Vascular of Humboldt Bay's and Wetlands Published by U.S. Fish and Wildlife Service G. Leppig and A. Pickart and Department of Fish Game Release 4.0 June 2014* www.fws.gov/refuge/humboldt_bay/

Habitat- Habitat - Occurs on Status Occurs within Synonyms Common name specific broad Lanphere- Jepson Manual (2012) (see codes at end) refuge (see codes at end) (see codes at end) Ma-le'l Units UD PW EW Adoxaceae Sambucus racemosa L. red elderberry RF, CDF, FS X X N X X Aizoaceae Carpobrotus chilensis (Molina) sea fig DM X E X X N.E. Br. Carpobrotus edulis ( L.) N.E. Br. Iceplant DM X E, I X Alismataceae lanceleaf water Alisma lanceolatum With. FM X E plantain northern water Alisma triviale Pursh FM X N plantain Alliaceae three-cornered Allium triquetrum L. FS, FM, DM X X E leek Allium unifolium Kellogg one- onion CDF X N X X Amaryllidaceae Amaryllis belladonna L. belladonna lily DS, AW X X E Narcissus pseudonarcissus L. daffodil AW, DS, SW X X E X Anacardiaceae Toxicodendron diversilobum Torrey poison oak CDF, RF X X N X X & A. Gray (E. Greene) lucida L. seacoast angelica BM X X N, C X X caucalis M. Bieb bur chevril DM X E Cicuta douglasii (DC.) J. Coulter & western water FM X N Rose hemlock Conium maculatum L. poison hemlock RF, AW X I X Daucus carota L. Queen Anne's lace AW, DM X X I X American wild Daucus pusillus Michaux DM, SW X X N X X carrot Foeniculum vulgare Miller sweet fennel AW, FM, SW X X I X Glehnia littoralis (A. Gray) Miq. American DM X N, C X X ssp. leiocarpa (Mathias) Hultén silvertop Heracleum maximum Bartram Cow parsnip FM, AW X N X occidentalis J. Coulter & western lilaeopsis BM, SW X X N X X Rose Page 1 Vascular Plants of Humboldt Bay's Dunes and Wetlands Published by U.S. Fish and Wildlife Service G. Leppig and A. Pickart and California Department of Fish Game Release 4.0 June 2014* www.fws.gov/refuge/humboldt_bay/

Habitat- Habitat - Occurs on Species Status Occurs within Synonyms Common name specific broad Lanphere- Jepson Manual (2012) (see codes at end) refuge (see codes at end) (see codes at end) Ma-le'l Units Oenanthe sarmentosa J.S. Presl water parsely FM, BM, FS, RF X N X X Sanicula arctopoides Hook. & Arn. footsteps of spring DM X X N Sanicula crassicaulis DC. Pacific sanicle CDF X N X X spreading hedge- Torilis arvensis (Hudson) Link CDF X E parsley Apocynaceae

Vinca major L. bigleaf periwinkle DM, FM, CDF, DS X X E, I X X Aquifoliaceae Ilex aquifolium L. English holly CDF X X E, I X X Araceae Lemna minuta Kunth least duckweed OW, FM X N X X Lysichiton americanus Hultén & St. yellow skunk RF, FS X N X X John cabbage Zantedeschia aethiopica (L.) callalily FM, DM, DS X X E, I Sprengel Araliaceae Hedera helix L. English ivy CDF, RF X X E, I X X Hydrocotyle ranunculoides L.f. marsh pennywort FM X N X X L. common yarrow DM, CDF, SW X X N X X Ambrosia chamissonis (Less.) E. beach-bur DM, DS X N X X Greene Anaphalis margaritacea (L.) Benth. pearly everlasting CDF, DM, SW X X N X X & Hook. Anthemis arvensis L. corn chamomile DM E Anthemis cotula L. mayweed AW, FM X E, I X Arctotheca calendula (L.) Levyns cape weed CDF,FS X X E,I pycnocephala (Less.) coastal sagewort DM X N X X DC. Baccharis salicifolia (Ruiz & saltmarsh Baccharis glutinosa Pers. BM, FM X X N X X Pav.) Pers. baccharis DM, AW, BM, FM, Baccharis pilularis DC. coyote brush X X X N X X FS, RF Bellis perennis L. lawn daisy AW X E X nodding Bidens cernua L. var. cernua FM, BM X N X beggartick

Page 2 Vascular Plants of Humboldt Bay's Dunes and Wetlands Published by U.S. Fish and Wildlife Service G. Leppig and A. Pickart and California Department of Fish Game Release 4.0 June 2014* www.fws.gov/refuge/humboldt_bay/

Habitat- Habitat - Occurs on Species Status Occurs within Synonyms Common name specific broad Lanphere- Jepson Manual (2012) (see codes at end) refuge (see codes at end) (see codes at end) Ma-le'l Units Cichorium intybus L. chicory AW X E Cirsium arvense (L.) Scop. Canada thistle AW X E, I X Cirsium vulgare (Savi) Ten. bullthistle SW, DM, AW, FM X E, I X X Corethrogyne filaginifolia (Hook & California sand- Arn.) Nutt var. californica (DC.) DM X N X X aster J.P. Saroyan Australian Cotula australis (Sieber) Hook. f. DM, DS X E waterbuttons Cotula coronopifolia L. brass buttons BM, SM X X E X X Crepis capillaris (L.) Wallr. hawksbeard AW X E,I Delairea odorata Lem. Cape ivy DM, FS X X E Conyza canadensis (L.) Cronq. canadensis L. Canada horseweed DM, CDF, AW X X E, I X X var. canadensis Erigeron glaucus Ker-Gawler seaside daisy DM, DS X N X X Erigeron sumatrensis Retz. Conyza floribunda Kunth asthmaweed CDF, DM, AW X X E X X western flat- Euthamia occidentalis Nutt. FM X N X topped goldenrod ustulata (Nutt.) purple cudweed DM X N X X Nesom corn Glebionis segetum (L.) Fourr. DM X E, I chrysanthemum western marsh palustre Nutt. SW, DM, AW X X N X X cudweed Grindelia stricta DC. var. stricta coastal gumplant SM, BM X N X X Helminthotheca echioides (L.) Picris echioides L. bristly oxtongue FM, CDF, AW X X E, I X Holub Hesperevax sparsiflora (A. Gray) shortleaf dwarf- E. Greene var. brevifolia (A. Grazy) DM X N, C cudweed J. Morefield Hieracium albiflorum Hook. white hawkweed CDF X N X X Hypochaeris glabra L. smooth cat's ear FM, DM, SW X X E X X Hypochaeris radicata L. rough cat's ear DM, CDF, AW X X E X X Jaumea carnosa (Less.) A. Gray marsh jaumea SM X N X X Layia carnosa (Nutt.) Torrey & A. beach layia DM X N, C X X Gray Leontodon taraxacoides (Villars) SW, DM, BM, FM, lesser hawkbit X X E, I X X Mérat ssp. taraxacoides DS, AW

Page 3 Vascular Plants of Humboldt Bay's Dunes and Wetlands Published by U.S. Fish and Wildlife Service G. Leppig and A. Pickart and California Department of Fish Game Release 4.0 June 2014* www.fws.gov/refuge/humboldt_bay/

Habitat- Habitat - Occurs on Species Status Occurs within Synonyms Common name specific broad Lanphere- Jepson Manual (2012) (see codes at end) refuge (see codes at end) (see codes at end) Ma-le'l Units Leucanthemum vulgare Lam. oxe-eye daisy FM, DM, AW X X E, I X X Madia sativa Molina coast tarweed SW, BM, DM X X N X X Matricaria discoidea D.C. pineapple weed AW, CDF, R X X E X Petasites frigidus (L.) Fries var. coltsfoot RF X N X palmatus (Aiton) Cronq. Pseudognaphalium ramosissimum pink cudweed CDF X N X X (Nutt.) Anderb. Pseudognaphalium stramineum cotton-batting DM, SW, AW X X N X X (Kunth) Anderb. Senecio elegans L. purple ragwort DM, DS X E, I X X Erechtites glomerata (Desf. ex DM, SW, FM, DS, Senecio glomeratus Desf. ex Poir. cutleaf burnweed X X E, i X X Poir.) DC. RF, CDF Senecio jacobaea L. tansy ragwort BM X X E,I CDF, DM, BM, FM, Senecio minimus Poir Erechtites minima (Poiret) DC. coastal burnweed X X E, I X X DS, RF Senecio sylvaticus L. woodland ragwort CDF, AW X X E, I X Senecio vulgaris L. old man of spring DM, DS X E X X Silybum marianum (L.) Gaetner milkthistle AW X E, I Solidago canadensis L. var. meadow Solidago elongata Nutt. BM X N X salebrosa (Piper) M.E. Jones goldenrod Solidago simplex Kunth ssp. Solidago spathulata DC. simplex var. spathulata (DC.) goldenrod DM, CDF X N X X Cronquist Soliva sessilis Ruiz & Pavon field burnweed DM, CDF X E, I

Sonchus asper (L.) Hill ssp. asper spiny sow-thistle AW, DS, CDF X E, I X common sow Sonchus oleraceus L. DM, AW X X E, I X X thistle Symphyotrichum chilense (Nees) common SW, BM, FM X N X X G.L. Nesom var. chilense California aster Tanacetum bipinnatum (L.) Sch. dune tansy DM X N X X Bip red-seeded Taraxacum officinale Wigg. AW X E X dandelion Azollaceae Pacific mosquito Azolla filiculoides Lam. FM, SW, OW X N X X fern Mexican mosquito Azolla microphylla Kaulf OW, FM X N X X fern Betulaceae Page 4 Vascular Plants of Humboldt Bay's Dunes and Wetlands Published by U.S. Fish and Wildlife Service G. Leppig and A. Pickart and California Department of Fish Game Release 4.0 June 2014* www.fws.gov/refuge/humboldt_bay/

Habitat- Habitat - Occurs on Species Status Occurs within Synonyms Common name specific broad Lanphere- Jepson Manual (2012) (see codes at end) refuge (see codes at end) (see codes at end) Ma-le'l Units Alnus rubra Bong. red alder RF, FS, SW X N X X spectabilis Fischer & C. Amsinckia spectabilis Fischer & seaside fiddleneck DM X X N X Meyer var. spectabilis C. Meyer Borago officinalis L. common AW X E leiocarpa (Fischer & C. coastal cryptantha DM X N X X Meyer) E. Greene Myosotis discolor Pers. forget-me-not AW, FM X E asperum Lepchin rough comfrey AW X E

Barbarea verna (Mill.) Asch. early yellowrocket SW X E common Barbarea vulgaris R. Br. AW, SW X E wintercress Brassica rapa L. common mustard AW, FM X E, I X American sea edentula (Bigelow) Hook. DM X E X X rocket European sea Cakile maritima Scop. DM X E X X rocket Capsella bursa-pastoris (L.) shepherd's purse DM, DS, AW X X E X X Medikus Cardamine oligosperma Torrey & bitter cress DM, AW, DS, SW X X N X X Gray Coincya monensis (L.) Greuter & star-mustard DM X E,I Burdet small fruiting Draba verna L. DM, SW X X N spring draba Menzies' (Hook) Wettst. DM X N, C X X wallflower Hirschfeldia incana (L.) Lagr.- shortpod mustard DM, AW X X E,I X X Fossat Lepidium nitidum Nutt. var. shining Lepidium nitidum Torrey & A. Gray DM X N X X nitidum pepperweed maritima (L.) Desv. sweet DS X E Raphanus raphanistrum L. wild radish DM, AW, RF X X E X X Nasturtium officinale W.T. Aiton water cress FM, RF X N Sisymbrium officinale L. hedge mustard AW, CDF X X E, I X X

Page 5 Vascular Plants of Humboldt Bay's Dunes and Wetlands Published by U.S. Fish and Wildlife Service G. Leppig and A. Pickart and California Department of Fish Game Release 4.0 June 2014* www.fws.gov/refuge/humboldt_bay/

Habitat- Habitat - Occurs on Species Status Occurs within Synonyms Common name specific broad Lanphere- Jepson Manual (2012) (see codes at end) refuge (see codes at end) (see codes at end) Ma-le'l Units Caprifoliaceae Lonicera hispidula (Douglas ex pink honeysuckle CDF, RF X N X X Torr. & A. Gray Lonicera involucrata (Richardson) Banks var. ledebourii (Eschsch.) coast twinberry CDF, FS, RF X X N X X Zabel ramosissimum (J. A. sandmat DM, DS X N X X weinm.) Nelson & J.F. Macbr. DM, DS, SW, AW, Cerastium glomeratum Thuill. sticky chickweed X X E X X FM four-leaved Polycarpon tetraphyllum (L.) L. DM X E manyseed Sagina decumbens (Elliot) Torrey & Gray ssp. occidentalis (S. western pearlwort SW X N X X Watson) G. Crow Sagina procumbens L. birdeye pearlwort AW X E Silene gallica L. common catchfly DM X E X X Spergula arvensis L. spurry DM, DS, FM X X E

Spergularia canadensis (Pers.) G. western sand- BM, SM X X N, C X X Don var. occidentalis R. Rossbach spurrey

Spergularia salina (L.) (Griseb.) salt marsh sand Spergularia marina (L.) Griseb. SM, BM, AW X X N X X J. Presl & C. Presl spurrey Spergularia macrotheca (Hornem.) sticky sand SM, SW X X N X X Heynh. var. macrotheca spurrey Spergularia rubra (L.) J.S. Presl. & red sand spurry BM, AW X E C. Presl. Stellaria nitens Nutt. northern starwort BM, SW X N X Stellaria longipes Goldie. var. meadow starwort FM X N longipes common AW, DM, FM, DS, Stellaria media (L.) Villars X X E X X chickweed SW Chenopodiaceae Atriplex leucophylla (Moq.) D. beach salt bush DM, SM X X N Dietr. Atriplex patula L. spear saltbush BM, SM X N X X Atriplex prostrata DC. triangle oracle BM, SM, AW X X E, I X X

Page 6 Vascular Plants of Humboldt Bay's Dunes and Wetlands Published by U.S. Fish and Wildlife Service G. Leppig and A. Pickart and California Department of Fish Game Release 4.0 June 2014* www.fws.gov/refuge/humboldt_bay/

Habitat- Habitat - Occurs on Species Status Occurs within Synonyms Common name specific broad Lanphere- Jepson Manual (2012) (see codes at end) refuge (see codes at end) (see codes at end) Ma-le'l Units Chenopodium album L. lamb's quarters DM, CDF, BM X X E, I X Dysphania ambrosioides (L.) Chenopodium ambrosioides L. Mexican tea R, CDF X X E, I X Mosyakin & Clemants Chenopodium foliosum (Moench) leafy goosefoot BM, AW X X E, I Asch. Salicornia bigelovii Torrey dwarf saltwort SM X N (range ext.) X X Sarcocornia pacifica (Standl.) common Salicornia pacifica Standl. SM, BM, AW X X N X X A.J. Scott pickleweed Clusiaceae Hypericum anagalloides Cham. & tinker's penny AW, FM X N Schdl. common St. Hypericum perforatum L. DM, DS, BM X X X E, I Johnswort beach morning- soldanella (L.) R. Br. DM X N X X glory Calystegia silvatica (Kit.) griseb. shortstalk false RF, FS, AW X E, I X ssp. disjuncta Brummitt bindweed Convolvulus arvensis L. field bindweed AW, FM X E X Cuscuta pacifica Costea & M. Cuscuta salina Engelm. var. saltmarsh dodder SM, BM X X N X X Wright var. pacifica major Yuncker Crassulaceae Crassula connata Ruiz Lopez & sand pygmy-weed DM N X Pavon oreganus (Torr. ex S. coast wild- Marah oregana (S. Watson) Howell CDF, RF X X N X X Watson) Howell Cupressaceae Hesperocyparis macrocarpa Monterey cypress CDF, AW, DM X X E X X Gordon Sequoia sempervirens (D. Don) coastal redwood AW, RF X N Endl. Thuja plicata D. Don red cedar RF X N Schoenoplectus maritimus (L.) maritimus (L.) Palla seacoast bulrush BM X X N X X Lye Bolboschoenus robustus Pursh Schoenoplectus robustus (Pursh) sturdy bulrush BM X X N Soják M.T. Strong

Page 7 Vascular Plants of Humboldt Bay's Dunes and Wetlands Published by U.S. Fish and Wildlife Service G. Leppig and A. Pickart and California Department of Fish Game Release 4.0 June 2014* www.fws.gov/refuge/humboldt_bay/

Habitat- Habitat - Occurs on Species Status Occurs within Synonyms Common name specific broad Lanphere- Jepson Manual (2012) (see codes at end) refuge (see codes at end) (see codes at end) Ma-le'l Units cusickii Mackenzie Cusick's sedge FS X N Dewey's taper- Carex leptopoda Mack. CDF N sedge Carex lyngbyei Hornem Lyngbye's sedge BM, SM X X N, C X X Carex obnupta L. Bailey slough sedge SW, FM X N X X Carex pansa L. Bailey sand-dune sedge SW, DM X X N X X Cyperus eragrostis Lam. tall flatsedge FM, SW, BM, DM, R X X X N X

Eleocharis macrostachya Britton common spikerush SW, FM X N X X broad-fruit pachycarpa Desv. FM X E spikerush Eleocharis parvula (Roemer & small spikerush FM X N C Schultes) Link Isolepis cernua (Vahl) Roem. & low bulrush SW, FM, BM, SM X X N X X Schult. Schoenoplectus acutus (Muhl. ex Bigelow) A Löve & D. Löve var. hard-stemmed tule FM, BM X X N X X occidentalis (S. Watson) S.G. Sm.

Schoenoplectus americanus (Pers.) bulrush FM, BM X X N Volkart ex Schinz & R. Keller

Schoenoplectus pungens (Vahl) three-square FM, BM X X N X Palla var. longispicatus

small-fruited microcarpus C. Presl FM, RF X N X X bulrush Dennstaedtiaceae Pteridium aquilinum (L.) Kuhn var. western bracken CDF, DM, DS X N X X pubescens L. Underw. fern Dipsacaceae Dipsacus fullonum L. wild teasel AW, FM X E, I Dryopteridaceae

Dryopteris arguta (Kaulf.) Maxon coastal wood fern RF, FM, FS X N X X

Polystichum munitum (Kaulf.) C. sword fern CDF, RF X X N X X Presl

Page 8 Vascular Plants of Humboldt Bay's Dunes and Wetlands Published by U.S. Fish and Wildlife Service G. Leppig and A. Pickart and California Department of Fish Game Release 4.0 June 2014* www.fws.gov/refuge/humboldt_bay/

Habitat- Habitat - Occurs on Species Status Occurs within Synonyms Common name specific broad Lanphere- Jepson Manual (2012) (see codes at end) refuge (see codes at end) (see codes at end) Ma-le'l Units Equisetaceae

Equisetum arvense L. common horsetail FM, RF X N

smooth scouring- Equisetum laevigatum A. Braun RF, SW X N rush Equisetum telmateia Ehrh. ssp. Equisetum telmateia Ehrh. var. giant horsetail FM, RF, AW, CDF X X N X X braunii (Milde) R.L.Hauke braunii (Milde) R.L.Hauke virgata A. Gray sugar-stick CDF X N X X Arbutus menziesii Pursh Pacific madrone CDF X N X X Arctostaphylos columbiana Piper hairy manzanita CDF X N X X Arctostaphylos uva-ursi (L.) bearberry CDF, DM X N X X Sprengel Arctostaphylos x media E. Greene hybrid manzanita CDF X N X X

Chimaphila umbellata (L.) Bartram pipsissewa CDF X N X X

Erica lusitanica Rudolphi Spanish heath FM/FS X E,I Pursh salal CDF X N X X white-veined Pyrola picta Smith CDF X N X X wintergreen evergreen Vaccinium ovatum Pursh CDF, FS X X N X X huckleberry Acacia dealbata Link silver wattle FS, CDF X X E, I Acmispon americanus (Nutt.) Rydb. Lotus unifoliatus (Hook) Benth. American bird's- SW, DM X X N X X var. americanus var. unifoliatus foot trefoil

Cytisus scoparius (L.) Link scotch broom CDF, RF, FS, DS, SW X X E, I X

Genista monspessulana (L.) L. French broom FS X E, I Johnson

Hosackia gracilis Benth. Lotus formosissimus Greene coastal lotus FM, FS X N perennial sweet latifolius (L.) BM, DM X X E pea Lathyrus littoralis (Nutt.) Endl. silky beach pea DM X N X X Lathyrus palustris L. marsh pea FS, FM X N, C

Page 9 Vascular Plants of Humboldt Bay's Dunes and Wetlands Published by U.S. Fish and Wildlife Service G. Leppig and A. Pickart and California Department of Fish Game Release 4.0 June 2014* www.fws.gov/refuge/humboldt_bay/

Habitat- Habitat - Occurs on Species Status Occurs within Synonyms Common name specific broad Lanphere- Jepson Manual (2012) (see codes at end) refuge (see codes at end) (see codes at end) Ma-le'l Units Lathyrus vestitus Nutt. ssp. Lathyrus vestitus Nutt. var. vestitus hillside pea CDF X N X X vestitus Lotus corniculatus L. bird's-foot trefoil FM, SW, AW X E, I X X Lotus uliginosus Schk. Lotus pedunculatus Cav. big trefoil AW, FM X E, I X arboreus Sims yellow bush lupine DM, DS X E,I X X Lupinus bicolor Lindley miniature lupine DM X N X X Lupinus littoralis Douglas seashore lupine DM X N X X Lupinus polyphyllus Lindley var. Lupinus polyphyllus Lindley ssp. bigleaf lupine AW X N polyphyllus polyphyllus Lupinus rivularis Lindley riverbank lupine AW, DM, DS X X N X Medicago polymorpha L. bur-clover DS, BM, DM X X E X Melilotus albus Medikus Melilotus officinalis (L.) Lam. white sweetclover DM, DS, AW X E X yellow Melilotus officinalis (L.) Pall. DM. DS, AW X E X X sweetclover Trifolium arvense L. rabbitfoot clover DM, DS X E Trifolium campestre Schreber hop clover DM, DS X E Trifolium dubium Sibth. shamrock DM, DS, CDF, AW X X E, I Trifolium fragiferum L. strawberry clover AW X E, I X Trifolium glomeratum L. clustered clover CDF X E X X Trifolium hirtum All. rose clover DM, DS X E Trifolium macraei Hook. & Arn. Macrae's clover DM X N X X small-headed Trifolium microcephalum Pursh DM X N X X clover

Trifolium microdon Hook. & Arn. thimble clover DM X N X X Trifolium pratense L. red clover AW X E X Trifolium repens L. white clover AW, SW X E, I X subterranean Trifolium subterraneum L. SW, AW X E, I X clover Trifolium wormskioldii Lehm. coast clover SW, AW, FM X N X X Vicia benghalensis L. purple vetch DM, DS, BM X X X E, I X Vicia nigricans Hook. & Arn. Vicia gigantea Hook. ssp. gigantea (Hook.) Lassetter & giant vetch FS, RF X N X X C.R. Gunn. Vicia hirsuta (L.) S.F. Gray hairy vetch CDF, DM, DS X E X X Vicia sativa L. ssp. sativa spring vetch FM, RF X E

Page 10 Vascular Plants of Humboldt Bay's Dunes and Wetlands Published by U.S. Fish and Wildlife Service G. Leppig and A. Pickart and California Department of Fish Game Release 4.0 June 2014* www.fws.gov/refuge/humboldt_bay/

Habitat- Habitat - Occurs on Species Status Occurs within Synonyms Common name specific broad Lanphere- Jepson Manual (2012) (see codes at end) refuge (see codes at end) (see codes at end) Ma-le'l Units Vicia sativa L. ssp. nigra (L.) common vetch AW, DM X E X X Erhart Vicia villosa Roth ssp. villosa hairy vetch CDF X E, I X X Garryaceae Garrya elliptica Lindley coast silk tassel CDF X N X X davyi (Jepson) davyi (Jepson) G. Mans. Davy's centaury BM X N Abrams Zeltnera muehlenbergii (Griseb.) G. Centaurium muehlenbergii Muhlenberg's FM, SW X N X X Mans. (Griseb.) W. Wight ex Piper centaury Geraniaceae long-beaked Erodium botrys (Cav.) Bertol. DS, DM X E filaree red-stemmed Erodium cicutarium (L.) L’Hér. AW, DM X X E filaree Geranium dissectum L. cutleaf geranium AW X E X dove's-foot Geranium molle L. AW, DM, DS X X E geranium Geranium robertianum L. Robert's geranium RF, FS X E, I Grossulariaceae Ribes divaricatum Douglas var. spreading CDF X N X X pubiflorum Koehne gooseberry trailing black Ribes laxiflorum Pursh CDF X N X X currant Ribes sanguineum Pursh var. pink-flowering CDF, FS, RF X X N X X glutinosum (Benth) Loudon currant Hippuridaceae common mares- Hippuris vulgaris L. FM X N tail Hydrocharitaceae Egeria densa Planchon Brazilian elodea OW X E, I Iridaceae Crocosmia xcrocosmiiflora (Burb. monbretia DM, DS, FM X X E, I & Dean) N.E. Br. Iris douglasiana Herbert Douglas Iris CDF, FM X X N Iris germanica L. var. florentina orris-root DS X E (L.) Dykes

Page 11 Vascular Plants of Humboldt Bay's Dunes and Wetlands Published by U.S. Fish and Wildlife Service G. Leppig and A. Pickart and California Department of Fish Game Release 4.0 June 2014* www.fws.gov/refuge/humboldt_bay/

Habitat- Habitat - Occurs on Species Status Occurs within Synonyms Common name specific broad Lanphere- Jepson Manual (2012) (see codes at end) refuge (see codes at end) (see codes at end) Ma-le'l Units Iris pseudacorus L. water iris FM X E Sisyrinchium bellum S. Watson blue-eyed grass FM X N Sisyrinchium californicum (Ker California golden- SW, FM X N X X Gawler) Dryander eyed grass ambiguus Guss. saline toad rush BM, SM, X X N Juncus articulatus L. joinleaf rush FM X N Juncus arcticus Willd. ssp. Juncus balticus Willd. Baltic rush SM, BM X X N littoralis (Engelm.) Hultén Juncus bolanderi Engelm. Bolander's rush AW, FM X N X Juncus breweri Engelm. Brewer's rush SW, DM X X N X X Juncus bufonius L. var. bufonius toad rush SW, AW, FM X N X X Juncus bufonius L. var. western toad rush DM, SW X X N X X occidentalis F.J. Herm Juncus effusus L. var. brunneus Juncus hesperius (Piper) Lint bog rush AW, FM X N X Engelm. Juncus effusus L. ssp. pacificus Juncus effusus L. var. pacificus Pacific rush AW X N X (Fern. & Wieg.) Piper & Beattie Fern. & Wieg. three-stamened Juncus ensifolius Wikstrom AW, FM X N rush Juncus falcatus E. Meyer ssp. sickle-leafed rush SW X N X X falcatus Juncus gerardii Loisel. salt-meadow rush SM X E Juncus lescurii Bol. Juncus lesueurii Bol. salt rush BM, SM X X N X X common wood- comosa E. Meyer CDF, DM, DS X N X X rush Luzula subcongesta (S. Watson) Donner wood rush CDF, DM N X X Jepson Juncaginaceae Triglochin concinna Burtt Davy Triglochin concinna Burtt Davy arrow-grass SM X N X X var. concinna seaside arrow- Triglochin maritima L. BM, SM X X N X X grass Triglochin scilloides (Poir.) Mering flowering Lilaea scilloides (Poiret) Hauman SM, BM X N & Kadereit quillwort Triglochin striata Ruiz Lopez & three-ribbed arrow- BM, FM X N X Pavon grass

Page 12 Vascular Plants of Humboldt Bay's Dunes and Wetlands Published by U.S. Fish and Wildlife Service G. Leppig and A. Pickart and California Department of Fish Game Release 4.0 June 2014* www.fws.gov/refuge/humboldt_bay/

Habitat- Habitat - Occurs on Species Status Occurs within Synonyms Common name specific broad Lanphere- Jepson Manual (2012) (see codes at end) refuge (see codes at end) (see codes at end) Ma-le'l Units purpureum L. purple dead-nettle FM, DM, DS X X E X X Mentha pulegium L. pennyroyal SW, AW, FM X E, I X X Prunella vulgaris L. var. vulgaris Prunella vulgaris L. ssp. vulgaris self-heal AW X E Clinopodium douglasii (Benth.) yerba buena CDF X N X X Kuntze Stachys chamissonis Benth. coast hedge-nettle RF, FS X N X X FM, BM, RF, SW, Stachys ajugoides Benth. rigid hedge-nettle X N X AW Laxmanniaceae Cordyline australis (G. Forst.) cabbage tree FM, BM E Endl. Liliaceae Fritillaria affinis (Schultes) Sealy checker lily CDF X N X X common grape Muscari botryoides (L.) Miller CDF X E hyacinth

Linum bienne Miller narrow-leaved DM, FM, BM X X E Linnaeaceae Linnaea borealis L. var. longiflora Linnaea borealis L. ssp. N twin CDF X X X Torrey longiflora (Torrey) Hultén planted Lythraceae Lythrum hyssopifolium L. hyssop loosestrife SW, AW, FM X E, I X X Malvaceae Malva neglecta Wallr. common mallow AW X E X Montiaceae Calandrinia ciliata (Ruíz Lopez & red-maids DM, DS, SW X X N X X Pavón) DC. Claytonia exigua Torrey & A. Gray little spring beauty DM, DS X N X X ssp. exigua Claytonia perfoliata Willd. ssp. miner's lettuce DM, DS, RF X X N X X perfoliata

Page 13 Vascular Plants of Humboldt Bay's Dunes and Wetlands Published by U.S. Fish and Wildlife Service G. Leppig and A. Pickart and California Department of Fish Game Release 4.0 June 2014* www.fws.gov/refuge/humboldt_bay/

Habitat- Habitat - Occurs on Species Status Occurs within Synonyms Common name specific broad Lanphere- Jepson Manual (2012) (see codes at end) refuge (see codes at end) (see codes at end) Ma-le'l Units Claytonia rubra (Howell) red-stemmed Tidestrom ssp. depressa (A. Gray) DM X N X spring beauty John M. Miller & Chambers Myrtaceae globulus Labill. blue gum RF, AW, CDF X E X Myricaceae

Morella californica (Cham.) Wilbur wax myrtle CDF, RF, FS, SW X X N X X

Myrsinaceae

Anagallis arvensis L. scarlet pimpernel FM, DM, BM, SW X X X E

Glaux maritima L. sea milk-wort BM, SM X X N X X yellow sand latifolia Eschsch DM X N X X verbena Abronia umbellata Lam. var. Abronia umbellata Lam. ssp. pink sand verbena DM X N, C X X breviflora (Standl.) L.A. Galloway breviflora (Standley) Munz Nymphaeaceae Nuphar lutea (L.) Sm. ssp. Nuphar polysepala Engelm. yellow pond-lily FM X N X polysepala (Engelm.) E.O. Beal cheiranthifolia (Sprengel) Raim ssp. beach-primrose DM X N X X cheiranthifolia Camissonia strigulosa (Fischer & strigose sun-cup CDF, DM X N X X C. Meyer) Raven davyi (Jespon) Harlan Davy's clarkia DM, DS X N X X Lewis & M. Lewis Clarkia purpurea (Curtis) Nelson & J.F. Macbr ssp. quadrivulnera purple clarkia SW X N X X (Douglas) Harlan Lewis & M. Lewis. Chamerion angustifolium (L.) Holub ssp. circumvagum (Mosquin) fireweed FM, RF X N X X Hoch Epilobium ciliatum Raf. ssp. Watson's SW, FM, BM, AW X X N X X watsonii (Barbey) P. Hoch & Raven willowherb

Page 14 Vascular Plants of Humboldt Bay's Dunes and Wetlands Published by U.S. Fish and Wildlife Service G. Leppig and A. Pickart and California Department of Fish Game Release 4.0 June 2014* www.fws.gov/refuge/humboldt_bay/

Habitat- Habitat - Occurs on Species Status Occurs within Synonyms Common name specific broad Lanphere- Jepson Manual (2012) (see codes at end) refuge (see codes at end) (see codes at end) Ma-le'l Units magellanica Lam. hardy fuchsia FS X E

red-sepaled Oenothera glazioviana Micheli DM, DS, FM X X E, I evening-primrose

Ophioglossaceae Sceptridium multifidum (S.G. Botrychium multifidum (S. common grapefern FM, FS, SW X N X X Gmel.) Tagawa Gmelin) Trevis. Orchidaceae Calypso bulbosa (L.) Oakes fairy slipper CDF X X N X X Corallorhiza maculata Raf. summer coral root CDF X N X X rattlesnake Goodyera oblongifolia Raf. CDF X N X X plantain heart-leaved Listera cordata (L.) R. Br. CDF X N X X twayblade Piperia elegans (Lindley) Rydb. coast piperia SW, CDF X X N X X Piperia transversa Suksd. rein orchid CDF X N X X hooded ladies- Spiranthes romanzoffiana Cham. SW, CDF X X N X X tresses Orobanchaceae Mediterranean Bellardia trixago (L.) All. FM, DM X X E, I lineseed Castilleja ambigua Hook & Humboldt Bay Arn.ssp. humboldtiensis (Keck) SM X X N, C X X owl's-clover Chuang & Heckard Castilleja attenuata (A. Gray) narrow-leaved DM X N X X Chuang & Heckard owl's clover Castilleja exserta (A.A. Heller) purple owl's Chuang & Heckard ssp. latifolia (S. DM X N X X clover Watson) Chuang & Heckard Chloropyron maritimum (Benth.) Point Reyes bird's- A. Heller ssp. palustre (Behr) Tank SM X N, C X X beak & J.M. Egger

Parentucellia viscosa (L.) Caruel yellow glandweed SW, FM, AW X E, I X X

Triphysaria eriantha (Benth.) butter 'n' eggs DM X N Chuang & Heckard ssp. eriantha

Page 15 Vascular Plants of Humboldt Bay's Dunes and Wetlands Published by U.S. Fish and Wildlife Service G. Leppig and A. Pickart and California Department of Fish Game Release 4.0 June 2014* www.fws.gov/refuge/humboldt_bay/

Habitat- Habitat - Occurs on Species Status Occurs within Synonyms Common name specific broad Lanphere- Jepson Manual (2012) (see codes at end) refuge (see codes at end) (see codes at end) Ma-le'l Units Triphysaria pusilla (Benth.) little owl's clover DS, DM, CDF X N Chuang & Heckard Triphysaria versicolor Fischer & C. yellow owl's AW X N Meyer ssp. versicolor clover Oxalidaceae Oxalis articulata Savigny ssp. Oxalis rubra A. St. Hil. red oxalis DM, DS X E, I rubra (A. St. -Hil.) Lourteig creeping Oxalis corniculata L. FS, DS X X E X X woodsorrel Oxalis pes-caprae L. DM, DS X E, I buttercup Papaveraceae Eschscholzia californica Cham. California poppy AW, DM X X N X Fumaria officinalis (L.) fumitory AW X E Platystemon californicus Benth. cream cups DM, DS X N X X Phrymaceae common Mimulus guttatus DC. FM, AW, RF X N X monkeyflower Pinaceae Abies grandis (Douglas) Lindley grand fir CDF, RF X X N X X Picea sitchensis (Bong.) Carriere Sitka spruce CDF, FS, SW X X N X X Pinus contorta Loudon ssp. Pinus contorta Douglas ex shore pine CDF, SW X X N X X contorta Louden var. contorta Pinus radiata D. Don Monterey pine AW, CDF X X E X Pseudotsuga menziesii (Mirbel) Douglas-fir CDF X N X X Franco var. menziesii Pittosporaceae

Pittosporum tenuifolium Gaertner short-leaf box CDF X E, I X X

Plantaginaceae Callitriche heterophylla Pursh Callitriche heterophylla Pursh var. ssp. bolanderi (Hegelm.) Calder water starwort DS, AW, FM X X N bolanderi (Hegelm.) Fassett & Roy L. Taylor pond water- Callitriche stagnalis Scop. OW, FM X E starwort Digitalis purpurea L. fox glove FM, DS X E, I Nuttallanthus texanus (Scheele) blue toad-flax DM X E X X D.A. Sutton Page 16 Vascular Plants of Humboldt Bay's Dunes and Wetlands Published by U.S. Fish and Wildlife Service G. Leppig and A. Pickart and California Department of Fish Game Release 4.0 June 2014* www.fws.gov/refuge/humboldt_bay/

Habitat- Habitat - Occurs on Species Status Occurs within Synonyms Common name specific broad Lanphere- Jepson Manual (2012) (see codes at end) refuge (see codes at end) (see codes at end) Ma-le'l Units Plantago coronopus L. cut-leaf plantain DM, SW X X E Plantago erecta E. Morris California plantain DM, SW, DS X X N X X narrow-leaved Plantago lanceolata L. CDF, DM, AW X X E X X plantain Plantago major L. common plantain SW, AW, FM X E X Plantago maritima L. salt marsh plantain SM X N X X tall coastal Plantago subnuda Pilger AW, SW, FM X N plantain Veronica americana (Raf.) American FM X N Schwein. speedwell bird's-eye Veronica persica Poiret AW X E X X speedwell Armeria maritima (Miller) Willd. California sea- DM, DS X N X X ssp. californica (Boiss.) pink californicum (Boiss.) western marsh- SM X N X X A.A. Heller rosemary exarata Trin. spike bentgrass AW, BM X N X Agrostis hallii Vasey Hall's bentgrass CDF X N X X little-leaf Agrostis microphylla Steudel SW X N X X bentgrass

Agrostis stolonifera L. creeping bentgrass BM, AW, FM X E, I X X

Aira praecox L. yellow hairgrass DM, DS, SW X X E, I X X Aira caryophyllea L. silver hairgrass DM, DS, SW X X E, I X X Alopecurus aequalis Sobol. short-awn foxtail FM X N X Alopecurus geniculatus L. water foxtail FM, AW X N X Alopecurus pratensis L. meadow foxtail AW X E European beach (L.) Link DM, DS X E, I X X grass Anthoxanthum occidentale western Hierochloe occidentalis Buckley CDF X N X (Buckley) Veldkamp sweetgrass DM, DS, SW, AW, Anthoxanthum odoratum L. sweet vernal grass X X E, I X X FS, CDF Avena fatua L. wild oats AW, DM X E Page 17 Vascular Plants of Humboldt Bay's Dunes and Wetlands Published by U.S. Fish and Wildlife Service G. Leppig and A. Pickart and California Department of Fish Game Release 4.0 June 2014* www.fws.gov/refuge/humboldt_bay/

Habitat- Habitat - Occurs on Species Status Occurs within Synonyms Common name specific broad Lanphere- Jepson Manual (2012) (see codes at end) refuge (see codes at end) (see codes at end) Ma-le'l Units Avena sativa L. common oats AW X E X Briza maxima L. big quaking grass DM, DS, SW X X E , I X X little quaking Briza minor L. SW, FM X E X X grass carinatus Hook & Arn. var. Bromus carinatus Hook & Arn. mountain brome DM X N X X carinatus Bromus diandrus Roth ripgut brome DM, DS, SW X E, I X X Bromus hordeaceus L. soft chess DM, BM , SW X X X E, I X X

Bromus maritimus (Piper) Hitchc. maritime brome DM, DS X N X X

Calamagrostis nutkaensis (J. Presl) reedgrass CDF, BM X X N X X Steudel Cortaderia jubata (Lemoine) Stapf jubata grass DM, SW, CDF, RF X X E, I X X Cortaderia selloana (Schultes) pampas grass DM X E, I Asch & Graebner Cynodon dactylon (L.) Pers. bermuda grass DM, DS, AW X X E, I Cynosurus echinatus L. dogtail grass DM, DS, CDF X E, I X X Dactylis glomerata L. orchard-grass CDF, AW X X E, I X X Deschampsia cespitosa (L.) Beauv. coastal tufted hair- ssp. holciformis (C. Presel) W.E. Deschampsia holciformis J. Presl BM,SM X X N X X grass Lawrx Distichlis spicata (L.) E. Greene saltgrass SM, BM, SW, AW X X N X X Echinochloa crus-galli (L.) P. barnyard grass FM X E, I X Beauv. glaucus Buckley ssp. Jepson's blue CDF, AW X N glaucus wildrye mollis (Trin.) Pilger ssp. Elymus mollis Trin. ssp. mollis dune wild-rye DM X N X X mollis Leymus xvancouverensis (Vasey) Vancouver wild- Elymus xvancouverensis Vasey AW, DM, BM X X X N X X Pilger rye Schedonorus arundinaceus arundinacea Schreber tall fescue FM, BM, AW X E, I X (Schreb.) Dunmort., nom. cons. Festuca bromoides L. Vulpia bromoides (L.) Gray brome fescue DM, DS X E, I X X Festuca myuros L. Vulpia myuros (L.) C. Gmelin six-weeks fescue CDF, R X E X Festuca occidentalis Hook. western fescue CDF X N X X

Page 18 Vascular Plants of Humboldt Bay's Dunes and Wetlands Published by U.S. Fish and Wildlife Service G. Leppig and A. Pickart and California Department of Fish Game Release 4.0 June 2014* www.fws.gov/refuge/humboldt_bay/

Habitat- Habitat - Occurs on Species Status Occurs within Synonyms Common name specific broad Lanphere- Jepson Manual (2012) (see codes at end) refuge (see codes at end) (see codes at end) Ma-le'l Units Festuca perennis (L.) Columbus & Lolium perenne L. ssp. Italian rye-grass AW, RF X E, I X X J.P. Sm. multiflorum (Lam.) Husnot Lolium perenne L. ssp. perenne perennial ryegrass AW, CDF X X E X L. red fescue DM, CDF X N X X Glyceria elata (Nash) A. Hitchc. Glyceria striata (Lam.) Hitchc. tall mannagrass FM X N X Glyceria xoccidentalis (Piper) J.C. western FM, RF X N X X Nelson mannagrass common Holcus lanatus L. AW, SW, FM, CDF X X E, I X X velvetgrass

Hordeum brachyantherum Nevski meadow barley BM X N X ssp. brachyantherum

Hordeum jubatum L. foxtail barley BM, SM X X N X X

Hordeum marinum Hudson ssp. Mediterranean AW, SM X X E gussoneanum (Parl.) Thell. barley

Hordeum murinum L. ssp. glaucum smooth barley DM X E (Stuedel) Tzvelev Hordeum murinum L. ssp. wall barley DM, DS X E murinum Parapholis incurva (L.) C.E. Hubb. curved sicklegrass SM X E X X

Parapholis strigosa (Dumort) C.E. hairy sickle grass SM X E, I X X Hubb. Phalaris aquatica L. Harding grass FM, AW X E, I X Phalaris arundinacea L. reed canarygrass FM, AW, RF, FS X E, I X

Phragmites australis (Cav.) Steudel common reed BM, SM X X E, I Pleuropogon californicus (Nees) semaphore grass AW X N Vasey Poa annua L. annual blue grass CDF, DM, DS, AW X X E X X

Poa confinis Vasey coastline bluegrass DM X N X X large-flowered Poa macrantha Vasey sand-dune blue DM X N X X grass Kentucky blue Poa pratensis L. ssp. pratensis CDF X E X grass

Page 19 Vascular Plants of Humboldt Bay's Dunes and Wetlands Published by U.S. Fish and Wildlife Service G. Leppig and A. Pickart and California Department of Fish Game Release 4.0 June 2014* www.fws.gov/refuge/humboldt_bay/

Habitat- Habitat - Occurs on Species Status Occurs within Synonyms Common name specific broad Lanphere- Jepson Manual (2012) (see codes at end) refuge (see codes at end) (see codes at end) Ma-le'l Units Polypogon interruptus Kunth ditch beard grass AW, FM X E X Mediterranean Polypogon maritimus Willd. SW, FM X E, I X beard grass Polypogon monspeliensis (L.) Desf. rabbitsfoot grass SW, FM X E, I X X

Polypogon viridis (Gouan) Breistr. water beard grass AW, BM X X E X

Puccinellia nutkaensis (J.S. Presl.) Alaska alkali SM X N X X Fern. & Weath. grass dense-flowered Spartina densiflora Brongn. SM, BM X X E, I X X cordgrass Torreyochloa pallida (J.S. Presl) Church var. pauciflora (J.S. Presl) weak mannagrass FM X N J.I. Davis Trisetum canescens Buckley nodding oatgrass CDF X N X X Zea mays L. corn AW X E millefoliata Fischer & C. dark-eyed gilia DM X N, C X X Meyer Navarretia squarrosa (Eschsch.) skunkweed DM, AW X X N Hook. & Arn. latifolium Smith coast buckwheat DM X N X X Fallopia japonica (Houtt.) Ronse Polygonum cuspidatum Sieb. & Japanese DM X E, I Decr. Zucc. knotweed Persicaria hydropiperoides Polygonum hydropiperoides water pepper FM X N X (Michx.) Small Michaux Persicaria lapathifolia (L.) Gray Polygonum lapathifolium L. willow weed FM, AW X N X Persicaria maculosa Gray Polygonum persicaria L. lady's thumb AW, FM, BM, RF X E, I X Polygonum aviculare L. ssp. Polygonum arenastrum Boreau common knotweed AW X E depressum (Meisn.) Arcangeli Polygonum paronychia Cham. & beach knotweed DM, DS X N X X Schldl. DM, DS,SW, CDF, Rumex acetosella L. sheep sorrel X X E, I X X RF Rumex conglomeratus Murr. green dock AW X E X Rumex crispus L. curly dock FM, SW, AW, BM X X E, I X X Rumex fueginus Phillippi Rumex maritimus L. golden dock BM X X N X Page 20 Vascular Plants of Humboldt Bay's Dunes and Wetlands Published by U.S. Fish and Wildlife Service G. Leppig and A. Pickart and California Department of Fish Game Release 4.0 June 2014* www.fws.gov/refuge/humboldt_bay/

Habitat- Habitat - Occurs on Species Status Occurs within Synonyms Common name specific broad Lanphere- Jepson Manual (2012) (see codes at end) refuge (see codes at end) (see codes at end) Ma-le'l Units Rumex aquaticus L. var. Rumex occidentalis S. Watson western dock FM, BM X N X X fenestratus (Greene) Dorn Rumex pulcher L. fiddle dock AW X E X

Rumex salicifolius Weinm. var. willow-leaved Rumex crassus Rech. f. SW, BM X X N X X crassus (Rech. f.) J.T. Howell dock

Rumex salicifolius Weinm. var. Rumex transitorius Rech. f. transitorius (Rech. f.) J.C. willow dock FM, BM X N X Hickman Polypodiaceae Polypodium calirhiza S. Whitmore licorice fern CDF, DS X N X X & A. R. Smith Polypodium scouleri Hook. & leather-leaf fern CDF X N X X Grev. Potamogetonaceae long-leaved Potamogeton nodosus Poiret OW X N pondweed

Stuckenia pectinata (L.) Böerner fennel pondweed OW X N X

Pteridaceae

Pentagramma triangularis (Kaulf.) G. Yatskievych, Windham & gold-back fern CDF, DM X N X X Wollenweber ssp. triangularis

Ranunculaceae California Ranunculus californicus Benth. SW X buttercup Ranunculus muricatus L. spiny buttercup AW X E Ranunculus repens L. creeping buttercup FM, RF, AW X E X X Rhamnaceae

Frangula purshiana (DC.) Cooper cascara CDF X N X X

Aphanes occidentalis (Nutt.) Rydb. arvensis L. lady's mantle DM, SW X X N

Crataegus monogyna Jacq. hawthorn AW X E, I

Page 21 Vascular Plants of Humboldt Bay's Dunes and Wetlands Published by U.S. Fish and Wildlife Service G. Leppig and A. Pickart and California Department of Fish Game Release 4.0 June 2014* www.fws.gov/refuge/humboldt_bay/

Habitat- Habitat - Occurs on Species Status Occurs within Synonyms Common name specific broad Lanphere- Jepson Manual (2012) (see codes at end) refuge (see codes at end) (see codes at end) Ma-le'l Units Francheti Cotoneaster franchetii Boiss. CDF X E, I X X cotoneaster

Fragaria chiloensis (L.) Duchesne beach strawberry DM, CDF X N X X

Holodiscus discolor (Pursh.) oceanspray FS X N X Maxim. Malus fusca ( Raf.) C. Schneider Oregon crabapple RF, CDF X X N X X

Malus pumila Mill. common apple CDF, FS X X E X X Oemleria cerasiformis (Hook. oso berry RF X N X &Arn.) J.W. Landon Potentilla anserina L. ssp. pacifica egedii (Wormsk.) silverweed SW, BM, FM X X N X X (T.J. Howell) Rousi Rydb. ssp. egedii Rosa nutkana C. Presl ssp. Nootka rose CDF X N X X nutkana Himalayan Rubus armeniacus Focke FS, SW, RF, AW, X E, I X blackberry Rubus parviflorus Nutt. thimbleberry FS, RF X N X X Rubus spectabilis Pursh salmonberry RF, FS X N X X California CDF, RF, FS, DM, Rubus ursinus Cham. & Schdl. X X N X X blackberry DS, SW Spiraea douglasii Hook. Douglas' spirea FM, FS, AW X N CDF, RF, BM, FM, aparine L. common bedstraw X X N X X AW Lamarck's Galium divaricatum Lam. FM X E X bedstraw Galium trifidum L. ssp. three-petaled BM X X N X X columbianum (Rydb.) Hultén bedstraw Ruppiaceae Ruppia maritima L. ditch grass SM, BM X N X X Ruscaceae Maianthemum dilatatum (Alph. false lily of the CDF, RF X X N X X Wood) Nelson & J.F. Macbr. vally Salicaceae Populus balsamifera L. ssp. Populus trichocarpa Hook. trichocarpa (Torry & A. Gray) black cottonwood FS, RF X N Brayshaw

Page 22 Vascular Plants of Humboldt Bay's Dunes and Wetlands Published by U.S. Fish and Wildlife Service G. Leppig and A. Pickart and California Department of Fish Game Release 4.0 June 2014* www.fws.gov/refuge/humboldt_bay/

Habitat- Habitat - Occurs on Species Status Occurs within Synonyms Common name specific broad Lanphere- Jepson Manual (2012) (see codes at end) refuge (see codes at end) (see codes at end) Ma-le'l Units Salix alba L. white willow FS X E, I Salix hookeriana Hook. Hoooker willow SW, FS, DS X X N X X Salix lasiandra Benth. var. Salix lucida Muhl. Ssp. lasiandra Pacific willow FS, RF X N X lasiandra (Benth.) A.E. Murray Salix sitchensis Bong. Coulter Willow FS X N Sapindaceae Acer macrophyllum Pursh big leaf maple FS X N X Saxifragaceae

Tellima grandiflora (Pursh) Lindley fringe cups RF X N X X

Tolmiea menziesii (Pursh) Torrey & pig-a-back plant RF X N A. Gray Scrophulariaceae Scrophularia californica Cham. & California figwort DS, SW, CDF X X N X X Schldl. Solanaceae Solanum americanum Miller white nightshade FM, CDF X X N X X Solanum aviculare Forst. F. Solanum laciniatum G. Forst. CDF X E, I X X nightshade greenspot Solanum douglasii Dunal FS X N nightshade Solanum nigrum L. black nightshade CDF, AW, FM X X E, I X X Themidaceae early harvest Brodiaea coronaria (Salisb.) Engl. DM, CDF X N X X brodiaea Dichelostemma capitatum Alph. blue dicks DM , CDF X N X X Wood Dichelostemma congestum (Sm.) fork-toothed CDF X N X X Kunth ookow Triteleia hyacinthina (Lindley) E. white brodiaea DM, CDF X N X X Greene Typhaceae Sparganium eurycarpum A. Gray Sparganium eurycarpum Engelm. bur-reed BM X X N X X var. eurycarpum Typha latifolia L. cattail SW, BM, FM X X N X X Urticaceae Urtica dioica L. ssp. gracilis stinging nettle RF, AW, FM X N (Aiton) Selander Page 23 Vascular Plants of Humboldt Bay's Dunes and Wetlands Published by U.S. Fish and Wildlife Service G. Leppig and A. Pickart and California Department of Fish Game Release 4.0 June 2014* www.fws.gov/refuge/humboldt_bay/

Habitat- Habitat - Occurs on Species Status Occurs within Synonyms Common name specific broad Lanphere- Jepson Manual (2012) (see codes at end) refuge (see codes at end) (see codes at end) Ma-le'l Units Valerianaceae Plectritis congesta (Lindley) A. sea blush DM, SW, DS X X N X DC. Plectritis congesta (Lindl.) DC. short-spur sea ssp. brachystemon (Fisch. & C.A. AW X N blush Mey) Morey Woodsiaceae Athyrium filix-femina (L.) Roth var. Athyrium filix-femina (L.) Roth lady fern CDF, FS X X N X X cyclosorum Rupr. ssp. cyclosorum (Rupr.) C. Chr. Zosteraceae Zostera japonica Aschers. & Nanozostera japonica (Asch. & dwarf eelgrass MF X E, I Graebn. Graebn.) P. Toml. & U. Posl. Zostera marina L. eelgrass MF X N X X

Status: Conservation concern (C); Exotic (E); Invasive (I); California Native (N); Broad Habitat: Upland dune (UD), Palustrine wetland (PW), Estuarine wetland (EW). Specific Habitat: Dune mat (DM), Dune scrub (DS), Dune swale (SW), Coniferous dune forest (CDF), Riparian forest (RF), Freshwater swamp (FS), Freshwater marsh (FM), Open water (OW), Brackish marsh (BM), Salt marsh (SM), Mudflat (MF) Agricultural wetland (AW).

* This list for Humboldt Bay dunes and wetlands is still in progress. It is based on collections made by the authors primarily between 2001 and 2008. Please email comments to [email protected] or [email protected]. Nomenclature follows the Jepson Manual II, Higher Plants of California (2012).

Synonyms are current names accepted by USDA PLANTS

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