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Download Full Article in PDF Format comptes rendus palevol 2021 20 7 iles — Jean- pt Cl re au d d e n a R s a n g a e i — b i h P p a l a m e a o f b o i o y l h o p g a y r g a o n e d g p o i a l b a o e DIRECTEURS DE LA PUBLICATION / PUBLICATION DIRECTORS : Bruno David, Président du Muséum national d’Histoire naturelle Étienne Ghys, Secrétaire perpétuel de l’Académie des sciences RÉDACTEURS EN CHEF / EDITORS-IN-CHIEF : Michel Laurin (CNRS), Philippe Taquet (Académie des sciences) ASSISTANTE DE RÉDACTION / ASSISTANT EDITOR : Adeline Lopes (Académie des sciences ; [email protected]) MISE EN PAGE / PAGE LAYOUT : Fariza Sissi (Muséum national d’Histoire naturelle; [email protected]) ÉDITEURS ASSOCIÉS / ASSOCIATE EDITORS (*, took charge of the editorial process of the article/a pris en charge le suivi éditorial de l’article): Amniotes du Mésozoïque/Mesozoic amniotes Hans-Dieter Sues* (Smithsonian National Museum of Natural History, Washington) Lépidosauromorphes/Lepidosauromorphs Hussam Zaher (Universidade de São Paulo) Métazoaires/Metazoa Annalisa Ferretti (Università di Modena e Reggio Emilia, Modena) Micropaléontologie/Micropalaeontology Maria Rose Petrizzo (Université de Milan) Palaeoanthropology R. Macchiarelli (Université de Poitiers, Poitiers) Paléobotanique/Palaeobotany Evelyn Kustatscher (The Museum of Nature South Tyrol, Bozen/Bolzano) Paléoichthyologie/Palaeoichthyology Philippe Janvier (Muséum national d’Histoire naturelle, Académie des sciences, Paris) Palaeomammalogy (small mammals) L. van den Hoek Ostende (Naturalis Biodiversity Center, CR Leiden) Palaeomammalogy (large and mid-sized mammals) L. Rook (Università degli Studi di Firenze, Firenze) Prehistorical archaeology M. Otte (Université de Liège, Liège) Tortues/Turtles Juliana Sterli (CONICET, Museo Paleontológico Egidio Feruglio, Trelew, Argentine) COUVERTURE / COVER : Tooth specimens of the anacoracid shark Squalicorax pristodontus (Agassiz, 1843) Comptes Rendus Palevol est indexé dans / Comptes Rendus Palevol is indexed by: - Cambridge Scientific Abstracts - Current Contents® Physical - Chemical, and Earth Sciences® - ISI Alerting Services® - Geoabstracts, Geobase, Georef, Inspec, Pascal - Science Citation Index®, Science Citation Index Expanded® - Scopus®. Les articles ainsi que les nouveautés nomenclaturales publiés dans Comptes Rendus Palevol sont référencés par / Articles and nomenclatural novelties published in Comptes Rendus Palevol are registered on: - ZooBank® (http://zoobank.org) Comptes Rendus Palevol est une revue en flux continu publiée par les Publications scientifiques du Muséum, Paris et l’Académie des sciences, Paris Comptes Rendus Palevol is a fast track journal published by the Museum Science Press, Paris and the Académie des sciences, Paris Les Publications scientifiques du Muséum publient aussi / The Museum Science Press also publish: Adansonia, Geodiversitas, Zoosystema, Anthropozoologica, European Journal of Taxonomy, Naturae, Cryptogamie sous-sections Algologie, Bryologie, Mycologie. L’Académie des sciences publie aussi / The Académie des sciences also publishes: Comptes Rendus Mathématique, Comptes Rendus Physique, Comptes Rendus Mécanique, Comptes Rendus Chimie, Comptes Rendus Géoscience, Comptes Rendus Biologies. Diffusion - Publications scientifiques Muséum national d’Histoire naturelle CP 41 - 57 rue Cuvier F-75231 Paris cedex 05 (France) Tél. : 33 (0)1 40 79 48 05 / Fax : 33 (0)1 40 79 38 40 [email protected] / https://sciencepress.mnhn.fr Académie des sciences, Institut de France, 23 quai de Conti, 75006 Paris. © Publications scientifiques du Muséum national d’Histoire naturelle / © Académie des sciences, Paris, 2021 ISSN (imprimé / print) : 1631-0683/ ISSN (électronique / electronic) : 1777-571X Stratigraphy, age, and vertebrate palaeontology of the latest Cretaceous Quintanilla la Ojada locality (Basque-Cantabrian Region, northern Spain): a synthesis José-Carmelo CORRAL Museo de Ciencias Naturales de Álava/Arabako Natur Zientzien Museoa, Dpto. Geología, Siervas de Jesús 24, 01001 Vitoria-Gasteiz (Spain) [email protected] (corresponding author) Ana BERRETEAGA Universidad del País Vasco/Euskal Herriko Unibertsitatea, Facultad de Ciencia y Tecnología, Dpto. Estratigrafía y Paleontología, Apdo. 644, 48080 Bilbao (Spain) [email protected] Francisco José POYATO-ARIZA Universidad Autónoma de Madrid, Dpto. de Biología, Unidad de Paleontología, 28049 Madrid (Spain) [email protected] Nathalie BARDET CR2P, Centre de Recherche en Paléontologie - Paris, UMR 7207, CNRS-MNHN-SU, Muséum national d’Histoire naturelle, CP 38, 57 rue Cuvier, 75005 Paris (France) [email protected] Henri CAPPETTA Institut des Sciences de l’Evolution de Montpellier (ISEM, UMR 5554, NRS/UM/IRD/EPHE), Université de Montpellier, Place E. Bataillon 34095, Montpellier Cedex 5, (France) [email protected] Marc FLOQUET Centre Européen de Recherche et d’Enseignement de Géosciences de l’Environnement, CEREGE - UM 34 Aix Marseille Université, CNRS, IRD, Collège de France, INRA, OSU Institut Pythéas, Technopole de l’Arbois, BP 80, 13545 Aix-en-Provence (France) [email protected] Humberto ASTIBIA Ainara BADIOLA Xabier PEREDA-SUBERBIOLA Universidad del País Vasco/Euskal Herriko Unibertsitatea, Facultad de Ciencia y Tecnología, Dpto. de Geología, Apdo. 644, 48080 Bilbao (Spain) [email protected] [email protected] [email protected] Submitted on 15 October 2019 | Accepted on 21 February 2020 | Published on 22 February 2021 urn:lsid:zoobank.org:pub:CEBC9E4D-3ED4-4330-BE40-67B037B63A37 COMPTES RENDUS PALEVOL • 2021 • 20 (7) © Publications scientifiques du Muséum et/and Académie des sciences, Paris. www.cr-palevol.fr 91 Corral J.-C et al. Corral J.-C., Berreteaga A., Poyato-Ariza F. J., Bardet N., Cappetta H., Floquet M., Astibia H., Badiola A. & Pereda- Suberbiola X. 2021. — Stratigraphy, age, and vertebrate palaeontology of the latest Cretaceous Quintanilla la Ojada locality (Basque-Cantabrian Region, northern Spain): a synthesis, in Folie A., Buffetaut E., Bardet N., Houssaye A., Gheerbrant E. & Laurin M. (eds), Palaeobiology and palaeobiogeography of amphibians and reptiles: An homage to Jean-Claude Rage. Comptes Rendus Palevol 20 (7): 91-117. https://doi.org/10.5852/cr-palevol2021v20a7 ABSTRACT The Quintanilla la Ojada section (Basque-Cantabrian Region, northern Spain) has yielded two assemblages of Late Cretaceous vertebrates, deposited during the Maastrichtian in coastal environ- ments and related to a transgressive lag at the base of the Valdenoceda Formation. Numerous teeth of Elasmobranchii and Actinopterygii are the most prevailing fossil material, although scarce teeth of marine reptiles (Mosasauridae) and dinosaurs (Hadrosauridae) also occur. The presence of one KEY WORDS Noselachians, hadrosaurian tooth, a terrestrial taxon, constitutes the first report of ornithischians in the Valdenoceda actinopterygians, Formation. The fossil vertebrate association of Quintanilla la Ojada is similar to that discovered in mosasaurids, Albaina (Treviño County, Burgos), also located in the Basque-Cantabrian Region, although relatively hadrosaurians, Maastrichtian, younger in age. Both fossil sites are characterised by a mixture of taxa from the northern and southern Iberian Peninsula. margins of the Mediterranean Tethys (north-European and north-African outcrops). RÉSUMÉ Stratigraphie, âge et paléontologie des vertébrés du gisement finicrétacé de Quintanilla la Ojada (Région Basco-cantabrique, Nord de l’Espagne) : une synthèse. La coupe de Quintanilla la Ojada (Région Basco-Cantabrique, Nord de l’Espagne) a livré deux assem- blages de vertébrés d’âge Crétacé supérieur, déposés durant le Maastrichtien dans des environnements côtiers et liés à un lag transgressif situé à la base de la Formation Valdenoceda. De nombreuses dents d’Elasmobranchii et d’Actinopterygii sont les plus fréquentes, bien que de rares dents de reptiles marins (Mosasauridae) et de dinosaures (Hadrosauridae) aient aussi été trouvées. La présence d’une dent d’hadrosaure, taxon terrestre, constitue la première mention d’ornithischiens dans la Formation MOTS CLÉS Néosélaciens, Valdenoceda. L’association de vertébrés fossiles de Quintanilla la Ojada est similaire à celle découverte actinoptérygiens, à Albaina (Comté de Treviño, Burgos), situé également dans la région Basco-Cantabrique, bien que mosasauridés, d’âge légèrement plus jeune. Les deux sites fossilifères sont caractérisés par un mélange de taxons des hadrosauridés, Maastrichtien, marges nord et sud de la Téthys méditerranéenne (affleurements du Nord de l’Europe et du Nord Péninsule ibérique. de l’Afrique). INTRODUCTION ried intermittently for silica sand, an exploratory trench and a road cut, all located near the village. More specifically, the Until relatively recently, our palaeontological knowledge of the pits lie in a narrow valley, southeast of the village, flanked by Valdenoceda Formation was generally sparse, both in terms of the Vienda Mountain Range to the east and the hilly area micro- and macroinvertebrate studies. Microvertebrate fossils of the periclinal closure of the La Hoz-Lalastra anticline to were described for the first time by Berreteaga (2008) from the west. Nearby towns to the site are Villarcayo, Medina de this Maastrichtian formation near the small village of Quin- Pomar and Trespaderne (see Fig. 1). tanilla la Ojada (Burgos, northern Spain). A few years later, The aim of this paper is twofold. First, it is to introduce the this area was studied by Berreteaga et al. (2011) and Corral vertebrate
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