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Neuroptera of Arabia: Fam. Sisyridae, Hemerobiidae, Chrysopidae (Part 2) and Myrmeleonidae (Part 3)

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Neuroptera of Arabia: Fam. Sisyridae, Hemerobiidae, Chrysopidae (Part 2) and Myrmeleonidae (Part 3) Neuroptera of Arabia: Fam. Sisyridae, Hemerobiidae, Chrysopidae (Part 2) and Myrmeleonidae (Part 3) H. Holzel Abstract: For the first time species of the families Sisyridae and Hemerobiidae are recorded from Saudi Arabia. Supplements to the annotated lists of Chrysopidae and I\/lyrmeleonidae are made; nine species new to science, one Flemcrobiitlae, three Chrysopidae and five Myrmeleonidae, are described and illustrated. ICeywords: Neuroptera, Sisyridae, llcmerobiidae, Chrysopidae, Myrmeleonidae, taxonomy, zoogeogr;~phy ~emerobiidae%I '-Yl .% Sisyridae +Yl ,&dl : i;%b '&awl '-+I '-1 4 '-$I ij- Nyrrneleonidae JLJI &Ig Cilrysopidae 'I+! -91 wg INTRODUCTION The attempt at a comprehensive annotated list of the Neuroptera of Saudi Arabia is supplemented here, introducing the results of further collecting and studies. Two families, not hitherto noted in Saudi Arabia, are now included: Sisyridae represented by a single, and Hemerobiidae by seven species. This increases the number of Planipennia to 156 species known to occur in Saudi Arabia. The terminology of the wing venation and of genital structures as used herein follows rZsl~)cI\ et al. (1980). Abbreviations: A: anal vein C: costa Cua: anterior branch of the cubitus Cup: posterior branch of the cubitus KAU-NHMB: King Abdulaziz University - Naturhistorisches Museum Base1 expedition to N Hijaz, 1979 FAUNA OF SAUDI ARABIA 9.1988 Mp: posterior branch of the media NHMB: Naturhistorisches Museum Base1 R: radial vein Rs: radial sector Sc: subcosta Fam. Sisyridae This very small family has about 50 described species. These are distributed all over the inhabited world and were recorded from most European countries, too. However, reports from the arid and semiarid parts of northern Africa and the Arabian peninsula are scarce, because the larvae are aquatic and feed on freshwater sponges. So far, only two species have been described from this region: Siyra nilotica in Sudan and Sis_yra tdobata in Palestine. The following represents the first evidence of sisyrid spe- cies in Saudi Arabia. Sisyra nilotica Tjeder, 1957 SYsyru terminuIis. - Esben-Petersen (nec Curtis), 1915: 83. Sisyru niloticu Tjeder, 1957: 161. Material: Saudi Arabia: 1 d, Wadi 'rurabah, Camp 2, 1580 m alt., 7.X.1979, W. Biittiker. With some hesitation EsBEN-P~:I'I-:KsEN(1915) listed the species from southern Sudan as S. terminalis Curtis; TJ~IIEK(1957) stated that these specimens did not belong to 3. terminalis - a species known from Europe only - and proposed a new name, S. nilotica. I have examined one male paratype from Bahr el Ghazal and found that the Arabian specimen belongs to the same species. Figs 1 and 2 show the male terminal segments and should serve to distinguish this from the other species of the region. BUTTIKER(1980) characterized the sampling site as follows: "Over long distance permanent river with fishes, amphibians and rich aquatic insect life. Also on hillsides considerable growth of trees, bushes and shrubs." Distribution: Saudi Arabia, Sudan. Fam. Hemerobiidae This family comprises about 600 described species distributed all over the inhabited world with dis- tinct centres in the arborean parts. Only a few records are known from the Arabian peninsula, none from Saudi Arabia. The available material consists of 23 specimens belonging to seven species. 'Three of them, one Wesmaelizis sp. and two Hemerobizrs spp., each represented only by a single female, cannot be identified. One Wesmaelitrs sp. proved to be new and is described as follows. Wesmaelius (Kimminsia) saudiarabicus n. sp. Holotype: 8,Saudi Arabia, Village Qaraah, 2000 malt., 16.IV.1976, W. Hiittiker (NIIMB). - Paratype: 1 P, same data as holotype. A small, pale yellowish Wesmaelizrs species, length of forewings 5.5-6 mm. Head pale yellowish with small fuscous spots laterally on clypeus; vertex brownish posteriorly. Thorax: pronotum with fuscous spots laterally on scutum, and brown lines along the sutures between scutum and prescutum; metanotum fuscous laterally on scutum. Legs entirely pale. Wings: membrane hyaline, venation pale; three branches of Rs in forewings. Abdomen pale yellowish with lateral bands of fuscous spots. Apex (8) as in figs 3 and 4; ectoprocts bandlike with a rather long apex; rows of lamel- lae rather long; gonarcus (figs 6 and 7) with a long mediuncus which is bent downwards; entoprocessus rather long with oblique apex. Parameres (figs 8 and 9) very long and slender, slightly curved and fused apically. Apex of 9 abdomen see fig. 5; subgenitale and spermatheca are very weakly sclerotized and can- not be figured. The genital structures of W. saudiarabicui n. sp. resemble very much those of W. nubihs (Kimmins); both the ectoprocts and the gonarcus are confusingly alike. However, W. nubih is a dark brown species with heavily spotted forewings, and, so far, only known from South Africa and Angola (TJIIII*I~ 1961); confusions are therefore not likely to occur. From W. navasi (Andreu), the only other Wesmaelius-species of the region, W. sazldiarabiczls can easily be distinguished by its pale yellowish colouration. Figs 1-9: 1-2, Siya nilotica Tjeder; 1, apex of 8 abdomen, side view; 2, same, caudal view; 3-5, Wesmaelius (Kimminsia) suudtirrubicus n. sp.; 3, apex of abdornen, holotype 8, side view; 4, same, apex of ectoproct, inside; 5, apes of abdomen, paratype P, side view. Scale: 0.5 mm. 6-9, Wesmuelius (Kimminsia) saudiarabicus n. sp., holotype 8; 6, gonarcus, dorsal view; 7, gonarcus, side view; 8, parameres, dorsal view; 9, parameres, side view. Scale: 0.25 mm. FAUNA OF SAUDI ARABIA 9,1988 Wesmaelius (Kimminsia) navasi (AndrCu, 19 11) Buriomyia navasi Andrttu, 1911: 58. Wesmaelius (Kimminsia) navasi (AndrCu). - Aspock et al. 1980: 205. Material: Saudi Arabia: 1 8, Wadi Salbukh, 15.1V.1977; 1 P, Hofuf, 24.V.1978, W. Buttiker. Distribution: Spain, Greece, Malta; Madeira, Canary Islands, Morocco; Anatolia, South Russia, Saudi Arabia, Iraq, Lebanon, Palestine, Iran, Afghanistan, Pakistan (Asi~~cltet al. 1980). Micromus sjoestedti Weele, 1910 Micrumus sjoestedti Weele, 1910: 18. Material: Saudi Arabia: 1 8, Wadi Dhi Khul, 20.11.1980, W. Biittiker. TJEDER(1961) has examined the type specimen from Tanzania and published figures of the 8 and P genital structures. I have no doubts that the Arabian specimen belongs to this species, which in the meantime also has been recorded from the Cape Verde Islands (OHM& HOLZEL1982). Distribution: Saudi Arabia; South Africa, Zimbabwe, Uganda, Tanzania, Cape Verde Archipela- go. Sympherobius (Sympherobius)fallax Navas, 1908 .liynpberubiusfallax Navas, 1908: 408. Material: Saudi Arabia: 15 exs, Najran, 20.1X.1978, W. Riittiker. Distribution: Saudi Arabia, Palestine, Lebanon, Turkey; Spain, Greece, Malta; Canary Islands, Madeira, Morocco, Egypt, Sudan, Ethiopia (ASPOCKet al. 1980); Cape Verde Archipelago (OI~I& H~I~ZEL1984). Fam. Chrysopidae In HOI.ZEL(1980) 17 species were recorded from Saudi Arabia. Due to the receipt of new material, the total can now be increased to 23; 3 species are described as new. Subfam. Chrysopinae Tribe Italochrysini ltalochrysa pittawayi n. sp. Iiolotype: P, Saudi Arabia, Muhayl (Asir), 11.1V.1983, A.R. Pittaway (in coll. Holzel). A relatively small, stout Italocbr_~a-specieswith spotted wings; length of forewings 13.5 mm, hindwings 12 mm. Head yellowish without spots. Palpi and other mouthparts of the same colour. Antennae stout and rather short (distinctly shorter than forewings); scape and pedicel yellow, flagellum black. Thorax yel- low. Pronotum with broad reddish brown lateral margins; mesonotum: brown spots on prescutum along the anterior margin, over the sutures between prescutum and scutum and between scutum and scutellum; brown patches laterally on scutum. Metanotum: brown spots forming broad lateral bands, and a roundish spot on anterior margin of scutellum. Wings: membrane hyaline with blackish spots as shown in fig. 10. Pterostigma elongate, pale brownish. Forewings: intramedian cell triangular; C and Sc entirely pale, all other longitudinal veins pale at base and dark brown in apical half; base of C with a small brown spot, base of R with a fine brown line. Crossveins: costals near the base of the wings brown, in apical half pale; all other crossveins dark brown. Hindwings: all veins near the base of wings pale, gradually darkening apically. Hairs on veins dark brown, marginal fringes short, black. Legs yellowish with short dark hairs, brownish on apical parts of femora and tibiae; tarsi brown, claws with basal tooth. Abdomen reddish brown with yellow patches laterally and ventrally; hairiness short, pale. Apex as shown in fig. 11: sternite 7 at apex square with a broad list-like apodeme along the posterior margin; spermatheca and subgenitale as in figs 12 and 13. Fig. 10: Italocbtysapittawayi n. sp., holotype P. I. pittaw+ n. sp. can be distinguished from all other known Italochya-species of the region by the markings on head and pronotum (head unspotted, pronotum with broad reddish brown lateral bands) and, particularly, by the striking broad apodeme along the hind margin of sternite 7. 'I'he triangular intramedian cell is quite unusual within this genus, but, as I had only one single female at hand, I am not able to decide whether this represents the standard pattern or not. The species is dedicated to its collector, A.R. Pittaway, London. Italochysa arabica n. sp. Holotype: 9, Saudi Arabia, Wadi Shuquh, 1440 m alt., 6.1V.1980, W. Biittiker (NHhfH). - Paratype: 1 9 in alcohol, sarne data as holotype. A medium-sized Italochrya-species with spotted wings; length of forewings 16 mm, hintlwings 15 mm. Head yellowish with broad areas of reddish brown on vertex, between the antennae, on genae and Figs 11-17: 11-13, ItaLocb5ysapittawayi n. sp.; 11, apex of abdomen, holotype P, side view; 12, spermatheca, side view; 11, apex of abdomen with suhgenitale, ventral view; 14-17, ItaIocbyu arabica n. sp.; 14, apex of abdomen, holotype P, ventral view; 15, same, side view; 16, spermatheca, side view; 17, claw of foreleg. Scale: 0.5 mm.
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