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(Early Palaeocene), Withers, 1914 Meded. Werkgr. Tert. Kwart. Geol. 25(2-3) 175-196 2 figs, 1 pi. Leiden, oktober 1988 The biostratigraphy of the Geulhem Member (Early Palaeocene), with reference to the occurrence of Pycnolepas bruennichi Withers, 1914 (Crustacea, Cirripedia) by J.W.M. Jagt Venlo, The Netherlands and J.S.H. Collins London, United Kingdom Jagt, &J.S.H. Collins. The biostratigraphy of the Geulhem reference of Member (Early Palaeocene), with to the occurrence Pyc- nolepas bruennichi Withers, 1914 (Crustacea, Cirripedia).—Meded. Werkgr. Tert. Kwart. Geol., 25(2-3): 175-196, 2 figs, 1 pi. Leiden, October 1988. Valves of the brachylepadomorph cirripede Pycnolepas bruennichi Withers, 1914 are reported from the Early Palaeocene of the environs of Maastricht (The Netherlands and NE Belgium). The occurrenceof this distinctive species provides additional proof of a correlationof the with the in Maastricht Danish Danian Early Palaeocene the area. A discussion of the biostratigraphy of the Geulhem Member (Houthem Formation) from which the cirripede remains were collected is presented. It is concluded that P. bruennichi is restricted to strata of Palaeocene in Denmark well in the Early (Danian) age as as Maastrichtian type area (SE Netherlands, NE Belgium). John W.M. Jagt, 2de Maasveldstraat 47, 5921 JN Venlo, The Netherlands; J. S.H. Collins, 63 Oakhurst Grove, East Dulwich, Lon- don SE22 9AH, United Kingdom. Contents 176 Samenvatting, p. Introduction, p. 176 177 Systematic description, p. and distribution of P. 178 Stratigraphic geographic bruennichi, p. of the Geulhem 182 Stratigraphy Member, p. Conclusion, p. 190 191 Acknowledgements, p. References, p. 191. 176 Samenvatting De de Geulhem Kalksteen voorkomen biostratigrafie van (Vroeg Paleoceen) naar aanleiding van het 1914 van Pycnolepas bruennichi Withers, (Crustacea, Cirripedia). de eendemossel bruennichi worden uit het onderste Geisoleerde platen van Pycnolepas gemeld en deel de de Houthem Formatie bovenste van Geulhem Kalksteen van (Vroeg Paleoceen, Danien) de zoals die ontsloten is in twee sleutelprofielen in de omgeving van Maastricht, te weten voormalige de groeve Curfs bij Geulhem (gemeente Valkenburg aan de Geul) en de secties die bij verbreding het Albertkanaal Vroenhoven-Riemst P. van bij (Belgisch Limburg) toegankelijk waren. bruennichi is die het een soort tot nog toe alleen uit Midden Danien van Fakse, Sjaelland, Denemarken gemeld werd. Het in Paleoceen optreden het Vroeg van Maastricht en omgeving is een bijkomend argument de Danien uit het om Geulhem Kalksteen te correleren met het type gebied. In dit kader wordt hier discussie verschillende ook een vrij uitvoerige gepresenteerd over fossielgroepen die voor de Deze de biostratigrafie en correlaties over grotere afstanden van belang zijn. fossielgroepen zijn volgende: planktonische en benthonische foraminiferen, echinodermen, ostracoden, kalkschalig nan- en enkele kleinere noplankton, dinoflagellaten/sporomorfen groepen (bryozoen, serpuliden). Geconcludeerd wordt dat betrouwbare de Kalk- voor een biostratigrafische indeling van Geulhem steen, en zodoende van het Danien in de omgeving van Maastricht, uitgebreid èn gecombineerd onderzoek is alle nodig van bovengenoemde groepen in zoveel mogelijk profielen, die van te voren andere op lithologische en op gronden (bv. gamma ray logging) aan elkaar gekoppeld zijn. Introduction In 1839 Steenstrup described and figured a cirripede scutum, tergum and carina from the Danian of Bryozoan Limestone Fakse, Denmark, as Pollicipes rigidus J. de C. Sowerby, 1836. These elements described Darwin were subsequently by (1851) as a new species, Pollicipes elegans, but this name had been used Lesson for already by (1830) a Recent cirripede species. Maintaining Darwin's name, Nielsen of his however, Brunnich (1912) published additional information, although some figured the later referred specimens attributed to species were to Calantica (Scillaelepas) dorsata (Steenstrup, 1839) by Withers (1914, 1935). When Withers to include this which he the his (1914) came species, to gave name brünnichi, in erected the studied Darwin could newly genus Pycnolepas, original specimens by Steenstrup and no be longer found and he 'reluctantly' selected as type a scutum (Withers, 1914, pi. 7, fig. 6) from a large number of elements recently collected by Brunnich Nielsen from the Bryozoan Limestone of Fakse. In 1935, Withers included Pycnolepas in his suborder Brachylepadomorpha (Withers, 1923) and thoroughly reviewed the description of P. bruennichi. The is known from several species now new localities. It is considered opportune, therefore, to amend the of this occurrences species as they are known now. The of this presence distinctive species of cirripede provides an opportunity to critically review the of the Geulhem Member of the Houthem Formation in the Maastrichtian stratigraphy type area, and a discussion of several of fossils of value with the is groups biostratigraphic occurring cirripede presented. 177 SYSTEMATIC DESCRIPTION Ordo Thoracica Darwin, 1854 Subordo Brachylepadomorpha Withers, 1923 Familia Brachylepadidae Woodward, 1901 Genus Pycnolepas Withers, 1914 Type species—Pollicipes rigidus J. de C. Sowerby, 1836, by original designation. Diagnosis —See Withers (1914, 1935) and Newman et al. (1969). Pycnolepas bruennichi Withers, 1914 Plate 1 1839 Pollicipes rigidus J. de C. Sowerby.—Steenstrup, p. 404, pi. 5, figs 24-26 (non Sowerby). 1851 Pollicipes elegans Darwin, Darwin, p. 76, pi. 4, fig. 9 [non Pollicipes elegans Lesson, 1830 (Cirripedia)] (Partim, not the specimens from Ignaberga, Sweden = Pycnolepas ignabergensis Carlsson, 1953). 1912 Darwin—Briinnich Nielsen, 32, 1, 18-20; 2, 1-8, 11, 12 2, Pollicipes elegans p. pi. figs pi. figs [non pl. figs 9, 10, 13-18 = Calantica (Scillaelepas) dorsata (Steenstrup, 1839)]. 1914 Pycnolepas brünnichi Withers, p. 181, pi. 7, figs 5-9; pi. 8, fig. 6 (with synonymy). 1935 brünnichi Withers 7-15 Pycnolepas —Withers, p. 47, 73, 359, pi. 47, figs (with synonymy). 1953 Pycnolepas brünnichi Withers—Carlsson, p. 24. 1980 Pycnolepas brünnichi Withers—Collins, p. 21, 25. the Middle Danian Type—Withers (1914, pi. 7, fig. 6; 1935, pi. 47, fig. 8) designated a scutum from which has Bryozoan Limestone of Fakse, Sjaelland, Denmark, as 'type specimen', to be considered the holotype. It is kept in the British Museum (Natural History), Department of Palaeontology, Lon- don, catalogue number I. 16.625. studied for this collected three localities in the Maastricht Materialstudied—The material paper was at area (Fig. 1). 43 15 1 and from —34 scuta, terga, rostra/carinae, imbricating plate several fragments of valves the of 1000 western bank of the Albert Canal at Vroenhoven(municipality Riemst, Belgian Limburg), c. N of the the Geulhem Member m bridge, from the lowermost 0.5 m of (Houthem Formation). Locality 4 in Fig. 1. Jagt and Collins collections. 4 and —29 scuta, 17 terga, 21 rostra/carinae, imbricating plates 2 fragmentary upper latera from bank Canal N of from 8- 9 the eastern of the Albert at Vroenhoven, c. 2000 m the bridge, c. m above the Vroenhoven Horizon of Geulhem Member. 5 in 1. (see Fig. 2), upper part Locality Fig. Jagt and Collins collections. — from the Member in the former 1 fragmentary, worn tergum the uppermost metre of Geulhem at Geulhem of de The 1 Curfs quarry (municipality Valkenburg aan Geul, Netherlands). Locality in Fig. 1. Jagt collection. Description —The reader is referred to Brünnich Nielsen (1912) and Withers (1914, 1935) for com- prehensive descriptions of P. brüennichi. 178 in the Remarks—Only isolated elements are present samples studied. They are at times more or less but of excellent Some elements but this is strongly worn, some are preservation. are fragmentary, due the method of found. partly to sample processing. Several growth stages were The number lateral valves in of almost of present a sample Pycnolepas is, invariably, dispropor- the elements. reasonable for be tionate to number of other capitular So far, no explanation this can The minimum number of individuals in assessed the given. present a sample can be by counting number of the more common side (i.e. left or right) of the most common valve - usually a scutum collection of from 5 the Albert Canal or tergum. The made by one us (JWMJ) locality along (see Fig. 1) contains 18 scuta of several growth sizes, yet both left and right latera together number only much of 2. This proportion appears to be the same as encountered in similar samples P. rigidus. STRATIGRAPHIC AND GEOGRAPHIC DISTRIBUTION OF P. BRUENNICHI Withers (1935) reviewed the taxonomic history of P. bruennichi, thoroughly revised the description and documented the geological and stratigraphical data as known at that time. Since 1935 the stratigraphy of the Late Cretaceous and its boundary with the Danian has been considerably revised and recorded P. bruennichi all of Danian will be discussed below. Withers's occurrences for are age, as P. from the Limestone Fakse bruennichi was first recorded (as Pollicipes rigidus) Bryozoan of by Steenstrup (1839). Subsequently, Darwin (1851) united, under the new taxon Pollicipes elegans, two of different viz. Fakse and species totally stratigraphic provenance, (Danian Bryozoan Limestone) Ignaberga (Skane, Sweden; latest Early Campanian, see Christensen, 1975, 1986). The specimens from Fakse were later rightly isolated and described by Withers (1914) as P. bruen- = 1839 de 1851 nichi [ Pollicipes rigidus Steenstrup, non J. C. Sowerby, 1836; Pollicipes elegans Darwin, (partim)]. The Ignaberg
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