Smithsonian Fellowships Te Smithsonian provides several fellowships through its Ofce of Fellowships and Internships (OFI) to Diptera Collection at the enable graduate students, postdocs, and others to work at the NMNH and utilize the collection and facilities for research. Te application deadline for each of the National Museum of Natural Samuel Wendell Williston was a distinguished biologist programs below is 1 December annually. who made signifcant contributions to paleontology, History 10-week Graduate Student Fellowship: M.Sc. or early Ph.D. entomology, medicine, and education. He was the frst students can spend 10 weeks at the NMNH (stipend of native dipterist in the U. S., the frst to produce generic US $7,000). monographs of Nearctic Diptera, the frst to curate and study the Diptera of the USNM, and the frst to make 3 – 24 month Predoctoral Fellowship: Ph.D. students a contribution to that collection (his types of Nearctic who have fulflled the requirements of candidacy (or its Syrphidae). equivalent internationally) can spend up to 24 months Te S.W. Williston Diptera Research Fund is a small at the NMNH (stipend of US $32,700 annually plus an endowment fund that has been established for “the annual research budget of US $4,000). increase and difusion of knowledge about Diptera” and 12 – 24 month Postdoctoral Fellowship: Scientists who have welcomes applications with an annual deadline of 30 completed their Ph.D. within the past fve years can apply November (asiloidfies.si.edu/williston-diptera-research-fund). to the competitive postdoctoral fellowship program in About US $6,000 are available from the endowment Evolutionary and Systematic Biology and conduct cutting- annually. To this day, the fund has supported the travel edge research in close collaboration with one of the of graduate students and dipterists to the International curators (stipend of US $48,000 annually plus an annual Congresses of Dipterology and to our museum for research budget of US $4,000). collections-based research as well as feld work. USNM pinned Diptera collection at NMNH Smithsonian Short-Term Visitor Program: Tis program is Please consider donating to this endowment fund to available for a scholarly visit to the NMNH for research or support the increase and difusion of knowledge about collaboration of up to 21 days (funding of US $2,000) or Diptera and particularly the research and travel of a new 30 days for scientists from developing regions of the world generation of dipterists. (funding of up to US $4,000). Te application deadline is not fxed and submissions are welcome year round.

Further information and contacts SI Department of Entomology entomology.si.edu SIEntomology @NMNH_entomology SEL www.sel.barc.usda.gov WRBU www.wrbu.org Diptera collection entomology.si.edu/Collections_Diptera.html search Diptera collection collections.nmnh.si.edu/search/ento

SI Fellowship Ofce www.smithsonianof.com USNM Diptera slide collection at MSC

Contact the current research staf SEL: Allen L. Norrbom [email protected] SI: Torsten Dikow [email protected] @TDikow #asiloidfies #USNMDiptera Smithsonian Credits photographs by S. Marshall, illustration by T. Litwak, Williston photograph from Aldrich (1918), and Williston logo by B. Muller.

Samuel Wendell Williston (1852 – 1918) fyer version 2, 2016-09-09 Collection Te collection comprises some 3,200,000 pinned specimens representing some 55,900 species, more than 600,000 slide-mounted specimens, and some 26,000 primary types. A genomic collection with currently more than 2,000 specimens preserved in liquid Nitrogen is growing steadily. Although there is material from all biogeographical regions, the New World is best represented. All of the 159 currently recognized families of Diptera (sensu Pape et al. 2011) are represented except for Inbiomyiidae and . A species inventory is being fnalized and soon a list of species with country data for the pinned collection will be available on our Curtonotum trypetipenne Hendel, 1913, Ecuador departmental website. Pantophthalmus planiventris (Wiedemann, 1821), Bolivia Organization Acquisitions that greatly expanded the collection are Highlights of the collection those of C.P. Alexander (1,600,000, 1981), P.H. Arnaud, Te Diptera Collection at the Department of “”: crane fies (Limoniidae and Tipulidae, Jr. (700,000, 2000), A.L. Melander (250,000, 1961), Entomology, National Museum of Natural History coll. of C.P. Alexander), mosquitoes (Culicidae, J.N. Belkin (200,000, 1980), S.W. Bromley (35,000, (NMNH), Smithsonian Institution in Washington, DC, collections of J.N. Belkin, J.P. Duret, L.E. Rozeboom, 1955), A.E. Pritchard (27,000, 1962), J.P. Duret (14,000, is one of the fnest in the world. Te ofcial abbreviation SEL dipterist A. Stone, WRBU dipterists), biting midges 1990), and L.E. Rozeboom (12,000, 1998) as well as for citing this museum as a repository is USNM (, SEL dipterist W.W. Wirth), and gall collections of C.H.T. Townsend (), R.H. Painter (referring to the former United States National Museum). (), and C.F. Baker (1928, Philippine Diptera). midges (, SEL dipterist R.J. Gagné). Te collection is administered by the Smithsonian Orthorrhapha, , and : soldier Institution, but three separate U. S. government agencies Of-site Collections Enhancement Program fies (, SEL dipterist N.E. Woodley), bee provide research and curatorial staf, i.e., the Smithsonian Te USNM provides the opportunity for established fies (Bombyliidae, coll. of R.H. Painter), assassin fies (SI), the Systematic Entomology Laboratory (SEL) of curators at other natural history museums to take over (, coll. of S.W. Bromley, A.E. Pritchard, SI dipterist the Agricultural Research Service, U. S. Department of the curatorial obligations of entire families at their T. Dikow), dance fies (, coll. of A.L. Melander, Agriculture, and the Walter Reed Biosystematic Unit home institution. For Diptera, the following families are P.H. Arnaud, Jr.), and fower fies (Syrphidae, SEL dipterist (WRBU) of the Walter Reed Army Institute of Research, part of the Of-site Enhancement Program and are not F.C. Tompson). U. S. Department of Defense. housed at the NMNH: Bombyliidae and Mythicomyiidae : Te three agencies have a diferent mission: SEL at BPBM (N. Evenhuis), at CNC (J. fruit fies (, SEL dipterists dipterists study primarily taxa of agricultural Skevington), and , , and R.H. Foote, A.L. Norrbom, A. Stone), eye gnats importance and provide identifcation services, WRBU at CSCA (S. Gaimari). Please contact these (, SEL dipterist C.W. Sabrosky), snail- dipterists focus on medically important taxa, and SI colleagues for access to specimens. killing fies (, SEL dipterists L.V. Knutson, dipterists are not bound to study applied taxa. G.C. Steyskal), lauxaniid fies (), kelp fies (), sap fies (), as well as surf Te SI and SEL dipterists are located at the NMNH in fies and shore fies ( and , SEL downtown Washington, DC, but the WRBU dipterists dipterist W.W. Wirth, SI dipterist W.N. Mathis). are working a few miles southeast of Washington, DC at the Museum Support Center (MSC), which also houses : tachinid fies (, USDA dipterists all of the medically important biting fy families as well D.W. Coquillett and C.H.T. Townsend, SEL dipterists C.W. as , Limoniidae, and Tipulidae. Tese Sabrosky and N.E. Woodley, SI dipterist J.M. Aldrich, coll. latter three families are currently “de-activated” and of P.H. Arnaud, Jr.), fesh fies (Sarcophagidae), house fies accessible through visits and loan of type material only. (), and blow fies (, USDA dipterist D.G. Hall, SI dipterist J.M. Aldrich). Te responsibilities for curation and loan requests are divided among the three agencies and a current list of the Field work by D.W. Coquillett, C.H.T. Townsend, J.R. contact people to obtain a loan or other information can Malloch, A. Stone, W.W. Wirth, C.W. Sabrosky, R.J. be found on the Department of Entomology website. Gagné, G.C. Steyskal, W.N. Mathis, F.C. Tompson, N.E. Oxycera fumipennis (Kertész, 1923), India Woodley, and A.L. Norrbom added notable taxa.