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Crustaceans of the Eastern Tropical Pacific Macro BENTHIC MACRO- MACRO- CRUSTÁCEOS CRUSTACEANS BENTÓNICOS OF THE EASTERN DEL PACIFICO TROPICAL PACIFIC ESTE TROPICAL San Diego r Instituto de Ciencias del Mar y Limnología Natural History Museum Universidad Nacional Autónoma de México Proceedings of the San Diego Society of Natural History 9-27 This volume is the result of a joint effort between the San Diego Natural History Museum, San Diego, California, United States and the Instituto de Ciencias del Mar y Limnología, Universidad Nacional Autónoma de México Este volumen es el resultado de la colaboración entre el Instituto de Ciencias del Mar y Limnología, Universidad Nacional Autónoma de México México, y el San Diego Natural History Museum, San Diego California, Estados Unidos de América Scientific editors Editores científicos Spanish contributions English contributions Contribuciones en Español Contribuciones en Inglés Michel E. Hendrickx Richard C. Brusca Technical editor Editor técnico Philip Unitt San Diego Natural History Museum ISBN No. 0-918969 -03 -4 BENTHIC MACRO- MACRO- CRUSTÁCEOS CRUSTACEANS BENTÓNICOS OF THE EASTERN DEL PACIFICO TROPICAL PACIFIC ESTE TROPICAL San Diego r Instituto de Ciencias del Mar y Limnología Natural History Museum Universidad Nacional Autónoma de México Proceedings of the San Diego Society of Natural History 9-27 i Introduction Introducción The crustacean fauna of the tropical eastern Pacific constitutes a La fauna crustácea del Pacifico Este Tropical constituye un rich and varied biological resource. The past ten years have wit- recurso biológico rico y variado. Los últimos diez años han visto un nessed a virtual flowering of research on this fauna, and hundreds florecimiento de investigaciones sobre esta fauna y cientos de of papers have appeared treating various aspects of systematics, artículos han aparecido tratando varios aspectos sistemáticos, biology, ecology, and fisheries research for the region. Much of this biológicos, ecológicos, y investigaciones pesqueras de esta región. work has focused on the Gulf of California, the northern outpost of La mayoría de este trabajo se ha enfocado en el Golfo de California the tropical Pacific New World marine fauna. Many researchers en el punto mas al norte de la fauna Pacifico-tropical del nuevo studying this fauna, from Canada to Chile, have had little contact mundo. Investigadores desde Canada hasta Chile que estudian esta with or were unaware of similar projects in other labs and in other fauna han tiendo poco contacto o conocimiento de proyectos countries. The time seemed ripe to bring these scattered researchers similares en otros laboratorios y en otros paises. El tiempo era together, especially so that Latin American workers could interact propicio para reunir a todos estos científicos, especialmente a los de closely for a few days with their U.S. colleagues. America Latina de manera de que junto con sus colegas From March 28-30, 1990, the Instituto de Ciencias del Mar y norteamericanos interactuaran conjuntamente durante varios días. Limnología of the Universidad Nacional Autónoma de México Del 28 al 30 marzo de 1990, el Instituto de Ciencias del Mar y (UNAM) hosted an international meeting at its Mazatlán Research Limnología de la Universidad Nacional Autónoma de México Station (Estación Mazatlán) titled "Colloquium on Benthic (UNAM) fue sede de una reunión internacional en sus instalaciones Macrocrustaceans of the Eastern Tropical Pacific." The meeting de la Estación Mazatlán: el 1 Coloquio sobre Macro-crustáceos was co-sponsored by UNAM, the San Diego Natural History Bentónicos del Pacifico Este Tropical. La reunión fue copatrocinada Museum, and The Crustacean Society, and it brought together por la Universidad Nacional Autónoma de México, el San Diego crustacean researchers from North and Latin America to share ideas Natural History Museum, y The Crustacean Society. El objectivo and exchange information. As plans developed we realized partici- principal era juntar científicos de America del Norte y de America pation would be substantial and that it would be worthwhile to Latina, tanto los más expertos como los principiantes, para que publish the papers presented at the meeting. pudieran reunirse, compartir ideas y intercambiar informaciones In addition to the support of UNAM's Instituto de Ciencias del acerca de sus trabajos. A medida que esta idea se concretizaba, nos Mar y Limnología, the San Diego Natural History Museum, and dimos cuenta que existia un interés genuino para este tipo de The Crustacean Society, we were fortunate to obtain additional reunión, que la participación al mismo se anticipaba muy buena y financial sponsorship from Consejo Nacional de Ciencias y que existia, además, una muy buena posibilidad de publicar los Tecnología (CONACyT) for meeting costs, and from the Regional trabajos presentados durante la reunión. Office for Science and Technology for Latin America and the Afortunadamente, pudimos contar con el apoyo indispensable Caribbean (ROSTLAC) of UNESCO for airfare for some South del CONACyT (Consejo Nacional para Ciencias y Tecnología, American participants. In all, 80 scientists and students from six México) para cubrir parte de los gastos de organización del evento, countries (Chile, Colombia, Costa Rica, Peru, Mexico, and the y de la Oficina Regional de Ciencia y Tecnología para America U.S.) attended the meeting, contributing 42 presentations to the Latina y el Caribe (ORCYT), UNESCO, para contribuir en el gasto program. The meeting enjoyed great success, and a follow-up mail de viaje de algunos participantes de America Latina. En total, 80 assessment indicated that more meetings of a similar nature would científicos y estudiantes de 6 paises (Chile, Colombia, Costa Rica, be welcomed by the research community. Nineteen of the papers Perú, México, y los Estados Unidos de America) participaron de that were submitted for publication are included in the present manera activa en la reunión y disfrutaron de un programa con 42 volume. We wish to thank the following persons for serving as contribuciones. El encuentro fue todo un éxito, y una encuesta reviewers for these papers: Larry Abele (Florida State University, posterior sobre el desarollo del mismo indicó el deseo por parte de U.S.A.), F. Alberto Abreu Grobois (UNAM, Mazatlán, Mexico), los participantes de reunirse nuevamente en un evento de la misma Daniel Adkison (Macon, Georgia, U.S.A.), Ricardo Alvarez Leon naturaleza. Diecinueve de los artículos sometidos a este coloquio (1NDERENA, Cartagena, Colombia), E. E. Boschi, (Insituto han sido incluidos en el presente volumen. Queremos agradecer a Nacional de Investigación y Desarrollo Pesquero, Mar del Plata, las siguientes personas que revisaron estos trabajos: Larry Abele Argentina), Thomas E. Bowman (Smithsonian Institution, Wash- (Florida State University, U.S.A.), F. Alberto Abreu Grobois ington, D.C., U.S.A.), Ramond Bauer (University of Southwestern (UNAM, Mazatlán, México), Daniel Adkisoy (Macón, Georgia, Louisiana, U.S.A.), Jose Antonio Calderón Pérez (UNAM, U.S.A.), Ricardo Alvarez Leon (INDERENA, Cartegena, Colom- Mazatlán, Mexico), Ernesto Campos (Universidad Autonoma de bia), E. E. Boschi, (Insituto Nacional de Investigación y Desarrollo Baja California, Ensenada, Mexico), Alberto Carvacho (Instituto Pesquero, Mar del Plata, Argentina)» Thomas E. Bowman Profesional de Osorno, Osomo, Chile), Elva Escobar (UNAM, (Smithsonian Institution, Washington, D.C., U.S.A.), Ramond Mexico, D.F.), Adolfo Gracia Gasea (UNAM, Mexico, D.F.), John Bauer (University of Southwestern Louisiana, U.S.A.), José Anto- R. Holsinger (Old Dominion University, Virginia, U.S.A.), David nio Calderón Peréz (UNAM, Mazatlán, Mexico), Ernesto Campos Jablonski (University of Chicago, Illinois, U.S.A.), Hans Kuck (Universidad Autónoma de Baja California, Ensenada, Mexico), (Los Angeles County Natural History Museum, California, U.S.A.), Alberto Carvacho (Instituto Profesional de Osomo, Osomo, Chile), Enrique Lozano-Alvarez (UNAM, Cancún, Mexico), Joel W. Mar- Elva Escobar (UNAM, México, D.F.), Adolfo Gracia Gasea tin (Los Angeles County Natural History Museum, California, (UNAM, México, D.F.), John R. Holsinger (Old Dominion Univer- U.S.A.), William A. Newman (Scripps Institution of Oceanogra- sity, Virginia, U.S.A.), David Jablonski (University of Chicago, phy, California, U.S.A.), Francisco Sarda (Instituto de Ciencias del Illinois, U.S.A.), Hans Kuck (Los Angeles County Natural History Mar, Barcelona, Spain), Marie Simovich (University of San Diego, Museum, California, U.S.A.), Enrique Lozano-Alvarez (UNAM, California, U.S.A.), Henry R. Spivey (Florida State University, Cancún, México), Joel W. Martin (Los Angeles County Natural U.S.A.), James Weinberg (Woods Hole Oceanographic Institution, History Museum, California, U.S.A.), William A. Newman (Scripps ii Massachusetts, U.S.A.), Regina Wetzer (San Diego Natural History Institution of Oceanography, California, U.S.A.), Francisco Sarda Museum, California, U.S.A.), Mary K. Wicksten (Texas A&M (Instituto de Ciencias del Mar, Barcelona, España), Marie Simovich University, U.S.A.), Deborah L. Zmarzly (San Diego Natural His- (University of San Diego, California, U.S.A.), Henry R. Spivey tory Museum, California, U.S.A.), Victor A. Zullo (University of (Florida State University, U.S.A.), James Weinberg (Woods Hole North Carolina. U.S.A.). Special thanks go to Philip Unitt (Scien- Oceanographic Institution, Massachusetts, U.S.A.), Regina Wetzer tific Editor, San Diego Natural History Museum), who served as (San Diego Natural
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