulter’s plement, additions and emendations” (pp. 657–698, published 1835 map. It is sewn into his small book between pages 30 and in June 1840) but no Asteraceae are in that volume. Volume 31. In neither of the two copies I have seen of this work are most II published in parts between 1841 and 1843 does not include of the lines depicting Coulter’s travel routes visible. However, a supplement but pages 502–504 published in February 1843 most of the “x” notations from the original are visible. are identified as “Additions to Compositae”. Not surprisingly none of Coulter’s Californian exsiccata taxa are cited in that sec- Bancroft (1885, Vol. 3: 407) Version of Coulter’s Map tion, likely because his specimens had not yet been distributed. This map was redrawn from Coulter’s map with labeling and Gray recognized that the names attributed to “Torr. & Gray, Fl. route lines differing from the original. It does not include the N. Amer. Supp.” had never been published. This is clear from southern portion of Coulter’s map south of San Diego and San his notation in Plantae Fendlerianae under Polypappus sericeus Felipe or any reference to the Colorado River. The “x” nota- Nutt. [now Pluchea sericea Nutt.] where Gray cites as synony- tions are changed to “o” for the mission sites but all others are mous his own combination of Tessaria as “(T. borealis Torr. & missing. Gray, Fl. Suppl. Compos. ined.)” (Gray 1849: 75). Another in- stance is under Riddellia tagetina “Torr. & Gray, Fl. 2: 362, & Nelson and Probert (1994: 101) Suppl. Compos. ined.” (Gray 1849: 93). Several taxa with these On this copy of Coulter’s 1835 map, the lines depicting Coul- labels were validly published in other works such as Plantae ter’s travel routes are clear. The “x” marks for the missions and VOLUME 37(1–2) Plantae Coulterianae 21

Fig. 17. Type material of Malacothrix coulteri Harv. & A. Gray ex A. Gray, Coulter 252 (TCD). 22 Wallace ALISO villages are mostly not apparent. The accompanying redrawn ter” or “T.C.” Several database entries mistakenly attribute col- version is helpful and does indicate most of the notations on the lections to J. M. Coulter. This is John Merle Coulter (1851– original map. 1928) but the dates and localities of collections make discrimi- nation of the Coulters relatively easy. Data about Coulter speci- mens may be confused by other collections mounted on the same Notes sheets. Coulter specimens mounted on sheets with other collec- Thomas Coulter’s specimens have been designated type ma- tions are occasionally filed under the name of other specimens. terial (Figure 17) for about 60 plant taxa. Type specimens are Likewise, images may be associated with one of the non-Coulter usually the first specimens of a taxon ever collected. As noted specimens on the sheet. earlier, Coulter’s Californian collections remained relatively un- known for about 10 years after his death because the duplicates had not yet been distributed. Consequently, later collectors gath- acknowledgements ered specimens of some of the same taxa that would become type specimens of taxa first collected by Coulter. Sixteen such First, I must thank Dr. John Parnell of the Department of taxa are noted as being first collected by Coulter, Coulter and Botany at Trinity College, Dublin, for his generosity of infor- Douglas, or Coulter and Nuttall in Gray’s Synoptical Flora of mation and assistance in all aspects of this multi-year endeavor. North America (Appendix V). As evidence that Douglas’s speci- Access to the herbarium on my numerous trips to Dublin made mens were more readily known than Coulter’s, even though both this work possible. The assistance and enthusiasm of Dr. Daniel men collected at about the same time in California, Torrey and Kelly, recently retired from the Botany Department of Trinity Gray (1838–1843) make reference to over 300 Douglas collec- College, was a pleasant and rewarding experience. He brought tions but no mention of any Coulter specimens. It was probably the bryophytes up to date with confirmed identifications, even several years later that Gray saw a Coulter specimen. Bentham enlisting the aid of his colleagues Drs. Len Ellis at BM and D. G. (1839–1857) also mentioned specimens collected by Coulter in Long at E. The kindness of the staff is also greatly appreciated, California and Mexico in his enumeration of species collected as was the assistance and pleasure of invitation to tea from Mrs. by Theodor Hartweg in Mexico. Sixty-one vascular plants (22 Marcella Campbell. Clarification of taxonomic and specimen from California), one moss and four marine algae are named for questions for particular taxa were provided by Naomi Fraga, Thomas Coulter (Appendix IV). Most of these were first noted David Keil, Walter Kittredge, J. Mark Porter, Fred Roberts, Jr., by Coville (1895). and Gordon Tucker. Many identifications of images of speci- It is known that Coulter met David Douglas while in Mon- mens in TCD were suggested by J. Travis Columbus, LeRoy terey. They may have collected together near Monterey, Califor- Gross and Victor Steinmann. Taxonomic and bibliographic as- nia. There are some sheets at TCD with a Coulter collection and sistance was provided by David Boufford, J. Travis Columbus, a Douglas collection of the same species mounted side by side. Kanchi Gandhi, Irene Holiman, Walter Kittredge, Simon Lang As an example, there is a sheet at TCD with two collections of and John Parnell. Charles Nelson and John Parnell provided Dendromecon rigida on it. Dendromecon rigida Benth. is writ- valuable background information on Thomas Coulter and his ten at the bottom of the sheet. Both specimens have handwrit- association with Trinity College, Dublin. Kathy Ann Miller pro- ten labels. One reads “Monterey, California T. Coulter 1832” vided significant clarifications to the list of marine algae inAp- and the other reads “California D. Douglas”. There is a simi- pendix II. In their reviews of the manuscript, Charles Nelson and larly composed sheet of Viola pedunculata at TCD.AtK there John Parnell provided information that greatly improved the ac- are sheets of Notholithocarpus densiflorus (Quercus densiflora) curacy and scholarship of this paper. A warm thank you to the and Cryptantha (Myosotis) muricata with Douglas and Coulter barmen at The Stag’s Head and The Ginger Man in Dublin for specimens mounted side by side although neither has more than serving up a pint or two of the black stuff at the end of the day California for a locality. in the herbarium and contributing to the always friendly atmo- Specimens lacking exsiccata numbers are identified here as sphere of Ireland. Thank you to the late Reid Moran for showing “s.n.”. Harvey may have failed to affix numbers to them or they us all the full breadth of possibilities and importance of acknowl- got separated from the numbered specimens of that taxon. Mul- edgements (Moran 1962). Editorial guidance prior to submis- tiple “s.n.” specimens attributed to a particular taxon may repre- sion of the manuscript was graciously provided by Vanessa Ash- sent duplicates of a single collection or unicates of two or more worth. A special thanks to my wife, Marianne, for her patience, collections. In a few instances specimens bearing the same ex- interest and understanding during our many visits to Dublin to siccata number are reported to be different taxa, e.g., Coulter look at and for Coulter specimens at TCD. 744 in TCD is Toxicoscordium (Zygadenus) while Coulter 744 in GH is Iris. There are instances where the handwritten exs- iccata number has been misread and entered into a database or literature cited where the database indicated that the specimen had some type status but the specimen bore no annotation to that effect. I sus- Abrams, L. 1923–1960. Illustrated flora of the Pacific States, vols. 1–3, pect this might be the reason for the GH exsiccata specimen vol. 4 with R. S. Ferris. Stanford University Press, Stanford, Califor- nia. of Iris, of which no image was provided: the Iris specimen is Ball, R. 1846. The Dublin University Museum, Jan 1846. Dublin Uni- entered in the GH database as Coulter 744 instead of Coulter versity Press, Dublin, Ireland. 16 p. 724 as written on the TCD Iris specimen label. These sorts of Bancroft, H. H. 1885. History of California vol. III. 1825–1840. A. discrepancies are rare when images of labels are available. Al- L. Bancroft & Co., San Francisco. though the printed labels read “Dr. Coulter” the attributions on Barneby, R. C. 1964. Atlas of North American Astragalus.Memoirs the handwritten labels variously reference “Dr. Coulter”, “Coul- of the New York Botanical Garden 13: pt. I: 1–596, pt. II: 597–1188. VOLUME 37(1–2) Plantae Coulterianae 23

Beeler, M. S. 1970. Harmony in Chumash. International J. Amer. Lin- ———. 1884. Synoptical flora of North America, vol. I. The Gamopeta- guistics 36(1): 14–17. lae, part II. Smithsonian Institution, New York. Bentham, G. 1839–1857. Plantas Hartwegianas [alternative title: Plan- ———. 1888. Synoptical flora of North America. The Gamopetalae. A tae Hartwegianae]. Gulielmus Pamplin, London, UK. second edition of vol. I, part II, and vol. II, part I, collected. Smith- ———. 1844. The botany of the voyage of the H.M.S. Sulphur, un- sonian Misc. Publ. 591. Washington, D.C. der the command of captain Sir Edward Belcher … during the years ———. 1897. Synoptical flora of North America, vol. I, part I, fascicle 1836–42. Smith, Elder and Co., London, UK. II, Caryophyllaceae to Polygalaceae. University Press, Cambridge, Brewer, W. H., S. Watson and A. Gray. 1876–1880. Botany [of Cal- Massachusetts. ifornia], 2 vols. Cambridge, Massachusetts. ———, S. Watson and B. L. Robinson. 1895–1897. Synoptical flora Brummitt, R. K. and C. E. Powell (editors). 1992. Authors of plant of North America. Polypetalae from the Ranunculaceae to the Poly- names. Royal Botanic Gardens, Kew, UK. 732 p. galaceae, vol. I, part I. American Book Company, Cambridge, Mas- Carter, A. 1982. Lectotypification of Cercidium floridum (Legumi- sachusetts. nosae: Caesalpinioideae). Taxon 31(2): 333–335. Gray, J. (editor). 1893. The letters of Asa Gray, 2 vols. Houghton Mif- Coulter, T. 1832. Letter dated 16 May 1832 written from the desert flin & Co., Boston, Massachusetts. of southeastern California by Coulter to his mentor Alphonse de Harvey, W. H. 1845a. Description of a new genus of Papaveraceae, Candolle in Geneva transcribed by Nelson and Probert (1994: detected by the late Dr. Coulter, in California. London J. Bot. 4: 106–108). 73–76, Tab. III. ———. 1835. Notes on Upper California. Communicated by Dr. ———. 1845b. Characters of two new genera of Cruciferae, discovered Thomas Coulter. Read 9th March 1835. J. Royal Geog. Soc., Lon- by the late Dr. Coulter, in California. London J. Bot. 4: 76–78, Tab. don. Proc. Roy. Geogr. Soc. London 5: 59–70, 1 map. IV & V. Coville, F. V. 1895. The botanical explorations of Thomas Coul- ———. 1846. Description of a new genus of Hydrophyllaceae, from ter in Mexico and California. Bot. Gaz. 20(12): 519–531. [In- California. London J. Bot. 5: 311–312, Tab. XI & XII. cludes copy of Coulter’s map and a list of species named for ———. 1852. Nereis Boreali-Americana: or Contributions to a history Coulter]. of the marine algae of North America. I Melanospermae. Smithso- Dawson, G. 1951. Notes on upper California: a journey from Monterey nian Contr. Knowl. (preprint) 3(4): 1–150, 12 pl. to the Colorado River in 1832 by Dr. Thomas Coulter. Printed with- ———. 1853. Nereis Boreali-Americana: or Contributions to a history out changes or additions. Los Angeles, California. of the marine algae of North America. II Rhodospermae. Smithsonian De Candolle, A. P. (editor). 1823–1873. Prodromus systematis natu- Contr. Knowl. 5(5): 1–258, 36 pl. ralis regni vegetabilis, 17 vols. Paris, France. ———. 1858. Nereis Boreali-Americana: or Contributions to a history Don, D. 1836. Descriptions of five new species of the genus Pinus, of the marine algae of North America. III Chlorospermae. Smithso- discovered by Dr. Coulter in California. Trans. Linn. Soc. London nian Contr. Knowl. 10(2): 1–140, 49 pl. 17: 439–444. Hemsley, W. B. 1879–1888. Biologia Centrali-Americana. 3 vols. Lon- Durand, T. 1888. Index generum phanerogamorum. Usque ad finem don, UK. anni 1887 promulgatorum in Benthami et Hookeri “Genera Plan- Keil, D. J. 1975. Revision of Pectis sect. Heteropectis (Compositae: tarum” fundatus. Dulau & Co., London, UK. Tageteae). Madroño 23(4): 181–190. Engelmann, G. and A. Gray. 1845. Plantae Lindheimerianae; an enu- Lightner, J. 2014. San Diego native plants in the 1830s. The collec- meration of Lindheimer’s collection of Texan plants … Freeman & tions of Thomas Coulter, Thomas Nuttall, and H.M.S. Sulphur with Bolles, Boston, Massachusetts. George Barclay and Richard Hinds. San Diego Flora, San Diego, Cal- Farjon, A. 2010. A handbook of the world’s conifers, vol. 1. Brill, Lon- ifornia. don, UK. McKelvey, S. D. 1955. Botanical exploration of the Trans-Mississippi ———. 2017. A handbook of the world’s conifers, vol. 1, 2nd revised west. Arnold Arboretum of Harvard University, Jamaica Plain, Mas- edn. Brill, London, UK. sachusetts. Gray, A. 1849. Plantae Fendlerianae Novi-Mexicanae: an account of McVaugh, R. 1943. The travels of Thomas Coulter (1824–1827). J. a collection of plants made chiefly in the vicinity of Santa Fe, New Washington Acad. Sci. 33: 65–70. Mexico, by Augustus Fendler; with descriptions of new species, crit- Moran, R. 1962. Cneoridium dumosum (Nuttall) Hooker f. collected ical remarks, and characters of other undescribed or little-known 26 Mar 1960, at an elevation of about 1400 meters on Cerro Que- plants from the surrounding regions. Mem. Amer. Acad. Arts,new mazón, 15 miles south of Bahía de Los Angeles, Baja California, series, vol. 4, pt. 1: 1–116. Mexico, apparently for a southeastward range extension of some 140 ———. 1852. Plantae Wrightianae Texano–Neo-Mexicanae: an ac- miles. Madroño 16(8): 272. count of a collection of plants made by Charles Wright, A.M., in an Nelson, E. C. 1983. Thomas Coulter (1793–1843) in North America: expedition from Texas to New Mexico, in the summer and autumn of some bibliographic problems and some solutions. In Wheeler, A. 1849, with critical notices and characters of other new or interesting [ed.], Contributions to the history of North American natural history. plants from adjacent regions etc. Part I. Smithsonian Contr. Knowl. Society for the Bibliography of Natural History, London, UK. 3, Art. 5: 1–146, Tab. 1–10. ———. 1988. Trinity’s miner-botanist Dr. Thomas Coulter (1793– ———. 1854. Plantae Novae Thurberianae: the characters of some new 1843). Hermathena 145: 7–21. genera and species of plants in a collection made by George Thurber, ——— and A. Probert. 1994. A man who can speak of plants. Dr. Esq., of the late Mexican Boundary Commission, chiefly in New Thomas Coulter (1793–1843) of Dundalk in Ireland, Mexico and Mexico and Sonora. Mem. Amer. Acad. Arts,newseries,vol.5,pt.2: Alta California. Privately published by E. Charles Nelson, Dublin, 297–328. Ireland. ———. 1864a.OnStreptanthus Nutt., and the plants which have been Parnell, J. A. N. 1982. The lichen herbarium of Trinity College, referred to that genus. Proc. Amer. Acad. Arts 6: 182–188. Dublin. Lichenologist 14: 280–281. ———. 1864b. A revision and arrangement (mainly by the fruit) of the Powell, A. M. 1974. Taxonomy of Perityle section Perityle North American species of Astragalus and Oxytropis. Proc. Amer. (Compositae—Peritylinae). Rhodora 76: 229–306. Acad. Arts 6: 188–236. Renkema, H. W. and J. Ardagh. 1930. Aylmer Bourke Lambert and ———. 1878–1897. Synoptical flora of North America, vol. I, part II, his ‘Description of the genus Pinus’. J. Linn. Soc., Bot. 48: 439–466. and vol. II, part I, & supplement, 2nd edn. (1886), collated in: Smith- Robinson, T. R. 1843. Biography of Dr. Coulter. Proc. Roy. Irish Acad. sonian Misc. Collect. 31 (1888). 2: 553–557. 24 Wallace ALISO

Scouler, J. 1841. Observations on the indigenous tribes of the N. W. appendix i coast of America. J. Roy. Geogr. Soc. 11: 215–251. coulter’s californian vascular and nonvascular Steudel, E. G. 1840–1841. Nomenclator botanicus, 2nd edn., 2 vols. plant exsiccata J. G. Cotta, Stuttgart, Germany. Torrey, J. 1848. Appendix by Professor Torrey, pp. 137–155, pl. 1–12. This list of Coulter’s Californian exsiccata includes the vascular plant In W. H. Emory, Notes on a reconnaissance from Fort Leavenworth, taxa followed by the mosses. The vascular plant families are listed in alpha- in Missouri, to San Diego, in California. Wendell and Van Benthuy- betical sequence starting with ferns followed by gymnosperms, magnoliids, sen, Washington, D.C. eudicots, and monocots with genera in alphabetical sequence within each ——— and A. Gray. 1838–1843. A flora of North America, 2vols. family. Wiley & Putnam, New York. Going on the optimistic assumption that the original determinations were Turner, M. W. 2006. Baileya, pp. 444–447. In Flora of North America correct, the specimens are listed under their currently recognized names. Editorial Committee [eds.], Flora of North America north of Mexico, Some of the specimens have been annotated since that time. In most cases vol. 21, Magnoliophyta: Asteridae, part 8: Asteraceae, part 3. Oxford the current name was determined by simple taxonomic equivalence, under- University Press, New York. standing full well that many specimens may have been originally misidenti- Watson, S. 1889. XVI Contributions to American Botany. 1. Upon a fied. Online databases (e.g., Tropicos and IPNI [International Plant Names collection of plants made by Dr. E. Palmer, in 1887, about Guaymas, Index]) as well as Abrams (1923–1960) and the current volumes of the Flora Mexico, at Muleje and Los Angeles Bay in Lower California, and on of North America were among the references useful in tracking older names. the Island of San Pedro Martin in the Gulf of California. Proc. Amer. Complications arising from the use of an older taxonomic scheme occasion- Acad. Arts 24: 36–82. ally necessitated looking for specimens under genera that were one or two iterations removed from their currently recognized taxonomic status. Place- Webb, D. A. 1991. The herbarium of Trinity College, Dublin: its history ment of specimens under these current names is not intended to be an annota- and contents. J. Linn. Soc., Bot. 106: 295–327. tion or acceptance of the original determinations but rather a convenience of Wright, E. P. 1896. The herbarium of Trinity College Dublin: a retro- organization. Specimens from herbaria other than TCD for which no image spect. Notes Bot. School Trinity Coll. 1: 1–14. was available are listed in accordance with the name provided in the insti- tution’s database. Specimens seen in TCD that lack a name are generally listed as “sp.” under the genus in which the specimen was filed. Similarly, if a currently recognized infraspecific taxon could not be determined orwas an innovation since the mid-1800s, the specific epithet is followed by just database references “subsp.” or “var.” An asterisk (*) preceding a taxon’s name indicates that the specimen was AlgaeBase: http://www.algaebase.org/search/species/ likely collected in Sonora, Mexico, by Coulter before he arrived in California Biodiversity Heritage Library: https://www.biodiversitylibrary.org but distributed with his “California” label (see Locality errors California). Conservatoire et Jardin botaniques Genève, Geneva Herbaria Cat- Some specimens that bore names of taxa that were conspicuously out of alogue: http://www.ville-ge.ch/musinfo/bd/cjb/chg/index.php?lang = en range have occasionally been relegated to the unverified notations (e.g., sp.) Consortium of California Herbaria (CCH): http://ucjeps.berkeley. above. edu/consortium/ Species entries start with a current name and authorities, all in bold. Au- Consortium of North American Bryophyte Herbaria: http:// thor abbreviations follow those in Brummitt and Powell (1992). Names of bryophyteportal.org/portal/index.php introduced taxa are prefixed by a dagger (†). Species entries are followed by Consortium of Northeastern Herbaria: http://portal.neherbaria.org/ “Coulter” and an exsiccata number or “s.n.” (sine numero) underlined. Any portal/collections/list.php?db = all&collector = Coulter&page = 2 applicable abbreviation for Dr. Thomas Coulter on the specimen is written Google Books: https://books.google.com here as “Coulter”. I assigned the “s.n.” notation—it does not appear on the Harvard University Herbaria and Libraries: https://kiki.huh.harvard. exsiccata labels. If there is no exsiccata number on the label, it is left blank. edu/databases/ Multiple collections of the same taxon are arranged in ascending numerical Index Herbariorum: http://sweetgum.nybg.org/science/ih/ order of the exsiccata followed by any “s.n.” specimens. Each specimen of Integrated Digitized Biocollections (iDigBio): https://www.idigbio. this latter group is listed separately because there is no way to determine org/portal whether they represent duplicates of the same collection. Intermountain Region Herbarium Network: http:// Next are the acronyms, in bold, of herbaria housing a specimen bear- intermountainbiota.org/portal/collections/index.php ing that exsiccata number, or s.n. These start with TCD,followedbyany International Plant Names Index (IPNI): http://www.ipni.org/ other herbaria in alphabetical order of their acronyms. Herbaria identified ipni/plantnamesearchpage.do by acronyms may be found in Index Herbariorum online (see Database Macroalgal Herbarium Portal: http://macroalgae.org/portal/ References). collections/list.php Information on the labels was recorded mostly verbatim to what was writ- Natural History Museum (British Museum): https://data.nhm.ac.uk/ ten on the original labels. I did not use quotation marks because it would dataset/56e711e6-c847-4f99-915a-6894bb5c5dea/resource/05ff2255- render the text unreasonably cluttered and difficult to read. Abbreviations, c38a-40c9-b657-4ccb55ab2feb?view_id = 203a0ae5-6a14-480a-a407- initials and spelling were written as they appear on the original labels, except 27eeb9373858&filters = collectionCode%3Abot for “California”. Occasionally I wrote out the word in brackets. However, Naturalis Biodiversity Center: https://bioportal.naturalis.nl/?language the information was arranged in a consistent format for the entries as fol- = en&back lows: name or names of the specimen with any author citation as provided, New York Botanical Garden, Steere Herbarium: http:// followed by California, then any additional locality reference, and any ref- sweetgum.nybg.org/science/vh/ erence to a date of collection. Next, I added one of two abbreviations, either Royal Botanic Garden, Edinburgh, Herbarium Catalogue: https:// “PL” meaning that the specimen had a printed label or “SL” meaning that data.rbge.org.uk/search/herbarium/ the specimen had a handwritten label. If no name was indicated on the label Royal Botanic Gardens, Kew, Herbarium Catalogue: http:// “[no name]” is inserted in the name position. apps.kew.org/herbcat/navigator.do After information from the label, information from annotation labels fol- Smithsonian National Museum of Natural History, Botany Col- lows “Annotation 1:”. This information is nearly verbatim. These labels are lections: https://collections.nmnh.si.edu/search/botany/ physically or stylistically associated with the labeling of the specimen. In Tropicos: https://www.tropicos.org/ some cases, there are also simple printed labels, e.g., “Fl. N. Amer., Torr. & VOLUME 37(1–2) Plantae Coulterianae 25

Gray” affixed to the sheet, presumably indicating that Gray saw the speci- Coulter 822. men at some point. Following annotations there are occasionally references BRU: Adiantum capillus-veneris, California [no image]. PBRU to “Written on sheet:”. This precedes plant names or notes handwritten at 00064754. the bottom of the sheet lacking an annotator attribution and typically apart from any other labeling. These are possibly names under which the spec- Adiantum aleuticum (Rupr.) C.A. Paris imens were to be filed. Lastly is a reiteration of the herbarium’s acronym Coulter 822. followed by an identification number if available. TCD has identification TCD: [no name], California (PL). TCD! numbers only for those few specimens that have been digitized. All notations Aspidotis californica (Hook.) Copel. of “TCD!” indicate that I saw the specimen in the Trinity College herbar- ium, Dublin, Ireland. Additional information about some entries is included Coulter 820. TCD Hypolepis californica under the heading “Note”. : Hook., California (PL). Annotation 1. ISOSYNTYPE Hypolepis californica Hooker by G. Wallace in 2016. Written on sheet: Aspidotis californica. TCD! GH: Cheilanthes coulteri [sic], California (PL). Annotation 1: ISOSYN- FERNS TYPE Hypolepis californica Hook. and = Aspidotis californica (Hook.) blechnaceae Copel. by Kittredge in 2014. GH 00518245. K: Cheilanthes coulteri Harv., California (PL). Annotation 1: Woodwardia fimbriata Sm. TYPE SPECIMEN. Written on sheet: Hypolepis californica Hook. Coulter 825. K 001057971. TCD: Woodwardia radicans, California (PL). TCD! BRU: Woodwardia radicans, California [no image]. PBRU 00067177. Pellaea andromedifolia (Kaulf.) Fée Coulter 823. TCD: [no name], California (PL). TCD! dryopteridaceae BRU: [no name], California [no image]. PBRU 00066095. Dryopteris arguta (Kaulf.) Maxon Pellaea mucronata (D.C. Eaton) D.C. Eaton var. californica (Lemmon) Coulter 826. Munz & I.M. Johnst. TCD: [no name], California (PL). TCD! Coulter 824. TCD: [no name], California (SL). TCD! TCD: [no name], California (PL). TCD! BRU: Dryopteris felix-mas, California [no image]. PBRU 00072928. Pentagramma triangularis (Kaulf.) Yatsk., Windham & E. Wollenw. Polystichum munitum (Kaulf.) C. Presl subsp. Coulter s.n. Coulter 817. TCD: [no name], California (PL). TCD! TCD: [no name], California (PL). Written on sheet: Pentagramma. TCD! TCD: [no name], California (PL). TCD! Coulter 827. TCD: [no name], California (SL). TCD! BRU: Pentagramma triangularis [no image]. PBRU 00026845. BRU: Polystichum munitum [no image]. PBRU 00073125.

GYMNOSPERMS equisetaceae cupressaceae Equisetum hyemale L. subsp. affine (Engelm.) Calder & R.L. Taylor Coulter 829. Calocedrus decurrens (Torr.) Florin TCD: [no name], California (SL). TCD! Coulter 710. TCD: [no name], California (PL). TCD! Equisetum telmateia Ehrh. subsp. braunii (J. Milde) Hauke Coulter s.n. Coulter 830. TCD: [no name], California (PL). TCD! TCD: [no name], California (PL). TCD! BRU: [no name], California [no image]. PBRU 00064186. ephedraceae marsileaceae Ephedra sp. Coulter s.n. Marsilea vestita Hook. & Grev. TCD: Ephedra, California (PL). TCD! Coulter 828. TCD: M. vestita H. & G., California (PL). TCD! pinaceae polypodiaceae Pinus coulteri D. Don Coulter s.n. Polypodium californicum Kaulf. TCD: [no name]. Annotation 1: Pinus coulteri D. Don, Trans. Linn. Soc., Coulter 818. London 17: 440. 1836. Coll. T. Coulter 1832/33. U.S.A., California, Sta. TCD: [no name], California (PL). TCD! Lucia Mtns., Cone Peak, by Farjon in 1995. TCD! Note: cone only. Coulter 819. TCD: [no name], California (PL). TCD! Pinus muricata D. Don Coulter 712. pteridaceae TCD: P. bracteata??, P. muricata??, California (PL). Note attached to stem: cone small hooked near Sta. San Luis Obispo. Annotation 1: Pinus Adiantum capillus-veneris L. muricata D. Don Trans. Linn. Soc. London 17: 441. 1936. ISOTYPE col- Coulter 821. lected 1833? Coon Creek, San Luis Obispo County, California, U.S.A. TCD: [no name], California (SL). TCD! by Farjon in 1995. Annotation 2: Pinus muricata D. Don var. muricata 26 Wallace ALISO

by J.A. Perez De Larosa in 2001. Annotation 3: TYPUS. TCD! Note: see adoxaceae Appendix III for discussion of this specimen. Sambucus nigra L. subsp. caerulea (Raf.) Bolli Coulter 211. TCD: [no name], California (PL). TCD! ANGIOSPERMS MAGNOLIIDS amaranthaceae lauraceae †Alternanthera caracasana Kunth Umbellularia californica (Hook. & Arn.) Nutt. Coulter 681. Coulter 708. TCD: [no name], California (PL). TCD! GH: Umbellularia californica, California. [no image]. GH 381286. Coulter 709. Amaranthus sp. GH: Tetranthera sp., California [no image]. GH 381277. Coulter 682. K: [no name], California (SL). Written on sheet: Oreodaphne califor- TCD: Amaranthus, California (PL). TCD! nica Nees, Tetranthera californica Hook. & Arn., DC. Prod. XV, 192. K 000815469. anacardiaceae Malosma laurina (Nutt.) Abrams saururaceae Coulter 238. Anemopsis californica (Nutt.) Hook. & Arn. TCD: Rhus laurina, California (SL). TCD! [on same label as 549 below] Coulter 702. Coulter 549. TCD: Anemopsis, California (PL). TCD! TCD: Rhus laurina, California (SL). TCD! [on same label as 238 above] Rhus aromatica Aiton Coulter 127. EUDICOTS TCD: Rhus, California (PL). TCD! acanthaceae Coulter 128. TCD: Rhus, California (PL). TCD! *Holographis virgata (Harv. ex Benth. & Hook. f.) T.F. Daniel Coulter 130. Coulter 603. TCD: Rhus, California (PL). TCD! TCD: Berginia virgata, California (PL). Annotation 1: Holographis vir- gata subsp. virgata, ISOTYPE Berginia virgata Harv. ex Benth. & Hook. Rhus ovata S. Watson by Daniel in 2015. TCD! Coulter 550. BM: Berginia virgata, Mexico. [no image]. BM 000796114. Note: BM GH: Rhus ovata, California. [no image]. Annotation 1: Rhus ovata by database lists Mexico. Young in 1971. GH 00347143. BRU: Bergenia virgata Gr. & Harv. n. gen, California (PL). Annotation 1: Herbarium Olneyanum. Annotation 2: ISOTYPE Bergenia virgata Harv. Toxicodendron diversilobum (Torr. & A. Gray) Greene ex Benth. & Hook. f. by M.D. Cooper in October 2016. PBRU 00044618. Coulter 126. GH: Berginia virgata Harv., California. Script on label: coulteri n. sp. TCD: Rhus, California (PL). TCD! (PL). Annotation 1: ISOTYPE Berginia virgata Harv. ex Benth. & Hook. GH: R. diversiloba, California (PL). Annotation 1: Syn. Fl. N. Amer. and Holographis virgata (Harv. ex Benth. & Hook.) Daniel ssp. virgata Annotation 2: Toxicodendron lobatum (Hook.) Gillis by Gillis in 1968. by Daniel in 1981. GH 00093746. GH 00241341. K: Bergenia virgata Harv. nov. gen., California (PL). Annotation 1: Holo- Coulter 129. graphis virgata (Harv. ex Benth. & Hook.) Daniel ssp. virgata (ISO- TCD: Rhus lobata Hook., California (PL). TCD! TYPE of Berginia virgata Harv. ex Benth. & Hook.) by Daniel in 1981. GH: Rhus diversiloba, California [no image]. GH 00347304. K 000534561. Coulter s.n. K: Berginia virgata Benth. & Harv., California (PL). Annotation 1: NY: Rhus diversiloba Torr. & Gr., California (SL). Annotation 1: Torr. & HOLOTYPE Berginia virgata Harv. ex Benth. & Hooker by Daniel in Gray, Fl. N. Amer. NY 00050824. 1981. Annotation 2: Holographis virgata (Harv. ex Benth. & Hooker) Daniel ssp. virgata by Daniel in 1981. Written on sheet: Arizona or north- west Mexico in A. Gray Fl. Calif. Since rediscovered in Sonora W.B.H. apiaceae K 000534562 Apiastrum angustifolium Nutt. K: [no name], California. Note on label: nov. gen. Benth. (SL). Annota- Coulter 210. tion 1: Holographis virgata (Harv. ex Benth. & Hook.) Daniel ssp. vir- TCD: Apiastrum latifolia?, S. Francisco, California. Ex Herb. Hook. gata (ISOTYPE of Berginia virgata Harv. ex Benth. & Hook.) by Daniel (PL). TCD! in 1981. K 000534563. Coulter 213. TCD: Apiastrum angustifol[ium], California (PL). TCD! *Ruellia occidentalis (A. Gray) Tharp & F.A. Barkley Coulter 556. Bowlesia incana Ruiz & Pav. TCD: Ruellia? [another name illegible], California (PL). Annotation 1: Coulter 665. ISOSYNTYPE Ruellia tuberosa var. occidentalis A. Gray by G. Wallace TCD: Bowlesia lobata Dougl., California (PL). TCD! in 2016. TCD! BRU: Bowlesia lobata Dougl., California (PL). PBRU 00048134. GH: Ruellia tuberosa, California (PL). Annotation 1: as to foliage = var. occidentalis Syn. Fl. N. Amer. Annotation 2: R. nudiflora glabrata by Eryngium pendletonense K.L. Marsden & M.G. Simpson E.C. Leonard in 1928. Annotation 3: Ruellia tuberosa var. occidentalis Coulter 226. by Kittredge in 2008. GH 94183. Note: GH database lists as SYNTYPE. TCD: Eryngium, California (PL). TCD! VOLUME 37(1–2) Plantae Coulterianae 27

K: Eryngium cf. diffusum Torr., California (SL). Filed as E. vaseyi. K Sanicula laciniata Hook. & Arn. 000686497. Coulter 206. TCD: Sanicula laciniata, California (PL). TCD! Lomatium caruifolium (Hook. & Arn.) J.M. Coult. & Rose var. Coulter 218. Sanicula sp. TCD: Ferula caruifolium Hook. & Arn., S. Miguel, California (PL). Coulter 207. TCD! TCD: Sanicula, S. Francisco, California (PL). TCD! TCD: Sanicula, S. Miguel, California (PL). TCD! Lomatium parvifolium (Hook. & Arn.) Jeps. Coulter s.n. Coulter 222. TCD: Sanicula, Monterey to El Sur, California (PL). TCD! TCD: Ferula parvifolia Hook. & Arn., California (PL). TCD! GH: Lomatium parvifolium, California [no image]. GH 00424295. Yabea microcarpa (Hook. & Arn.) Koso-Pol. K: Euryptera lucida Nutt.?, Peucedanum parvifolium Torr. & Gray, Cal- Coulter 220. ifornia (SL). Written on sheet: Peucedanum parvifolium Torr. et Gray. K TCD: Caucalis microcarpa, S. Miguel to S. Barbara, California (PL). 000685914. Note: K database lists J.M. Coulter as collector in error. Annotation 1: Yabea microcarpa by S.S. Jwy. in 1983. TCD! Coulter 222x. Coulter 221. TCD: Ferula parvifolia, California (PL). TCD! TCD: Caucalis microcarpa, California (PL). Annotation 1: Yabea micro- carpa (Hook. & Arn.) Koso-Pol. by S.S. Jwy. in 1983. TCD! Lomatium sp. Coulter 223. Coulter 219. TCD: Caucalis microcarpa, California (PL). Annotation 1: Yabea micro- TCD: Ferula, California (PL). TCD! carpa (Hook. & Arn.) Koso-Pol. by S.S. Jwy. in 1983. TCD! Lomatium utriculatum (Torr. & A. Gray) J.M. Coult. & Rose Coulter 224. apocynaceae TCD: Ferula foeniculacea?, California (PL). TCD! BM: Ferula foeniculacea Nutt., California (PL). Annotation 1: possi- Asclepias californica Greene ble type Lomatium caruifolium by Mathias and Constance in 1962. BM Coulter s.n. 001042947. TCD: Acerates tomentosa, California (PL). TCD! K: Peucedanum caruifolium Torr. et Gray, California (SL). Annotation 1: Lomatium utriculatum (Nutt.) C. & R. by Constance in 1963. K araliaceae 000685888. Note: K database lists J.M. Coulter as collector in error. Hydrocotyle sp. Oenanthe sarmentosa DC. Coulter 215. Coulter 209. TCD: Hydrocotyle interrupta Ml. [Muhl.], California (PL). TCD! TCD: Helosciad[ium] californicum?, California (PL). TCD!

Osmorhiza sp. asteraceae Coulter 214. TCD: Glyc[osma], California (PL). TCD! Acamptopappus sphaerocephalus (Harv. & A. Gray ex A. Gray) A. Gray var. sphaerocephalus Perideridia oregana (S. Watson) Mathias Coulter 281. Coulter 212. TCD: [no name], California (PL). Annotation 1: Aplopappus (Acamp- TCD: Edosmia, California (PL). TCD! topappus) sphaerocephalus Harv. & Gr., Torr. & Gray, Fl. N. Amer., K: Edosmia gairdneri Nutt., Atenia, California (SL). K 000697456. Suppl. Annotation 2: ISOTYPE Haplopappus sphaerocephalus A. Gray by Lane in 1986. Annotation 3: ISOTYPE Haplopappus sphaerocephalus Sanicula arctopoides (Hook. & Arn.) Harv. & A. Gray ex A. Gray by G. Wallace in 2017. TCD! Coulter 204. GH: Aplopappus, Acamptopappus sphaerocephalus Ex no. 281 Calif. TCD: Sanicula arctopoides H. & A., California (PL). TCD! Coulter. (Herb. A. Gray label). Annotation 1: TYPE material (Coulter Coulter s.n. material) Aplopappus sphaerocephalus by Boufford in 1982. Annotation TCD: Sanicula arctopoides, California (SL). TCD! 2: Acamptopappus sphaerocephalus var. sphaerocephalus by M. Lane in BRU: Sanicula arctopoides, California (PL). PBRU 00048549. 1986. In GH database as Haplopappus sphaerocephalus TYPE MATE- NY: Sanicula arctopoides, California [no image]. Annotation 1: S. arc- RIAL. GH 00008699. topoides by Mathias in 1933. Annotation 2: Sanicula arctopoides by Shan K: Aplopappus (Acamptopappus) sphaerocephalus Harv. & Gray, Pl. & Constance in 1948–1949. NY 00214439. Fendl., California (SL). Annotation 1: ISOTYPE Haplopappus sect UC: Sanicula arctopoides, California [no image]. UC 973556. Acamptopappus sphaerocephalus Gray. Annotation 2: Acamptopappus Sanicula arguta J.M. Coult. & Rose sphaerocephalus (A. Gray) A. Gray var. sphaerocephalus by Lane in Coulter 216. 1986. K 000890922. Note: K database lists collection date as 1867 in TCD: [no name], Mont’y [Monterey] to El Sur, California (PL). TCD! error. K: Deweya arguta? Nutt., California (SL). K 000697371. Achillea millefolium L. Sanicula bipinnata Hook. & Arn. Coulter 348. Coulter 208. TCD: Achillea millefolium, Monterey to S. Gabriel, California (PL). TCD: Sanicula bipinnata, California (PL). TCD! TCD! K: S. bipinnata, California (SL). Annotation 1: S. bipinnata Hook. & Arn. by Lee in 1999. K 000697318. Ambrosia confertiflora DC. Coulter 259. Sanicula crassicaulis DC. TCD: Ambrosia, California (PL). Annotation 1: Franseria (Hanthiop- Coulter 205. sis?) tenuifolia Harv. & Gray n. sp., Torr. & Gray Fl. N. Amer. Suppl. TCD: Sanicula menziesii, California (PL). TCD! TCD! 28 Wallace ALISO

GH: Franseria tenuifolia Harv. & A. Gray, California [no image]. GH GH: [no name], California (PL). Written on fragment folder: B. paucira- 00283089. diata. Annotation 1: Baileya pauciradiata Harv. & Gray HOLOTYPE by K: Ambrosia artemisiaefolia [different handwriting on label, likely A. Brown in 1973. Annotation 2: HOLOTYPE Baileya pauciradiata Harv. Gray] = “F. tenuifolia Harv. & Gray”, California (SL). K 000502089. & Gray by Turner in 1991. GH 00002440. K: Baileya pauciradiata Harv. & Gray, California (PL). K 000943134. Ambrosia dumosa (A. Gray) W.W. Payne [Filed as Hemenoxys richardsonii]. Coulter 255. K: Bahia? Baileya pauciradiata Harv. & Gray. Pl. Fendl., California TCD: Franseria, California (PL). Annotation 1: Franseria coulteri n. sp. (SL). [Filed as Hemenoxys richardsonii]. K 000943135. [coulteri is crossed out] dumosa Gray in Frem. Rep., Torr. & Gray, Fl. N. Coulter s.n. Amer. TCD! NY: [no name] Annotation 1: New genus Baileya, California, Coulter., Torr. & Gray, Fl. N. Amer. Annotation 2: ISOTYPE Baileya pauciradiata Ambrosia sp. Harv. & Gray ex Gray by M. Turner in 1991. Written on sheet: Bailey’s Coulter 260. pauciradiata. Written on Torr. & Gray, Fl. N. Amer. label = “new genus”. TCD: Ambrosia longistylis Nutt.?, California (PL). Annotation 1: “I pos- NY 00162450. sess no specimen of A. longistylis Nutt. but this agrees pretty well with the descr. (except for the styles?) A. Gr.”, Torr. & Gray, Fl. N. Amer. TCD! Baileya pleniradiata Harv. & A. Gray ex A. Gray Coulter 312. Artemisia californica Less. GH: “Ex 312 Calif. Coulter (Hb. Trin. Col.) Baileya pleniradiata n. sp. Coulter 257. Torr. & Gray, Fl. N. Amer., Suppl.” (SL). Annotation 1: Baileya pleni- TCD: Artemisia californica Herb. Hook., California (PL). TCD! radiata Harv. & Gray LECTOTYPE by Brown in 1973. Annotation 2: Coulter 264. LECTOTYPE Baileya pleniradiata Harv. & A. Gray by Turner in 1991. TCD: Artemisia foliosa Dougl., Monterey to S. Gabriel, California (PL). GH 00002441. TCD! Bebbia juncea (Benth.) Greene var. aspera Greene Artemisia pycnocephala (Less.) DC. Coulter 297. Coulter 256. TCD: Carphephora chondrilloides H. & Gr. MS, near C. junceus, Cali- TCD: Artemisia pachystachya, Monterey to El Sur, California (PL). fornia (PL). Annotation 1: C. junceus β chondrilloides Harv. & Gray, Torr. TCD! & Gray, Fl. N. Amer., suppl. 2. TCD! K: Carphephorus junceus Benth., California (SL). K 000497511. Artemisia sp. Bidens laevis (L.) Britton, Sterns & Poggenb. Coulter 254. Coulter 324. TCD: Artemisia vulgaris, Monterey to S. Gabriel, California (PL). TCD! TCD: [no name], Monterey to S. Gabriel, California (PL). Written on Baccharis glutinosa Pers. sheet: B. laevis (L.) Britton et al. TCD! Coulter 265. Bidens sp. TCD: [no name], Monterey to San Gabriel, California (PL). TCD! Coulter 295. Baccharis sp. TCD : Bidens, California (PL). Annotation 1: B. leucantha Torr. & Gray, Coulter 282. Fl.N.Amer.TCD! TCD: Baccharis glomuliflora, Monterey to S. Gabriel, California (PL). Coulter 296. TCD! TCD: Bidens, California (PL). TCD!

Baileya multiradiata Harv. & A. Gray ex Torr. Blennosperma nanum (Hook.) S.F. Blake var. nanum Coulter 312. Coulter 346. TCD: Baileya multiradiata, California (PL). [Mixed sheet, right side TCD: Blennosperma californica, Monterey to El Sur, California (PL). specimens] Annotation 1: Baileya multiradiata H. & Gr. Torr. & Gray, TCD! Fl.N.Amer.TCD! Note: this may the sheet from which type material of BRU: Blennosperma californicum, California (PL). Annotation 1: B. pleniradiata was taken by A. Gray, ref. GH 00002441. Herbarium Olneyanum. PBRU 00049561. Coulter 326. Coulter s.n. TCD: Baileya nudicaulis, California (PL). Annotation 1: Baileya nudi- NY: Coniothele californica, California (PL). NY 00167627. caulis Harv. & Gr., Torr. & Gray, Fl. N. Amer. Annotation 1: ISOLEC- UC: Blennosperma nanum var. nanum, California [no image]. UC TOTYPE Baileya multiradiata Harv. & A. Gray ex Torr. by G. Wallace 973802. in 2016. TCD! GH: Baileya multiradiata Harv. & Gr., California (PL). Annotation 1: *Brickellia coulteri A. Gray LECTOTYPE Baileya multiradiata Harv. & A. Gray by Brown in 1973. Coulter 293. Annotation 2: HOLOTYPE Baileya multiradiata Harv. & A. Gray by TCD: Acourtia microcephala, California (SL). Annotation 1: “Though Turner in 1991. GH 00002436. the leaves are petioled I think this must be Acourtia microcephala DC. K: Bahia? Baileya nov. gen. Baileya multiradiata Harv. & Gray, (which I have not seen) and a congener of A. hebeclada DC. Deless. Ic. California (SL). [Filed as Baileya nervosa]. K 000943138. But in neither is the “pappus apicie panicellatus” I leave it untouched not having the means to study the Labiatiflorae. A. Gr., Torr. & Gray Fl. N. Baileya pauciradiata Harv. & A. Gray ex A. Gray Amer.” Annotation 2: ISOTYPE of Brickellia coulteri by G. Wallace in Coulter 312. 2017. TCD! TCD: Baileya multiradiata, California (PL). [Mixed sheet, left side spec- GH: Brickellia bulbostylis; Brickellia coulteri Gray MS, California (SL). imens] Annotation 1: Baileya pauciradiata H. & Gr. Torr. & Gray, Fl. N. Annotation 1: Fragment of type from Kew by S.F. Blake in 1915. Amer. TCD! Annotation 2: fragment of Holotype Brickellia coulteri A. Gray by BM: [no name], California (PL). Annotation 1: ISOTYPE Baileya pau- Boufford in 1982. Annotation 3: Brickellia coulteri var. coulteri by ciradiata Harv. & Gray by Brown in 1974. Annotation 2: ISOTYPE. BM Turner in 1990. Annotation 4: Brickellia coulteri by Scott in 2014. 001025628. GH 00004337. VOLUME 37(1–2) Plantae Coulterianae 29

K: Brickellia bulbostylis; Brickellia coulteri Gray MS, California (SL). GH: Eupatorium (Phyllaero.) sagittatum, California (PL). Annotation 1: K 001067146. Syn. Fl. N. Amer. Annotation 2: Eupatorium sagittatum Gray by Robin- son. Annotation 3: SYNTYPES Eupatorium sagittatum A. Gray Smith- Centromadia parryi (Greene) Greene subsp. australis (D.D. Keck) B.G. son. Contr. Knowl. 3: 88. 1850 (“California 294; Mexico, Sonora Alta, Baldwin Coulter 252; between Mexico and Mazatlan, Gregg”). GH 00007094. Coulter 290. GH: Hemizonia parryi subsp. australis, California [no image]. GH Cirsium occidentale (Nutt.) Jeps. var. coulteri (Harv. & A. Gray ex A. 414573. Gray) Jeps. Coulter 367. Centromadia pungens (Hook. & Arn.) Greene subsp. pungens TCD: Cirsium coulteri Harv. & Gray, California (PL). Annotation 1: Cir- Coulter 290. sium occidentale by Howell in 1958. Annotation 2: Type material Cirsium TCD: Hartmannia pungens, Monterey to S. Gabriel, California (PL). coulteri by G. Wallace in 2017. TCD! TCD! [two sheets] K: Cirsium? California (SL). Written across label and onto sheet: = 366 Coulter s.n. = Cirsium coulteri Gray = Carduus occidentalis Nutt. = Cnicus occi- US: Hartmannia pungens, California (SL). Written on sheet: Hemizonia dentalis Gray. K 000778242. Note: K database lists collection date of pungens (Hook. & Arn.) Torr. & Gray typica by D.D. Keck in 1935. US 30-07-1871 in error. 00223948. RSA: Cirsium coulteri, California [photo]. Annotation 1: TYPE. RSA 201381. Chaenactis artemisiifolia (Harv. & A. Gray ex A. Gray) A. Gray Coulter 313. Cirsium undulatum (Nutt.) Spreng. TCD: Acarphaea artemisiaefolia Harv. & Gr., California (PL). Annota- Coulter 289. tion 1: Acarphaea artemisiaefolia [sic] nov. gen. Harv. & Gr. in Torr. & TCD: Cirsium undulata [sic], S. Miguel to S. Barbara, California (PL). Gray Fl. N. Amer. Suppl. Annotation 2: ISOTYPE Acarphaea artemisi- Written on sheet: not discolor but will do for undulatum AG [ = A. Gray]. aefolia Harv. & A. Gray ex A. Gray by G. Wallace in 2016. TCD! TCD! GH: Herb. A. Gray label: “Ex no. 313 California Coulter Chaenactis Coulter s.n. artemisiaefolia, Acarphaea artemisiaefolia Harv. & Gray in Torr. & Gray TCD: Cirsium undulatum, California (PL). Written on sheet: [illegible] Fl. N. Amer. Suppl.” Annotation 1: Syn. Fl. N. Amer. Written on sheet: undulatum A. Gr. TCD! TYPE. GH 00004662. K: Chaenactis, Acarphaea artemisiaefolia Harv. & Gray, P. Fendl., Cal- Corethrogyne filaginifolia (Hook. & Arn.) Nutt. ifornia (SL). Written on sheet: “Written Acicarphaea in Walpers by mis- Coulter 267. take”. K 001065668. TCD: Corethrogyne tomentella, California (PL). TCD! GH: Corethrogyne filaginifolia, California (SL). Annotation 1: Chaenactis glabriuscula DC. Corethrogyne filaginifolia by Saroyan in 1976. GH 00387137. Coulter 364. K: Corethrogyne tomentella, California (SL). Annotation: Corethrogyne TCD: Chaenactis filifolia H. & G., California (PL). Annotation 1: Chae- filaginifolia Hook. & Arn. subsp. filaginifolia by Saroyan in 1976. K nactis filifolia Harv. & Gr., Torr. & Gray, Fl. N. Amer. Annotation 2: Type 000884801. material Chaenactis filifolia Harv. & A. Gray ex A. Gray by G. Wallace Coulter 268. in 2016. TCD! TCD: Corethrogyne leucophylla Lindl., Monterey, S. Gabriel, El Sur, K: Chaenactis, C. filifolia, California (SL). K 001065656. California (PL). TCD! Coulter 465. GH: Corethrogyne filaginifolia, California [no image]. Annotation 1: TCD: Chaenactis glabriuscula, California (PL). Annotation 1: C. Corethrogyne filaginifolia by Saroyan in 1976. GH 387138. glabriuscula DC. ex char. Torr. & Gray, Fl. N. Amer. Annotation 2: ISO- K: Corethrogyne filaginifolia¸ California (SL). K 000884804. TYPE Chaenactis filifolia Harv. & A. Gray ex A. Gray by G. Wallace in Coulter 270. 2017. TCD! TCD: Corethrogyne californica, Monterey to El Sur, California (PL). GH: Ex no. 465, Calif. Coulter C. glabriuscula DC. ex char. Torr. & Gray, TCD! Fl. N. Amer. Written on Herb. A. Gray label. Annotation 1: Type. Anno- Coulter 271. GH tation 2: Syn. Fl. N. Amer. Annotation 3: [in database but not on TCD: Corethrogyne, California (PL). Annotation 1: C. filaginifolia, foliis Chaenactis filifolia GH specimen] TYPE by Kettridge in 2017. 961327. gracilioribus,Torr.&Gray,Fl.N.Amer.TCD! K Chaenactis K : ? “by mistaken omitted” (SL). 001065665. Note: on Coulter 273. sheet with Coulter 365 below. TCD: [no name], California (PL). Annotation 1: Corethrogyne virgata Chaenactis glabriuscula DC. var. lanosa (DC.) H.M. Hall Benth. Voy. Sulph., Torr. & Gray, Fl. N. Amer. TCD! Coulter 365. GH: Corethrogyne filaginifolia, California [no image]. Annotation 1: TCD: Chaenactis lanosa, Chaenactis glabriuscula, California (PL). An- Corethrogyne filaginifolia by Saroyan in 1976. GH 387063. notation 1: C. lanosa, more glabrous than Dougl. spec., Torr. & Gray, Fl. Deinandra corymbosa (DC.) B.G. Baldwin N. Amer. TCD! Coulter 300. K: Chaenactis glabriuscula? (SL). Written on sheet: Ch. filifolia Gr. K TCD: Hemizonia angustifolia, Monterey to S. Gabriel, California (PL). 001065664. Annotation 1: Hemizonia angustifolia Torr. & Gray, Fl. N. Amer. TCD! Chaenactis stevioides Hook. & Arn. Coulter 337. Deinandra fasciculata (DC.) Greene TCD: Chaenactis stevioides Hook. & Arn., California (PL). TCD! Coulter 361. TCD: Hemizonia ramosissima?, California (PL). Annotation 1: An Hem- *Chromolaena sagittata (A. Gray) R.M. King & H. Robinson izonia ramosissima Benth. Sulph.? [illegible] vix glandulosa seu viscosa Coulter 294. Torr. & Gray, Fl. N. Amer. TCD! TCD: Eupatorium, California (PL). Annotation 1: Eupatorium (Phyl- laeron) sagittatum Harv. & Gr., Torr. & Gray, Fl. N. Amer. Suppl. Anno- Eclipta prostrata (L.) L. tation 2: ISOSYNTYPE Eupatorium sagittatum A. Gray by G. Wallace Coulter 341. in 2016. TCD! TCD: Eclipta erecta, California (PL). TCD! 30 Wallace ALISO

Encelia californica Nutt. aea canescens Torr. & A. Gray ex A. Gray by Blake in 1913. Annota- Coulter 308. tion 4: Geraea canescens Torr. & A. Gray ex Gray by Nishida in 1989. TCD: Encelia?, California (PL). Written on sheet: cf. E. parvifolia GH 00306798. Note: listed as PARATYPE in CCH database. Not listed Kunth, E. conspersa Benth. TCD! Note: looks like A. Gray’s handwrit- as TYPE in GH database. ing. Coulter 309. Gnaphalium sp. TCD: Encelia californica, California (PL). TCD! Coulter 354. K: Encelia californica, California (SL). Annotation 1: E. californica by TCD: Gnaphalium, Monterey to El Sur, California (PL). TCD! S.F.B. [ = S.F. Blake] in 1914. K 001065979. *Helenium laciniatum A. Gray Encelia farinosa Torr. Coulter 356. Coulter 327. TCD: Helenium laciniatum H. & G., California (PL). Annotation 1: H. TCD: Encelia nivea, California (PL). TCD! laciniatum Harv. & Gr., Torr. & Gray, Fl. N. Amer. Annotation 2: Type GH: Encelia farinosa, California [no image]. Annotation 1: Encelia fari- material Helenium laciniatum A. Gray by G. Wallace in 2017. TCD! nosa by Blake in 1913. GH 414449. GH: Helenium laciniatum, California (PL). Annotation 1: H. laciniatum Gray. Torr. & Gray, Fl. N. Amer. Suppl. Annotation 2: Syn. Fl. N. Amer. Ericameria ericoides (Less.) Jeps. Annotation 3: SYNTYPE Helenium laciniatum A. Gray by H.F.L. Rock Coulter 269. in 1956. Annotation 4: ISOSYNTYPE Helenium laciniatum A. Gray by TCD: Ericameria microphylla, Mont’y. [ = Monterey] to S. Gabriel, Cal- Gandhi and Ferreras in 2007. GH 00008769. Note: listed in GH database ifornia (PL). TCD! as ISOSYNTYPE, SYNTYPE. K: Helenium, California (SL). K 000974774. Erigeron canadensis L. MO: Helenium, California (PL). Annotation 1: Type Specimen Herb. Coulter 284. M.B.G. Annotation 2: Helenium laciniatum A. Gray Syntype by H.F.L. TCD: Erigeron canadense, Monterey to S. Gabriel, California (PL). Rock. Annotation 3: “Helenium laciniatum by Bierner in 1971. One of TCD! the original syntypes. Lectotype Coulter 358”. MO 1987329. Coulter 358. Erigeron glaucus Ker Gawl. TCD: Helenium laciniatum H. & G., California (PL). Annotation 1: H. Coulter 325. laciniatum Harv. & Gr., Torr. & Gray, Fl. N. Amer. Annotation 1: Type TCD: Erigeron glaucum [sic], California (PL). TCD! material Helenium laciniatum A. Gray by G. Wallace in 2017. TCD! K: Erigeron glaucum, Aster californicum, [California] (SL). Annotation GH: Helenium laciniatum Gray, California (PL). Annotation 1: H. 1: Erigeron glaucus videtur ex adm. Hook. & Arn. Bot. Beechey Suppl., laciniatum Torr. & Gray, Fl. N. Amer. Suppl. Annotation 2: LECTOTYPE Torr. & Gray, Flora N. Amer. K 000885072. Helenium laciniatum A. Gray by H.F.L. Rock in 1956. Annotation 3: Helenium laciniatum Erigeron philadelphicus L. var. philadelphicus ISOLECTOTYPE A. Gray Proc. Amer. Acad. Arts 9: 203. 1874, Lectotypified by Bierner in Brittonia 24(4): 346. 1972 by Coulter 287. Gandhi & Ferreras in 2007. Written on sheet: Coulter’s 358 in Ball Herb. TCD: Erigeron purpureum [sic], California (PL). TCD! from Monterey! GH 00008768. Note: listed in GH database as ISOLEC- Eriophyllum confertiflorum (DC.) A. Gray var. TOTYPE, LECTOTYPE. Coulter 362. US: Helenium. Monterey, California (SL). Annotation 1: Helenium TCD: Bahia confertiflora, California (PL). TCD! laciniatum A. Gray (2nd number cited by Gray) by S.F.B. [ = S.F. Blake] K: Bahia confertiflora DC., California (SL). K 001092076. in 1934. Annotation 2: Helenium laciniatum Gray Isotype by H.F.L. Rock in 1956. Annotation 3: Helenium laciniatum Gray Lectotype by M.W. Eriophyllum multicaule (DC.) A. Gray Bierner in 1971. US 00124879. Coulter 343. TCD: Actinolepis multicaulis, Monterey to S. Miguel, California (PL). *Helenium puberulum DC. TCD! Coulter 357. TCD: H[elenium] puberulum, California (PL). Annotation 1: H. mexi- Eriophyllum staechadifolium Lag. canum ex chars.Torr.&Gray,Fl.N.Amer.TCD! Coulter 340. K: Helenium puberulum DC., California (SL). Written on sheet: A spec- TCD: Eriophyllum artemisiaefolia, Monterey to S. Gabriel, California imen sub. nom. H. brachyglossum (Herb. Kew 8/70) comes very near this (PL). TCD. and is probably identical, J.B. K 000974776. Coulter s.n. TCD: [no name], Monterey to El Sur, California (PL). TCD. Helenium sp. Coulter 355. Euthamia occidentalis Nutt. TCD: Helenium, California (PL). Written on sheet: “Rays none, pappus Coulter 266. very short. New to me: but I dare not take it up. Is it Cephalophora aro- TCD: [no name], Monterey to S. Gabriel, California (PL). Annotation 1: matica?A.Gr.”[= A. Gray]. TCD! S. occidentalis Nutt. Torr. & Gray, Fl. N. Amer. Written on sheet: occi- Coulter s.n. dentalis. TCD! TCD: Helenium, Helenium mexicanum, California (PL). Annotation 1: K: Solidago occidentalis Nutt., California (SL). K 000890796. H. mexicanum DC. ex char. Torr. & Gray, Fl. N. Amer. TCD!

Geraea canescens Torr. & A. Gray *Helenium thurberi A. Gray Coulter 304. Coulter 359. TCD: [no name], California (PL). Annotation 1: Geraea canescens Torr. TCD: Helenium puberulum, California (PL). Annotation 1: Type material & Gray, Fl. N. Amer. Annotation 2: ISOSYNTYPE Geraea canescens Helenium thurberi A. Gray by G. Wallace in 2017. Written on sheet: H. Torr. & A. Gray by G. Wallace in 2016. TCD! puberulum DC. but pappus longer [ = A. Gray]. TCD! GH: Geraea canescens Torr. & A. Gray, California [no image]. An- GH: Helenium thurberi, puberulum but pappus long, California (PL). notation 1: Encelia eriocephala Gray. Annotation 2: Simsia canescens Annotation 1: Helenium thurberi Gray. Syn. Fl. N. Amer. Annotation 2: (Torr. & A. Gray ex A. Gray) A. Gray. Annotation 3: COTYPE Ger- SYNTYPE Helenium thurberi A. Gray by Rock in 1956. GH 00022610. VOLUME 37(1–2) Plantae Coulterianae 31

K: Helenium puberulum Benth., Desc. vin DC., California (SL). Written GH: Lasthenia var. coulteri [sic], California (PL). Annotation 1: var. on sheet: Helenium puberulum DC. Prod. 5, p. 667. K 000974775. coulteri. Syn. Fl. N. Amer. Annotation 2: LECTOTYPE Lasthenia glabrata Lindl. subsp. coulteri (A. Gray) Ornduff by Ornduff in 1960. Helianthella sp. Annotation 3: LECTOTYPE Lasthenia glabrata var. coulteri A. Gray. Coulter 303. Syn. Fl. N. Amer. 1(2): 324. 1884. Lectotypified by R. Ornduff, Univ. TCD: Helianthella, California (PL). Annotation 1: Helianthus lenticu- Calif. Pub. Bot. 40: 74. 1966 by W. Kittredge in 2009. GH 00009588. laris Torr. & Gray, Fl. N. Amer. TCD!

Hemizonia congesta DC. subsp. luzulifolia (DC.) Babc. & H.M. Hall Lasthenia gracilis (DC.) Greene Coulter 298. Coulter 345. TCD: Hemizonia rudis Benth.?, California (PL). TCD! TCD: Burriela [sic] gracilis, Monterey to El Sur, California (PL). TCD! Coulter s.n. Hemizonia sp. BM: Burrielia gracilis, California (PL). Annotation 1: Lasthenia Coulter 448. chrysostoma by Ornduff 1964. Annotation 2: ISOTYPE Lasthenia gra- TCD: Hemizonia, California (PL). TCD! cilis by Chan in 2001. BM 690270.

Heterotheca grandiflora Nutt. Lasthenia microglossa (DC.) Greene Coulter 275. Coulter 328. TCD: Heterotheca floribunda Benth., California (PL). TCD! TCD: Burrielia microglossa, S. Miguel, California (PL). TCD! BRU: [no name] California (PL). Annotation 1: Herbarium Olneyanum. K: Burrielia microglossa, California (SL). Annotation 1: Lasthenia mi- PBRU 00051153. croglossa (DC.) Greene by Ornduff in 1964. K 001092121. Heterotheca sessiliflora (Nutt.) Shinners subsp. echioides (Benth.) Sem- Coulter s.n. ple NY: Burrielia microglossa, California [no image]. NY 00162829. Coulter 349. TCD: Chrysopsis near villosa, Monterey to S. Gabriel, California (PL). Layia glandulosa (Hook.) Hook. & Arn. Annotation 1: Chrysopsis echioides Benth., Torr. & Gray, Fl. N. Amer. Coulter 314. Suppl. TCD! TCD: Layia glandulosa, California (PL). [Filed in Madroglossa folder]. TCD! Iva axillaris Pursh Coulter 258. TCD: Iva axillaris? S. Gabriel, California (PL). Annotation 1: Certainly Layia heterotricha (DC.) Hook. & Arn. I. axillaris Torr. & Gray, Fl. N. Amer. TCD! Coulter 320. K: Iva axillaris? California (SL). K 001065924. Note: K database lists TCD: Layia heterotricha, California (PL). TCD! as a type in error. GH: Layia heterotricha, California [no image]. Annotation 1: Layia het- erotricha by Keck in 1935. GH 274836. Laënnecia coulteri (A. Gray) G.L. Nesom Coulter 285. Layia platyglossa (Fisch. & C.A. Mey.) A. Gray TCD: [no name], California (PL). Annotation 1: Type material Conyza Coulter 315. coulteri A. Gray by G. Wallace in 2017. Written on sheet: Conyza diver- TCD: Callichroa platyglossa, California (PL). Annotation 1: C. platy- sifolia. TCD! glossa Torr. & Gray, Fl. N. Amer. TCD! Coulter 286. K: Layia galardioides [sic] Hook. & Arn., California (SL). K 000251305. TCD: Conyza diversifolia, California (PL). Annotation 1: Type material Coulter 318. Conyza coulteri TCD A. Gray by G. Wallace in 2017. ! TCD: Layia elegans, California (PL). Note on label: near glandulosa but Coulter 285, 286. pappus diff[eren]t. Annotation 1: Layia elegans Nutt. (wool of the pappus GH : [no name], California. Script on fragment packet: Coulter 285, 286 finely crisped inside) Torr. & Gray, Fl. N. Amer. Annotation 2: “2.3. like GH (Hb. Harvey). [Just small pieces in a fragment folder]. 00006095. no. 1 with all the inner crisped wool wanting. As it is, it has less hairs on GH Note: in database as SYNTYPE but no annotations on sheet. the pappus than any species I know. I have not L. glandulosa (send me L. Lasthenia californica Lindl. subsp. some achenes of that). If not that, it might be safer to call it a state of elegans. A. Gr. Torr. & Gray, Fl. N. Amer.” TCD! Coulter 344. Coulter 319. TCD: Burriela [sic] chrysostoma, Monterey to S. Francisco, California TCD: Callichroa platyglossa, California (PL). TCD! (PL). Annotation 1: Burrielia (Baeria) chrysostoma Torr.&Gray,Fl.N. Coulter 322. Amer. TCD! TCD: Callichroa platyglossa, S. Miguel, California (PL). Annotation 1: Coulter 566. Callichroa platyglossa Torr. & Gray, Fl. N. Amer. TCD! K: Burrielia¸ California. [Note on label]: omitted compositae (SL). K 001065993. Leptosyne douglasii DC. Lasthenia coronaria (Nutt.) Ornduff Coulter 339. Coulter 350. TCD: Leptosyne douglasii, Monterey to S. Francisco, California (PL). TCD: Dichaeta tenella Tor. & Gr. vol. 2, p. 379, California (PL). TCD! TCD! K: Dichaeta Nutt., California (SL). Annotation 1: Lasthenia coronaria GH: Leptosyne, California (PL). Annotation 1: Coreopsis douglasii (Nutt.) Ornduff by Ornduff in 1964. K 001092102. (DC.) H.M. Hall by Smith in 1984. GH 00009637. Note: listed as possible SYNTYPE in GH database. Lasthenia glabrata Lindl. subsp. coulteri (A. Gray) Ornduff Coulter s.n. Coulter 338. NY: Written on annotation label: Leptosyne douglasii, California. Torr. TCD: Lasthenia glabrata, San Miguel to S. Barbara, California (PL). & Gray, Fl. N. Amer. (SL). Annotation 1: All C. californica (Nutt.) Annotation 1: ISOLECTOTYPE Lasthenia glabrata var. coulteri by G. H.K. Sharsm. by D.D. Keck in 1956. Annotation 2: Coreopsis californica Wallace in 2016. TCD! (Nutt.) H.K. Sharsmith by S. Welsh in 1991. NY 02055261. 32 Wallace ALISO

Leptosyne gigantea Kellogg Osmadenia tenella Nutt. Coulter 316. Coulter 360. GH: Leptosyne gigantea, San Miguel to San Barbara, California [no im- TCD: Calycadenia near C. tenella, California (PL). Written on sheet: age]. Annotation 1: Coreopsis gigantea by Smith in 1982. GH 00415552. Calycadenia tenella. TCD! K: L. gigantea fid AG [ = A. Gray] The Bot. Calif (SL). Annotation 1: Coulter s.n. Coreopsis gigantea (Kellogg) H.M. Hall by Smith in 1983. K 000943080. NY: Osmadenia tenella, California [no image]. NY 230331. Note: filed as Coreopsis maritima because of a Nuttall collection on the same sheet. Palafoxia arida B.L. Turner & M.I. Morris var. arida Coulter 347. Lessingia pectinata Greene var. TCD: Palafoxia linearis Lag., California (PL). Annotated: P. linearis Coulter 291. Torr. & Gray, Fl. N. Amer. Suppl. TCD! TCD: Lessingia germanorum, Monterey, California (PL). TCD! GH: Lessingia germanorum Cham., California [no image]. Annotation 1: Palafoxia arida B.L. Turner & M.I. Morris var. gigantea (M.E. Jones) Lessingia glandulifera A. Gray by Gray. Syn. Fl. N. Amer. Annotation 2: B.L. Turner & M.I. Morris Lessingia glandulifera var. pectinata by Spence in 1963. GH 00283125. Coulter 347. TCD: Palafoxia macrocephala, California (PL). Annotation 1: P. m a c ro - Logfia filaginoides (Hook. & Arn.) Morefield cephala n. sp., Torr. & Gray, Fl. N. Amer. Suppl. TCD! Coulter 352. GH: Palafoxia arida var. gigantea, California [no image]. GH 00427278. TCD: Filago californica, California (PL). TCD! Note: in GH database as 367.

Madia elegans D. Don *Pectis coulteri Harv. & A. Gray ex A. Gray Coulter 299. Coulter 330. TCD: Madaria elegans, California (PL). TCD! TCD: Pectis (Heteropectis) coulteri n. sp., California (PL). Annotation 1: Pectis coulteri Harv. & Gr. in Torr. & Gray, Fl. N. Amer. Suppl. An- notation 2: HOLOTYPE Pectis coulteri Harv. & A. Gray by D.J. Keil in Malacothrix californica DC. 1984. TCD! Coulter 342. GH: Pectis (Heteropectis) coulteri (SL). Annotation 1: 330. Calif. Coul- TCD: Malacothrix californica, Monterey to S. Miguel, California (PL). ter. Annotation 2: Syn. Fl. N. Amer. Annotation 3: revision by M.L. F. [ Annotation 1: M. californica Torr. & Gray, Fl. N. Amer. TCD! = Fernald]. GH 00010916. Note: just pieces in fragment folder. K: Malacothrix californica, California (SL). Annotated: Malacothrix K: Pectis, Tagetes? California (SL). Annotation 1: ISOTYPE Pectis coul- californica DC. by Davis in 1987. K 000796917. teri by D.J. Keil in 1984. Written on sheet: Pectis coulteri Gray. K 001065561. Malacothrix coulteri Harv. & A. Gray ex A. Gray Coulter 252. *Pectis filipes Harv. & A. Gray ex A. Gray TCD: Malacothrix coulteri H. & G., California (PL). Annotation 1: Coulter 329. Malacothrix?(Malacolepis) coulteri Harv. & Gr., Torr. & Gray, Fl. N. TCD: Pectis (Heteropectis) filipes Harv. & A. Gray, California (PL). An- Amer. Annotation 2: Type material Malacothrix coulteri Harv. & A. Gray notation 1: HOLOTYPE Pectis filipes Harv. & Gray in Gray by D.J. Keil ex A. Gray by G. Wallace in 2016. TCD! in 1972. TCD! GH: Malacothrix (Malacolepis) coulteri Harv. & Gray, California Herb. GH: Pectis (Heteropectis) coulteri Harv. & Gray in Torr. & Gray, Fl. N. A. Gray label (SL). Annotation 1: Syn. Fl. N. Amer. GH 00010011. Note: Amer. Suppl. (SL) Annotation 1: Syn. Fl. N. Amer. Annotation 2: Frag- in GH database as POSSIBLE TYPE. ment of HOLOTYPE Pectis filipes Harv. & Gray var. filipes by D.J. Keil K: Malacothrix (Malacolepis) coulteri Gray, California (SL). Annotation in 1973. Annotation 3: Holotype fragment Pectis filipes Harvey & A. Gray 1: Malacothrix coulteri ISOTYPE by Davis in 1987. K 000796912. Mem. Amer. Acad. Arts n.s. 4: 62. 1849. T. Coulter 329 Holotype TCD by I. Ferreras in 2008. GH 00010952. Note: just pieces in fragment folder. Malacothrix saxatilis (Nutt.) Torr. & A. Gray var. saxatilis K: Pectis caliscana, Pectis filipes Harv. & Gray, this is the one at the upper Coulter 247. right hand corner, California (SL). Annotation 1: ISOTYPE Pectis filipes TCD: Malacothrix saxatilis, California (PL). Annotation 1: M. saxatilis Harv. & Gray var. filipes by D.J. Keil in 1973. K 001065565 Torr. & Gray Fl. N. Amer. TCD! Pectis papposa Malacothrix saxatilis (Nutt.) Torr. & A. Gray var. tenuifolia (Nutt.) A. Harv. & A. Gray ex A. Gray Gray Coulter 331. TCD: Pectis (Pectothrix) papposa, California (PL). Annotation 1: Pectis Coulter 246. (Pectothrix) papposa Harv. & Gray [in] Torr. & Gray, Fl. N. Amer. Suppl. TCD: Malacothrix tenuifolia, Monterey to S. Gabriel, California (PL). Annotation 1: HOLOTYPE Pectis pappposa Harv. & A. Gray by D.J. Keil TCD! in 1972. TCD! K: Hieraceum californicum DC., California (SL). Annotation 1: Mala- GH: Pectis papposa, Lorentea n. sp. Coulter, California (PL). Annotation cothrix saxatilis var. tenuifolia (Nutt.) A. Gray by Ferris in 1954. Anno- 1: 331 Calif. Coulter, Pectis (Pectothrix) papposa Harv. & Gray. Torr. & tation 2: Malacothrix saxatilis var. tenuifolia (Nutt.) A. Gray by Davis in Gray, Fl. N. Amer. Annotation 2: Syn. Fl. N. Amer. Annotation 3: Re- 1988. K 000796897. vision by M.L. F. [ = Fernald]. Annotation 4: ISOTYPE Pectis papposa Malacothrix sp. Harv. & A. Gray by D.J. Keil in 1973. GH 00010965. Coulter 251. K: Pectis coulteri A. Gray, California (SL). Annotation 1: ISOTYPE Pec- TCD: Malacothrix sonchoides, California (PL). TCD! tis papposa Harv. & A. Gray by D.J. Keil in 1984. K 001965562. Coulter s.n. K: Pectis papposa Harv. & Gray in Pl. Fendl., California (PL). Annota- GH: Malacothrix saxatilis, California [no image]. GH 00263184. tion 1: ISOTYPE Pectis papposa Harv. & A. Gray by D.J. Keil in 1973. K 001065570. Monolopia major DC. K: Pectis (Pectothrix) papposa Harv. & Gray, Pl. Fendl., California (SL). Coulter 323. Annotation 1: ISOTYPE Pectis papposa Harv. & Gray by D.J. Keil in TCD: Monolopia major, S. Miguel, California (PL). TCD! 1973. K 001065571. VOLUME 37(1–2) Plantae Coulterianae 33

*Perityle californica Benth. GH: Senecio flaccidus var. douglasii, Monterey to S. Gabriel, California Coulter 279. [no image]. GH 427771. TCD: [no name], California (PL). Annotation 1: Perityle plumiger [sic] Coulter 334. Harv. & Gray, Torr. & Gray Fl. N. Amer. Suppl. Annotation 2: ISOTYPE TCD: Senecio douglasii var.?, California (PL). Annotation 1: S. douglasii Perityle plumigera Harv. & A. Gray ex A. Gray by G. Wallace in 2016. var., Torr. & Gray, Fl. N. Amer. TCD! TCD! US: Perityle plumigera type, California (SL). Annotation 1: Perityle cal- Senecio sp. ifornica by A.M. Powell in 1970. US 00047189. Note: filed as Perityle Coulter 332. plumigera. TCD: Senecio coulteri Harv. & Gray, California (PL). Annotation 1: S. coulteri Harv. & Gr. Torr. & Gray, Fl. N. Amer. TCD! *Perityle leptoglossa Harv. & A. Gray ex A. Gray Coulter 336. Coulter 277. TCD: Senecio multilobatus, Monterey to S. Gabriel, California (PL). An- TCD: Perityle californica Benth., California (PL). Annotation 1: ISO- notation 1: S. multilobatus β Torr. & Gray, Fl. N. Amer. TCD! TYPE Perityle leptoglossa Harv. & A. Gray ex A. Gray by G. Wallace in 2016. TCD! Solidago spathulata DC. GH: Perityle leptoglossa, California (PL). Annotation 1: P. leptoglossa Coulter 288. Torr. & Gray, Fl. N. Amer. Annotation 2: Syn. Fl. N. Amer. Annotation TCD: [no name], California (PL). Annotation 1: S. spiciformis Torr. & 3: HOLOTYPE Perityle leptoglossa Harv. & Gray in Gray by Powell in Gray, Fl. N. Amer. Written on sheet: spiciformis. TCD! 1970. GH 00011080. K: Perityle leptoglossa Harv. & Gray, California (PL). Annotation 1: Solidago velutina DC. subsp. californica (Nutt.) Semple TYPE of Perityle leptoglossa Harv. & Gray. Annotation 2: Herbarium Coulter 263. Careyanum. Annotation 3: Perityle leptoglossa Harv. & Gray by Ryd- TCD: Solidago, California (PL). Annotation 1: S. californica near β,cf. berg. K 001079928. in Herb. Hook., Torr. & Gray, Fl. N. Amer. Written on sheet: californica. K: [no name], MEXICO Coulter (SL). Annotation 1: TYPE Perityle lep- TCD! toglossa Harv. & Gray. Written on sheet: Perityle leptoglossa Harv. & Gray, coll. type descr. Hab.: Sonora, fide Rydberg. K 000582187. Stephanomeria virgata Benth. subsp. US: Perityle leptoglossa Harv. & Gray, ex herb. Gray, California (SL). Coulter 250. Initials on label: J.N.R. [ = J.N. Rose]. Annotation 1: U.S. Nat. Herbarium TCD: Stephanomeria virgata, California (PL). TCD! TYPE No. 931. Type of Perityle leptoglossa Gray. Annotation 2: Type. US 00131458. Symphyotrichum chilense (Nees) G.L. Nesom Coulter 274. *Perityle microglossa Benth. var. microglossa TCD: Aster chilensis?, Monterey to S. Gabriel, California (PL). Written Coulter 278. on sheet: in Torr. & Gr. fl. A. Gr. TCD! TCD: Spilanthes, California (PL). Annotation 1: Perityle acmella Harv. & Gray, Torr. & Gray, Fl. N. Amer., suppl. Annotation 2: ISOTYPE Per- Uropappus lindleyi (DC.) Nutt. ityle acmella Harv. & A. Gray ex A. Gray by G. Wallace in 2016. TCD! Coulter 253. GH: Perityle acmella Gray, California (PL). Annotation 1: Perityle TCD: Calais linearifolia, S. Miguel to S. Barbara, California (PL). TCD! acmella Harv. & Gray. Annotation 2: P. microglossa Benth. Syn. Fl. N. K: Calais linearifolia, California (SL). Annotation 1: Microseris linear- Amer. Annotation 3: Perityle microglossa Benth. var. microglossa by ifolia (DC.) Schultz Bip. by Chambers in 1953. K 000796775. Powell in 1970. GH 00014292. Note: in GH database as POSSIBLE TYPE. Venegasia carpesioides DC. K: Perityle acmella Harv. & A. Gr., California (PL). Annotation 1: Coulter 317. TYPE of Perityle acmella Harv. & Gray = P. microglossa Benth. var. TCD: Venegasia carpesioides, S. Miguel to S. Barbara, California (PL). microglossa. K 001079924. TCD! K: [no name] California (PL). Annotation 1: TYPE of Perityle acmella Wyethia angustifolia (DC.) Nutt. Harv. & Gray = Perityle microglossa Benth. var. microglossa. K Coulter 310. 001079925. TCD: Wyethia angustifolia, California (PL). TCD! Pluchea sericea (Nutt.) Coville K: Wyethia angustifolia Nutt., California (SL). Written on sheet: Wieythia Coulter 280. [sic] angustifolia Nutt. Torr. et Gray Fl. N. Am. 2. p. 300. K 001065772. TCD: Baccharis?, California (PL). Annotation 1: Tessaria borealis. Coulter 311. TCD! TCD: Helianthella, California (PL). Annotation 1: Wyethia robusta Nutt. GH: Tessaria borealis Torr. & A. Gray, California [no image]. Annota- Torr. & Gray, Fl. N. Amer. TCD! tion 1: Pluchea borealis (Torr. & A. Gray) Gray. Annotation 2: Pluchea Xanthisma spinulosum (Pursh) D.I. Morgan & R.L. Hartm. var. good- sericea (Nutt.) Coville. GH 00415662. dingii (A. Nelson) D.I. Morgan & R.L. Hartm. K: [no name], California (SL). Written on sheet: cf. Tessaria borealis Coulter 276. Gray, Pl. Fendl. K 00974654. TCD: Aplopappus coulteri H. & G., California (PL). Annotation 1: Aplopappus (Blepharodon) coulteri Harv. & Gray, Torr. & Gray, Fl. N. Senecio californicus DC. Amer. TCD! Note: This species ined. Coulter 335. TCD: Senecio californicus, Monterey to S. Miguel, California (PL). TCD! K: Sen. californicus β, California (SL). K 000843675. betulaceae Senecio flaccidus Less. var. douglasii (DC.) B.L. Turner & T.M. Barkley Coulter 333. Alnus rhombifolia Nutt. TCD: Senecio douglasii, Monterey to S. Gabriel, California (PL). Anno- Coulter 649. tation 1: S. douglasii var? Torr. & Gray, Fl. N. Amer. TCD! TCD: [no name], California (PL). TCD! 34 Wallace ALISO

GH: Alnus rhombifolia, California [no image]. Annotation 1: Alnus ser- same label: I doubt now that these Coulter things are Greene’s plant – rulata var. rugosa (Du Roi) Regel. GH 401364. these seem to be A. eastwoodiae 1918. On same sheet a fragment packet labeled: “ex Herb. Hook. – Kew Amsinckia microcarpa Greene deter- Alnus rubra Bong. mined by Greene and probably type. California, Coulter 497.” Written on Coulter 649. sheet: “ = spectabilis”. GH 00094013. Note: in GH database as TYPE. K: Alnus rubra Bong., California (SL). K 000859952. Note: lacks leaves K: Amsinckia microcarpa [E.L.G.], California (SL). Annotation 1: Am- for identification but is on sheet with A. rubra from Alaska. sinckia spectabilis Fisch. & Mey. (var. microcarpa (Greene) Jeps.). Anno- tation 2: evidently the type (LECTOTYPE) specimen of Amsinckia micro- Corylus cornuta Marshall subsp. californica (A. DC.) E. Murray carpa although he cited no number. This is the only specimen of Coulter’s Coulter 649. that matched his description (very short calyx, etc.), by Lassen in 1986. GH Corylus cornuta californica : subsp. , California [no image]. Anno- K 001067016. tation 1: Corylus cornuta subsp. californica by Furlow in 2003. GH Coulter s.n. 401475. NY: Myosotis fulva Hook., California [no image]. Annotation 1: Probable type of Amsinckia microcarpa Greene. Erythea 2: 191. 1894. Written on bignoniaceae sheet: cf. A. microcarpa Greene, Eryth. 2: 191. 1894. NY 00334937.

Chilopsis linearis (Cav.) Sweet subsp. arcuata (Fosberg) Henrickson Amsinckia tessellata A. Gray var. Coulter 634. Coulter 502. TCD: Chilopsis linearis (C. saligna Don), California (PL). TCD! E: Amsinckia tessellata, California [no image]. E 00225291. GH: Chilopsis linearis subsp. arcuata, California [no image]. Annotation 1: Chilopsis linearis subsp. arcuata by Henrickson in 1985. GH 384759. Amsinckia vernicosa Hook. & Arn. Coulter 496. TCD Amsinckia vernicosa TCD boraginaceae : , California (PL). ! E: Amsinckia vernicosa, California (PL). Annotation 1: Amsinckia verni- Amsinckia douglasiana A. DC. cosa Hook. & Arn. by J. Francis Macbride. E 888050. Coulter 504. K: Amsinckia angustifolia vernicosa H. & A., California (SL). K GH: Amsinckia douglasiana, California [no image]. GH 424699. 001067009. Coulter 498 in part. *Amsinckia inepta J.F. Macbr. TCD: Amsinckia angustifolia, California (PL). TCD! Coulter 501. Coulter 498. E: Amsinckia, California (PL). Annotation 1: Amsinckia inepta by Lasen E: Amsinckia angustifolia, California (PL). Written on sheet: Amsinckia in 1985. E 00225293. vernicosa Hook. & Arn. Label: Herbarium of John Ball, F.R.S. Presented to the Trustees 1891. E 888049. Amsinckia sp. Coulter 495. Cryptantha angustifolia (Torr.) Greene TCD: Amsinckia, Monterey to El Sur, California (PL). TCD! Coulter 500. Coulter 498? TCD: [no name], California (PL). TCD! TCD: Amsinckia [illegible], California (PL). TCD! E: Eritrichium hirtum, California (SL). Annotation 1: Cryptantha angus- Coulter 498 in part. tifolia (Torr.) Greene by Johnston in 1925. E 00893171. TCD: Amsinckia, California (PL). TCD! Coulter 501. Cryptantha flaccida (Lehm.) Greene TCD: Amsinckia, California (PL). TCD! Coulter 499. E: Amsinckia, California (PL). Annotation 1: “1: ?A. inepta,2,3:A. in- TCD: Myosotis flaccida, California (PL; Ex is crossed out). TCD! epta Macbr. [both] wrongly sorted” by Per Lassen in 1985. E 00225293. Coulter 513. Coulter s.n. TCD: Myosotis flaccida, California (PL; Ex is crossed out). TCD! TCD: [no name], California (PL). TCD! Coulter 514. TCD: Myosotis flaccida, California (PL; Ex is crossed out). TCD! Amsinckia spectabilis Fisch. & C.A. Mey. var. Coulter 515. Coulter 504. TCD: Myosotis flaccida, California (PL; Ex is crossed out). TCD! TCD: Amsinckia spectabilis, California (PL). TCD! BRU: Amsinckia spectabilis, California (PL). Annotation 1: Herbarium Cryptantha muricata (Hook. & Arn.) A. Nelson & J.F. Macbr. var. Olneyanum Amsinckia spectabilis Fisch. & Meyer. PBRU 00042172. Coulter 511. E: Amsinckia spectabilis, California (PL). Annotation 1: Amsinckia mi- TCD: Myosotis muricata, El Sur, California (PL). Written on sheet: Er- crocarpa Greene by J. Francis MacBride. E 00888028. itrichum. TCD! GH: Amsinckia douglasiana, California [no image]. Annotation 1: Am- BRU: Myosotis muricata, California (PL). Annotation 1: Herbarium Ol- sinckia douglasiana by Suksdorf in 1931. GH 00424669. neyanum. PBRU 00043139. K: Amsinckia spectabilis F. & M., California (SL). Annotation 1: A. dou- E: Cryptantha muricata, California [no image]. E 893490. glasiana by Ray in 1957. K 001067024. Coulter 523. TCD: [no name], California (PL). TCD! Amsinckia spectabilis Fisch. & C.A. Mey. var. microcarpa (Greene) Jeps. K: Myosotis muricata H. & Arn., California (SL). K 000895827. & Hoover Coulter 497. Cryptantha sp. TCD: Amsinckia, California (PL). Annotation 1: Type material Am- Coulter s.n. sinckia microcarpa Greene by G. Wallace in 2017. TCD! TCD: [no name], California (PL). TCD! E: Amsinckia, California (PL). Annotation 1: Amsinckia microcarpa Greene by J. Francis MacBride. Written on sheet: n. sp. E 00225295. Cynoglossum grande Lehm. GH: [no name], California (PL). Annotation 1: Nomen. Rev. Acc. To In- Coulter 520. ternat. Rules Amsinckia microcarpa Greene by J.F.M. in 1916. Note on TCD: Cynoglossum grande, Monterey, California (PL). TCD! VOLUME 37(1–2) Plantae Coulterianae 35

Coulter s.n. Coulter 479. BRU: Cynoglossum grande, California (PL). Annotation 1: Herbarium TCD: Nemophila insignis, California (PL). TCD! Olneyanum Cynoglossum grande Dougl. PBRU 00042982. TCD: Nemophila insignis, California (PL). TCD! Note: This is cited as Nemophila menziesii by Constance, 1941. Cynoglossum sp. Coulter s.n. Coulter 512. UC: Nemophila menziesii, California [no image]. UC 973557. TCD: Cynoglossum, San Francisco, California (PL). TCD! UC: Nemophila menziesii var. integrifolia, California [no image]. UC Coulter s.n. 973661. TCD: [no name], California (PL). TCD! TCD: [no name], California (PL). [Filed in Cynoglossum] TCD! Nemophila parviflora Benth. var. Coulter 480. Emmenanthe penduliflora Benth. var. TCD: Nemophila parviflora, California (PL). TCD! [two sheets] Coulter 494. TCD: Emmenanthe penduliflora, California (PL). TCD! Pectocarya penicillata (Hook. & Arn.) A. DC. Coulter 516. Eriodictyon crassifolium Benth. var. TCD: Cynoglossum penicillatum H. & A., California (PL). TCD! Coulter 601. BRU: Cynoglossum penicillatum, California (PL). Annotation 1: Pecto- TCD: Eriodictyon crassifolium, California (PL). TCD! carya penicillata A. DC. Herbarium Olneyanum. PBRU 00044603. GH: Eriodictyon crassifolium, California [no image]. Annotation 1: Eri- E: Cynoglossum penicillatum, California (PL). Annotation 1: Pectocarya odictyon crassifolium by Macbride in 1917. GH 00402820. penicillata by B. Veno in 1981. E 00782096. Eucrypta chrysanthemifolia (Benth.) Greene var. Coulter s.n. NY: Cynoglossum penicillatum H. & A., California [no image]. NY Coulter 487. 00547997. TCD: Ellisia chrysanthemifolia, California (PL). TCD! K: Ellisia chrysanthemifolia Benth., Pterophyllum foliosum Nutt., Cali- fornia (SL). Annotation 1: Eucrypta chrysanthemifolia (Benth.) Greene Phacelia ciliata Benth. by Constance in 1938. Annotation 2: ? ISOTYPE Eucrypta foliosa Coulter 489. Nutt. a synonym of Eucrypta chrysanthemifolia (Benth.) Greene [Ellisia TCD: Phacelia ciliata, California (PL). TCD! chrysanthemifolia Benth.] by Constance in 1938. K 001070902. K: Phacelia ciliata Benth., Eutoca, California (SL). K 001070997. Coulter s.n. TCD: Ellisia chrysanthemifolia, Eutoca?, California (PL). TCD! Phacelia distans Benth. Coulter 482. Heliotropium curassavicum L. var. oculatum (A. Heller) Tidestr. TCD: Phacelia tenuifolia, California (PL). TCD! [two sheets] Coulter 521. BRU: Phacelia tenuifolia¸ California (PL). Annotation 1: Herbarium Ol- TCD: Heliotropium curassavicum, California (PL). TCD! neyanum. PBRU 00045425. Coulter 522. Coulter 483 bis. TCD: Heliotropium, California (PL). TCD! UC: Phacelia distans, California [no image]. UC 973558.

Nama aretioides (Hook. & Arn.) Brand var. Phacelia douglasii (Benth.) Torr. Coulter 463 (bis). Coulter 478. TCD: Eutoca aretioides, California (PL). Annotation 1: Type material TCD: Eutoca douglasii, California (PL). TCD! Nama coulteri A. Gray by G. Wallace in 2017. TCD! BM: Eutoca douglasii, California (PL). BM 001024934. Coulter 463. GH: Nama coulteri, Nama dichotoma R. & P.var., California (PL). Anno- Phacelia minor (Harv.) F. Zimm. tation 1: N. coulteri Gray. Annotation 2: Nama coulteri A. Gray by Taylor Coulter 492. in 2015. Annotation 3: Nama coulteri Gray by C.L.H. [ = C.L. Hitch- TCD: Whitlavia grandiflora, California (PL). Annotation 1: ISOTYPE cock] in 1932. Annotation 4: Nama hispidum Gray var. coulteri (Gray) Whitlavia grandiflora Harv. by G. Wallace 2017. TCD! Brand “passes directly into the narrow leaved forms with soft pubescence K: [no name], California (SL). Annotation 1: Phacelia minor (Harv.) and branched simple forms” by J.F.M. [ = Macbride] in 1917. Annota- Thell. by Gillett in 1954. Written on sheet: Whitlavia grandiflora Harv. tion 5: LECTOTYPE Nama coulteri A. Gray by Kittredge in 2016. GH in Hook. Lond. J. Bot. t. 11. K 001070937. 00521989. Coulter 493. K: Eutoca aretioides? California (SL). K 001096149. TCD: W[hitlavia] minor, California (PL). Annotation 1: ISOTYPE Whit- lavia minor Harv. by G. Wallace in 2017. TCD! Nama sp. K: [no name], California (SL). Annotation 1: TYPE of Phacelia minor Coulter 627. (Harv.) Thell. by Gillett in 1954. Written on sheet: Whitlavia minor Harv. TCD: Nama, California (PL). TCD! in Hook. Lond. Journ. Bot. 5, t12. Written on sheet: Phacelia Whitlavia A. Gr. No. Am. Hydroph. no. 22. K 001070935. Nemophila menziesii Hook. & Arn. var. Coulter 473. Phacelia ramosissima Lehm. K: Nemophila parviflora, California (SL). Annotation 1: Nemophila men- Coulter 484. ziesii var. integrifolia Parish (N. rotata Eastw.) by Constance in 1938. K TCD: Phacelia ramosissima, California (PL). TCD! 001070894. Coulter 475. Phacelia sp. TCD: Nemophila insignis, California (PL). TCD! Coulter 483. Coulter 477. TCD: Eutoca, California (PL; Ex is crossed out). TCD! [two sheets] TCD: Nemophila menziesii, California (PL). TCD! TCD: Phacelia, California (PL; Ex is crossed out). TCD! K: N. menziesii, California (SL). Annotation 1: Nemophila menziesii var. Coulter 485. integrifolia Parish by Constance in 1938. K 001070866. TCD: Phacelia, California (PL). TCD! 36 Wallace ALISO

Phacelia viscida (Lindl.) Torr. var. Coulter 510. Coulter 490. BRU: Myosotis fulva, California (PL). Annotation 1: Herbarium Ol- TCD: Eutoca viscida, California (PL). Written on sheet: Cosmanthus vis- neyanum. PBRU 00043134. cida. TCD! Coulter s.n. K: Eutoca viscida, California (SL). Annotation 1: Phacelia viscida TCD: Myosotis fulva (PL; Ex is crossed out). TCD! (Benth.) Torr. by Gillett in 1954. K 001070944. PH: Myosotis fulva, California (SL). PH 00018838. Note: PH database Coulter 491. lists as a type in error. TCD: Eutoca viscida, California (PL). TCD! K: Eutoca viscida, California (SL). K 001070946. Note: on sheet with Plagiobothrys sp. Coulter 490 above. Coulter 507. TCD: Myosotis, California (PL). TCD! BRU: Myosotis, California (PL). Annotation 1: Herbarium Olneyanum. Pholistoma auritum (Lindl.) Lilja var. PBRU 00043150. Coulter 471. Coulter 510. TCD: Nemophila aurita, S. Miguel to S. Barbara, California (PL). TCD! TCD: [no name], California (PL). TCD! TCD: Nemophila aurita, California (PL). TCD! BRU: Nemophila aurita, California (PL). Annotation 1: Herbarium Plagiobothrys tenellus (Hook.) A. Gray Olneyanum Nemophila aurita Lindl. Annotation 2: Pholistoma aurita Coulter 507. (Lindl.) Lilja by Constance in 1948. PBRU 000447792. K: Myosotis (Dasymorpha) tenella Nutt., California (SL). K 000895925. Coulter s.n. PH: Eritrichium tinctorum, California (PL). PH 00018851. UC: Pholistoma auritum, California [no image]. UC 973662.

Pholistoma membranaceum (Benth.) Constance Coulter 469. brassicaceae TCD: Ellisia membranacea, California (PL). TCD! [two sheets] BRU: Ellisia membranacea, California (PL). Annotation 1: Herbarium Athysanus pusillus (Hook.) Greene Olneyanum Ellisia membranaceum Pursh. PBRU 00042215. Coulter 30. Coulter 470. TCD: Thysanocarpus pusillus, California (PL). TCD! TCD: Ellisia membranacea, California (PL). TCD! K: Thysanocarpus pusillus, California (SL). K 000697075. Coulter 472. Coulter s.n. TCD: Ellisia membranacea, California (PL). TCD! UC: Athysanus pusillus, California [no image]. UC 973828. Coulter s.n. Boechera sp. TCD: Ellisia membranacea, California (PL). TCD! Coulter 43. TCD: Arabis canadensis, California (PL). TCD! Plagiobothrys chorisianus (Cham.) I.M. Johnst. var. Coulter 51. Coulter 518. TCD: Turritis stricta, Monterey to El Sur, California (PL). TCD! TCD: Myosotis chorisiana Cham., California (PL). TCD! Coulter 519. †Brassica nigra (L.) W.D.J. Koch TCD: Myosotis chorisiana Cham., California (PL). Written on sheet: Er- Coulter 57. itrichum scouleri and Myosotis chorisiana. TCD! TCD: Brassica nigra, California (PL). TCD! BRU: Myositis chorisiana, California (PL). Annotation 1: Herbarium Ol- neyanum. PBRU 00043133. Cardamine californica (Nutt.) Greene E: Plagiobothrys chorisianus, California (PL). Annotation 1: Herbarium Coulter s.n. of John Ball, F.R.S. Presented by the Trustees 1891. Written on sheet: GH: Cardamine californica var. integrifolia, California [no image]. GH Eritrichum chorisanum. E 896352. 00378990. GH: Plagiobothrys chorisianus, California [no image]. Annotation 1: Plagiobothrys chorisianus by Higgins in 1972. GH 420105. Cardamine oligosperma Nutt. Coulter s.n. Coulter 53. GH: Plagiobothrys chorisianus, California [no image]. Annotation 1: TCD: Cardamine oligosperma Nutt, C. hirsuta?, California (PL). TCD! Plagiobothrys chorisianus by Higgins in 1972. GH 420021. Coulter s.n. BM: Card. oligosperma, California (PL). BM 001024438. Note: BM database identified specimen as Type in error. Plagiobothrys fulvus (Hook. & Arn.) I.M. Johnst. Coulter 502. Cardamine sp. TCD: Myosotis fulva, California (PL). TCD! Coulter 45. Coulter 503. TCD: C. angulata. Monterey, California (PL). TCD! TCD: Myosotis fulva, El Sur, California (PL). TCD! TCD: Cardamine angulata, Monterey to S. Miguel, California (PL). Coulter 505. TCD! TCD: Myosotis fulva, California (PL). TCD! Coulter 45*. Coulter 506. TCD: [no name], California (PL). TCD! Note: on sheet with Coulter 45 TCD: Myosotis fulva, S. Francisco, California (PL). TCD! above. E: Plagiobothrys fulvus, California (PL). Annotation 1: Herbarium of John Ball, F.R.S. presented by his trustees in 1891. E 896367. Note: E Caulanthus coulteri (S. Watson) Payson database lists exsiccata number as 500 in error. Coulter s.n. Coulter 508. GH: Streptanthus heterophyllus, California (PL). Annotation 1: Caulan- TCD: Myosotis fulva, California (PL). TCD! thus coulteri by Payson. Annotation 2: HOLOTYPE Caulanthus coulteri Coulter 509. by Buck in 1996. Annotation 3: LECTOTYPE Caulanthus coulteri by TCD: Myosotis fulva, California (PL). TCD! Kittredge in 2017. GH 00061150. VOLUME 37(1–2) Plantae Coulterianae 37

K: Streptanthus heterophyllus Nutt.?, California (PL). Written on sheet: Erysimum sp. Streptanthus coulteri n. sp. (S. heterophyllus p.p.) (A.G. [ = A. Gray] Coulter 60. 1868). K 000653778. TCD: [no name], Between Monterey & S. Miguel, California (PL). TCD! NY: Streptanthus heterophylla Nutt., California (PL). Annotation 1: Iso- TCD: [no name], Monterey to S. Francisco, California (PL). TCD! Note: type Caulanthus coulteri S. Watson var. coulteri by Roy Buck in 1996. on sheet with Coulter 60 above. Annotation 2: “Streptanthus heterophyllus Nutt. You do not notice the hispid pedicels in your description of S. heterophyllus, which makes me †Hornungia procumbens (L.) Hayek doubt whether our plants be the same. If mine be new I propose to call it Coulter 36. S. scapripes. Harvey” “Will you compare with Nuttall’s spec. and see if BRU: Hymenolobus divaricatus, California [no image]. PBRU it agrees in this respect. California Coulter. A. Gr.” NY 00023503. Note: 00073977. NY database lists J.M. Coulter as collector in error. Coulter s.n. BM: Hymenolobus divaricatus, California (PL). BM 1024419. Caulanthus heterophyllus (Nutt.) Payson BRU: Hymenolobus divaricatus, California [no image]. PBRU Coulter 42. 00073976. TCD: Streptanthus heterophyllus, California (PL). TCD! GH: Hymenolobus divaricatus Nutt. ex Torr. & A. Gray, California [no BRU: Strept. heterophyllus Nuttall, California (PL). Annotation image]. Annotation 1: Capsella elliptica C.A. Mey. Syn. Fl. N. Amer. 1: Herbarium Olneyanum Streptanthus heterophyllus Nutt. PBRU Annotation 2: Hutchinsia procumbens by Fernald in 1917. GH 263025. 00039255. GH: Hymenolobus divaricatus Nutt. ex Torr. & A. Gray, California [no K: Streptanthus n. sp., S. heterophyllus Nutt., Gray Monog. Strep. p. 185 image]. Annotation 1: Capsella elliptica C.A. Mey. Syn. Fl. N. Amer. GH [Gray 1864a] (SL). Written on sheet: S. coulteri A.G [ = Gray] 1868, 263025. California. K 000653779. UC: Hornungia procumbens, California [no image]. UC 973605.

Caulanthus lasiophyllus (Hook. & Arn.) Payson Lepidium nitidum Nutt. Coulter 54. Coulter 38. TCD: [no name], Between Monterey & S. Miguel, California (SL). TCD! TCD: Lepidium nitidum Nutt., — leiocarpum H. & A., California (PL). Coulter 55. TCD! TCD: S[isymbrium] deflexum, Between Monterey & San Francisco, Cal- K: Lepidium leiocarpum Herb. Hooker, California (SL). Written on sheet: ifornia (SL). TCD! Note: on sheet with Coulter 54 and Coulter 56. nitidum Nutt., Lepidium leiocarpum Hooker. K 000642813. Coulter 56. Coulter s.n. TCD: Sis[ymbrium] deflexum H., California (SL). TCD! BM: Lepidium nitidum, California (PL). BM 1024413. K: Lepidium nitidum, California (PL). Annotation 1: Herbarium Caulanthus sp. Careyanum. K 000642809. Coulter s.n. K: Lepidium nitidum, California [no label; script on sheet]. K 000642811. UC: Caulanthus, California [no image]. UC 973823. Lepidium oxycarpum Torr. & A. Gray Descurainia pinnata (Walter) Britton Coulter 29. Coulter 59. TCD: Lepidium oxycarpum Tor. & Gr., California (PL). TCD! TCD: Sisymb. canescens Nutt., California (PL). TCD! BRU: Lepidium oxycarpum, California (PL). Annotation 1: Herbarium TCD: [no name], Between Monterey & S. Miguel, California (SL). TCD! Olneyanum. PBRU 00038552. Note: on sheet with Coulter 59 above. Coulter s.n. NY: Lepidium oxycarpum, California [no image]. NY 00076581. Dithyrea californica Harv. NY: Lepidium oxycarpum Hook., California [no image]. Annotation 1: Coulter 37. Lepidium oxycarpum Torr. & Gray by Hitchcock in 1935. NY 00076580. TCD: Dithyrea californica Harv., Monterey, California (PL). Annotation NY: Lepidium oxycarpum, California [no image]. Annotation 1: Lepid- 1: ISOTYPE Dithyrea californica Harv. by G. Wallace in 2017. TCD! ium oxycarpum T. & G. by Hitchcock in 1935. NY 00076576. K: Dithyrea californica Harv., California (PL). Annotation 1: HOLO- TYPE Dithyrea californica Harvey by Al-Shehbaz in 2003. K Lepidium sp. 000484155. Coulter 39. TCD: Lepidium ruderale L., Monterey, California (PL). TCD! Erysimum capitatum (Hook.) Greene var. capitatum Coulter s.n. Coulter 61. TCD: Lepidium sativum, California (SL). TCD! TCD: Cheiranthus capitatus B, Monterey, California (PL). TCD! Coulter s.n. Lyrocarpa coulteri Hook. & Harv. GH: Cheiranthus capitatus α lanceolatus, California (PL). Annotation 1: Coulter 40. E. asperum DC. Syn. Fl. N. Amer. Annotation 2: Erysimum arkansanum TCD: Lyrocarpa coulteri Hook. & Harv., California (PL). Annotation Nutt. by Rossbach 1937. Annotation 3: Erysimum capitatum (Dougl.) 1: Type material Lyrocarpa coulteri Hook. & Harv. by G. Wallace Nov. Greene by G. Rossbach 1940. Annotation 4: Erysimum capitatum (Dougl. 2017. TCD! ex Hook.) Greene subsp. capitatum by R. Price in 1987. GH 00379568. Coulter s.n. GH: Lyrocarpa coulteri Hook. & Harv., California (PL). Annotation 1: Erysimum menziesii (Hook.) Wettst. L. coulteri Hook. & Harv. Syn. Fl. N. Amer. Written on sheet: ISOTYPE Coulter s.n. Lyrocarpa coulteri Hook. & Harv. By R.C.R. [ = Rollins] in 1939. GH GH: Cheiranthus capitatus β spathulatum, California (PL). Annotation 00019360. 1: Erysimum grandiflorum Nutt. Syn. Fl. N. Amer. Annotation 1: Erysi- mum menziesii (Hook.) Wettst., unusually many leaves on stem by Ross- Nasturtium officinale W.T. Aiton bach 1940. Annotation 2: Erysimum menziesii (Hook.) Wettst. by R. Price Coulter 44. 1990. GH 00387488. TCD: Nasturtium officinale [crossed out], California (PL). TCD! 38 Wallace ALISO

†Sisymbrium sp. Coulter 62. Coulter 58. TCD: Tropidocarpum gracile, California (PL). Annotation 1: ISOTYPE TCD: [no name], California (SL). TCD! Tropidocarpum macrocarpon Hook. & Harv. ex Greene by G. Wallace Nov. 2017. TCD! Streptanthus glandulosus Hook. subsp. K: Tropidocarpum macrocarpon Hook. & Harv., California (SL). K Coulter 41. 000642942. Note: K database lists B.L. Coulter as collector in error; this TCD: Streptanthus glandulosus, between Monterey & San Francisco, is the HOLOTYPE but is not so noted in database. California (PL). TCD! Coulter s.n. BRU: Streptanthus glandulosus, California (PL). Annotated: Herbarium GH: Tropidocarpum gracile, California [no image]. GH 00422572. Olneyanum Streptanthus glandulosus Hook. PBRU 00039254. UC: [no name], California [no image]. Annotation 1: Tropidocarpum GH: Streptanthus glandulosus, California [no image]. Annotation 1: gracile Hook. var. scabriusculum (Hook.) Greene. Annotation 2: Tropi- Streptanthus glandulosus by Al-Shehbaz in 2007. GH 422108. docarpum gracile Hook. UC 973822. Coulter s.n. E: Euklisia glandulosa, California [no image]. E 195926. UC: Streptanthus glandulosus subsp. glandulosus, California [no image]. caprifoliaceae UC 973824. Lonicera hispidula (Lindl.) Torr. & A. Gray Thysanocarpus curvipes Hook. Coulter 201. Coulter 31. TCD: Lonicera hispidula Bot. Bch., California (PL). TCD! TCD: T. pulchellus, California (PL). TCD! K: Lonicera hispidula, L. microphylla Herb. Hook., California (SL). An- TCD: Thysanocarpus pulchellus, between San Miguel & San Barbara, notation 1: Lonicera denudata Davidson & Moxley. = L. subspicata var. California (PL). TCD! johnstonii Keck and var. denudata Rehder in Munz So. Cal. Bot. p. 496 Coulter 33. by H.E. McMinn in 1937. Annotation 2: Lonicera subspicata Hook. & TCD: Thysanocarpus elegans, California (PL). TCD! Arn. var. denudata Rehder by C. Perino 1975. K 000797814. Coulter 33xx. K: Th. curvipes, California (SL). Annotation 1: Thysanocarpus curvipes Lonicera involucrata (Richardson) Spreng. var. ledebourii (Eschsch.) Hooker ssp. curvipes by Thomas and Parnell in 1971. Written on sheet: Jeps. Thysanocarpus curvipes Hooker. K 000484482. Coulter 594. Coulter 33xxx. TCD: Lonicera involucrata Banks, California (PL). TCD! K: Th. curvipes, California (SL). Written on sheet: Thysanocarpus curvipes Hooker Fl. Bor. Amer. K 000484483. Symphoricarpos mollis Nutt. Coulter s.n. Coulter 202. TCD: Thysanocarpus elegans, California (PL). TCD! TCD: Symphoricarpos mollis, California (PL). TCD! UC: Thysanocarpus curvipes, California [no image]. UC 973829. GH: Symphoricarpos mollis, California [no image]. Annotation 1: Sym- phoricarpos mollis by Jones in 1939. GH 00386321. Thysanocarpus laciniatus Nutt. var. K: Symph. mollis Nutt., California (SL). K 000797846. Coulter 33x. K: Thysanocarpus n. sp? T. crenatus Nutt.?, California (SL). Annota- Symphoricarpos sp. tion 1: Thysanocarpus laciniatus Nutt. by Thomas and Parnell in 1971. Coulter 203. K 000484489. TCD: Symphoricarpos occidentalis, California (PL). TCD! Coulter 32xx. K: [no name], California (SL). Annotation 1: Thysanocarpus laciniatus Nutt. by Thomas and Parnell in 1971. K 000484487. caryophyllaceae Coulter 35. Arenaria sp. TCD: [no name], California (PL). TCD! Coulter 73. Coulter s.n. TCD: Arenaria tricoidea Nutt., California (SL). TCD! TCD: Thysanocarpus crenatus, California (PL). TCD! BRU: Arenaria tricoidea Nutt., California (PL). Annotation 1: Arenaria UC: Thysanocarpus laciniatus, California [no image]. UC 973606. trichoidea Nutt. PBRU 00040862. Coulter s.n. Thysanocarpus sp. TCD: Arenaria trinervia (SL). TCD! Coulter 32x. BM: Arenaria trichophylla Nutt., California (PL). BM 1024695. TCD: Thysanocarpus angustus, California (PL). TCD! [two sheets] Minuartia californica (A. Gray) Mattf. Tropidocarpum gracile Hook. Coulter s.n. Coulter 46. UC: Minuartia californica, California [no image]. UC 973815. TCD: [no name], Between Monterey & San Francisco, California (PL). TCD! Minuartia douglasii (Torr. & A. Gray) Mattf. Coulter 47. Coulter s.n. TCD: [no name], California (PL). TCD! TCD: Alsine douglasii Fenzl, California (SL). TCD! K: Tropidocarpum gracile, California (SL). K 000642941 Note: K database lists as Coulter s.n. and B.L. Coulter as collector in error. Minuartia tenella (J. Gay) Mattf. Coulter 48. Coulter 83. TCD: [no name], Monterey to San Miguel, California (PL). TCD! Note: TCD: Arenaria tenella Nutt., California (SL). TCD! on sheet with Coulter 47 above. BRU: Arenaria tenella Nutt., California (PL). Annotation 1: Herbarium Coulter 49. Olneyanum Arenaria tenella. PBRU 00040851. TCD: Tropidocarpum scabriusculum, California (PL). TCD! Coulter 84. Coulter 50. TCD: Arenaria tenella, California (SL). TCD! Note: on sheet with Coul- TCD: Tropidocarpum gracile, California (PL). TCD! ter 83 above. VOLUME 37(1–2) Plantae Coulterianae 39

Coulter s.n. TCD: Obione coulteri Moq., California (PL). Annotation 1: ISOTYPE BM: Arenaria tenella, Greniera tenella, California (PL). BM 1024705. Obione coulteri Moq. by G. Wallace in 2017. TCD! K: Obione coulteri Moq., California (SL). Annotation 1: HOLOTYPE Sagina decumbens (Elliott) Torr. & A. Gray subsp. occidentalis (S. Wat- Obione coulteri Moquin-Tandon by Welsh in 1999. K 000898684. Note: son) G.E. Crow K database lists J.M. Coulter as collector in error. Coulter 82. TCD: Sagina decumbens, Spergula saginoides, California (SL). TCD! Atriplex sp. Coulter 669. Sagina sp. TCD: Atriplex, California (PL). TCD! Coulter s.n. UC: Sagina, California [no image]. UC 973814. †Beta vulgaris L. subsp. maritima (L.) Arcang. Coulter 684. Silene antirrhina L. TCD: Beta, California (PL). TCD! Coulter 85. TCD: Silene antirrhina L., California (SL). TCD! Chenopodium californicum (S. Watson) S. Watson Coulter 683. Silene douglasii Hook. var. douglasii BRU: [no name], California (PL). Annotation 1: Herbarium Olneyanum. Coulter 72. PBRU 00039619. TCD: Silene douglasii, San Miguel to San Barbara, California (SL). K: [no name] [no location] (SL). Annotation 1: Ch. californicum S. Wats. TCD! Annotation 2: Blitum bonus-henrius Reichenb. var. erosus. K 000898460.

†Spergularia rubra (L.) J. Presl & C. Presl Chenopodium sp. Coulter 77. Coulter 688. TCD: Arenaria rubra, California (SL). TCD! TCD: Chenopodium, California (PL). TCD! Coulter 78. Coulter 689. TCD: Arenaria rubra, California (SL). TCD! TCD: Chenopodium, California (PL). TCD! Coulter 79. Coulter 690. TCD: [no name], Monterey, California (SL). TCD! Note: on sheet with TCD: Chenopodium, California (PL). TCD! Coulter 77 above. Coulter 80. †Dysphania anthelmintica (L.) Mosyakin & Clemants TCD: Arenaria rubra, Monterey, California (SL). TCD! Coulter 683. TCD: [no name], San Miguel to San Barbara (SL). TCD! Note: on sheet K: Boubiea anthelmintica, California (SL). K 000898460. with Coulter 80 above. Coulter 81. Suaeda sp. TCD: [no name], San Miguel to San Barbara (SL). TCD! Note: on sheet Coulter 686. with the two Coulter 80 above. TCD: [no name], California (PL). TCD! Coulter s.n. BRU: Arenaria rubra, California (PL). Annotation 1: Herbarium Ol- cistaceae neyanum Spergularia rubra Pers. PBRU 00040788. Helianthemum scoparium Nutt. †Stellaria media (L.) Vill. Coulter s.n. Coulter 76. TCD: Helianthemum scoparium, San Miguel to S. Barbara, California TCD: Stellaria media, Monterey, California (SL). TCD! (SL). TCD! TCD: Hel. scoparium, Monterey, California (SL). TCD! Stellaria nitens Nutt. GH: Helianthemum scoparium, California [no image]. Annotation 1: He- Coulter 74. lianthemum scoparium by Wilbur & Daoud in 1963. GH 420299. K: Stell. nitida [sic], California (SL). Annotation 1: Stellaria nitens Nutt. by E.L. Greene in 1889. K 000723607. Note: K database lists S. Coulter as collector in error. cleomaceae Coulter s.n. Peritoma arborea (Nutt.) H.H. Iltis var. TCD: Stellaria nitens, Monterey to El Sur, California (SL). TCD! Coulter 65. TCD: [no name], Monterey, California (SL). TCD! K: Isomeris arborea Nutt., St. Diego (SL). [Filed as Cristatella arborea]. K 000643879. Coulter s.n. chenopodiaceae TCD: Isomeris arborea Nutt., St. Diego (SL). TCD! GH: Isomeris arborea, California [no image]. Annotation 1: Isomeris ar- Arthrocnemum sp. borea by Vanderpool in 1988. GH 00378637. Coulter 691. TCD: Arthrocnemum fruticosum var. californicum, California (PL). TCD! convolvulaceae

Atriplex californica Moq. Calystegia sp. Coulter 685. Coulter 554. TCD: Atriplex californica, California (PL). TCD! TCD: [no name], California (PL). TCD! K: [no name], California. Annotated: Atriplex californica Moq. (SL). K 000197038. Note: K database lists J. Coulter as collector in error. cornaceae Atriplex coulteri (Moq.) D. Dietr. Cornus sericea L. subsp. sericea Coulter 687. Coulter 553. 40 Wallace ALISO

TCD: Viburnum, California (PL). TCD! Coulter s.n. TCD: Gaultheria shallon, California (PL). TCD! UC: Gaultheria shallon, California [no image]. UC 973747.

cucurbitaceae Vaccinium ovatum Pursh *Ibervillea sonorae (S. Watson) Greene Coulter 245. Coulter 200. TCD: Vaccinium ovatum, California (PL). TCD! [two sheets] TCD: Sicydium lindheimeri, California (PL). TCD! GH: Vaccinium ovatum, California [no image]. GH 00363214. Coulter s.n. Marah oregana (S. Watson) Howell BRU: Vaccinium ovatum, California (PL). Annotation label: Herbarium Coulter 199. Olneyanum. PBRU 00007700. TCD: Sycios [Sicyos] oregana, California (PL). TCD! UC: Vaccinium ovatum, California. [no image]. UC 973819. euphorbiaceae datiscaceae Croton californicus Müll. Datisca glomerata (C. Presl) Baill. Coulter 666. Coulter 706. TCD: Croton¸ California (PL). Written on sheet: Hendecandra [sic]. TCD: [no name], California (PL). Written on sheet: Tricerastes Presl. TCD! TCD! Euphorbia cf. melanadenia Torr. Coulter 673. ericaceae TCD: Euphorbia, California (PL). TCD! Note: V. Steinmann referred image to E. melanadenia in 2016. Arbutus menziesii Pursh Coulter 714. Euphorbia spathulata Lam. TCD: Arbutus menziesii, California (SL). TCD! Coulter 672. K: Euph. platyphylla L, California (SL). Annotation 1: Euphorbia Arctostaphylos crustacea Eastwood subsp. crinata (J.E. Adams) V.T. spathulacea by Berry and Peirson in 2012. K 001080375. Parker, M.C. Vasey & J.E. Keeley Coulter 678. Coulter s.n. TCD: Euphorbia, California (PL). TCD! Note: image identified by V. UC: Arctostaphylos tomentosa subsp. crinata, California [no image]. UC Steinmann in 2016. 973818. K: E. platyphylla, California (SL). K 001080376. Arctostaphylos hookeri G. Don subsp. Coulter 679. Coulter 243. TCD: [no name], California (PL). TCD! Note: image identified by V. GH: Arctostaphylos hookeri, California [no image]. GH 00350453. Steinmann in 2016. fabaceae Arctostaphylos pumila Nutt. Coulter 240. Acmispon americanus (Nutt.) Rydb. var. americanus TCD: Arbutus pungens? California (PL). TCD! Coulter 427. K: A. pungens?, California (SL). K 000780442. TCD: Hosackia purshiana, Monterey to San Miguel, California (SL). TCD! Arctostaphylos sp. K: Hosackia purshiana, Monterey to San Miguel, California (PL). Writ- Coulter 243. ten on sheet: H. purshiana. K 000873278. TCD: Arbutus pungens, California (PL). TCD! K: [no name], California (SL). Written on sheet: Hosackia purshiana K: Arb. pungens (SL). Annotation 1: Xerobotrys venolosus Nutt. by C.C. Benth. Gray Syn. Hosack. p. 352. K 000873276. Parry and on label A. hookeri Don. K 000780445. Coulter 427x. TCD: Hosackia purshiana, California (SL). TCD! Arctostaphylos tomentosa (Pursh) Lindl. subsp. Coulter s.n. Coulter 241. K: Hosackia purshiana, California (PL). K 000873281. TCD: Arbutus tomentosa β, El Sur, California (PL). TCD! GH: Arctostaphylos tomentosa, California [no image]. GH 00362269. Acmispon cytisoides (Benth.) Brouillet Coulter 241x. Coulter 425. TCD: Arbutus tomentosus β, California (PL). TCD! TCD: Hosackia cytisoides Benth., California (SL). TCD! Coulter 242. Coulter s.n. TCD: Arbutus tomentosus α, California (PL). TCD! BM: Hosackia cytisoides Benth., California (PL). BM 001042515. GH: Arctostaphylos tomentosa, California [no image]. GH 00362268. Coulter s.n. Acmispon glaber (Vogel) Brouillet var. BRU: Arbutus tomentosa, California (PL, top line cut off). Annotation 1: Coulter 424. Herbarium Olneyanum. PBRU 00006858. K: [no name], California (SL). Written on sheet: Hosackia scoparia T. & NY: Arbutus tomentosa α, California (PL). Annotation 1: A. tomentosa G. K 000873247. Torr. & Gray, Fl. N. Amer. NY 02520537. Coulter s.n. NY: Arbutus tomentosa β, California (PL). NY 02520538. Note: on sheet TCD: Hosackia scoparia Nutt., California (SL). TCD! with Coulter s.n. above and filed as Arctostaphylos vestita. Acmispon maritimus (Nutt.) D.D. Sokoloff var. Gaultheria shallon Pursh Coulter 421. Coulter 237. TCD: [no name], California (SL). Annotation 1: I think all these n. 421 TCD: Gaultheria shallon, California (PL). TCD! must be Hosackia maritima Nutt. though there are more leaflets than he BRU: Gaultheria shallon, California (PL). PBRU 00007093. describes. TCD! VOLUME 37(1–2) Plantae Coulterianae 41

BRU: Hosackia maritima Nutt., California (PL). Annotation 1: Herbar- K: [no name], California (SL). Written on sheet: Astragalus didymocar- ium Olneyanum Hosackia maritima Nutt. PBRU 00034533. pus, Gray Monog. Astrag. p. 198 [Gray 1864b]. K 000999254. Note: K Coulter s.n. database lists B.L. Coulter as collector in error. BM: Hosackia maritima, California (PL). BM 000901403. Coulter 440. TCD: Astragalus didymocarpus Hook. & Arn. var. major, S. Miguel to Acmispon rigidus (Benth.) Brouillet S. Barb. (SL). TCD! Note: on sheet with Coulter 338 above. Coulter 420. K: [no name], California (SL). K 000999253. [On sheet with Coulter 438 K: Hosackia rigida Benth., Monterey, California (PL). Annotation 1: above]. Lectotype of Hosackia rigida Benth. by Sokoloff in 1994. K 001276907. K: Astragalus didymocarpus Hook. et Arn. var. major. Caules basi Coulter s.n. ramosi, S. Miguel to Sa. Barbara, California (PL). K 000999252. TCD: Hosackia rigida sp. n., California (SL). Annotation 1: Type mate- Astragalus lentiginosus Douglas var. borreganus M.E. Jones rial Hosackia rigida Benth. by G. Wallace in 2016. TCD! Coulter 431. TCD: Astragalus coulteri sp. n., Monterey, California (SL). Annotation Acmispon strigosus (Nutt.) Brouillet 1: Type material Astragalus coulteri Benth. by G. Wallace in 2017. TCD! Coulter 429. K: [no name], California (SL). Annotation 1: This is apparently part of TCD: [no name], California (SL). Annotation 1: I believe this includes type of A. coulteri Benth. by Barneby in 1962. K 000478311. Note: filed Hosackia strigosa and H. rubella Nutt. which are most probably one as Astragalus candidissimus, the other specimen on sheet. species. Written on sheet: strigosa & rubella. TCD! K: Astragalus coulteri β, Monterey, California (PL). Written on sheet: K: [no name], California (SL). Written on sheet: Hosackia strigosa Nutt., never grow [sic] at Monterey; syn. A. arthu-schottii, Gray Monog. Astrag. Gray Syn. Hosack. p. 351. K 000873310. p. 209 [Gray 1864b]. K 000263892.

Acmispon wrangelianus (Fisch. & C.A. Mey.) D.D. Sokoloff Astragalus nuttallii (Torr. & A. Gray) J.T. Howell var. Coulter 442. Coulter 432. TCD: Hosackia subpinnata T. & G., California (SL). TCD! TCD: [no name], Monterey to San Gabriel, California (SL). Annotation 1: Phaca densifolia var. [illegible]. TCD! Amorpha californica Nutt. var. californica Coulter 435. Coulter 399. TCD: [no name], Monterey to El Sur, California (SL). Annotation 1: GH: Amorpha californica, California [no image]. Annotation 1: Amor- Phaca densifolia Sm. var. Written on sheet: densifolia. TCD! pha californica by Wilbur in 1966. GH 00366199. Coulter 437. Coulter s.n. K: [no name], California (SL). Written on sheet: Astragalus menziesii, TCD: Amorpha californica Nutt., Monterey to S. Miguel, California Gray Monog. Astrag. p. 217 [Gray 1864b] K 000999371. Note: filed as (PL). Note on label: Probably Amorpha californica but not yet in flower. Astragalus menziesii; K database lists B.L. Coulter as collector in error. TCD! Astragalus sp. Astragalus asymmetricus E. Sheld. Coulter 439. Coulter 434. TCD: Homolobus multiflorus Torr. et Gr. var. foliolis floribusque numero- TCD: [no name], S. Miguel to San Barbara, California (SL). Annotation sis, California (SL). TCD! 1: Phaca leucophylla Hook. & Arn. Written on sheet: leucophylla. TCD! K: Phaca canescens, California (SL). Annotation 1: Astragalus leuco- Dalea mollis Benth. phyllus Torr. & Gray, Gray Monog. Astrag. p. 217 [Gray 1864b]. K Coulter 430. 000999373. Note: filed as Astragalus curtipes; K database lists B.L. Coul- GH: Dalea mollis Benth. n. sp., California (PL). Annotation 1: D. mol- ter as collector in error. lis Benth. Syn. Fl. N. Amer. Det. B.L.R. [ = Robinson]. Annotation 2: K: Astragalus leucophylla Hook. & Arn., California (PL). K 000999381. ISOTYPE Dalea mollis Benth. Pl. Hartw. p. 306. 1848. Barneby Mem. Note: K database lists B.L. Coulter as collector in error. NYBG 27: 162. 1977. Written on sheet: TYPE. GH 00053604. Coulter s.n. K: Dalea mollis Benth., Monterey, California (PL). Annotation 1: TYPE TCD: Astragalus leucophylla Hook. & A., California (SL). TCD! Note: SPECIMEN Herb. Kew. Written on sheet: Dalea mollis Benth. K on sheet with Coulter 434. 000082207. K: [no name], California (SL). Written on sheet: Dalea mollis Benth. K Astragalus crotalariae (Benth.) Torr. 000082207. [On sheet with Coulter 430 above] Coulter 436. Coulter s.n. TCD: Phaca crotalariae sp. n., Monterey, California (SL). Annotation 1: TCD: Dalea mollis sp. n., California (SL). Annotation 1: Type material ISOTYPE Phaca crotalariae Benth. by G. Wallace in 2017. TCD! Dalea mollis Benth. by G. Wallace in 2017. TCD! K: Phaca crotalaria, Monterey, California (PL). K 000263923. Hosackia crassifolia Benth. var. crassifolia Astragalus curtipes A. Gray Coulter 428. Coulter 433. TCD: Hosackia crassifolia Benth., California (SL). Written on sheet: K: Astragalus leucophylla Hook. & Arn., California (PL). Annotation 1: crassifolia Benth. TCD! A. curtipes Gray by Barneby in 1962. K 000999379. Note: K database K: [no name], California (SL). Written on sheet: Hosackia crassifolia lists B.L. Coulter as collector in error. Benth. Gray Syn. Hosack. p. 350. K 000873197. K: Phaca leucophylla Hook., Ast. leucophyllus Torr. & Gray, California Coulter s.n. (SL). Written on sheet: Astragalus leucophyllus Torr. & Gr., Gray Monog. TCD: Hosackia leptocarpa, H. crassifolia Nutt. non Kunth, California Astrag. p. 218 [Gray 1864b]. K 000999376. Note: K database lists B.L. (SL). [2 sheets] TCD! Coulter as collector in error. Hosackia gracilis Benth. Astragalus didymocarpus Hook. & Arn. var. Coulter 423. Coulter 438. TCD: Hosackia gracilis Benth., Monterey, California (SL). Written on TCD: Astragalus didymocarpus Hook. & Arn., California (SL). TCD! sheet: gracilis TCD! 42 Wallace ALISO

K: [no name], California (Script on sheet). K 000873326. Note: filed as TCD: Lupinus douglasii Ag. scarcely different from L. albifrons, Cali- Lotus formosissimus fornia (SL). TCD! UC: Lupinus albifrons var. douglasii, California [no image]. UC Hosackia oblongifolia Benth. var. oblongifolia 1713155. Coulter s.n. Coulter 384, 394. TCD: Hosackia oblongifolia sp. n., California (SL). Note on label: unique K: Lupinus douglasii Ag., Monterey, California (PL). K 000829310. specimen. Annotation 1: Type material by G. Wallace in 2016. Written on Coulter 394. sheet: oblongifolia. TCD! BRU: Lupinus douglasii, California (PL). Annotation 1: Herbarium Ol- neyanum Lupinus douglasii Agardh. PBRU 00034813. Hosackia stipularis Benth. var. stipularis Coulter s.n. Coulter 441. TCD: Lupinus douglasii, California (PL). Written on sheet: L. douglasii. K: Hosackia stipularis, Monterey to El Sur, California (PL). Annotation TCD! 1: Hosackia stipularis Benth. by Sokoloff in 1999. K 000873199. TCD: [no name], California (SL). Written on sheet: Lupinus douglasii. K: Hosackia stipulacea, California (SL). Written on sheet: Hosackia stip- TCD! ularis Bentham Gray Syn. Hosackia p. 329. K 000873202. NY: Lupinus douglasii, California (SL). Annotation 1: Ex Herbarium S.T. Coulter s.n. Olney. NY 02586456. TCD: Hosackia stipulacea, Monterey to El Sur, California (SL). TCD! NY: Lupinus douglasii, California (PL). NY 02586457. Lathyrus sp. Lupinus arboreus Sims Coulter s.n. Coulter 380. TCD: Lathyrus venosus Muhl. var (SL). TCD! GH: Lupinus arboreus, California [no image]. Annotation 1: Lupinus TCD: Lathyrus venosus Muhl., California (SL). TCD! rivularis. GH 00366755. TCD: Lathyrus venosus var. angustifolia, California (SL). TCD! K: [no name], California (SL). Written on sheet: arboreus Sims. A.G. = K Lupinus affinis J. Agardh [ A. Gray]. 000829408. Coulter 382. Lupinus benthamii Heller K Lupinus affinis : [no name], California (SL). Annotation 1: Agardh. by Coulter 370. K Dunn in 1964. 000829544. TCD: Lupinus leptophyllus Benth., Monterey to San Miguel, California Coulter 386. (SL). TCD! TCD: Lupinus affinis Ag., Monterey to El Sur, California (SL). TCD! TCD: Lupinus leptophyllus, California (SL). TCD! UC: Lupinus affinis, California [no image]. UC 1713153. GH: Lupinus leptophyllus, Monterey to San Miguel, California [no im- Lupinus benthamii GH Lupinus albifrons Benth. var. age]. Annotation 1: by Dziekanowski. 00065559. K: Lupinus leptophyllus Benth., California (PL). Annotation 1: Lupinus Coulter 375, 379, 390, 391. benthamii Heller by Dunn in 1966. K 000829377. K: Lupinus albifrons, California (PL). K 000829453. K: Lupinus leptophyllus Benth., California (SL). Annotation 1: Lupinus Coulter 379, 390, 392, 395, 396. benthamii Heller by Dunn in 1966. Label: Herb. J. Ball, F.R.S. August TCD: Lupinus albifrons, Monterey, S. Miguel, S. Barbara, California 1800. K 000829378. (PL). K 000829453. K: Lupinus leptophyllus Benth., Monterey to S. Miguel, California (PL). Coulter 390. Annotation 1: Lupinus benthamii Heller by Dunn in 1966. K 000829379. TCD: [no name], Monterey to S. Barbara (SL). Written on sheet: L. alb- ifrons Benth. TCD! Lupinus bicolor Lindl. Coulter 391. Coulter 377. TCD L. : [no name], Monterey to San Miguel (SL). Written on sheet: TCD: Lupinus micranthus, Monterey to San Francisco, California (SL). albifrons. TCD! TCD! Coulter 392. TCD: [no name], Monterey to San Miguel, California (SL). TCD! TCD: L. albifrons, San Miguel to San Francisco (SL). TCD! Note: on TCD: Lupinus micranthus, California (SL). TCD! Note: on sheet with sheet with Coulter 390 above. Coulter 377 [no name] above. Coulter 395 . TCD: [no name], San Miguel to San Barbara, California (SL). Annotation TCD L. albifrons = : Benth. Monty. [ Monterey] to S. Francisco, Califor- 1: Lupinus micranthus Douglas var. TCD! nia (PL). Written on sheet: L. albifrons. TCD! Note: on sheet with Coulter Coulter s.n. 391 and Coulter s.n. above. TCD: Lupinus bicolor Lindl., California (SL). TCD! Coulter 396. BRU: Lupinus micranthus Lindl., California (PL). Annotation 1: Herbar- TCD: Lupinus albifrons Benth., Mt [ = Monterey] to San Miguel (SL). ium Olneyanum Lupinus micranthus Dougl. PBRU 00034855. TCD! MEL: Lupinus polycarpus Greene, California [no image]. MEL GH: Lupinus albifrons, California [no image]. Annotation 1: Lupinus 0308004a. albifrons var. eminens by Piper. GH 366709. NY: Lupinus micranthus Lindl., California [no image]. NY 02599428. K: [no name], California (SL). Note on label: (Gordon). Written on sheet: NY: Lupinus micranthus Lindl., California [no image]. NY 02599540. albifrons. K 000829454. NY: Lupinus micranthus Lindl., California [no image]. NY 02599541. Coulter s.n. TCD: [no name], San Miguel to San Barbara (SL). Written on sheet: L. Lupinus concinnus J. Agardh albifrons. TCD! Coulter 372. [Coulter s.n.] TCD: Lupinus concinnus Ag., Monterey to S. Francisco, California (SL). TCD: [no name], San Miguel to San Barbara (SL). Written on sheet: L. TCD! albifrons. TCD! Note: on sheet with Coulter 391 [no name] above. BRU: Lupinus concinnus Ag., California (PL). Annotation 1: Herbarium Olneyanum Lupinus concinnus Ag. PBRU 00034794. Lupinus albifrons Benth. var. douglasii (J. Agardh) C.P. Sm. K: L. concinnus Ag., Monterey to S. Francisco and S. Miguel, California Coulter 384. (PL). K 000829336. VOLUME 37(1–2) Plantae Coulterianae 43

K: L. concinnus Ag., California (SL). Annotation 1: Lupinus concinnus Lupinus microcarpus Sims var. densiflorus (Benth.) Jeps. Agardh subsp. concinnus by Dunn in 1966 (Aliso 1966). K 000829333. Coulter 371. Coulter 372x. TCD: Lupinus densiflorus,My[= Monterey] to S. Francisco, California K: L. concinnus Ag., California (SL). Annotation 1: Lupinus concinnus (SL). TCD! Agardh subsp. concinnus by Dunn in 1966 (Aliso 1966). K 000829332. Coulter 381. Note: K database lists as Coulter s.n. in error. TCD: Lupinus menziesii Ag., California (SL). Written on sheet: L. men- Coulter s.n. ziesii. TCD! TCD: [no name], California [no label]. Written on sheet: Lupinus concin- K: [no name], California (SL). Annotation 1: Lupinus microcarpus Sims nus Ag., TCD! var. densiflorus (Benth.) Jeps. by Riggins in 1989. Written on sheet: L. K: Lupinus concinnus Ag., California (PL). Annotation 1: Lupinus menziesii according to Mr. Bentham Pl. Hartw. p. 303. K 000829603. concinnus Agardh subsp. concinnus by Dunn in 1966. K 000829337. K: Lupinus menziesii Ag., S. Miguel to S. Barbara, California (PL). An- NY: Lupinus concinnus Ag., California (PL). NY 00015825. notation 1: Lupinus densiflorus Benth. by Wainwright in 1972. Annota- UC: Lupinus concinnus, California [no image]. UC 973804. tion 2: Lupinus microcarpus Sims var. densiflorus (Benth.) Jeps. by Rig- gins in 1989. K 000829609. Lupinus excubitus M.E. Jones var. hallii (Abrams) C.P. Sm. Coulter s.n. Coulter 394. TCD: [no name], My [ = Monterey] to S. Miguel. TCD! Note: on sheet GH: Lupinus excubitus var. hallii, California [no image]. Annotation 1: with Coulter 371. Lupinus excubitus var. hallii by Smith in 1928. GH 00369017. Lupinus microcarpus Sims var. microcarpus Coulter 396. Coulter 371. GH: Lupinus excubitus var. hallii, California [no image]. Annotation 1: K: L. densiflorus Benth., “California (Douglas) Coulter” (SL). Annota- Lupinus excubitus var. hallii by Smith in 1928. GH 00369018. tion 1: Lupinus microcarpus Sims var. microcarpus by Riggins in 1989. K 000829611. Lupinus hirsutissimus Benth. Coulter 373. Lupinus nanus Benth. TCD: L. hirsutissimus B., Monterey to El Sur, California (SL). Written Coulter 374. on sheet: L. hirsutissimus. TCD! TCD: [no name], San Miguel to San Barbara (SL). TCD! TCD: [no name], Monterey to San Miguel, California (SL). Written on Coulter 375. sheet: L. hirsutissimus. TCD! TCD: L. nanus, M[onterey] to S. Fco. [Francisco] (SL). TCD! Note: on BRU: Lupinus hirsutissimus, California (PL). Annotation 1: Herbarium sheet with Coulter 374 above. Olneyanum Lupinus hirsutissimus Benth. PBRU 00034819. Coulter 376. K: L[upinus] hirsutissimus Benth., California (SL). K 000829479. TCD: Lupinus nanus, San Miguel to San Barbara, California (SL). TCD! K: Lupinus hirsutissimus Benth. Monterey to S. Miguel and El Sur, Cal- Coulter 374, 375, 376. ifornia (PL). Written on sheet: L. hirsutissimus. K 000829296. K: Lupinus nanus Dougl., Monterey to S. Francisco, S. Miguel to S. Bar- NY: Lupinus hirsutissimus Benth., California [no image]. NY 02598222. bara, California (PL). K 000829597. Coulter s.n. Coulter s.n. TCD: Lupinus hirsutissimus Benth., California (SL). TCD! TCD: Lupinus nanus Dougl., California (SL). TCD! Note: on sheet with NY: Lupinus hirsutissimus, California. On label: Ex Herbarium S.T. Ol- Coulter 374 and 375 above. ney. Note on label: Ex Herb. Chapman. NY 02610933. TCD: [no name], Monterey to San Miguel (SL). TCD! Note: on sheet with Coulter 376 above. BRU: Lupinus nanus, California (PL). Annotation 1: Herbarium Ol- Lupinus latifolius J. Agardh var. neyanum Lupinus nanus Dougl. PBRU 00034873. Coulter 382. NY: Lupinus nanus, California (SL). NY 02599541. Note: in NY TCD : [no name], San Miguel to San Barbara, California (SL). Written database as L. micrantha. Lupinus affinis latifolius TCD on sheet: var. . ! UC: Lupinus nanus, California [no image]. UC 973806. Coulter 383. TCD: Lupinus cytisoides Ag., California (SL). TCD! Lupinus rivularis Lindl. TCD: [no name], San Miguel to San Barbara (SL) [on sheet with L. cyti- Coulter 380. soides]. TCD! TCD: Lupinus rivularis, California (SL). TCD! BRU: Lupinus cytisoides Ag., California (PL). Annotated: Herbarium Ol- TCD: [no name], Monterey to San Gabriel, California (SL). TCD! Note: neyanum Lupinus cytisoides Ag. PBRU 00034796. on sheet with Coulter 380 above. UC: Lupinus latifolius, California. UC 00170322. TCD: [no name] San Miguel to San Barbara, California (SL). TCD! Coulter 387. Note: on sheet with Coulter 385 L. rivularis. TCD: Lupinus affinis var. latifolia, California (SL). TCD! BRU: Lupinus rivularis Lindl., California (PL). Annotation 1: Herbarium Coulter 388. Olneyanum Lupinus rivularis Lindl. PBRU 00034925. TCD: Lupinus nanus Dougl. var. latifolius an sp. distincta?, St. Miguel UC: Lupinus rivularis, California [no image]. UC 00170316. to S. Barbara, California (SL). TCD! Coulter 385. [Coulter] 389. TCD: L. rivularis Lindl., Monterey to El Sur, California (SL). TCD! TCD: [no name], California (SL). TCD! Note: on sheet with Coulter 387 Note: on sheet with Coulter 380, San Miguel to San Barbara, above. and [Coulter s.n.] above. Coulter s.n. Coulter s.n. TCD: [no name], Monterey to El Sur (SL). TCD! Note: on sheet with TCD: L. affinis var. latifolia,My[= Monterey] to S. Francisco (SL). two Coulter 380 above. TCD! Note: on sheet with Coulter 382 above. TCD: Lupinus cytisoides Ag. var., California (SL). TCD! Note: on sheet Lupinus sparsiflorus Benth. with Coulter 383 [no name] above. Coulter s.n. TCD: [no name], Monterey to El Sur (SL). TCD! Note: on sheet with TCD: Lupinus sparsiflorus sp., n. California (SL). Note on label: was Coulter 387 above. mixed with L. truncatus. Written on sheet: L. sparsiflorus Benth. TCD! UC: Lupinus latifolius, California [no image]. UC 973803. [needs annotation as Isotype] 44 Wallace ALISO

GH: Lupinus sparsiflorus Benth. n. sp., California (PL). Stamped Prosopis pubescens Benth. on sheet: ISOTYPE. Listed in GH database as possible TYPE. GH Coulter s.n. 00065672. TCD: Prosopis pubescens sp. n., Mont[erey] to S. Miguel, California K: Lupinus sparsiflorus, California (PL). Annotation 1: Bentham’s TYPE (SL). Annotation 1: Type material Prosopis pubescens Benth. by G. Wal- Lupinus sparsiflorus subsp. sparsiflorus, see Aliso 1966 by Dunn in 1966. lace in 2017. TCD! Written on sheet: sparsiflorus. K 000829553. UC: Lupinus sparsiflorus, California [no image]. UC 00693878. Prosopis sp. Coulter s.n. Lupinus sp. TCD: Prosopis dulcis Kunth, California (SL). TCD! Coulter 377. K: L. bicolor? [just a fragment folder with pencil script below]. K Psorothamnus spinosus (A. Gray) Barneby 000829433. Coulter s.n. Coulter 398. TCD: Psoralea, California (PL). TCD! K: [no name], California [no image]. K 000829365. Note: filed as Lupi- nus bicolor . Trifolium albopurpureum Torr. & A. Gray Coulter 394x. Coulter 417. TCD: Lupinus ornatus Dougl. L. argenteus Ag., California (SL). TCD! TCD: [no name], California (SL). Annotation 1: Trifolium albopur- Coulter s.n. pureum T. & G. Written on sheet: albopurpureum. TCD! UC: Lupinus, California [no image]. UC 973805. GH: Trifolium albopurpureum, California [no image]. Annotation 1: Tri- Lupinus succulentus K. Koch folium macraei var. albopurpureum (Torr. & A. Gray) Greene by Martin GH Coulter 387x. in 1942. 00245594. K Trifolium albopurpureum K: [no name], California (SL). Annotation 1: Lupinus succulentus Dou- : Torr. & Gray, Monterey to S. Miguel, Cali- fornia (PL). K 001051207. glas ex Koch by Eastwood in 1911. K 000829543. K: [no name], California (SL). Annotation 1: T. neolagopus Loja by = Lupinus truncatus Nutt. P.B.K. [ Kennedy] in 1911. K 001051202. Coulter 369. Coulter s.n. TCD: [no name], Monterey to S. Miguel (SL). Written on sheet: L. trun- TCD: Trifolium albopurpureum, California (SL). Annotation 1: scarcely catus. TCD! the same as T. macraei. TCD! Coulter 378. BRU: Trifolium albopurpureum, California (PL). Annotation 1: Herbar- TCD: Lupinus truncatus, San Miguel to San Barbara, California (SL). ium Olneyanum Trifolium albopurpureum T. & G. PBRU 00035603. Written on sheet: L. truncatus. TCD! Trifolium ciliolatum Benth. TCD: [no name], San Miguel to San Barbara, California (SL). Written on Coulter 414. sheet: Lupinus truncatus. TCD! Note: on sheet with 369 and 378 above. K: Trifolium ciliolatum, S. Miguel to S. Barbara, California (PL). Written Coulter 376x & 397. on sheet: Type. K 001051164. TCD: Lupinus truncatus, California (SL). TCD! K: [no name], California (SL). Annotation 1: Trifolium ciliolatum Benth. Coulter 369, 378, 397. by P.B.K. in 1911. K 001051170. K: Lupinus truncatus Nutt., Monterey to S. Miguel & S. Miguel to S. Coulter s.n. Barbara, California (PL). Written on sheet: L. truncatus. K 000829537. TCD: Trifolium ciliolatum sp. n. San Miguel to San Barbara, California Coulter s.n. (SL). Annotation 1: Type material Trifolium ciliolatum Benth. by G. Wal- BM: [no name], California (PL). Annotation 1: Lupinus truncata Nutt. lace in 2017. Written on sheet: T. ciliolatum. TCD! BM 001042527. Trifolium depauperatum Desv. var. amplectens (Torr. & A. Gray) Mc- Lupinus variicolor Steud. Dermott Coulter 393. Coulter 416. TCD: Lupinus versicolor var., Monterey to El Sur, California (SL). TCD! TCD: Trifolium amplectens Hook. & Arn. var., S. Miguel to S. Barbara, UC: Lupinus variicolor, California [no image]. UC 00170312. California (SL). Written on sheet: Trifolium amplectens. TCD! [Coulter] 393x. BRU: Trifolium amplectens, California (PL). Annotation 1: Herbarium TCD: [no name], Monterey, [California] (SL). Written on sheet: L. ver- Olneyanum Trifolium amplectens T. & G. PBRU 00035608. sicolor. TCD! K: [no name], California (SL). Annotation 1: Trifolium stenophyllum TCD: Lupinus versicolor Lindl. (SL). TCD! [On sheet with Coulter 393x Nutt. by P.B.K. in 1911. Annotation 2: vidi by Vincent in 1996. K (no name) above] 001051260. †Medicago polymorpha L. Coulter 416x. K: [no name], California (SL). Annotation 1: Trifolium depauperatum Coulter 403. Desv. by P.B.K. in 1911. K 001051264. TCD: [no name], San Miguel to San Barbara, California (SL). Annotation Coulter 418. 1: Medicago denticulata Willd. TCD! TCD: Trifolium amplectens, California (SL). Written on sheet: T. am- †Melilotus sp. plectens. TCD! Coulter 404 & 405. K: [no name], California (SL). Annotation 1: Trifolium amplectens T. & TCD: Melilotus parviflora Desv., California (SL). TCD! G. by P.B.K. in 1911. K 001051259. Note: on sheet with Coulter 416 and 416x above. *Mimosa polyantha Benth. K: Trifolium amplectens Hook. & Arn., California (PL). Annotation Coulter s.n. 1: [plant] (1) T. amplectens (2) T. hydrophilum Greene by E.L.G. [ = TCD: Mimosa polyantha Benth. in Hook. Journ. 4, p. 410. n. spec. valde Greene]. Annotation 2: (2) not T. hydrophyllum Greene, = T. stenophyl- affinis (SL). TCD! lum. K 001051252. VOLUME 37(1–2) Plantae Coulterianae 45

[Coulter s.n.] BRU: Trifolium tridentatum, California (PL). Annotation 1: Herbarium TCD: [no name], Monterey to San Miguel, California (SL). Written Olneyanum Trifolium tridentatum Lindl. PBRU 00035874. Note: BRU on sheet: Trifolium amplectens. TCD! Note: on sheet with Coulter 416 database lists as Coulter 108 in error. above. Trifolium wormskioldii Lehm. Trifolium fucatum Lindl. Coulter 412. K T. wormskioldii Coulter 406. : [no name], California (SL). Annotation 1: Lehm. by = T. wormskioldii TCD: Trifolium fucatum, San Miguel to S. Barbara (SL). Written on P.B.K. [ Kennedy] in 1911. Annotation 2: Lehm. by Vincent in 1993. Annotation 3: T. wormskioldii Lehm. by Gillett in 1963. sheet: Trifolium fucatum. TCD! K 001051316. TCD: T[rifolium], San Miguel to San Barbara, California (SL). Written Coulter s.n. on sheet: Trifolium fucatum. TCD! Note: on sheet with Coulter 406 above. TCD: Trifolium fimbriata Lindl. forma latifolia, California (SL). Written TCD: [no name], Monterey to San Gabriel, California (SL). Written on on sheet: T. fimbriatum. TCD! sheet: Trifolium fucatum. TCD! Note: on sheet with two other Coulter TCD: Trifolium fimbriata Lindl., California (SL). Written on sheet: T. 406 above. fimbriatum. TCD! Note: on sheet with Coulter s.n., Trifolium fimbriata K: Trifolium fucatum Lindl., Monterey, S. Gabriel, S. Miguel, S. Barbara, forma latifolia above. California (PL). Written on sheet: T. fucatum. K 001051248. Coulter s.n. Vicia gigantea Hook. TCD: Trifolium fucatum Lindl., Monterey, California (SL). Written on Coulter 443. sheet: T. fucatum Lindl. TCD! K: [no name], California (SL). K 001051443. BRU: Trifolium fucatum, California (PL). Annotation 1: Herbarium Ol- K: Vicia gigantea Hook., Monterey, California (PL). K 001051445. neyanum Trifolium fucatum Lindl. PBRU 00035686. Coulter s.n. NY: Trifolium fucatum Lindl., California (PL). Annotation 1: Trifolium TCD: Vicia gigantea Hooker, California (SL). TCD! fucatum in 1942. NY 00033715. Note: attributed to J.M. Coulter in NY database in error. Vicia ludoviciana Torr. & A. Gray subsp. ludoviciana UC: Trifolium fucatum, California [no image]. UC 00973628. Coulter 447. TCD: Vicia exigua Nutt., Monterey, California (SL). Written on sheet: Trifolium gracilentum Torr. & A. Gray exigua. TCD! Coulter 415. TCD: Trifolium gracilentum Torr. & Gr., Monterey to S. Miguel, Cali- †Vicia sativa L. subsp. fornia (SL). Written on sheet: gracilentum T. & G. TCD! Coulter 445. K: [no name], California (SL). Written on sheet: T. gracilentum T. & G. TCD: Vicia sativa L., S. Miguel to S. Barbara, California (SL). TCD! K 001051231. Vicia sp. Coulter s.n. Coulter s.n. TCD: Trifolium gracilentum var. major, California (SL). Note on label TCD: Vicia sparsifolia Nutt., California (SL). Note on label: not suffi- (was mixed with 414). Written on sheet: gracilentum. TCD! cient to ascertain with certainty. TCD! Trifolium microcephalum Pursh Coulter 411 & 419. TCD: Trifolium microcephalum Pursh, California (SL). Written on sheet: fagaceae microcephalum. TCD! Notholithocarpus densiflorus (Hook. & Arn.) P.S. Manos, C.H. Cannon Trifolium obtusiflorum Hook. & S.H. Oh Coulter 410. Coulter 663. K: T. roscidum Greene, California (SL). Annotation 1: T. obtusiflorum TCD: Quercus densiflora Hook., California (PL). TCD! [two sheets] by P.B.K. in 1911. Annotation 2: Trifolium tridentatum by Gillett in K: Q. densiflora Hook. & Arn., California (SL). Written on sheet: DC. 1963. Annotation 3: Trifolium obtusiflorum Hook. by Vincent in 1995. Prod. xvi 82. K 000512505. K 001051311. Quercus agrifolia Née Coulter 662. Trifolium variegatum Nutt. var. TCD: Quercus agrifolia, California (PL). TCD! Note: may be mixed Coulter 413. sheet. TCD: Trifolium variegatum, S. Miguel to S. Barbara, California (SL). Coulter s.n. Annotation 1: Trifolium variegatum this was mixed with T. micro- TCD: [no name], California (PL). TCD! cephalum. Written on sheet: Tr. variegatum. TCD! K: Trifolium variegatum Nutt., S. Miguel to S. Barbara, California (PL). Quercus berberidifolia Liebm. Annotation 1: T. trilobatum by P.B.K. in 1911. K 001051288. Coulter 661. TCD: Quercus agrifolia, California (PL). Annotation 1: Type material Trifolium willdenovii Spreng. Quercus berberidifolia Liebmann by G. Wallace in 2017. TCD! Coulter 407. GH: Quercus agrifolia, California [no image]. GH 00344520. K: [no name], California (SL). Annotation 1: Trifolium tridentatum by K: Qu. agrifolia Nee, California (SL). Annotation 1: Quercus berberid- Gillett in 1963. K 001051300. ifolia. Annotation 2: Quercus dumosa HOLOTYPE of Q. berberidifolia Coulter 407 & 409. by Muller in 1958. Written on sheet: DC. Prod. xvi 36. K 000832346. TCD: Trifolium tridentatum, California (SL) [single label for both num- MO: Quercus berberidifolia Lieb. Script on fragment folder. Annotation bers]. TCD! 1: Ex Herb. A. Gray 1876. Annotation 2: Isotype Quercus berberidifo- Coulter 408. lia Liebm. by KCN I 1983. Annotation 3: Type Specimen Herb. M.B.G. TCD: Trifolium tridentatum Lindl., Monterey to San Miguel, California MO 03461523. Note: identified as Q. beccariana and attributed to J.M. (SL). TCD! Coulter as collector in the MO database in error. 46 Wallace ALISO

Quercus dumosa Nutt. GH: Erythraea venusta, California (PL). Annotation 1: Centaurium caly- Coulter s.n. cosum (Buckl.) Fernald (TYPE of Erythraea venusta A. Gray by Martin in GH: Quercus dumosa, California [no image]. GH 00344519. 1939. Annotation 2: Centaurium venustum (Gray) Robinson Coulter 561 is the TYPE of the basionym by Dunn in 1970. Written on sheet: Centau- Quercus engelmannii Greene rium venustum (Gray) Robinson. GH 00072252. Note: in GH database Coulter 659. as SYNTYPE. TCD: Quercus agrifolia?, California (PL). TCD! K: [no name], California (SL). Annotation 1: Centaurium venustum (Gray) B.L. Robinson by Broome in 1979. Annotation 2: Ex num. Coul- Quercus lobata Née ter 561. Two plants on right: ISOTYPE of Erythraea venusta A. Gray Coulter 660. now = Centaurium venustum (A. Gray) B.L. Rob. by Pringle in 1998. K TCD Quercus douglasii TCD : , California (PL). ! 000438600. Note: attributed to Mexico in K database in error. GH: Quercus lobata, California [no image]. GH 00343132. Coulter 562. Quercus sp. TCD: [no name], California (PL). Annotation 1: Erythraea venusta Harv. TCD Coulter s.n. & Gr., Torr. & Gray, Fl. N. Amer. ! GH Erythraea venusta TCD: [no name], California (PL). TCD! : , California (PL). Annotation 1: Syn. F. N. Amer. Written on sheet: Flowers larger on pl. Trin. Coll. [Note: on sheet with Coulter 561 but no separate GH number; in database as GH 00415266 frankeniaceae and annotated as: Zeltnera venusta by Kittredge in 2013 but annotations not evident on specimen. Frankenia salina (Molina) I.M. Johnst. K: [no name], California (SL). Annotation 1: Centaurium venustum Rob. Coulter 150. By W.L.J. [ = Jepson]. K 000438601. Note: attributed to Mexico in K TCD: Frankenia grandifolia Cham., California (PL). TCD! database in error. BRU: Frankenia grandifolia, California (PL). Annotation 1: Herbarium Olneyanum Frankenia grandifolia C. & Schlect. PBRU 00041904. Note: in BRU database as Coulter 157 in error. geraniaceae GH: Frankenia salina, California [no image]. Annotation 1: Frankenia salina by Welsh in 1990. GH 00376189. †Erodium cicutarium (L.) Aiton Coulter s.n. TCD: Erodium cicutarium, San Miguel to S. Barbara, California (SL). garryaceae TCD! Garrya elliptica Lindl. Coulter 647. TCD: Garrya elliptica, California (PL). TCD! Note: two sheets, each grossulariaceae with male and female plant specimens. Ribes aureum Pursh var. GH: Garrya elliptica, California [no image]. GH 00401725. Coulter s.n. US: Garrya elliptica, California [no image]. US 40349. TCD: Ribes tenuiflorum Lindl., California (PL). TCD! [two sheets] Coulter s.n. TCD: Ribes tenuiflorum Lindl., S. Miguel, California (PL). TCD! BRU: Garrya elliptica, California (PL). Annotation 1: Herbarium Ol- BRU: Ribes tenuiflorum Lindl., California (PL). Annotation 1: Herbar- neyanum. PBRU 00041894. ium Olneyanum Ribes tenuiflorum Lindl. PBRU 00030912. L: Garrya elliptica, California [no image]. Annotation 1: Garrya elliptica GH: Ribes tenuiflorum, California (PL). GH 363959. by Wangerin in 1909. L 1557438. GH: [no name], California (PL). GH 363960. UC: Garrya elliptica, California [no image]. UC 973612. Coulter s.n. UC: Ribes aureum, California [no image]. UC 973799. gentianaceae Ribes californicum Hook. & Arn. var. Frasera parryi Torr. Coulter 228x. Coulter 558. K: R. occidentale California (Douglas), intermediate with R. occidentale TCD: Frazera [sic] n. sp., California (PL). Annotation 1: Frasera selen- (SL). K 000762746. odes Harv. & Gr., Torr. & Gray, Fl. N. Amer. TCD! Coulter 233. GH: Frasera n. sp. Glands peculiar, California (PL). Annotated: BRU: Ribes occidentale Hook. & Arn., California (PL). Annotation ISOSYNTYPE Frasera parryi Torrey Rep. U.S. Mex. Bound., Bot. 156. 1: Herbarium Olneyanum Ribes occidentale Hook. & Arn. PBRU 1859 by Kittredge in 2008. GH 00267289. 00030876. K: Frasera hippocrepis, California (SL). Written on sheet: Frasera parryi GH: Ribes californicum, California [no image]. GH 00380021. Torr. in Bot. Whipp. Exp. p. 126, Mex. Bound. Surv. p. 156. K 000854336. K: R[ibes] occidentale, “California (Douglas) Dr. Coulter” (SL). K Coulter s.n. 000762745. TCD: Frasera, California (PL). Annotation 1: Frasera selenodes Harv. Coulter s.n. & Gr., Torr. & Gray, Fl. N. Amer. TCD! TCD: Ribes occidentale, California (PL). TCD!

Zeltnera venusta (A. Gray) G. Mans. Ribes divaricatum Douglas var. Coulter 560. Coulter s.n. TCD: Erithrea [sic] trichantha?, California (PL). Written on sheet: same TCD: Ribes divaricatum, Monterey, California (SL). TCD! as 562, 561 E. venusta. TCD! TCD: Ribes divaricatum, California (PL). TCD! Coulter 561. TCD: Ribes villosum Nutt., San Miguel to San Barbara, California. Note TCD: Erythraea, California (PL). Annotation 1: E. venusta n. sp. Harv. on label: spines often but one, sometimes 3, the lateral small (SL). TCD! & Gr., Torr. & Gray, Fl. N. Amer. Annotation 2: Type material Erythraea TCD: Ribes villosum Nutt., California (PL). TCD! venusta Harv. & A. Gray ex A. Gray by G. Wallace in 2016. Written on TCD: Ribes villosum Nutt., California (PL). Written on sheet: Ribes di- sheet: E. venusta T. & G. TCD! varicatum. TCD! VOLUME 37(1–2) Plantae Coulterianae 47

BRU: Ribes divaricatum, California [no image]. Annotation 1: Herbar- krameriaceae ium Olneyanum Ribes divaricatum. PBRU 00030667. Krameria erecta Willd. in Schult. & Schult. f. Ribes divaricatum Douglas var. pubiflorum Koehne Coulter 71*. Coulter s.n. K: [no name] California (SL). K 000471038. Note: Gray (1852, p. 42) GH: Ribes divaricatum var. pubiflorum, California [no image]. Annota- mentions having seen this specimen along with an original specimen of tion 1: Ribes divaricatum var. pubiflorum by Sinnott in 1982. GH 380167. Bentham’s Krameria parvifolia. The Coulter 71*. The specimen was dif- GH: Ribes divaricatum var. pubiflorum, California [no image]. Annota- ficult to locate because it did not appear in the K database and shares the tion 1: Ribes divaricatum var. pubiflorum by Sinnott in 1982. GH 380174. same accession number as the co-mounted Barclay specimen from Mag- GH: Ribes divaricatum var. pubiflorum, California [no image]. Annota- dalena Bay, Mexico. Based on the text in Gray (1852, p. 42) and Bentham tion 1: Ribes divaricatum var. pubiflorum by Sinnott in 1982. GH 380176. (1844, p. 6) the Barclay specimen is likely an isotype of Krameria parv- UC: Ribes divaricatum var. pubiflorum, California [no image]. UC ifolia Benth. 973798.

Ribes malvaceum Sm. var. lamiaceae Coulter 231. TCD: Ribes malvaceum, California (PL). TCD! Lepechinia calycina (Benth.) Munz Coulter s.n. Coulter 525. BRU: Ribes malvaceum, California (PL). Annotation 1: Herbarium Ol- TCD: Sphacele calycina, California (PL). TCD! neyanum Ribes malvaceum Smith. PBRU 00030868. GH: Lepechinia calycina, California [no image]. Annotation 1: Lepe- GH: Ribes malvaceum, California [no image]. GH 0038096. chinia calycina by Hart in 1984. GH 00400232. UC: Ribes malvaceum, California [no image]. UC 973608. Coulter s.n. BM: Sphacele calycina, California (PL). BM 001025195. Ribes menziesii Pursh var. menziesii Coulter 236. †Mentha pulegium L. TCD: Ribes subvestitum, California (PL). Written on sheet: Ribes men- Coulter 527. ziesii. TCD! TCD: Mentha pulegium?, California (PL). TCD! Coulter s.n. TCD: Ribes menziesii Pursh, San Miguel to San Barbara, California (SL). Monardella breweri A. Gray subsp. TCD! Coulter 542. GH: Ribes menziesii var. leptosum, California [no image]. Annotation 1: K: Monardella undulata, California (SL). Annotation 1: Monardella Ribes subvestitum Hook. & Arn. GH 00380573. lanceolata Gray by Epling in 1927. K 000248491.

Ribes sanguineum Pursh var. glutinosum (Benth.) Loudon Monardella sp. Coulter 230. Coulter 541. TCD: Ribes glutinosum, California (PL). TCD! [three sheets] TCD: Monardella, California (PL). TCD! Coulter s.n. Monardella undulata Benth. subsp. crispa (Elmer) Elvin & Sanders TCD: Ribes glutinosum, California (PL). TCD! Coulter 536. BRU: Ribes glutinosum, California [no image]. Annotation 1: Herbarium TCD: Monardella, S. Miguel to S. Barbara, California (PL). TCD! Olneyanum Ribes glutinosum Benth. GH: Monardella undulata subsp. crispa, California [no image]. Anno- GH: Ribes sanguineum var. glutinosum, California [no image]. Anno- tation 1: Monardella undulata subsp. crispa by Sanders and Elvin 2009. tation 1: Ribes sanguineum var. glutinosum by Reuter in 1950. GH GH 400655. 00380678. Coulter 542. UC: Ribes sanguineum var. glutinosum, California [no image]. UC TCD: Monardella undulata Benth., California (PL). TCD! 973607. Salvia apiana Jeps. Ribes speciosum Pursh Coulter 532. Coulter 227. TCD: Audibertia polystachya, California (PL). TCD! TCD: Ribes speciosum, California (PL). TCD! GH: Salvia apiana, California [no image]. GH 00403231. Coulter s.n. BRU: Ribes speciosum, California (PL). Annotation 1: Herbarium Ol- Salvia carduacea Benth. neyanum Ribes speciosum Pursh. PBRU 00030909. Coulter 539. UC: Ribes speciosum, California [no image]. UC 973609. GH: Salvia gossypina, California [no image]. Annotation 1: Salvia car- duacea by Epling in 1936. GH 00403278. Ribes sp. Coulter s.n. Coulter 229. TCD: Salvia carduacea Benth., S. Miguel to S. Barbara, California (PL). TCD: Ribes brevipes MS n. sp., Monterey to San Miguel, California (SL). TCD! TCD! Coulter s.n. Salvia columbariae Benth. TCD: [no name], California (PL). TCD! Coulter 537. TCD: Salvia columbariae, S. Miguel, California (PL). TCD! Coulter 538. TCD: Salvia columbariae, S. Francisco, California (PL). TCD! juglandaceae BRU: Salvia columbariae, California (PL). Annotation 1: Herbarium Ol- neyanum. PBRU 00044873. Juglans californica S. Watson GH: Salvia columbariae, California [no image]. GH 00403293. Coulter 131. Coulter s.n. TCD: Rhus, California (PL). TCD! TCD: Salvia columbariae, California (PL). TCD! 48 Wallace ALISO

Salvia leucophylla Greene lythraceae Coulter 540. Lythrum californicum Torr. & A. Gray TCD: Audibertia nivea Benth., California (PL). TCD! Coulter 149. GH: Salvia leucophylla, California [no image]. Annotation 1: Salvia leu- TCD: Lythrum californicum, Between Monterey & S[a]n Gabriel, Cali- cophylla by Epling. GH 403371. fornia (SL). TCD! K: Audibertia nivea Benth. (SL). K 000929936. malvaceae Salvia mellifera Greene Abutilon sp. Coulter 535. Coulter 105. TCD: Audibertia stachyoides, California (PL). TCD! TCD: Abutilon, California (SL). TCD! BRU: Audibertia stachyoides, California (PL). Annotation 1: Herbarium Olneyanum Audibertia stachyoides Benth. PBRU 00044945. †Anoda cristata (L.) Schltdl. GH: Salvia mellifera, California [no image]. GH 00403454. Coulter 98. TCD: Anoda cristata?, Sida sp. nov.?, San Gabriel, California (SL). Salvia spathacea Greene TCD! Coulter 526. TCD: Audibertia grandiflora, California (PL). TCD! Malacothamnus fasciculatus (Torr. & A. Gray) Greene var. GH: Salvia spathacea, California [no image]. GH 00403510. Coulter 671. K: Malva fasciculata Nutt., California (SL). Annotation 1: Malacotham- Scutellaria tuberosa Benth. nus fasciculatus (Nutt. ex T. & G.) Greene ssp. jonesii (Munz) Bates nov. Coulter 524. comb. by Bates in 1963. K 000569253. TCD: Scutellaria tuberosa, California (PL). TCD! Malva assurgentiflora (Kellogg) M.F. Ray BRU: Scutellaria tuberosa, California (PL). Annotation 1: Herbarium Ol- Coulter s.n. neyanum Scutellaria tuberosa Benth. PBRU 00045659. TCD: Lavatera, California (SL). TCD! K: Scutellaria tuberosa Benth., Monterey to El Sur, California (SL). TCD: [no name] no label. Written on sheet: Herb. Coulter; Lavatera. Annotation 1: Scutellaria tuberosa Benth. by R. Olmstead in 1987. K Annotation 1: probably Lavatera assurgentiflora, California by Webb in 000906381. 1988. TCD! Coulter s.n. BM: Scutellaria tuberosa Benth., California (PL). BM 001025217. †Malva parviflora L. Coulter 108. Stachys bullata Benth. TCD: Malva obtusa Tor. & Gr., Monterey (SL). TCD! Coulter 531. TCD: Stachys bullata, California (PL). TCD! †Malva sp. GH: Stachys bullata, California [no image]. Annotation 1: Stachys bul- Coulter 97. lata by Mulligan in 1987. GH 00402193. TCD: Malva debilis Harv., Monterey, California (SL). TCD! Coulter 101. Stachys chamissonis Benth. TCD: Malva new?, California (PL). TCD! Coulter 533. Coulter 104. TCD: Stachys chamissonis Benth., California (PL). TCD! TCD: Malva, Monterey, California (SL). TCD!

Trichostema lanceolatum Benth. Malvella leprosa (Ortega) Krapov. Coulter 528. Coulter 109. TCD: Trichostema lanceolatum, California (PL). TCD! TCD: Malva hederacea Dougl., Sida? obliqua Nutt. Tor. & Gr. 1: 233, GH: Trichostema lanceolatum, California [no image]. Annotation 1: Tri- Monterey, California (SL). TCD! chostema lanceolatum by Lewis in 1941. GH 00402432. K: [no name], California (SL). Written on sheet: M. californica Presl., Malva hederacea Dougl. MS in Hook. Fl. Bor. Amer. 1, p. 107, Sida hed- eracea Torr. & Gray. K 000659579. Note: in K database as Coulter 169 in error. loasaceae Coulter 671. TCD: Sida hederacea?, California (PL). TCD! Mentzelia albicaulis (Hook.) Torr. & A. Gray Coulter s.n. Coulter 174. GH: Malvella leprosa, California [no image]. Annotation 1: Sida leprosa TCD: B[artonia] albicaulis?, California (PL). TCD! var. hederacea by Clement in 1948. GH 00385052. BM: Mentzelia, California [no image]. BM 000929395. K: [no name] (SL). Written on sheet: M. californica Presl., Malva heder- acea Dougl. MS in Hook. Fl. Bor. Amer. 1, p. 107, Sida hederacea Torr. Mentzelia gracilenta Torr. & A. Gray &Gray.K 000659582. Coulter 165. TCD: Mentzelia gracile?, California (PL). TCD! Sidalcea malachroides (Hook. & Arn.) A. Gray Coulter 171. Coulter 107. TCD: Bartonia gracile?, Mentzelia, S[an] Miguel, California (PL). TCD! TCD: Malva malachroides,My[= Monterey] to El Sur, California (PL). TCD! Mentzelia sp. Coulter 163. Sidalcea malviflora (DC.) A. Gray subsp. malviflora TCD: Mentzelia, California (PL). TCD! Coulter 102. Coulter s.n. TCD: Sida malvaflora?orS. oregana, California (PL). Written on sheet: TCD: Mentzelia, California (PL). TCD! Sidalcea. TCD! TCD: Mentzelia, California (PL). TCD! Coulter 103. VOLUME 37(1–2) Plantae Coulterianae 49

TCD: S. malvaflora? Sida n. sp.? California (SL). Written on sheet: Sidal- Coulter s.n. cea malviflora. TCD! TCD: Boerhavia, California (PL). TCD!

Sphaeralcea coulteri (S. Watson) A. Gray Mirabilis laevis (Benth.) Curran var. crassifolia (Choisy) Spellenb. Coulter 96. Coulter 600. TCD: M[alvastrum] grossularifolia, Monterey, California (SL). Annota- GH: Mirabilis laevis var. crassifolia, California [no image]. Annotation tion 1: Type material Malvastrum coulteri S. Watson by G. Wallace Nov. 1: Mirabilis laevis var. crassifolia by Spellenberg in 1996. GH 00589078. 2017. Written on sheet: Malvastrum grossularifolia. TCD! K: [no name], California (SL). K 000779331. GH: Malvastrum munroana, California (PL). Annotation 1: Sphaeralcea coulteri Gray. Syn. Fl. N. Amer. Annotation 2: HOLOTYPE Malvastrum Mirabilis multiflora (Torr.) A. Gray var. coulteri by Kittredge in 2007. GH 00260913. Coulter 559. TCD: Lysianthus?, California (PL). Written on sheet: Mirabilis, Quamo- Sphaeralcea munroana (Lindl.) A. Gray clidion multiflorum Torr. TCD! Coulter 106. K: Malva munroana Dougl., California (SL). K 000582080. Sphaeralcea sp. oleaceae Coulter 94. TCD: Sphaeralcea n. sp.?, California (PL). TCD! Fraxinus dipetala Hook. & Arn. Coulter 95. Coulter 563. TCD: [no name], California (PL). TCD! TCD: Fraxinus dipetala, California (PL). TCD! GH: Fraxinus dipetala, California [no image]. GH 00401140. montiaceae Calandrinia ciliata (Ruiz. & Pav.) DC. onagraceae Coulter 87. Camissonia contorta (Douglas) Kearney TCD: Calandrinia menziesii, California (SL). TCD! Coulter 172. Coulter 88. TCD: Oe. contorta or dentata, California (SL). TCD! TCD: Calandrinia menziesii Hook., California (SL). TCD! Coulter s.n. Camissonia sp. UC: Calandrinia menziesii, California [no image]. UC 973613. Coulter 180. Claytonia perfoliata Willd. subsp. perfoliata TCD: Oenothera n. sp., Monterey, California (SL). TCD! Coulter 86. Camissonia strigulosa (Fisch. & C.A. Mey.) P.H. Raven TCD: Claytonia perfoliata, Monterey to San Francisco (SL). TCD! Coulter 160. TCD: Oe. dentata, California (SL). TCD! Note: Coulter 172 on same myrsinaceae label. †Anagallis arvensis L. Coulter 172. Coulter 564. TCD: Oe. dentata, California (SL). TCD! Note: Coulter 160 on same TCD: Anagallis arvensis, California (PL). TCD! label. BRU: Anagallis arvensis, California (PL). Annotation 1: Herbarium Ol- Camissoniopsis bistorta (Torr. & A. Gray) W.L. Wagner & Hoch neyanum. PBRU 00041039. Coulter 158. Coulter s.n. TCD: [no name], California (SL). TCD! UC: Anagallis arvensis, California. [no image]. UC 973811. K: [no name], California (SL). K 000742400.

nyctaginaceae Camissoniopsis cheiranthifolia (Spreng.) W.L. Wagner & Hoch subsp. suffruticosa Abronia sp. (S. Watson) W.L. Wagner & Hoch Coulter 581. Coulter 152. K: Oe. viridescens Hook. Bor. Am., California (SL). Annotation 1: Oe. TCD: Abronia, S. Miguel to S. Barbara, California (PL). TCD! cheiranthifolia var. typica by Munz in 1931. K 000742406. Coulter s.n. Coulter 153. TCD: [no name], California [no label]. Written on sheet: Abronia glauca. GH: Camissonia cheiranthifolia, California [no image]. Annotation 1: TCD! Camissonia cheiranthifolia by Raven in 1965. GH 00367577. Abronia umbellata Lam. var. K: [no name], California (SL). Written on sheet: Oenothera viridescens Coulter 574. Hook. & Arn. K 000742404. TCD: Abronia, California (PL). TCD! Coulter 164. Coulter 578. TCD: Oe. viridescens, California (SL). TCD! TCD: Abronia, California (PL). TCD! BRU: Abronia, California (PL). Annotation 1: Herbarium Olneyanum. Camissoniopsis sp. PBRU 00041452. Coulter 152. NY: Abronia umbellata, California [no image]. Annotation 1: Abronia TCD: Oenothera, California (SL). TCD! umbellata by Tillett in 1960. NY 0679443. Coulter 168. TCD: [no name], California (SL). TCD! Boerhavia sp. K: Oe. lepida, California (SL). K 000712412. Coulter 699. TCD: [no name], California (PL). TCD! Note: in Boerhavia folder. Clarkia epilobioides (Torr. & A. Gray) A. Nelson & J.F. Macbr. Coulter 701. Coulter 157. TCD: Boerhavia, California (PL). TCD! TCD: Oenothera (Godetia), California (SL). TCD! 50 Wallace ALISO

K: Oe. epilobioides, California (SL). K 000742235. K: [no name], California (SL). K 000195546. Note: identified in K database as Castilleja purpurea in error and filed as that species because Clarkia purpurea (Curtis) A. Nelson & J.F. Macbr. subsp. mounted on sheet with type material of that species. Coulter s.n. TCD: Oenothera purpurea, California (PL). TCD! Castilleja affinis Hook. & Arn. subsp. affinis Coulter 610. Clarkia sp. E: Castilleja affinis subsp. affinis, California [no image]. Annotation 1: Coulter 154. Castilleja affinis subsp. affinis by M. Egger in 2008. E 399121. TCD: Oenothera (Godetia), California (SL). TCD! Coulter 622. Coulter 159. E: Castilleja affinis subsp. affinis, Monterey to El Sur, California [no im- TCD: Clarkia, California (SL). TCD! age]. Annotation 1: Castilleja affinis subsp. affinis by M. Egger in 2008. K: Oe[nothera] pallida Nutt. in Herb. Hook. Oe./Godetia, California E 272499. (SL). Written on sheet: This Calif. pl. is a Godetia. K 000742446. Coulter 161. Castilleja densiflora Benth. subsp. densiflora TCD: Godetia, Calif (SL). TCD! Coulter 609. TCD: Orthocarpus densiflorus, California (PL). Annotation 1: O. densi- Epilobium canum (Greene) P.H. Raven subsp. canum florus Torr. & Gray, Fl. N. Amer. TCD! Coulter 181. BRU: [no name], California (PL). PBRU 00045068. TCD: Zauschneria californica, California (PL). TCD! E: Castilleja densiflora subsp. densiflora, California [no image]. Anno- GH: Epilobium canum, California [no image]. Annotation 1: Epilobium tation 1: Castilleja densiflora subsp. densiflora by M. Egger in 2008. E canum by Hoch in 1995. GH 00367742. 272672.

Eremothera boothii (Douglas) W. Wagner & Hoch subsp. decorticans Castilleja exserta (A. Heller) T.I. Chuang & Heckard subsp. (Hook. & Arn.) W. L. Wagner & Hoch Coulter 608. Coulter 151. TCD: Orthocarpus purpurascens, California (PL). TCD! TCD: Oenothera gauraeflora, California (SL). TCD! TCD: Orthocarpus purpurascens, California (PL). Annotation 1: O. pur- Coulter s.n. purascens Torr. & Gray, Fl. N. Amer. TCD! TCD: Oenothera boothii, California (SL). TCD! BRU: [no name], California (PL). Annotation 1: Herbarium Olneyanum. PBRU 00045067. Eulobus californicus Torr. & A. Gray Coulter s.n. Castilleja exserta (A. Heller) T.I. Chuang & Heckard subsp. exserta TCD: Eulobus californicus Nutt., Monterey, California (SL). TCD! Coulter 609. BRU: [no name], California (PL). Annotation 1: Herbarium Olneyanum. Ludwigia sp. PBRU 00045068. Coulter 175. E: Castilleja exserta subsp. exserta, California [no image]. Annotation 1: TCD: Justicia, Monterey (SL). TCD! Castilleja exserta var. exserta by M. Egger in 2008. E 00335383. Coulter s.n. Oenothera elata Kunth subsp. hookeri (Torr. & A. Gray) W. Dietr. & E: Castilleja exserta subsp. exserta, Monterey to San Francisco, Califor- W.L. Wagner nia [no image]. E 00272673. Coulter 155. TCD: Oenothera hookeri Tor. & Gr., California (SL). TCD! Castilleja foliolosa Hook. & Arn. K: Oe. biennis, California (SL). K 000742437. Coulter 617. TCD: Castilleja foliosa [sic], California (PL). Annotation 1: C. foliosa Oenothera sp. Torr. & Gray, Fl. N. Amer. TCD! Coulter 156. BRU: [no name], California (PL). Annotation 1: Herbarium Olneyanum. TCD: [no name], California (SL). TCD! PBRU 00045062. Coulter 166. E: Castilleja foliolosa, California [no image]. Annotation 1: Castilleja TCD: Oenothera, California (PL). TCD! foliolosa by M. Egger. E 00272302. K: Gaura parviflora Douglas, California (SL). K 000742339. Coulter 627. E: Castilleja foliolosa, California [no image]. E 272302. Taraxia ovata (Torr. & A. Gray) Small Coulter 179. Castilleja latifolia Hook. & Arn. TCD: Oenothera ovata, California (PL). TCD! Coulter 618. TCD: Castl. latifolia, California (PL). TCD! Tetrapteron graciliflorum (Hook. & Arn.) W.L. Wagner & Hoch K: [no name], California (SL). K 000195501. Coulter 173. Coulter 620. TCD: Oenothera graciliflora, S. Francisco, California (PL). TCD! TCD: Castilleja latifolia, California (PL). Annotation 1: C. latifolia Torr. TCD: [no name], S. Miguel, California (PL). TCD! &Gray,Fl.N.Amer.TCD! K: Oe. graciliflora Hook., California (SL). K 000742418. K: [no name], Monterey to San Gabriel (SL). K 000195500. Coulter s.n. NY: Oe. graciliflora Hook., California [no image]. Annotation 1: Castilleja miniata Hook. subsp. miniata Camissonia graciliflora (Hook. & Arn.) Raven by Raven in 1964. NY Coulter 621. 00233552. K: [no name], California (SL). Written on sheet: Cast. pallida var. mini- orobanchaceae ata. K 000195527. Castilleja affinis Hook. & Arn. subsp. Castilleja sp. Coulter 613. Coulter 610. TCD: Castl. affinis Benth., California (PL). Annotation 1: C. affinis TCD: Castilleja douglasii, California (PL). Annotation 1: C. douglasii Benth. Torr. & Gray, Fl. N. Amer. TCD! Torr. & Gray, Fl. N. Amer. TCD! VOLUME 37(1–2) Plantae Coulterianae 51

K: [no name], California (SL). K 000195510. [Filed as Castilleja angus- paeoniaceae tifolia] Paeonia californica Nutt. Coulter 610x. Coulter 17. K: [no name], California (SL). Written on sheet: parvifolia Bong. K TCD: [no name], Monterey to San Miguel, California (SL). Written on 000195511. sheet: Paeonia brownii and P. californica. TCD! Coulter 615. Coulter s.n. TCD: Castilleja hispida, California (PL). Annotation 1: C. hispida Torr. TCD: Paeonia californica Monterey to San Francisco, Californica (SL). &Gray,Fl.N.Amer.TCD! Written on sheet: Paeonia brownii and P. californica. TCD! Note: on K: [no name], (SL). K 000195547. Note: identified in K database as sheet with Coulter 17 above. Castilleja purpurea in error and filed as that species because mounted on TCD: [no name], Monterey to San Miguel, California (SL). Written on sheet with type material of that species; on sheet with Coulter 613 above. sheet: Paeonia brownii Dougl., P. californica Nutt. TCD! Coulter 619. TCD: Paeonia, near Monterey, California. 1832 (SL). Written on sheet: TCD: Castilleja hispida, California (PL). Annotation 1: C. hispida Torr. P. brownii. TCD! &Gray,Fl.N.Amer.TCD! BRU: Paeonia californica, California (PL). Annotation 1: Herbarium Ol- Coulter 621. neyanum Paeonia californica. PBRU 00031011. TCD: Castl. hispida, California (PL). Annotation 1: C. hispida Torr. & UC: Paeonia californica, California [no image]. UC 973816. Gray, Fl. N. Amer. TCD! Coulter 622. TCD: Castl. hispida, El Sur, California (PL). Annotation 1: C. hispida papaveraceae Torr. & Gray, Fl. N. Amer. TCD! Argemone munita Durand & Hilg. Coulter 18. Orobanche cooperi (A. Gray) A. Heller TCD: Argemone mexicana, Monterey, California, 1832 (SL). TCD! Coulter 646. Coulter s.n. TCD: Orobanche ludoviciana, California (PL). TCD! TCD: [no name], Monterey (SL). TCD!

Pedicularis densiflora Hook. Dendromecon rigida Benth. Coulter 605. Coulter 23. TCD: Pedicularis densiflora, California (PL). TCD! BRU: Dendromecon rigidum, California (PL). Annotation 1: Herbarium TCD: Pedicularis densiflora, California (PL). Annotation 1: P.densiflora Olneyanum Dendromecon rigidum Benth. PBRU 00029201. Torr. & Gray, Fl. N. Amer. TCD! Coulter s.n. Coulter 606. TCD: [no name], between San Miguel & San Barbara, California, 1832 TCD: [no name], California (PL). Annotation 1: P. attenuata Torr. & (SL). TCD! Gray, Fl. N. Amer. Annotation 2: ISOTYPE Pedicularis attenuata Benth. TCD: [no name], Monterey California (SL). Written on sheet: Den- by G. Wallace in 2017. TCD! dromecon rigidum. TCD! TCD: [no name], California (PL). Annotation 1: P. attenuata Benth. TCD: [no name], Between Monterey & El Sur, California. 1832 (SL). Torr. & Gray, Fl. N. Amer. Annotation 2: ISOTYPE Pedicularis atten- TCD! uata Benth. by G. Wallace in 2017. TCD! E: Dendromecon rigidum, California [no image]. E 00060071. BRU: [no name], California (PL). Annotation 1: Herbarium Olneyanum. GH: Dendromecon rigida, California [no image]. Annotation 1: Den- PBRU 00043940. Note: BRU database lists as 101 in error. dromecon rigida by Klier in 1977. GH 381412. K: Pedicularis attenuata, California (SL). Annotation 1: TYPE SPECI- K: Dendromecon rigidum, California (PL). K 00010746. MEN. K 000741011. UC: Dendromecon rigida, California [no image]. UC 973617. Coulter s.n. TCD: Pedicularis attenuata, California (PL). TCD! Ehrendorferia chrysantha (Hook. & Arn.) Rylander NY: Pedicularis attenuata Benth., California (PL). Written on sheet: Coulter 52. ISOTYPE. NY 00090880. TCD: Dielytra chrysantha H. & A., Monterey, California (SL). Annota- UC: Pedicularis densiflora, California [no image]. UC 973627. tion 1: Ehrendorferia chrysantha by M. Liden 2009. TCD! Eschscholzia caespitosa Benth. Triphysaria eriantha (Benth.) T.I. Chuang & Heckard subsp. eriantha Coulter 24. Coulter 612. TCD: Ch[ryseis] caespitosa?, Monterey to San Francisco, California, TCD: Orthocarpus erianthus, California (PL). Annotation 1: O. eri- 1832 (SL). Written on sheet: C[hryseis] tenuifolia Benth. TCD! anthus Torr. & Gray, Fl. N. Amer. TCD! K: Ch[ryseis] caespitosa (SL). Annotation 1: Eschscholzia caespitosa BRU: Orthocarpus, California [no image]. PBRU 00045001. Benth. by Ernst 1964. K 000653328. E: Triphysaria eriantha var. eriantha, California [no image]. E 272870. Coulter 24*. Coulter s.n. TCD: Ch[ryseis] caespitosa Lindl., Monterey to San Miguel, California UC: Triphysaria eriantha subsp. eriantha, California [no image]. UC (SL). TCD! 973623. Coulter s.n. K: [no name], California [script on sheet]. Annotation 1: Eschscholzia tenuifolia Benth. [ = E. caespitosa sensu lat.] by Ernst in 1964. K Triphysaria pusilla (Benth.) T.I. Chuang & Heckard 000653325. Coulter 611. TCD: Orthocarpus pusillus, California (PL). Annotation 1: Orthocarpus Eschscholzia californica Cham. pusillus Benth. Torr. & Gray, Fl. N. Amer. TCD! Coulter 25. BRU: Orthocarpus, California (PL). PBRU 00045000. TCD: Ch[ryseis] douglasii H. & A., Monterey, California, 1832 (SL). E: Triphysaria pusilla, California [no image]. E 272567. TCD! Coulter s.n. K: Ch[ryseis] douglasii? California (SL). Annotation 1: Eschscholzia UC: Triphysaria pusilla, California [no image]. UC 973622. californica by Ernst in 1964. K 000340213. 52 Wallace ALISO

Coulter 26. BRU: Platystemon californicum, California (SL). Herbarium label: Herb. TCD: Chryseis californica Hook. & Arn., Monterey (SL). TCD! A. Gray, Torr. & Gray, Fl. N. Amer. Annotation 1: Herbarium Olneyanum Coulter s.n. Platystemon californicum Benth. PBRU 00029652. TCD: C[hryseis] californica Hook. & Arn., Mt [ = Monterey] to El Sur, GH: Platystemon californicus, California [no image]. GH 00295021. California (SL). TCD! [On sheet with Coulter 26 above] UC: Platystemon californicus, California [no image]. UC 973616. TCD: C. californica Hook. & Arn., Monterey, California (SL). TCD! TCD: Ch. californica Hook. & Arn., Monterey to S. Miguel, California Romneya coulteri Harv. (SL). TCD! Coulter 20. TCD: Ch. californica Hook. & Arn., San Gabriel, California (SL). TCD! K: Romneya coulteri Harv., California (PL). Annotated: Romneya coul- Note: on sheet with Coulter s.n., Monterey to S. Miguel above. teri Harv. by Ernst 1964. K 000653099. TCD: Ch. californica H. & A., between San Miguel & San Barbara, Cal- K: Romneya coulteri Harv., California (PL). Annotation 1: Romneya ifornia, California. 1832 (SL). TCD! coulteri Harv. by Ernst 1964. K 000653101. BRU: Chryseis, California (PL). Annotation 1: Herbarium Olneyanum. Coulter s.n. PBRU 00029246. TCD: Romneya coulteri Harv., California (PL, top line cut off). TCD BRU: Chryseis, California (PL). Annotation 1: Herbarium Olneyanum. 0016618! PBRU 00029247. GH: Romneya coulteri Harv., California (PL). Written on label: “Dscrips. Harvey!” Annotation 1: Syn. Fl. N. Amer. Annotated: ! by Welsh in Eschscholzia hypecoides Benth. 1992. Annotation 2: ISOTYPE Romneya coulteri Harv. by Schwarzbach Coulter 24x. in 1996. GH 00004589. K: C. hypecoides Benth., California (SL). Annotation 1: E. caespitosa var. hypecoides Gray Syn. Fl. N. Amer. Annotation 2: Eschscholzia hy- pecoides Benth. by Ernst in 1964. K 000340218. Coulter 27. phrymaceae TCD: [no name], Monterey to San Francisco, California (SL). Written Mimetanthe pilosa (Benth.) Greene on sheet: Echl. [sic] hypecoides B[enth.]. TCD! Coulter 623. K: Chryseis pilosa [crossed out] hypecoides var., California (SL). K BRU: [no name], California (PL). Annotation 1: Herbarium Olneyanum. 000340219. Note: on sheet with Coulter 24x above. [Filed as Herpestis]. PBRU 00046655. Coulter 28. TCD: [no name], California (SL). Written on sheet: E. hypecoides. TCD! Mimulus aurantiacus Curtis var. Coulter s.n. Coulter 638. GH: Eschscholzia hypecoides, California [no image]. Annotation 1: TCD: Diplacus glutinosus, California (PL). Annotation 1: D. glutinosus Eschscholzia hypecoides by Ernst in 1961. GH 381490. β brachypus,Torr.&Gray,Fl.N.Amer.TCD! GH: Mimulus aurantiacus var. pubescens, California [no image]. Anno- Hesperomecon linearis (Benth.) Greene tation 1: Mimulus aurantiacus var. pubescens by Thompson in 1993. GH Coulter s.n. 00549472. BRU: Platystigma lineare, California (PL). Annotation 1: Herbarium Ol- Coulter 639. neyanum Platystigma lineare Benth. PBRU 00029672. TCD: Diplacus glutinosus, California (PL). Annotation 1: D. glutinosus PH: Platystigma lineare Benth., California (PL). Annotation 1: Hesper- inter α and β Torr. & Gray, Fl. N. Amer. Annotation 1: Type material omecon linearis (Benth.) Greene by Ernst in 1968. PH 00030654. Mimulus glutinosus var. brachypus A. Gray by G. Wallace in 2017. TCD! UC: Platystigma lineare, California (PL). UC 973615. GH: [no name], California (PL). Annotation 1: LECTOTYPE Mimulus glutinosus var. brachypus by Thompson in 1993. Annotation 2: Mimulus aurantiacus var. pubescens by Thompson in 1993. GH 00022822. Papaver heterophyllum (Benth.) Greene K: M. (Diplacus) glutinosus var. brachypus T. & Gr., California (PL). An- Coulter 19. notation 1: ISOLECTOTYPE Mimulus glutinosus Wendland var. brachy- TCD: [no name], between Monterey & San Francisco, California (SL). pus A. Gray in W.H. Brewer, S. Watson, & A. Gray Bot. California 1 (ed. Written on sheet: Meconopsis heterophylla. TCD! 1) p. 566. 1876. Lectotype GH, other isolectotypes E, K (another sheet), TCD: [no name], Monterey, California, 1832 (SL). Written on sheet: by Thompson 1993. Annotation 2: Mimulus aurantiacus var. pubescens Meconopsis heterophylla. TCD! (Torrey) D. Thompson by Thompson in 1993. K 001079313. Coulter s.n. K: [no name], California (PL). Annotation 1: ISOLECTOTYPE Mimu- TCD: [no name], between San Miguel & San Barbara, California (SL). lus glutinosus Wendland var. brachypus A. Gray in W.H. Brewer, S. Wat- TCD! son, & A. Gray Bot. California 1 (ed. 1) p. 566. 1876. Lectotype GH, BRU: Meconopsis heterophylla, California (PL). Annotation 1: Herbar- other isolectotypes E, K (another sheet) by Thompson 1993. Annotation ium Olneyanum Meconopsis heterophylla Benth. PBRU 00029258. 2: Mimulus aurantiacus var. pubescens (Torrey) D. Thompson by Thomp- UC: Stylomecon heterophylla, California (PL). UC 973614. son in 1993. K 001079314. Note: K database lists J.M. Coulter as collec- tor in error. Platystemon californicus Benth. Coulter 641. Coulter 22. TCD: Diplacus glutinosus, California (PL). Annotation 1: Diplacus BRU: Platystemon californicum, California (PL). Annotation 1: Herbar- glutinosus Torr. & Gray, Fl. N. Amer. TCD! ium Olneyanum Platystemon californicum Benth. PBRU 00029655. GH: Mimulus aurantiacus, California [no image]. Annotation 1: Mimulus Coulter s.n. aurantiacus by Thompson in 1993. GH 365936. TCD: Platystemon californicum, Monterey, California, 1832 (SL). TCD! K: [no name], California (SL). Annotation 1: Mimulus aurantiacus var. TCD: [no name], Monterey to San Miguel, California, 1832 (SL). TCD! aurantiacus by Thompson in 1993. Written on sheet: D. glutinosus. K TCD: Platystemon californicum, between San Miguel & San Barbara, 001079311. Note: K database lists J.M. Coulter as collector in error. California, 1832 (SL). TCD! Note: on sheet with Coulter s.n. [no name] Coulter 642. above. K: [no name], California (SL). Annotation 1: forma linearis Syn. Fl. N. TCD: [no name], Monterey, California, 1832 (SL). Written on sheet: Amer. Annotation 2: Mimulus aurantiacus var. aurantiacus by Thompson Platystemon californicum. TCD! in 1993. K 001079312. [Filed as Mimulus aurantiacus var. pubescens]. VOLUME 37(1–2) Plantae Coulterianae 53

Note: K database lists as Coulter s.n. and J.M. Coulter as collector in Antirrhinum nuttallianum A. DC. subsp. error. Coulter 599. TCD: Linaria, California (PL). Annotation 1: Type material Antirrhinum Mimulus aurantiacus Curtis var. puniceus (Nutt.) D.M. Thomps. nuttallianum A. DC. by G. Wallace in 2017. Written on sheet: Antir- Coulter 642. rhinum nuttallianum Benth. TCD! TCD: Diplacus glutinosus, California (PL). Annotation 1: D. glutinosus GH: Antirrhinum nuttallianum Benth., California (PL). Annotation 1: β puniceus Torr. & Gray, Fl. N. Amer. TCD! SYNTYPE Antirrhinum nuttallianum Benth. by Thompson in 1986. GH 00077907. Mimulus brevipes Benth. K: Antirrhinum nuttallianum, California (SL). Annotation 1: Antir- Coulter 640 . rhinum nuttallianum subsp. subsessile (A. Gray) D. Thompson. K TCD Mimulus brevipes β Mimulus bre- : , California (PL). Annotation 1: 000979994. vipes Benth. Torr. & Gray, Fl. N. Amer. TCD!

Mimulus cardinalis Benth. Collinsia heterophylla Graham var. Coulter 637. Coulter 602. TCD: Mimulus cardinalis, California (PL). Annotation 1: M. cardinalis TCD: Collinsia bicolor Benth., California (PL). TCD! Torr. & Gray, Fl. N. Amer. TCD! TCD: Collinsia bicolor Benth. Douglas, California (PL). TCD! BRU: Collinsia, California (PL). Annotation 1: Herbarium Olneyanum. Mimulus douglasii (Benth.) A. Gray PBRU 00046768. Coulter 636. Coulter s.n. TCD: [no name], California (PL). TCD! UC: Collinsia heterophylla, California [no image]. UC 973625. BRU: Eunanus douglasii, California (PL). Annotation 1: Herbarium Ol- neyanum. PBRU 00043615. Collinsia parryi A. Gray GH: Mimulus douglasii, California (PL). Annotation 1: Syn. Fl. N. Amer. Coulter 597. Annotation 2: Mimulus douglasii (Benth.) A. Gray by D. Thompson in GH: Collinsia parryi, California [no image]. Annotation 1: Collinsia par- 1992. GH 00549505. ryi by Newsom in 1927. GH 00365623. K: Mimulus nanus H. & A. (SL). Annotation 1: Coll. Coulter M. douglasii Collinsia parviflora Lindl. A. Gray fide Grant. Annotation 2: Mimulus douglasii (Benth.) A. Gray by Thompson in 1992. K 001079300. Note: K database lists J.M. Coulter as Coulter 604. TCD Collinsia parviflora Collinsia collector and collection date as 1833 in error. : , California (PL). Annotation 1: parviflora TCD Coulter s.n. Torr. & Gray, Fl. N. Amer. ! UC Mimulus douglasii UC : , California [no image]. 973626. Collinsia sparsiflora Fisch. & C.A. Mey. var. Mimulus fremontii (Benth.) A. Gray var. Coulter 597. TCD: Collinsia sparsiflora, California (PL). Annotation 1: Collinsia Coulter 631. sparsiflora F. & Meyer. TCD! TCD: Eunanus fremontii β major, California (PL). TCD! GH Mimulus fremontii : [no name], California (PL). Annotation 1: by Collinsia sparsiflora Fisch. & C.A. Mey. var. collina (Jeps.) Newsom Thompson in 1992. GH 00549314. Coulter 604. BRU Veronica virginica Mimulus guttatus DC. : , California (PL). Annotation 1: Herbarium Ol- neyanum. PBRU 00047924. Coulter 623. UC: Collinsia sparsiflora var. collina, California [no image]. UC 973624. BRU: [no name], California (PL). [Not sure of genus, pedicels flexuous]. PBRU 00046655. Keckiella antirrhinoides (Benth.) Straw var. antirrhinoides Coulter 643. Coulter 633. TCD: Mimulus luteus, Monterey to S. Gabriel, California (PL). TCD! TCD: Penstemon antirrhinoides, California (PL). Annotation 1: P. an- Coulter 644. tirrhinoides Benth. Torr. & Gray, Fl. N. Amer. Annotation 2: ISOTYPE TCD: Mimulus luteus, Mimulus parviflorus, California (PL). Written on Penstemon antirrhinoides Benth. ex DC. by G. Wallace in 2017. TCD! sheet: M. luteus A. Gr. TCD! BM: [no name], California (PL). Annotation 1: P. antirrhinoides by Mimulus tricolor Lindl. Ewan in 1954. Annotation 2: ISOTYPE Penstemon antirrhinoides. BM Coulter 614. 1025006. GH Keckiella antirrhinoides Keck- TCD: Eunanus coulteri, California (PL). TCD! : , California [no image]. Annotation 1: iella antirrhinoides by Michener in 1984. GH 00365676. [This is type material] K: Penstemon antirrhinoides Benth., California (SL). Annotation 2: TYPE SPECIMEN. K 000979800. plantaginaceae K: Penstemon sp., California (PL). Annotation 1: Herbarium Careyanum. Annotation 2: Penstemon antirrhinoides Benth. ssp. antirrhinoides by Antirrhinum coulterianum A. DC. Crosswhite in 1969. Annotation 3: ISOTYPE Penstemon antirrhinoides Coulter 607. Benth. K 000979801. TCD: Antirrhinum coulterianum, California (PL). Annotation 1: Antir- K: Penstemon antirrhinoides Benth., California (PL). K 000979802. rhinum coulterianum Torr. & Gray, Fl. N. Amer. Annotation 2: ISOTYPE Antirrhinum coulterianum A. DC. by G. Wallace in 2017. TCD! Keckiella corymbosa (A. DC.) Straw GH: Antirrhinum coulterianum Benth., California (PL). Annotation 1: Coulter 629. ISOTYPE Antirrhinum coulterianum Benth. LT: K! by Thompson in TCD: Penstemon corymbosus, California (PL). Annotation 1: P. c o r y m - 1986. Written on sheet: perhaps an Isotype. GH 00077900. bosus Benth. Torr. & Gray, Fl. N. Amer. Annotation 2: Type material Pen- K: Antirrhinum coulterianum, California (SL). Annotation 1: A. coulteri- stemon corymbosus Benth. ex DC. by G. Wallace in 2017. TCD! anum Benth. TYPUS by Rothmaler in 1957. Annotation 2: TYPE SPEC- BRU: [no name], California (PL). Annotation 1: Herbarium Olneyanum. IMEN. K 000979997. PBRU 00042376. [Name listed as Ehretia in BRU database] 54 Wallace ALISO

GH: [no name], California (PL). Annotation 1: Probably ISOTYPE P. c . platanaceae Benth. D.D. Keck in 1935. Written on sheet: sterile filament erroneously Platanus racemosa Nutt. described as glabrous. GH 00091228. Note: in GH database as Penstemon Coulter 703. corymbosus, probable ISOTYPE. TCD: Platanus Nutt., S. Miguel to S. Barbara, California (PL). Written K: Penstemon corymbosus Benth., California (SL). Annotation 1: TYPE on sheet: Bot. Beechey. TCD! SPECIMEN. K 000979805. K: Penstemon sp., California (PL). Annotation 1: P. corymbosus Benth. plumbaginaceae by Crosswhite in 1971. Written on sheet: type number. K 000979806. NY: Penstemon corymbosus Benth. in DC. Prod. X, p. 593, California Armeria maritima (Mill.) Willd. subsp. californica (Boiss.) A.E. Porsild (PL). Annotation 1: ISOTYPE of Penstemon corymbosus Benth. in DC. Coulter 577. Prod. 10: 593. 1846. Written on sheet: ISOTYPE Penstemon corymbosus TCD: Armeria vulgaris Hook. & Arn., California (PL). Annotation 1: Benth. NY 00091053. ISOTYPE Armeria andina var. californica Boiss. by G. Wallace in 2017. UC: Keckiella corymbosa, California [no image]. UC 525524. TCD! BRU: Armeria, California (PL). Annotation 1: Herbarium Olneyanum. Mohavea confertiflora (A. DC.) A. Heller PBRU 00041054. Coulter 616. GH: Armeria maritima subsp. californica, California [no image]. Anno- TCD: [no name], California (PL). Note fixed to sheet: “Bates [?] This tation 1: Armeria maritima var. californica by Lawrence in 1940. GH is my Mohavea and I would have given anything for the fruit but never 00376103. Note: this is type material of Armeria maritima var. califor- thought of your parcel. To much amends, I give you (herewith) my best nica. flowering specimen; and then to complete my knowledge of thegenusI K: [no name], California (SL). Annotation 1: TYPE A. andina Poep- β mutilate your fruiting specimen a little, that I may complete my knowl- pig californica Prod. 12, 682 by E.B. [Boiss.]. Annotation 2: TYPE edge of the genus. A.G.” [A. Gray]. Annotation 1: ISOTYPE Antirrhinum Armeria maritima var. californica (Boiss.) Lawr. by Lawrence in 1950. confertiflorum Benth. ex DC. by G. Wallace in 2016. Written on sheet: K 000704630. Mohavea viscida Gray. TCD! Coulter s.n. K: Antirrhinum confertiflorum, California (SL). Annotation 1: TYPE UC: Armeria maritima subsp. californica, California [no image]. UC SPECIMEN. K 000979975. [Filed as Schweinfurthia confertiflora] 973611. polemoniaceae Penstemon centranthifolius Benth. Allophyllum glutinosum (Benth.) A.D. Grant & V.E. Grant Coulter 630. Coulter 461. TCD: Penstemon centranthifolius, California (PL). Annotation 1: Pen- TCD: Collomia glutinosa Benth., Monterey (SL). Annotation 1: Collo- stemon centranthifolius Torr. & Gray, Fl. N. Amer. TCD! mia glutinosa Torr. & Gray, Fl. N. Amer. TCD! Coulter 635. TCD: Penstemon centranthifolius, California (PL). Annotation 1: Pen- Eriastrum densifolium (Benth.) H. Mason subsp. elongatum (Benth.) H. stemon centranthifolius Torr. & Gray, Fl. N. Amer. TCD! Mason Coulter 453. TCD: Gilia (Huegelia) elongata, California (PL; Ex is crossed out). An- Penstemon heterophyllus Lindl. var. notation 1: Eriastrum densifolium subsp. elongatum by De Groot in 2007. Coulter 626. TCD! BRU: Conobea multifida, California (PL). Annotation 1: Herbarium Ol- BRU: Gilia elongata, California (PL). Annotation 1: Herbarium Ol- neyanum Cocobaea multifida. PBRU 00046670. neyanum. PBRU 00042680. Coulter 632. GH: [no name], California [no image]. GH 00303692. TCD: Penstemon heterophyllus (SL). Note on label: The form with the Coulter s.n. α narrow calyx ( Hook. & Arn.). Torr. & Gray, Fl. N. Amer. TCD! BM: Huegelia elongata, California (PL). BM 000939588.

†Plantago major L. Eriastrum virgatum (Benth.) H. Mason Coulter 571. Coulter 452. TCD: [no name], California (PL). Annotation 1: Plantago major L. by TCD: Gilia (Huegelia) virgata B., California (PL). TCD! Parnell in 2002. TCD! Gilia achilleifolia Benth. subsp. Coulter 468. Plantago sp. TCD: Gilia capitata, California (PL). Annotation 1: Gilia achilleifolia Coulter 568. Bentham by Porter in 2007. Written on sheet: capitata. TCD! TCD: Pl. mexicana, California (PL). TCD! BRU: Gilia capitata, California (PL). Annotation 1: Herbarium Ol- Coulter 568 bis. neyanum Gilia capitata Douglas. PBRU 00042849. TCD: [no name], California (PL). TCD! Coulter 468x. Coulter 569. TCD: Gilia capitata, California (PL). Annotation 1: Gilia achilleifolia TCD: Pl. mexicana, California (PL). TCD! Benth. by M. Porter in 2007. TCD! BRU: Plantago mexicana, California (PL). Annotation 1: Herbarium Ol- neyanum. PBRU 00046838. Gilia achilleifolia Benth. subsp. multicaulis (Benth.) V.E. Grant & A.D. Coulter 569*. Grant TCD: [no name], California (PL). TCD! Coulter 466. Coulter 570. TCD: [no name], California (PL). Annotation 1: Gilia multicaulis B. de- TCD: Pl. mexicana, California (PL). TCD! pauperata,Torr.&Gray,Fl.N.Amer.TCD! Coulter 572. GH: Gilia achilleifolia subsp. multicaulis, California [no image]. GH TCD: Pl. mexicana, California (PL). TCD! 01083906. Coulter 573. K: [no name], California (SL). Written on sheet: G. multicaulis var. ten- TCD: Pl. mexicana, California (PL). TCD! era A. Gr. K 000769103. VOLUME 37(1–2) Plantae Coulterianae 55

K: Gilia? Collomia gilioides, California (PL). Written on sheet: Gilia BRU: Gilia parvifolia [sic], California (PL). Annotation label: Herbar- multicaulis forma. tenera A. Gr. K 000769108. ium Olneyanum. PBRU 00042188. Coulter 467. Coulter s.n. TCD: Gilia, California (PL). Written on sheet: multicaulis. TCD! UC: Linanthus parviflorus, California [no image]. UC 973812. BRU: [no name], California (PL). Annotation 1: Herbarium Olneyanum. PBRU 00043125. Linanthus californicus (Hook. & Arn.) J.M. Porter & L.A. Johnson GH: Gilia multicaulis var. tenera, California [no image]. Torr. & Gray, Fl. Coulter 481. N. Amer. Annotation 1: Gilia multicaulis Benth. by R.H.W. Annotation TCD: [no name], California (PL). Annotation 1: Linanthus californicus 2: Gilia multicaulis var. tenera Syn. Fl. N. Amer. Annotation 3: Gilia (Hook. & Arn.) J.M. Porter & L.A. Johnson by Porter in 2007. Written achilleifolia subsp. multicaulis by Kittredge in 2017. Written on sheet: on sheet: Gilia californica and Leptodactylon californicus. TCD! Gilia achilleifolia subsp. multicaulis (Benth.) V.E. Grant & A.D. Grant. BRU: Gilia californica, California (PL). Annotation 1: Herbarium Ol- GH 01083905. neyanum Gilia californica. PBRU 00042839. K: [no name], California (SL). Written on sheet: G. multicaulis var. ten- GH: Linanthus californicus (Hook. & Arn.) J.M. Porter & L.A. John- era A. Gr. K 000769104. son, California [no image]. Annotation 1: Leptodactylon californicum by Coulter s.n. Gordon in 1983. GH 1104507. TCD: G. capitata, California (PL). Annotation 1: G. multicaulis Torr. & Gray, Fl. N. Amer. TCD! Linanthus dianthiflorus (Benth.) Greene Coulter 463. Gilia sp. TCD: Gilia dianthiflora, S. Miguel to S. Barbara, California (PL). An- Coulter 474. notation 1: Linanthus dianthiflorus E.L. Greene by Porter in 2007. TCD! BRU: [no name], California (PL). Annotation 1: Herbarium Olneyanum. BRU: Gilia dianthiflora, California (PL). Annotation 1: Herbarium Ol- PBRU 00043126. neyanum Gilia dianthoides Endl. PBRU 00042667. GH GH Langloisia setosissima (Torr. & A. Gray) Greene : [no name], California (PL). 00872604. Coulter 463x. Coulter 449. TCD: Gilia dianthiflora, California (PL). Annotation 1: Linanthus di- TCD: Navarretia setosissima, Aegochloa, California (PL). TCD! anthiflorus E.L. Greene by Porter in 2007. TCD! Leptosiphon androsaceus Benth. Coulter 464. Coulter 460. TCD: Gilia coulteri n. sp., California (PL). Annotation 1: Linanthus (Di- GH: Gilia micrantha, California [no image]. Annotation 1: Linanthus anthoides) coulteri Harv. & Gr. Torr. & Gray, Fl. N. Amer. Annotation 1: androsaceus subsp. micranthus. Annotation 2: Gilia lutea by R.H.W. in Linanthus dianthiflorus E.L. Greene (sic) by Porter in 2007. Written on 1922. GH 00967846. sheet: Possibly one of Brand’s in D.C. [illegible]. TCD! Coulter s.n. K: [no name], California. Written on sheet: Leptosiphon (Fenzelia) di- BRU: Gilia androsacea, California (PL). Annotation 1: Herbarium Ol- anthiflorus (Gilia Benth.) var. coulteri:Gray,floribunda corolla lobis vix neyanum Gilia androsacea Steud. PBRU 00042868. erosis A. Gray, 1869. K 000614080. Coulter s.n. Leptosiphon floribundus (A. Gray) J.M. Porter & L.A. Johnson subsp. UC: Linanthus dianthiflorus, California [no image], UC 973809. floribundus Coulter 454. Linanthus dichotomus Benth. TCD: Gilia, California (PL). Annotation 1: Linanthus (Dactylophyllum) Coulter 462. floribundus Harv. & Gray. Torr. & Gray, Fl. N. Amer. Annotation 2: Lep- TCD: Gilia dichotomum, Benth., California (PL). Annotation 1: Linan- tosiphon floribundus (A. Gray) J.M. Porter & L.A. Johnson by Porter in thus dichotomus Bentham by Porter in 2007. TCD! 2007. TCD! BRU: Gilia dichotoma, California (PL). Annotation 1: Herbarium Ol- GH: [no name], fragment folder only (SL). Annotation 1: Linanthus neyanum Gilia dichotoma Benth. PBRU 00042669. floribundus (Gray) Greene ex Millikan subsp. floribundus HOLOTYPE GH: Linanthus dichotomus, California [no image]. GH 945148. by Patterson in 1974. GH 78872. Note: listed as LECTOTYPE in GH database. Linanthus sp. K: [no name], California (SL). Written on sheet: Leptosiphon (Siphonella Coulter s.n. Nutt.) montanus fere var. acensus A. Gray 1864. Gilia nuttallii A. Gray. UC: Linanthus, California [no image]. UC 973808. K 000769043. Loeseliastrum schottii (Torr.) Timbrook Leptosiphon grandiflorus Benth. Coulter 449. Coulter 455. TCD: Navarretia setosissima, California (PL). TCD! TCD: Gilia densiflora, S. Miguel to S. Barbara, California (PL). Annota- GH: Navarretia schottii, California [no image]. Annotation 1: Loeselias- tion 1: Linanthus (Dactylo.) densiflorus. Torr. & Gray, Fl. N. Amer. An- trum schottii. GH 267259. notation 2: Leptosiphon grandiflorus Benth. by Porter in 2007. TCD! GH: Linanthus grandifloras, San Miguel to Santa [likely corrected from Microsteris gracilis (Hook.) Greene original “San”] Barbara, California [no image]. GH 00385233. Coulter 459. TCD: Collomia gracilis, S. Francisco, California (PL). TCD! Leptosiphon liniflorus (H. Mason) R. Battaglia BRU: Collomia gracilis, California (PL). Annotation 1: Herbarium Ol- Coulter 457. neyanum. PBRU 00041508. TCD: Gilia pharnaceoides Benth., California (PL). TCD! GH: Collomia gracilis, California (PL). GH 967221. GH: Leptosiphon liniflorus, California [no image]. GH 1105541. Coulter s.n. UC: Phlox gracilis, California [no image]. UC 973807. Leptosiphon parviflorus Benth. Coulter 461. Navarretia atractyloides (Benth.) Hook. & Arn. TCD: Gilia parviflora?, California (PL). Written on sheet: G. micrantha. Coulter 450. TCD! TCD: Navarretia atractyloides β, California (PL). TCD! 56 Wallace ALISO

K: Aegochloa atractyloides Benth., A. viscida Nutt. (SL). Annotation Polygonum sp. 1: Navarretia atractyloides (Benth.) H. & A. by Day in 1992. Written Coulter 696. on sheet: Aegochloa atractyloides Benth., A. Gr. in Proc. Am. Acad. K TCD: [no name], California (PL). TCD! 000190137. Coulter 697. Coulter 456. TCD: [no name], California (PL). TCD! TCD: Navarretia atractyloides var., Monterey (SL). TCD! Coulter 697x. TCD: [no name], California (PL). TCD! Coulter 698. TCD: [no name], California (PL). TCD! polygonaceae Pterostegia drymarioides Fisch. & C.A. Mey. Chorizanthe fimbriata Nutt. var. fimbriata Coulter 89. Coulter 574 in part. TCD: Pterostegia drymarioides, California (PL). Annotation 1: Pteroste- TCD: Chorizanthe staticoides, California (PL). Annotation 1: Chorizan- gia drymarioides Fisch. & Meyer by Reveal 1988. TCD! the fimbriata Nutt. var. fimbriata by Reveal in 1987. TCD! K: Pterostegia drymarioides (SL). Annotation 1: Pterostegia drymari- Chorizanthe fimbriata Nutt. var. laciniata (Torr.) Jeps. oides Fisch. & Meyer by Reveal 1987. K 000830390. Coulter 574*. Rumex sp. TCD: Chorizanthe fimbriata, California (PL). Annotation 1: Chorizanthe Coulter 692. fimbriata Nutt. var. laciniata (Torr.) Jeps. by Reveal in 1987. TCD! TCD: Rumex, California (PL). TCD! Chorizanthe leptotheca Goodman Coulter 695. Coulter 579. TCD: [no name], Monterey to S. Gabriel, California (PL). TCD! TCD: Chorizanthe staticoides, California (PL). Annotation 1: Chorizan- primulaceae the leptotheca Goodman by Reveal in 1987. TCD! Dodecatheon hendersonii A. Gray Chorizanthe sp. Coulter s.n. Coulter 575. UC: Dodecatheon hendersonii, California [no image]. Annotation 1: Do- TCD: Eriogonum, California (PL). TCD! decatheon hendersonii A. Gray by Reveal in 2006. UC 973610.

Chorizanthe staticoides Benth. Dodecatheon sp. Coulter 576. Coulter 567. TCD: Chorizanthe staticoides, California (PL). Annotation 1: Chorizan- TCD: Dodecatheon meadia [crossed out] D. frigidum?, California (PL). the staticoides Benth. by Reveal in 1987. TCD! TCD! K: [no name], California (SL). Note on label: no leaves. Annotation BRU: Dodecatheon integrifolium, California (PL). Annotation 1: Herbar- 1: Chorizanthe xanti S. Watson by Eastwood in 1911. Annotation 2: ium Olneyanum PBRU 00040884. Chorizanthe staticoides Benth. by Reveal in 1987. K 000830305. Coulter s.n. Coulter s.n. TCD: [no name], California (PL). Note on label: something [illegible] TCD: Mucronea, California (Part of PL). Annotation 1: Chorizanthe from the other specimens. TCD! staticoides by Reveal in 1987. TCD! BRU: Pterygodium catholicum [Orchidaceae], California (PL). Annota- tion 1: Herbarium Olneyanum. PBRU 00043303. Eriogonum angulosum Benth. Coulter 193. ranunculaceae TCD: Eriogonum angulosum Benth., California (PL). TCD! Aquilegia formosa DC. Eriogonum elatum Benth. var. elatum Coulter 7. Coulter 693. TCD: A. formosa Monterey, California (SL). TCD! TCD: Eriogonum elatum, California (PL). TCD! Clematis lasiantha Nutt. Eriogonum fasciculatum Benth. var. Coulter s.n. Coulter 191. TCD: Clematis lasiantha Nutt., Monterey, California (SL). TCD! TCD: Eriogonum fasciculatum, California (PL). TCD! Delphinium californicum Torr. & A. Gray subsp. californicum Eriogonum gracile Benth. var. Coulter 9. Coulter 694. TCD: D. californicum Tor. & Gr., Monterey to El Sur, California, 1832 TCD: Eriogonum gracile Benth., California (PL). TCD! (SL). TCD!

Eriogonum parvifolium Sm. Delphinium hesperium A. Gray subsp. Coulter 700. Coulter 11. TCD: Eriogonum parvifolium, California (PL). TCD! K: [no name], California (SL). Name written on sheet: D. hesperium. K 000692540. Lastarriaea coriacea (Goodman) Hoover Coulter 190 (in part). Delphinium nudicaule Torr. & A. Gray TCD: Mucronea californica, California (SL). Annotation 1: Lastarriaea Coulter 10. coriacea (Goodman) Hoover by Reveal in 1987. TCD! TCD: Delphinium nudicaule Tor. & Gr., Monterey, California, 1832 (SL). TCD! Mucronea californica Benth. Coulter 190 (in part). Delphinium sp. TCD: Mucronea californica, California (SL). Annotation 1: Mucronea Coulter 11. californica Benth. by Reveal in 1987. TCD! TCD: Delphinium simplex, Monterey, California, 1832 (SL). TCD! VOLUME 37(1–2) Plantae Coulterianae 57

TCD: Delphinium simplex, Between Monterey and San Francisco, Cali- K: Ceanothus divaricatus Nutt., California (SL). Annotation 1: Cean- fornia, 1832 (SL). TCD! othus leucodermis Greene = C. divaricatus of American authors not Nut- Coulter 12. tall by H.E. McMinn in 1935. K 000729083. TCD: Delphinium menziesii, Between Monterey & San Francisco, Cali- Coulter 123. fornia, 1832 (SL). TCD! TCD: Ceanothus divaricatus, California (PL). TCD! TCD: Delphinium menziesii. Monterey, California, 1832 (SL). TCD! K: C. divaricatus? var., California (SL). Annotation 1: Ceanothus leuco- Coulter 13. dermis Greene = C. divaricatus of American authors not Nuttall by H.E. TCD: Delphinium menziesii? Monterey to El Sur, California, 1832 (SL). McMinn in 1937. K 000729087. TCD! BRU: Delphinium menziesii, California (PL). Annotation 1: Herbarium Ceanothus oliganthus Nutt. var. Olneyanum Delphinium. PBRU 00030027. Coulter 122. Coulter 14. BRU: Ceanothus divaricatus, California (PL). Annotation 1: Herbarium TCD: Delphinium menziesii, Monterey, California, 1832 (SL). TCD! Olneyanum Ceanothus divaricatus. PBRU 00006226. Coulter 15. GH: Ceanothus oliganthus, California [no image]. Annotation 1: Cean- TCD: D. menziesii, Between Monterey & San Miguel, California (SL). othus divaricatus. [no image] GH 00349897. TCD! Ceanothus rigidus Nutt. Delphinium variegatum Torr. & A. Gray subsp. variegatum Coulter 125. Coulter 8. TCD: Ceanothus rigidus, California (PL). TCD! TCD: D. variegatum? Between San Miguel and San Barbara, California, BRU: Ceanothus rigidus, California (PL). Annotation 1: Herbarium Ol- 1832 (SL). TCD! neyanum Ceanothus rigidus Nutt. PBRU 00006277 Ranunculus californicus Benth. var. GH: Ceanothus rigidus, California [no image]. GH 00425228. Coulter 3. K: C[eanothus] rigidus, California (SL). K 000729173. TCD: [no name], Written on sheet: R. delphinifolius, Monterey (SL). Ceanothus sp. TCD! Coulter 5. Coulter 113. TCD TCD TCD: Ranunculus delphinifolius T. & G., R. dissectus H. & A., Mon- : [no name], California (PL). ! terey, California, 1832 (SL). Note on label: agrees with specimen in Herb. Ceanothus thyrsiflorus Eschsch. var. Hooker W.H.H. [ = Harvey, Herbarium Curator at TCD]. TCD! Coulter 124. TCD Ceanothus thyrsiflorus TCD Ranunculus hebecarpus Hook. & Arn. : , California (PL). ! [three sheets] Coulter 6. Ceanothus thyrsiflorus Eschsch. var. griseus Trel. TCD R. parviflorus : L. Monterey to San Miguel, California, 1832 (SL). Coulter 124. R. hebecarpus. TCD Written on sheet: ! GH: Ceanothus thyrsiflorus var. griseus, California [no image]. GH Coulter s.n. 00425494. TCD R. parviflorus : , Between Monterey & San Francisco, California, K: C. thyrsiflorus Esch., California (SL). Written on sheet: var. griseus. R hebecarpus 1832 (SL). Written on sheet: . . [On sheet with Coulter 6 K 000729063. above]. TCD! Coulter 124x. BRU: Ranunculus parviflorus, California (PL). Annotation 1: Herbarium K: [no name], California (SL). Written on sheet: var. griseus. K Olneyanum Ranunculus parviflorus Sims. PBRU 00030317. 000729062. UC: Ranunculus hebecarpus, California [no image]. UC 973817. Coulter 718. TCD C eanothus thyrsiflorus TCD Thalictrum fendleri A. Gray var. : [ ] Eschsch., California (PL). ! Coulter 16. Ceanothus verrucosus Nutt. TCD: Thalictrum dioicum L., Monterey to El Sur, California, Mar 1832 Coulter s.n. (SL). TCD! UC: Ceanothus rigidus, California (PL). UC 973620. Coulter s.n. TCD: Thalictrum dioicum L., Between San Miguel & San Barbara, Cal- Frangula californica (Eschsch.) A. Gray subsp. ifornia, 1832 (SL). TCD! [On sheet with Coulter 16 above] Coulter 117. TCD: Rhamnus oleifolius, California (PL). TCD! resedaceae GH: Rhamnus californica, California [no image]. Annotation 1: Fran- Oligomeris linifolia (Hornem.) J.F. Macbr. gula californica. GH 00425913. Coulter s.n. K: Rh. oleifolius, California (SL). K 000729125. TCD: Oligomeris subulata, Monterey, California (SL). TCD! Rhamnus crocea Nutt. rhamnaceae Coulter 116. TCD: Rhamnus croceus Nutt., California (PL). TCD! Ceanothus cuneatus Nutt. var. BRU: Rhamnus crocea Nutt., California (PL). Annotation 1: Herbarium Coulter 110. Olneyanum Rhamnus crocea Nutt. PBRU 00036343. TCD: Ceanothus cuneatus, S. Miguel, California (PL). TCD! BM: Rhamnus crocea, California (PL). BM 001024172. GH: Ceanothus cuneatus, California [no image]. GH 00349168. K: [no name], California (SL). K 000729136. Coulter 110x. Coulter s.n. K: Cean. cuneatus, California (SL). K 000729181. UC: Rhamnus crocea, California [no image]. UC 973619.

Ceanothus leucodermis Greene Rhamnus ilicifolia Kellogg Coulter 122. Coulter 114. TCD: Ceanothus divaricatus, California (PL). TCD! TCD: Rhamnus, California (PL). TCD! 58 Wallace ALISO

rosaceae GH: [no name], California [no image]. Annotation 1: Rosa californica Cham. & Schlec. by Cole in 1950. GH 00345904. Acaena pinnatifida Ruiz & Pav. var. californica (Bitter) Jeps. Coulter 135. Rubus parviflorus Nutt. TCD: Acaena, California (PL). TCD! Coulter 147. TCD: Rubus velutinus, Monterey to El Sur, California (PL). TCD! Adenostoma fasciculatum Hook. & Arn. var. Coulter 132. Rubus ursinus Cham. & Schltdl. K: Adenostoma fasciculata H. & A., San Francisco, California (SL). K Coulter 143. 000639489. TCD: Rubus ursinus?, California (PL). TCD! Coulter s.n. GH: Rubus ursinus, California [no image]. GH 00346318. TCD: Adenostoma fasciculatum, California (PL). TCD! Coulter 144. GH: Adenostoma fasciculatum, California [no image]. GH 00345397. TCD: Rubus ursinus?, California (PL). TCD! Coulter 145. Cercocarpus betuloides Nutt. var. betuloides TCD: Rubus ursinus?, California (PL). TCD! Coulter 141. Coulter 166. TCD: Cercocarpus parvifolius Nutt., S. Francisco, California (PL). GH: Rubus ursinus, California [no image]. GH 00346254. TCD! Coulter 142. rubiaceae TCD: Cercocarpus, California (PL). TCD! Galium angustifolium A. Gray subsp. angustifolium Drymocallis glandulosa (Lindl.) Rydb. var. Coulter 197. Coulter 138. TCD: Galium trichocarpum T. & G., California (PL). TCD! TCD: Potentilla arguta, El Sur, California (PL). TCD! GH: Galium angustifolium, California [no image]. GH 00385687. Coulter s.n. Galium aparine L. TCD: Potentilla arguta, California (PL). TCD! Coulter 195. TCD: Galium aparine, California (PL). TCD! Fragaria chiloensis (L.) Mill. GH: Galium aparine var. spurium, California [no image]. GH 00417118. Coulter 139. TCD: Fragaria chilensis [sic], El Sur, California (PL). TCD! Galium californicum Hook. & Arn. subsp. Coulter 140. Coulter 196. TCD: Fragaria chilensis?, California (PL). TCD! TCD: Galium californicum, California (PL). TCD! GH: Galium californicum, California [no image]. Annotation 1: Galium Heteromeles arbutifolia (Lindl.) M. Roem. californicum by Ehrendorfer in 1952. GH 00385808. Coulter 239. K: Galium californicum H. & A. (SL). K 000772723. Note: K database TCD: Photinia arbutifolia, California (PL). TCD! lists J. Coulter as collector in error. Horkelia cuneata Lindl. var. Galium sp. Coulter 133. Coulter 192. BRU: Horkelia cuneata, California (PL). Annotation 1: Herbarium Ol- TCD: Galium, California (PL). TCD! neyanum Horkelia cuneata Lindl. PBRU 00036596. K: Horkelia cuneata, California (SL). Written on sheet: Horkelia califor- nica Cha. & Sch. var. cuneata. K 000730047. Note: K database lists as salicaceae Coulter s.n. in error. Populus fremontii S. Watson subsp. fremontii Oemleria cerasiformis (Hook. & Arn.) J.W. Landon Coulter 648. Coulter 148. TCD: Populus, California (PL). TCD! TCD: Nuttallia cerasiformis, California (PL). TCD! BRU: [no name], California (PL). Annotation 1: Herbarium Olneyanum. BRU: Nuttallia cerasiformis, California (PL). Annotation 1: Herbarium PBRU 00032470. Olneyanum Nuttallia cerasiformis T. & G. PBRU 00036636. Coulter s.n. Salix exigua Nutt. var. TCD: Nuttallia cerasiformis, California (PL). TCD! Coulter 651. GH: Oemleria cerasiformis, California (PL). GH 00954996. TCD: [no name], California (PL). TCD! F: Salix taxifolia, California [no image]. F V038211F. Potentilla anserina L. subsp. pacifica (Howell) Rousi Coulter 651x. Coulter 137. TCD: [no name], California (PL). TCD! TCD: Potentilla anserina, California (PL). TCD! Salix lasiolepis Benth. Prunus ilicifolia (Hook. & Arn.) D. Dietr. subsp. ilicifolia Coulter 657. Coulter 115. TCD: [no name], California (PL). TCD! TCD: Cerasus ilicifolia Nutt., California (PL). TCD! GH: Salix lasiolepis, California [no image]. GH 00383524. Coulter 716. TCD: Cerasus ilicifolia Nutt., California (SL). Annotation 1: Torr. & Salix scouleriana Hook. Gray, Fl. N. Amer. TCD! Coulter 656. TCD: [no name], California (PL). TCD! Note: needs annotation label as Rosa californica Cham. & Schltdl. type material of Salix capreoides. Coulter 134. GH:1.Salix brachystachys Benth. 2. var. capreoides, California (PL). TCD: Rosa californica Cham., California (PL). TCD! Annotation 1: 656 (Coulter) (2) TYPE of S. capreoides And. (et S. VOLUME 37(1–2) Plantae Coulterianae 59

brachystachys scouleriana crassijulis And.) by Schneider in 1918. GH K: Tellima heterophylla, California (SL). K 000732609 and K 00030925. 000732608. Coulter s.n. Salix sitchensis Bong. TCD: Tellima heterophylla, California (PL). TCD! Coulter 655. TCD: Tellima heterophylla, California (PL). TCD! TCD: [no name], California (PL). TCD! Note: needs annotation label as type material of Salix coulteri. Micranthes californica (Greene) Small A: Salix coulteri, California (SL). Annotation 1: TYPE. A 30945. Coulter 184x. GH: Salix coulteri Ands., California (SL). Annotation 1: Type of Salix TCD: Saxifraga integrifolia, California (PL). TCD! coulteri; Isotype of S. coulteri. GH 0000030945. Coulter 185. GH: Salix coulteri And., California (PL). Annotation 1: Salix coulteri TCD: Sax. virginiensis?, California (PL). TCD! probable ISOTYPE by Dorn in 1997. GH 0030946. Coulter 186. K: Salix coulteri And., Salix hookerianus pro[illegible], California (SL). GH: Saxifraga californica, California [no image]. Annotation 1: Sax- Annotation 1: Salix coulteri Anderss., Proc. Amer. Acad. Arts. 4: 58. ifraga californica by Piper GH 381146. 1858. HOLOTYPE. = S. sitchensis Sanson ex Bong by Dorn in 1998. Written on sheet: Sal. Bor. Desc. p. 13; Salix coulteri And. in DC. Prod. scrophulariaceae XVI II 264. K 001079478. Scrophularia californica Cham. & Schltdl. Salix sp. Coulter 595. Coulter 650. TCD: Scrophularia, California (PL). TCD! TCD: [no name], California (PL). TCD! Coulter 596. Coulter 652. TCD: Scrophularia nodosa, S. Miguel to El Sur, California (PL). TCD! TCD: [no name], California (PL). TCD! Coulter 653. †Verbascum virgatum Stokes TCD: [no name], California (PL). TCD! Coulter 580. Coulter 654. TCD: Verbascum, California (PL). TCD! TCD: [no name], California (PL). TCD! simmondsiaceae sapindaceae Simmondsia chinensis (Link) C.K. Schneid. Acer macrophyllum Pursh Coulter 717. Coulter 93. TCD: Simmondsia californica Nutt., “looks like Ebenacea”, California TCD: Acer macrophyllum, California (PL). TCD! (SL). TCD! GH: Acer macrophyllum, California [no image]. GH 00363551. K: Simmondsia californica Nutt., California (SL). Written on sheet: see also Mexican Boundary Survey t. 49. K 000880448. Acer negundo L. Coulter 704. TCD: Acer negundo, California (PL). TCD! solanaceae GH: Acer negundo var. californicum, California [no image]. GH Lycium sp. 00363638. Coulter 582. NY: Negundo fraxinifolium Nutt., California (PL). Written on sheet: Acer TCD: [no name], California (PL). TCD! californicum (Torr.) Dietr. NY 00337739. Coulter 583. Coulter 704 [706 in CCH]. TCD: [no name], California (PL). Written on sheet: Lycium. TCD! UC: Acer negundo var. californicum, California [no image]. UC 973800. Nicotiana sp. saxifragaceae Coulter 460. TCD: [no name], California (PL). TCD! Boykinia occidentalis Torr. & A. Gray Coulter 579. Coulter 183. TCD: Nicotiana, San Gabriel, California (PL). TCD! TCD: Boykinia occidentalis, S. Gabriel, California (PL). TCD![two TCD: Nicotiana, California (PL). TCD! sheets] Petunia parviflora Juss. Heuchera pilosissima Fisch. & C.A. Mey. Coulter 626. Coulter 182. TCD: Leptophragma prostrata, California (PL). Written on sheet: Petu- TCD: Heuchera hispida, California (PL). TCD! [two sheets] nia parviflora. TCD! BRU: Heuchera hispida, California (PL). Annotation 1: Herbarium Ol- E: Petunia parviflora, California [no image]. E 00019754. neyanum Heuchera hispida Pursh. Written on sheet: not hispida. PBRU 00031110. Physalis acutifolia (Miers) Sandwith GH: Heuchera pilosissima, California [no image]. GH 00363371. Coulter 593. TCD: Solanum, California (PL). Annotation 1: Saracha acutifolia Miers, Lithophragma heterophyllum (Hook. & Arn.) Torr. & A. Gray part of the type collection = Physalis wrightii A. Gr., and hence Physalis Coulter 186. acutifolia (Miers) mihi, comb. nov. N.Y. Sandwith, 3 iv 1959. See Kew TCD: Tellima heterophylla, California (PL). TCD! Bulletin 14, pp. 231–233 (1960). Annotation 2: ISOTYPE Saracha acu- TCD: Tellima heterophylla, El Sur, California (PL). TCD! tifolia Miers by G. Wallace in 2017. Written on sheet: Physalis. TCD! BRU: Tellima heterophylla, California (PL). Annotation 1: Herbarium BM: Saracha acutifolia (SL). Annotation 1: ISOTYPE Saracha acutifo- Olneyanum Lithophragma heterophyllum. PBRU 00031414. lia by Knapp. BM 001070467. GH: Lithophragma heterophyllum, California (PL). Annotation 1: K: [no name], California (SL). Annotation 1: Physalis the name of the Lithophragma heterophylla [sic] by Taylor in 1962. GH 00363841. species I shall send later Bitter. Annotation 2: Cited as possibly syn- 60 Wallace ALISO

onymous with Chamaesaracha coronopus (Don) A. Gr.; see A. Gr. Bot. styracaceae Cal. 540; Rydberg Mem. Torr. Club iv, 370 (1896). Surely Physalis Styrax redivivus (Torr.) L.C. Wheeler wrightii A. Gr. N.Y. Sandwith 6.1.1959. Annotation 3: Physalis acutifo- Coulter 551. lia (Liers) mihi in Kew Bull. 14, p. 232 (1960) HOLOTYPUS. (ISOTY- TCD: [no name], California (PL). TCD! PUS in Hb. Trinity Col., Dublin by Sandwith in 1960. Written on sheet: Saracha acutifolia Miers, U.S. Am. Pls. II: 19. Seems to be Physalis. K 000759429. urticaceae Physalis sp. Urtica sp. Coulter 584. Coulter 705. TCD: Physalis, California (PL). TCD! TCD: [no name], California (PL). TCD! Coulter 584x. TCD: Physalis, California (PL). TCD! valerianaceae Solanum sp. Coulter s.n. Plectritis macrocera Torr. & A. Gray TCD: [no name], California (PL). Annotation 1: Solanum by Vickery in Coulter 534. 2005. TCD! TCD: Valeriana, El Sur, California (PL). Written on sheet: see Plectritis macrocera. TCD! Solanum umbelliferum Eschsch. Coulter 624. Coulter 586. TCD: [no name], S. Miguel, California (PL). Written on sheet: Plectritis E: Solanum, S. Miguel to San Barbara, California (SL). Annotation 1: macrocera Tor. & Gr. TCD! Solanum umbelliferum by Sarkinen in 2015. E 526617. TCD: [no name], California (SL). TCD! K: [no name], California (SL). Annotation 1: S. xantii Gr. by [illegi- ble]. Annotation 2: Solanum umbelliferum Eschsch. by Knapp in 2010. K 000658200. Note: K database lists J.M. Coulter as collector in error. verbenaceae Coulter 587. TCD: Solanum umbelliferum, California (PL). TCD! Phyla lanceolata (Michx.) E. Greene K: S. umbelliferum, California [no image]. Annotation 1: Solanum um- Coulter 547. belliferum Eschsch. by Knapp in 2010. Written on sheet: Solanum um- TCD: [no name], California (PL). TCD! belliferum DC 193. K 000658271. Phyla nodiflora (L.) E. Greene Coulter 588. Coulter 546. TCD: Solanum umbelliferum, S. Miguel, California (PL). TCD! TCD: Zapania [sic] nodiflora, California (PL). Written on sheet: Lippia K: S. umbelliferum California [no image]. Annotation 1: Solanum um- nodiflora. TCD! belliferum Eschsch. by Knapp in 2010. Written on sheet: Solanum um- Coulter 548. belliferum DC 193. K 000658272. TCD: Lippia chamaedrioides [sic], California (PL). TCD! Coulter 589. TCD: Solanum umbelliferum, California (PL). TCD! Verbena sp. GH: Solanum umbelliferum, California [no image]. Annotation 1: Coulter 543. Solanum umbelliferum by Knapp in 2010. GH 00267040. TCD: Verbena, California (PL). TCD! K: S. umbelliferum [no image]. Annotation 1: Solanum umbelliferum Es- Coulter 544. chsch. by Knapp in 2010. Written on sheet: Solanum umbelliferum DC TCD: Verbena, California (SL). TCD! 193. K 000658273. Note: on sheet with Coulter 587 and 588 above. Coulter 590. TCD: Solanum umbelliferum, California (PL). TCD! BRU: Solanum umbelliferum, California (PL). Annotation 1: Herbarium violaceae Olneyanum. PBRU 00047611. Viola adunca Sm. subsp. adunca E: Solanum umbelliferum, California (PL). Annotation 1: Solanum um- Coulter 69. belliferum by Knapp in 2010. E 298934. K: Viola longipes Nutt., California (SL). K 000327852. GH: Solanum umbelliferum, California [no image]. Annotation 1: Coulter s.n. Solanum umbelliferum by Knapp in 2010. GH 00267043. TCD: Viola longipes Nutt. var., Monterey, California (SL). TCD! K: S. umbelliferum, California (SL). Annotation 1: Solanum umbel- GH: [no name], California [no image]. Annotation 1: Viola adunca Sm. liferum Eschsch. by Knapp in 2010. Written on sheet: Solanum umbel- f. glabra (Brainard) G.N. Jones by Ballard in 1992. GH 381865. liferum DC 193. K 000658274. Note: on sheet with Coulter 587, 588 and GH: Viola adunca f. glabra, California [no image]. GH 381875. 598 above. Coulter s.n. Viola douglasii Steud. BM: Solanum umbelliferum, California (PL). BM 000846437. Coulter 67. E: Solanum, California (PL). Annotation 1: Solanum umbelliferum by TCD: Viola chrysantha Hook., Monterey to S. Miguel, California (SL). Sarkinen in 2015. E 00526616. Written on sheet: V. douglasii Steud., V. chrysantha Hook. TCD! Note: K: Solanum umbelliferum, California. Written on label: “ = 590” (PL). Douglas s.n. mounted on same sheet may be type material of Viola Annotation 1: Solanum umbelliferum Eschsch. by Knapp in 2010. K chrysantha Hook. 000658264. BRU: Viola chrysantha, California (PL). Annotation 1: Herbarium Ol- UC: Solanum umbelliferum, California [no image]. UC 973820. neyanum Viola chrysantha Hook. PBRU 00032677 K: Viola chrysantha Hook., California (SL). Annotation 1: Viola dou- Solanum wallacei (A. Gray) Parish glasii Steud. by Ballard in 1995. K 000327831. Coulter 586. Coulter s.n. GH: Solanum wallacei, California [no image]. GH 402703. UC: Viola douglasii, California [no image]. UC 973827. VOLUME 37(1–2) Plantae Coulterianae 61

Viola pedunculata Torr. & A. Gray TCD: [no name], California (PL). TCD! Coulter 68. Coulter 807. TCD: Viola pedunculata T. & G., San Miguel to S. Barbara, California TCD: [no name], California (PL). TCD! (SL). TCD! TCD: Viola pedunculata T. & G., Between Monterey & St. [sic] Miguel, *Cyperus articulatus L. [not known from California but is known from California (SL). TCD! Sonora, Mexico] TCD: Viola pedunculata Tor. & Gr., Monterey, California (SL). TCD! Coulter 795. Note: Douglas s.n. is on same sheet. TCD: [no name], California (PL). Annotation 1: Cyperus articulatus L. BRU: Viola pedunculata, California (PL). Annotation 1: Herbarium Ol- by Tucker in 2002. TCD! neyanum Viola pedunculata T. & Gr. PBRU 00032715. Cyperus erythrorhizos Muhl. K: Viola pedunculata T. & G., California (SL). Written on sheet: Viola Coulter 796 pedunculata Torr. & Gray. [and] Viola pedunculata Torr. & Gray Fl. of N. . TCD Cyperus erythrorhizos Am. 1, p.141. K 000370194. : [no name], California (PL). Annotation 1: Muhl. by Tucker in 2000. TCD! Coulter s.n. NY: Viola pedunculata T. & G., California [no image]. NY 03576980. Eleocharis sp. UC Viola pedunculata UC : , California [no image]. 973821. Coulter 799. TCD TCD Viola quercetorum M.S. Baker & J.C. Clausen : [no name], California (PL). ! BRU: [no name], California (PL). PBRU 00010672. Coulter s.n. GH Viola purpurea quercetorum : var. , California [no image]. Annotation Schoenoplectus californicus C.A. Mey. & Soják 1: Viola quercetorum by Baker and Clausen in 1936. GH 381996. Coulter 803. TCD:cf.Scirpus riparius (Elytrospermum C.A. Mey.), California (PL). TCD! MONOCOTS agavaceae Schoenoplectus sp. Coulter 797. Agave deserti Engelm. TCD: Scirpus aff. triqueter, California (PL). TCD! Coulter s.n. Coulter 798. TCD: Agave, California (PL). TCD! TCD: [no name], California (PL). TCD!

alliaceae Scirpus sp. Allium sp. Coulter 800. Coulter 745. TCD: [no name], California (PL). TCD! TCD: [no name], S. Miguel to S. Barbara, California (PL). TCD! Coulter 802. TCD: [no name], California (PL). TCD! cyperaceae hydrocharitaceae Bolboschoenus maritimus (L.) Palla subsp. paludosus (A. Nelson) T. Koyama Najas marina L. Coulter 804. Coulter 188. TCD: Scirpus maritimus, California (PL). TCD! TCD: Near Ruppia?, California (PL). TCD!

Carex douglasii Boott iridaceae Coulter 805. Iris longipetala Herb. TCD: [no name], California (PL). Annotation 1: Type material Carex Coulter 744. douglasii var. brunnea by G. Wallace in 2016. TCD! GH: Iris longipetala, California [no image]. GH 00357724. BRU: [no name], California (PL). Annotation 1: SYNTYPE Carex dou- th glasii var. brunnea Olney, Botany 40 Parallel p. 363. 1871 by M. Cooper Iris sp. June 24, 2015. Annotation 2: Brown University bequest of Stephen T. Ol- Coulter 724. ney 1878. PBRU 00027078. TCD: Iris tenax, California (PL). TCD! GH: Carex douglasii, California (PL). Written on sheet: C. douglasii C. BRU: [no name], California (PL). Written on sheet: probably Iris dou- Boott; C. usta Bailey. GH 222548. Note: In GH database as SYNTYPE glasiana. Annotation 1: Herbarium Olneyanum. PBRU 00009189. Carex douglasii var. brunnea Olney. Coulter 726. K: C. douglasii, California (SL). Annotation 1: C. douglasii Boott by TCD: [no name], California (PL). TCD! Clarke in 1899. Annotation 2: Coulter 805 is the “type” of Carex dou- BRU: [no name], California (PL). Annotation 1: Herbarium Olneyanum. glasii var. brunnea Olney in King’s Report (1871) 363. This again is the PBRU 00009203. [illegible] of Carex usta sp. nov. Bailey in Mem. Torrey Club v 1 (1889) p. 20. Bailey however says that his C. usta bears no immediate relation to Sisyrinchium bellum S. Watson C. douglasii and that its reference to that species is [illegible] erroneous. Coulter 723. If this n. 805 is the number Bailey had I do not agree with him. But I TCD: [no name], California (PL). TCD! think it is probable that Bailey had something else. By Clarke in 1900. K Coulter 725. 000907696. TCD: [no name], California (PL). TCD! K: [no name], (SL). Annotation 1: Coulter 805 from California was BRU: [no name], California (PL). Annotation 1: Herbarium Olneyanum. the “type” of Carex douglasii var. brunnea Olney which Bailey made a PBRU 00009302. species Carex usta Bailey. By Clarke in 1902. K 000907695. Sisyrinchium californicum (Ker Gawl.) W.T. Aiton Carex sp. Coulter 723. Coulter 806. UC: Sisyrinchium californicum, California [no image]. UC 973618. 62 Wallace ALISO

Sisyrinchium sp. BM: Calochortus, California (PL). BM 1009899. Note: filed as C. luteus, Coulter s.n. ”possible type” because mounted on sheet with Douglas specimen. BRU: [no name], California (Pl). Annotation 1: Herbarium Olneyanum. Coulter 734. PBRU 00009303. TCD: [no name], California (PL). TCD! BRU: Calochortus, California (PL). Annotation 1: Herbarium Ol- juncaceae neyanum. PBRU 00008053. Coulter 736. Juncus bufonius L. var. TCD: [no name], California (PL). TCD! Coulter 810. TCD: Juncus bufonius, California (PL). TCD! Calochortus uniflorus Hook. & Arn. BRU: [no name], California (PL). Annotation 1: Herbarium Olneyanum Coulter 735. Juncus bufonius. PBRU 00011802. TCD: Calochortus, California (PL). TCD! Juncus sp. BRU: [no name], California (PL). Annotation 1: Herbarium Olneyanum Coulter 808. [corm is included on sheet] PBRU 00008055. TCD: Juncus, California (PL). Note on label: Bot. Beechey. TCD! K: [no name], California (SL). Written on sheet: Calochortus uniflorus BRU: [no name], California (PL). Annotation 1: Herbarium Olneyanum Hook. K 000203629. Juncus falcatus.E.Mey.PBRU 00011954. MO: Juncus menziesii, California (SL). Note on fragment folder: Herb. Fritillaria affinis (Schult. & Schult. f.) Sealy A. Gray. MO 100303927. Coulter 740. Coulter 809. TCD: [no name], California (PL). Written on sheet: F. lanceolata. TCD! TCD: af. Juncus scirpoides, California (PL). TCD! MO: Juncus megacephalus, California (SL). MO 3266550. Fritillaria biflora Lindl. var. Coulter 812. Coulter 737. TCD: Juncus, California (PL). Note on label: Bot. Beechey. TCD! TCD: [no name], California (PL). Annotation 1: Fritillaria biflora Lindl. by Macfarlane in 1982. TCD! Luzula subsessilis (S. Watson) Buchenau BRU: [no name], California (PL). Annotation 1: Herbarium Olneyanum. Coulter 811. PBRU 00008146. TCD: Luzula campestris, California (PL). TCD! GH: Fritillaria biflora, California [no image]. Annotation 1: Fritillaria BRU: [no name], California (PL). Annotation 1: Herbarium Olneyanum. biflora by McFarlane in 2014. GH 00357601. PBRU 00012245. Coulter 738. GH: Luzula campestris (L.) DC., California [no image]. Annotation 1: L. TCD: [no name], California (PL). Annotation 1: Fritillaria biflora. Lindl. campestris var. congesta (Thuillier) E. Meyer by Fernald in 1912. Anno- by Macfarlane in 1982. TCD! tation 2: L. congesta (Thuillier) Lejeune by Yelton in 2003. Annotation BRU: [no name], California (PL). Annotation 1: Herbarium Olneyanum. 3: L. subsessilis (S. Watson) Buchenau by Zika in 2012. GH 00415204. PBRU 00008142. UC: Luzula comosa, California [no image]. UC 973801. GH: Fritillaria biflora, California [no image]. Annotation 1: Fritillaria biflora by Santana in 1982. GH 00357596. juncaginaceae Triglochin scilloides (Poir.) Mering & Kadereit Fritillaria liliacea Lindl. Coulter 813. Coulter 739. TCD: Lilaea H. & Bonpl., Heterostylus Hook., California (PL). TCD! TCD: [no name], California (PL). Written on sheet: Frit. liliacea. TCD! Coulter 739x. TCD: [no name], California (PL). Written on sheet: Frit. liliacea. TCD! liliaceae GH: Fritillaria liliacea, California [no image]. GH 357417. Coulter s.n. Calochortus albus (Benth.) Benth. GH: Fritillaria liliacea, California [no image]. GH 00357417. Coulter 730. TCD: [no name], California (PL). TCD! Lilium sp. Coulter s.n. Calochortus catalinae S. Watson TCD: [no name], California. Written on sheet: L. martagon. TCD! Coulter 732. TCD: [no name], California (PL). TCD! BRU: Calochortus, California (PL). Annotation 1: Herbarium Ol- neyanum. PBRU 00008052. melanthiaceae GH: Calochortus catalinae, California [no image]. Annotation 1: Calo- chortus catalinae by F.M. Ownbey in 1938 GH 00357302. Toxicoscordion fontanum (Eastw.) Zomlefer & Judd Coulter 733. Coulter 744. TCD: [no name], California (PL). TCD! TCD: Zygadenus [sic] glaucus, California (PL). TCD! BRU: Calochortus, California (PL). Annotation 1: Herbarium Ol- Coulter 746. neyanum. PBRU 00008050. TCD: Zygadenus [sic] glaucus, California (PL). TCD! GH: Calochortus venustus Douglas ex Benth., California (PL). Annota- tion 1: Calochortus catalinae by F.M. Ownbey in 1938. GH 00443365. Trillium chloropetalum (Torr.) Howell NY: Calochortus, California [no image]. Annotation 1: not determinable Coulter s.n. by Ownbey in 1939. NY 1104591. TCD: [no name], California (PL). Written on sheet: Trillium sessile. TCD! Calochortus sp. TCD: Trillium sessile, California (PL). TCD! Coulter 731. BRU: Trillium sessile, California (PL). PBRU 00008437. TCD: [no name], California (PL). TCD! UC: Trillium chloropetalum, California (PL). UC 973621. VOLUME 37(1–2) Plantae Coulterianae 63

poaceae TCD: Eragrostis aff. pilosa, S. Gabriel, California (PL). Written on sheet: Eragrostis pilosa. TCD! Note: image identified as E. pectinacea by J.T. Anthoxanthum occidentale (Buckley) Veldkamp Columbus. Coulter 786. GH: Hierochloë occidentalis, California [no image]. Annotation 1: Hie- Eragrostis sp. rochloë macrophylla. GH 360155. Coulter 772. Anthoxanthum sp. TCD: [no name], California (PL). Written on sheet: Eragrostis poaoides. Coulter 784. TCD! Note: image identified to genus by J.T. Columbus. TCD: [no name], California (PL). TCD! Note: in Hierochloë folder. BRU: Eragrostis cilianensis, California [no image]. PBRU 00026969. Coulter 774. Aristida adscensionis L. TCD: Eragrostis, California (PL). TCD! Note: image identified to genus Coulter 768. by J.T. Columbus. GH: Aristida adscensionis, California [no image]. GH 341039. †Avena barbata Link Eriochloa acuminata (J. Presl) Kunth var. acuminata Coulter 767. Coulter 787. TCD: [no name], California (PL). TCD! TCD: [no name], California (PL). TCD! Note: image identified by J.T. Columbus. Bromus carinatus Hook. & Arn. var. Coulter 760. Festuca microstachys Nutt. K: Bromus carinatus, California (SL). K 000913858. Coulter 762. Coulter 764. TCD: Vulpia, California (PL). TCD! Note: image identified by J.T. TCD: Bromus, California (PL). TCD! Note: image verified to genus by Columbus. J.T. Columbus. BRU: [no name], California (PL). Annotation 1: Herbarium Olneyanum BRU: [no name], California (PL). Annotation 1: Herbarium Olneyanum V. microstachya Nutt. PBRU 00027047. Bromus compare carinatus. PBRU 0025961. GH: Vulpia microstachya var. pauciflora, California [no image]. Anno- GH: Bromus carinatus, California [no image]. GH 00361739. tation 1: Festuca pacifica. GH 341875. K: B. carinatus, California (SL). K 000913857. Coulter 763. Coulter 766. TCD: Festuca, California (PL). TCD! Note: image identified by J.T. GH: Bromus carinatus, California [no image]. Annotation 1: Bromus car- Columbus. inatus var. hookerianus (Thurber) Shear. GH 00361757. Coulter 763 bis. †Cenchrus sp. TCD: no name], California (PL). TCD! Note: image identified by J.T. Coulter 758. Columbus. TCD: Cenchrus tribuloides?, California (PL). TCD! Hilaria rigida (Thurb.) Benth. ex Scribn. †Digitaria ciliaris (Retz.) Koeler var. ciliaris Coulter 752. Coulter s.n. TCD: [no name], California (PL). TCD! GH: Digitaria ciliaris, California [no image]. Annotation 1: Digitaria GH: Hilaria rigida, California [no image]. GH 00341005. ciliaris by Webster. GH 00355505. Hordeum depressum (Scribn. & J.G. Sm.) Rydb. †Digitaria sanguinalis (L.) Scop. Coulter 756. Coulter 786. GH: Hordeum depressum, California [no image]. Annotation 1: Hordeum TCD: Digi. sang., California (PL). TCD! depressum by Jacobsen in 1993. GH 00365196. Distichlis spicata (L.) Greene Hordeum jubatum L. subsp. jubatum Coulter 779. Coulter 750. TCD: Poa, Brizopyrum, California (PL). TCD! Note: image identified as TCD: Hordeum jubatum, California (PL). TCD! Distichlis spicata by J.T. Columbus. BRU: [no name], California (PL). PBRU 00026412. Hordeum pusillum Nutt. GH: Distichlis spicata var. nana Beetle, California [no image]. GH Coulter 756. 00355665. TCD: Hordeum, California (PL). Written on sheet: Hordeum pusillum. †Echinochloa colona (L.) Link TCD! Coulter 794. Leptochloa fusca (L.) Kunth subsp. uninervia (J.Presl)N.Snow TCD: Panicum crus-galli, California (PL). TCD! Note: image identified Coulter 776. by J.T. Columbus. GH: Leptochloa fusca subsp. uninervia, California [no image]. Annota- Elymus multisetus (J.G. Sm.) Burtt Davy tion 1: Leptochloa fusca subsp. uninervia by Snow in 1997. GH 355751. Coulter 750. GH: Elymus multisetus, California [no image]. Annotation 1: Sitanion Leptochloa sp. jubatum by Wilson in 1956. GH 00365089. Coulter 776. TCD: Leptochloa, California (PL). Written on sheet: Diplachne obtusi- Elymus sp. flora, Leptochloa dubia H.B.K. TCD! Coulter 748. Coulter 785. TCD: Elymus, California (PL). TCD! TCD: Leptochloa, California (PL). TCD! Coulter 755. TCD: Elymus, California (PL). TCD! Melica californica Scribn. Coulter 781. Eragrostis pectinacea (Michx.) Nees var. pectinacea TCD: Melica, California (PL). Written on sheet: Melica speciosa Much. Coulter 773. [? illegible] nov. sp. TCD! Note: image identified by J.T. Columbus. 64 Wallace ALISO

GH: Melica californica, California [no image]. Annotation 1: Melica cal- †Setaria pumila (Poir.) Roem. & Schult. subsp. pumila ifornica by Boyle in 1943. GH 360266. Coulter 749. TCD: Setaria glauca, California (SL). TCD! Melica imperfecta Trin. Coulter 771. †Setaria viridis (L.) P. Beauv. TCD: [no name], California (PL). Note on label: same as 780. Written Coulter 791. Image on sheet: Reboulea nov. sp. R. californica (Muns). TCD! Note: filed in TCD: Setaria imberbis, California (PL). TCD! Note: image identified by Reboulea folder. J.T. Columbus. Coulter 780. TCD: Diplachne same as 771 (PL). Written on sheet: Reboulea nov. sp. Sphenopholis sp. say [sic] R. californica. TCD! Coulter 770. GH: Melica imperfecta, California [no image]. Annotation 1: Melica im- TCD: [no name], California (PL). TCD! Note: filed in Reboulea folder. perfecta by Boyle in 1943. GH 00360377. Stipa sp. Muhlenbergia microsperma (DC.) Trin. Coulter 765. Coulter 761. TCD: Stipa, California (PL). TCD! TCD: Muhlenbergia/Indochloa, California (PL). TCD! Note: image identified by J.T. Columbus. †Triticum aestivum L. GH: Muhlenbergia microsperma, California [no image]. Annotation 1: Coulter 747. Muhlenbergia microsperma by Peterson in 1988. GH 00355850. TCD: Triticum, California (PL). TCD! Coulter s.n. †Paspalum dilatatum Poir. TCD: Triticum, California (PL). TCD! Coulter 788. TCD: [no name], California (PL). TCD! potamogetonaceae Paspalum sp. Stuckenia pectinata (L.) Börner Coulter 790. Coulter 719. TCD: [no name], California (PL). TCD! Note: image identified by J.T. TCD: Potamogeton pectinata, California (PL). TCD! Columbus.

Phalaris californica Hook. & Arn. ruscaceae Coulter 754. Maianthemum dilatatum (A.W. Wood) A. Nelson & J.F. Macbr. TCD: [no name], California (PL). Annotation 1: Phalaris californica Coulter 728. Hook. & Arn. by Baldwin in 1992. TCD! BRU: Maianthemum, California (PL). Annotation 1: Herbarium Ol- GH: Phalaris californica, California [no image]. Annotation 1: Phalaris neyanum. PBRU 00009084. californica by Anderson in 1959. GH 00360496. Coulter 729. Poa bolanderi Vasey BRU: Maianthemum, California (PL). Annotation 1: Herbarium Ol- Coulter 783. neyanum. PBRU 00009087. GH: Poa bolanderi, California [no image]. GH 00360669. themidaceae Poa douglasii Nees Bloomeria crocea (Torr.) Coville var. Coulter 753. TCD: Poa douglasii, Brizopyrum, California (PL). TCD! Coulter 741. TCD TCD BRU: [no name], California (PL). Annotated: Herbarium Olneyanum Poa : [no name], California (PL). ! douglasii PBRU (Hook.). 00028032. Brodiaea minor (Benth.) S. Watson GH: Poa douglasii, California [no image]. Annotation 1: Poa douglasii Coulter 742. by Marsh in 1942. GH 00360701. TCD: Brodiaea, California (PL). TCD! GH: Poa douglasii, California [no image]. Annotation 1: Poa douglasii K: Brodiaea minor, California [no image]. Annotation 1: Brodiaea minor by Marsh in 1942. GH 00360706. by Niehaus in 1968. K 000457455. NY: Brizopyrum douglasii H. & A., California (PL). Annotation 1: Poa douglasii Nees by Marsh in 1942. NY 01789773. Dichelostemma capitatum (Benth.) Alph. Wood Coulter 743. Poa secunda J. Presl subsp. TCD: [no name], California (PL). Annotation 1: ISOSYNTYPE Brodi- Coulter 782. aea capitata Benth. by G. Wallace in 2017. TCD! GH: Poa secunda, California [no image]. GH 00360892. BRU: Brodiaea congesta, California (PL). Annotation 1: Herbarium Ol- Poa sp. neyanum. PBRU 00008800. Coulter 777. TCD: Poa, California (PL). TCD! typhaceae Coulter 783. TCD: Poa, California (PL). TCD! Typha angustifolium L. Coulter 816. †Polypogon viridis (Gouan) Breistr. TCD: Typha angustifolia [sic], California (PL). TCD! Coulter 775. TCD: Agrostis verticillata?, California (PL). TCD! Note: image identi- zanichelliaceae fied by J.T. Columbus. Coulter 778. Zanichellia palustris L. TCD: Agrostis alba, California (PL). TCD! Note: image identified by Coulter 815. J.T. Columbus. TCD: Zanichellia, California (PL). TCD! VOLUME 37(1–2) Plantae Coulterianae 65

BRYOPHYTES BM: Orthotr[ichum](nr.pallens & roellii) California (SL). BM mosses 000982229. TYPE of Orthotrichum coulteri Mitt. Filed as Orthotrichum tenellum var. coulteri. brachytheciaceae FH: Orthotrichum coulteri, California [no image]. Lectotype. Annotation Brachythecium sp. 1: Orthotrichum cylindrocarpum Lesq. by Medina and Lara in 2011. FH Coulter 837. 00301491. TCD: [no name], California (SL). Written on sheet: Brachythecium as- Coulter 840. perimum [sic]. TCD! (This specimen does not seem to fit this species de- TCD: [no name], California (SL). Annotation 1: Identified as Or- termination according to L. Ellis [BM] October 2018), and other experts, thotrichum tenellum by Daniel Kelly (TCD) 20 September 2018. Filed D. Kelly [TCD], J. Shevock and D. Toren [CAS] are not in agreement as as ISOLECTOTYPE Orthotrichum coulteri. TCD! to a species name). BM: Orthot[richum], California (SL). Note on label: (see 839). BM 000982230. Note: Identified as TYPE of Orthotrichum coulteri Mitt. Homalothecium aureum (Spruce) H. Rob. Filed as Orthotrichum tenellum var. coulteri. Coulter 836. FH: Orthotrichum cylindrocarpum, California [no image]. Lectotype. TCD: [no name], California (SL). Annotation 1: most likely Homalothe- Annotated: Orthotrichum cylindrocarpum Lesq. By Medina and Lara in cium aureum (Spruce) H. Rob. by L. Ellis (BM) October 2018. TCD! 2011. FH 00301491. FH: Hypnum nuttallii, California. 1831 [no image]. FH 01139016. Note: Coulter 839, 840 was designated the LECTOTYPE of Or- thotrichum coulteri Mitt. by Vitt in 1973.

bryaceae Orthotrichum lyellii Hook. & Taylor Ptychostomum sp. Coulter 838. Coulter 832. TCD: [no name], California (PL). Annotation 1: Orthotrichum lyellii by TCD: [no name], California (SL). Annotation 1: as 832-II by D. Kelly Daniel Kelly (TCD) 24 September 2018. TCD! (TCD) to differentiate this from 832 identified as Orthothrichum. It is ei- FH: Orthotrichum lyellii, California [no image]. FH 00913628. ther Bryum or a related taxon by D. Kelly (TCD, October 2018). Seems Coulter s.n. to belong to the segregate of Bryum now widely recognized as Ptychos- NY: [no name], California [no image]. Annotation 1: Orthotrichum lyellii tomum (L. Ellis [BM] October 2018). TCD! Hooker & Taylor by Vitt in 1970. NY 453283.

daltoniaceae polytrichaceae Daltonia splachnoides (Smith) Hook. & Taylor Polytrichum juniperinum Hedw. Coulter s.n. Coulter 835. TCD: Daltonia heteromala, [no location] TCD! TCD: [no name], California (PL). Annotation 1: Polytrichum juniper- inum by Daniel Kelly (TCD) 6 October 2018. TCD! FH: Polytrichum juniperinum, California [no image]. FH 00913303. ditrichaceae

Ceratodon purpureus (Hedw.) Brid. pottiaceae Coulter 834. TCD: [no name], California (SL). Annotation 1: Ceratodon purpureus Timmiella anomala (Bruch & Schimp.) Limpr. (Hedw.) Brid. by L. Ellis (BM) October 2018. TCD! Coulter 833. TCD: [no name], California (SL). Annotation 1: Timmiella anomala by funariaceae Daniel Kelly (TCD) 24 September 2018. TCD! TCD: [no name], California (SL). Annotation 1: Timmiella anomala by Funaria hygrometrica Hedw. Daniel Kelly (TCD) 24 September 2018. TCD! Coulter 831. FH: Barbula anomala, California [no image]. FH 01137572. TCD: [no name], California (SL). Annotation 1: Funaria hygrometrica by Daniel Kelly (TCD) 24 September 2018. TCD! FH: Funaria hygrometrica, California [no image]. FH 01138623. liverworts aytoniaceae leptodontaceae Asterella bolanderi (Austin) Underw. Alsia californica (Hook. & Arn.) Sull. Coulter 841. Coulter s.n. TCD: Note: Coulter 841 two mounted packets at TCD. The numbers NY: [no name], California [no image]. NY 845788. were amended to 841a and 841b by D. Kelly in 2018. BM: Asterella tenella [no image]. BM 000676410. NY: Asterella bolanderi, California [no image]. Specimen from NY cited orthotrichaceae by Evans (1920) in Contributions from the U.S. National Herbarium 20: Orthotrichum alpestre Hornsch. ex Bruch & Schimp. 304. NY 00266108. Note: no collector given in NY database but fits Coul- Coulter 832. ter’s numbers. TCD: Orthotrichum watsonii James: Sull. Icones, California (SL). Anno- Coulter 841a. tation 1: Orthotrichum by Daniel Kelly (TCD) October 10, 2018. TCD! TCD: Fimbraria, California (SL). TCD! Coulter 841b. Orthotrichum cylindrocarpum Lesq. TCD: [no name?] (SL). Identified as Asterella bolanderi (Austin) Un- Coulter 839. derw. By Daniel Kelly (TCD) 10 October 2018, and confirmed by D.G. TCD: [no name], California (SL). Annotation 1: Orthotrichum tenellum Long (E) 20 November 2018. TCD! by Daniel Kelly (TCD) 20 September 2018. To be filed as ISOLECTO- Coulter s.n. TYPE Orthotrichum coulteri. TCD! FH: Fimbraria violacea, California [no image]. FH 1137041 66 Wallace ALISO

appendix ii TCD: [no name]. Written on sheet: Macrocystis pyrifera, California. TCD! coulter’s california marine algae exsiccata S: Macrocystis pyrifera, California [no image]. S-Algae A 11260. This list is arranged alphabetically by phylum, family, and genus. The format of the entries follows that in Appendix I. There are few attributions of type status among Coulter’s California marine algae. Most specimens sargassaceae lack annotations and there are no exsiccata numbers assigned to most of Stephanocystis osmundacea (Turner) Trevis. the specimens which would aid in the identification of duplicates and type Coulter s.n. material. Several of these specimens are likely misidentified and/or repre- TCD: Halidrys osmundacea Harv., Monterey, Dr. Coulter (v.s. in Herb. sent taxa not known to occur in California. A special acknowledgement and T.C.D.) (Harvey 1852, p. 64). thanks are due Kathy Ann Miller (KAM), Curator of Algae, UC Berkeley TCD: Cystoseira expansa C. Agardh, at Monterey, California, Mr. Dou- for her suggestions and corrections to this appendix. My search of the ma- glas; Dr. Coulter (v.s. in Herb. T.C.D.) (Harvey 1852, p. 66). rine algae collections was not complete. Some additional algal taxa based BM: Cystoseira osmundacea (Turner) J. Agardh, California (PL). An- on references to Coulter’s Californian specimens are included with citations notation 1: Isotype of Cystoseira douglasii Harv. by Silva in 1974. BM to Harvey (1852, 1853, 1858). 000563385. Note: BM database incorrectly lists S.D. Coulter as collector. BRU: Halidrys osmundacea H., California (PL). ABRU 00000229. BRU: Cystoseira douglasii Harvey., California (PL). Annotation 1: CHLOROPHYTA Herbarium Olneyanum. ABRU 00000093. cladophoraceae L: Cystoseira douglasii Harv., California (PL). Annotation 1: Cystoseira osmundacea. L 3995662. L database incorrectly lists J.W. Coulter as the Cladophora microcladioides Collins collector. Coulter s.n. L: Halidrys osmundacea Harv., California (PL). Annotation 1: Cysto- TCD: Cladophora laetevirens Dillw., California, Dr. Coulter (v.v.) (Har- seira osmundacea. L 3995664. Note: L database incorrectly lists J.W. vey 1858, p. 82). Coulter as the collector. NY: Cystoseira osmundacea (W.B. Turner) C. Agardh, California. NY codiaceae 02229663. NY: Halidrys osmundacea H., California (PL). NY 02243131. Codium fragile (Suringar) Har. subsp. californicum, (J. Agardh) Maggs PC: Halidrys osmundacea Harv., California (PL). PC 0044669. & Kelly PC: Halidrys osmundacea Harv., Fucus osmundaceus Turn., California Coulter s.n. (PL). Annotation 1: Halidrys osmundacea Harv. PC 0535424. TCD: Written on sheet: Codium tomentosum, California. Annotation 1: S: Stephanocystis osmundacea (Turner) Trevis., California [no image]. Codium mucronatum by S.D. Colton. Annotation 2: Codium fragile (Sur.) S-Algae A 3954. Har. by P. Silva in 1955. TCD! S: Stephanocystis osmundacea (Turner) Trevis., California [no image]. L: Codium tomentosum Agardh, California (PL). Annotation 1: Codium S-Algae A 14392. fragile (Sur.) Har. by P.C. Silva in 1959. L 4121113. Note: L database incorrectly lists J.M. Coulter as collector. NY: Codium tomentosum, California (PL). Annotation 1: Codium fragile lessoniaceae subsp. californicum (J. Ag.) Silva by Silva in 1960. NY 02130970. Egregia menziesii (Turner) Aresch. Codium sp. Coulter s.n. Coulter s.n. TCD: [no name]. Written on sheet: Phyllospora menziesii, California. BRU: Codium tomentosum, California (PL). ABRU 00000623. TCD! PC: Codium tomentosum, California (PL). Annotation 1: “Both probably Codium vermilara (Olivi) Delle Chiaje (NOT from California!)” det. P.C. Silva June 1957. PC 0037400. RHODOPHYTA ceramiaceae ulvaceae Ceramium sp. Coulter s.n. Ulva taeniata (Setch.) Setch. N.L. Gardner TCD: [no name]. Written on sheet: Ceramium coulteri H., California. Coulter s.n. TCD! TCD: Ulva fasciata Delile, California, Dr. Coulter …(v.v.) (Harvey 1858, BRU: Ceramium obsoletum, California (PL). ABRU 00001141. p. 59). Microcladia coulteri Harv. Coulter s.n. OCHROPHYTA fucaceae TCD: Microcladia coulteri H., California. Annotation 1: Type. TCD 0012510! Silvetia compressa (J. Agardh) E. Serrão, T.O. Cho, S.M. Boo & Brawley UC: Microcladia coulteri Harv., California, Dr. Coulter. Ex. Herb. W.H. Coulter s.n. Harvey, Trinity College, Dublin, Ireland. Annotation 1: Type material Mi- L: Fucus fastigiatus J. Agardh, California (PL). Annotation 1: Pelvetia crocladia coulteri Harv. AHFH 53940 in UC. fastigiata. L 4013729. Note: L database incorrectly lists J.M. Coulter as collector. S: Silvetia compressa, California [no image]. S-Algae A 13665. delesseriaceae Cryptopleura ruprechtiana (J. Agardh) Kylin laminariaceae Coulter s.n. TCD: Botryoglossum platycarpum Kütz., Monterey, California, Mr. Macrocystis pyrifera (L.) C. Agardh Douglas!, Dr. Coulter!, Dr. Sinclair!, Capt. Wilkes! (v.v.) (Harvey 1853, Coulter s.n. p. 100). VOLUME 37(1–2) Plantae Coulterianae 67

NY: Hymenena fimbriata, California, N. B.-A. 2: 101, J. Ag. 2: 674 (PL). PC: Gelidium lanceolatum Harv., California (PL). Annotation 1: Ge- Written on sheet: Nitrophyllum ruprechtiana. NY 02184796. lidium lanceolatum California. Ex Hook. Herb. Annotation 1: Prionitis lanceolata (collected with type material from Monterey California det. Cryptopleura sp. I.A. Abbott [?]. PC 0048777. Coulter s.n. PC: Delesseria platycarpa var. an sp. nov.? vidi Postels & Ruppt., Cali- gigartinaceae fornia (PL). PC 0048318. Chondracanthus canaliculatus (Harv.) Guiry Hymenena sp. Coulter 856. Coulter s.n. BRU: Coulter 856. Gigartina canaliculata, California (PL). Annotation BRU: Hymenena, California (PL). ABRU 00001546. 1: Herbarium Olneyanum. ABRU 00001427. Note: in BRU database as canaliculata. F: Gigartina canaliculata, Algae Californicae. F C05000997F. endocladiaceae Coulter s.n. Endocladia muricata (Endl.) J. Agardh TCD: [no name]. Written on sheet: Gigartina canaliculata Harv. In type Coulter s.n. folder TCD 00012212! TCD: Endocladia muricata J. Ag., California, Beechey! Coulter! Wos- FH: Gigartina canaliculata, California. FH 907423. nessensky! Capt. Pike! (v.s. in Herb. T.C.D.) (Harvey 1853, p. 182). NY: Gigartina canaliculata, California [no image]. NY 02209943. PC: Gigartina canaliculata Harv. in Beechey, G. lindleyana J. Ag. in Lin- Gloiopeltis furcata (Postels & Rupr.) J. Agardh naea, California (PL). Filed as Gigartina acicularis (Roth) J.V. Lamour. Coulter s.n. PC 0048848. PC: Dumontia saccata, California (Coulter) amiciss.Harvey(SL).PC 0048516. Chondracanthus corymbiferus (Kütz.) Guiry Coulter s.n. L: Iridaea papillata, Mastocarpus corymbiferus Kütz. Tab. Phyc. xvii, gelidiaceae 46 (PL). Annotation 1: TYPE Mastocarpus corymbiferus Kütz. Anno- tation 2: Mastocarpus corymbiferus Kützing TYPE. 1847 Bot. Zeit. 5: Gelidium coulteri Harv. 24, 1867 Tab. Phyc. xvii, 46; = Chondracanthus corymbiferus (Kützing) Coulter 872. Guiry 1993 Hydrobiologia 260/261: 115 by M. Hommersand in 1997. BRU: Gel. cartilagineum, California (PL). ABRU 00001417. Annotation 3: TYPE. L 4051094. BRU: Gelidium corneum, California (PL). ABRU 00001397. PC: Iridaea papillata Grev., California (PL). PC 0059302. F: Gelidium corneum, Algae Californicae. Annotation 1: Gelidium coul- teri by D.C. Eaton. F C0502757F. Note: F database incorrectly lists J.M. Chondracanthus harveyanus (Kütz.) Guiry Coulter as collector. Coulter s.n. F: Gelidium corneum, Algae Californicae. Annotation 1: Gelidium coul- TCD: Gigartina microphylla Harv., Monterey, California, Douglas, Dr. teri by D.C. Eaton. F C0502680F. Note: F database incorrectly lists J.M. Coulter (v.s. in Herb. T.C.D.) (Harvey 1853, p. 176). Coulter as collector. BRU: Iridaea microphylla, California (PL). Annotation 1: Herbarium Ol- YU: Gelidium corneum, Algae Californicae. Annotation 1: Gelidium neyanum, Gigartina microphylla. ABRU 00001436. coulteri by Eaton. YU 245966. L: Iridaea microphylla Harv. Mastocarpus microphylla, California (PL). Coulter s.n. Annotation 1: TYPE Mastocarpus harveyanus Kützing. Annotation 2: TCD: [no name]. Written on sheet: Gelidium coulteri H., California. Herbarium Kützing Mastocarpus harveyanus Kg. Annotation 3: Gi- Type. TCD 0011891! gartina harveyanus (Kützing) Setchell & Gardner. Type of Mastocarpus BM: Gelidium coulteri, California (SL). Type. BM 000610919. harveyanus Kützing. 1847 Bot. Zeit. 5: 24, nom. nud., 1849 Sp. Alg. P. L: Gelidium polyacanthum Kg., California (PL). Annotation 1: Herbar- 734, 1867 Tab. Phyc. xvii, 45; = Chondracanthus harveyanus (Kützing) ium Kützing Gelidium polyacanthum Kg. Tab. Phyc. xviii, 64, California. Guiry 1993 Hydrobiologia 260/261: 115 by M. Hommersand in 1997. L Annotation 2: Gelidium coulteri Harv. L 4044574. Note: L database in- 4051095. correctly lists J.M. Coulter as collector. NY: Mastocarpus harveyanus Kg., Iridaea [crossed out] Gigartina mi- NY: Gelidium cartilagineum (SL). Annotation card: Gelidium coulteri crophylla Harv. N.A.B. [Nereis Boreali-Americana] II, p. 176, California. Harv. NY 02226327. Note: NY database incorrectly lists J.M. Coulter as PL. Written on sheet: Gigartina microphylla Harv. J. Ag. Vol. III, p. 200. the collector. NY 02196623. NY: Gelidium cartilagineum, California (PL). Written on sheet: Gelid- PC: Iridaea microphylla Harv. Rhodomela microphylla H. in Beechey, ium purpurascens N.L. Gardner. NY 02209478. California (PL). PC 0059296. PC: Gelidium corneum 3 varieties, California (PL). PC 0048651. UC: Gelidium coulteri Harv., California. Annotation 1: Type material Ge- Chondracanthus spinosus (Kütz.) Guiry lidium coulteri Harv. AHFH 53929 in UC. Coulter s.n. TCD: [no name]. Written on sheet: 1847 Mastocarpus spinosus, 1849 Prionitis lanceolata (Harv.) Harv. Gigartina ornithorhynchos, California. TCD! Coulter 871. L: Iridaea stiriata, Mastocarpus stiriata Kg., California (PL). Annota- BM: Gelid. lanceolatum Harv. in Beechey. Monterey (SL). BM tion 1: TYPE Gigartina spinosa (Kütz.) J.G. Ag. Annotation 2: TYPE 000530327. Mastocarpus spinosus Kützing. Annotated: Herbarium Kützing Masto- BRU: Gelidium lanceolatum Harv., California (PL). Annotation 1: carpus spinosus Kg. Annotation 3: Gigartina spinosa (Kützing) J. Agardh Herbarium Olneyanum. ABRU 00001421. TYPE. This is the type of Mastocarpus spinosus Kützing 1847 Bot. Zeit. Coulter s.n. 5: 24, 1867 Tab. Phyc. xvii, pl. 47. = Chondracanthus spinosus (Kützing) TCD: Prionitis lanceolata (Harv.) Harv., California, Douglas, Dr. Coulter Guiry 1993 Hydrobiologia 260/261: 115 by M. Hommersand in 1997. L …(v.s. in Herb. T.C.D.) (Harvey 1853, p. 197). 4051117. L: Gelidium lanceolatum Harv., California (PL). Annotation 1: Herbar- NY: Gigartina spinosa H. Mastocarpus spinosus Kg. [sic]. N.A.B. 2 p. ium Kützing, Gelidium lanceolatum Harv. Tab. Phyc. xviii, 48. Note: L 177, California. (PL). Written on sheet: Gigartina spinosa J. Ag. vol. III, database incorrectly lists J.M. Coulter as collector. L 4073630. p. 200. NY 0216876. 68 Wallace ALISO

Mastocarpus sp. plocamiaceae Coulter 854. Plocamium pacificum Kylin PC: Chondrus mamillosus?, California (PL). PC 0047625. Coulter s.n. Coulter s.n. TCD: Gelidium cartilagineum Grev., California, D. Douglas, Dr. Coulter; TCD: [no name]. Written on sheet: Gigartina radula Harv.Ner.Bor.- Lieut. Wood (v.v.) (Harvey 1853, p. 117). Amer. TCD! NY: Gelidium cartilagineum, California. NY 02209478. TCD: [no name]. Written on sheet: Gigartina radula New. Bor. Amer., Mastocarpus corymbiferus, Iridaea papillata Harvey in Beechey not of Ag. TCD! TCD: Gigartina mammillosa J. Ag., California, Dr. Coulter! (v.v.) (Har- rhodomelaceae vey 1853, p. 175). NY: Mastocarpus corymbiferus Kg., M. papillatus Harv., California [no Laurencia pacifica Kylin image]. NY 02196768. Coulter s.n. TCD: Laurencia virgata J. Ag., Monterey, California, Dr. Coulter (v.v.) Mazzaella affinis (Harv.) Fredericq (Harvey 1853, p. 71). Coulter s.n. TCD: Chondrus affinis Harv., California, Douglas, Coulter (v.s. in Herb. Neorhodomela larix (Turner) Masuda T.C.D.) (Harvey 1853, p. 181). Coulter s.n. L: Chondrus affinis Harvey in Beechey, California (PL). Annotated: TCD: Rhodomela larix Ag., …to Monterey, Coulter (v.s. in Herb. Chondrus affinis Harvey Kützing 1867 Tab. Phyc. xvii, p. 53, c. d. This T.C.D.) (Harvey 1853, p. 24). is probably not type material as it was collected by Dr. Coulter and not PC: Rhodomela larix, California (PL). PC 0061854. by Douglas. = Mazzaella affinis (Harvey) Fredericq 1993 Hydrobiologia 260/261: 110 by M. Hommersand in 1997. L 4081350. Note: L database Odonthalia floccosa (Esper) Falkenb. incorrectly lists J.M. Coulter as the collector. Coulter s.n. PC: Chondrus affinis Harv. in Beechey, California (PL). Annotated: TCD: Rhodomela floccosa Ag., …Monterey, Dr. Coulter …(v.s. in Herb. Chondrus affinis Harv. California. Ex Herb. Hook. PC 0047599. T.C.D.) (Harvey 1853, p. 25).

Mazzaella sp. Osmundea spectabilis (Postels & Rupr.) K.W. Nam Coulter 851. Coulter s.n. PC: Iridaea?, California (PL). PC 0059318. TCD: Laurencia pinnatifida Lamour, Monterey, California, D. Douglas, Coulter s.n. Dr. Coulter (v.v.) (Harvey 1853, p. 71). TCD: [no name]. Written on sheet: Iridaea laminarioides, California. BRU: Laurencia pinnatifida, California (PL). Annotation 1: Herbarium TCD! Olneyanum. ABRU 00001639. BRU: Iridaea, California (PL). ABRU 00001567. NY: Laurencia pinnatifida, California. Stamp: The F.S. Collins Herbar- BRU: Gigartina stiriata (Turner) J. Agard, California. ABRU 00001450. ium presented to the New York Botanical Garden by N.L. Britton 1922. F: Iridaea (I. laminaroides or minor det. by Eaton). Algae Californicae. Stamp: Boston Society of Natural History. NY 02182197. F C0506257F. L: Laurencia pinnatifida, Laurencia californica (Kütz.), Tab. Phyc. xv, NY: Iridaea, California (PL). Filed as: Iridaea laminarioides Bory. NY 67, California. (PL). Annotation 1: Osmundea pinnatifida. L 4070014. 02215517. Note: L database incorrectly lists J.M. Coulter as the collector. PC: Iridaea stiriata, California (PL). PC 0059311M. PC: Laurencia pinnatifida, California (PL). PC 0059658. UC: [no name] California, Dr. Coulter. Annotation 1: Algae from the collections of W.H. Harvey in the herbarium of Trinity College, Dublin described in Neris [Nereis] Boreali-Americana. Written on sheet: Un- lomentariaceae labelled but undoubtedly Laurencia pinnatifida N.B.-A. 2: 70, 71. UC Neogastroclonium subarticulatum (Turner) L. Le Gall, J. Dalen & G. 68345. Saunders Coulter 858. Polysiphonia sp. PC: Chylocladia coulteri Harv. MS., California (PL). PC 0059866. Coulter 865. Coulter s.n. BRU: Polysiphonia, California (PL). Annotated: Herbarium Olneyanum. TCD: Lomentaria ovalis Endl., β at Monterey, Dr. Coulter (v.s. in Herb. ABRU 00001954. T.C.D.) (Harvey 1853, p. 78). F: Polysiphonia [crossed out] Rhodomela. Algae Californicae. F C0511173F. YU: Polysiphonia [crossed out] Rhodomela? Algae Californicae. YU 087616. palmariaceae Coulter s.n. Halosaccion glandiforme (S.G. Gmel.) Rupr. TCD: [no name]. Written on sheet: Polysiphonia urceolata, California. Coulter s.n. TCD! TCD: Halosaccion fucicola Post. & Rupr., on Laurencia at Monterey, Dr. TCD: Polysiphonia urceolata Grev., β at Monterey, California, Dr. Coul- Coulter (v.v. in Herb. T.C.D.) (Harvey 1853, p. 194). ter (v.v.) (Harvey 1853, p. 32). S: Halosaccion glandiforme, California [no image]. S-Algae: A6776. NY: Polysiphonia, California (PL). NY 02221950. PC: Polysiphonia urceolata var. β, California (SL). Filed as P. tenella (C. Agardh) Moris & De Notaris. PC 0061480. phyllophoraceae PC: Polysiphonia, California (PL). PC 0061608. UC: Polysiphonia urceolata β. California [no label]. Annotation 1: Al- Mastocarpus papillatus (C. Agardh) Kütz. gae from the collections of W.H. Harvey in the herbarium of Trinity Col- Coulter s.n. lege, Dublin described in Neris [Nereis] Boreali-Americana. Annotation BRU: Gigartina papillata, California (PL). Annotation 1: Herbarium Ol- 2: Polysiphonia pasifera var. determinata Hollenberg G.J.H. Dec. 1941. neyanum. ABRU 00001439. UC 68396. VOLUME 37(1–2) Plantae Coulterianae 69

solieriaceae mixed collection of the two subspecies and their intermediates).” Thomp- son 1988. Syst. Bot. Monogr. 22: 92 “Bentham listed both Coulter 599 Sarcodiotheca gaudichaudii (Mont.) P.W. Gabrielson (GH! K!) and Nuttall’s collection from “St. Diego. U-Cal” (K! PH!) in Coulter 857. the protologue for A. nuttallianum, and indicated he saw the specimens BM: Rhabdonia coulteri barren frond of …California (SL). BM in Hooker’s herbarium, now at K (the two collections are mounted on the 000054003. same sheet). Specimens of these two collections are therefore syntypes.” Coulter 861. NOTE: Thompson misquoted Bentham, see above. He also miscited the BM: Laurencia sp. nov. Rhabdonia coulteri, California (SL). Note: BM authority as Bentham in de Candolle: A. De Candolle is the author of the database lists J.M. Coulter as collector in error. BM 000937085. Addenda et Corrigenda section of the volume that starts on p. 587 and Coulter s.n. includes p. 591. TCD: Rhabdonia coulteri Harv., Monterey, California, Dr. Coulter …(v.s. in Herb. T.C.D.) (Harvey 1853, p. 154). Armeria andina Poep. var. californica Boiss. De Candolle Prodromus 12: PC: Rhabdonia coulteri, California [no image]. PC 0061815. 682. 1848. “In California (Dr. Coulter in h. Hook.!).” = Armeria maritima (Mill.) Willd. subsp. californica (Boiss.) A.E. Porsild. appendix iii TCD!: Coulter 577. Annotated: Isotype by G.D. Wallace November K new taxa based on coulter specimens from california 2017. Also, at (presumed type material). and mexico Astragalus coulteri Benth. in Herb. cit. [this means same as Phaca crota- With emphasis on specimens at TCD lariae that is just above this species on p. 307 = Herb. Coll. SS. Trin., Dubl.]. Plantas Hartwegianas p. 307. 1848. “Juxta Monterey legit Coul- The entries here begin with the basionyms and author name(s), publica- ter.” = Astragalus lentiginosus Douglas var. borreganus Jones. tion citation and relevant specimen citation from the protologue. This is fol- TCD!: Coulter 431. Annotated: Type material by G.D. Wallace Novem- lowed by the current name and author(s) of the combination if it is different ber 2017. Also, at K. Barneby 1964 Mem. NYBG 13(2): 954: “Holotypus, and a note of the taxon’s range if it is not known to occur in California. Coul- Coulter 431, labeled “Monterey” but collected probably near the mouth ter’s specimen at TCD is indicated with an exsiccata number or s.n. This is of the Gila River in 1832, K (Herb. Bth.)!” followed by reference to any annotations on the TCD specimen. Coulter’s TCD exsiccata are followed by the acronyms of additional herbaria with on- Baileya multiradiata Harv. & A. Gray ex Torr. In Emory: Notes of a Mil- line catalogues known to hold the same Coulter exsiccata material and any itary Reconnaissance from Fort Leavenworth …144. 1848. “California, note of type status provided by the institution. The type status of Coulter Coulter.” specimens at several institutions has not been recorded by annotation of the TCD!: Coulter 326. Annotated: Isolectotype by G.D. Wallace August specimens or in the databases. In a few instances additional references and 2016. Also, at GH (type), K. comments are included. I encourage specialists in these groups to compare all of the type material available and make the appropriate designations of Baileya pauciradiata Harv. & A. Gray ex A. Gray. Mem. Amer. Acad. Arts type status. and Sciences II, 4: 105. 1849. “California, Coulter”. TCD!: Coulter 312 Baileya multiradiata. Annotated: Baileya pauciradi- Acarphaea artemisaefolia Harv. & A. Gray ex A. Gray. Mem. Amer. Acad. ata H. & Gr., Torr. & Gray, Fl. N. Amer. Also, at GH (holotype), NY. Arts and Sciences II, 4: 98–99. 1849. “California Coulter.” = Chaenactis artemisiifolia (Harv. & A. Gray ex A. Gray) A. Gray. Baileya pleniradiata Harv. & A. Gray ex A. Gray. Mem. Amer. Acad. Arts TCD!: Coulter 313. Annotated: Isotype of Acarphaea artemisaefolia by and Sciences II, 4: 105–106. 1849. “California, Coulter.” G.D. Wallace August 2016. Also, at GH (possible type), K. TCD!: Coulter 312 Baileya multiradiata [likely source of fragment at Amsinckia microcarpa Greene. Erythea 2: 191. 1894. “Plant collected long GH]. Also, Lectotype fragment at GH. ago by Dr. Coulter, probably in the southern part of California, but possi- Berginia virgata Harv. ex Benth. & Hook.f. Gen. Pl. 2: 1097. 1876. “Species bly in Mexico.” [no collections cited]. = Amsinckia spectabilis Fisch. & 1, B. virgata, Harv., California incola. Coulter n. 603.” [Note the genus C.A. Mey. var. microcarpa (Greene) Jeps. & Hoover. Berginia was described on the preceding page 1096 “Berginia, Harv. in TCD!: Coulter 497. Annotated: Type material of Amsinckia microcarpa Herb. Trin. Coll. Dubl ….”] = Holographis virgata (Harv. ex Benth. & by G.D. Wallace November 2017. Also, at GH (type), K, NY. Hook.f.) T.F. Daniel. Known from Mexico, Sonora. Antirrhinum confertiflorum Benth. ex A. DC. Prodromus 10: 592. 1846. TCD!: Coulter 603. Annotated: Isotype by Tom Daniel in 2015. Also, at “ …in California, (Coulter!)” “(v. in Herb. Hook.).” = Mohavea conferti- GH (isotype), K (holotype and two isotypes). flora (Benth. ex A. DC.) A. Heller. Brickellia coulteri A. Gray. Smithsonian Contr. Knowl. 3, article 5. Plantae TCD!: Coulter 616. Annotated: Isotype by G.D. Wallace August 2016. Wrightianae Texano-Neo-Mexicanae part 1, pp. 1–146, t. 1–10. March Also, at K (presumed type material). 1852. p. 86. “California, (Coulter 293), in Herb. Hook.” = Brickellia coul- Antirrhinum coulterianum Benth. ex A. DC. Prodromus 10: 592. 1846. teri A. Gray var. coulteri. Known from Arizona, Texas and Mexico. “ …in California (Coulter!)” and “(v. in Herb. Hook.).” = Antirrhinum TCD!: Coulter 293. Annotated: Isotype by G.D. Wallace November coulterianum Benth. ex A. DC. 2017. Also, at GH (holotype), K. TCD!: Coulter 607. Annotated: Isotype by G.D. Wallace November 2017. Also, at GH (isotype), K (type). Thompson 1988. Syst. Bot. Brodiaea capitata Benth. Plantas Hartwegianas p. 339. 1848. “In sylvis Monogr. 22: 98 cites holotype K!; photos: DS!, MO!, NY!, RSA!; isotype prope Monterey. Occurrit etiam inter plantas Californicas Douglasii, = GH! Coulteri (n. 743), Jefroyi (n. 1026), et Bigelowii.” Dichelostemma cap- itatum (Bentham) A. Wood. Antirrhinum nuttallianum Benth. ex A. DC. Prodromus 10: 592. 1846. “An- TCD!: Coulter 743. Annotated: Isosyntype by G.D. Wallace November tirrhinum nuttallianum (Benth. in litt.) in California (Coulter!) ad S. 2017. Diego (Nuttall) Sairocarpus californicus Nutt.! mss.” TCD!: Coulter 599. Annotated: Type material by G.D. Wallace Novem- Carex douglasii W. Boott. var. brunnea Olney. Olney Carex pp. 361–375. In ber 2017. Also, at GH (syntype), K (presumed type material). Thomp- S. Watson, Botany of the King Expedition. 1871. p. 363 Carex douglasii son 1988. Syst. Bot. Monogr. 22: 90: “Type: “St. Diego. U-Cal.” Nuttall Boott var. brunnea Olney “California, (805 Coulter, 4503 Bolander).” = s.n. (lectotype), here designated: K-Herb. Hooker! photos: DS!, POM!; a Carex praegracilis W. Boott. 70 Wallace ALISO

TCD!: Coulter 805. Annotated: Isosyntype by G.D. Wallace August TCD!: Coulter 281. Annotated: Isotype by G.D. Wallace November 2016. Also, at GH (syntype), K (presumed type material). 2017. Also, at GH (type material), K (isotype).

Caulanthus coulteri S. Watson. Botany by Sereno Watson, aided by Daniel Helenium laciniatum A. Gray. Proc. Amer. Acad. Arts and Sciences 9: 203. C. Eaton and others, In: United States Geological Exploration of the For- 1874. “California (probably on the southeastern borders), Coulter, no. tieth Parallel. Clarence King, Geologist in charge. [Botany of the King 356, 358.” Known from Mexico. Expedition]. 27. 1871. “(Streptanthus, Gray MSS, S. heterophyllus,Gray. TCD!: Coulter 356. Annotated: Type material by G.D. Wallace Novem- Proc. Amer. Acad. 6: 185 in part, not of Nuttall).” S. California. ber 2017. Also, at GH (isosyntype). GH: Holotype. TCD!: Coulter 358. Annotated: Type material by G.D. Wallace Novem- ber 2017. Also, at GH (isolectotype). Chaenactis filifolia Harv. & A. Gray ex A. Gray. Mem. Amer. Acad. Arts and Sciences II, 4: 98. 1849. “California Coulter.” = Chaenactis glabriuscula Helenium thurberi A. Gray. Proc. Amer. Acad. Arts and Sciences 19: 32–34. DC. var. glabriuscula. TCD!: Coulter 465. Annotated: Isotype by G.D. Wallace November 1883. “S. Arizona, Coulter (359, distributed as of Californian collection), H puberulum 2017. Also, at GH (type material), K. Thurber (346 wrongly referred to . in Bot. Mex. Bound.), Pringle (137 was referred to H. ooclinium): a well-marked and apparently Cirsium coulteri Harv. & A. Gray ex A. Gray. Mem. Amer. Acad. Arts and always rayless species.” Known from Arizona and Mexico. Sciences II, 4: 110. 1849. “California Coulter.” = Cirsium occidentale TCD!: Coulter 359. Annotated: Type material by G.D. Wallace Novem- (Nuttall) Jepson var. coulteri (Harv. & A. Gray ex A. Gray) Jepson. ber 2017. Also, at GH (syntype). TCD!: Coulter 367. Annotated: Type material by G.D. Wallace November 2017. Hosackia oblongifolia Benth. (Benth. in Herb. Coll. SS. Trin. Dubl. MSS). Plantas Hartwegianas p. 305. 1849. “specimen unicum florens legit Coul- Conyza coulteri A. Gray. Proc. Amer. Acad. 7: 355. 1868. “This is Coul- ter in vicinibus Monterey.” = Hosackia oblongifolia Benth. var. oblongi- ter’s no. 285 and 286 and has recently been collected near Fort Mohave folia. [Mohave Desert] by Dr. Cooper.” = Laënnecia coulteri (A. Gray) G.L. TCD!: Coulter s.n. “unique specimen” on label. Annotated: Type mate- Nesom. rial by G.D. Wallace August 2016. [This is likely the holotype]. TCD!: Coulter 285 and Coulter 286. Annotated: Type material by G.D. Wallace November 2017. Also, at GH (syntype). Hosackia rigida Benth. (Benth. in Herb. cit. MS) Plantas Hartwegianas p. 305. 1849. “Juxta Monterey legit Coulter.” = Acmispon rigidus (Benth.) Dalea mollis Benth. in Herb. Coll. SS. Trin., Dubl. MS. Plantas Hartwe- Brouillet. gianas p. 306. 1848. “In vicinibus Monterey [California] legit Coulter.” TCD!: Coulter s.n. Annotated: Type material by G.D. Wallace August TCD!: Coulter s.n. Annotated: possible Type material by G.D. Wallace 2016. November 2017. Also, at GH (isotype), K (holotype). Hypolepis californica Hook. Species Filicum 2: 71. 1851 (or 8). “Hab. Sa. Dithyrea californica Harv. London J. Bot. 4: 77. 1845. “In California legit Barbara California, Nuttall, Dr. Coulter.” Lists as synonyms: “Aspidotis T. Coulter (N. 37).” californica Nutt. MSS. in Herb. nostr., Cheilanthes coulteri Harv. MSS. TCD!: Coulter 37. Annotated: Isotype by G.D. Wallace November 2017. n. 820 in Herb. nostr.” = Aspidotis californica (Hook.) Nutt. ex Copel. Also, at K (holotype). TCD!: Coulter 820. Annotated: Isosyntype by G.D. Wallace August Erythraea venusta A. Gray. Geological Survey of California, Botany 1: 479. 2016. Also, at GH (isosyntype), K. “The name given in Coulter’s collect &c., 25 years ago, is now resumed Lasthenia glabrata Lindl. var. coulteri A. Gray. Synoptical Flora of North for this …” = Zeltnera venusta (A. Gray) G. Mans. America 1(2): 324. 1884. “Saline marshes, S. California Coulter (no. TCD!: Coulter 561. Annotated: Type material by G.D. Wallace August 338), Brewer, Cleveland, Pringle.” = Lasthenia glabrata subsp. coulteri 2016. Also, at GH (holotype annotation on sheet). (A. Gray) Ornduff. Eupatorium sagittatum A. Gray. Smithsonian Contr. Knowl. 3(5): 88. 1850. TCD!: Coulter 338. Annotated: Isolectotype by G.D. Wallace August “California, No. 294; and Sonora Alta, Mexico, No. 252 Coulter. Between 2016. Also, at GH (lectotype). Mexico and Mazatlan, Gregg.” = Chromolaena sagittata (A. Gray) R.M. King & H. Robinson. Not in CA, or USA. Lupinus sparsiflorus Benth. Plantas Hartwegianas p. 303. 1849. “inter Lupi- TCD!: Coulter 294. Annotated: Isosyntype by G.D. Wallace August nis Coulterianis.” 2016. Also, at GH (syntype). TCD!: Coulter s.n. Lupinus sparsiflorus sp., n. California. Note on la- bel: was mixed with L. truncatus. Written on sheet: L. sparsiflorus Benth. Geraea canescens Torr. & A. Gray ex A. Gray. Proc. Amer. Acad. Arts and TCD! [Needs annotation as type material]. Also, at GH (possible type), Sciences 1: 49. 1847. “California Fremont, Coulter.” K (presumed type material). TCD!: Coulter 304. Annotated: Isosyntype by G.D. Wallace August 2016. Also, at GH (cotype written on sheet). Lyrocarpa coulteri Hook. & Harv. London J. Bot. 4: 76. 1845. “In California legit T. Coulter, 1832. (No. 40).” Gilia floribunda A. Gray. Proc. Amer. Acad. Arts and Sciences 8: 267. 1870. TCD!: Coulter 40. Annotated: Type material by G.D. Wallace November “California, probably on S.E. borders, Coulter, no. 454. Lower California 2017. Also, at GH (isotype). 50 miles S of San Diego, E.W. Morse, 1866, ex A. Wood.” = Leptosiphon floribundus (A. Gray) J.M. Porter & L.A. Johnson subsp. floribundus. Malacothrix coulteri Harv. & A. Gray ex A. Gray. Mem. Amer. Acad. Arts TCD!: Coulter 454 Annotated: Linanthus (Dactylophyllum) floribundus Sci., n.s. 4: 113–114. 1849. [Pl. Fendlerianae Novi-Mexicanae] “Califor- Harv. & Gr., Torr. & Gray, Fl. N. Amer. Annotated: Leptosiphon floribun- nia, Coulter.” dus (A. Gray) J.M. Porter & L.A. Johnson June 2007. Needs annotation TCD!: Coulter 252. Annotated: Type material by G.D. Wallace August label as Isolectotype. Also, at GH (holotype or lectotype). 2016. Also, at GH (possible type).

Haplopappus sphaerocephalus Harv. & A. Gray ex A. Gray. Mem. Amer. Malvastrum coulteri S. Watson. Proc. Amer. Acad. Arts and Sci. 11: 125. Acad. Arts and Sciences II, 4: 76. 1849 as Aplopappus “California, Coul- 1876. “Collected probably in south-eastern California by Coulter (n. 96) ter.” = Acamptopappus sphaerocephalus (Harv. & A. Gray ex A. Gray) and in the valley of the Gila by Schott, on the Mexican Boundary survey.” A. Gray var. sphaerocephalus. = Sphaeralcea coulteri (S. Watson) A. Gray. VOLUME 37(1–2) Plantae Coulterianae 71

TCD!: Coulter 96. Annotated: Type material by G.D. Wallace November ityle microglossa Benth. var. microglossa. Known from Arizona and Mex- 2017. Also, at GH (holotype). ico. TCD!: Coulter 278. Annotated: Isotype by G.D. Wallace August 2016. Mimulus glutinosus var. brachypus A. Gray. Brewer, W.H., S. Watson, & A. Also, at GH (possible type), K (presumed type material). Gray Botany of California 1: 566. 1876. “The last variety collected by According to Powell 1974, p. 270 “Type Mexico: “California”, Coulter Coulter (No. 639), near Santa Barbara by Nuttall, and a form of it, con- 278 (Holotype GH; isotypes NY).” necting with ordinary M. glutinosus, in San Luis Obispo Co. by Brewer.” = Mimulus aurantiacus Curtis var. pubescens (Torr.) D.M. Thomps. Perityle leptoglossa Harv. & A. Gray ex A. Gray Mem. Amer. Acad. Arts TCD!: Coulter 639. Annotated: Type material by G.D. Wallace Novem- and Sciences II, 4: 77. 1849. “ …in Coulter’s Californian collection;”. ber 2017. Also, at GH (lectotype). Known from Mexico. TCD!: Coulter 277. Annotated: Isotype by G.D. Wallace August 2016. Nama coulteri A. Gray. Proc. Amer. Acad. Sci. 8: 283. 1870. “California Also, at GH (holotype), K (presumed type material). (perhaps Arizona), Coulter 463, Nazas Valley, Bolson de Mapimi, Chi- huahua, Mexico, Gregg.” Known from Mexico. According Powell 1974, p. 292 “Type: Mexico: Sonora: “vicinity of Her- TCD!: Coulter 463 bis. Annotated: Type material by G.D. Wallace mosillo,” 1829–1830, Coulter 277 (Holotype GH!, fragment US!)”. November 2017. Also, at GH (lectotype). [There is no mention of Mexico or Hermosillo in the protologue]. Obione coulteri Moq. De Candolle Prodromus Systematis Naturalis Regni Perityle plumigera Harv. & A. Gray ex A. Gray Mem. Amer. Acad. Arts Vegetabilis 13(2): 113. 1849. “In California (Coulter! no. 687)” and “(v.s. and Sciences II, 4: 77. 1849. “ …in Coulter’s Californian collection;”. in h. Hook.)” = Atriplex coulteri (Moq.) D. Dietr. Gray 1888, p. 321 “California,” probably Arizona, Coulter”. = Perityle TCD!: Coulter 687 . Annotated: Isotype by G.D. Wallace November californica Benth. 2017. Also, at K (holotype). TCD!: Coulter 279. Annotated: Isotype by G.D. Wallace August 2016. Powell 1974, p. 298. “Type: Mexico: Sonora: “California” (vicinity of Orthotrichum coulteri Mitten. Journal of the Proceedings of the Linnean Hermosillo) Coulter 279. Holotype GH; fragm. UC!, US!” Society, Botany 8: 25. 1865. “Hab. California, Coulter.” [Mitten, W. 1864. Bryologia of the survey of the 49th parallel of latitude.] Phaca crotalariae Benth. in Herb. Coll. SS. Trin., Dubl. MS. Plantas TCD!: Coulter 839 and Coulter 840. Annotated: Isolectotype Or- Hartwegianas p. 307. “Juxta Monterey legit Coulter.” = Astragalus cro- thotrichum coulteri Mitten. = Orthotrichum tenellum Bruch ex Bridel by talariae D. Kelly (TCD) September 2018. Also, at FH: (lectotypes), BM: (types). (Benth.) A. Gray. TCD!: Coulter 436. Annotated: Isotype by G.D. Wallace November Pectis coulteri Harv. & A. Gray ex A. Gray Mem. Amer. Acad. Arts and 2017. Also, at K (identified as Holotype by Barneby 1964, p. 580). Sciences II, 4: 62. 1849. “California Coulter (no. 330).” Not known from Pinus bracteata D. Don Trans. Linn. Soc. London 17: 442–443. 1836. California, likely from Sonora Mexico. (Keil 1975). [David Don Libr. L.S., XXIV Descriptions of five new species of the TCD!: Coulter 330. Annotated: Holotype by D.J. Keil 1984. Also, at GH genus Pinus discovered by Dr. Coulter in California.] “Habitat in Cali- (type), K (isotype). fornia in montibus Sanctae Luciae, alt. 3000 ped. Coulter. . (v.s.sp.).” Pectis filipes Harv. & A. Gray ex A. Gray. Mem. Amer. Acad. Arts and “Species of Fir- sea side of mtn. range. 1000 ft lower than P. coulteri” = Sciences II, 4: 62. 1849. “California Coulter (no. 329).” = Pectis filipes Abies bracteata (D. Don) A. Poit. Harv. & A. Gray ex A. Gray var. filipes. Known from Mexico. TCD: No specimens found. TCD!: Coulter 329. Annotated: Holotype by D.J. Keil in 1972. Also, at Pinus coulteri D. Don Trans. Linn. Soc. London 17: 440–441. 1836. [David GH (fragment of holotype). Don Libr. L.S., XXIV Descriptions of five new species of the genus Pinus Pectis papposa Harv. & A. Gray ex A. Gray Mem. Amer. Acad. Arts and discovered by Dr. Coulter in California.] “Habitat in California, in mon- Sciences II, 4: 62. 1849. “California Coulter (no. 331).” (Keil 2006, p. tibus Sanctae Luciae, alt. 3000–4000 ped. Coulter. . (v.s.sp.).” “near the 223 incorrectly cites Coulter 331 as holotype of P. coulteri, D.J. Keil, mission of San Antonio within sight of the sea, intermingled with P. l a m - pers. comm.) bertiana”. TCD!: Coulter 331. Annotated: Holotype by D.J. Keil in 1972. Also, at TCD!: Coulter s.n. [1832/33] HOLOTYPE Annot label = “Pinus coul- GH (isotype), K (two isotypes). teri D. Don Trans. Linn. Soc. London 17: 440. 1836. HOLOTYPE coll. T. Coulter 1832/33. Det. A. Farjon 26 April 1995. USA, California, Sta. Pedicularis attenuata Benth. Prodromus 10: 574. 1846. “ …in California Lucia Mts., Cone Peak.” (Coulter!)” and “(v. in Herb. Hook.)”. = Pedicularis densiflora Hook. TCD!: 2 sheets, Coulter 606. Annotated: Isotypes by G.D. Wallace Pinus muricata D. Don Trans. Linn. Soc. London 17: 441. 1836. [David November 2017. Also, at K (presumed type material). Don Libr. L.S., XXIV Descriptions of five new species of the genus Pinus discovered by Dr. Coulter in California.] “Habitat in California ad locum Penstemon antirrhinoides Benth. ex A. DC. Prodromus 10: 594. 1846. “In San Luis Obispo Hispanice dictum, alt. 3000 ped. Coulter. . (v.s.sp.)” California (Coulter!)” and “(v. in Herb. Hook.!).” = Keckiella antirrhi- “about 10 miles from the sea, latitude 35.” noides (Benth.) Straw var. antirrhinoides. TCD!: Coulter 712. Annotated: Isotype [in error] by Farjon and cited as TCD!: Coulter 633. Annotated: Isotype by G.D. Wallace November isotype by Farjon (2017, p. 741). Note: this specimen, consisting of stem 2017. Also, at GH, K. and leaves and lacking an associated cone, cannot be an isotype because as stated by Farjon (2010, p. 718 and 2017, p. 741) “(holotype not located)” Penstemon corymbosus Benth. ex A. DC. Prodromus 10: 593. 1846. “Cal- and an isotype is defined as a duplicate of the holotype. Additionally, ifornia (Coulter!)” and “(v. in Herb. Hook.).” = Keckiella corymbosa the TCD specimen may not be suitable as type material because in the (Benth. ex A. DC.) Straw. protologue Don states “I regret that I have not seen the leaves of this [P. TCD!: Coulter 629. Annotated: Type material by G.D. Wallace Novem- muricata D. Don] and the two following species [P.radiata D. Don and P. ber 2017. Also, at GH (probable isotype), K (two specimens, presumed tuberculata D. Don]. Coulter was unable to locate them at the time among type material). his “vast collections” (Don 1836, p. 440, 441).

Perityle acmella Harv. & A. Gray ex A. Gray Mem. Amer. Acad. Arts and Pinus radiata D. Don Trans. Linn. Soc. London 17: 442. 1836. [David Don Sciences II, 4: 77. 1849. “ …in Coulter’s Californian collection;”. = Per- Libr. L.S., XXIV Descriptions of five new species of the genus Pinus 72 Wallace ALISO

discovered by Dr. Coulter in California.] “Habitat in California in maris Whitlavia minor Harv. London J. Botany 5: 312, pl. 12. 1846. (Tab. XII.) littore ad Monterey. Coulter. . (v.s.sp.)” “about Monterey 36d near sea “Hab. California, Dr. Coulter, (N. 493).” = Phacelia minor (Harv.) Thel- level almost close to beach.” lung. TCD: No specimen found. TCD!: Coulter 493. Annotated: Isotype by G.D. Wallace November 2017. Also, at K (label on sheet, type). Pinus tuberculata D. Don Trans. Linn. Soc. London 17: 442. 1836. [David Don Libr. L.S., XXIV Descriptions of five new species of the genus Pinus appendix iv discovered by Dr. Coulter in California.] “Habitat in California in maris littore ad Monterey. Coulter. . (v.s.sp.)” “along with the preceding [P. basionyms of plants and marine algae named for radiata].” thomas coulter TCD: No specimen found. The basionym is followed by current name, author and notation of whether the protologue refers to Coulter’s collection from California, Mex- Prosopis pubescens Benth. London J. Bot. 5: 82–83. 1846. “California be- ico or elsewhere. Most of the vascular plant taxa were previously listed by tween San Miguel [in the northern part of San Luis Obispo County] and Coville (1895). Monterey, Coulter.” TCD!: Coulter s.n. Mont. to San Miguel. Annotated: Type material by Flowering Plants G.D. Wallace November 2017. Acacia coulteri Benth. = Acacia macilenta Rose (Mexico) Quercus berberidifolia Liebmann. Oversigt over det Kongelige Dansk vi- Antirrhinum coulterianum Benth. ex A. DC. (California) denskabernes Selskabs Forhandlinger Medlemmers Arbeider 1854: 172. Asclepias coulteri A. Gray (Mexico) = 1854. “Q. agrifolia Nee var. n. 661 leg Dr. Coulter (hb. Hook!—Coll Fre- Astragalus coulteri Benth. Astragalus lentiginosus Dougl. ex Hook. var. mont in itinere ad Californiam (1843–44) in hb. Hook!).” borreganus M.E. Jones (California) TCD!: Coulter 661. Annotated: Type material by G.D. Wallace Novem- Bauhinia coulteri J.F. Macbr. (Mexico) = ber 2017. Also, at GH (isotype), K (holotype). Berendtia coulteri A. Gray Hemichaena coulteri (A. Gray) Thieret (Mex- ico) Romneya coulteri Harv. London J. Bot. 4: 75, pl. 3. 1845. “In California Bolanosa coulteri A. Gray (Mexico) boreali legit T. Coulter, 1832.” Brickellia coulteri A. Gray (Mexico) TCD!: Coulter s.n. [Needs annotation as type material]. Caesalpinia coulterioides Griseb. (Argentina) Calliandra coulteri S. Watson (Mexico) Ruellia tuberosa var. occidentalis A. Gray. Fl. N. Amer. 2(1): 325. 1878. “— Carex coulteri Boott ex Hemsl. (Mexico) W. & S. Texas, Berlandier, Wright. S. Arizona, Rothrock. “California” (or Caulanthus coulteri S. Watson = Streptanthus coulteri (S. Watson) Greene probably Arizona), Coulter.” = Ruellia nudiflora (Engelm. & A. Gray) (California) Urb. Cirsium coulteri Harv. & A. Gray ex A. Gray = Cirsium occidentale var. TCD!: Coulter 556. Annotated: Isosyntype by G.D. Wallace August coulteri (Harv. & A. Gray ex A. Gray) Jeps. (California) 2016. Also, at GH (syntype). Cleomella coulteri Harv.exA.Gray= Cleomella longipes Torr. (Mexico) Conyza coulteri A. Gray = Laënnecia coulteri (A. Gray) G.L. Nesom (Cal- Salix capreoides Andersson. Oefversigt af Kongl. Vetenskaps-Akademiens ifornia) forhandlingar. (Oefver. Kongl. Vet. Akad. Förh.) 15: 119. 1858. “In Cali- Coulteria Kunth = Caesalpinia L. (Mexico) fornia (Coulter) et Oregon (Hb. Hooker).” = Salix scouleriana Barratt ex Cupressus coulteri Forbes (Mexico) Hooker. Decatropis coulteri Hook. f. ex Benth. & Hook. f. May be nom. nud. [just TCD!: Coulter 656 [Needs annotation as Isotype], Also, at GH (type). name, no description] (Mexico) Salix coulteri Andersson. Oefversigt af Kongl. Vetenskaps-Akademiens Echinocactus coulteri Don (Mexico) forhandlingar (Oefver. Kongl. Vet. Akad. Förh.) 15: 119. 1858. “In Cali- Echites coulteri S. Watson = Mandevilla torosa (Jacq.) Woodson (Mexico) fornia (Coulter) Hb. Hooker.” = Salix sitchensis Sanson ex Bongard. Elaterium coulteri A. Gray = Echinopepon coulteri (A. Gray) Rose (Mex- TCD!: Coulter 655 [Needs annotation as Isotype], Also, at GH (probable ico) isotype), K. (holotype). Eunanus coulteri Harv. & A. Gray ex Benth. (California) Exostema coulteri Hook. f. = Syringantha coulteri (Hook. f.) T. McDowell Saracha acutifolia Miers. Ann. & Mag. Natural History series 2, 3(18): 449. (Mexico) 1849. “California v. in Herb. Hook. (Dr. Coulter, no. 593).” = Physalis Geniostemon coulteri Engelm. & A. Gray (Mexico) acutifolia (Miers.) Sandwith. Guaiacum coulteri A. Gray (Mexico) TCD!: Coulter 593. Annotated: Isotype by G.D. Wallace November Gutierrezia coulteri Hemsl. = Gutierrezia sphaerocephala A. Gray (Mex- 2017. Also, at K. (holotype). ico) Helianthemum coulteri S. Watson (Mexico) Trifolium ciliolatum Benth. Plantas Hartwegianas p. 304. 1849. “In pascuis Hibiscus coulteri Harv.exA.Gray(Mexico) vallis Sacramento …Coulter legit inter San Miguel et Santa Barbara.” Kosteletzkya coulteri A. Gray = Kosteletzkya hispidula (Spreng.) Garcke TCD!: Coulter 414. Not found at TCD but Coulter s.n.-labeled Trifolium (Mexico) ciliolatum sp. n. Annotated: Type material by G.D. Wallace November Lasthenia glabrata Lindl. var. coulteri A. Gray = Lasthenia glabrata subsp. 2017. Also, at K (presumed type material). coulteri (A. Gray) Ornduff (California) = Tropidocarpum macrocarpum Hook. & Harv. ex Greene. Proc. Acad. Nat. Leptoglossis coulteri A. Gray Hunzikeria coulteri (A. Gray) D’Arcy Sciences, Philadelphia 47: 553. 1895. = Tropidocarpum gracile Hook. (Mexico) = TCD!: Coulter 62. Annotated: Isotype by G.D. Wallace November 2017. Linum coulterianum Planch. Linum schiedeanum Schltdl. & Cham. Also, at K. (Mexico) Lyrocarpa coulteri Hook. & Harv. (California) Whitlavia grandiflora Harv. London J. Botany 5: 312, pl. 11. 1846. (Tab. Malacothrix coulteri Harv. & A. Gray ex A. Gray (California) XI.) “Hab. California, Dr. Coulter, (N. 492).” = Phacelia minor (Harv.) Malvastrum coulteri S. Watson = Sphaeralcea coulteri (S. Watson) A. Gray Thellung. (Mexico) TCD!: Coulter 492. Annotated: Isotype by G.D. Wallace November Marsdenia coulteri Hemsl. = Marsdenia selerorum Loes. (Mexico) 2017. Also, at K. Menodora coulteri A. Gray (Mexico) VOLUME 37(1–2) Plantae Coulterianae 73

Microsplenium coulteri Hook. f. in Benth. & Hook. f. (Mexico) Hemizonia fasciculata DC.: “first coll. by Coulter, Douglas, etc.” (Gray Nama coulteri A. Gray (California) 1884, 1(2): 310). Obione coulteri Moq. (California) Hymenoclea monogyra Torr. & A. Gray: “first coll. by Coulter” (Gray 1884, Oserya coulteriana Tul. = Noveloa coulteriana (Tul.) C.T. Philbrick (Mex- 1(2): 248). ico) Isomeris arborea Nutt.: “first coll. by Th. Coulter” (Gray 1895, 1(1): 181). Pectis coulteri Harv. & A. Gray ex A. Gray (Mexico) Leptosyne gigantea Kellogg: “first coll. by Coulter” (Gray 1884, 1(2): 300). Peperomia coulteri C. DC. (Mexico) Malacothrix saxatilis (Nutt.) Torr. & A. Gray: “first coll. by Coulter” (Gray Perezia coulteri A. Gray = Acourtia coulteri (A. Gray) Reveal & R.M. King 1884, 1(2): 423). (Mexico) Malacothrix saxatilis var. tenuifolia (Nutt.) A. Gray: “first coll. by Coulter” Phacelia coulteri Greenm. (Mexico) (Gray 1884, 1(2): 423). Philadelphus coulteri S. Watson (Mexico) Rhus ovata S. Watson: “first coll. by Th. Coulter” (Gray 1897, 1(1): 385). Pinus coulteri D. Don (California) Simsia (Geraea) canescens A. Gray: “first coll. by Coulter” (Gray 1884, Psilactis coulteri A. Gray = Arida coulteri (A. Gray) D.R. Morgan & R.L. 1(2): 282). Hartm. (Mexico) Symphoricarpos mollis Nutt.: “first coll. by Coulter and Nuttall” (Gray 1884, Pterocephalus coulteri Boiss. (Mexico) 1(2): 14). Romneya coulteri Harv. (California) Venegasia carpesioides DC.: “first coll. by Douglas and Coulter” (Gray Salix coulteri Andersson = Salix sitchensis Sanson ex Bong. (California) 1884, 1(2): 317). Salvia coulteri Fernald (Mexico) Wislizenia refracta Engelm.: “first coll. by Th. Coulter” (Gray 1895, 1(1): Schoenocaulon coulteri Baker = Schoenocaulon caricifolium (Schltdl.) A. 186). Gray var. caricifolium (Mexico) Senkenbergia coulteri Hook. f. = Boerhavia coulteri (Hook. f.) S. Watson (Mexico) Sisymbrium coulteri Hemsl. (Mexico) appendix vi Spigelia coulteriana Benth. = Spigelia hedyotidea A. DC. (Mexico) Tauschia coulteri A. Gray & Harv. ex A. Gray = Arracacia atropurpurea lesser-known generic epithets and the equivalents (Lehm.) Benth. & Hook. f. ex Hemsl. (Mexico) used here Tetraclea coulteri A. Gray (Mexico) Uragoga coulteriana Kuntze (Brazil) Genus on label Genus here Family Verbesina coulteri A. Gray (Mexico) Zaluzania coulteri Hemsl. = Zaluzania megacephala Sch. Bip. var. mega- Acarphaea Chaenactis Asteraceae cephala (Mexico) Acerates Asclepias Apocynaceae Acourtia Brickellia Asteraceae Mosses Burriel[i]a Lasthenia Asteraceae Orthotrichum coulteri Mitt. = Orthotrichum tenellum Bruch ex Brid. (Cal- Calais Uropappus Asteraceae ifornia) Callichroa Layia Asteraceae Carphephora Bebbia Asteraceae Marine Algae Chryseis Eschscholzia Papaveraceae Gelidium coulteri Harv. (California) Conobea Penstemon Plantaginaceae Hypnea coulteri Harv. = Sarcodiotheca gaudichaudii (Mont.) P.W. Gabriel- Deweya Sanicula Apiaceae son (California) Lomentaria ovalis var. coulteri Harv. = Neogastroclonium subarticulatum Dichaeta Lasthenia Asteraceae (Turner) L. Le Gall, Dalen & G.W. Saunders (California) Edosmia Perideridia Apiaceae Microcladia coulteri Harv. (California) Eunanus Mimulus Phrymaceae Eutoca Phacelia Boraginaceae appendix v Ferula Lomatium Apiaceae Glycosma Osmorhiza Apiaceae plant taxa first collected by thomas coulter Helosciadium Oenanthe Apiaceae Herpestis Mimetanthe Phrymaceae Taxa reported to have been first collected by Coulter in Gray’s Synoptical Hymenolobus Hornungia Brassicaceae Flora of North America vol. 1 part 1 (fasc.1 pp. 1–208, 1895; fasc. 2 pp. 209– Leptophragma Petunia Solanaceae 506, 1897) and part 2 (1884), and vol. 2 part 1 (1878). These determinations would have been made based on specimens or information available to Gray Lysianthus Mirabilis Nyctaginaceae at the time. Taxa for which a Coulter Californian specimen is type material Madaria Madia Asteraceae are not included in this list. Obione Atriplex Chenopodiaceae Rhus Juglans Juglandaceae Burrielia microglossa DC.: “first coll. by Coulter and Douglas” (Gray 1884, Sicydium Ibervillea Cucurbitaceae 1(2): 324). Spilanthes Perityle Asteraceae Encelia farinosa A. Gray: “first coll. by Coulter” (Gray 1884, 1(2): 282). Sycios [Sicyos] Marah Cucurbitaceae Eriophyllum multicaule (A. DC.) A. Gray: “first coll. by Coulter & Douglas” Tetranthera Umbellularia Lauraceae (Gray 1884, 1(2): 328). Turritis Boechera Brassicaceae Franseria dumosa A. Gray: “first coll. by Coulter” (Gray 1884, 1(2): 251). Zapania [sic] Phyla Verbenaceae Gilia schottii S. Watson: “first coll. by Coulter” (Gray 1878, 2(1): 409).