Chironomus Cornpertus, Disclusus, Min~Isculus and Morulus), Has Been Found in the British Museum (Nat

Home , Acricotopus, Diamesini, Krenopelopia, Paramerina, Potthastia, Rheopelopia

Eltsabeth Stur

MlTTElLUN GEN AUS DE R CHIRONOMIDENKUNDE NEWSLETTER OF CH l RONOMID RESEARCH

A LISr' OF TYPES OF P.~LI\E~RCTICCHIRONONID:\E

IN THE RRITISlI MUSEUM (NATUR4L HISTORY). by P.S. Cranston

The earliest British worker to publish on the Chironomidae was F. Walker (1856) who recorded 297 species, including 82 now recognised as Cerntopogoni- dae. Of the remaining 215 species 99 ~vere new to science, and type-material of all but 4 species, (Chironomus cornpertus, disclusus, min~isculus and morulus), has been found in the British Museum (Nat. Hist.). .4 syntype of Chirononus moturus Walker was rediscovered in the preparation of this list, and the name is synonymised with Acricotopus lucens Zetterstedt. In the list Walker's types are referred to as Holotypes unless there is definite evldence from either the collection pr his description that he had a syntypic series. In some cases, marked with an asterisk, (+I,the syntypic series is mixed, With the exception of Verrnll's list (1901), little taxonomic work was undertaken until F.W. Edwards, who published widely from 1919 until 1941. In his major work on the British'~id~esEdwards (1929) described 87 new species and 1 new named variety, and some type-material of all of these, with the exception of Metriocnemus grimshawi, has been found in the Museum. Of equal im- portance was Edwards' recognition of Walker's species; this interpretation is largely followed in this list. In many cases workers have incorrectly regarded specimens which Edwards had separated from his type-series as holotypes; however Edrvards often did not designate a holotype, and these series are therefore syntypic, as pointed out in the list. In this list type localities have been reduced to the country only, but further data for all except Walker's species is available. The current name for the species follows Cranston in Kloet and Iiinclcs (1976) for the British fauna, and a variety of works for the non-British species. Further type-material of Palaearctic Chironomidae may be found in the col- lections of the Hope Department, University Museum, Oxford, The Royal Scottish Museum, Edinhurgh and The National Museum of Ireland, Dublin. It is hoped that a study of this material may be made in the future. abiskoensis Xdwards, 1937a:144, (~rthocladius) Orthocladius nbiskoensis Syntypes 26 (1 slide, 1 spirit). Sweden Edwards ad junctus !Talker, 1656 : 190, (~hironomus) Metriocnemus fuscipes Pieigen Holotype 9. England. aff inis Walker, 1856: 162, (~hironomus) ,Paratanytarsus tenuis Meigen Holotype 9. England. alligatus Walker, 1856 :190, (~hironornus) Metriocnemus fuscipes Meigen Ilolotype d. England. angusta Edwnrds, 1929:364, (~~aniotoma) Pseudoomittiii angusta Edwards Syntypes 9d, England, Wales. apicalis Kief'f er, :!;1:806, (Netriocnemus) Heterotanytarsus apicalis Syntypes 2d (slide). Czechoslovakia. Kief f er appositus Walker, 1856 : 191, (Chironornus ) Micropsectra apposita Walker Holotype d. England. barbimanus Edwards, 1929:373, (spaniotoma) Psectroclndius barbimanus Holotype d. England. Edwards belingi Fittkau, 1954:84,(Pseudodiamesa) Pseudodiamesn braniclci Nowick Paratypes (18 slide, 19 spirit). Germany. bertrami Edwards, 1975:470, (~iarnesa) Diamesn bertrnmi Ed~vards Holotype 6. Greenland. betuletorum Edwards, 1941:226, (~inittia) Srnittia betuletorum Edwards Syntypes %d (slide). Sweden. biformis Eduvnrds, 1929:325, (~ricoto~us) Cricotopus ? fuscus Kieffer Holotype 6. England. bituberculatus Edwards, 1929:408, (Tan~tarsus) Parntanytnrsus bituberculatus Synt~pes96, 19. England. Edwards bohemnni Goetghebuer, 1932: 18 1, (~iamesa) Dismesn bohernnni Edwards Lectotype d (fixed Edwiprds, 1977:617). SpTt zbergen. borealis F. souamatus Andersen, 1937:70, Limnophyes ? minimus Meigen (Limnophyes) Syntype 8. Greenland. brachylabis Edwards, 1929:406, (~hironomus) Paralnuterbvrniella Holotype 6. England. nigrohnlternlis Malloch brayi Goetghebuer, 1921: 177, (~hironornus) Nilothnuma brayi Goetghebuer Syntype d. Belgium. brevicosta Edwnrds, 19372:146, (Tanytarsus) Phnenopelnn brevicostn Edwards Syntypes (36 slide). Sweden. brevifurcata Edwnrds, 1926:781, (Spaniotoma) Orthosrnittia brevifurcata Syntypes 156. England. Edwards brevis Edwards, 1929:420, (Tanytnrsus) Stempellinelln brevis Edwards Syntypes 226, h? (18 slide). England, Ireland, Scotland. brevitarsis Edrvards, 1929:716, (Metriocnemus) Gymnometriocnemus brevitarsis Holotype 9. England. Edwnrds britteni Edwnrds, 1929:399, ,(Chironomus) Microtendipes britteni Edwards Syntypes 98. England. brumnlis Edwnrds, 1929: 316, (Metriocnemus) Gymnornetriocnemus brumalis Syntypes 36, 49. England. Edwnrds calcnrata Edwards, 1929:733, (Spaniotoma) Psectrocladius cnlcaratus syntypes 36, England. Edwards caledonicus Edwnrdif.*1929: 47, (~hirono!nus) Microtendipes cnledonicus Syntypes 3b, 29 (18 slide). Scotland. Edwards calvescens EdrvarBs, 1929:753, (Spaniotoma) Eukiefferiella calvescens Syntypes ljs, 19 (33 slide). England. Edwards campestris Edwnrds, 1929: 507, (Diamcsn) Potthastia longimana Kieffer Syntypes yd. EnglanA, Scotland. camptophleps Edrvards , 1929: 357, (~~a~ziotona) Krenosmitt ia cnrnpt ophleps Syntypes 18, 19. England. Edwards cnrriana Edwnrds , 1924b : 189, (cor~no~?eura) Corynoneurn carriana Edwards Syntypes ?d, 19. England. caudntws Edwnrrls , 1924~:122, (krotanypus ) Protnnypus caudntus Edrvards Syntypes 66, 4?. Nor~rny. celtica Edwards, 192kb: 186, (Corynoncura) Corynoneura celtica Edwards .Holotype 8. Wales. 30 Parntypes. England, lfales. chalybenta Edwards, 1929:331, (~~aniotoma) Rheocricotopus chalybeatus Syntypes 108, 59. England. Edwards - 139 7 cheethami Edwards, 1929:359, (~paniotoma) Parakiefferiella bathophila Holotype 6, Paratypes 26, 19. England. Kief fer chionophila Edwards, 1933b:90, (Tanytarsus) Parapsectra chionophila Edwards Syntypes l&, 29. Scotland. cinctipes Edwards, 1928: 171, (~eptag~iaj- Boreoheptagyia cinctipes Syntypes 10d. Corsica. Edwards 'cingulatus Walker, 1856 : 172, (Chironomu-s) Paramerina cingulata Walker Holotype b. England. clavata Edwards, 1933a:615, (~iamesa) Diamesa clavata Edwards Syntypes lb, 19. N. Canada. claviforceps Edwards, 1929:389, (Chironomus) Parachironomus claviforceps Syntypes 26. England. Edwards coaecluatus Walker, 1056 : 190, (Chironomus ) Acricotopus lucens Zetterstedt Holotype 9. England. coerulescens Kief fer, 1926:279, (~richocladius) Eukiefferiella coerulescens Syntype (id, pupa, larva on slide). Kieff er Czechoslovakia. conformis Holmgren, 1869:42, (~hironomus) Orthocladius conformis Syntypes 18, Spitzbergen. Holmgren conjuncta Edwards ,'p~29: 365, (~~aniotoma) Pseudosmittia con juncta Syntypes 6~.England. Edwards contingens Walker, 1856 : 191, (Chironomus ) Smittia contingens Walker Syntypes 28. England. convectus Walker, 1856: 171, (Chironomus ) Einfeldia dissidens Walker Holotype d. England. conversus Walker, 1856:175, (~hironomus) Paracladius conversus Walker Syntype 9. England. convestitus Walker, 1856:177, (~hironomus) Prodiamesa olivacea Meigen Holotype b. England. convictus Walker, 1856 : 161, (~hironomus) Polypedilum convicturn Walker +syntypes 26. England. Parachironomus parilis Walker coronata Edwards, 1924b:187, (~or~noneura) Corynoneura coronata Edwards Holotype 8, Paratypes 66, 79. England. coronata Edwards, 1929:354, (~~aniotoma) Parakiefferiella coronata Paratype 8 (no abdomen). Wales. Edwards crassimanus Strenzke, 1959:24, (Chironomus) Chironomu's crassimanus Paratypes id, 1 (spirit). West Germany. Strenzke cuneatus Edwards , 19x9: 3 15, (Metriocnemus ) Paraphaenocladius cuneatus Syntypes 29 (l only thorax). England. Edwards cuneipennis Edwards, 1929:421, (Tanytarsus) Stempellinella cuneipennis Holotype 9. England. Edwards curticosta Edwards , 1929 :364, (Spaniotoma) ~seedosmittiacukticosta Syntypes 26. Scotland, Wales. Edwards davisi Edwards, 1933a:614, (Diamesa) Diamesa davisi Edwards Syntypes 26, 19. N. Canada. denotatus Walker, 1856: 190, (Chironomus ) Paracladius conversus Walker Syntype d. England. dentiforceps Edwards, 1929:346, (Spaniotoma) Chaetocladius dentiforceps Syntypes 5&, 59. England. Edwards deproperans Walker, 1856:188, (Chironomus) Metriocnemus fuscipes Meigen Holotype Q. England. deribae-Freeman, 1957:395, (Chironomus) Leptochironomus deribae Holotype 8, Paratypes lb, 29: Sudan. Freeman devonica Edwards, 1929:349, (Spanlotoma) Eukiefferiella ,devonica Holotype d (slide). England. Edwards diffinis Edwards, 1929:397, (Chironomus) Microtendipes diffinis Syntypes 43, 29. England. Edwards ,digitalis Edwards, 1929:389, (Chironomus) Parachironomus digitalis Syntypes 28, England, Scotland. Edwards discrepans llalker, 'p656 : 169, (Chironomus) Microtendipes chloris Meigen Holotype b. England. d\ispessus Walker, 1856 : 16 1, (~hironomus) Limnochironomus pulsus Walker Syntype c?. England. dissidens Walker, 1856:154, (Ehironomus) Einfeldia dissidens Walker Holotype b. England. dissipata Edwards,1929:378, (S~aniotoma) Chaetocladius dissipatus Syntypes 36, 19. England. Edwards divisus Walker, 1856 :192, (~hironomus) Paramerina divisa Walker Holotype b. England. dolens Walker, 1856 :178, (Chironomus) Chironomus pilicornis Holotype b. England. Fabricius effusus Walker, 1856: 180, (Chironomus Rheocricotopus effusus Walker. - +syntypes 26. England. Paraphaenocladius impensus Ifalker ejuncidus Walker, 1856:155, (Chironomus) Tanytarsus ejuncidus Walker . Holotype b. England. eltoni Edwards, 1922:203, (~am~tocladius) Limnophyes eltoni Edwards Syntypes 48, 19. Bear Island. eltoni f. asquamatus Andersen, 1937:71, Limnophyes asquamatus (Limnophyes) Syntype b. Greenland. Andersen (see Saethar, 1975: 1030 eminulus Walker, 1856:156, (Chironomus) Tanytarsus eminulus Walker Holotype 9. England. enotatus Walker, 1856:170, (Chironomus) Polypedilum nubeculosum Holotype d. England. Mevigen excavatus Edwards, 1929:416, (Tan~tarsus) Tanytarsus excavatus Edwards Syntypes 58. England, Wales. excerptus Walker, 1856 : 179, (Chironomus ) Paratrissocladius excerptus Holotype d. England. Wallcer eximia Edwards, 1929:290, (Pentaneura) Rheopelopia eximia Edwards Paratype 9. cotla land). expalpans Walker, 1856 : 18 1, (~hironomus) Arctopelopia barbitarsis Holotype b. England.. Zetterstedt exspatians Walker,i856:182, (Chironomus) Psectrocladius sordidellus Holotype 9. England. Zetterstedt feminea Edwards; 1929: 343, (Spaniotoma) Bryophaenocladius feminea Holotype b. England. Edwards fertus Walker, 1856:192, (Chironomus) Metriocnemus fuscipes Meigen Syntype b. England. festivus Holmgren, 1869:43, (Chironomus) Orthocladius festivus Syntypes ld, 19. Spitzbergen. Holmgren flavescens Edwards, 1929:367, (Corynoneura) Thienemanniella flavescens Holotype 6, Paratypes 26. England. Edwards flavidulus Edwards, 1929:420, any-tarsus) Stempellinella flavi,dula Syntypes 38, 29 (2d slides) England. Edwards flavifrons Edwards, 1929:302, (~rocladius) Psilotanypus flavifrons Syntypes 49. England. Edwnrds flexuella Edwards, 1929:349, (~~aniotoma) Mes~gmittiaflexuella Syntypes 53. England. Edwards foveata Edwards, 1929:330, (~~aniotorna) Rheocricotopus atripes Kieffer Syntypes 56, 59. ( 18 slide). England, Scotland. fucicola Edwards, 1926:782, (Trichocladius) Halocladius fucicola Edwards Holotype d, Paratypes 12&, 29. England, Scotland, Wales. furcata Edwards, 1933a:617, (~iamesa) Diamesa furcata Edwards Syntypes Id, 39. N. Canada. fusciceps Edrvards , 1929 :290, (Pentaneura) Thienemannimyin fusciceps Syntypes 26. England. Edwards. fuscihalter Edwards, 1929:369, (Cor~noneura) Corynoneura fuscihalter Synt~esz!. England. Edwards futilis Walker, 1856: 156, (chironomus) Limnochironornus nervosus ? Holotype d" (described by Wallcer as Staeger England. 8) glabrescens Edwards, 1929:416, (Tan~tarsus) Tanytarsus glabrescens Holotype b. England. Edwards ' glacialis Edrvards, 1922:209, (Cricotopus) Cricotopus glacialis Edwards Lectotype d (~irvenoja1973:284), Para- lectotypes 913,79. Spitzbergen. gotchi Goetghebuer, 1928:143, (Tan~tarsus) Tanytarsus heusdensis Holotype d. England. Goetghebuer . gracei Edwards, 1929:712, (~etriocnemus) Metriocnemus gracei Edwards Syntypes 26. England. gracei Edwards, 1929:346, (Spaniotoma) Eukiefferiella gracei Syntypes 26. England. Edwards divisus Walker, 1856: 192, (Chironomus) Paramerina d ivisa Walker Holotype b. England. dolens Walker. 1856:178, (~hironomus) Chironomus pilicornis Holotype d. England. . Fabricius effusus Walker, 1856: 180, (Chironomus ) Rheocricotopus effusus Walker *Syntypes 26. England. Paraphaenocladius impensus Walker ejuncidus Walker, 1856:155, (Chironoms) Tanytarsus ejuncidus Walker . Holotype d. England. eltoni Edwards, 1922:207, (Camptocladius) Limnophyes eltoni Edwards Syntypes 46, 19. Bear Island. eltoni f. asquamatus Andersen, 1937:71, Limnophyes asquamatus (Limnophyes) Syntype d. Greenland. Andersen (see Saether, 1975: 1030) eminulus Walker, 1856 : 156, (Chironomus) Tanytarsus eminulus Walker Holotype p. England. enotatus Walker, 1856:170, (~hironomus) Polypedilum nubeculosum Holotype d. England. Meigen excavatus Edwards, 1929:416, anyta tars us) Tanytarsus excavatus Edwards Syntypes 5d. England, Wales. excerptus Walker, 1856:179, (Chironomus) Paratrissocladius excerptus Holotype d. England. liallcer eximia Edwards , 1929: 290, (Pentaneura) Rheopelopia eximia Edwards Paratype 9. (Scotland). expalpans Walker, 1856:181, (~hironomus) Arctopelopin barbitarsis Holotype d. England; Zetterstedt exspatians Walker,1856:182, (~hironomus) Psectrocladius sordidellus Holotype 9. England. Zetterstedt feminea Edwards; 1929:343, (Spaniotoma) Bryophaenocladius feminea Holotype 6. England. Edwards fertus Walker, 1856 :192, (Chironomus) Netriocnemus fuscipes Meigen Syntype d. England. festivus Holmgren, 1869:43, (Chironomus) Orthocladius f est ivus Syntypes ld, 19. Spit zbergen. Holmgren flavescens Edwards, 1929:367. (~ory-noneura) Thienemanniella flavescens Holotype 13, Paratypes 26. England. Edwards flavidulus Edwards, 1929:420, (Tan~tarsus) Stempellinella flavi,dula Syntypes 38, 29 (26 slides) England. Edwards flavifrons Edwards, 1929:302, (~rocladius) Psilotanypus flavifrons Syntypes 49. England. Edwards flexuella Edwards , 1929: 349, (Spaniotoma) Mesappittia flexuella Syntypes 56. England. Edwards foveata Edwards, 1929:770, (Spaniotoma) Rheocricotopus atripes Kieffer Syntypes 56, 59. (l6 sllde). England, Scotland. fucicola Edwards, 1926:782, (Trichocladius) Halocladius fucicola Edwards Holotype 8, Paratypes 128, 29. England, Scotland, Wales. furcata Edwards, 1937a:617, (~iamesa) Diamesa furcata Edwards Syntypes 18, 39. N. Canada. fusciceps Edrvards, 1929: 290, (~entaneura) Thienemannimyia fusciceps Syntypes 26. England. Edwards . fuscihalter Edwards, 1929:369, (~or~noneura) Corynoneura fuscihalter Syntves 2b. England. Edwards futilis Walker, 1856: 156, (~hironoms) Limnochironomus nervosus ? Holotype 8 (described by Wallcer as Staeger England. P' glabrescens Edwards, 1929:416, (Tanytarsus) Tanytarsus glabrescens Holotype b. England. Edwards glacialis Edwards, 1922:209, (~ricoto~us) Cricotopus glacialis Edwards Lectotype S (Hirvenoja 1973:284), Para- lectotypes 96, 79. Spitzbergen. gotchi Goetghebuer, 1928: 143, (~an~tarsus) Tanytarsus heusdensis Holotype b. England. Goetghebuer gracei Edwards, 1929:312, (~etriocnemus) Metriocnemus gracei Edwa-rds Syntypes 25. England. gracei Edwards, 1929:746, (Spaniotoma) Eukiefferiella gracei Synty-pes 26. England. Edwards lacnum Edwards, 1929:324, (~ricoto~us) Cricotopus ephippium Syntypes 48, 39. England, Scotland. Zetterstedt laricomalis Edwards, 1932:46, (Cricotopus) Cricotopus laricornalis Syntypes 36, Scotland., Edwards latiforceps Edwa:y. 1941:235. (Tanytarsus) Not recognised with certainty . Syntypes id, l?. (hypopygium of 8 ~oorly Reiss & Fittkau,' mounted.) Sweden. 1971:81 lentulus Walker, 1856:178, (~hironomus) Microtendipes confinis Meigen Syntype d. England. lestralis Edwards, 1924a:172, (~ricoto~us) Cricotopus lestralis Edwards Lectotype 8 (~irvenoja, 1973: 1531, Paralectotypes h#,' l?. Spit zbergen. lobata Edwards , 1924b : 186, (~or~noneura) Corynoneura lobata Edwards Holotype b, Paratypes 38, 2y. England. longicosta Edwards, 1922:201, (~amptocladius) Smittia lengicosta Edwards ' Syntype 49. Bear Island. luridus Strenske, 1959:26, (chironoms) Chirnnomus luridus Strsnzke Paratypes 28 (spirit). West Germany. ' luridus f. semicinctus Strenzke, 1959227, Chironomus luridus (Chironomus) Paratypes 18, 19 (spirit). f. semicinctus Strenzke West Germany. lutea Edwards, 1924b: 184, (~orynoneura) Thienemanniella lutea Edwards Holotype 8, Paratypes 96, 39. England. lygropis Edwaeds, 1929:325, (C~ICO~OPUS) Cricotopus lygropjs Edwards Holotype 9. England. macani Freeman, 1948:50, (Chironoms) Einfeldia macani Freeman Holotype d, Paratypes 29. England. majuscula Edwards, 1924b:185, (Cor~noneura) Thienernanniella majuScula Holotype J,.Paratypes 7d, 39. England. Edwards malacus Walker, 1856: 167, (~hironomus) Parachironomus biannulatus Syntype 9. England . Staeger mancunianus Edwards, 1929:393, (Chironomus) Glyptotendipes mancunianus Syntypes 46. England. Edwards mancus Walker, 1856 :161, (~hironomus) Cladotanytarsus mancus Walker Holotype d. England. marcidus Walker, 1856:177, (~hironomus) Heterotrissocladius karcidus Holotype b. England. Walker minor Edwards, 1929: 748, (~~aniotorna) Eukiefferiella minor Edwards Syntypes 56. England. minor Edwards, 1929:420, (~an~tarsus) Stempellinella minor Edwards Syntypes 40, 10 (slide). England, Wales. modicellus Walker, 1856: 162, (Chironomus ) Cladotanytarsus mancus Walker Holotype 9. England. moerens Walkey, 1856 :178, (Chironomus) Chironomus pilicornis Holotype d. England. Fabricius monticola Edwards, 1929:408, (Tan~tarsus) Parapsectra nana Meigen Holotype 8, Paratypes 18, 19. Scotland. montium Edwards, 1929:307, (~iamesa) Potthastia montium Edwards Syntypes 36. England, Scotland. morosa Edwards, 1924b:185, (Corynoneura) Thienemanniella morosa Edwards Holotype d. England. motitator f. subcoeruleus Edwards, 1929:727, Cricotopus annulator (Cricotopus) Syntypes 58, 69. England. Goetghebuer - moturus Walker, 1856 : 188, (Chironomus ) Acricotopus lucens Syntype d (slide). England. Zetterstedt. m. -. nactus Walker, 1856:179, (Chironomus) Micropsectra praecox Meigen Holotype 8. England. nemorosus Edwards, 1929:416, (Tan~tarsus) Tanytarsus nernorosus Edwards Holotype d. England. nexilis Yalker, 1856: 184, (Chironomus) Diamesa incallida Walker Holotype d. England. nidorum Edwards , 1929 :742, (Spaniotoma) Bryophaenocladius nidorum Holotype d. England. Edwards niger Andersen, 1937:40, (~an~tarsus) Tanytarsus niger Andersen Syntype d. Greenland. nigratus Walker, 1856:18P, (Chironomus) Smittia aterrirna Meigen Holotype 9. England. nigroclnctus Freeman, 1957:220, (Tan~tarsus) Tanytarsus horni Goetghebuer Holotype 6, Paratypes 5d, 29. Uganda. nigronitens Edwards, 1929:390, (~hironomus) Cryptotendipes nigronitens Syntypes 26. England. Edwards nitidlcollls Walker, 1856:187, (Chironomus) Lkcricotopus lucens Zetterstedt , Holotype 6. England. northumbrica Edwards, 1929:291, (Pentaneura) Thienemannimyia northumbrica Syntype b. England. Edwards notescens Walker, 1856: 156, (Chlronomus) Micropsectra praecox Meigen , Holotype b. 4ngland. novatus Walker, 1856:184, (Chironomus) Chaetocladius psrennis Meigen Syntype 9 (no abdomen). England. nubens Edwards, 1929:376, (Penta~edilum) Pentapedilum nubens Edwards Syntypes 156,+ 49. England, Ireland, Scotland. nudisquarna Edwards, 1929:396, (chironomus) Paratendipes nudisquama Syntypes ldj 10. England, Ireland. Edwards nugax Walker, 1856:186, (chironomus) Natarsia nugax Walker Holotype d. England. obditus Walker, 1856: 181, (Chironomus) Prodiimesa olivacea Meigen Syntype d. England. ob jectans Walker, 1856 :157, (chironomus ) Limnochironomus pulsus walker Holotype d. Eqgland. obl~densWalker, 1856 : 180, (chironomus ) ~rthocladius~oblidens Walker Holotype d (silde). England. obnixus Walker, 1856:165, (Chironornus) Cricotopus obnixus Walker Holotype d. England. obreptans Walker, 1856 :168, (chironomus ) Cryptochironomus obreptans Holotype 9. Eqgland. Walker obscur~pesHolmgren, 1869:38, (Chironomus) Metriocnemus obscuripes Syntypes l&, 10. Spitzbergen. Holmgren obsepiens Walker, 1856 :183, (Chironomus) Acricotopus,lucens ZetCerstedt Syntype d. England. obsistens Walker, 1856:188, (~hironomus) Metriocnemus fuscipes Meigen Holotype b. England. obtexens Walker, 1856 :188, (chironomus ) Paracladius conversus'Walker Holotype 9. England. obvertens Walker, 1856 :177, (Chironomus ) Microtendipes confinis Meigen Syntypes 18, 19. England. obvius Walker, 1856 : 174, (Chironomus ) Psectrocladius obvius Walker Holotype 8. England. occipiens Walker, 1856:165, (Chironomus) Mic~wsectrapraecox Meigen Holotype d. England. offectas Walker, 1856:185, (Chironomus) Nicropsectra praecox Meigen Holotype d. England. olivaceus Goetghebuer, 1928:143, (Tan~tarsus) Cladotanytarsus.atridorsum. Holotype d. England. Kieffer oppertus Walker, 1856:183, (Chironomus) Psectrocladius oppertus Walker Holotype d England. oppetitus Walke , 1856:176, (Chironomus) Stictochironomus,rosenshoeldi' Holotype 6 (slide). England. Zetterstedt opplens Walker,f 1856:187, (Chironomus) Paratrichocladius rufiventris Holotype bl England. Meigen ornaticollis Edwards, 1929:359, (Spaniotoma) Heleniella ornaticollis Holotype d, Paratypes 2d, 20. England. Edwards orophilus Edwards, 1929:405, (Chironomus) Pagastiella orophila Edwards SY~~YP~S36, England, Ireland, Wales. oxonianus Edwards , 'P1922 :204, (Camptocladius ) Pseudosmittia 'oxoniana Syntypes 39. Bear Island. Edwards paganicus Walker, 1856:183, (Chironomus) Metriocnemus ? picipes Meigen Holotype 9. England. pallidicornis Wa ker, 1856:169, (Chironomus) Tanytarsus pallidicornis Syntype 6. Ingland. Walker pallidipes Edwards, 1929:324, (Cricotopus) Cricoto us pallddipes Edwards Syntypes 86, 10. England, Scotland, Wales. B parilis Walker, 1856 : 16 1, (Chironomus Parachironomus parilis Walker Syntypes ld, 19. England. paripes Edwards, 1929:392, (Chironomus) Glyptotendipes paripes Syntypes 46, 19. England. Edwards ' parva Edwards, j932: 245, (~iamesa) Pseudokiefferiella p.arva Edwards Holotype 3. Scotland. Paratypes 46, 19. England. parvibarba Edwards, 1939~259,(Smittia) Lapposmittin parvibarba Holotype d (slide). Sweden. Edward s pastoris Edwards, 1977h:88, (~iamesn) Potthastia pastoris Edwards Syntypes 106, 109. Scotland. qatens Walker, 1856: 170, (Chironomus ) Polypedilum pedestre Meigen Holotype Q. Englnnd. patibilis Walker, 1856 : 174, (Chxronomus) Metriocnemus fuscipes Meigen +Syntypes 26. Englnnd. hcricotopus lucens Zetterstedt pedestris Wollaston, 1858:114, (Cheironomus sicl) Thalassomva frauenfeldi Syntypes 96, 39. Madeira. Schiner penerasus Edwards, 1929:315, (~etriocnemus) Parnphnenocladius penerasus Syntypes 106, 29. England, Scotland, Wales. Edwnrds peregrinus Edwnrdg, 1929:296, (~odonomus) Parochlus kiefferi Gnrrett Syntypes 26. England (no abdomen), Scotland. perexilis Walker, 1856 :19 1, (Chironomus) Microcricotopus bicolor Holotype 9. England. Zetterstedt pergens Walker, 1856 : 187, (Chironomus ) Diamesa tonsa Walker Holotype d. England. perlevis Walker, 1856:158, (~hironomus) Par.~tanytarsustenuis Meigen Holotype 9. England. permacer Walker, 1856:183, (Chironomus) g Dinmesa permacra Walker +Syntypes 18, 19. England. y Chaetocladius perennis M.eigen perpessus Walker, 1856:178, (~hironomus) Prodiamesa olivacea Meigen Holotype 6. Englnnd. perreptans Walker, 1856: 177, (~hironomus), Microtendipes confinis Holotype d (no abdomen). England. Meigen persidens IfAlker, 1856: l8 l, (Chironomus ) Orthocladius rubicundus Holotype d. England. Meigen pertenuis Walker, 18561182, (Chironomus) Bryophaenocladius ictericus Holotype 9. England. Meigen pertractus Walker, 1856:188, (~hironomus) Diamesa tonsa Walker Holotype d. England. pervulsus Walker, 1856: 179, (Chironomus) Acricotopus lucens Zetterstedt Holo'type d. England. platypus Edwards, 1929:373, (S~aniotoma) Psectrocladius platypus Syntypes 46, 29. England, Scotland. Edward's polaris Kirby; 1824: 218, (? Chironomus) Chironomus polaris Kirby Syntype (thorax, l :ring). 'Ross voyage'. poultoni .Edwards, 1922:213, (~iamesa) . Diamesa arctica Boheman Syntypes 46, 19. Spitzbergen. pseudothummi Strenzke, 1959:27, (~hironomus) Chironomus pseudothummi Paratypes Id, 19 (spirit). West Germany Strenzke pulsus Walker, 1856 :165, (~hironomus) Limnochironomus pulsus walker Syntype d. England. recta Edwards, 1929:362, (Spaniotoma) Pseudosmittia recta Edwards Holotype 6, Paratypes 4d, 29. England. reflexens Edwards, 1929:411, (Tanytarsus) Tanytarsus triangularis Syntypes 86, 39. England, Scotland, Wales. Goetghebuer richmondensis Edwards, 1929:413, (Tanytarsus) Tanytarsus arduennensis Syntypes 86, 49. England, Scotland, 1Jales. Goetghebuer rydalensis Edwards, 1929:404, (Chironomus) Microtendipes rytialensis Syntypes 26. England. Edwards samboni Edwards, 1929: 416, (Tanytarsus ) Tanytarsus debilis Meigen Syntypes 2d. England. scotica Edwards, 1929:363, (Spanioto.m& - ,Pseudosmittia scotica Edwards Holotype 9. Scotland. separatus Walker, 1856:171, (Chironomus; Microtendipes conf inis.Meigen Holotype d. England. serpentinus Edwards , 1937b : 105, (Lasiodiamesa) Lasiodiamesa gracilis kieffer Holotype d (slide). Sweden. sima Edwards, 1929:342, (Spaniotoma) Bryophaenocladius simus ' Holotype d. England. Edwards simplicistilis Freeman, 1948:49, (Procladius ) Procladius simplicistilis Holotype d. Paratypes 26, l?. Englapd. Freeman skirwithensis Edwards, 1929:329, (spani0t0iii&) Paratrichocladius skirwithensis Syntypes 26. Engaand . Edwards steinb6cki Goetghebuer, 1973:56, (Diamesa) Diamesa steinbocki Paratypes 28 (slide). Austria. Goetghebuer strlatus Strenzke, 1959: 17, (Chlrollomus) Chironomus striatris Strenzke Paratypes 18, l? (spirit). West Germany. submontana Edwards, 1932:46, (~paniotoma) Acamptocladius submontanus Syntypes l&, 19. Scotland. Edwards subnudus Edwards , 1929 :3 16, (Metriocnemus ) Gymnometriocnernus subnudus Syntypes 10d, England, Scotland. Edwards subvernalis Edwards:?;929: 341, (~paniotoma) Bryophaenocladius subvernalis Holotype d. England. Edwards tarsalis Wallter, 1856 :170, (Chironomus ) Microtendipes tarsalis Walker Holotype d. England. thienemanni Edwards, 1938:153, (Boreochlus) Boreochlus thienemanni Paratypes 76, 79 (spirit 1. Sweden. Edwards Type & Allot~pe (in B.M.N.H., according to Edwards & Brundin, have not been found.] thummi piger Strenzke, 1959:29, (~hironomus) Chironomus thumrni f. piger Paratypes 16, 19 (spirit), West Germany. Strenzke tornetrZskensis Edwards, 1941:211, (Orthocladius) Trissocladius tornetraskensis Syntypes 56 (slide). Sweden. Eri~vards transgressus Holmgren, 1883:180, (~hironornus) Metriocnemus transgressus Syntype d. U.S.S.R., Novaya Zemlya Is. Holmgren trichopus Walker, 1856: 169, (Chironomus) Endochironomus tendens Holotype cf. England. Fsbricius trifascia Edwards, 1929:322, (~ricoto~us) Cricotopus trifascia Edwards Syntypes 6d, 29. England, trilobata Edx-ards, 1929: 364, (~paniotornn) Pseudosmittia trilobata Holotype d. England. Edwards tripilata Edwards, 1929:348, (~paniotoma) Synorthocl~dius setnivirens Syntypes 208, 149. England. ICief fer tristellus Edwards, 1929:312, (Hetriocnemus) Metriocnemus tristellus Syntypes 36, l? (16 slide). England. Edwards tritus Walker, 1856: 162, (~hironornus) Pentapedilum tritum Walker tlolotype 9. England. tuberculata Edwards , 1929:341, (~~aniotoma) Bryophnenocladius tuberculata Syntypes 28. England. vanderwulpi Edwards, 1929:418, (Tnn~tnrsus) Cladotanytarsus vanderwulpi Syntypes 5d, l?. Englnnd. Edwards variabills Staeger, 1879: 571, (Chironornus) Haloclndius vnrinbilis Paralectotypes 16, 1 . Denmark. Staeger verr~lliEdwards, 1929: 348, (Spaniotomn) Eukiefferiella verralli Holbtype d (slide). Wales. Edwards verralli Goetghebuer, 1928 : 143, (Tanytarsus) Tawtarsus verralli ? Holotype d. England. Goetghebuer virgo Edwnrdo, 1937a: 141, (.4biskomyia) .4biskomyin virgo Edwsrds Syntype Q (sli e). Sweden. vittata Edwards , l9q1b: 184, (Corynoneura) Thienemanniella vittata Holotype 2d. Wa es. Edwards woodi Edwards, 1929:292, (Pentnneura) Th'enemannirnyia woodl Syntypes 26 (1 no abdomen). Englnnd. Edwards xanthogyne Edwards, 1929:3&2, (Sprniotoma) Bryophnenocladius xanthogyne Syntypes lid, 6?. England. Edwards zernyi Edwards, 1933a:617, (Diamesn) Diantesc zernyi Edwards Holotype d. Austria. zonarius Walker, 1856:176, (Chlronomus) Microtendipes confinis Holotne 6. Ellgland. Meigen

Andersen, F.S., 1937, Meddr, Grdnland. 116 ($86-96. Cranston, P.S.,1976, in Kloet, G.S. & Hincks, W.D. Handbk Ident. Br. Insects G (~):12-21. Edwards, F.W., 1919, Ann. Mag. nat. Hist. (9)2:222-228. 1922, Ann. MRS. nat. Hist. (2) -10: 193-215. 1924a, .Inn. Flag. nat. Hist. (2)14: 162-174. 1924b, Entomologist's mon. Mng. e:203-207. 1924c, Annls. Biol. lncustre 1/7:119-122. 1926, Proc. zool. Soc. Lond. -:779-,?06. 1928, Encycl. ent. (B) IT, k:157-189. 1972, Scott. Nat. =:47-52. 1937a, Ann. Mag. nat, Hist. (10)~:611-620. 1933b, Scott. Nat. -195:87-92, 113-117. 1935, Ann. Mag. nat. Hist. (2)2:46?-4?5. 1937a,,Ann. Mag. nat. Hist. (2)=:140-1/18. 1937b, Int. Revue ges. Hydrobiol. Hydrgr. 2:99-107. 197g, Zo01. Anz. ~:152-154.1939, 2001. Anz. 127:259-264. 1941, in Thienemann, A., 1941, Arch. Iiydrobiol. Supp1.-Bd. x:1-253. Fittkau, E.J., 1954, Beitr. Ent. -4:84-98. Freeman, P., 1948, Entomologistls mon. Mag. %:49-50. 1957a, Bull. Br. Mus. nat. Hist. (Ent.) 3 (2):323-426. 1Y57b, Result.scient. EqLlor. h~drobiol. Lacs Kivu, Edouard, Albert 2 (2):203 -221, Goetghebuer, M. ~6m.Mus. Hist. nat. Belg. 8 (4):l-208. 1928, Faune Fr. 1831-171. 1932, Fame Fr. a:1-204. 1933, Bull. Annls. Soc. r. ent. Belg. z:111-120. 1934, Arb. morph. taxon. Ent. Berl. ~:36-39. IIirvenoja, M-, 1973, Ann. Zool. Fenn. 10:1-363. Holmgren, A.E., 1869, K. svenska Vetensk Akad. Handl. 8 (2): 1-55. 1887, Ent. Tidskr. 2: 162-190. Kieffer,, J. J., 1921, Aruh. Hydrobiol. Supp1.-Bd. 2:785-808. 1926, in Zavrel, J., 1926, Acta Soc-Scient. nat. moravo-siles. 2:251-282. Kirby, W., 1824, in Parry, W.E., A supplement to the appendix of Captain Parry's voyage for the discovery of a North-West Passage, in the years 1819-20. Pp. ccxiv-ccxix (~otseed). Saether, O.A., 1975, Can, Ent. =:1029-1056. Staeger,.C., 1839, Naturh. Tidsskr. 2:549-600. Strenzke, K., edi- tion, 47 pp. Cambridge. Walker, F., 1856, Insecta britannica. Diptera. 2 352 pp. London. Wollaston, T.V., 1858, Ann. Mag. nat. Hist. (3) ~:113-124.

DIE CHIRONOMIDENTYI'EN UND IHR ERHALTUNGSZUSTAND IN DER SAMPlLUNG DES MUS%UM NATIONAL D~HISTOIRENATURELLE, PARIS.

von Ernst Josef Fittkau und Friedrich Reiss

Die Sammlung Maigen enthalt genadeltes Material von 134 nominellen Chironomiden- arten,das unter den Nummern 22-156 inventarisiert ist. ES wird in seiner Gesamt- heit als Typusmaterial betrachtet und ist in jiingerer ieit nur zu einem geringen Teil revidierbnd bearbeitet worden. AnlaRlich der Revision europiiisch verbreiteter Tanytarsus-Arten mdedie Samm- lung von E.J. Fittkau durchgesehen. Die dabei entstandenen Notizepl zclr Anzahl, dem Geschlecht und Erhaltungazustand der Individuen der jeweiligen Arten warden in der folgenden Liste angefiihrt, die auRerdem Hinweise auf die ZugehSrigkeit der Arten zu den heute giiltigen Gattungen oder Unterfamilien enthalt.

Abd. = Abdomen, dnt. = Antenne, Fl. = Fliigel, Hyp. = Hypopyg,

Chironomus

22 obscuripes, 28 (beide Hyp. fehlen), 19 Glyptot endipes 23 aprilinus, 26 (1 Hyp. fehlt), 29 Chironomus 24 dllaveiibus, id: 1 nicht definierbarer Rest Chironomus 25 prasinus, 33 (2 Hyp. fehlen), l$(~bd. fehlt) Chironomus 26 plumosus, l# (Ant. fehlen) , 12 Chironomus 27 sticticus'(histrio) 18:(Fragment mit ~einenj Stictochironomns 28 tendens 16 (Ant, fehlen) Endochironomus 29 flerilis (gibbua) 28 (1 ~bd. und l Hyp. fehlen) Zavreliella 30 litorellurn 1 Fragment ohne Kopf und Abd. Chironomini 31 notatus l? Chironomus 32 laetus loa (~bd. fehlt ),l$ (~nt. fehlen) Poypedilum 33 maculipennis 13 (Abd. fehkt) Stictochironomus 34 chloris 2d* (l Hyp. und 1 Abd. priipariert) Microtendipes 35 lividus 48 (2 Hyp. f ehlen, 2dd' Prodiamasa olivacea) Orthocladiinae 36 olivaceus id' (Hyp. prapariert) Prodiamesa 37 psittacinus id (Hyp. priipariert) Cryptochironomus 38 lepidus 13 (Kopf fehlt ) Endochironomus 39 cingulatus 36 (1 ~bd. fhlt1, 19 Chironomus 40 riparius 28 (beide Hyp. prapariert), l? Chironomus 41 sticticus 18, 29 Stictochironomus 42 tentana l? Camptochironomus 43 nigrinue (pilicornis) I# (H-~.fehlt), 17 Chironomus 44 ecalaenus 14 1% Polypedilum 45 melaleucus 19 (~bd. fehlt) Orthocladiinae 46 scriptus 1% (Abd. fehlt) Orthocladiinae 47 pedellus 23 2~ Microtendipes 48 fuscipennis Orthocladiinae 2$$ 49 dispar 16 (Hyp. fehlt) , 12 (~bd. fehlt) Endochironomus 50 rufipes 2c? ( l Hp. f ehlt 1, 2% Glyptotendipes 51 dorsalis 28 (1 Hyp. prapariert, 1 Kopf und 1 Abd. fehlen) , 1y (Kopf und Abd. fehlen) Einfeldia 52 obscurimanus l$ (Abd. fehlt), l? (Fragment) Chironomus 53 aterrimus 201(1 Exemplar prxpariert, l Exemplar ohne Abd. ) Othocladiinae 54 vagans 18 Paratendipes 55 supplicans IC? (Ant. fehlen) Chironomini 56 albipennis 2d ( 1 Abd. f ehlt 1, l? Chironomus 57 nitidus 1 Fragment ohne Kopf und Abd. Microtendipes 58 pallidulus 2d (1 Abd. fehlt 1, l? Micropsectra 59 flavipes 24 (1 Hyp. und 1 Abd. fehlen) , 22 Pentaneurini 60 fuscus I? Micropsectra 61 opacus 20' ( l Exemplar ohne Abd. und Beine) Camptocladius 62 albolineatus 2rf (1 Hyp. und 1 Abd. fehlen), 28 (1 Abd. fehlt, 1 F1. fehlt) Orthocladiinae 63 scutellatus 2$ (1 Hyp. fehlt), 7$ (1 Abd. fehlt) Prodiamesa 64 plebeius 1s (Hyp. fehlt) Paratendipes 65 proximus 13 (Ant. fehlen) Orthocladiinae 66 albimanus 12(~bd. fehlt ) , 41pl ( 1 Exemplar ohne Abd. , 2 ohne F1. und Beine) Paratendipes 67 glabricollis 4$ ( 1 Antennenpaar fehlt ) , 2q Orthocladiinae 68 obscurus 23 (1 Abd. fehlt) , 19 (Kopf fehlt) Microtendipes 69 nubeculosus l# (~~p.przpariert ) , 2s Polypedilum . 70 bicinrtus ion (~bd. fehlt) , ly (~nt. fehlt) Cricotopus Cricotopus 72 pulchellus l$ Cricotopus

73 dizenias 32 (l? Cricotopus tremulus, 10t Cricotopus flavocinctus det. M. Hirvenoja) Cricotopus 74 perennis 3dl (2 Hyp. und 1 Kopf fehlen) Orthocladiinae 75 oscillator 16 (Hyp. fehlt), 12 (Cricotopus silvestris, det. M. Hirvenoja) Cricot opus 76 tremulus 12(Abd. fehlt), l? (~bd. fehlt) Cricot opus 77 carbonarius 1 Fragment (Thorax und l F1.) Orthocladiinae 78 bilineatus lc? (Abd. und Ant. fehlen) Diamesini 79 virescens l? (Fragment mit Kopf,Thorax, l F1. und 1Bein) Orthocladiinae 80 silvestris 33 (1 Abd. und 1 Hyp. fehlen), 2% Cricotopus 81 tricinctus 1 Fragment Cricotopus 82 trifasciatus 17 Cricot opus 83 amasia 36 (1 Exemplar ohne Abd., l Exemplar ohne Kopf und Beine) Cricotopus ornatus \ 84 flabellatus 10' (ICopf und Abd. fehlen) , ly(ohne ~bd.) 2 Arten (d~hironominio ?) t 85 dirnidiatus l7 Tanytarsini 86 bicolor 107 Microcricotopus 87 chiopterus lon(~bd. und fast alle Beine fehlen) Orthocladiinae 88 picipes 23( 1 Abd. fehlt 1, 49 Orthocladiinae 89 occultans 19 (Kopf und 4 Beine fehlen) Orthocladiinae 90 pusillus 3d'(alle Abd. fehlen) Orthocladiinae 91 confinis lon (Hyp. fehlt ) Microt endipea 92 pusio 12(Hyp. fehlt) Orthocladiinae 93 stercorarius 52(3 Abd. fehlen, 1o7pr6pariert), 10 Camptocladius 94 byssinus 52 (2 Hyp. fehlen) Orthocladiinae 95 rubicundus 28(1 Abd. fehlt), 19 (~bd. fehlt) Orthocladiinae 96 vernus 267 (1 Exemplar ohne Hyp.. 1 Exemplar ohne ~nt.),'l~ (Abd. fehlt) Orthocladiinae 97 junci 20' (1 Abd. fehlt, 1 Hype prapariert) l? Micropsectra 98 nanus 2oa ( l Hyp. fehlt 1, 2 Fragmente, l? (andere. Art) Parapsectra 99 flexilis (gibbus) nicht vorhanden 100 miniatus 12(Abd. fehlt ) Orthocladiinae l01 flavimanus l# (Abd. fehlt'), gro8e dunkle Art Chironominae 102 gibbosus 10 (nur Thorax erhalten) Cricotopus i- 103 leucopogon l$ (Fragment mit F1. und Beinen) Orthocladiinae 104 albicornis l Fragment mit Thorax, F1. und Beinen Tanytarsini 105 niveipennis 1 Fragment mit Thorax, F1; und div. Beinen Tanytarsini 106 punctipes l Gnbrauchbares Fragment Phaenopsectra (nach Etikett) 107 fuscipes 12(~bd. fehlt) , 2~ (l ~bd. fehlt) O~thocladiinae 108 debilis 26(1 Kopf und 1 Abd. fehlen, &p. prapariert ) Tanytarsus 109 campestris 26 (beide Abd. fehlen, 1 Antennenpaar fehlt) Micropsectra 110 praecox 201(1 zerfressenes Abd., 1 fehlender Kopf) , 1 Fragment (Thorax U. 1 Fl, ) ( l Hyp. / Prodiamesa 111 intersectus nicht vorhanden 112 minimus 48(3 Abd. fehlen, Hyp. priipariert) 2 (~r- Limnophyes ten) 113 rusticus I# Ch ironomus 114 tibialis 1s(Hyp. prapariert) Cricot opus 115 indulgens Ceratopogonidae 116 ictericus 10% (Kopf und Abd. rahlen) Orthocladiinae 117 sulphuricollis l? (Fragment mit Thorax,F1. und Beinen) Chironomini 118 albipes ldt (Fragment, Thorax mit l ~1.1, 17 (~bd. f ehlt ) Tanytarsini 119 albidus - l8(Fragment, Thorax mit FI. 1, lF($opf 7 .- und Abd. fehlen) Tanytarsini 120 tenuis 2on (2 Abd. und l Kopf fehlen) , l? Paratanytarsus 121 nubilipennis l# (Abd. fehlt) Polypedilum ? 122 denigratus nicht vorhanden lZJ modestus 1 Fragment ohne Kopf und Abd. Brillia 124 longimanus nicht vorhanden 125 festivus l? ? Cricotopus St ictochironomus 126 nubilus 19 (Abd. schlecht prapariert) , l? 127 annulipes nicht vorhanden 128 leucopus 3d (l Kopf, 2 Ant., 1 F1. und 2 Abd. fehlen, 1 Hyp. prapdriert) Polypedilum l ,~raparat), 129 vitripennis jc? ( 1 Abd. f ehlt , 2? (1 Abd. fehlt) Cricotopus 130 elegans 20' l Abd, fehlt), l? Orthocladiinae 131 pallens l$ (Fragment, Thorax mit l F1. und 1 Bein) Glyptotendipes 132 thoracicus l? Orthocladiinae -- rufiventris 18 (prapariert) Paratrichocladius Diamesa 133 waltli 2$ Diamesa Tany~us 134 varius 12 (~bd. fehlt 1, 1 Fragment Psectrotanypus 175 zonatus l? (~bd. und 1 F1. fehlen) Tanypodinae 136 melanurus 28(l Abd. und 1 F1. fehlen) Zavrelimyia ? 137 nebulosus (culiciformis) 18, 19 (~bd. und l F1. fehlen) Macropelopia 138 notatus 28 (l Hyp. fehlt) , l? (Abd. fehlt) Macropelopia 139 carneus Praparat r Pentaneurini 140 flabellicornis 13 (Hyp. und Ant. fehlen) Zavrelimyia ? 14 1 pallidulus 1s(4 Beine fehlen) , l? (Fl. and Abd. fehlen) Conchapelopia 142 ferruginicollis ld1 (3 Beine fehlen) , l? (Beine und Abd. fehlen) Zavrelimyia 143 punctipennis id' (psapariert) Tanypus 144 fasciculatus (nervosus ?) 22( 1 Abd. f ehlt 1, 2~ Clinot anypus 145 laetus 19 (Fragment mit Thorax und Fliigelrest! Tanypodinae 146 choreus l? ( 1 F1. fehlt) , 1 Fragment Procladius 147 nervosus nicht vorhanden 148 culiiiformis (lineola) l$ (Fragment 1, l?, .l? (Fragment) Procladius 149 monilie 14 lg Ablabesmyia 150 punctatus lc?, IF (Fragment , 1% Natarsia - 15 1 binotatus 2J (1 Prsparat) Krenopelopia 152 praecox 105 Diamesa 153 incarnatus id (~yp. fehlt) Rheopelopia 154 melanops lc?(pr~pariert), 3y (1 Abd. fehlt) Conchapelopia 155 nubilus 3p (lotThienemannhyia) Zavrelimyia 156 vitripennis lc?(~bd. und Ant. fehlen) ? (keine ~nny~odine)-

Literaturhinweis

Unter den in den letzten Jahren verijffentlichten Monographien ve~dientbe- sondere Beachtung die 1973 erschienene Arbeit van M. HIRVENOJA : Revision del- Gattung Cricotopus van der Wulp und ihrer Verwandten (Diptera, Chironomidae). - Ann. 2001. Fenn. g:1-363. Wir geben hier das Autorreferat aus der Originalarbeit wieder, auch wenn die Arbeit weit mehr Informationen bringt, als hier angedeutet sind.

A taxonomic revision of the species of the genus Cricotopus van der Wulp : and its closest relatives from the western part of the Palaearctic region is presented, and the data on the distribution and habitats of the different species are didcussed. To evaluate the numerous characters studied, the adult, pupal and larval morphology of the Chironomidae has been reviewed. The aim has been to find the most suitable names and assess the homologies of many important structures in the different subgroups of the Chironomidae and in some other Nematocera. It is suggested that a strictly monophyletic group consists of the genus Cricotopus (with subgen. Cricotopus S. str. and Isocladius Kieffer status n=) and its nearest relatives, the genera Acricotopus Kieffer, Paratrichocladius San- tos Abreu (syn. Syncricotopus Brundin, syn. n.), Paracladius Hirvenoja (nom. nov. for Paratrichocladius Thienemann) and Halocladius S. str. subgen. n.) About 50 species from the western part of the Palaearctic region have been redescribed, mostly from new material. As far as possible, the descriptions are based on all developmental stages (larva, pupal exuviae and adult). In addition, the following 21 new species are described: Paracladius quadrinodosus, Halocladius (H.) mediter- raneus, Cricotopus (C.) pilidorsum, C. (C.) pirifer, C. (C.) magus, C. (C.) sep- tentrionalis, C. (C.) claripes, C. (C.) tristis, C. (C.) beckeri, C. (C.) patens, C. (C.) coronatus, C. (C.) guttatus, C. (C.) caducus, C. (I.) laetus, C. (I.) pi- licauda, C. (I.) relucens, C. (I.) suspiciosus, C. (I.) reversus, C. (I.) arcua- tus, C. (I.) obtusus and C. (I.) reductus. In addit4on to these, C. (C.) tokuna- gai Hirvenoja is the new name for C. bituberculatus Tokunaga, which is a secon- dary homonym of Trichocladius bituberculatus Goetghebuer, and C. meunieri Hirve- noja is the new name for C. parvulus Meunier, which is a primary homonym of C. parvulus Kieffer. In the light of present knowledge, an attempt has been made to place the species from all different parts of the world that have been ascribed to the genus Cricotopus up to 1970 in the genera to which they probably belong. In addition, it is suggested that a number of names existing in the literature should be regarded as nomina dubia. Keys to the males, females, pupal exuviae and larvae are presented, as far as possible down to species level. Several hundreds of new figures of taxonomi- cally important characteristics are1presented.

Request for information

I have started work on the genus Paracladopelma Harnisch. Th29 work will initially concentrate on the immature stages and may later lead to a revision. The study will include nearctic and possibly palaearctic species. I would like to know the location of associated material and type specimens particularly for the palaearctic region. Information regarding the existence of such material would be greatly appreciated. George Jackson 3479 Lake Lansing Rd. East.Lansing, Michigan 40823 u.s.A.\ Hace algo mhs de 120 aiios que se comenzaron en Chile las investigaciones sobre chironomidos. En efecto, E. Blanchard en 1852 describib las cinco primeras especies y en 1865 R.A. Philippi aportb a las ya conocidas otra docena de especies de la fauna de chironomidos. .Desde entonces y hasta 10s a+s treinta de 1900 se su- man a la lista de especies otra decena de descripciones product0 del trabajo de diversos cientificos tales como J. Bigot, J. Kieffer, J. BrLthes y G. Enderlebn, entre otros. Sin embargo, la mayoria de las especies descritas en aquellos tiem- pos, no pueden ser hoy dia identificadas 0 sblo con muchas dificultades, debido' a Pa insuficiente descri~cibne ilustracibn o por la fa1ta.de especimen tipo. .El primer aporte taxonbmico de consideraci6n a1 conocimiento de la fauna de chironomidos chilenos, fue realizado por el cientifico F.W. Edwards en 1931, el que en base a modernos criterios de clasificadibn y a1 anblisis del genero y parcial de las especies, citb alrededor de 80 especies para el sur de Chile y la Patago- nia. Quince aiios mhs tarde, en relacibn a1 trabajo de Edwards, aparece el "Cat?- logo de 10s Dipteros de Chile" de C. Stuardo, en 1946, con un examen de todas las especies descritas hasta entonces. En 1952 y 1957, W. ~irthpubli'ca dos trabajos sobre 10s chironomidos del Archi- pielago de Juan Ferngndez. Los aportes mhs recientes a la fauna de chironomidos de Chile, se deben a L. Brundin en 1966 y a F. Reiss en.1972, ocupandose el primer0 de 10s Podonominae, Aphroteniinae y Heptagyiae y el segundo de 10s Tanytarsini. En conjunto ambos investigadores citan mhs de un centenar de especies nuevas para Chile y la Cien- cia. El trabajo'de Brundin marca un importante hito en las investigaciones sobre chironomidos de Chile y tambibn en la aplicacibn prgctica de 10s modernos M6to- doe ~ilo~enbticospara determinar las relaciones de parentesco entre organismos circumanthrcticos. - El conjunto de las aproximadamente 200 especies citadas de chironomidos que ha- bitan las aguas continentales de Chile, puede ser porcentualmente desglosado en las Subfamilias: Podonominae 50 %; Tanypodinae 8 %, Orthocladiinae 29 %, Chirono- minae 11 % y Aphroteniinae 2 %. Sin embargo, estos valores porcentuales sblo pueden ser condiderados como indicadores relativos y variables de la composicibn de la fauna de chironomidos de Chile, ya que a pesar de ser Qsta, en opinibn de F. Reiss, hasta hoy dsa la mejor conocida de Fudambrica, dista mucho de estar com- pleto su estudio. Especialmente en las Subfamilias Tanypodinae, Orthocladiinae y Chironominae, de las cuales es posible esperar un nGmero de especies nuevas no ' inferior a1 hasta ahora conocido,. La intensa colecta de material llevada a cab0 por L. Brundin hacen altamente pro- bable que en estos momentos se conozcan todas las especies de 10s grupos traba,ja- dos por 61. Una comparacibn a groso modo entre la composicibn ~or'centual de la fauna de chironomidos chilena con la europea - (E.J. Fittkau et al., 1967, in: Illies, J. ( ed.) Limnofauna Europea, G. Fischer Verlag, Stuttgart) - en la que 10s Podonomi- nae no alcanzan a1 1 % sobre un total de 1523 especies, contrasta con el porcen- taje (50 %) que presenta esta Subfamilia en Chile. Esto vendria a demostrar una vez m&s el marcado endemism0 que junto a algunas singularidades morfolbgicas, que entre 10s chironomidos se encuentran en el Genero Rhinocladius, con su larga pro- boscis, caracterizan en general a la fauna de Chile. En razbn de esto parece acon- sejable la revisibn a nivel de Genero de por 10 menos el 40 % de las especies hasta ahora descritas, especialmente en Los grupos Tanypodinae, Orthocladiinae y Chironomini. Sin embargo, para ell0 es previamente necesario completar el estudi; Y descripci6n de la metamorfosis de la gran mayoria de las especies citadas hasta hoy. Sblo con el anslisis precis0 de las caracteristicas de 10s tres estados de las especies, se haria m& factible dicha tarea. Es preocupacibn del autor realizar en el futuro aportes en este sentido y para 10 cual realiza actualmente un periodo de perfeccionamiento junto a 10s Drs. E.J. Fittkau y F. Reiss, en el Max-Planck-Institut fiir Limnologie en Pion, Alemania, con especial interes en la Subfamilia Tanypodinae, con ell0 espera hacer posible estudios ecolbgicos relacionados con chironomidos en las aguas continentales, ge- nerosamente abundantes, en el sur de Chile. La casi totalidad de la literatura relativa a 10s chironomidos chilenos esta ci- tada en la obra de L. Brundin, 1966. "Transantarctic relationships and their sig- nificance, as evidenced by Chironomid midges. With a-monograph of the Subfamilies Podonominae and Aphroteniinae and the Austral Heptagyae." K. svenska Vetensk. Akad. Handl. l1 (1):l - 472. Stockholm. Una bibliografia completa sobre Chironomidae del sur de ~udamericaha sido prepa- rada por F. Reiss y se espera su aparicibn en "~ibliografiasTaxonbmicas de la Flora y Fauna de Aguas Continentales del Sur de ~udam&rica" a cargo de S. H. Hurl- bert, San Diego, California. J. N. Arenas, Valdivia, Chile Instituto de Zoologia, Universidad Austral Zum erfolgreichen Abschlul3 ihres Dr.-Examens begliickwiinschen wir folgende Koliegen: Claude Dejoux, Universitb Paris Sud: "Syn&cologie des Chironomides du Lac Tschad (~i~t6res-~&matocdres)".1974 Dean Cyrus Hansen, University of Minnesota, St. Paul: "Systematics and morphology of the nearctic species of Diamesa Meigen, 1835 (Diptera:Chironomidae)." 1977

Friedhelm Ringe, Christian-Albrechts-Universit5t Kiel: "Chironomiden-Emergenz 1970 in Breitenbach und Rohrwiesenbach." 1977 Amele R. Soponis, Carleton University Ottawa: "A Revision of the Nearctic Species of Orthocladius (Orthocladius) van der liulp (Kieffer) ." 1974 Dirk-Heinrich Stechmann, Christian-Albrechts-Universitiit Kiel: "Biologisch-Gkologische Untersuchungen zum Entwicklungszyklus einheimischer Arrenurus-Arten (Hydrachnellae, Acari)." 1975 Jean-Noel Tourenq, Universit6 Paul Sabatier de Toulouse: "Recherches ecologiques sur les Chironomides (~i~tcra)de Camargue." 1975

hdressen: ~nderunlfenund ErgZnzungen

ELFENBEINKUSTE SPANIEN

DEJOUX, Dr. C., STOCKERT, Dr. J.C,. O.R.S.T.O.M. Laboratoire Departamento de Citologia d1 Hydrobiologie Instituto de Biologia Celular B.P. 1474, Bouakh, C8te d' Ivoire, Afrika Veljzquez 144 Madrid - 6 ENGLAND -USA CARTER, Dr. C. BAY, B.C., Limnology Laboratory, Superintendent, Traad Point,'Drumenagh, Western Washington, Magherafelt, Co. Derry. Research & Extension Center, Pioneer Way, INDIEN Puyallup, WA 98371 KULSIIRESTHA, A.K. , School of Entomology, ENGLISH, Prof. Dr. D.S., St. John's College, Northern Arizor~aUniv. , Dept. of Biological Sciences, Agra - 282002 (U.P.) Box 5640, Flagstaff, Arizona 86001 NIEDERLANDE WINGO, P.S., Biologist, PARMA, Dr. S., Duke Power Company Delta Institute for Environmental Laboratories Hydrobiological Research, Rt. 7, Box 90 Vierstraat 28, Huntersville, N. C- 28978 Yerseke

CHIRONOMUS-Mitteilungen aus der Chironomidenkunde. - Erscheint mehrmals jzhrlich. Herausgegeben von E. J. Fittlrnu und F. Reiss , Ilnx-l'lnnclc- Institut fur Limnologie, Abt. Tropenokologie, 272 Plon, Postfach 165, Deutschland (BRD).