Phuket mar. biol. Cent. Res. Bull. 70: 1–6 (2011)

ON THE IDENTITY OF THE MANGROVE CRAB, PARACLEISTOSTOMA ERIOPHORUM NOBILI, 1903 (CRUSTACEA: BRACHYURA: CAMPTANDRIIDAE)

Peter K. L. Ng 1, 2, Cheryl G. S. Tan 2 and Rueangrit Promdam 3

1Tropical Marine Science Institute and Raffles Museum of Biodiversity Research, National University of Singapore, 14, Science Drive 4, Singapore 117543, Republic of Singapore. 2Department of Biological Sciences, National University of Singapore, 14, Science Drive 4, Singapore 119260, Republic of Singapore. 3Reference Collection, Phuket Marine Biological Center, Phuket Marine Biological Center, P.O. Box 60, Phuket, 83000, Thailand. ([email protected]) Corresponding author: P. K. L. Ng, e-mail: [email protected]

ABSTRACT: The identity of the poorly known camptandriid mangrove crab Paracleistostoma eriophorum Nobili, 1903, is clarified. It is shown to be a senior synonym of Paracleistostoma tweediei Tan & Humpherys, 1995, and the taxonomy of the species is discussed, with the range of the species extended to Thailand. Notes on its ecology are also provided.

INTRODUCTION Paracleistostoma eriophorum Nobili, 1903: 23. - Manning & Holthuis, 1981: 209 (list) - Ng The camptandriid genus Paracleistostoma et al., 2008: 233, 234. De Man, 1895, is currently represented by eight Paracleistostoma wardi - Yang, 1979: 39 (list). species (Rahayu & Ng, 2003; Ng et al., 2008). - Harminto, 1988: 88 (nec P. wardi Rathbun, Ng et al. (2008: 233, 234) commented that the 1926). - Tan & Ng, 1994: 83 (list). poorly known Paracleistostoma eriophorum Paracleistostoma tweediei Tan & Humpherys, Nobili, 1903, was actually a senior synonym of 1995: 251, figs. 1–3. Paracleistostoma tweediei Tan & Humpherys, Paracleistostoma tweediei Tan & Ng, 1995: 608. 1995, but did not elaborate. We here redescribe and figure the type of Material examined. - Holotype male of P. Paracleistostoma eriophorum Nobili, 1903, and eriophorum (7.5 x 10.0 mm) (MZUT Cr1200, ex comment on its taxonomic relationships. 1531), Buntal, Sarawak, East Malaysia, don. R. Shelford, 1902. Others - SINGAPORE: 1 male MATERIALS AND METHODS (holotype of P. tweediei) (7.9 x 10.2 mm) (ZRC 1987.57), 8 males (paratypes of P. tweediei) (5.3 The terminology follows that in Tan & x 6.7 mm, 6.4 x 8.3 mm, 6.6 x 8.4 mm (ZRC Ng (1999). Measurements provided, in millimetres, 1987.60–62), 7.4 x 9.4 mm, 7.5 x 9.7 mm, 7.5 x are of the carapace length and width respectively. 10.0 mm, 8.3 x 10.7 mm, 8.8 x 11.5 mm (ZRC Specimens are deposited in the Museo Regionale 1987.57–59), Lim Chu Kang mangrove, coll. P. di Scienze Naturali (MZUT), Sezione di Zoologia, K. L. Ng & S. Harminto, 13 September 1986; 1 Turin, Italy; Zoological Reference Collection (ZRC) male (7.2 x 9.3 mm) (ZRC 1987.1382), of the Raffles Museum of Biodiversity Research, Woodlands, Singapore, coll. S, Harminto, 2 June National University of Singapore, Singapore; 1987. PENINSULAR MALAYSIA: 18 males (ZRC Phuket Marine Biological Centre (PMBC), Phuket, 1998.1007), Matang mangroves, Perak, coll. C. Thailand; and Queensland Museum (QM), G. S. Tan, 8 June 1995; 4 males (5.0–6.9 x 6.1– Brisbane, Australia. 8.8 mm) (ZRC 1998.998), Matang mangroves, Perak, coll. A. Ramli, M. Faizal, C. G. S. Tan, 10 Paracleistostoma eriophorum Nobili, 1903 June 1995. BRUNEI: 1 male (9.3 x 11.9 mm) (QM (Figs. 1, 2) W19967), Batu Marang, coll. N. A. Mackenzie, 2 Phuket mar. biol. Cent. Res. Bull.

Figure 1. Paracleistostoma eriophorum Nobili, 1903. A, holotype male (7.5 x 10.0 mm) (MZUT Cr1200, ex 1531), Sarawak; B, C, male (7.6 x 9.6 mm) (ZRC), Thailand. A, B, overall views; C, left G1 in situ.

March 1993. THAILAND: 1 male (7.6 x 9.6 mm), Type locality. - Buntal, Sarawak. 1 female (6.5 x 8.3 mm) (ZRC 2010.0071), 1 male (6.9 x 8.5 mm) (ZRC 2010.0072); 3 males (10.9 Diagnosis (holotype male of P. eriophorum). - x 8.3 mm, 10.4 x 8.0 mm, 10.0 x 7.7 mm), 3 Carapace surface smooth, slightly convex, broadly females (12.0 x 9.0 mm, 9.0 x 7.0 mm, 8.8 x 6.8 quadrangular, broader than long (1.2–1.3 times); mm) (PMBC 23738), mangroves, Khanom District, regions indistinct, epigastric lobes moderately low; Nakhon Srithammarat, Gulf of Thailand, coll. R. metagastric groove shallow. Front almost straight Promdam & A, Kaenphet, 23 June 2007. from dorsal view, upper surface slightly concave 3 Identity of the mangrove crab

Figure 2. Paracleistostoma eriophorum Nobili, 1903. A, G–I, holotype male (7.5 x 10.0 mm) (MZUT Cr1200, ex 1531), Sarawak; B–F, male (7.9 x 10.2 mm) (ZRC 1987.57), holotype of P. tweediei Tan & Humpherys, 1995, Singapore. A, B, male abdomen; C, frontal view showing epistome; D, anterior part of male thoracic sternum; E, outer view of left chela; F, G, dorsal view of right carpus of cheliped; H, I, left G1 (setae not drawn). Scales: A, G = 0.5 mm; H, I = 1.0 mm. medially, anterolateral angles of front rounded; dense setae; posterolateral margin lined with small frontomedial projection triangular, apex rounded; granules, completely obscured by pubescence; orbital margins setose; supraorbital margin weakly posterior carapace margin straight. Anterior and granular, sinuous; infraorbital margin straight, finely posterior halves of epistome separated by median granular along central third, inner infraorbital tooth transverse ridge; postero medial-tooth broadly well developed, distinct from rest of infraorbital triangular, ending in small, pointed extremity, margin; suborbital ridge defined by line of granules; posterior margin on either side almost straight; posterolateral border and surfaces of carapace with sulcus separating epistome from anterior 4 Phuket mar. biol. Cent. Res. Bull.

pterygostomian margin indistinct. Third maxilliped the appendages detached. Most of the setae on ischium broadly rectangular, outer distal margin the carapace have also been lost, although one can with row of long setae along outer half; merus discern where the main setose areas were (Fig. with well developed outer distal angle which is 1A). The distal part of the G1 of the holotype male bilobed, outer surface uneven, inverted V shaped is somewhat less swollen (Fig. 2H, I) compared sulci on outer surface indistinct. Chelipeds to the recent material (Fig. 1C), but this is almost subequal, enlarged; carpus subglobular, outer certainly because of its age and some dehydration. surface almost smooth, inner surface sparsely Paracleistostoma eriophorum appears to granular, inner dorsal edge finely beaded with be close to P. depressum and is distinguished from granules, appearing subcarinate, inner ventral edge all its congeners by the unique structure of the with large triangular granules, with short row of G1, which has the distal appendage modified to long setae.Ambulatory legs relatively long, densely form a spine like process. It is one of the two setose; first and last pairs of ambulatory legs Paracleistostoma species having the inner ventral almost equal in length, first pair with setae only on edge of the carpus lined with sharp granules, the dorsal surface of merus. Abdominal somites 2–5 only other Paracleistostoma species exhibiting this completely fused, sutures not discernible, suture condition being P. longimanum. Also, in P. between somites 2 and 3 visible only as short, eriophorum, the ridge below the infraorbital margin shallow grooves on either side of abdomen; somite is formed by a raised line of granules. This 1 broader than somite 2, with transverse mdian condition is also found in P. wardi. ridge; somite 6 shorter than telson, lateral margin Neither Nobili (1903) nor Tan & sinuous. G1 recurved, apex large, globular, with Humpherys (1991) described the epistome and the short but distinct submedian spine-like distal anterior sternal structure, and these are included appendage. in the present description (Fig. 2C, D). Only one female specimen of this species (from Thailand) Remarks. - Nobili’s (1903) description of is known. It closely resembles the male in all non- Paracleistostoma eriophorum was relatively sexual features except that the chelae are even detailed but he did not provide a figure. He more slender, and the patches of setae on the distinguished P.eriophorum from P.depressum and posterolateral regions are less clearly demarcated, P. cristatum by the unusually tomentose partially because there are also short setae on the ambulatory legs and the presence of two carinae dorsal parts of the carapace which are normally of long granules on the inferior and superior glabrous in males. The female abdomen is margins of the outer surface of the cheliped palm. subcircular, covers the thoracic sternum, and all Nobili (1903) also mentioned that the posterior the somites and telson are mobile. carapace, including the median branchial region Harminto (1988) recorded this species as and the area around the intestinal region, bears a P. wardi (Rathbun, 1926). His specimens were thick, short felt like tomentum, and the ambulatory examined and compared withAustralian specimens legs were covered with long, woolly setae with of P. wardi in the ZRC and were found to differ the dactyli bearing sulci and setae. From his significantly from the latter species in terms of comparisons, Serène (1974:64) noted “... it is the structures of the G1, epistome and male possible that eriophorum is a Leipocten or Baruna”. cheliped carpus. Tan & Ng (1994) had incorrectly After examining the type, there is no doubt recorded this species as P. wardi in their list of that P.eriophorum is a species of Paracleistostoma. mangrove Brachyura. AsnotedbyNget al. (2008), P.eriophorum Nobili, The figure of the frontal region and 1903, is a senior synonym of P. tweediei Tan & abdomen provided by Tan & Humpherys (1995) Humpherys (1995). The type specimens of both is somewhat inaccurate in that the inner infraorbital species are of similar size, and agree closely in all tooth is not clearly shown (Tan & Humpherys, characters. The holotype male of P. eriophorum 1995: Fig. 2A) and the sutures between abdominal is not in a good condition (Fig. 1A) with most of segments 3–5 are shown to be present as notches 5 Identity of the mangrove crab

on either side of the abdomen (Tan & Humpherys, 1995, priority over P. tweediei C. G. S. Tan & 1995: Fig. 2G). In fact, only the suture between Ng, 1995.” somites 3 and 4 is visible as a notch on either side of the abdomen. The above features are refigured Ecology. - In Perak, Malaysia, Paracleistostoma here (Fig. 2A, B). eriophorum was collected from the Matang As discussed by Ng et al. (2008), mangroves along with specimens of P. depressum there is a nomenclatural problem with the names and the varunid Metaplax elegans De Man, 1888. Paracleistostoma tweediei Tan & Humpherys, These were all found in burrows in soft, brown 1995, and Paracleistostoma tweediei Tan & Ng, mud, close to the edge of the river. In the Khanom 1995. Ng et al. (2008: 234) noted that “Both are District, Thailand, P. eriophorum were found in objective synonyms as they are based on the same mangrove habitats near rivers, with burrows holotype. The problem arose because C. G. S. Tan typically occurring in the areas between nearby and P. K. L. Ng originally intended to publish the mangrove trees. Sympatric species there, include new species, and the paper was prepared for a the sesarmids Episesarma palawanense (Rathbun, regional symposium in 1994 (Tan & Ng, 1995). 1914), Parasesarma ungulatum (H. Milne At the same time, P. J. F. Davie and A. Humpherys Edwards, 1853), Perisesarma eumolpe (De Man, had independently also discovered the species. As 1895) and P. indiarum (Tweedie, 1940). it was clear that the two taxa in question were conspecific, P. J. F. Davie and P. K. L. Ng left it Distribution. - Buntal, Sarawak, East Malaysia; to C. G. S. Tan and A. Humpherys to finish the Brunei, Peninsular Malaysia, Singapore, and now new species description on their own (Tan & for the first time from the eastern coast of Thailand. Humpherys, 1995). In early 1995, P. K. L. Ng asked the editors of the symposium volume to ACKNOWLEDGEMENTS have the paragraph on the new Paracleistostoma in their paper deleted, but although they agreed, The authors thank Giovanni Balma of the the changes were not made, no proofs were sent, University of Turin for his help in obtaining the and the volume was published in 1995 with the type of P. eriophorum. Part of the study was problem paragraph still intact. This mistake supported by research grants to the first author nevertheless validates P. tweediei C. G. S. Tan & from the National University of Singapore. The Ng, 1995. In any case, the symposium article third author was supported by the TOTAL came out later than the paper in the Raffles Bulletin Corporate Foundation, TOTAL E&P Thailand and of Zoology. Since the symposium volume did not the TRF/BIOTEC Special Program for Biodiversity have a publication date, under the Code, it should Research and Training Grant BRT R_149023 and be regarded as published on 31 December 2005. he would like to thank Arwut Kaenphet, staff of This gives P. tweediei C. G. S. Tan & Humpherys, Walailak University, who worked closely with him during the field trips to the mangroves of Khanom.

REFERENCES

Harminto, S. 1988. Systematics of the family Ocypodidae Rafinesque, 1815, s. lat. (Crustacea: Decapoda: Brachyura) of Southeast Asia. Unpublished M.Sc. thesis, Department of Zoology, National University of Singapore. Manning, R.B. & L.B. Holthuis. 1981. WestAfrican Brachyuran Crabs (Crustacea: Decapoda). Smithson. Contr. Zool. 306: 1–379. Ng, P.K.L., D. Guinot & P.J.F. Davie. 2008. Systema Brachyurorum: Part I. An annotated checklist of extant brachyuran crabs of the world. Raffles Bull. Zool., Suppl. 17: 1–286. Nobili, G. 1903. Contributo alla fauna carcinologica di Borneo. Boll. Mus. Zool. Anat. comp. Torino. 18(447): 1–32, 3 figs. 6 Phuket mar. biol. Cent. Res. Bull.

Rahayu, D.L. & P.K.L. Ng, 2003. The Camptandriidae of Irian Jaya, Indonesia, including the descriptions of two new species (Decapoda: Brachyura). J. Crust. Biol. 23(4): 951–962. Rathbun, M.J. 1926. Brachyuran crabs from Australia and New Guinea. Rec. Aust. Mus. 15(2): 177– 182, pls. 14–16. Serène, R. 1974. Note on the genera and species of the Camptandriinae Stimpson, 1858 (Decapoda, Brachyura: Ocypodidae). Treubia. 28(3): 59–117. Tan, C.G.S. & A. Humpherys. 1995. Paracleistostoma tweediei, a new species of camptandriine crab, from Singapore and Brunei mangroves (Crustacea: Decapoda: Brachyura: Ocypodidae). Raffles Bull. Zool. 43(1): 251–256. Tan, C.G.S. & P.K.L. Ng. 1995. The Camptandriinae of Singapore and Malaysia with description of one new species of Paracleistostoma De Man, 1895 (Crustacea: Decapoda: Brachyura: Ocypodidae). Proc 3rd ASEAN-Australia Symp. Living Coastal Resources Vol. 2: research papers. C. R. Wilkinson, S. Sudara & L. M. Chou (Eds.), Bangkok, pp. 607–611. Tan, C.G.S. & P.K.L. Ng. 1994. An annotated checklist of mangrove brachyuran crabs from Malaysia and Singapore. Hydrobiologia. 285:75–84. Tan, C.G.S. & P.K.L. Ng. 1999. A revision of the genus Camptandrium Stimpson, 1858 (Crustacea: Decapoda: Brachyura: Camptandriidae). Raffles Bull. Zool. 47(1): 193–219. Yang, C. M. 1979. A list of Brachyura in the Zoological Reference Collection of the Department of Zoology. Unpublished checklist, Department of Zoology, University of Singapore, 60 pp. (mimeographed).

Manuscript received: 20 October 2009 Accepted: 9 April 2010 Phuket mar. biol. Cent. Res. Bull. 70: 7–14 (2011)

NEW RECORDS OF SPIDER CRABS OF THE GENERA CYRTOMAIA MIERS, 1886, AND PLATYMAIA MIERS, 1886 (DECAPODA: MAJOIDEA: INACHIDAE) FROM THE ANDAMAN SEA, THAILAND

Rueangrit Promdam

Department of Biology, Faculty of Science, Prince of Songkla University, Hatyai, Songkhla 90112, Thailand Email: [email protected]

ABSTRACT Two species of deep-water spider crabs, Cyrtomaia suhmii Miers, 1886, and Platymaia alcocki Miers, 1886, based on the materials of the BIOSHELF Project (1996–2000) are recorded for the first time from Thailand. It is also the first record of these two genera from the Andaman Sea coast of Thailand. Adult and juvenile forms of C. suhmii collected from the same locality confirm that C. curviceros Bouvier, 1915, is a junior synonym, the characters previously used to separate them explained by size and age.

Key words: new records, spider crabs, Decapoda, Majoidea, Inachidae, Cyrtomaia, Platymaia, Thailand

INTRODUCTION TAXONOMY

Two rare spider crabs, Cyrtomaia suhmii Family Inachidae MacLeay, 1838 Miers, 1886, and Platymaia alcocki Miers, 1886, Genus Cyrtomaia Miers, 1886 were collected by the R.V. Chakratong Tongyai Cyrtomaia suhmii Miers, 1886 from the Andaman Sea during the BIOSHELF (Figs. 1, 2A–F) project surveys of the west coast of Thailand in 1996–2000. Based on Ng and Davie (2002), and Cyrtomaia suhmii Miers, 1886: 16, pl. 3, fig. 2, Naiyanetr (2007), the species as well as the genera 2a–2c; Rathbun, 1918: 6; Richer de had not been previously reported from Thai Forges and Guinot, 1990: 525; Richer de waters. The present report serves to formally Forges and Ng, 2007: 56 (list); Ng et al., record these species, as well as to discuss variation 2008: 111 (list). in adult and juvenile specimens. Cyrtomaia suhmi - Rathbun, 1893: 230; Griffin, 1974: 9; Guinot and Richer de Forges, MATERIALS AND METHODS 1982a: 1096; 1982b: 21, fig. 10, 11A–B, 23B; Griffin and Tranter, 1986b: 352, figs. Specimens examined are deposited in the 1, 2; Richer de Forges and Guinot, 1988: Reference Collection of Phuket Marine Biological 42–43 fig. 2B, C, pl. 2F–G; Poore, 2004: Center (PMBC). All measurements provided are 360; 361 fig. 108. pre-rostral lengths and carapace widths Cyrtomaia Suhmi var. curviceros Bouvier, 1915b: respectively. For details on all BIOSHELF stations, 9–15, pls. 1. see Aungtonya et al. (2000). The classification Cyrtomaja suhmi typica Serène and Lohavanijya, used here follows Ng et al. (2008). 1973: 46 (in key). 8 Phuket mar. biol. Cent. Res. Bull.

Figure 1. Cyrtomaia suhmii, adult male (60.2 × 67.1 mm) (PMBC 19999), dorsal view of whole animal. Scale = 20 mm.

Cyrtomaja suhmi curvicornis Serène and 488 m, coll. S. Bussarawit and C. Aungtonya, Lohavanijya, 1973: 45, 46 (in key). 06.02.1999; PMBC 20000, 2 juv. (11.8 × 11.5 mm, Cyrtomaia curviceros Sakai, 1976: 181; Guinot and 8.6 × 8.2 mm), BIOSHELF St. J10, 0715´N, Richer de Forges, 1982a: 1096; 1982b: 09715´E, Agassiz trawl, 695–683 m, coll. C. 24, fig. 12A–D; Richer de Forges and Aungtonya & V. Vongpanich, 19.02.2000; PMBC Guinot, 1988: 42–43, fig. 2A; Ng et al., 19929, 1 juv. (7.7 × 7.4 mm), 1 female (56.3 × 2001: 13; 81 fig. 3h; Richer de Forges 61.2 mm), BIOSHELF St. L10, 0645´N, 09723´E, and Ng, 2007: 56 (list). Agassiz trawl, 707–651 m, coll. C. Aungtonya & not Cyrtomaia suhmi - Griffin and Brown. 1976: V. Vongpanich, 21.02.2000 252, fig. 6; Griffin and Tranter, 1986a: 30, fig. 91g [= C. griffini Richer de Diagnosis Forges and Guinot, 1990]. Carapace smooth, not granular or not Cyrtomaia suhmi typica Doflein, 1904 : 54– spinulate, with 2 long, subparallel protogastric 55, pl. 19: figs. 1, 2 [= C. gaillardi Guinot spines. Pseudorostral spine divergent, arched. and Richer de Forges, 1982]. Basal antennal article cylindrical, armed with 3–5 spines (include internal spine). Orbital border Material examined smooth, intercalated orbital granule present (in PMBC 19999, 1 male (60.2 × 67.1 mm), young) or absent, without preocular spines (in BIOSHELF St. E8, 0832´N, 09604´E, Otter trawl, adults). Ocular spine blunt. 9 New records on spider crabs

Figure 2. Cyrtomaia suhmii: A–C, juvenile (11.8 × 11.5 mm) (PMBC 20000); D–F, adult male (60.2 × 67.1 mm) (PMBC 19999); A, D, dorsal view; B, E, anterior part of carapace showing rostrum; C, F, lateral view of carapace. Scale = 10 mm.

Distribution Remarks Indo-West Pacific: Japan, Philippines, Cyrtomaia suhmii was described by Miers Indonesia, Australia, Thailand (Andaman Sea) and (1886) on the basis of an imperfect crushed young India. male specimen collected from Talaud Islands (Tulur 10 Phuket mar. biol. Cent. Res. Bull.

Islands) (see Guinot & Richer de Forges 1982b: are similar to the original description and figure of 22, fig. 10), Indonesia, by the Challenger C. suhmii by Guinot and Richer de Forges (1982b: Expedition. Subsequently, Cyrtomaia suhmii 22, 23, fig. 10, 11A, B), especially with regards to curviceros Bouvier, 1915, was described from Japan the protogastric spines which are divergent (Miers, and recognized as a distinct species by Sakai 1886: 16, pl. 3, fig. 2). On the other hand, large (1976). The status of C. curviceros has long been male specimens resemble C. curviceros (Fig. 1, uncertain. Griffin and Tranter (1986) treated both 2D–F), although the number of the spines on the names as synonyms, while Richer de Forges and basal antennal article is slightly different. This Guinot (1982a, 1988) argued that it may be a good difference, however, is not significant and can species. Ng and Huang (1997) treated Taiwanese easily be explained by variation. The present series material of the species as C. curviceros (see also of specimens confirm that the differences which Ng et al., 2001), and until recently, was still have been observed in the past between these two regarded as a distinct species (see Richer de Forges species are due to size and age, and that C. suhnii and Ng, 2007). In their annotated checklist of extant curviceros Bouvier, 1915, clearly is a junior brachyuran crabs of the world, Ng et al. (2008) synonym of C. suhmii Miers, 1886. treated C. suhmii curviceros as a synonym of C. suhmii but without comments. Examination of Genus Platymaia Miers, 1886 more material of other allied species like C. griffini Richer de Forges & Ng, 1990, has shown that the Platymaia alcocki Rathbun, 1916 characters used to separate C. curviceros from C. (Figs. 3, 4A–F) suhmii are not valid at the species level (P. K. L. Ng, pers. comm., see also Richer de Forges and Platymaia wyville-thomsoni. - Wood-Mason and Ng, 2008). Alcock, 1891: 258; Alcock, 1895: 181– Specimens of various sizes were collected 182; Doflein, 1904: 59–69, 160–165, figs. from the same Thai locality. The young specimens 2–6; pls. 2, 20–23, 39, 43: figs. 5, 6; pl.

Figure 3. Platymaia alcocki, adult male (71.0 × 77.5 mm) (PMBC 20367), dorsal view of whole animal. Scale = 20 mm. 11 New records on spider crabs

Figure 4. Platymaia alcocki: A–C, adult male (71.0 × 77.5 mm); D–F, young male (25.9 × 30.4 mm) (PMBC 20367); A, D, dorsal view; B, E, anterior part of carapace showing rostrum; C, F, lateral view of carapace. Scale = 10 mm.

50: figs. 2, 5, 6 [not Platymaia wyville- 1986a: 48 (Pl. 4b; figs. 10a, b; 11c); thomsoni Miers, 1886]. 1986b: 357; Takeda and Webber, 2006: Platymaia alcocki Rathbun, 1916: 530; 1918: 8; 194. Serène and Lohavanijya, 1973: 48 (in key); Not Platymaia alcocki - Takeda and Miyake, 1969: Griffin, 1974: 27; Griffin and Tranter, 498–500, figs. 10a, b, 11a–c; Sakai, 1976: 12 Phuket mar. biol. Cent. Res. Bull.

176–177, fig. 94a; Dai and Yang, 1991: Remarks 120 [=Platymaia wyville-thomsoni Miers, Platymaia alcocki can sometimes be 1886]. misidentified with P. wywillethomsoni because young and juveniles of P. alcocki also have Material examined prominent spines on all the carapace regions PMBC 20367, 4 male (71.0 × 77.5 mm, 25.9 × (Alcock, 1895). Although all the spines on the 30.4 mm, 20.6 × 22.7 mm, 20.2 × 22.8 mm), 5 dorsal carapace surface of adult P. alcocki are female (19.4 × 21.4 mm, 19.0 × 21.4 mm, 19.0 × reduced to only tubercles (Fig. 4A–C), adult P. 22.1 mm, 18.7 × 21.2 mm, 18.4 × 20.3 mm), wywillethomsoni are still armed with one or two BIOSHELF St. J8, 0715´N, 09730´E, Agassiz spines on each side of the protogastric regions trawl, 490–479 m, coll. S. C. Aungtonya & V. even as adults (Griffin and Tranter, 1986a: 307, Vongpanich, 18.02.2000 fig. 5b). The main diagnostic feature of P. alcocki is the deep groove on each side of mesogastric Diagnosis region, which separates the two adjacent branchial Carapace tranversly subcircular, surface regions (Rathbun, 1916) (Fig. 4A, D). This nearly smooth in adults, rostral spine subequal to character is useful even for young and adult interantennular spine, preocular spine absent; specimens of P. alcocki and is clearly evident on protogastric ridge with 2 tubercles, lacking spines the series of Thai specimens here. (in adults); groove on each side of mesogastric region deep, clearly separating adjacent branchial ACKNOWLEDGEMENTS regions; palm of male chelipeds more than twice as long as high, distally increasing in width; The author would like to thanks Peter K. propodus of ambulatory legs flattened, 7.5 times L. Ng of the Raffles Museum of Biodiversity width of fourth ambulatory leg. Research, National University of Singapore, for reviewing the manuscript. Distribution Indian Ocean: Andaman Sea to eastern Africa

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Manuscript received: 9 February 2011 Accepted: 1 August 2011 Phuket mar. biol. Cent. Res. Bull. 70: 15–27 (2011)

NEW RECORDS OF THE BOBTAIL SQUID, EUPRYMNA HYLLEBERGI NATEEWATHANA, 1997 WITH DESIGNATION OF A NEOTYPE

Charatsee Aungtonya1, Anuwat Nateewathana2, Ole Secher Tendal3 and Jaruwat Nabhitabhata4

1Phuket Marine Biological Center, P.O. Box 60, Phuket 83000, Thailand 2Department of Marine and Coastal Resources, Bangkok, Thailand 3Zoological Museum, SNM, University of Copenhagen, Universitetsparken 15, DK-2100 Copenhagen, Denmark 4Centre for Biodiversity of Peninsular Thailand (CBIPT), Faculty of Science, Prince of Songkla University, Hatyai, Songkhla 90112, Thailand Corresponding author: Charatsee Aungtonya Email: [email protected]

ABSTRACT: A total of 265 specimens, 118 males and 147 females, of the bobtail squid, Euprymna hyllebergi, are registered in the Reference Collection, Phuket Marine Biological Center, Thailand. Nearly all specimens were collected from Thai waters, the Andaman Sea and the Gulf of Thailand. The species is here reported as a new record for Myanmar waters and the distribution in the Indian Ocean has been extended to the west of the Andaman Islands. A male neotype is designated on a specimen collected from the original locality in order to replace the lost male holotype specimen. The neotype is deposited in the Reference Collection of Phuket Marine Biological Center, Thailand, as is the remaining female paratype, whereas selected specimens are deposited in National Science Museum Tokyo, Japan, U.S. National Museum, Washington D.C., USA and the Zoological Museum, SNM, University of Copenhagen, Denmark. Measurements of the neotype, paratype, and selected specimens are presented and discussed.

Key word: bobtail squid, Euprymna hyllebergi, Thai waters, neotype

INTRODUCTION placed as invalid while four species, E. albatrossae Voss, 1963, E. bursa (Pfeffer, 1884), E. phenax Bobtail squids are small benthic Voss, 1963, and E. stenodactyla (Grant, 1833), cephalopods, belonging to the family Sepiolidae were treated as unresolved species (Norman & Leach, 1817, which contains about 14 genera and Lu, 1997). E. albatrossae, E. bursa, and E. phenax over 50 species. Sepiolid squids are distributed were known only from the types. The original type worldwide, ranging from tropical to temperate and material of E. stenodactyla is lost, no further sub-polar waters of all oceans (Nateewathana, material has been found in Mauritius yet and the 1997a). description of the species is poor (Norman & Lu, According to the most recent revision by 1997). Furthermore, a nominal species, E. similis Norman & Lu (1997) and a later paper on the genus Sasaki, 1913, was considered a junior synonym by Nateewathana (1997b), 13 species have been of E. morsei (Norman & Lu, 1997). referred to Euprymna Steenstrup, 1887. Six The species E. morsei, E. stenodactyla, species, E. berryi Sasaki, 1929, E. hoylei Adam, E. berryi and E. hyllebergi, have been reported 1986, E. hyllebergi Nateewathana, 1997, E. morsei from Thai waters, the Andaman Sea and the Gulf (Verrill, 1881), E. scolopes Berry, 1913, and E. of Thailand, by Chotiyaputta et al. (1992), tasmanica (Peffer, 1884) are currently accepted Chotiyaputta (1993), Supongpan (1995), in the genus. Two species, E. pusilla (Peffer, Nateewathana (1997b) and Nateewathana et al. 1884) and E. schneehageni (Peffer, 1884), were (2001). 16 Phuket mar. biol. Cent. Res. Bull.

According to Nateewathana et al. (2001) the male neotype of Euprymna hyllebergi in order the occurrence of E. morsei, E. stenodactyla, and to replace the lost male holotype and paratypes E. berryi in Thai waters was still uncertain since previously listed in Aungtonya et al. (2006, 2007), the voucher material could not be located. and describes specimens deposited in other However, E. morsei and E. stenodactyla from the museums. This study aims to facilitate future Andaman Sea and Myanmar waters have been revision and taxonomic work on this genus and to registered at the PMBC Reference Collection in update the known distribution of the species. 2007. Future taxonomic study on those specimens is pending. MATERIALS AND METHODS All specimens of Euprymna hyllebergi deposited in the Reference Collection of Phuket The bobtail squids were obtained from Marine Biological Center, were checked and fishing boats, research vessels, fish markets, and measured, but none were in accordance with the fish landings along the Andaman Sea coast of holotype. The first and the fourth authors Thailand and the Gulf of Thailand as described by contacted the cephalopod taxonomists who had Nateewathana (1997b) and Nateewathana et al. previously studied the type material, but they all (2001). Additional material for the present study denied any knowledge about the fate of those types. was primarily obtained from the R.V. Chulaphorn We conclude that the holotype is lost. of the Deep Sea Fishery Technology Research and Euprymna hyllebergi is close to E. hoylei, Development Institute operating off the Andaman but differs in the arrangement of suckers on the Sea coast of Thailand, and from fishing boats, fish hectocotylized arm, the size and distribution of markets and fish landings along the Andaman Sea enlarged suckers on arms II–IV, and in the length coast of Thailand and the Gulf of Thailand. of the tentacles (Nateewathana, 1997b: Fig. 3). Taxonomic studies are based on Nateewathana E. hoylei is known from the tropical (1997b). Western Pacific to North-West Australia (Norman The specimen designated as the neotype & Lu, 1997). It is characterized by lacking enlarged and five lots of specimens were selected from the suckers on arm II in mature males, having 3–4 samples in the Reference Collection of Phuket enlarged suckers in dorsal and ventral rows of arm Marine Biological Center. The neotype was, like pair IV and ventral row only of arm pair III the holotype, collected from small scale fishing (Norman & Lu, 1997, Nesis, 1987). The suckers areas in Trang Province, at a depth of less than 40 on arm II–IV of E. hyllebergi do not have any m off the Andaman Sea Coast of Thailand. significantly or abruptly enlarged suckers Measurements follow those of (Nateewathana, 1997b). E. hyllebergi has a high Nateewathana (1997b). Definitions of number of stout papillae with a slitlike aperture in measurements and indices are summarized in Table the distal half of the hectocotylus. The papillae of 1 and in Figure 1. The difference of the means of E. hyllebergi are crowded in rows of 4–6 suckers all the measurements and indices is compared with in the middle part, and reduced to two rows at the those listed by Nateewathana (1997b). Differences tip of the hectocotylus. were considered significant when The male characters are the most important features distinguishing the two species, E. hoylei and E. hyllebergi. Furthermore, the Specimens of the species, the newly arrangement of the sessile suckers of the male is designated neotype, and selected materials are the only diagnostic feature to characterize the deposited in the Reference Collection of Phuket species of the genus, as well as other genera in Marine Biological Center, Thailand (PMBC), the Family Sepiolidae. Hence it is necessary to National Science Museum Tokyo, Japan (NSMT), select a male neotype of E. hyllebergi. U.S. National Museum, Washington, D.C., USA The present report lists all specimens of (USNM), and the Zoological Museum, SNM, this species in the Reference Collection, designates University of Copenhagen, Denmark (ZMUC). 17 New records of the bobtail squid

Table 1. Definition of counts, measurements and indices (modified after Nateewathana, 1997b).

ML Mantle Length dorsal mantle length measured from the anterior-most point of mantle to posterior tip, along dorsal mid-line. MWI Mantle Width Index greatest straight-line (dorsal) width of mantle as a percentage of mantle length. FLI Fin Length Index greatest length of fins as a percentage of mantle length. FWI Fin Width Index greatest width (dorsally) across both fins as a percentage of mantle length. FBI Fin Base Index length of fin base as a percentage of mantle length. HLI Head Length Index dorsal length of head measured from point of fusion of dorsal arms to anterior tip of nuchal locking cartilage as a percentage of mantle length. HWI Head Width Index width of head across midpoint of eyes as a percentage of mantle length. ALI Arm Length Index length of arm measured from first basal (proximal-most) sucker to tip of arm as a percentage of mantle length. ASI Arm Sucker Index diameter of largest normal arm sucker on each designated arm as a percentage of mantle length. TtLI Tentacle Length Index total length of tentacular stalk and club as a percentage of mantle length. ClLI Club Length Index length of club as a percentage of mantle length. HcLI Hectocotylized Length Index length of modified portion of arm measured from proximal-most modified sucker to tip of arm as a percentage of total length of hectocotylized arm (left arm I). EDI Eye Diameter Index diameter of eye across bulbus as a percentage of mantle length. LnDI Lens Diameter Index diameter of eyelens as a percentage of mantle length.

RESULTS and ZMUC CEP-25 have been selected from PMBC 11742, PMBC 11748 and PMBC 11747, A total of 265 specimens, 118 males and respectively. Two females of PMBC 11734 have 147 females, were registered in the Reference been selected and registered as NSMT-Mo 76425 Collection (see Table 2). Nearly all of the specimens and ZMUC CEP-51. The male neotype, PMBC were collection from Thai waters, the Andaman accession number 21339, is deposited in the Sea and the Gulf of Thailand, except for PMBC Reference Collection of Phuket Marine Biological 11781 and PMBC 11775 which were collected Center, as is the remaining female paratype, PMBC from the Indian Ocean, west of the Andaman 11731. Islands and Myanmar waters, respectively. The Selected measurements and indices of the latter specimens are the first records of E. type materials and selected specimens of E. hyllebergi from the Indian Ocean, west of the hyllebergi are presented in Table 3. Mean of indices Andaman Islands and Myanmar waters, (in percent), standard deviation of five males and respectively. four females are presented in Table 4. The t-test The neotype was selected from PMBC comparing the means of all the measurements and 11742 and registered as PMBC 21339 (Fig. 2). indices of the present material and those given by Three lots of NSMT-Mo 76424, USNM 1150474, Nateewathana (1997b) are presented in Table 5. 18 Phuket mar. biol. Cent. Res. Bull.

Figure 1. Diagrammatic illustration of measurements in sepiolids (modified after Nateewathana, 1997b). AL = arm length, C1L = club length, ED = eye diameter, FB = fin base, FL = fin length, FW = fin width, HL = head length, HW = head width, LD = lens diameter, ML = mantle length, MW = mantle width, TtL = tentacle length.

Euprymna hyllebergi Nateewathana, 1997 Phangnga Bay, push net, 18 June 1988; USNM (Figs. 2A–B, 3A–B, 4A–C, Table 2–5) 1150474, 1 male and 1 female, Phuket fish market, Phuket Province, 23 Aug 1995; ZMUC CEP-25, 2 Euprymna hyllebergi – Nateewathana, 1997b: 466– males, Krabi fish market, Krabi province, 17 Dec 474, figs. 2–5, Table 2. 1994; ZMUC CEP-51, 1 female, Chong Pra, Phangnga Bay, push net, 18 June 1988. Neotype. PMBC 21339, 1 male, Kantang fish landing, Trang province, 14 Feb 1996 (Figs. 2– Material examined. 265 specimens have been 3). examined and registered (see Table 2). Paratype. PMBC 11731, 1 female, Tha Lane, Phangnga Bay, push net, 23 July 1989 Description. Colour in alcohol yellowish; numerous large and conspicuous purplish brown Selected specimens. NSMT-Mo 76424, 1 male, chromatophores cover mantle, head and arms, Kantang fish landing, Trang Province, 14 Feb more dense in middle part of dorsal and ventral 1996; NSMT-Mo 76425, 1 female, Chong Pra, sides. Mantle dome-shaped, thick, broad 19 New records of the bobtail squid

anteriorly and rounded posteriorly, slightly longer with smooth horny ring, rounded aperture on than wide (Figs. 2–3, 4A–B); antero-dorsal margin normal sucker, but more ovate on enlarged one. fused with head (Figs. 2, 4A). Large portion of Left arm I of male hectocotylized, stout, shorter the tubular funnel visible, median ventral margin than other arms and curved outwards slightly concave (Figs. 2B, 4B–C). Fins rounded, (Nateewathana, 1997b: figs. 3a, 4a–b). The size almost circular in outline, separate and attached at of arm suckers on different arms varied. Tentacles middle of mantle; anterior margin to posterior round, flattened on oral surface, edges of flat area margin nearly straight. Head broad, dorsoventrally angled, dorsal margin elevated into a low keel flattened and compact, width slightly less than towards club. Club short, curled, and rounded with mantle width; eyes large, oval with almost circular numerous rows of very small suckers. Gladius pupil. Funnel long, tubular, free for most of its absent. Buccal membrane smooth without suckers. length, reaching to gap between ventral arms (Fig. 4B–C). Funnel valve triangular to rounded flaplike. Type locality. Small scale fishing areas in Trang Funnel organs with a roughly triangular dorsal pad Province, in the depth less than 40 m, theAndaman and two oval ventral pads. Anterior end of dorsal Sea coast of Thailand. pad with a low crest projected in midline. Funnel locking cartilage ovate, rounded at both ends with Distribution. Indian Ocean (Andaman Sea and deep groove (Fig. 4C); mantle cartilage straight, Myanmar waters with range extension to the west ridge-like, two times longer than funnel cartilage. of the Andaman Islands) and Gulf of Thailand. Arms long, subequal, rounded in cross section, tapering distally. Web C (between arms II and III) Remarks. The characters of all specimens match developed and D distinct (between arms III and well with those of E. hyllebergi, as described and IV). Web D extends up about one-fourth of arm figured by Nateewathana (1997b). Description of length. Suckers almost equal-sized, subglobular, beak, radula, ink sac, and spermatophores of the with a small, smooth aperture; pedicel thin. In species are not repeated herein. females, arm suckers quadriserial but biserial at The right tentacle in NSMT-Mo 76424 and base and distally. In male, arm suckers globular the left tentacle in ZMUC CEP-51 are missing.

A B

Figure 2: Euprymna hyllebergi Nateewathana, 1997 (PMBC 21339). Neotype, male: (A) dorsal and (B) ventral. 20 Phuket mar. biol. Cent. Res. Bull.

A B

Figure 3: Euprymna hyllebergi Nateewathana, 1997 (PMBC 11731). Paratype, female: (A) dorsal and (B) ventral.

A B

C

Figure 4. Euprymna hyllebergi Nateewathana, 1997 (PMBC 21339). Neotype, male: (A) dorsal view; (B) ventral view; (C) funnel and funnel locking-cartilages. Scales = 2 cm (A–B) and 0.5 cm (C). 21 New records of the bobtail squid R e ma r ks o r i g n a l re c d e as E up r ym na s te noda c tyla ( G r a n t , 1 8 3 ) b y C . F E. R o p e r i n 1 9 79 I d en t if i er A . N a t e w h na A . N a t e w h na A . N a t e w h na A . N a t e w h na A . N a t e w h na A . N a t e w h na A . N a t e w h na A . N a t e w h na A . N a t e w h na A . N a t e w h na A . N a t e w h na A . N a t e w h na A . N a t e w h na A . N a t e w h na A . N a t e w h na A . N a t e w h na A . N a t e w h na A . N a t e w h na A . N a t e w h na A . N a t e w h na A . N a t e w h na A . N a t e w h na A . N a t e w h na A . N a t e w h na A . N a t e w h na A . N a t e w h na A . N a t e w h na A . N a t e w h na A . N a t e w h na A . N a t e w h na A . N a t e w h na A . N a t e w h na A . N a t e w h na A . N a t e w h na P huk et Kra bi T r a ng T r a ng Ra n ong P h a ngnga T r a ng Kra bi P h a ngnga P h a ngnga P h a ngnga T r a ng A nd a m n Sea Kra bi P h a ngnga P h a ngnga T r a ng Kra bi Kra bi Ra n ong Kra bi P huk et P huk et P h a ngnga P h a ngnga P huk et Kra bi S a t ul P h a ngnga Ra n ong A nd a m n Sea P h a ngnga P huk et Kra bi P r ov i n ce L o c a l i ty – K r a b i f s h m rket K a n ta g fi sh l nd i ng K a n ta g fi sh l nd i ng Ka pur T h a L n e , P g B ay R o k I s l a nd K r a b i f s h m rket Chong Pra, P h a ng n g Bay Ra y a Ri n g I s l nd T h a L n e , P g B ay K a n ta g fi sh l nd i ng 8 ° 0 2 ´ N , 9 3 ´E S o u t h w e s f L a n I l nd T a e ng C p e, P h n g Bay P h a ng n g Bay K a n ta g fi sh l nd i ng K r a b i f s h m rket K r a b i f s h m rket – K r a b i f s h m rket P h u k e t fi s m a r et T u k a e C pe R a y i n g I s l d , P h B ay M a k I s l n d , P h g B ay P h u k e t fi s m a r et S o u t h w e s f L a n I l nd T a m l ng f i s h nd – J a m I s l nd 7 ° 4 8 ´ N , 9 3 0 ´E Na k ae I s l a nd S a p u m B ay D a m H o k I s l nd re ma ining i n the Re f ere ce C oll ectio o Ph u k et M ar e Bi ol gi cal nt r. 2 1 4 6 1 1 3 2 1 3 5 7 6 17 6 1 1 1 7 1 1 1 2 7 3 1 1 1 1 F e m a le 3 2 5 3 1 1 2 3 6 4 5 1 2 3 6 1 1 2 Ma le Eupr y m na h yllebe r gi 5 1 6 11 1 1 6 3 2 3 5 9 9 3 2 15 12 1 1 5 12 2 1 1 4 3 13 4 1 1 1 3 1 N o . of s p e c. T y pe H O L T Y P E s p e c i m n l o st P AR A TYPE P A R T Y E s p e c i m n l o st P A R T Y E s p e c i m n l o st Det a ils o f s p eci m e n 1 6 07 1 727 1 728 1 729 1 730 1 731 1 732 1 733 1 734 1 735 1 736 1 737 1 738 1 739 1 740 1 741 1 742 1 744 1 745 1 746 1 747 1 748 1 749 1 750 1 751 1 752 1 753 1 755 1 756 1 757 1 758 1 759 1 760 1 761 P MB C N o. 1 2 3 4 5 6 7 8 9 10 11 12 13 14 15 16 17 18 19 20 21 22 23 24 25 26 27 28 29 30 31 32 33 34 T ab l e 2. N o. 22 Phuket mar. biol. Cent. Res. Bull. R e ma r ks s p e c i m n w a l t ed f ro m P M B C 1 7 42 s p e c i m n f o u d t he b o t l e f P M BC 1 7 3 , b u t n ot i d e n ti c a l t o ri g al p a rat y pe s p e c i m n f o u d t he b o t l e f P M BC 1 7 3 9 , b u t n ot i d e n ti c a l t o ri g al p ara t yp e spe c i m ns I d en t if i er A . N a t e w h na A . N a t e w h na A . N a t e w h na A . N a t e w h na A . N a t e w h na A . N a t e w h na A . N a t e w h na A . N a t e w h na A . N a t e w h na A . N a t e w h na A . N a t e w h na A . N a t e w h na A . N a t e w h na A . N a t e w h na A . N a t e w h na A . N a t e w h na A . N a t e w h na A . N a t e w h na A . N a t e w h na A . N a t e w h na T . K u b o d e ra T . K u b o d e ra A . N a t e w h na A . N a t e w h na A . N a t e w h na T r a ng P h a ngnga Kra bi – Kra bi A nd a m n Sea A nd a m n Sea P huk et Kra bi Ra n ong T r a ng M y a n m r wa t ers P a t ni T r a ng S a m u t P r k an In d ia n Oc e an A nd a m n Sea T r a ng T r a ng A nd a m n Sea A nd a m n Sea A nd a m n Sea T r a ng – – P r ov i n ce L o c a l i ty T r a n g f i s h m rket o f Y a i I s l nd E a s t o f L n I l nd – – 6 ° 5 8 ´ N , 9 4 ´E 6 ° 5 7 ´ N , 9 8 4 ´E Ra n g Y a i Is l nd – – K a n ta g fi sh l nd i ng 1 3 ° 2 7 ´ N , 9 0 ´E P a t n i f s h l d ng K a n ta g fi sh l nd i ng Sa m u t Pra k a n f is h la nd i ng 7 ° 1 ´ N , 9 0 ´E 7 ° 0 4 ´ N , 9 8 5 3 ´E K a n ta g fi sh l nd i ng K a n ta g fi sh l nd i ng 9 ° 3 4 ´ N , 7 ´E 7 ° 5 1 ´ N , 9 8 3 ´E 9 ° 0 ´ N , 7 5 4 ´E K a n ta g fi sh l nd i ng – – 1 2 1 1 1 2 1 1 1 1 1 13 1 1 1 1 6 9 4 1 1 2 F e m a le 1 1 2 5 2 10 17 1 6 1 3 3 Ma le 1 2 2 1 1 1 2 1 3 1 1 1 18 1 1 1 3 16 26 1 10 1 1 4 5 N o . of s p e c. T y pe NEOTYPE t h e n u m b r o f s p c i P M B C a g d , l A x 1 N w (1 9 7 b), me (c on t inued) 1 762 1 763 1 764 1 765 1 766 1 767 1 768 1 770 1 772 1 773 1 774 1 775 1 776 1 777 1 780 1 781 1 782 1 785 1 787 1 4 420 1 9 903 1 9 904 2 1 339 2 3 937 2 5 004 P MB C N o. 35 36 37 38 39 40 41 42 43 44 45 46 47 48 49 50 51 52 53 54 55 56 57 58 59 T ab l e 2. N o. R e m a r k s : P M B C a c e s i o n u m b r w t l d h p x. 23 New records of the bobtail squid 51 39.94 57.74 28.42 169. 11 f r o m the A nd a ma n S ea,

28.65 81.85 76.18 48.94 57.74 134 . 90 1 3 5.50 37.63 34.24 58.71 57.35 64.96 99.34 84.87 3.66 2.43 5.34 3.20 4.54 2.81 41.64 60.94 0.00 26.35 25.97 14.52 13.62 124.15 102.39 25 Ma l e Female Z M U C EP 2 7 . 21 8 3 . 83 5 2 . 11 5 3 . 55 6 8 . 83 8 . 06 98.46 101.22 88.53 2.94 4.01 2.53 2.94 4.41 4.59 4 2 . 08 5 8 . 01 3 0 . 10 1 5 . 40 1 10.66 Eupr y m na h yllebe r gi S e lec t ed sp cim ns F e m a le 2 6 . 79 8 . 28 6 4 . 05 6 7 . 19 9 1 . 75 99.22 93.77 101.14 108.34 91.87 9 6 . 90 2.24 2.61 2.43 2.43 6 2 . 64 2 4 . 15 1 5 . 57 U S NM 1 5 0 4 74 1 1 1 M a le 2 9 . 02 9 0 . 45 54.51 50.09 35.70 39.94 32.97 5 3 . 62 6 4 . 13 9 1 . 94 3.96 3.79 5.34 5.00 6 2 . 93 2 7 . 08 1 3 . 20 290.18 246.62 285.34 229.6 0 124.91 105.10 1 1 N S M T - Mo 7 6 425 F e m a le 35.22 7 4 . 82 5 1 . 02 3 . 19 5 0 . 03 5 9 . 71 5 9 . 00 74.76 69.96 104.31 67.18 2.13 2.30 2.98 2.73 84.44 31.63 31.74 2 4 . 11 1 2 . 12 1 2 = l e f t ten acle du to the ri ght o ne lost. Th v a u es in b old tex are ou th r ng N S M T - Mo 7 6 424 M a le 33.63 7 . 19 4 8 . 41 2 8 . 49 5 2 . 81 6 1 . 11 9 7 . 71 4.16 4.46 4.46 4 6 . 65 5 9 . 99 1 9 . 80 1 3 . 56 98.33 2 234.67 1 P a rat y pe P M BC 1 7 31 F e m a le 2 . 27 9 1 . 02 5 8 . 96 4 6 . 74 4 6 . 43 7 3 . 15 6 9 . 33 86.30 114.93 77.19 73.37 110.70 2.92 2.97 3.37 2.92 2.25 2 5 . 82 1 5 . 67 34.98 1 19.98 1 P M BC 2 1 339 M a le 3 0 . 98 8 4 . 83 4 . 09 5 4 . 71 6 3 . 52 3.23 4.03 5.23 6.17 42.32 6 5 . 73 2 . 60 1 2 . 33 Ne o ty pe 26.44 327.73 129.50 152.09 119.92 1 3 5.15 146.7 6 159.69 132.82 106.75 143.54 120.27 116.17 = t ip o f he ri ght ar m bro ke n; 1 M ea s u re m e nt ( mm ) an d indice % o f n otyp , p arat y pe a nd lected eci I I I I I I I I I II I II IV IV III II I R e gis t rat i on N o. I n d e x\ Sex ML ( m m) M WI FLI FWI FBI HLI H WI AL AL AL AL AS AS AS AS TtLI CI LI H c LI E DI Ln DI T ab l e 3. Th a ila n d. Rem a r k s: m easure e nt s a nd ind i ces re p o rte d by Nateewa t h n ( 1 9 97b ). 24 Phuket mar. biol. Cent. Res. Bull.

Table 4. Mean (X) of indices (%) and standard deviation of type materials and selected specimens of Euprymna hyllebergi from the Andaman Sea, Thailand. Male Female Index n Mean SD Range n Mean SD Range

ML (mm) 5 29.90 2.48 27.21–33.63 4 28.18 5.37 22.27–35.22 MWI 5 83.63 4.81 77.19–90.45 4 82.57 8.27 74.82–91.02 FLI 5 49.61 3.96 44.09–54.51 4 54.45 4.54 50.09–58.96 FWI 5 132.78 9.70 119.92–146.76 4 145.61 12.27 135.15–159.69 FBI 5 32.24 4.72 26.44–37.63 4 38.53 6.23 33.19–46.74 HLI 5 54.68 2.35 52.81–58.71 4 54.47 7.84 46.43–64.05 HWI 5 64.51 2.81 61.11–68.83 4 64.45 7.09 57.74–73.15

ALII 5 96.76 7.18 88.06–106.75 4 76.24 14.83 59.00–91.75

ALIII 5 123.64 12.67 110.66–143.54 4 90.67 12.68 74.76–102.39

ALIIII 5 106.48 8.56 98.33–120.27 4 83.20 11.52 69.96–93.77

ALIVI 5 106.33 7.17 98.46–116.17 4 81.50 13.64 67.18–96.90

ASII 5 3.42 0.53 2.94–4.01 4 2.46 0.35 2.13–2.92

ASIII 5 3.72 0.48 2.94–4.16 4 2.68 0.48 2.30–3.37

ASIIII 5 4.96 0.48 4.41–5.34 4 2.88 0.33 2.43–3.20

ASIVI 5 4.95 0.71 4.46–6.17 4 2.55 0.26 2.25–2.81 TtLI 5 273.50 41.21 229.60–327.73 4 155.04 70.23 84.44–246.62 CILI 5 40.89 5.50 31.74–46.65 4 42.30 13.98 31.63–62.64 HcLI 5 61.52 2.95 58.01–65.73 4 EDI 5 25.19 4.02 19.98–30.10 4 25.01 1.02 24.11–25.97 LnDI 5 13.80 1.19 12.33–15.40 4 14.24 1.70 12.12–15.67

Table 5. t-test of the mean of all measurements and indices by Nateewathana (1997b) and this study. Male Nateewathana (1997b) present study

Index n Mean SD n Mean SD T-test

ML (mm) 20 27.20 4.20 5 29.90 2.48 No diff MWI 20 81.20 11.30 5 83.63 4.81 No diff FLI 20 49.10 7.10 5 49.61 3.96 No diff FWI 20 131.10 13.80 5 132.78 9.70 No diff FBI 20 31.50 4.00 5 32.24 4.72 No diff HLI 20 52.80 8.50 5 54.68 2.35 No diff HWI 20 65.20 7.10 5 64.51 2.81 No diff

ALII 20 103.70 20.00 5 96.76 7.18 No diff

ALIII 20 141.00 15.70 5 123.64 12.67 Sig diff

ALIIII 20 118.20 14.10 5 106.48 8.56 No diff

ALIVI 20 118.60 18.00 5 106.33 7.17 No diff

ASII 20 3.50 0.50 5 3.42 0.53 No diff

ASIII 20 4.40 0.60 5 3.72 0.48 Sig diff

ASIIII 20 5.10 0.70 5 4.96 0.48 No diff

ASIVI 20 5.30 0.60 5 4.95 0.71 No diff TtLI 20 282.80 47.10 5 273.50 41.21 No diff CILI 20 48.40 7.10 5 40.89 5.50 Sig diff HcLI 20 63.40 8.30 5 61.52 2.95 No diff EDI 20 23.30 5.60 5 25.19 4.02 No diff LnDI 20 14.40 2.80 5 13.80 1.19 No diff 25 New records of the bobtail squid

Table 5. (continued)

Female Nateewathana (1997b) present study

Index n Mean SD n Mean SD T-test

ML (mm) 20 24.90 5.50 4 28.18 5.37 No diff MWI 20 80.70 8.90 4 82.57 8.27 No diff FLI 20 52.50 7.80 4 54.45 4.54 No diff FWI 20 142.40 12.80 4 145.61 12.27 No diff FBI 20 35.80 5.00 4 38.53 6.23 No diff HLI 20 55.30 6.20 4 54.47 7.84 No diff HWI 20 67.70 7.90 4 64.45 7.09 No diff

ALII 20 85.40 11.40 4 76.24 14.83 No diff

ALIII 20 110.60 12.30 4 90.67 12.68 Sig diff

ALIIII 20 93.20 8.80 4 83.20 11.52 No diff

ALIVI 20 89.70 9.20 4 81.50 13.64 No diff

ASII 20 2.70 0.40 4 2.46 0.35 No diff

ASIII 20 2.80 0.50 4 2.68 0.48 No diff

ASIIII 20 3.00 0.40 4 2.88 0.33 No diff

ASIVI 20 2.80 0.40 4 2.55 0.26 No diff TtLI 19 266.60 60.70 4 155.04 70.23 Sig diff CILI 19 53.30 10.90 4 42.30 13.98 No diff EDI 20 23.50 3.80 4 25.01 1.02 No diff LnDI 20 14.90 2.10 4 14.24 1.70 No diff

DISCUSSION the t-test of the mean of all measurements and indices by Nateewathana (1997b) and in this study

The Euprymna hyllebergi arm length were considered significant for ALII in both male formula is II > III > IV > I, but occasionally II > and female, for ASIII and ClLI in male only, and III = IV > I (Nateewathana, 1997b). The arm length for TtLI in female (Table 5). formulas of the present material of males and These differences may occur because the females (Table 4, II > III > IV > I), matches those suckers on arms easily fall off and are lost in of Nateewathana (1997b) as well as those in the preserved material and the tip of the first right arm neotype and paratype (Table 3). Differences are, in male specimen was broken. Therefore some however, found in some of the selected specimens; errors occurred in estimation of the length of the four specimens show II > III > IV > I but three first right arm, which was measured from the first other specimens show II > IV > III > I (Table 3). basal (proxima-most) sucker to tip of arm as a These differences occur because the tips of the percentage of mantle length. right arms were broken. Furthermore, according to C.C. Lu (pers. All measurements and indices of the comm., in Nateewathana, 1997b), the length of neotype are in the ranges reported by the tentacles in cephalopods may not be a usable Nateewathana (1997b). Some measurements and taxonomic character since it varies greatly indices in the paratype and selected specimens are, depending on how the specimen is fixed and however, out of these ranges, i.e., Mantle Length preserved. Live fixed animals have short tentacles (ML), Arm Length Index (ALI), Arm Sucker Index because of contraction and, on the other hand, (ASI), Tentacle Length Index (TtLI), and Club specimens thawed from frozen condition have long Length Index (ClLI) (Tables 3–4). Furthermore, tentacles. In order for this character to be of some 26 Phuket mar. biol. Cent. Res. Bull.

use, specimens should be fixed under identical ACKNOWLEDGEMENTS conditions. Specimens from the same batch may vary greatly (C.C. Lu, pers. comm., in We express our appreciation to Dr. Nateewathana, 1977b). Kongkiat Kittiwattanawong, Phuket Marine The mantle length, the first arm length Biological Center (PMBC) for his kind assistance index, the tentacle length index, and club length in t-test Analysis, and Dr. Janek von Byern, index are the main characters/indices used for University of Vienna, Austria, for providing valuable distinguishing the species from other members of comments on the draft of this manuscript. Sincere the genus. Although the tentacle length is thanks are given to Ms. Kanjana Bussabong, the particularly important in recognizing this species, Reference Collection, Phuket Marine Biological it is not necessarily an autopomorphy for the Center, and Ms. Thitima Boonkaew, training species. It is the combination of the arrangement student, PSU, for their kind assistance. Our sincere of suckers on the hectocotylized arm, the size and thanks also go to Mr. Rueangrit Promdam, (former distribution of enlarged suckers on arms II–IV, staff of Reference Collection), and Mr. Patirat and in the length of the tentacles that distinguishes Singdum, the PMBC artist, for photographing and Euprymna hyllebergi. drawings.

REFERENCES

Adam, W. 1986. Contribution à la connaissance du genre Euprymna Steenstrup, 1887 (Mollusca: Cephalopoda). Bulletin de l’Institut Royal des Science Naturelles de Belgique, Biologie. 56: 131–136. Aungtonya, C., K. Bussabong, K. Wongkamhaeng, I. Eedkerd, and S. Ubonsuwan. 2006. Type specimens in the Reference Collection of Phuket Marine Biological Center. Reference Collection and Aquarium Contributions no. 4: 45 p. (in Thai). Aungtonya, C., S. Sungjeen, W. Tripanichkul, K. Bussabong, and P. Kongthaworn. 2007. Recent records of the Class Cephalopoda (Phylum Mollusca) in the Reference Collection of Phuket Marine Biological Center. Reference Collection and Aquarium Contributions no. 5: 69 p. (in Thai). Berry, S. S. 1913. Some new Hawaiian cephalopods. Proc. U. S. natn. Mus. 45: 563–566. Chotiyaputta, C. 1993. Cephalopod resources of Thailand. Pp. 71–80. In: Okutani,T., R.K. O’Dor & T. Kubodera (eds.). Recent Advances in Fisheries Biology. Tokai University Press, Tokyo. Chotiyaputta, C., T. Okutani, and S. Chaitiamvong. 1992. Systematic study of cephalopods in Thailand. A Research Report submitted to the National Research Council of Thailand under JSPS-NRCT Cooperation Project. 100 pp. (In Thai with English abstract). Grant, R. E. 1883. On a new species of Sepiola (Sepiola stenodactyla) from the Mauritius, presented by C. Telfair, Esq. Proceedings of the Zoological Society of London. 1: 42–43. Nateewathana, A. 1997a. Systematics of the Cephalopoda of the Andaman Sea, Thailand. Ph.D. Thesis. Institute of Biological Sciences, Faculty of Natural Sciences, University of Aarhus. 343 pp. Nateewathana, A. 1997b. The Sepiolidae (Cephalopoda) of the Andaman Sea, Thailand, with description of Euprymna hyllebergi sp. nov. Phuket Marine Biological Center Special Publication. 17: 465–481. Nateewathana, A., J. Nabhitabhata, and P. Nilaphat. 2001. A new record of bobtail squid, Euprymna hyllebergi Nateewathana, 1997 in the Gulf of Thailand. Phuket Marine Biological Center Special Publication. 25(2): 501–506. Nesis, K.N. 1987. Cephalopods of the world. T.F.H. Publications, Inc. Ltd., New Jersey. 351 pp. 27 New records of the bobtail squid

Norman, M.D. & C.C. Lu. 1997. Redescription of the southern dumpling squid Euprymna tasmanica and a revision of the genus Euprymna (Cephalopoda: Sepiolidae). J. Mar. Biol. Ass. U.K. 77: 1109–1137. Pfeffer, G.J. 1884. Die Cephalopoden des Hamburger Naturhistorischen Museums. Abhandlungen des Naturwissenschaftlichen Vereins in Hamburg. 8(2): 63–90. Sasaki, M. 1913. Decapod cephalopods found in Japan: Sepiolidae. Zool. Mag., Tokyo. 25: 247–252. Steenstrup, J.J. 1887. Notae teuthologicae, no. 6. Species generic Sepiolae Maris Mediterranei. Oversi. K. Danske Vidensk Selsk Forh. Copenhagen. 1887: 47–66. Supongpan, M. 1995. Cephalopod resources in the Gulf of Thailand. pp. 191–203. In: Nabhitabhata, J. (ed). Biology and Culture of Cephalopods. Rayong Coastal Aquaculture Station, Contribution No. 18. (In Thai). Verrill, A.E. 1881. The cephalopods of the north-eastern coast of America: part 2. The smaller cephalopods, including the squids and the octopi, with other allied forms. Trans. Conn. Acad. Arts. Sci. 5: 259–446. Voss, G.L. 1963. Cephalopods of the Philippines. Smithsonian Institution Bulletin. 234: 1–180.

Manuscript received: 17 August 2009 Accepted: 12 June 2011 28 Phuket mar. biol. Cent. Res. Bull. Phuket mar. biol. Cent. Res. Bull. 70: 29–77 (2011)

THE FISHES OF SOUTHWESTERN THAILAND, THE ANDAMAN SEA – A REVIEW OF RESEARCH AND A PROVISIONAL CHECKLIST OF SPECIES

Ukkrit Satapoomin

Phuket Marine Biological Center, P.O. Box 60, Phuket 83000, Thailand Email: [email protected]

ABSTRACT:Abrief review of recent ichthyological research and compiled results on the records of fishes from the Andaman Sea coast of Thailand are presented. A total of 1,746 species in 198 families of fishes are currently known from the area. The 10 most speciose families are the Gobiidae (227 species), Labridae (78), Pomacentridae (71), Serranidae (61), Apogonidae (60), Blenniidae (52), Carangidae (52), Scorpaenidae (49), Lutjanidae (39) and Chaetodontidae (37), and these together account for 42% of the total fish fauna. The ichthyofauna is dominated by reef-associated fishes (983 species) and pelagic/benthic fishes inhabiting offshore habitats (971 species). A zoogeographic analysis reveals a peculiarity of the fauna of this marginal-sea region, viz., being one of the areas of sympatry of Indian and Pacific Ocean fishes, as well as harbouring regional endemics. The PMBC Reference Collection currently contains examples of only about 63% of the fishes known from the area, indicating a need for further local sampling campaigns and/or international collaborative exploratory research programs in the Andaman Sea to build up the Reference Collection.

Keywords: Thailand, Andaman Sea, fishes, checklist, diversity.

INTRODUCTION northwesterly through-flow in the Strait of Malacca. It is thus an area of distributional The Andaman Sea is an approximately confluence of the biotas of both the Indian Ocean 1,200 km long x 650 km wide body of water to and the Pacific Ocean (Satapoomin, 2002a, 2007; the southeast of the Bay of Bengal. It is bounded Kimura and Satapoomin, 2009). Being a marginal by the Gulf of Mataban in the north, the Malay sea separated from the main Indian Ocean by the Peninsula in the east, the Andaman and Nicobar Andaman-Nicobar island arcs, it has been subjected Islands in the west and the northern tip of Sumatra to a profound effect of sea-level lowering, especially in the south. At its southeastern corner, the during major glaciations when it became a Andaman Sea narrows to form the Strait of separated, enclosed basin. Such isolation processes Malacca, which separates the Malay Peninsula from are believed to have promoted speciation and/or the island of Sumatra. Along an approximate 1,000 endemism of the fish fauna in the Andaman Sea km length of the Andaman Sea coast of Thailand, (e.g., McManus, 1985; Randall, 1998; Randall and a variety of marine and coastal habitats are present, Satapoomin, 1999). The area is thus characterized viz., mangroves, mud-flats, seagrass beds, rocky by its high diversity and the unique faunal shores, intertidal sand-flats, coral reefs, open composition of the fishes. However, knowledge pelagic zones, and complex seabed systems on the and information on the overall diversity of marine continental shelf. The integrity of different peculiar fishes in the area is still limited, because very few environmental settings and/or characteristics attempts had been made to compile a list of all among those habitat types provides suitable refuges known fish species from the area. for a vast variety of aquatic biota, including fishes. The purposes of the present paper are to Although, biogeographically, it belongs to the provide a brief review of recent ichthyological eastern Indian Ocean, it is strongly influenced by research, and to establish a comprehensive the western Pacific Ocean biota via the checklist of all known marine and brackish-water 30 Phuket mar. biol. Cent. Res. Bull.

fishes from the Andaman Sea coast of Thailand. information on habitats and zoogeographic The checklist was compiled from various sources distribution of each species. The availability of of information, including published literature, voucher specimens lodged at the Reference unpublished data made available under several Collection of the PMBC was also emphasized in sampling programs/projects, and the fish collection order to draw the attention of ichthyologists who data repository of the Phuket Marine Biological may wish to examine these specimens. The Center (PMBC). It should be noted that the overall designation of habitat type(s) for each species was list of species presented herein is only provisional based largely on either quoted information from as a number of species records and identifications the compiled literature or from the author’s in previously published literature (particularly those unpublished survey results and observations. Four of inventory, non-taxonomic works), cannot be principal habitat categories adopted herein included verified. Nonetheless, this provisional checklist will mangrove (Mg), coral reef (Cr), seagrass bed (Sg), serve as a significant baseline for further and other offshore habitats (OShr). These establishment of a conventional, annotated categories can be considered as good checklist or taxonomic review of the Andaman Sea representations of the major marine and coastal fishes. features along the southwestern Thailand coastline. Each of these habitat types, however, was more METHODS or less loosely defined in order to accommodate a range of associated minor biotopes of each habitat. The compilation of the species list was The offshore category, in particular, was the most based largely on accumulating information from inclusive term as it encompassed a range of the earlier literature, including inventory works and/ benthic- and pelagic-related habitats. This or faunistic surveys, field guides, and taxonomic designation includes both near-shore and far- works. Some unpublished data, including recent offshore fishes considered as either pelagic (i.e., unpublished survey results made by the author, coastal pelagic, offshore pelagic, epipelagic, and collection data of the Royal Ontario Museum’s mesopelagic) or demersal (i.e., coastal demersal, project on gobioid fishes of southwestern Thailand benthopelagic, and deep demersal) species. The in 1993 (under auspices of Dr. Richard mangrove habitat was considered to include certain Winterbottom), and the PMBC fish collection data inter-connected zones like mud-flats and the upper repository were also incorporated into the current reaches of the estuarine areas. Coral reef was checklist. Selected taxonomic papers pertaining to defined to include several nearby and/or associated records of fishes from the Andaman Sea coast of areas, namely intertidal rocky shore/rock platform, Thailand used to compile the current checklist are intertidal sand-flat, and subtidal bare-seafloor in listed in Appendix B. front of the reef (so called as forereef). The systematic arrangement of families Most of the well defined zoogeographic follows Nelson (2006). Species in each family are distribution categories (see, e.g., Myers, 1991) arranged alphabetically. The scientific names of were generally adopted or redefined with additional fishes used in the cited literature have been updated explanation, while some new categories were added where possible, following recent usage, and hence with their definitions. The definitions for those junior synonyms were omitted. Since the distribution categories and their acronyms are as compilation was also based, in part, on literature follows: Indo-Pacific (IP), widespread across the that provided only a list of names, the author has Indian Ocean and western portion of the Pacific only been able to check or correct with an eastern limit east of non-marginal areas of misidentifications to a limited extent. In this regard, the Pacific Plate; Indo-west Pacific (IWP), a subset only correct names were retained, synonyms were of the Indo-Pacific with northeastern limit on the allocated according to the latest usage, while Pacific Plate, marginally in the Carolines but not doubtful species records were excluded. reaching the Marshalls; West Pacific (WP), limited In addition to scientific names and to the area bound by the Andaman Island and classification, the checklist provides basic Cocos-Keeling Island in the west to the margin of 31 The fishes of southwestern Thailand

the Pacific Plate in the east, north to southern westerner was a reef-associated goby, Pipidonia Japan, south to western and eastern Australia; West quinquecincta Smith, 1931 [valid as Gobiopsis central Pacific (WCP), west Pacific and Pacific quinquecincta (Smith)]. During his term of service Plate distributions and may include the East Pacific; (1923–1935) as the Adviser in Fisheries to the Indo-Australian Archipelago (IA), a subset of the Department of Fisheries of Thailand (formerly West Pacific, north to southern Taiwan, east to Siamese Bureau of Fisheries), Smith published a the Solomons, south to the Great Barrier Reef; number of articles pertaining to freshwater fishes Indo-Malayan Archipelago (IM), a subset of Indo- and fishes of the Gulf of Thailand. Later, in 1945, Australian which is limited to the areas within he published a monograph entitled “The Fresh- bounds of Malay Peninsula, Sumatra, Java, New water Fishes of Siam, or Thailand” (Smith, 1954). Guinea and Philippines; Indian Ocean (IO), Suvatti (1950) was the first Thai ichthyologist to widespread throughout the Indian Ocean; Northern publish a list of Thai fishes. Several subsequent Indian Ocean (NIO), the extended range of eastern studies on marine fishes in Thai waters by the Indian Ocean to encompass the northwestern Thais were confined to groups of particular limits which include the west coast of India, interest, for instance Pleuronectiformes (by Pakistan, Oman and Persian Gulf; Eastern Indian Punpoka, 1964), Lutjanidae (Banasopit, 1968), Ocean (EIO), the eastern section of the Indian Serranidae (Banasopit, 1969; Monkolprasit, 1983), Ocean westwards to Laccadive, Maldive and Chaetodontidae (Banasopit, 1971), Leiognathidae Chagos archipelagoes; the eastern extremity of the (Monkolprasit, 1973), Elasmobranchii Indian Ocean (EEIO), a narrow stripe or transition (Monkolprasit, 1984), and Labridae (Monkolprasit, area along the joint margin of Indian-Australian 1989). The foremost comprehensive compiled and Asian lithospheric plates, including the checklist of the fishes of Thailand, which included Andaman Sea, Sumatra, Java, Bali, Flores, Cocos- both freshwater and marine fishes, was published Keeling/Christmas Islands and northwestern by Monkolprasit et al. in 1997. It listed 171 fish Australian Shelf; Andaman Sea province (AS), the families with a total of 1,741 species, about 30% area assigned to accommodate species having (516 species) of which can be considered as restricted distribution only to the Andaman Sea or freshwater fishes. The records of marine fishes ranging southwards only to western Sumatra; in the checklist were compiled from the literature Circumtropical (CIR), tropical seas worldwide; and published before 1990, except for two papers (one Cosmopolitan (COS), tropical and temperate seas on mangrove fishes by Monkolprasit (1994) and worldwide. The boundary of some distribution one on ammodytid fishes by Ida et al. (1994)). categories is presented (Fig. 1) for clarification of That checklist recorded 710 species representing the definitions of those areas. 129 families of fishes from the Andaman Sea. A few collections of reef fishes from the RESULTSAND DISCUSSION Similan Islands were made during the R/V Te Vega Cruise 2, eastern Indian Ocean cruise (from 1. Brief review of ichthyological research Singapore to Colombo, Ceylon), in late 1963 (http:/ The fish fauna of the Andaman Sea coast /tevega.stanford.edu/cruises.htm). These of Thailand was poorly documented until relatively collections were deposited at the U.S. National recent times. There were several well-known Museum of Natural History, and some were western ichthyologists/naturalists (e.g., Cuvier, designated as types of new fish species described Valenciennes, Bleeker, Günther, Fowler, Regan, subsequently (i.e., Melichthys indicus Randall & Boulenger, etc.) who reported, listed or described Klausewitz, 1973, Gymnothorax pseudoherrei Thai fishes between 1800 and the 1950s, but their Böhlke, 2000, and Helcogramma lacuna Williams works were restricted only to either the Gulf of & Howe, 2003). In February 1979, Randall Thailand (marine) or inland (freshwater) fishes (Bishop Museum, Hawaii) and Allen (Western (see Monkolprasit et al. (1997) for detailed Australian Museum) made a few collections and historical review). The first marine fish of the photographed reef fishes around Phuket and the Andaman Sea coast of Thailand described by a Similan Islands (see Allen et al., 2005). Although 32 Phuket mar. biol. Cent. Res. Bull.

Figure 1. Approximate range of some zoogeographic distribution categories. A: Indo-Pacific (IP), Indo-west Pacific (IWP), West central Pacific (WCP), and West Pacific (WP); B: Indo-Australian archipelago (IA) and Indo-Malayan Archipelago (IM); C: Northern Indian Ocean (NIO), Eastern Indian Ocean (EIO), and Eastern extremity of Indian Ocean (EEIO). no comprehensive list of fishes resulted from that Ctenochaetus truncatus Randall & Clements, 2001, effort, the collected materials had been used in Scorpaenopsis possi Randall & Eschmeyer, 2001, describing several new species (i.e., Leptojulis Helcogramma lacuna Williams & Howe, 2003, and chrysotaenia Randall & Ferraris, 1981, Chromis Opistognathus cyanospilotus Smith-Vaniz, 2009). flavipectoralis Randall, 1988, Pomacentrus adelus The most important collections of reef fishes for Allen, 1991, P. azuremaculatus Allen, 1991, P. the area were made in November 1993 under the polyspinus Allen, 1991, P. similis Allen, 1991, collaborative project between the Royal Ontario Neopomacentrus sororius Randall & Allen, 2005, Museum (ROM), Canada and the Phuket Marine Plesiops auritus Mooi, 1995, Plectorhinchus Biological Center (PMBC). From well over 35 macrospilus Satapoomin & Randall, 2000, sampling stations using the ichthyocide rotenone 33 The fishes of southwestern Thailand

around Phuket and adjacent islands, nearly eighteen surrounding area of Libong Island, Trang Province. thousand specimens representing about 570 species The most recent list of the fish fauna in the seagrass in 75 families of fishes were collected and beds of the Andaman Sea coast of Thailand, with documented. It was also the first time that 230 species in 62 families, was published by significant information on small cryptobenthic reef Satapoomin (2006). fishes for the area was obtained. The collected The first publication on demersal and/or materials have been used in the descriptions of 13 pelagic fishes of the Andaman Sea coast of new species to date (including 7 species in Gobiidae Thailand was published by Lohakarn (1968). This and 1 in each of the following families:Apogonidae, included taxonomic accounts of 109 species in 57 Serranidae, Pomacentridae, Opistognathidae, families. Fishes collected by bottom trawls during Tripterygiidae, and Soleidae), with several other the 5th Thai-Danish Expedition to the Andaman descriptions underway. Satapoomin (1993) Sea in 1966 (the first Thai-Danish marine established a first comprehensive checklist of coral expedition) were reported by Boonyanate and reef fishes of the west coast of Thailand by Hylleberg (1993) with a total records of 80 species compiling information from both available literature in 41 families. Other early studies on demersal (dated back to 1950) and the author’s survey fishes in the area were mostly focused on fisheries results carried out during 1990–1992. The list aspects and, in most cases, faunal composition revealed a total of 597 species belonging to 66 was reported mainly as higher taxonomic ranks families of reef fishes in the area. Allen et al. (2005) rather than individual species (e.g., Pokapunt and provided an extensive list, consisting of 764 species Sinoda, 1978; Boonragsa, 1988; Chantawong, in 70 families, of coral reef fishes being recorded 1996). Some studies on experimental operations and/or known for the area. The most recent list of of certain types of fishing gears reported not more coral reef fishes, with 888 species in 85 families, than 50 fish species each (e.g., Wongratana, was published by Satapoomin (2007). 1982b, 1983; Ruengpan, 1988). A more There are a limited number of studies on comprehensive list of demersal fishes collected by ichthyofauna of either mangroves or seagrass beds otterboard trawl nets and deep sea shrimp traps in the Andaman Sea coast of Thailand, most of along the Andaman Sea coast of Thailand at depths which were confined to particular sampling sites of about 30–420 m was published by Wongratana (e.g., Boonruang et al., 1994; Monkolprasit, 1994; (1982a), with 256 species in 91 families. Also, Janekitkarn and Monkolprasit, 1994). Satapoomin Pokapunt and Tantivala (1987) reported 182 and Poovachiranon (1997) provided the first species of demersal fishes (in 66 families) in the comprehensive list of the fish fauna from several area from surveys using otterboard trawls at water major mangrove and seagrass bed sites along the depths less than 90 m. Deep-sea fishes of the coast of southwestern Thailand. That study Andaman Sea were poorly known. The results of revealed a total of 280 species representing 75 surveys during 1975 and 1976 on fishes inhabiting families of fishes, which included 232 species (69 the continental shelf within the depth range of 200– families) and 149 species (51 families) found in 550 m was reported by Pokapunt et al. (1983), mangroves and seagrass beds, respectively. In with taxonomic accounts of 31 species in 20 2002, a study by Fish Team of the Trang Project families. Under the Scientific Cooperation (2002) provided a list, together with illustrations, Programme between Denmark and Thailand during of 134 species in 49 families of fishes inhabiting a 1996–2000, the Biodiversity of the Andaman Sea mangrove estuary at Sikao, southwestern Thailand. Shelf (BIOSHELF) Project was undertaken by the Under the JSPS (Japan Society for the Promotion staff of the PMBC together with their Danish of Science) Multilateral Project on Coastal Marine counterparts. The benthic fauna, including fishes, Science, Matsuura and Kimura (2005) published was sampled at depths of 40–900 m. Preliminary a guide book with taxonomic accounts and results revealed a total of 55 families in 20 orders illustrations of 128 species in 46 families of fishes of fishes, most of which were deep-water fishes inhabiting mangroves, seagrass beds and that have not yet been identified (Aungtonya et 34 Phuket mar. biol. Cent. Res. Bull.

al., 2000). Under the JSPS project, a Myxini, Chondrichthyes, and Actinopterygii, comprehensive guide book entitled “Fishes of respectively. Andaman Sea, West Coast of Southern Thailand” covered 778 species in 106 families of fishes 2. Faunal composition and zoogeography (Kimura et al., 2009). The present checklist contains 1,746 The Reference Collection of the PMBC is species, representing 696 genera, 198 families, 40 one of the major sources of information on marine orders, and 3 classes, of marine and coastal fishes biodiversity of theAndaman Sea coast of Thailand. from theAndaman Sea coast of Thailand (Appendix The collection of fishes, in particular, was largely A). The figure is relatively high as compared to established as a result of the FAO/DANIDA those reported from several other localities in the Seminar on Fish Taxonomy in Southeast Asia held Indian Ocean. For example, 533 species were at the PMBC in 1972. Apart from the Andaman reported from the Cocos (Keeling) Islands (Allen Sea fishes, a number of specimens obtained from and Smith-Vaniz, 1994), 603 from the Laccadive the Gulf of Thailand, as well as from neighboring Archipelago (Jones and Kumaran, 1980), 622 from countries (i.e., India, Malaysia, Singapore, Christmas Island (Hobbs et al., 2010), 784 from Indonesia, Philippines, and Hong Kong) were the Chagos Archipelago (Winterbottom and brought in during the seminar and subsequently Anderson, 1999), 880 from Seychelles and deposited in the PMBC Reference Collection. Some adjacent areas (Smith and Smith, 1969), 885 from deposited material of fishes from Australia was the Reunion Island (Letourneur et al., 2004), 930 donated by the Australian Museum under a past species from Oman (Randall, 1995), 1,078 from exchange programme. One lot of fishes collected the Red Sea (Golani and Bogorodsky, 2010), 1,090 from Myanmar during the first joint Myanmar-Thai from the Maldives (Anderson et al., 1998), and Fishery Exploratory Survey in 1990 was also 1,800 (as an estimate) from southern Africa deposited in the Reference Collection (Heemstra and Heemstra, 2004). There may be (Nateewathana et al., 1983). A revised checklist the real differences in fish diversity among those of fishes in the PMBC Reference Collection was localities, but it should be noted that the number published by Nateewathana et al. (1983), of species recorded probably reflect biases or comprising a total of 629 species in 119 families. differences in any or all of the following aspects: The collection has gradually increased following a sampling effort, habitat/depth representation, and number of subsequent researches (e.g., geographical coverage. Sirimontraporn et al., 1993; Satapoomin and The 10 most speciose families (Gobiidae, Poovachiranon, 1997; Satapoomin, 1999, 2002b) Labridae, Pomacentridae, Serranidae,Apogonidae, and intensive sampling programmes (e.g., a Blenniidae, Carangidae, Scorpaenidae, Lutjanidae collaborative project between ROM and PMBC in and Chaetodontidae) collectively account for about 1993 and the PMBC’s past and ongoing projects, 42% of the total fish fauna. The cumulative namely Coral Reef Management Project (1995– percentage contribution when the top 20 families 1999), Seagrass Resources Management and are taken into account (Table 1) is over half the Dugong Conservation Project (1997–2000), and total (56%), suggesting that the fauna is not Assessment of Coastal Marine Resources Project predominated by only a few families but by a set (since 2004)). According to the recent PMBC data of several families. This seems to reflect the repository (http://pmbc.go.th/pmbc-rc/ diversity of habitats encompassed by the current Taxonomy.php/), the most reliable records of fishes faunal list. Each habitat usually contains a unique (excluding doubtful records and/or composition of the predominant fish families, misidentifications) deposited in the Reference except for the Gobiidae, which is the most diverse Collection total at least 1,200 species belonging to group in nearly all coastal habitats (e.g., coral reefs, 170 families. TheAndaman Sea fishes, in particular, seagrass beds and mangroves – Satapoomin, 2006, are well represented with a total number of 1,093 2007; Fish Team of the Trang Project, 2002). The species, comprising 1, 39, and 1,053 species in current checklist, which includes species from 35 The fishes of southwestern Thailand

Table 1. Twenty most speciose families of fishes of southwestern Thailand, the Andaman Sea.

Family Number of genera Number of species % of total fish fauna

Gobiidae 62 227 13.0 Labridae 30 78 4.5 Pomacentridae 18 71 4.1 Serranidae 17 61 3.5 Apogonidae 14 60 3.4 Blenniidae 21 52 3.0 Carangidae 20 52 3.0 Scorpaenidae 23 49 2.8 Lutjanidae 10 39 2.2 Chaetodontidae 7 37 2.1 Acanthuridae 5 32 1.8 Muraenidae 7 32 1.8 Syngnathidae 11 26 1.5 Carcharhinidae 7 25 1.4 Nemipteridae 3 25 1.4 Engraulidae 5 24 1.4 Lethrinidae 5 22 1.3 Leiognathidae 8 21 1.2 Tetraodontidae 8 21 1.2 Scaridae 6 20 1.1

Subtotal 287 974 55.8

Other 176 families 409 772 44.2

Total 696 1,746 100.0 various habitats, thus evens out the significance diverse families include 25 families with a range contribution of certain predominant families in the of species numbers of 10–19 species, 27 families analysis. The relative abundance of these families with 5–9 species, 59 families with 2–4 species, is similar, although with variable ranking, to those and the remaining 66 families each with single in studies of inshore fishes at many other Indo- species. Pacific localities, for example, Reunion Island With respect to habitat association, the fish (Letourneur et al., 2004), Maldive Islands (Randall fauna consists overwhelmingly of reef fishes, and Anderson, 1993), Chagos Archipelago which comprise about 56% (983 species) of the (Winterbottom andAnderson, 1999), northwestern total species. The coastal species from mangroves Australia (Allen, 1993), Calamianes Islands, and seagrass beds represent about 17% (296 Palawan Province (Allen, 2000), southern Taiwan species) and 14% (246 species) of the total fauna, (Shao et al., 2008), Yaku-shima Island, southern respectively. The other major component, which Japan (Motomura et al., 2010), and Mariana is loosely categorized to accommodate fishes Islands (Myers and Donaldson, 2003). The inhabiting a variety of offshore habitats (spanning Gobiidae, in particular, was usually the leading pelagic to benthic zones at various depths), family when it was adequately sampled. Other less comprises about 55% (971 species) of the total 36 Phuket mar. biol. Cent. Res. Bull.

fauna. The figures accounting for the proportion primarily Indian Ocean range, with lesser values of species shared between habitats vary (Table 3) for either the fauna confined to the eastern considerably and are relatively low, i.e., 42% extremity of the Indian Ocean (about 1%) or the between mangroves and seagrass beds, 20.5% fauna restricted only to the Andaman Sea province between offshore habitats and coral reefs, 16.8% (1.7%). The remaining species (4.9%) have pan- between offshore habitats and seagrass beds, oceanic ranges, including circumtropical and 15.6% between offshore habitats and mangroves, cosmopolitan distributions. The presence of fauna 9.4% between coral reefs and seagrass beds, and having both an Indian Ocean distribution range and 5.3% between coral reefs and mangroves. The also a western Pacific distribution range in the area species shared among the 4 biotopes account for can be considered as compelling evidence of a only 2.3% of the total species recorded. Species distribution confluence of both Indian Ocean and shared between or among habitats, however, Pacific Ocean faunas in the region. The boundary comprise a number of species which are strongly where sympatry of the eastern (= Pacific Ocean) dependent on more than a single habitat type, and and western (= Indian Ocean) populations is most often exhibit a segregated habitat-use pattern with evident seems to encompass Christmas Island and respect to their different life-history stages (e.g., the Java–Bali region (see, Randall, 1998; Allen and Mumby et al., 2004; Shibuno et al., 2008). This Adrim, 2003; Hobbs et al., 2010). The strong seems to suggest an inter-system connection in influence of the Indian Ocean fauna in the area fish populations. Because unique assemblages of can be further highlighted by considering the species are restricted to each particular habitat number of sibling species that involve Indian and type, while others depend upon the existence of Pacific Ocean species pairs, and some geographic multiple habitat-types through connectivity, special variants that contain different Indian and Pacific attention must be paid to holistic approaches of Ocean colour morphs. While inconsistent multi-ecosystems management and conservation considerations among ichthyologists over if the overall diversity of the fishes is to be taxonomic status (i.e., variants, subspecies, or full maintained. At present, there is growing concern species) of the latter exist, concrete morphological over the science of connectivity because and/or genetic evidence is required to ascertain understanding of the linkage among ecosystems their taxonomic status (Randall, 1998; Hastings is considered as a key to the success of and Springer, 2009). As exemplified in Table 4, conservation and management of marine and which includes 68 sibling species-pairs (in 17 coastal areas, particularly when emphasizing the families) and 18 species (in 10 families) possessing design of marine protected areas (MPAs) and MPA geographic variation in colour morphs, the network (Botsford et al., 2009; Sale et al., 2010). Andaman Sea fauna is predominantly represented The zoogeographic analysis of species by the Indian Ocean forms. with adequately known distribution ranges is Since establishment of the PMBC presented in Table 2. The analysis reveals that the Reference Collection about 25 years ago, a great fish fauna consists primarily of species which are deal of knowledge has accumulated and well over widely distributed either in the Indo-Pacific or Indo- 190 new species (see also Aungtonya et al., 2006) west Pacific (50.7%). Approximately 17.1% of the have been described from the Andaman Sea. At fauna are found within portions of the western present, the Reference Collection maintains type Pacific and northeastern Indian Ocean (i.e., NIO– materials of 21 fish species, including 9 holotypes WCP, NIO–WP, NIO–IA, NIO–IM, EIO–WCP, and 27 lots of paratypes (Table 5). Reliable records EIO–WP, EIO–IA, and EIO–IM). Similarly 15.5% of Andaman Sea fishes currently deposited at the of species have a distribution range in the western Reference Collection account for about 63% of Pacific or portions of this region (i.e., WCP, WP, the total fish fauna. The fish collection thus still IA, and IM). This clearly confirms that eastern does not include a comprehensive representation most boundary of the Indian Ocean including the of theAndaman Sea fish fauna. In order to maintain Andaman Sea overlaps the biogeographic region its significant role in supporting biodiversity of the West Pacific. About 12% of the fauna has a research of the ichthyofauna of this particular 37 The fishes of southwestern Thailand

region, the fish collection needs to be upgraded Although the present checklist provides by additional sampling so that the missing 27% of the most extensive species records for the area, it species can be made available as voucher or should be considered as a provisional rather than reference materials for future researchers. Also a complete faunal list. Also, there are a number of worth mentioning here are the figures reflecting identifications that still need verification through biotope representation of the fishes catalogued in examination of voucher specimens. Furthermore, the Reference Collection, which vary considerably, the sampling efforts for several fish groups and/ i.e., 64% of total offshore fishes and 65% of the or in certain habitats remain deficient. The present reef-associated fishes, but 81% of the mangrove provisional checklist, however, can at least help fishes and 88% of the seagrass fishes. These to draw the attention of ichthyologists who are figures can form a basis for targeting future interested in fish fauna of the Andaman Sea to sampling programs for the Reference Collection. where there are knowledge gaps. Also, it can serve Small and cryptic reef fishes and particularly those as a baseline for a comprehensive taxonomic inhabiting the deep-reef habitats are in need of review and/or annotated checklist of the Andaman extensive sampling. For fishes dwelling among Sea fishes. It is hoped that, joint research or various offshore habitats, deep demersal, collaborative efforts will be possible in the near benthopelagic, and mesopelagic fishes are among future to accomplish such goals. While the current the groups that have not been adequately sampled. limited effort and interest, especially with respect Deep sea exploratory research programs are to taxonomy, has slowed down the progress in generally costly and particularly require specific this research area, a concomitant obstacle techniques, equipment as well as taxonomic regarding decreasing personnel trained in expertise. The establishment of such programs is ichthyology in Thailand seems to be more critical likely to require international collaborative efforts and needs urgent action. and funding. Table 2. Zoogeographic analysis of the fish fauna of southwestern Thailand, the Andaman Sea.

Distribution No. species %

Indo-Pacific (IP) 417 24.5 Indo-West Pacific (IWP) 448 26.3 West central Pacific (WCP) 27 1.6 West Pacific (WP) 137 8.0 Indo-Australian Archipelago (IA) 48 2.8 Indo-Malayan Archipelago (IM) 53 3.1 Indian Ocean (IO) 99 5.8 Northern Indian Ocean (NIO) 10 0.6 Eastern Indian Ocean (EIO) 46 2.7 Northern Indian Ocean to West central Pacific (NIO–WCP) 2 0.1 Northern Indian Ocean to West Pacific (NIO–WP) 37 2.2 Northern Indian Ocean to Indo-Australian Archipelago (NIO–IA) 21 1.2 Northern Indian Ocean to Indo-Malayan Archipelago (NIO–IM) 29 1.7 Eastern Indian Ocean to West central Pacific (EIO–WCP) 12 0.7 Eastern Indian Ocean to West Pacific (EIO–WP) 112 6.6 Eastern Indian Ocean to Indo-Australian Archipelago (EIO–IA) 28 1.6 Eastern Indian Ocean Indo-Malayan Archipelago (EIO–IM) 50 3.0 Eastern extremity of Indian Ocean (EEIO) 15 0.9 Andaman Sea (AS) 28 1.7 Circumtropical (CIR) 38 2.2 Cosmopolitan (COS) 46 2.7 38 Phuket mar. biol. Cent. Res. Bull.

Table 3. List of fishes currently known to have restricted distribution in the Andaman Sea and adjacent areas: A) Andaman Sea endemics, including the species ranges southward to western Sumatra; B) The species found in Andaman Sea and some other localities along eastern edge of the eastern Indian Ocean (= eastern extremity of Indian Ocean (EEIO))

Family: Species Distribution

A)Andaman Sea endemics Myxinidae: Eptatretus indrambaryai Wongratana, 1983 Andaman Sea Proscylliidae: Proscyllium magnificum Last & Vongpanich,2004 Andaman Sea Rajidae: Anacanthobatis sp.1 Andaman Sea Acropomatidae: Acropoma argentistigma Okamoto & Ida, 2002 Andaman Sea Aploactinidae: Cocotropus echinatus (Cantor, 1850) Andaman Sea Apogonidae: Apogon sp.1 West coast of Thailand Archamia ataenia Randall & Satapoomin, 1999 Andaman Sea–W. Sumatra Blenniidae: Blenniella leopardus (Fowler, 1904) Andaman Sea–W. Sumatra Meiacanthus urostigma Smith-Vaniz, Satapoomin & Allen, 2001 Andaman Sea–W. Sumatra Bythitidae: Alionematichthys phuketensis Møller & Schwarzhans, 2008 West coast of Thailand Gobiidae: Callogobius andamanensis Menon & Chatterjee, 1974 Andaman Sea Scartelaos cantoris (Day, 1871) Andaman Sea Trimma fucatum Winterbottom & Southcott, 2007 West coast of Thailand Trimma sanguinellus Winterbottom & Southcott, 2007 Andaman Sea–W. Sumatra Haemulidae: Hapalogenys merguiensis Iwatsuki, Satapoomin & Amaoka, 2000 Andaman Sea Plectorhinchus macrospilus Satapoomin & Randall, 2000 West coast of Thailand Pomadasys andamanensis McKay & Satapoomin, 1994 West coast of Thailand Labridae: Cirrhilabrus joanallenae Allen, 2000 Andaman Sea–W. Sumatra Halichoeres kallochroma (Bleeker, 1853) Andaman Sea–W. Sumatra Monacanthidae: Paraluteres sp.1 Andaman Sea Ophichthidae: Allips concolor McCosker, 1972 Andaman Sea Phallostethidae: Phenacostethus posthon Roberts, 1971 Andaman Sea Plesiopidae: Plesiops thysanopterus Mooi, 1995 N. Sumatra (Andaman Sea) Pomacentridae: Chromis xouthos Allen & Erdmann, 2005 Andaman Sea–W. Sumatra Pomacentrus polyspinus Allen, 1991 West coast of Thailand Serranidae: Cephalopholis polyspila Randall & Satapoomin, 2000 Andaman Sea–W. Sumatra Meganthias filiferus Randall & Heemstra, 2007 Andaman Sea 39 The fishes of southwestern Thailand

Table 3. (continued)

Family: Species Distribution

Soleidae: Aseraggodes satapoomini Randall & Desoutter-Meniger, 2007 West coast of Thailand Sternoptychidae: Polyipnus asper Harold, 1994 Andaman Sea Tripterygiidae: Helcogramma lacuna Williams & Howe, 2003 West coast of Thailand

B) Limited to eastern extremity of Indian Ocean (EEIO) Urolophidae: Urolophus javanicus (Martens, 1864) Andaman Sea, Java Sea (W. Java) Blenniidae: Ecsenius lubbocki Springer, 1988 Andaman Sea, Bali Ecsenius paroculus Springer, 1988 Andaman Sea., W. Sumatra, Java Sea (Seribu Is.& N. Java) Praealticus triangulus (Chapman, 1951) Andaman Sea, Java Sea (N. Java) Labridae: Cirrhilabrus cyanopleura (Bleeker, 1851) Andaman Sea, W. Sumatra, Christmas Is., Java Sea (Pulau Putri & N. Java), Bali Malacanthidae: Hoplolatilus luteus Allen & Kuiter, 1989 Andaman Sea, Bali, Flores Opistognathidae: Opistognathus cyanospilotus Smith-Vaniz, 2009 Andaman Sea, Bali Polynemidae: Filimanus perplexa Feltes, 1991 Andaman Sea, W. Sumatra, S. Java, Bali Pomacentridae: Pomacentrus alleni Burgess, 1981 Andaman Sea, Christmas Is., Java Sea (Sunda Strait & Pulau Putri) Pomacentrus azuremaculatus Allen, 1991 Andaman Sea, Java Sea (Seribu Is.) Pomacentrus xanthosternus Allen, 1991 Andaman Sea, W. Sumatra, Java Sea (Seribu Is.), Bali Pseudochromidae: Pseudochromis andamanensis Lubbock, 1980 Andaman Sea, W. Sumatra, Bali, Timor Sea (Ashmore Reef & Cartier Is., N. Australia) Scaridae: Chlorurus troschelii (Bleeker, 1853) Andaman Sea, Java Sea (Seribu Is.)

Scarus maculipinna Westneat, Satapoomin & Randall, 2007 Andaman Sea, W. Sumatra, Java Sea (Pulau Putri) Siganidae: Siganus magnificus (Burgess, 1977) Andaman Sea, W. Sumatra, Java Sea (W. Java) 40 Phuket mar. biol. Cent. Res. Bull.

Table 4. Examples of Indian and Pacific Ocean species pairs and geographic variants that the Andaman Sea fishes represented by the Indian Ocean forms. A) Sibling species: Andaman Sea fish (vs. its Pacific sibling)

Acanthuridae Pinguipedidae Acanthurus tristis (A. pyroferus) Parapercis maculata (P. pulchella) Ctenochaetus truncatus (C. strigosus) Pomacanthidae Naso elegans (N. lituratus) Centropyge eibli (C. vroliki) Zebrasoma desjardinii (Z. veliferum) Genicanthus caudovittatus (G. melanospilos) Apogonidae Pomacentridae Archamia ataenia (A. zosterophora) Amphiprion akallopisos (A. sandarcinos) Blenniidae Amphiprion ephippium (A. melanopus) Aspidontus tractus (A. taeniatus) Amphiprion sebae (A. polymnus) Atrosalarias fuscus (A. holomelas) Chromis dimidiata (C. iomelas) Blenniella cyanostigma (B. caudolineata) Chromis opercularis (C. xanthura) Blenniella leopardus (B. interrupta) Dascyllus carneus (D. reticulatus) Meiacanthus urostigma (M. ditrema) Neopomacentrus sororius (N. azysron) Plagiotremus phenax (P. laudandus) Pomacentrus similis (P. coelestis) Chaetodontidae Pomacentrus xanthosternus (P. simsiang) Chaetodon andamanensis (C. plebeius) Ptereleotridae Chaetodon triangulum (C. baronessa) Ptereleotris arabica (P. hanae) Chaetodon falcula (C. ulietensis) Scaridae Chaetodon gardineri (C. selene) Chlorurus capistratoides (C. japanensis) Chaetodon interruptus (C. unimaculatus) Chlorurus rhakoura (C. oedema) Chaetodon madagaskariensis (C. mertensii) Chlorurus sordidus (C. spilurus) Chaetodon melannotus (C. ocellicaudus) Chlorurus strongylocephalus (C. microrhinos) Chaetodon trifasciatus (C. lunulatus) Chlorurus troschelii (C. bleekeri) Hemitaurichthys zoster (H. polylepis) Hipposcarus harid (H. longiceps) Heniochus pleurotaenia (H. varius) Scarus maculipinna (S. flavipectoralis) Gobiidae Scarus russelii (S. schlegeli) Amblygobius semicinctus (A. phalaena) Scarus scaber (S. oviceps) Trimma flammeum (T. macrophthalma) Scarus viridifucatus (S. spinus) Labridae Scorpaenidae Anampses lineatus (A. melanurus) Pterois miles (P. volitans) Cirrhilabrus cyanopleura (C. lyukyuensis) Serranidae Coris cuvieri (C. gaimard) Cephalopholis polyspila (C. microprion) Bodianus diana (B. dictynna) Pogonoperca ocellata (P. punctata) Gomphosus caeruleus (G. varius) Pseudanthias ignitus (P. dispar) Halichoeres cosmetus (H. claudia) Rabualichthys stigmaticus (R. altipinnis) Halichoeres kallochroma (H. leucurus) Siganidae Halichoeres leucoxanthus (H. chrysus) Siganus puelloides (S. puellus) Halichoeres vrolokii (H. melanurus) Siganus stellatus (S. punctatus) Halichoeres zeylonicus (H. hartzfeldii) Syngnathidae Paracheilinus mccoskeri (P. carpenteri) Doryramphus multiannulatus (D. pessuliferus) Pseudojuloides kaleidos (P. cerasinus) Tetraodontidae Mullidae Canthigaster smithae (C. epilampra) Parupeneus trifasciatus (P. crassilabris) 41 The fishes of southwestern Thailand

Table 4. (continued)

B) Geographic variants: Andaman Sea fish possessing Indian Ocean colour morph

Dasyatidae Nemipteridae *(4)Dasyatis kuhlii *(6, 7)Scolopsis bilineatus Acanthuridae Scolopsis vosmeri *(7)Paracanthurus hepatus Pomacanthidae Balistidae *(7, 8)Pomacanthus imperator *(5, 7)Balistapus undulatus *(5, 7)Pygoplites diacanthus Caesionidae Pomacentridae Caesio cuning *(2)Chrysiptera rollandi Labridae *(7)Neoglyphidodon nigroris *(9)Epibulus insidiator *(6)Pomacentrus chrysurus *(5, 7)Halichoeres hortulanus *(2, 8)Pomacentrus pavo *(5)Labroides dimidiatus Serranidae *(7)Thalassoma jansenii *(3)Cephalopholis urodeta Lutjanidae *(1, 7)Lutjanus russelli

*Note: Recognition of allopatric colour morphs in certain species can be referred to (1) Allen and Talbot, 1985; (2) Allen, 1991; (3) Randall and Heemstra, 1991; (4) Randall, 1995; (5) Randall, 1998; (6) Satapoomin, 2000; (7) Kuiter and Tonozuka, 2001; (8) Lieske and Myers, 2001; and (9) Carlson et al., 2008.

Table 5. List of fishes for which their type materials are deposited at the Reference Collection of the Phuket Marine Biological Center. Number in bracket denotes the number of catalogued lots for paratypes.

Family: Species Type materials

Apogonidae: Archamia ataenia Randall & Satapoomin, 1999 Paratypes (1) Blenniidae: Meiacanthus urostigma Smith-Vaniz, Satapoomin & Allen, 2001 Paratypes (2) Gobiidae: Cryptocentrus crocatus Wongratana, 1975 Holotype [valid as Myersina crocatus (Wongratana, 1975)] Gladiogobius rex Shibukawa & Allen, 2007 Holotype, Paratypes (3) Myersina adonis Shibukawa & Satapoomin, 2006 Holotype, Paratypes (1) Periophthalmus walailakae Darumas & Tantichodok, 2002 Paratypes (2) Trimma fucatum Winterbottom & Southcott, 2007 Paratypes (1) Trimma sanguinellus Winterbottom & Southcott, 2007 Paratypes (1) Haemulidae: Hapalogenys merguiensis Iwatsuki, Satapoomin & Amaoka, 2000 Paratypes (2) Plectorhinchus macrospilus Satapoomin & Randall, 2000 Paratypes (1) Pomadasys andamanensis McKay & Satapoomin, 1994 Holotype Nemipteridae: Nemipterus aurora Russell, 1993 Paratypes (1) Nemipterus thosaporni Russell, 1991 Paratypes (1) 42 Phuket mar. biol. Cent. Res. Bull.

Table 5. (continued)

Family: Species Type materials

Opistognathidae: Opistognathus variabilis Smith-Vaniz, 2009 Paratypes (2) Proscylliidae: Proscyllium magnificum Last & Vongpanich, 2004 Holotype, Paratypes (3) Pomacentridae: Pomacentrus azuremaculatus Allen, 1991 Holotype Pomacentrus polyspinus Allen, 1991 Holotype Pomacentrus similis Allen, 1991 Paratypes (1) Scaridae: Scarus maculipinna Westneat, Satapoomin & Randall, 2007 Holotype, Paratypes (1) Serranidae: Cephalopholis polyspila Randall & Satapoomin, 2000 Paratypes (4) Meganthias filiferus Randall & Heemstra, 2007 Holotype

ACKNOWLEDGEMENTS thanks are due to Dr. Keiichi Matsuura [KM] of the National Museum of Nature and Science, This paper was initially prepared as a Tokyo, for his considerable effort in leading the contribution to the concluding conference of the fish diversity project under the JSPS multilateral 10-year JSPS Multilateral Core University programme and encouraging me to write up this Programme on “Coastal Marine Science” in Tokyo paper. I am very grateful to Dr. Richard during 26–29 October 2010, but I finally could Winterbottom [RW] of the Royal Ontario Museum not make the trip. It was then presented as a plenary for sharing the research outcomes from the 1993 talk at the seminar on “Biodiversity of Southern expedition to Phuket and inspiring my interest in Thailand in Focus: Status and Research Direction” small cryptobenthic reef fishes. Both KM and RW at Walailak University, Nakorn Sri Thammarat, kindly gave their time for reviewing and improving Thailand, during 10–12 November 2010. I am the first draft of this manuscript. Several past and grateful for the generous support made available ongoing projects at PMBC which contribute to the by the JSPS for me to be able to join several fish information and knowledge on the Andaman Sea workshops conducted over the years, which fish fauna have been continually supported by the ultimately gave me a wonderful opportunity to learn Thai government budget. I am also grateful to the more about the fish faunas of SE Asia. Special many staff members of the PMBC for their assistance in the field.

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Manuscript received: 10 January 2011 Accepted: 1 August 2011 47 The fishes of southwestern Thailand

Appendix A Order Carcharhiniformes Scyliorhinidae Checklist of marine fishes of southwestern Atelomycterus marmoratus (Bennett, 1830): OShr; [IWP] Thailand, the Andaman Sea Cephaloscyllium umbratile Jordan & Fowler, 1903: OShr; [WP]; PMBC Abbreviations: Halaelurus buergeri (Müller & Henle, 1838): OShr; [WP] 1. Habitat types – Cr (coral reefs); Mg (mangroves); Sg Halaelurus hispidus (Alcock, 1891): OShr; [EIO-WP]; (seagrass beds); OShr (other coastal and offshore areas) PMBC 2. Distribution categories – IP (Indo-Pacific); IWP (Indo- Proscylliidae West Pacific); WCP (west central Pacific); WP (west Proscyllium magnificum Last & Vongpanich, 2004: OShr; Pacific); IA (Indo-Australian Archipelago); IM (Indo- [AS]; PMBC Malayan Archipelago); IO (Indian Ocean); NIO (northern Triakidae Indian Ocean); EIO (eastern Indian Ocean); EEIO (eastern Iago omanensis (Norman, 1939): OShr; [IO]; PMBC extremity of Indian Ocean); AS (Andaman Sea); CIR Mustelus manazo Bleeker, 1854: OShr; [IWP]; PMBC (circumtropical); COS (cosmopolitan) [see methods Mustelus mosis Hemprich & Ehrenberg, 1899: OShr; [IO]; section for definitions] PMBC 3. Voucher specimens – PMBC (denotes availability of Hemigaleidae voucher specimens at the Reference Collection of the Chaenogaleus macrostoma (Bleeker, 1852): OShr; [IWP] Phuket Marine Biological Center) Hemigaleus microstoma Bleeker, 1852: OShr; [EIO-WP]; PMBC CLASS MYXINI Hemipristis elongatus (Klunzinger, 1871): OShr; [IWP]; Order Myxiniformes PMBC Myxinidae Carcharhinidae Eptatretus indrambaryai Wongratana, 1983: OShr; [AS]; Carcharhinus albimarginatus (Rüppell, 1837): Cr, OShr; PMBC [IP] CLASS CHONDRICHTHYES Carcharhinus altimus (Springer, 1950): OShr; [COS] Order Chimaeriformes Carcharhinus amblyrhynchoides (Whitley, 1934): OShr; Chimaeridae [IWP] Hydrolagus sp.: OShr; PMBC Carcharhinus amblyrhynchos (Bleeker, 1856): OShr; [IP] Order Orectolobiformes Carcharhinus amboinensis (Müller & Henle, 1839): OShr; Parascylliidae [IWP] Parascyllium sp.: OShr; PMBC Carcharhinus brachyurus (Günther, 1870): OShr; [COS] Hemiscylliidae Carcharhinus brevipinna (Müller & Henle, 1839): OShr; Chiloscyllium griseum Müller & Henle, 1838: OShr; [NIO- [CIR] IM]; PMBC Carcharhinus dussumieri (Valenciennes, 1839): OShr; Chiloscyllium hasselti Bleeker, 1852: Sg, OShr; [IM]; [NIO-WP] PMBC Carcharhinus falciformis (Müller & Henle, 1839): OShr; Chiloscyllium indicum (Gmelin, 1788): OShr; [IWP]; [CIR] PMBC Carcharhinus leucas (Valenciennes, 1839): OShr; [CIR] Chiloscyllium plagiosum (Bennett, 1830): OShr; [IWP] Carcharhinus limbatus (Valenciennes, 1839): OShr; [COS] Chiloscyllium punctatum Müller & Henle, 1838: Cr, Sg, Carcharhinus longimanus (Poey, 1861): OShr; [COS] OShr; [EIO-WP] Carcharhinus macloti (Müller & Henle, 1839): OShr; [IWP]; Stegostomatidae PMBC Stegostoma fasciatum (Hermann, 1793): Cr, OShr; [IP]; Carcharhinus melanopterus (Quoy & Gaimard, 1824): Cr, PMBC OShr; [IP]; PMBC Ginglymostomatidae Carcharhinus obscurus (LeSueur, 1818): OShr; [COS] Nebrius ferrugineus (Lesson, 1830): Cr, OShr; [IWP] Carcharhinus plumbeus (Nardo, 1827): OShr; [CIR]; Rhincodontidae PMBC Rhincodon typus Smith, 1828: Cr, OShr; [COS] Carcharhinus sealei (Pietschmann, 1916): OShr; [IWP] Order Lamniformes Carcharhinus sorrah (Valenciennes, 1839): OShr; [IWP] Alopiidae Galeocerdo cuvier (Péron & LeSueur, 1822): OShr; [CIR] Alopias pelagicus Nakamura, 1935: OShr; [CIR] Loxodon macrorhinus Müller & Henle, 1839: OShr; Alopias superciliosus (Lowe, 1840): OShr; [CIR] [IWP] Alopias vulpinus (Bonnaterre, 1788): OShr; [COS] Negaprion acutidens (Rüppell, 1837): OShr; [IP] Lamnidae Rhizoprionodon acutus (Rüppell, 1837): OShr; [CIR]; Isurus oxyrinchus Rafinesque, 1810: OShr; [COS] PMBC Rhizoprionodon oligolinx Springer, 1964: OShr; [IWP] 48 Phuket mar. biol. Cent. Res. Bull.

Scoliodon laticuadus Müller & Henle, 1839: OShr; [IWP] Urolophidae Triaenodon obesus (Rüppell, 1835): Cr, OShr; [IP] Urolophus javanicus (Martens, 1864): OShr; [EEIO] Sphyrnidae Dasyatidae Eusphyra blochii (Cuvier, 1817): OShr; [NIO-WP]; PMBC Dasyatis kuhlii (Müller & Henle, 1841): Cr, OShr; [IWP]; Sphyrna lewini (Griffith & Smith, 1834): OShr; [COS]; PMBC PMBC Dasyatis zugei (Müller & Henle, 1841): OShr; [EIO-WP] Sphyrna mokarran (Rüppell, 1837): OShr; [CIR] Himantura fai Jordan & Seale, 1906: OShr; [IWP] Sphyrna zygaena (Linnaeus, 1758): OShr; [CIR]; PMBC Himantura gerrardi (Gray, 1851): Sg, OShr; [IP]; PMBC Order Hexanchiformes Himantura imbricata (Bloch & Schneider, 1801): Mg, Sg, Hexanchidae OShr; [IWP]; PMBC Heptranchias perlo (Bonnaterre, 1788): OShr; [COS] Himantura jenkinsii (Annadale, 1909): Cr, OShr; [EIO] Order Squaliformes Himantura undulata (Bleeker, 1852): OShr; [EIO-WP] Squalidae Himantura walga (Müller & Henle, 1841): Sg, OShr; [EIO- Etmopterus brachyurus Smith & Radcliffe, 1912: OShr; IM]; PMBC [WP] Pastinachus sephen (Forsskål, 1775): Cr, OShr; [IWP] Squalus acanthias Linnaeus, 1758: OShr; [COS]; PMBC Taeniura lymma (Forsskål, 1775): Cr; [IWP]; PMBC Squalus mitzukurii Jordan & Snyder, 1903: OShr; [CIR]; Taeniura meyeni Müller & Henle, 1841: Cr; [IP] PMBC Urogymnosus asperrimus (Bloch & Schneider, 1801): Cr; Order Squantiformes [IP] Squatinidae Gymnuridae Squatina sp.1: OShr Gymnura poecilura (Shaw, 1804): Mg, OShr; [IP]; PMBC Order Torpediniformes Myliobatidae Narcinidae Aetobatus narinari (Euphrasen, 1790): Cr, OShr; [COS] Narcine brunnea Annandale, 1909: OShr; [NIO-IM]; Aetomylaeus maculatus (Gray, 1832): OShr; [EIO-WP] PMBC Aetomylaeus milvus (Valenciennes, 1841): OShr; [IWP] Narcine indica Henle, 1834: OShr; [WP] Aetomylaeus nichofii (Bloch & Schneider, 1801): OShr; Narcine maculata (Shaw, 1804): OShr; [WP]; PMBC [IWP]; PMBC Narcine prodosalis Bessednov, 1966: OShr; [EIO-WP] Aetomylaeus vespertilio (Bleeker, 1852): OShr; [IWP] Narcine timlei (Bloch & Schneider, 1801): OShr; [EIO- Manta birostris (Donndorff, 1798): Cr, OShr; [CIR] WP] Mobula kuhlii (Valenciennes, in Müller & Henle, 1841): Narke dipterygia (Bloch & Schneider, 1801): OShr; [NIO- OShr; [IWP] WP] Rhinoptera javanica Müller & Henle, 1841: OShr; [IWP]; Temera hardwickii Gray, 1831: OShr; [IM]; PMBC PMBC Order Pristiformes Rhinoptera sewelli Misra, 1946: OShr; [EIO] Pristidae CLASS ACTINOPTERYGII Pristis microdon Latham, 1794: OShr; [CIR]; PMBC Order Elopiformes Order Rajiformes Elopidae Rhinidae Elops hawaiiensis Regan, 1909: Mg, Sg, OShr; [WCP]; Rhina ancylostoma (Bloch & Schneider, 1801): Cr, OShr; PMBC [IP]; PMBC Megalopidae Rhynchobatidae Megalops cyprinoides (Broussonet, 1782): Mg, OShr; [IP]; Rhynchobatus palpebratus Compagno & Last, 2008: Cr, PMBC OShr; [IA (presently known from west coast of Thailand Order Albuliformes and northern Australia)] Albulidae Rhinobatidae Albula glossodonta (Forsskål, 1775): OShr; [IP] Rhinobatos granulatus Cuvier, 1829: OShr; [NIO-IM]; Albula oligolepis Iwatsuki & Hidaka, 2008: OShr; [IO]; PMBC PMBC Rhinobatos thouin (Anonymous in Lacepède, 1798): OShr; Notacanthidae [IWP] Notacanthus sp.: OShr; PMBC Rhinobatos typus Bennett, 1830: OShr; [EIO-WP] Order Anguilliformes Rajidae Anguillidae Anacanthobatis sp.1 : OShr, PMBC Anguilla bicolor bicolor McClelland, 1844: Mg; [IWP]; Raja sp.: OShr PMBC Order Myliobatiformes Anguilla marmorata Quoy & Gaimard, 1824: OShr; [IP]; Plesiobatidae PMBC Plesiobatis daviesi Wallace, 1967: OShr; [IP]; PMBC Chlopsidae 49 The fishes of southwestern Thailand

Kaupichthys sp.: Cr Scolecenchelys macroptera (Bleeker, 1857): Sg, OShr; Muraenidae [IWP]; PMBC Echidna delicatula (Kaup, 1856): OShr; [EIO-WP]; PMBC Xyrias revulsus Jordan & Snyder, 1901: OShr; [IWP] Echidna nebulosa (Ahl, 1789): Cr; [IP]; PMBC Colocongridae Echidna polyzona (Richardson, 1844): Cr; [IP]; PMBC Coloconger scholesi Chan, 1967: OShr; [IWP]; PMBC Enchelycore bayeri (Schultz, 1953): Cr; [IP] Coloconger sp.: OShr; PMBC Gymnomuraena zebra (Shaw, 1797): Cr; [IP]; PMBC Muraenesocidae Gymnothorax annulatus (Smith & Bohlke, 1997): OShr; Congresox talabon (Cuvier, 1829): OShr; [EIO-IM]; [IM] PMBC Gymnothorax buroensis (Bleeker, 1857): Cr; [IP] Congresox talabonoides (Bleeker, 1853): OShr; [IWP]; Gymnothorax chilospilus Bleeker, 1865: Cr; [IP]; PMBC PMBC Gymnothorax favagineus (Bloch & Schneider, 1801): Cr; Muraenosox bagio (Hamilton, 1822): OShr; [IWP] [IWP]; PMBC Muraenosox cinereus (Forsskål, 1775): OShr; [IWP]; Gymnothorax fimbriatus (Bennett, 1832): Cr; [IP]; PMBC PMBC Gymnothorax flavimarginatus (Rüppell, 1828): Cr; [IP] Congridae Gymnothorax javanicus (Bleeker, 1859): Cr; [IP]; PMBC Ariosoma anago (Temminck & Schlegel, 1846): OShr; Gymnothorax margaritophorus Bleeker, 1864: Cr; [IP] [IWP]; PMBC Gymnothorax melatremus Schultz, 1953: Cr; [IP] Bathycongrus guttulatus (Günther, 1887): OShr; [IP] Gymnothorax meleagris (Shaw & Nodder, 1795): Cr; [IP] Bathyuroconger sp.: OShr; PMBC Gymnothorax moluccensis (Bleeker, 1864): Cr; [IWP] Conger cinereus Rüppell, 1828: OShr; [IP]; PMBC Gymnothorax monochrous (Bleeker, 1856): Cr; [IP] Gnathophis sp.: OShr; PMBC Gymnothorax permistus (Smith, 1962): Cr; [IO]; PMBC Gorgasia preclara Böhlke & Randall, 1981: Cr; [EIO-IA] Gymnothorax pindae Smith, 1962: Cr; [IP]; PMBC Heteroconger hassi (Klausewitz & Eibl-Eibesfeldt, 1959): Gymnothorax pseudoherrei Böhlke, 2000: Cr; [IWP] Cr; [IP] Gymnothorax pseudothyrsoideus (Bleeker, 1852): Cr; Rhynchoconger squaliceps (Alcock, 1894): OShr; [EIO] [NIO-WP]; PMBC Uroconger lepturus (Richardson, 1845): OShr; [IWP]; Gymnothorax reticularis Bloch, 1795: OShr; [IO]; PMBC PMBC Gymnothorax richardsoni (Bleeker, 1852): Cr; [IWP]; Nettastomatidae PMBC Nettastoma sp.: OShr Gymnothorax rueppelliae (Mcclelland, 1845): Cr; [IP] Order Clupeiformes Gymnothorax thyrsoideus (Richardson, 1844): Cr, OShr; Pristigasteridae [EIO-WCP]; PMBC Ilisha elongata (Bennett, 1830): OShr; [EIO-IM]; PMBC Gymnothorax undulatus (Lacepède, 1803): Cr; [IP]; PMBC Ilisha kampeni (Weber & Beaufort, 1913): Sg, OShr; [EIO- Gymnothorax zonipectis Seale, 1906: Cr; [IP]; PMBC IM] Rhinomuraena quaesita Garman, 1888: Cr; [IWP] Ilisha megaloptera (Swainson, 1839): Mg, OShr; [EIO- Scuticaria tigrina (Lesson, 1828): Cr; [IP] IM]; PMBC Uropterygius concolor Rüppell, 1838: Cr; [IP]; PMBC Ilisha melastoma (Schneider, 1801): Mg, OShr; [EIO-WP]; Uropterygius inornatus Gosline, 1958: Cr; [WCP] PMBC Uropterygius xanthopterus Bleeker, 1859: Cr; [IP] Ilisha sirishai Seshagiri Rao, 1975: OShr; [NIO-IM] Ophichthidae Opisthopterus tardoore (Cuvier, 1829): Mg, OShr; [NIO- Allips concolor McCosker, 1972: Cr, Mg; [AS] IM]; PMBC Apterichthus sp.: OShr; PMBC Pellona ditchela Valenciennes, 1847: Mg, OShr; [IWP]; Brachysomophis crocodilinus (Bennet, 1833): Cr, OShr; PMBC [IP] Raconda russeliana Gray, 1831: OShr; [EIO-IM] Echelus uropterus (Temminck & Schlegel, 1846): OShr; Engraulidae [IWP]; PMBC Coilia dussumieri Valenciennes, 1848: OShr; [EIO-IM]; Muraenichthys gymnopterus (Bleeker, 1853): OShr; [EIO- PMBC WP] Coilia macrognathos Bleeker, 1852: OShr; [IM] Myrichthys colubrinus (Boddaert, 1781): Cr; [IP]; PMBC Coilia neglecta Whitehead, 1968: OShr; [EIO-IM] Neenchelys sp.: OShr; PMBC Enchasicholina punctifer Fowler, 1938: OShr; [IP] Ophichthus evermanni Jordan & Richardson, 1909: OShr; Encrasicholina devisi (Whitley, 1940): OShr; [IWP]; PMBC [WP] Encrasicholina heteroloba (Rüppell, 1837): OShr; [IWP]; Ophichthus sp.: OShr; PMBC PMBC Pisonodophis cancrivorus (Richardson, 1844): Mg, Sg, Setipinna breviceps (Cantor, 1850): OShr; [EIO-IM] OShr; [IWP]; PMBC Setipinna taty (Valenciennes, 1848): Mg, OShr; [EIO-IM] Scolecenchelys gymnota (Bleeker, 1854): Sg; [IWP] Setipinna tenuifilis (Valenciennes, 1848): OShr; [EIO-WP] 50 Phuket mar. biol. Cent. Res. Bull.

Stolephorus andhraensis Babu Rao, 1966: OShr; [EIO- Spratelloides gracilis (Temminck & Schlegel, 1846): Cr, IA]; PMBC Mg, Sg, OShr; [IP] Stolephorus baganensis Hardenberg, 1933: Mg, OShr; Tenualosa reevesii (Richardson, 1846): OShr; [WP] [EIO-IM] Tenualosa toli (Valenciennes, 1847): OShr; [NIO-IM]; Stolephorus chinensis (Günther, 1880): Mg, OShr; [IM] PMBC Stolephorus commersonii Lacepède, 1803: Mg, Sg, OShr; Order Gonorynchiformes [IWP]; PMBC Chanidae Stolephorus dubiosus Wongratana, 1983: OShr; [EIO-IM] Chanos chanos (Forsskål, 1775): Cr, Mg, OShr; [IP] Stolephorus indicus (van Hasselt, 1823): Mg, Sg, OShr; Order Siluriformes [IWP]; PMBC Plotosidae Stolephorus insularis Hardenberg, 1933: Mg, OShr; [NIO- Plotosus canius Hamilton, 1822: Mg, Sg, OShr; [EIO-IA]; WP] PMBC Stolephorus waitei Jordan & Seale, 1926: OShr; [EIO-IA]; Plotosus lineatus (Thunberg, 1787): Cr, Mg, Sg, OShr; PMBC [IWP]; PMBC Thryssa baelama (Forster, 1775): OShr; [IWP] Ariidae Thryssa dussumieiri (Valenciennes, 1848): OShr; [NIO- Arius arius (Hamilton, 1822): OShr; [EIO-IM]; PMBC IM] Arius caelatus Valenciennes, 1840: Mg; [EIO-IM] Thryssa hamiltonii (Gray, 1835): Mg, Sg, OShr; [NIO- Arius gagora (Hamilton, 1822): OShr; [EIO] IA]; PMBC Arius oetik Bleeker, 1846: OShr; [WP] Thryssa kammalensis (Bleeker, 1849): Mg, OShr; [IM] Arius venosus Valenciennes, 1840: OShr; [EIO-IM] Thryssa mystax (Schneider, 1801): OShr; [NIO-IM]; Nemapteryx nenga (Hamilton, 1822): OShr; [EIO-IM] PMBC Nemuta thalassinus (Rüppell, 1837): OShr; [IWP]; PMBC Thryssa setirostris (Broussonet, 1782): Mg, Sg, OShr; Osteogeneiosus militaris (Linnaeus, 1758): OShr; [EIO- [IWP]; PMBC IM] Thryssa spinidens (Jordan & Seale, 1925): OShr; [EIO] Plicofollis nella Bleeker, 1846: OShr; [WP] Chirocentridae Bagridae Chirocentrus dorab (Forsskål, 1775): Mg, Sg, OShr; [IWP]; Mystus gulio (Hamilton, 1822): Mg; [EIO-IM]; PMBC PMBC Order Argentiniformes Chirocentrus nudus Swainson, 1839: OShr; [IWP] Alepocephalidae Clupeidae Alepocephalus bicolor Alcock, 1891: OShr; [IWP]; PMBC Amblygaster clupeoides Bleeker, 1849: OShr; [EIO-WP] Order Stomiiformes Amblygaster leiogaster (Valenciennes, 1847): OShr; [IWP] Sternoptychidae Amblygaster sirm (Walbaum, 1792): OShr; [IWP]; PMBC Argyropelecus affinis Garman, 1899: OShr; [CIR]; PMBC Anodontostoma chacunda (Hamilton, 1822): Mg, Sg, OShr; Argyropelecus sladeni Regan, 1908: OShr; [CIR]; PMBC [NIO-WP]; PMBC Polyipnus asper Harold, 1994: OShr; [AS]; PMBC Anodontostoma thailandiae Wongratana, 1983: OShr; [EIO- Sternoptyx obscura Garman, 1899: OShr; [IWP]; PMBC IM] Stomiidae Dussumieria acuta Valenciennes, 1847: OShr; [NIO-IM]; Astronesthes lucifer Gilbert, 1905: OShr; [IP] PMBC Chauliodus sloanei Bloch & Schneider, 1801: OShr; [COS] Dussumieria elopsoides Bleeker, 1849: Mg, OShr; [IWP] Eustomias sp.: OShr; PMBC Escualosa thoracata (Valenciennes, 1847): Mg, Sg, OShr; Order Ateleopodiformes [NIO-IA]; PMBC Ateleopodidae Herklotsichthys quadrimaculatus (Rüppell, 1837): Cr, Ateleopus indicus Alcock, 1891: OShr; [EIO]; PMBC OShr; [IP]; PMBC Order Aulopiformes Hilsa kelee (Cuvier, 1829): Mg, Sg, OShr; [IWP] Paraulopidae Nematalosa galatheae Nelson & Rothman, 1973: OShr; Paraulopus brevirostris (Fourmanoir, 1981): OShr; [WP] [EIO-IM] Synodontidae Nematalosa nasus (Bloch, 1795): OShr; [EIO-WP] Harpadon nehereus (Hamilton, 1822): OShr; [IWP] Sardinella albella (Valenciennes, 1847): Mg, Sg, OShr; Saurida elongata (Temminck & Schlegel, 1846): OShr; [IWP]; PMBC [WP]; PMBC Sardinella fimbriata (Valenciennes, 1847): OShr; [EIO-IA]; Saurida gracilis (Quoy & Gaimard, 1824): Cr, OShr; [IP]; PMBC PMBC Sardinella gibbosa (Bleeker, 1849): OShr; [IWP]; PMBC Saurida longimanus Norman, 1939: OShr; [NIO]; PMBC Sardinella lemura Bleeker, 1853: OShr; [WP] Saurida micropectoralis Shindo & Yamada, 1972: Mg, Spratelloides delicatulus (Bennett, 1831): Cr, OShr; [IP]; OShr; [WP]; PMBC PMBC Saurida nebulosa Valenciennes, 1849: Cr, Mg, Sg, OShr; [IP]; PMBC 51 The fishes of southwestern Thailand

Saurida tumbil (Bloch, 1795): OShr; [IWP]; PMBC Encheliophis homei (Richardson, 1846): Sg; [IWP]; PMBC Saurida undosquamis (Richardson, 1848): Cr, OShr; Onuxodon fowleri (Smith, 1955): Cr; [IP] [IWP]; PMBC Onuxodon parvibranchium (Fowler, 1927): Cr; [IP]; Synodus dermatogenys Fowler, 1912: Cr, OShr; [IP]; PMBC PMBC Pyramodon ventralis Smith & Radcliffe, 1913: OShr; [IP] Synodus hoshinonis Tanaka, 1917: OShr; [IWP]; PMBC Order Ophidiiformes Synodus indicus (Day, 1873): OShr; [IWP]; PMBC Ophidiidae Synodus jaculum Russell & Cressey, 1979: Cr; [IP]; PMBC Glyptophidium argenteum Alcock, 1889: OShr; [IM]; Synodus rubromarmoratus Russell & Cressey, 1979: Cr; PMBC [WP] Glyptophidium macropus Alcock, 1894: OShr; [NIO] Synodus variegatus (Lacepède, 1801): Cr, OShr; [IP]; Hypopleuron caninum Smith & Radcliffe, 1913: OShr; PMBC [IWP]; PMBC Trachinocephalus myops (Bloch & Schneider, 1801): Mg, Neobythites fasciatus Smith & Radcliffe, 1913: OShr; [WP] Sg, OShr; [CIR]; PMBC Bythitidae Chlorophthalmidae Alionematichthys phuketensis Møller & Schwarzhans, Chlorophthalmus corniger Alcock, 1894: OShr; [EIO-WP] 2008: Cr; [AS] Alepisauridae Alionematichthys piger (Alcock, 1890): Cr; [WCP] Alepisaurus ferox Lowe, 1833: OShr; [COS] Alionematichthys riukiuensis (Aoyaki, 1954): Cr; [WP] Paralepididae Brosmophyciops pautzkei Schultz, 1960: Cr; [IWP]; PMBC Lestolepis luetkeni (Ege, 1933): OShr; [IP] Dinematichthys iluocoeteoides Bleeker, 1855: Cr; [IWP]; Maculisudis longipinnis Kotthaus, 1967: OShr PMBC Paralepis philippinus Fowler, 1934: OShr Order Batrachoidiformes Order Myctophiformes Batrachoididae Neoscopelidae Allenbatrachus grunniens (Linnaeus, 1758): Mg, OShr; Neoscopelus macrolepidotus Jonhson, 1863: OShr; [COS] [NIO-IM]; PMBC Neoscopelus microchir Matsubara, 1943: OShr; [CIR]; Order Lophiiformes PMBC Lophiidae Myctophidae Lophiodes lugubris (Alcock, 1894): OShr; [EIO] Benthosema pterotum (Alcock, 1890): OShr; [IWP] Lophiodes mutilus (Alcock, 1893): OShr; [IWP]; PMBC Diaphus coeruleus (Kluzinger, 1871): OShr; [IWP] Antennariidae Diaphus splendidus (Brauer, 1904): OShr; [CIR] Antennarius coccineus (Cuvier, 1831): Cr; [IP] Diaphus thiollierei Fowler, 1934: OShr; [IWP] Antennarius commerson (Latreille, 1804): Cr, OShr; [IP]; Order Lampriformes PMBC Lophotidae Antennarius hispidus (Bloch & Schneider, 1801): Cr, OShr; Lophotus lecepede Giorna, 1809: OShr; [CIR]; PMBC [IWP] Trachipteridae Antennarius maculatus (Desjardins, 1840): Cr; [IP] Trachipterus jacksonensis (Ramsay, 1881): OShr; [COS] Antennarius nummifer (Cuvier, 1817): Cr; [IP]; PMBC Zu cristatus (Bonelli, 1820): OShr; [CIR]; PMBC Antennarius pictus (Shaw & Nodder, 1794): Cr; [IP]; PMBC Order Polymixiiformes Chaunacidae Polymixiidae Chaunax pictus Lowe, 1846: OShr; [COS] Polymixia berndti Gilbert, 1905: OShr; [IP] Ogcocephalidae Order Gadiformes Halieutaea coccinea Alcock, 1889: OShr; [IO] Bregmacerotidae Halieutaea indica Annandale & Jenkins, 1910: OShr; [IWP] Bregmaceros mcclellandi Thompson, 1840: OShr; [EIO]; Halieutaea stellata (Vahl, 1797): OShr; [WP]; PMBC PMBC Halieutopsis micropa (Alcock, 1891): OShr; [IWP] Macrouridae Order Mugiliformes Coelorinchus argentatus Smith & Radcliffe, 1912: OShr; Mugilidae [IWP] Chelon macrolepis (Smith, 1846): Mg; [IP]; PMBC Coelorinchus macrorhynchus Smith & Radcliffe, 1912: Chelon parsia (Hamilton, 1822): Mg; [EIO] OShr; [WP] Chelon subviridis (Valenciennes, 1836): Mg, Sg; [IWP]; Coelorinchus radcliffei Gilbert & Hubbs, 1920: OShr; [WP] PMBC Coryphaenoides asprellus (Smith & Radcliff, 1912): OShr; Crenimugil crenilabris (Forsskål, 1775): Cr; [IP] [IO]; PMBC Ellochelon vaigiensis (Quoy & Gaimard, 1825): Cr, Mg, Malacocephalus laevis (Lowe, 1843): OShr; [COS] Sg; [IP]; PMBC Moridae Moolgarda cunnesius (Valenciennes, 1836): Mg, Sg; [IWP]; Physiculus roseus Alcock, 1891: OShr; [EIO-WP] PMBC Carapidae Moolgarda engeli (Bleeker, 1859): Mg; [IP] 52 Phuket mar. biol. Cent. Res. Bull.

Moolgarda pedaraki (Valenciennes, 1836): Mg, Sg; [IWP]; Belonidae PMBC Ablennes hians (Valenciennes, 1846): Cr, OShr; [COS]; Moolgarda perusii (Valenciennes, 1836): Mg, Sg; [IWP]; PMBC PMBC Strongylura incisa (Valenciennes, 1846): OShr; [EIO-WP]; Moolgarda seheli (Forsskål, 1775): Cr, Mg; [IP]; PMBC PMBC Oedalechilus labiosus (Valenciennes, 1836): Cr; [IP]; Strongylura leiura (Bleeker, 1850): OShr; [IWP]; PMBC PMBC Strongylura strongylura (van Hasselt, 1823): Mg, Sg, OShr; Paramugil parmatus (Cantor, 1849): Mg; [IM]; PMBC [NIO-IA]; PMBC Order Atheriniformes Tylosurus acus melanotus (Bleeker, 1850): Cr, OShr; [IP] Phallostethidae Tylosurus crocodilus crocodilus (Peron & Lesueur, 1821): Neostethus bicornis Regan, 1916: Mg; [IM] Cr,Mg,Sg,OShr;[CIR] Neostethus lankesteri Regan, 1916: Mg; [IM]; PMBC Order Cyprinodontiformes Phenacostethus posthon Roberts, 1971: Mg; [AS] Aplocheilidae Atherinidae Aplocheilus panchax Day, 1875: Mg; [EIO-IM]; PMBC Atherinomorus duodecimalis (Valenciennes, 1835): Cr, Mg, Order Beryciformes Sg, OShr; [IP]; PMBC Diretmidae Atherinomorus lacunosus (Forster & Schneider, 1801): Diretmoides veriginae Kotlyar, 1987: OShr; [EIO-WP]; OShr; [EIO-WP]; PMBC PMBC Atherinomorus pinguis (Lacepède, 1803): Sg, OShr; [IWP] Monocentridae Atherion elymus Jordan & Starks, 1901: Cr, OShr; [WP]; Monocentris japonica (Houttuyn, 1782): OShr; [IWP]; PMBC PMBC Hypoatherina temminckii (Bleeker, 1853): Cr, OShr [IP]; Trachichthyidae PMBC Gephyroberyx darwinii (Johnson, 1866): OShr; [COS]; Hypoatherina valenciennei (Bleeker, 1853): Mg, Sg, OShr; PMBC [WP]; PMBC Berycidae Order Beloniformes Beryx splendens Lowe, 1834: OShr; [COS] Andrianichthyidae Centroberyx druzhinini (Busakhin, 1981): OShr; [IWP]; Oryzias javanicus (Bleeker, 1854): Mg; [IM]; PMBC PMBC Exocoetidae Holocentridae Cheilopogon arcticeps (Günther, 1866): OShr; [WP] Myripristis adustus (Bleeker, 1853): Cr, OShr; [IP] Cheilopogon spilopterus (Valenciennes, 1846): OShr; [WP]; Myripristis berndti (Jordan & Evermann, 1903): Cr, OShr; PMBC [IP]; PMBC Exocoetus volitans (Linnaeus, 1758): OShr; [COS]; PMBC Myripristis botche Cuvier, 1829: Cr, OShr; [IWP]; PMBC Hirundichthys speculiger (Valenciennes, 1847): OShr; [CIR] Myripristis chryseres Jordan & Evermann, 1903: Cr, OShr; Parexocoetus brachypterus (Richardson, 1846): OShr; [IP]; [IP] PMBC Myripristis hexagona (Lacepède, 1802): Cr, OShr; [IWP]; Hemiramphidae PMBC Dermogenys siamensis Fowler, 1934: Mg; [IM] Myripristis kuntee Cuvier, 1831: Cr, [IP] Hemiramphus far (Forsskål, 1775): Cr, Sg, OShr; [IWP]; Myripristis murdjan (Forsskål, 1775): Cr; [IP] PMBC Myripristis violacea Bleeker, 1851: Cr; [IP] Hyporhamphus balinensis (Bleeker, 1858): OShr; [IWP] Myripristis vittatus Valenciennes, 1831: Cr; [IP] Hyporhamphus dussumieri (Valenciennes, 1846): Sg, OShr; Neoniphon sammara (Forsskål, 1775): Cr, OShr; [IP]; [IP]; PMBC PMBC Hyporhamphus limbatus (Valenciennes, 1846): Mg, Sg, Ostichthys japonicus (Cuvier, 1829): OShr; [WP]; PMBC OShr; [NIO-WP] Sargocentron caudimaculatum (Rüppell, 1835): Cr; [IP]; Hyporhamphus melanopterus Collette & Parin, 1978: Mg, PMBC OShr; [IM]; PMBC Sargocentron cornutum (Bleeker, 1853): Cr; [WP]; PMBC Hyporhamphus quoyi Valenciennes, 1847: Mg, Sg, OShr; Sargocentron diadema (Lacepède, 1802): Cr; [IP] [IWP]; PMBC Sargocentron melanospilos (Bleeker, 1858): Cr; [IWP] Hyporhamphus unicuspis Collette & Parin, 1978: OShr; Sargocentron rubrum (Forsskål, 1775): Cr, OShr; [IWP]; [NIO] PMBC Zenarchopterus buffornis (Valenciennes, 1847): Mg, Sg, Sargocentron spiniferum (Forsskål, 1775): Cr; [IP] OShr; [EIO-IA]; PMBC Order Zeiformes Zenarchopterus dispar (Valenciennes, 1847): Mg, Sg, OShr; Grammicolepididae [IWP]; PMBC Xenolepidichthys dalgleishi Gilchrist, 1922: OShr; [CIR] Zenarchopterus dunckeri Mohr, 1926: Mg, Sg, OShr; [WP]; Zeidae PMBC Cyttopsis rosea (Lowe, 1843): OShr; [COS]; PMBC 53 The fishes of southwestern Thailand

Zenopsis nebulosa (Temminck & Schlegel, 1845): OShr; Trachyrhamphus serratus (Temminck & Schlegel, 1850): [IP] OShr; [EIO-WP]; PMBC Order Gasterosteiformes Aulostomidae Pegasidae Aulostomus chinensis (Linnaeus, 1766): Cr; [IP]; PMBC Eurypegasus draconis (Linnaeus, 1766): Cr, OShr; [IP]; Fistulariidae PMBC Fistularia commersonii Rüppell, 1835: Cr, Sg; [IP]; PMBC Pegasus volitans Linnaeus, 1758: Cr, Sg, OShr; [IWP]; Fistularia petimba Lacepède, 1803: OShr; [CIR]; PMBC PMBC Centriscidae Solenostomidae Aeoliscus strigatus (Günther, 1860): Cr, Sg, OShr; [IWP] Solenostomus cyanopterus Bleeker, 1852: Cr; [IP] Centriscus cristatus De Vis, 1885: OShr; [WP] Solenostomus paradoxus (Pallas, 1770): Cr, Sg; [IWP]; Centriscus scutatus Linnaeus, 1758: Cr, Sg, OShr; [IWP]; PMBC PMBC Syngnathidae Order Sybranchiformes Bhanotia fasciolata (Dumeril, 1870): Cr, OShr; [IA]; Sybranchidae PMBC Macrotrema caligans (Cantor, 1849): Mg; [IM] Choeroichthys brachysoma (Bleeker, 1855): Cr; [IP]; Order Scorpaeniformes PMBC Dactylopteridae Choeroichthys sculptus (Günther, 1870): Cr, OShr; [IP] Dactyloptena macracantha (Bleeker, 1854): OShr; [EIO- Corythoichthys amplexus Dowson & Randall, 1906: Cr, WP]; PMBC OShr; [IWP] Dactyloptena orientalis (Cuvier, 1829): Cr, OShr; [IP]; Corythoichthys flavofasciatus (Rüppell, 1838): Cr; [IP] PMBC Corythoichthys haematopterus (Bleeker, 1851): Cr, OShr; Dactyloptena perterseni (Nystrom, 1887): OShr; [IWP]; [IWP]; PMBC PMBC Corythoichthys schultzi Herald, 1953: Cr; [IP]; PMBC Scorpaenidae Doryrhamphus janssi (Herald & Randall, 1972): Cr, OShr; Ablabys taenianotus (Cuvier, 1829): Cr; [EIO-WP] [WP]; PMBC Apistus carinatus (Bloch & Schneider, 1801): OShr; [IWP] Dunckerocampus multiannulatus (Regan, 1903): Cr; [IO] Choridactylus multibarbus Richardson, 1848: OShr; [IWP] Halicampus grayi Kaup, 1856: OShr; [NIO-WP]; PMBC Dendrochirus biocellatus (Fowler, 1938): Cr; [IP] Halicampus mataafae (Jordan & Seale, 1906): Cr, OShr; Dendrochirus brachypterus (Cuvier, 1829): Cr, OShr; [IP]; [IP] PMBC Halicampus spinirostris (Dawson & Allen, 1981): Cr; Dendrochirus zebra (Cuvier, 1829): Cr, Sg, OShr; [IWP]; [EIO-WP] PMBC Hippichthys cyanospilus (Bleeker,1854):Sg,OShr;[IWP]; Inimicus cuvieri (Gray, 1832): Cr, OShr; [IM] PMBC Lioscorpius longiceps Günther, 1880: OShr; [WP] Hippichthys heptagonus (Bleeker, 1853): Sg, OShr; [IWP]; Minous coccineus Alcock, 1890: OShr; [IWP] PMBC Minous inermis Alcock, 1889: OShr; [IO] Hippichthys penicillus (Cantor, 1849): Mg, Sg, OShr; [NIO- Minous monodactylus (Bloch & Schneider, 1801): OShr; WP]; PMBC [IWP]; PMBC Hippichthys spicifer (Rüppell, 1840): Mg, OShr; [IP]; Minous trachycephalus (Bleeker, 1854): OShr; [EIO-WP] PMBC Neomerinthe amplisquamiceps (Fowler, 1938): OShr; [WP] Hippocampus comes Cantor, 1850: Cr, Sg; [IM]; PMBC Paracentropogon longispinis (Bleeker, 1829): Cr, OShr; Hippocampus kelloggi Jordan & Snyder, 1901: OShr; [EIO-WP] [IWP]; PMBC Parapterois heterurus (Bleeker, 1856): OShr; [IWP] Hippocampus kuda Bleeker, 1852: Mg, Sg, OShr;[NIO- Parascorpaena aurita (Rüppell, 1838): Cr, OShr; [IWP]; WCP]; PMBC PMBC Hippocampus spinosissimus Weber, 1913: OShr; [EIO-IA]; Parascorpaena mossambica (Peters, 1855): Cr; [IP] PMBC Pontinus macrocephalus (Sauvage, 1882): OShr; [WP]; Hippocampus trimaculatus Leach, 1814: OShr; [EIO-WP]; PMBC PMBC Pterois antennata (Bloch, 1787): Cr, OShr; [IP]; PMBC Phoxocampus belcheri (Kaup, 1856): Cr, OShr; [IWP] Pterois miles (Bennett, 1828): Cr, Sg, OShr; [IO]; PMBC Syngnathoides biaculeatus (Bloch, 1785): Sg, OShr; [IWP]; Pterois mombasae (Smith, 1957): Cr, OShr; [IO] PMBC Pterois radiata Cuvier, 1829: Cr, OShr; [IP]; PMBC Trachyrhamphus bicoarctatus (Bleeker, 1857): Cr, Sg, Pterois russellii Bennett, 1831: Sg, OShr; [IWP]; PMBC OShr; [IWP]; PMBC Rhinopias eschmeyeri Conde, 1977: Cr; [IWP] Trachyrhamphus longirostris Kaup, 1856: OShr; [IWP]; Rhinopias frondosa (Günther, 1891): Cr, OShr; [IWP] PMBC Richardsonichthys leucogaster (Richardson, 1848): OShr; [IWP] 54 Phuket mar. biol. Cent. Res. Bull.

Scorpaenodes albaiensis (Evermann & Seale, 1907): Cr; Bembridae [IWP]; PMBC Brachybembras sp.: OShr Scorpaenodes guamensis (Quoy & Gaimard, 1824): Cr, Plathycephalidae OShr; [IP]; PMBC Cociella punctata (Cuvier, 1829): Cr, Mg, Sg, OShr; [IWP]; Scorpaenodes hirsutus (Smith, 1957): Cr; [IP] PMBC Scorpaenodes minor (Smith, 1958): Cr; [IP] Elates ransonneti (Steindachner, 1877): OShr; [IA]; PMBC Scorpaenodes parvipinnis (Garrett, 1863): Cr, OShr; [IP]; Grammoplites scaber (Linnaeus, 1758): Mg, Sg, OShr; PMBC [NIO-IM]; PMBC Scorpaenodes scaber (Ramsay & Ogilby, 1886): Cr; [EIO- Inegocia japonica (Tilesius, 1812): Mg, Sg, OShr; [EIO- WCP] WP]; PMBC Scorpaenodes varipinnis Smith, 1957: Cr; [IWP] Kumococius rodericensis Cuvier in Cuvier & Valenciennes, Scorpaenopsis diabolus Cuvier, 1829: Cr, OShr; [IP]; 1829: OShr; [NIO-IA]; PMBC PMBC Platycephalus indicus (Linnaeus, 1758): Mg, Sg, OShr; Scorpaenopsis longispina Randall & Eschmeyer, 2001: Cr; [IWP]; PMBC] [IO] Rogadias pristiger (Cuvier, 1829): OShr; [IWP]; PMBC Scorpaenopsis neglecta Heckel, 1837: Sg, OShr; [EIO-WP]; Rogadias serratus (Cuvier, 1829): OShr; [IWP]; PMBC PMBC Sorsogona melanoptera (Knapp & Wongratana, 1987); Scorpaenopsis oxycephala (Bleeker, 1849): Cr, OShr; OShr; [NIO] [IWP]; PMBC Sorsogona tuberculata (Cuvier, 1829): OShr; [NIO-IA]; Scorpaenopsis possi Randall & Eschmeyer, 2001: Cr; [IP] PMBC Scorpaenopsis ramaraoi Randall & Eschmeyer, 2001: Cr, Suggrundus macracanthus (Bleeker, 1869): OShr; [EIO- OShr; [NIO-WP] IA]; PMBC Scorpaenopsis vesosa (Cuvier, 1829): OShr; [IWP] Sunagocia carbunculus (Valenciennes, 1833): Mg, Sg, OShr; Sebastapistes fowleri (Pietschmann, 1934): Cr; [IP] [EIO-IM]; PMBC Sebastapistes mauritiana (Cuvier, 1829): Cr, OShr; [IP] Sunagocia otaitensis (Cuvier, 1829): Cr, OShr; [IP]; PMBC Sebastapistes strongia (Cuvier, 1829): Cr; [IP] Thrysanophrys celebicus (Bleeker, 1854): Cr; [IWP] Setarches guentheri Johnson, 1862: OShr; [CIR] Thrysanophrys chitonae Schultz, 1966: Cr, OShr; [IP] Synanceia horrida (Linnaeus, 1766): Cr, OShr; [EIO-WP]; Hoplichthyidae PMBC Hoplichthys citrinus Gilbert, 1905: OShr; [WP] Synanceia verrucosa (Smith, 1956): Cr, OShr; [IP] Order Perciformes Taenianotus triacanthus Lacepède, 1802: Cr; [IP] Ambassidae Trachicephalus uranoscopus (Bloch & Schneider, 1801): Ambassis buruensis Bleeker, 1857: Mg; [WP]; PMBC Mg, Sg, OShr; [EIO-WP], PMBC Ambassis dussumieri Cuveir, 1828: Sg; [IWP] Vespicula trachinoides (Cuvier & Valenciennes, 1829): Mg, Ambassis gymnocephalus (Lacepède, 1802): Mg; [IWP]; Sg, OShr; [WP]; PMBC PMBC Caracanthidae Ambassis interruptus Bleeker, 1852: Mg, Sg; [IA]; PMBC Caracanthus unipinna (Gray, 1831): Cr; [IWP]; PMBC Ambassis macracanthus Bleeker, 1849: Mg, Sg; [EIO-IM]; Aploactinidae PMBC Acanthosphex leurynnis (Jordan & Seale, 1905): Mg, Sg, Ambassis nalua (Hamilton, 1822): Mg, Sg; [EIO-IA]; OShr; [EIO-IM]; PMBC PMBC Cocotropus echinatus (Cantor, 1850): Mg, Sg; [AS] Ambassis urotaenia Bleeker, 1852: Mg, Sg; [IWP]; PMBC Cocotropus larvatus Poss & Allen, 1987: OShr; [WP] Ambassis vachellii Richardson, 1846: Mg, Sg; [IWP]; Paraploactis obbesi (Weber, 1913): Cr, OShr; [WP] PMBC Sthenopus mollis Richardson, 1848: OShr; [WP] Latidae Xenaploactis cautes Poss & Eschmeyer, 1980: OShr; [IM Lates calcalifer (Bloch, 1790): Mg; [NIO-WP]; PMBC (presently known from AS and Gulf of Thailand)] Acropomatidae Triglidae Acropoma argentistigma Okamoto & Ida, 2002: OShr; [AS] Lepidotrigla punctipectoralis Fowler, 1938: OShr; [IWP] Synagrops japonicus (Döderlein, 1883): OShr; [IP]; PMBC Lepidotrigla spiloptera Günther, 1880: OShr; [IWP]; Synagrops malayanus Weber, 1913: OShr; [WCP] PMBC Synagrops philippinensis (Günther, 1880): OShr; [EIO- Peristediidae WP]; PMBC Peristedion liorhynchus Günther, 1871: OShr; [IWP]; Serranidae PMBC Aethaloperca rogaa (Forsskål, 1775): Cr, OShr; [IP]; Peristedion moluccense Bleeker, 1850: OShr; [WP]; PMBC PMBC Satyrichthys adeni (Lloyd, 1907): OShr; [IWP]; PMBC Anyperodon leucogrammicus (Valnciennes, 1828): Cr, OShr; Satyrichthys sp.: OShr; PMBC [IP]; PMBC 55 The fishes of southwestern Thailand

Cephalopholis argus Bloch & Schneider, 1801: Cr, OShr; Epinephelus octofasciatus Griffin, 1926: OShr; [IP]; PMBC [IP]; PMBC Epinephelus ongus (Bloch, 1790): Cr, OShr; [IWP]; PMBC Cephalopholis aurantia (Valenciennes, 1828): OShr; [IWP] Epinephelus polyphekadion (Bleeker, 1849): Cr, OShr; [IP]; Cephalopholis boenak (Bloch, 1790): Cr, Sg, OShr; [IWP]; PMBC PMBC Epinephelus quoyanus (Valenciennes, 1830): Cr, Sg, OShr; Cephalopholis formosa (Shaw & Nodder, 1812): Cr, OShr; [WP]; PMBC [EIO-WP]; PMBC Epinephelus radiatus (Day, 1867): OShr; [IWP]; PMBC Cephalopholis leopardus (Lacepède, 1801): Cr; [IP] Epinephelus sexfasciatus (Valenciennes, 1828): Mg, Sg, Cephalopholis miniata (Forsskål, 1775): Cr, OShr; [IP]; OShr; [IA]; PMBC PMBC Epinephelus spilotoceps Schultz, 1953: Cr; [IP] Cephalopholis polyspila Randall & Satapoomin, 2000: Cr, Epinephelus tauvina (Forsskål, 1775): Cr, OShr; [IP]; OShr; [AS]; PMBC PMBC Cephalopholis sexmaculata (Rüppell, 1830): Cr; [IP] Epinephelus tukula Morgan, 1959: Cr, OShr; [IWP] Cephalopholis sonnerati (Valenciennes, 1828): Cr, OShr; Epinephelus undulosus (Quoy & Gaimard, 1824): Cr, OShr; [IP]; PMBC [IWP]; PMBC Cephalopholis urodeta (Bloch & Schneider, 1801): Cr, Grammistes sexlineatus (Thunberg, 1792): Cr; [IP] OShr; [IP]; PMBC Grammistops ocellatus Schultz, 1953: Cr; [IP] Chelidoperca margaritifera Weber, 1913: OShr; [WP]; Liopropoma africanum (Smith, 1954): Cr; [IO]; PMBC PMBC Liopropoma susumi (Jordan & Seale, 1906): Cr; [IP]; Cromileptes altivelis (Valenciennes, 1828): Cr, OShr; [WP]; PMBC PMBC Meganthias filiferus Randall & Heemstra, 2007: OShr; [AS]; Diploprion bifasciatum Cuvier, 1828: Cr, OShr; [EIO-WP]; PMBC PMBC Plectropomus areolatus (Rüppell, 1830): Cr, OShr; [IP] Epinephelus amblycephalus (Bleeker, 1857): OShr; [WP]; Plectropomus laevis (Lacepède, 1801): Cr, OShr; [IP] PMBC Plectropomus pessuliferus Fowler, 1901: Cr, OShr; [IWP] Epinephelus areolatus (Forsskål, 1775): Cr, OShr; [IWP]; Pogonoperca ocellata Günther, 1859: Cr, OShr; [IO]; PMBC PMBC Epinephelus bleekeri (Vaillant, 1877): Cr, Mg, Sg, OShr; Pseudanthias evansi (Smith, 1954): Cr; [IO]; PMBC [NIO-IA]; PMBC Pseudanthias ignitus (Randall & Lubbock, 1981): Cr; [EIO] Epinephelus caeruleopunctatus (Bloch, 1790): Cr, OShr; Pseudanthias rubrizonatus (Randall, 1983): Cr; [WP] [IWP]; PMBC Pseudanthias squamipinnis (Peters, 1855): Cr; [IWP]; Epinephelus coioides (Hamilton, 1822): Cr, Mg, Sg, OShr: PMBC [IWP]; PMBC Pseudogramma polyacanthum (Bleeker, 1856): Cr; [IP]; Epinephelus epistictus (Temminck & Schlegel, 1842): OShr; PMBC [IWP]; PMBC Rabaulichthys stigmaticus Randall & Pyle, 1989: Cr, OShr; Epinephelus erythrurus (Valenciennes, 1828): Cr, Sg, OShr; [IO] [NIO-IM]; PMBC Variola albimarginata Baissac, 1953: Cr, OShr; [IWP] Epinephelus fasciatus (Forsskål, 1775): Cr, OShr; [IP]; Variola louti (Forsskål, 1775): Cr, OShr; [IP]; PMBC PMBC Ostracoberycidae Epinephelus flavocaeruleus (Lacepède, 1801): Cr, OShr; Ostracoberyx dorygenys Fowler, 1934: OShr; [IWP] [IO] Pseudochromidae Epinephelus fuscoguttatus (Forsskål, 1775): Cr, OShr; [IP]; Amsichthys knighti (Allen, 1987): Cr; [WP] PMBC Blennodesmus scapularis Günther, 1871: Cr; [IA]; PMBC Epinephelus heniochus Fowler, 1904: OShr; [WP] Pseudochromis andamanensis Lubbock, 1980: Cr; [EEIO]; Epinephelus hexagonatus (Bloch & Schneider, 1801): Cr; PMBC [IP] Pseudochromis caudalis Boulenger, 1898: Cr; [IO] Epinephelus lanceolatus (Bloch, 1790): Cr, OShr; [IP]; Pseudochromis coccinicauda Tickell in Day, 1888: Cr; PMBC [EIO] Epinephelus latifasciatus (Temminck & Schlegel, 1842): Pseudochromis fuscus Müller & Troschel, 1849: Cr; [EIO- OShr; [IWP] IA]; PMBC Epinephelus longispinis (Kner, 1864): Cr, OShr; [IO] Pseudochromis tapeinosoma Bleeker, 1853: Cr; [IA]; Epinephelus macrospilos (Bleeker, 1855): Cr; [IP]; PMBC PMBC Epinephelus malabaricus (Bloch & Schneider, 1801): Cr, Pseudoplesiops rosae Schultz, 1953: Cr; [EIO-WCP]; Mg, OShr; [IWP]; PMBC PMBC Epinephelus merra Bloch, 1793: Cr, OShr; [IP]; PMBC Plesiopidae Epinephelus morrhua (Valenciennes, 1833): OShr; [IP]; Calloplesiops altivelis (Steindachner, 1903); Cr; [IP] PMBC Plesiops auritus Mooi, 1995: Cr; [EIO] 56 Phuket mar. biol. Cent. Res. Bull.

Plesiops caeruleolineatus Rüppell, 1835: Cr; [IP]; PMBC Apogon striatodes Gon, 1996: OShr; [IM]; PMBC Plesiops corallicola Bleeker, 1853: Cr; [WCP]; PMBC Apogon striatus (Smith & Radcliffe, 1912): OShr; [NIO- Opistognathidae WP]; PMBC Opistognathus cyanospilotus Smith-Vaniz, 2009: Cr; Apogon taeniophorus Regan, 1908: Cr; [IP]; PMBC [EEIO] Apogon talboti Smith, 1961: Cr; [IWP] Opistognathus nigromarginatus Rüppell, 1830: Cr, OShr; Apogon thermalis Cuvier, 1829: Cr, Sg; [IWP]; PMBC [IWP]; PMBC Apogon trimaculatus Cuvier, 1828: Cr, OShr; [WP]; PMBC Opistognathus rosenbergii (Bleeker, 1856): Cr, OShr; Apogon truncatus Bleeker, 1854: OShr; [IWP]; PMBC [EIO-WP]; PMBC Apogonichthyoides nigripinnis (Cuvier, 1828): OShr; [EIO- Opistognathus variabilis Smith-Vaniz,2009: Cr; [EIO-WP]; IA]; PMBC PMBC Apogonichthyoides sialis (Jordan & Thompson, 1914): Cr; Priacanthidae [EIO-WP] Heteropriacanthus cruentatus (Lacepède, 1801): Cr, OShr; Apogonichthys sp.: OShr [COS]; PMBC Archamia ataenia Randall & Satapoomin, 1999: Cr; [AS]; Priacanthus blochii Bleeker, 1853: OShr; [IWP]; PMBC PMBC Priacanthus hamrur (Forsskål, 1775): Cr, OShr; [IP]; Archamia bleekeri Günther, 1859: Cr; [IWP] PMBC Archamia fucata (Cantor, 1850): Cr, Sg, OShr; [IP]; PMBC Priacanthus macracanthus Cuvier, 1829: Cr, OShr; [WP]; Archamia macroptera (Cuvier, 1828): Cr; [EIO-WP] PMBC Cheilodipterus artus Smith, 1961: Cr; [IP]; PMBC Priacanthus sagittarius Starnes, 1988: OShr; [IWP] Cheilodipterus macrodon (Lacepède, 1802): Cr; [IP]; Priacanthus tayenus Richardson, 1846: OShr; [NIO-WP]; PMBC PMBC Cheilodipterus quinquelineatus Cuvier, 1828: Cr; [IP]; Pristigenys niphonia (Cuvier, 1829): OShr; [IWP] PMBC Apogonidae Foa fo (Jordan & Seale, 1906): Sg, OShr; [IWP]; PMBC Apogon abrogramma Fraser & Lachner, 1985: Cr; [IWP] Fowleria aurita (Valenciennes, 1831): Cr; [IWP]; PMBC Apogon angustatus (Smith & Radcliffe, 1911): Cr; [IP]; Fowleria isostigma (Jordan & Seale, 1960): Cr; [IWP]; PMBC PMBC Apogon apogonides (Bleeker, 1856): Cr; [IWP] Fowleria vaiulae (Jordan & Seale, 1906): Cr; [IWP]; Apogon aureus (Lacepède, 1802): Cr, OShr; [IWP] PMBC Apogon bryx Fraser, 1998: OShr; [IWP] Fowleria variegata (Valenciennes, 1832): Cr, Mg, Sg; Apogon campbelli Smith, 1949: Cr; [IO] [IWP]; PMBC Apogon compressus (Smith & Radcliffe, 1911): Cr; [WP]; Gymnapogon urospilotus Lachner, 1953: Cr; [WCP] PMBC Neamia octospina Smith & Radcliffe, 1912: Cr; [IWP] Apogon cookii Macleay, 1881: Cr; [IWP]; PMBC Nectamia fusca (Quoy & Gaimard, 1824): Cr; [IWP]; Apogon cyanosoma Bleeker, 1853: Cr; [IWP]; PMBC PMBC Apogon endekataenia Bleeker, 1852: Cr; [IA]; PMBC Nectamia luxuria Fraser, 2008: Cr; [EIO-WP]; PMBC Apogon fasciatus (White, 1790): OShr; [IWP]; PMBC Nectamia savayensis (Günther, 1872): Cr; [IP] Apogon fleurieu (Lacepède, 1802): Cr, OShr; [IWP] Pseudamia hayashii Randall, Lachner, & Fraser, 1985: Cr; Apogon frenatus Valenciennes, 1832: Cr; [IP]; PMBC [IP] Apogon hyalosoma Bleeker, 1852: Mg, Sg; [WP]; PMBC Pseudamia sp.1: Cr Apogon kallopterus Bleeker, 1856: Cr; [IP]; PMBC Rhabdamia cypselurus Weber, 1909: Cr; [IWP]; PMBC Apogon kalosoma (Bleeker, 1852): Cr; [EIO-WP] Rhabdamia gracilis (Bleeker, 1856): Cr; [IWP]; PMBC Apogon lateralis Valenciennes, 1832: Cr, Mg, Sg; [IWP]; Siphamia tubifer Weber, 1909: Cr; [WP] PMBC Zoramia leptacantha (Bleeker, 1856): Cr; [IWP]; PMBC Apogon moluccensis Valenciennes, 1832: Cr, Sg; [IA]; Zoramia perlita (Fraser & Lachner, 1985): Cr; [IM] PMBC Sillaginidae Apogon nigrofasciatus Lachner, 1953: Cr; [IP]; PMBC Sillaginopsis panijus (Hamilton, 1822): OShr; [EIO]; Apogon novaeguinae Valenciennes, 1832: Mg, Sg; [IWP]; PMBC PMBC Sillago aeolus Jordan & Evermann, 1902: Cr, Mg, Sg, OShr; Apogon ocellicaudus Allen, Kuiter & Randall, 1994: Cr; [IWP]; PMBC [IA] Sillago chondropus Bleeker, 1849: OShr; [IWP]; PMBC Apogon pleuron Fraser, 2005: Sg, OShr; [EIO-WP]; PMBC Sillago intermedius Wongratana, 1977: OShr; [EIO-IM] Apogon poecilopterus Cuvier, 1828: OShr; [EIO-WP] Sillago lutea McKay, 1985: OShr; [EIO-IA] Apogon semiornatus Peters, 1876: Cr; [IWP] Sillago sihama (Forsskal, 1775): Mg, Sg, OShr; [IWP]; Apogon smithi (Kotthaus, 1970): OShr; [IWP]; PMBC PMBC Apogon sp.1: Cr; [AS] Malacanthidae 57 The fishes of southwestern Thailand

Branchiostegus sawakinensis Amirthalingam, 1969: OShr; Carangoides praeustus (Bennett, 1830): Mg, Sg, OShr; [IWP]; PMBC [NIO-IM]; PMBC Hoplolatilus cuniculus Randall & Dooley, 1974: Cr; [IP] Carangoides talamparoides Bleeker, 1852: OShr; [NIO- Hoplolatilus luteus Allen & Kuiter, 1989: Cr, OShr; [EEIO]; IA] PMBC Caranx heberi (Bennett, 1830): Cr, OShr; [NIO-WP] Malacanthus brevirostris Guichenot, 1848: Cr, OShr; [IP]; Caranx ignobilis (Forsskål, 1775): Cr, Mg, Sg, OShr; [IP]; PMBC PMBC Malacanthus latovittatus (Lacepède, 1801): Cr; [IP] Caranx melampygus Cuvier, 1833: Cr, OShr; [IP]; PMBC Lactariidae Caranx papuensis Alleyne & Macleay, 1877: Cr, OShr; Lactarius lactarius (Bloch & Schneider, 1801): OShr; [IP] [NIO-WP]; PMBC Caranx sexfasciatus Quoy & Gaimard, 1824: Cr, Mg, Sg, Coryphaenidae OShr; [IP]; PMBC Coryphaena equiselis Linnaeus, 1758: OShr; [COS]; PMBC Decapterus kurroides Bleeker, 1855: OShr; [IWP]; PMBC Coryphaena hippurus Linnaeus, 1758: OShr; [COS]; Decapterus macarellus (Cuvier, 1833): OShr; [CIR]; PMBC PMBC Rachycentridae Decapterus macrosoma Bleeker, 1851: Cr, OShr; [IP]; Rachycentron canadum (Linnaeus, 1766): Cr, Sg, OShr; PMBC [CIR]; PMBC Decapterus maruadsi (Temminck & Schlegel, 1843): OShr; Echeneidae [WP]; PMBC Echeneis naucrates Linnaeus, 1758: Cr, OShr; [COS]; Decapterus russelli (Rüppell, 1830): Cr, OShr; [IWP]; PMBC PMBC Phtheirichthys lineatus (Menzies, 1791): OShr; [COS] Elagatis bipinnulata (Quoy & Gaimard, 1825): Cr, OShr; Remora remora (Linnaeus, 1758): OShr; [COS] [CIR]; PMBC Carangidae Gnathanodon speciosus (Forsskål, 1775): Cr, OShr; [IP]; Alectis ciliaris (Bloch, 1788): Cr, OShr; [CIR]; PMBC PMBC Alectis indica (Rüppell, 1830): Cr, Sg, OShr; [IWP]; PMBC Megalaspis cordyla (Linnaeus, 1758): Cr, OShr; [IP]; Alepes djedaba (Forsskål, 1775): Mg, OShr; [IWP]; PMBC PMBC Alepes kleinii (Bloch, 1793): Mg, Sg, OShr; [NIO-WP]; Naucrates ductor (Linnaeus, 1758): OShr; [CIR]; PMBC PMBC Parastromateus niger (Bloch, 1795): OShr; [IWP]; PMBC Alepes melanoptera (Swainson, 1839): OShr; [NIO-IM]; Scomberoides commersonianus Lacepède, 1801: Cr, Mg, PMBC Sg, OShr; [IWP]; PMBC Alepes vari (Cuvier, 1833): Cr, Mg, OShr; [IWP] Scomberoides lysan (Forsskål, 1775): Cr, Mg, Sg, OShr; Atropus atropus (Schneider, 1801): OShr; [NIO-WP] [IP]; PMBC Atule mate (Cuvier, 1833): Cr, Mg, Sg, OShr; [IP]; PMBC Scomberoides tala (Cuvier, 1832): OShr; [EIO-WP]; Carangoides armatus (Rüppell, 1830): Mg, Sg, OShr; PMBC [IWP]; PMBC Scomberoides tol (Cuvier, 1832): Cr, Mg, Sg, OShr; [IWP]; Carangoides caeruleopinnatus (Rüppell, 1830): Cr, Sg, PMBC OShr; [IWP]; PMBC Selar boops (Cuvier, 1833): Cr, OShr; [IWP]; PMBC Carangoides chrysophrys (Cuvier, 1833): OShr; [IWP] Selar crumenophthalmus (Bloch, 1793): Cr, OShr; [CIR]; Carangoides equula (Temminck & Schlegel, 1844): Cr, PMBC OShr; [IP] Selaroides leptolepis (Cuvier, 1833): Cr, Sg, OShr; [NIO- Carangoides ferdua (Forsskål, 1775): Cr, OShr; [IP]; WP]; PMBC PMBC Seriola rivoliana Valenciennes, 1833: OShr; [CIR]; PMBC Carangoides fulvoguttatus (Forsskål, 1775): OShr; [IWP]; Seriolina nigrofasciata (Rüppell, 1829): Cr, OShr; [IWP]; PMBC PMBC Carangoides gymnostethus (Cuvier, 1833): OShr; [IWP]; Trachinotus bailloni (Lacepède, 1801): Cr, OShr; [IP]; PMBC PMBC Carangoides hedlandensis (Whitley, 1933): Cr, OShr; Trachinotus blochii (Lacepède, 1801): Cr, OShr; [IP] [IWP]; PMBC Trachinotus botla (Shaw, 1803): OShr; [IO]; PMBC Carangoides malabaricus (Bloch & Schneider, 1801): Ulua mentalis (Cuvier, 1833): Cr, OShr; [IWP]; PMBC OShr; [IWP]; PMBC Uraspis helvola (Forster, 1801): OShr; [IP]; PMBC Carangoides oblongus (Cuvier, 1833): OShr; [IWP]; Uraspis uraspis (Günther, 1860): OShr; [IWP]; PMBC PMBC Menidae Carangoides orthogrammus (Jordan & Gilbert, 1882): Cr, Mene maculata (Bloch & Schneider, 1801): OShr; [IWP]; OShr; [IP] PMBC Carangoides plagiotaenia Bleeker, 1857: Cr, OShr; [IP]; Leiognathidae PMBC Equulites berbis (Valenciennes, 1835): OShr; [IWP]; PMBC 58 Phuket mar. biol. Cent. Res. Bull.

Equulites elongatus (Günther, 1860): OShr; [IWP]; PMBC Lutjanus fulvus (Schneider, 1801): Cr, OShr; [IP]; PMBC Equulites leuciscus (Günther, 1860): Mg, Sg, OShr; [IWP]; Lutjanus gibbus (Forsskål, 1775): Cr, OShr; [IP]; PMBC PMBC Lutjanus johnii (Bloch, 1792): Cr, Mg, Sg, OShr; [IWP]; Equulites oblongus (Valenciennes,1835): Mg, OShr; [IWP]; PMBC PMBC Lutjanus kasmira (Forsskål, 1775): Cr, OShr; [IP]; PMBC Equulites stercorarius (Everman & Seale, 1907): Mg, Sg, Lutjanus lemniscatus (Valenciennes, 1828): Cr, Mg, OShr; OShr; [WP]; PMBC [EIO-IA]; PMBC Eubleekeria jonesi (James, 1971): Mg, Sg, OShr; [EIO- Lutjanus lunulatus (Park, 1797): Cr, Mg, Sg; [NIO-IA]; WP]; PMBC PMBC Gazza dentex (Valenciennes, 1835): OShr; [IO]; PMBC Lutjanus lutjanus Bloch, 1790: Cr, OShr; [IWP]; PMBC Gazza minuta (Bloch, 1797): Mg, Sg, OShr; [IWP]; PMBC Lutjanus madras (Valenciennes, 1831): Cr, Mg, OShr; Gazza rhombea Kimura, Yamashita & Iwatsuki, 2000: [IWP]; PMBC OShr; [EIO-WP]; PMBC Lutjanus malabaricus Schneider, 1801: Cr, OShr; [NIO- Karalla daura (Cuvier, 1829): Mg, OShr; [IWP]; PMBC WP]; PMBC Leiognathus equulus (Forsskål, 1775): Mg, Sg, OShr; Lutjanus monostigma (Cuvier, 1828): Cr, OShr; [IP]; [IWP]; PMBC PMBC Leiognathus fasciatus (Lacepède, 1803): Mg, OShr; [IWP]; Lutjanus quinquelineatus Bloch, 1790: Cr, OShr; [NIO- PMBC WP]; PMBC Leiognathus longispinis (Valenciennes, 1835): Mg, Sg, Lutjanus rivulatus (Cuvier, 1828): Cr, OShr; [IP]; PMBC OShr; [EIO-IA]; PMBC Lutjanus rufolineatus (Valenciennes, 1830): OShr; [WP] Nuchequula blochii (Valenciennes, 1835): Mg, OShr; [NIO- Lutjanus russelli (Bleeker, 1849): Cr, Mg, Sg, OShr; [IWP]; WP]; PMBC PMBC Nuchequula gerroides (Bleeker, 1851): Mg, Sg, OShr; [EIO- Lutjanus sebae (Cuvier, 1828): Cr, OShr; [IWP]; PMBC WP]; PMBC Lutjanus timorensis (Quoy & Gaimard, 1824): OShr; [WP] Photopectoralis bindus (Valenciennes, 1835): Mg, OShr; Lutjanus vitta (Quoy & Gaimard, 1824): Cr, Sg, OShr; [IWP]; PMBC [IWP]; PMBC Secutor hanedai Moshizuki & Hayashi, 1989: Mg, Sg, Macolor macularis Fowler, 1931: Cr, OShr; [EIO-WP] OShr; [IM]; PMBC Macolor niger (Forsskål, 1775): Cr, OShr; [IP]; PMBC Secutor indicius Monkolprasit, 1973: OShr; [IM]; PMBC Paracaesio xanthurus (Bleeker, 1869): OShr; [INP] Secutor interruptus (Valenciennes, 1835): Mg, OShr; [EIO- Pinjalo lewisi Randall, Allen &Anderson, 1987: Cr; [NIO- WP] WP] Secutor insidiator (Bloch, 1787): Mg, Sg, OShr; [IWP]; Pinjalo pinjalo (Bleeker, 1850): OShr; [IWP]; PMBC PMBC Pristipomoides flavipinnis Shinohara, 1963: OShr; [WCP] Secutor megalolepis Moshizuki & Hayashi, 1989: Mg, Sg, Pristipomoides multidens (Day, 1871): OShr; [IWP]; PMBC OShr; [IA]; PMBC Pristipomoides typus Bleeker, 1852: OShr; [IA]; PMBC Bramidae Symphorichthys spilurus (Günther, 1874): Cr, OShr; [WP] Brama dussumieri Cuvier, 1831: OShr; [CIR]; PMBC Symphorus nematophorus (Bleeker, 1860): Cr, Sg, OShr; Lutjanidae [WP]; PMBC Aphareus furca (Lacepède, 1801): OShr; [IP]; PMBC Caesionidae Aphareus rutilans Cuvier, 1830: Cr, OShr; [IP]; PMBC Caesio caerulaurea Lacepède, 1801: Cr, Mg, Sg, OShr; Aprion virescens Valenciennes, 1830: Cr, OShr; [IP]; [IP]; PMBC PMBC Caesio cuning (Bloch, 1791): Cr, Sg, OShr; [EIO-WP]; Lipocheilus carnolabrum (Chan, 1970): OShr; [NIO-WP]; PMBC PMBC Caesio lunaris Cuvier, 1830: Cr, OShr; [IWP]; PMBC Lutjanus argentimaculatus (Forsskål, 1775): Cr, Mg, Sg, Caesio teres Seale, 1906: Cr, OShr; [IP] OShr; [IP]; PMBC Caesio varilineata Carpenter, 1987: Cr, OShr; [IWP] Lutjanus bengalensis (Bloch, 1790): Cr, OShr; [IO] Caesio xanthonota Bleeker, 1853: Cr, OShr; [IO]; PMBC Lutjanus biguttatus (Valenciennes 1830): Cr, Mg, Sg, OShr; Dipterygonotus balteatus (Valenciennes, 1830): Cr, OShr; [EIO-IA]; PMBC [IWP]; PMBC Lutjanus bohar (Forsskål, 1775): Cr, OShr; [IP]; PMBC Gymnocaesio gymnoptera (Bleeker, 1856): Cr; [IWP]; Lutjanus decussatus (Cuvier, 1828): Cr, OShr; [EIO-IA]; PMBC PMBC Pterocaesio chrysozona (Cuvier, 1830): Cr, OShr; [IWP]; Lutjanus ehrengergii (Peters, 1869): OShr; [IWP] PMBC Lutjanus erythropterus Bloch, 1790: Cr, OShr; [NIO-IA] Pterocaesio pisang (Bloch, 1853): Cr, OShr; [IWP]; PMBC Lutjanus fulviflamma (Forsskål, 1775): Cr, Mg, Sg, OShr; Pterocaesio randalli Carpenter, 1985: Cr; [EIO-IM] [IWP]; PMBC Pterocaesio tessellata Carpenter, 1987: Cr, OShr; [EIO- IA]; PMBC 59 The fishes of southwestern Thailand

Pterocaesio tile (Cuvier, 1830): Cr, OShr; [IP]; PMBC Nemipterus bipunctatus (Ehrenberg, 1830): Cr, OShr; [IO]; Lobotidae PMBC Lobotes surinamensis (Bloch, 1790): Mg, Sg, OShr; [COS]; Nemipterus furcosus (Valenciennes, 1830): Cr, OShr; [EIO- PMBC WP]; PMBC Gerreidae Nemipterus hexodon (Quoy & Gaimard, 1824): OShr; [WP]; Gerres erythrourus (Bleeker, 1791): Mg, Sg, OShr; [EIO- PMBC WP]; PMBC Nemipterus japonicus (Bloch, 1791): OShr; [IWP]; PMBC Gerres filamentosus (Cuvier, 1829): Cr, Mg, Sg, OShr; Nemipterus marginatus (Valenciennes, 1830): OShr; [WP] [IP]; PMBC Nemipterus mesoprion (Bleeker, 1853): OShr; [IM]; Gerres limbatus Cuvier, 1830: Mg, OShr; [EIO-WP]; PMBC PMBC Nemipterus nematophorus (Bleeker, 1853): OShr; [EIO- Gerres longirostris (Lacepède, 1801): Cr, Mg, OShr; [IWP] IM]; PMBC Gerres lucidus (Cuvier, 1830): Cr; [EIO-WP] Nemipterus peronii (Valenciennes, 1830): Cr, Mg, OShr; Gerres macracanthus Bleeker, 1854: Mg, Sg, OShr; [IWP]; [IWP]; PMBC PMBC Nemipterus tambuloides (Bleeker, 1853): OShr; [IM]; Gerres oblongus (Cuvier, 1830): Cr, Mg, Sg, OShr; [IP]; PMBC PMBC Nemipterus thosaporni Russell, 1991: OShr; [IM]; PMBC Gerres oyena (Forsskål, 1775): Cr, Mg, Sg, OShr; [IWP]; Nemipterus zysron (Bleeker, 1856-57): Cr, OShr; [IWP]; PMBC PMBC Gerres phaiya Iwatsuki & Heemstra, 2001: Mg, Sg, OShr; Parascolopsis aspinosa (Rao & Rao, 1981): OShr; [NIO] [EIO]; PMBC Parascolopsis eriomma (Jordan & Richardson, 1909): Gerres setifer (Hamilton, 1822): OShr; [EIO] OShr; [IWP]; PMBC Gerres shima Iwatsuki, Kimura & Yoshino, 2007: OShr; Parascolopsis inermis (Schlegel, 1843): OShr; [EIO-WP]; [WP] PMBC Pentaprion longimanus (Cantor, 1850): OShr; [EIO-WP]; Scolopsis affinis Peters, 1877: Cr, Sg, OShr; [WP]; PMBC PMBC Scolopsis bilineatus (Bloch, 1793): Cr, Sg, OShr; [EIO- Haemulidae WP]; PMBC Diagramma pictum pictum (Thunberg, 1792): Cr, Sg, OShr; Scolopsis ciliatus (Lacepède, 1802): Cr, Mg, Sg, OShr; [WP]; PMBC [WP]; PMBC Hapalogenys merguiensis Iwatsuki, Satapoomin & Scolopsis lineatus Quoy & Gaimard, 1824: Cr, OShr; Amaoka, 2000: OShr; [AS]; PMBC [WCP]; PMBC Plectorhinchus chaetodonoides Lacepède, 1800: Cr, OShr; Scolopsis margaritifer (Cuvier, 1830): Cr, OShr; [WP] [IWP]; PMBC Scolopsis monogramma (Cuvier, 1830): Cr, Sg, OShr; Plectorhinchus gibbosus (Lacepède, 1802): Cr, Mg, Sg, [WP]; PMBC OShr; [IP]; PMBC Scolopsis taeniopterus (Valenciennes, 1830): Mg, Sg, OShr; Plectorhinchus macrospilus Satapoomin & Randall, 2000: [IA]; PMBC Cr; [AS]; PMBC Scolopsis vosmeri (Bloch, 1792): Cr, OShr; [IWP]; PMBC Plectorhinchus vittatus (Linnaeus, 1758): Cr, OShr; [IWP] Scolopsis xenochrous Günther, 1872: Cr, OShr; [EIO-WP] Pomadasys andamanensis Mckay & Satapoomin, 1994: Lethrinidae Cr; [AS]; PMBC Gnathodentex aurolineatus (Lacepède, 1802): Cr; [IP] Pomadasys argenteus (Forsskål, 1775): OShr; [IWP]; Gymnocranius elongatus Senta, 1973: OShr; [IWP] PMBC Gymnocranius grandoculis (Valenciennes,1830): Cr, OShr; Pomadasys argyreus (Valenciennes, 1833): Mg, OShr; [IP]; PMBC [NIO-IM]; PMBC Gymnocranius griseus (Schlegel, 1844): Cr, OShr; [EIO- Pomadasys auritus (Cuvier, 1830): OShr; [IA] WP]; PMBC Pomadasys furcatus (Bloch & Schneider, 1801): OShr; Gymnocranius microdon (Bleeker, 1851): Cr, OShr; [WCP] [IM]; PMBC Lethrinus conchyliatus (Smith, 1959): OShr; [IO] Pomadasys kaakan (Cuvier, 1830): Mg, Sg, OShr; [IWP]; Lethrinus crocineus Smith, 1959: Cr, OShr; [IO]; PMBC PMBC Lethrinus erythracanthus Cuvier, 1830: Cr, OShr; [IP] Pomadasys maculatus (Bloch, 1793): OShr; [IWP]; PMBC Lethrinus erythropterus Valenciennes, 1830: Cr, OShr; Pomadasys olivaceus (Day, 1875): OShr; [IO] [IWP] Pomadasys unimaculatus Tian, 1982: Mg, Sg, OShr; [WP]; Lethrinus harak (Forsskål, 1775): Cr, OShr; [IP]; PMBC PMBC Lethrinus lentjan (Lacepède, 1802): Cr, Sg, OShr; [IWP]; Nemipteridae PMBC Nemipterus balinensoides (Popta, 1918): OShr; [WP] Lethrinus mahsena (Forsskål, 1775): Cr; [IO]; PMBC Nemipterus bathybius Snyder, 1911: OShr; [IA]; PMBC Lethrinus microdon Valenciennes, 1830: Cr, OShr; [IWP] 60 Phuket mar. biol. Cent. Res. Bull.

Lethrinus nebulosus (Forsskål, 1775): Cr, OShr; [IP]; Mulloidichthys flavolineatus (Lacepède, 1801): Cr, Sg, OShr; PMBC [IP] Lethrinus obsoletus (Forsskål, 1775): Cr, OShr; [IP] Mulloidichthys vanicolensis (Valenciennes, 1831): Cr, OShr; Lethrinus olivaceus Valenciennes, 1830: Cr, OShr; [IP] [IP]; PMBC Lethrinus ornatus Valenciennes, 1830: Cr, Sg, OShr; [EIO- Parupeneus barberinus (Lacepède, 1801): Cr, OShr; [IP]; WP]; PMBC PMBC Lethrinus rubrioperculatus Sato, 1978: Cr; [IP]; PMBC Parupeneus cyclostomus (Lacepède, 1801): Cr, OShr; [IP]; Lethrinus variegatus Valenciennes, 1830: OShr; [IWP]; PMBC PMBC Parupeneus heptacanthus (Lacepède, 1801): Cr, Sg, OShr; Lethrinus xanthochilus Klunzinger, 1870: Cr, OShr; [IP] [IWP]; PMBC Monotaxis grandoculis (Forsskål, 1775): Cr, OShr; [IP]; Parupeneus indicus (Shaw, 1803): Cr, Sg, OShr; [IWP]; PMBC PMBC Wattsia mossambrica (Smith, 1957): OShr; [IWP]; PMBC Parupeneus macronema (Lacepède, 1801): Cr, OShr; [IWP]; Sparidae PMBC Acanthopagrus berda (Forsskål, 1775): Mg, OShr; [IWP] Parupeneus pleurostigma (Bennett, 1830): Cr, OShr; [IP]; Argyrops spinifer (Forsskål, 1775): Cr, OShr; [IWP]; PMBC PMBC Polynemidae Parupeneus trifasciatus Lacepède, 1801: Cr, OShr; [IO] Eleutheronema tetradactylum (Shaw, 1804): Mg, OShr; Upeneus asymmetricus Lachner, 1954: OShr; [IWP]; [NIO-IA]; PMBC PMBC Eleutheronema tridactylum (Bleeker, 1849): OShr; [IM] Upeneus guttatus (Day, 1868): OShr; [IWP] Filimanus perplexa Feltes, 1991: OShr; [EEIO]; PMBC Upeneus moluccensis (Bleeker, 1855): OShr; [IWP] Filimanus similis Feltes, 1991: OShr; [NIO] Upeneus sulphureus Cuvier, 1829: Mg, Sg, OShr; [IWP]; Filimanus xanthonema (Valenciennes, 1831): OShr; [EEIO] PMBC Leptomelanosoma indicum (Shaw, 1804): Mg, OShr; [NIO- Upeneus sundaicus (Bleeker, 1855): Mg, OShr; [NIO-IA]; IM]; PMBC PMBC Polydactylus microstomus (Bleeker, 1851): Mg, OShr; [EIO- Upeneus tragula Richardson, 1846: Cr, Mg, Sg, OShr; WP]; PMBC [IWP]; PMBC Polydactylus plebeius (Broussonet, 1782): Mg, OShr; [IWP]; Upeneus vittatus (Forsskål, 1775): OShr; [IP]; PMBC PMBC Pempheridae Polydactylus sexfilis (Valenciennes, 1831): OShr; [IP] Parapriacanthus ransonneti Steindachner, 1870: Cr; [WP]; Polydactylus sextarius (Bloch, 1801): OShr; [EIO-WP] PMBC Polydactylus siamensis Motomura, Iwatsuki & Yoshino, Pempheris mangula Cuvier, 1829: Cr; [IWP]; PMBC 2001: OShr; [IM (presently known from west coast of Pempheris otaitensis Cuvier, 1831: Cr; [IWP] Thailand and Gulf of Thailand)]; PMBC Pempheris vanicolensis Cuvier, 1831: Cr; [IM]; PMBC Sciaenidae Monodactylidae Dendrophysa russelli (Cuvier, 1830): Mg, Sg, OShr; [EIO- Monodactylus argenteus (Linnaeus, 1758): Cr, Mg, OShr; IM]; PMBC [IWP]; PMBC Johnius amblycephalus (Bleeker, 1855): OShr; [NIO-IA]; Toxotidae PMBC Toxotes chatareus (Hamiton, 1822): Mg; [EIO-IA]; PMBC Johnius belangerii (Cuvier, 1830): Mg, OShr; [NIO-IM] Toxotes jaculatrix (Pallas, 1766): Mg; [EIO-WP]; PMBC Johnius borneensis (Bleeker, 1851): OShr; [NIO-IA] Kyphosidae Johnius carutta Bloch, 1793: OShr; [NIO]; PMBC Kyphosus cinerascens (Forsskål, 1775): Cr; [IP]; PMBC Johnius coitor (Hamilton, 1822): OShr: [EIO-IM] Kyphosus vaigiensis (Quoy & Gaimard, 1825): Cr; [IP]; Johnius dussumieri (Cuvier, 1830): OShr; [NIO] PMBC Johnius latifrons Sasaki, 1992: OShr; [IM] Drepaneidae Macrospinosa cuja (Hamilton, 1822): OShr; [EIO] Drepane longimana (Bloch & Schneider, 1801): Mg, Sg, Nibea soldado (Lacepède, 1802): Mg, OShr; [EIO-IA] OShr; [IWP]; PMBC Otolithes cuvieri Trewavas, 1974: Mg, OShr; [IO] Drepane punctata (Linnaeus, 1758): Mg, Sg, OShr; [IWP]; Otolithes ruber (Bloch & Schneider, 1801): OShr; [IWP] PMBC Otolithoides biauritus (Cantor, 1850): OShr; [EIO-IM] Chaetodontidae Panna microdon (Bleeker, 1849): OShr; [IM] Chaetodon andamanensis Kuiter & Debelius, 1999: Cr; Pennahia anea (Bloch, 1793): Mg, Sg, OShr; [NIO-IM]; [EIO]; PMBC PMBC Chaetodon auriga Forsskål, 1775: Cr; [IP]; PMBC Protonibea diacanthus (Lacepède, 1802): OShr; [NIO-IA]; Chaetodon bennetti Cuvier, 1831: Cr; [IP] PMBC Chaetodon citrinellus Cuvier, 1831: Cr; [IP]; PMBC Mullidae Chaetodon collare Bloch, 1787: Cr; [NIO]; PMBC 61 The fishes of southwestern Thailand

Chaetodon decussatus Cuvier, 1831: Cr; [EIO]; PMBC Pomacanthus semicirculatus (Cuvier, 1831): Cr; [IP] Chaetodon ephippium Cuvier, 1831: Cr; [IP]; PMBC Pomacanthus sexstriatus (Cuvier, 1831): Cr; [WP] Chaetodon falcula Bloch, 1793: Cr; [IO]; PMBC Pomacanthus xanthometopon (Bleeker, 1853): Cr; [EIO- Chaetodon gardineri Norman, 1939: Cr; [IO]; PMBC WCP] Chaetodon guttatissimus Bennett, 1832: Cr; [IO]; PMBC Pygoplites diacanthus (Boddaert, 1772): Cr; [IP]; PMBC Chaetodon interruptus Ahl, 1922: Cr; [IO] Terapontidae Chaetodon kleinii Bloch, 1790: Cr; [IP]; PMBC Pelates quadrilineatus (Bloch, 1790): Mg, Sg; [IWP]; Chaetodon lineolatus Cuvier, 1831: Cr; [IP]; PMBC PMBC Chaetodon lunula (Lacepède, 1803): Cr; [IP]; PMBC Pseudoterapon theraps (Cuvier, 1829): Mg, Sg, OShr; Chaetodon madagaskariensis Ahl, 1923: Cr; [IO]; PMBC [IWP]; PMBC Chaetodon melannotus Bloch & Schneider, 1801: Cr; [IP]; Terapon jarbua (Forsskål, 1775): Cr, Mg, Sg, OShr; [IWP]; PMBC PMBC Chaetodon meyeri Bloch & Schneider, 1801: Cr; [IP]; Terapon puta (Cuvier, 1829): Mg, Sg, OShr; [IWP]; PMBC PMBC Kuhliidae Chaetodon octofasciatus Bloch, 1787: Cr; [EIO-WP]; Kuhlia mugil (Bloch & Schneider, 1801): Cr; [IP]; PMBC PMBC Cirrhitidae Chaetodon oxycephalus Bleeker, 1853: Cr; [EIO-WP] Cirrhitichthys falco Randall, 1963: Cr; [EIO-WP] Chaetodon rafflesi Bennett, 1830: Cr; [EIO-WCP]; PMBC Cirrhitichthys oxycephalus (Bleeker, 1855): Cr; [IP]; Chaetodon semeion Bleeker, 1855: Cr; [EIO-WCP] PMBC Chaetodon triangulum Cuvier, 1831: Cr; [IO]; PMBC Oxycirrhites typus Bleeker, 1857: Cr; [IP] Chaetodon trifascialis Quoy & Gaimard, 1825: Cr; [IP]; Paracirrhites arcatus (Cuvier, 1829): Cr; [IP] PMBC Paracirrhites fosteri (Schneider, 1801): Cr; [IP]; PMBC Chaetodon trifasciatus Park, 1797: Cr; [IO]; PMBC Cepolidae Chaetodon vagabundus Linnaeus, 1758: Cr; [IP]; PMBC Acanthocepola abbreviata (Valenciennes, 1835): OShr; Chaetodon xanthocephalus Bennett, 1832: Cr; [IO]; PMBC [NIO-IM]; PMBC Chelmon rostratus (Linnaeus, 1758): Cr; [WP]; PMBC Acanthocepola indica (Day, 1888): OShr; [IWP] Coradion altivelis Mcculloch, 1916: Cr; [WP] Pomacentridae Coradion chrysozonus (Cuvier, 1831): Cr, OShr; [WP]; Abudefduf bengalensis (Bloch, 1787): Cr; [EIO-WP]; PMBC PMBC Forcipiger flavissimus Jordan & Mcgregor, 1898: Cr; [IP]; Abudefduf notatus (Day, 1869): Cr; [IWP]; PMBC PMBC Abudefduf septemfasciatus (Cuvier, 1830): Cr; [IP]; PMBC Forcipiger longirostris (Broussonet, 1782): Cr; [IP] Abudefduf sordidus (Forsskål, 1775): Cr; [IP]; PMBC Hemitaurichthys zoster (Bennett, 1831): Cr; [IO]; PMBC Abudefduf vaigiensis (Quoy & Gaimard, 1825): Cr; [IP]; Heniochus acuminatus (Linnaues, 1758): Cr, Mg, Sg, OShr; PMBC [IP]; PMBC Amblyglyphidodon aureus (Cuvier, 1830): Cr; [WCP]; Heniochus diphreutes Jordan, 1903: Cr, OShr; [IP] PMBC Heniochus pleurotaenia Ahl, 1923: Cr; [EIO]; PMBC Amblyglyphidodon indicus Allen & Randall, 2002: Cr; [IO] Heniochus singularius Smith & Radcliffe, 1911: Cr; [EIO- Amblyglyphidodon leucogaster (Bleeker, 1847): Cr; [WP]; WCP]; PMBC PMBC Parachaetodon ocellatus (Cuvier, 1831): Cr, Sg: [EIO-WP]; Amphiprion akallopisos Bleeekr, 1853: Cr; [IO]; PMBC PMBC Amphiprion clarkii (Bennett, 1830): Cr, Sg; [IWP]; PMBC Pomacanthidae Amphiprion ephippium (Bloch, 1790): Cr; [EIO]; PMBC Apolemichthys trimaculatus (Lacapède, 1831): Cr; [IWP]; Amphiprion ocellaris Cuvier, 1830: Cr; [WP]; PMBC PMBC Amphiprion sebae Bleeekr, 1853: Cr, Sg; [EIO]; PMBC Apolemichthys xanthurus (Bennett, 1832): Cr; [IO]; PMBC Cheiloprion labiatus (Day, 1877): Cr; [IA]; PMBC Centropyge eibli Klausewitz, 1963: Cr; [EIO]; PMBC Chromis atripectoralis Welander & Schultz, 1951: Cr; Centropyge flavipectoralis Randall & Klausewitz, 1977: [IWP]; PMBC Cr; [EIO] Chromis cinerascens (Cuvier, 1830): Cr; [EIO-IA]; PMBC Centropyge multispinis (Playfair, 1866): Cr; [IO]; PMBC Chromis delta Randall, 1987: Cr; [EIO-WP] Chaetodontoplus mesoleucus (Bloch, 1787): Cr; [WP] Chromis dimidiata (Kluzinger, 1871): Cr; [IO]; PMBC Genicanthus caudovittatus (Günther, 1860): Cr, OShr; [IO]; Chromis elerae Fowler & Bean, 1928: Cr; [EIO-WP] PMBC Chromis flavipectoralis Randall, 1990: Cr; [EIO]; PMBC Pomacanthus annularis (Bloch, 1787): Cr, OShr; [IWP]; Chromis lepidolepis Bleeker, 1877: Cr; [IP] PMBC Chromis opercularis (Günther, 1866): Cr; [IO]; PMBC Pomacanthus imperator (Bloch, 1787): Cr; [IP]; PMBC Chromis ternatensis (Bleeker, 1856): Cr; [IP]; PMBC Pomacanthus navarchus (Cuvier, 1831): Cr; [IA] Chromis viridis (Cuvier, 1830): Cr; [IP]; PMBC 62 Phuket mar. biol. Cent. Res. Bull.

Chromis weberi Fowler & Bean, 1928: Cr; [IP]; PMBC Pomacentrus xanthosternus Allen, 1991: Cr; [EEIO]; Chromis xouthos Allen & Erdmann, 2005: Cr; [AS] PMBC Chrysiptera biocellata (Quoy & Gaimard, 1825): Cr; [IP]; Premnas biaculeatus (Bloch, 1790): Cr; [IA] PMBC Pristotis obtusirostris (Günther, 1862): Cr, OShr; [IWP]; Chrysiptera brownriggii (Bennett, 1828): Cr; [IP]; PMBC PMBC Chrysiptera glauca (Cuvier, 1830): Cr; [IP] Stegastes fasciolatus (de Vis, 1884): Cr; [IP]; PMBC Chrysiptera rollandi (Whitley, 1961): Cr; [IA]; PMBC Stegastes nigricans (Lacepède, 1803): Cr; [IP]; PMBC Chrysiptera unimaculata (Cuvier, 1830): Cr; [IWP]; PMBC Stegastes obreptus (Whitley, 1948): Cr; [EIO-WP]; PMBC Dascyllus aruanus (Linnaeus, 1758): Cr; [IP]; PMBC Stegastes punctatus (Quoy & Gaimard, 1825): Cr; [IWP]; Dascyllus carneus Fischer, 1885: Cr; [IO]; PMBC PMBC Dascyllus trimaculatus (Rüppell, 1828): Cr; [IP]; PMBC Teixeirichthys jordani (Rutter, 1897): OShr; [IWP] Dischistodus perspicillatus (Cuvier, 1830): Cr; [WP]; Labridae PMBC Anampses caeruleopunctatus Rüppell, 1829: Cr; [IP]; Hemiglyphidodon plagiometopon (Bleeker, 1852): Cr; [IA]; PMBC PMBC Anampses lineatus Randall, 1972: Cr; [IO]; PMBC Lepidozygus tapeinosoma (Bleeker, 1856): Cr; [IP]; PMBC Anampses meleagrides Valenciennes, 1840: Cr; [IWP]; Neoglyphidodon melas (Cuvier, 1830): Cr; [IWP]; PMBC PMBC Neoglyphidodon nigroris (Cuvier, 1830): Cr; [WP]; PMBC Anampses twistii Bleeker, 1856: Cr; [IP]; PMBC Neopomacentrus anabatoides (Bleeker, 1847): Cr; [IA]; Bodianus axillaris (Bennett, 1832): Cr; [IP] PMBC Bodianus diana (Lacepède, 1801): Cr; [IO]; PMBC Neopomacentrus bankieri (Richardson, 1846): Cr; [WP] Bodianus mesothorax (Bloch & Schneider, 1801): Cr; [WP]; Neopomacentrus cyanomos (Bleeker, 1856): Cr; [IWP]; PMBC PMBC Bodianus neilli (Day, 1867): Cr; [EIO]; PMBC Neopomacentrus filamentosus (Macleay, 1883): Cr; [IA]; Cheilinus chlorourus (Bloch, 1791): Cr, OShr; [IP]; PMBC PMBC Cheilinus fasciatus (Bloch, 1791): Cr; [IP]; PMBC Neopomacentrus sororius Randall &Allen, 2005: Cr; [IO]; Cheilinus oxycephalus Bleeker, 1853: Cr; [IP] PMBC Cheilinus trilobatus Lacepède, 1801: Cr; [IP]; PMBC Neopomacentrus taeniurus (Bleeker, 1856): Cr; [IWP]; Cheilinus undulatus Rüppell, 1835: Cr; [IP] PMBC Choerodon robustus (Günther, 1862): OShr; [IWP]; Neopomacentrus violascens (Bleeker, 1848): Cr; [IA] PMBC Plectroglyphidodon dickii (Leinard, 1839): Cr; [IP]; PMBC Cirrhilabrus cyanopleura (Bleeker, 1851): Cr; [EEIO]; Plectroglyphidodon johnstonianus Fowler & Ball, 1924: PMBC Cr; [IP]; PMBC Cirrhilabrus exquisitus Smith, 1957: Cr; [IP]; PMBC Plectroglyphidodon lacrymatus (Quoy & Gaimard, 1825): Cirrhilabrus joanallenae Allen, 2000: Cr; [AS] Cr; [IP]; PMBC Coris batuensis (Bleeker, 1856): Cr; [EIO-WCP]; PMBC Plectroglyphidodon leucozona (Bleeker, 1859): Cr; [IP]; Coris cuvieri (Bennett, 1831): Cr; [IO]; PMBC PMBC Diproctacanthus xanthurus (Bleeker, 1856): Cr; [IA] Pomacentrus adelus Allen, 1991: Cr; [IA]; PMBC Epibulus insidiator (Pallas, 1770): Cr; [IP]; PMBC Pomacentrus alleni Burgess, 1981: Cr; [EEIO]; PMBC Gomphosus caeruleus Lacepède, 1801: Cr; [IO]; PMBC Pomacentrus amboinensis Bleeker, 1868: Cr; [WCP]; Halichoeres argus (Bloch, 1791): Cr; [WP]; PMBC PMBC Halichoeres bicolor (Bloch & Schneider, 1801): Cr, Mg, Pomacentrus azuremaculatus Allen, 1991: Cr; [EEIO]; Sg; [EIO-IM]; PMBC PMBC Halichoeres chloropterus (Bloch, 1791): Cr; [IA]; PMBC Pomacentrus chrysurus Cuvier, 1830: Cr; [EIO-WP]; Halichoeres cosmetus Randall & Smith, 1982: Cr; [IO] PMBC Halichoeres hortulanus (Lacepède, 1801): Cr; [IP]; PMBC Pomacentrus lepidogenys Fowler & Bean, 1928: Cr; [WP]; Halichoeres kallochroma (Bleeker, 1835): Cr; [AS]; PMBC PMBC Pomacentrus moluccensis Bleeker, 1853: Cr; [WP]; PMBC Halichoeres leucoxanthus Randall & Smith, 1982: Cr; Pomacentrus nagasakiensis Tanaka, 1917: Cr; [EIO-WP]; [EIO]; PMBC PMBC Halichoeres margaritaceus (Valenciennes, 1839): Cr; Pomacentrus pavo (Bloch, 1787): Cr; [IP]; PMBC [WCP] Pomacentrus philippinus Evermann & Seale, 1907: Cr; Halichoeres marginatus Rüppell, 1835: Cr; [IP]; PMBC [EIO-WP]; PMBC Halichoeres nebulosus (Valenciennes, 1839): Cr; [IWP]; Pomacentrus polyspinus Allen, 1991: Cr; [AS]; PMBC PMBC Pomacentrus similis Allen, 1991: Cr; [EIO]; PMBC Halichoeres nigrescens (Bloch & Schneider, 1801): Cr, Sg; Pomacentrus tripunctatus Cuvier, 1830: Cr; [EIO-WP]; [IWP]; PMBC PMBC Halichoeres scapularis (Bennett, 1832): Cr; [IWP]; PMBC 63 The fishes of southwestern Thailand

Halichoeres timorensis (Bleeker, 1852): Cr; [EIO-IM]; Thalassoma purpureum (Forsskål, 1775): Cr; [IP] PMBC Thalassoma trilobatum (Lacepède, 1801): Cr; [IP]; PMBC Halichoeres vrolikii (Bleeker, 1855): Cr; [EIO]; PMBC Wetmorella nigropinnata (Seale, 1901): Cr; [IP]; PMBC Halichoeres zeylonicus (Bennett, 1832): Cr, OShr; [IO]; Xiphocheilus quadrimaculatus (Günther, 1880): OShr; PMBC [EIO-WP] Hemigymnus fasciatus (Bloch, 1792): Cr; [IP]; PMBC Scaridae Hemigymnus melapterus (Bloch, 1791): Cr; [IP]; PMBC Bolbometopon muricatum (Valenciennes, 1840): Cr; [IP]; Hologymnosus annulatus (Lacepède, 1801): Cr; [IP]; PMBC PMBC Calotomus carolinus (Valenciennes, 1840): Cr; [IP]; PMBC Hologymnosus doliatus (Lacepède, 1801): Cr; [IP] Cetoscarus bicolor (Rüppell, 1829): Cr; [IP] Iniistius baldwini (Jordan & Evermann, 1903): OShr; [IP]; Chlorurus capistratoides (Bleeker, 1847): Cr; [IO]; PMBC PMBC Chlorurus rhakoura Randall & Anderson, 1997: Cr; [EIO]; Iniistius bimaculatus (Rüppell, 1829): OShr; [IO]; PMBC PMBC Iniistius pavo (Valenciennes, 1840): Cr, OShr; [IP]; PMBC Chlorurus sordidus (Forsskål, 1775): Cr; [IP]; PMBC Labrichthys unilineatus (Guichenot, 1847): Cr; [IP]; PMBC Chlorurus strongylocephalus (Bleeker, 1854): Cr; [IO]; Labroides bicolor Fowler & Bean, 1928: Cr; [IP] Chlorurus troschelii (Bleeker, 1853): Cr; [EEIO] Labroides dimidiatus (Valenciennes, 1839): Cr; [IP]; Hipposcarus harid (Forsskål, 1775): Cr; [IO] PMBC Scarus frenatus Lacepède, 1802: Cr; [IP]; PMBC Labropsis xanthonota Randall, 1981: Cr; [IWP] Scarus ghobban Forsskål, 1775: Cr, Sg; [IP]; PMBC Leptojulis chrysotaenia Randall & Ferraris, 1981: Cr; [EIO] Scarus maculipinna Westneat, Satapoomin & Randall, Leptojulis cyanopleura (Bleeker, 1853): Cr, OShr; [NIO- 2007: Cr; [EEIO]; PMBC WP]; PMBC Scarus niger Forsskål, 1775: Cr; [IP]; PMBC Leptojulis urostigma Randall, 1996: OShr; [IA] Scarus prasiognathos Valenciennes, 1840: Cr; [EIO-WP]; Macropharyngodon ornatus Randall, 1978: Cr; [IWP]; PMBC PMBC Scarus quoyi Valenciennes, 1840: Cr; [EIO-WP]; PMBC Novaculichthys taeniourus (Lacepède, 1801): Cr; [IP] Scarus rubroviolaceus Bleeker, 1847: Cr; [IP]; PMBC Oxycheilinus areanatus (Valenciennes, 1840): Cr; [IP] Scarus russelii Valenciennes, 1840: Cr; [IO]; PMBC Oxycheilinus bimaculatus (Valenciennes, 1840): Cr; [IP] Scarus scaber Valenciennes, 1840: Cr; [IO]; PMBC Oxycheilinus celebicus (Bleeker, 1853): Cr; [WP]; PMBC Scarus tricolor Bleeker, 1847: Cr; [IWP]; PMBC Oxycheilinus digramma (Lacepède, 1801): Cr, OShr; [IP]; Scarus viridifucatus (Smith, 1956): Cr; [IO]; PMBC PMBC Champsodontidae Oxycheilinus orientalis (Günther, 1862): Cr; [IWP] Champsodon nudivittis (Ogilby, 1895): OShr; [IWP] Paracheilinus mccoskeri Randall & Harmelin-Vivien, 1977: Champsodon vorax Günther, 1867: OShr; [EIO-WP] Cr; [IO]; PMBC Pinguipedidae Pseudocheilinus evanidus Jordan & Evermann, 1903: Cr; Parapercis alboguttata (Günther, 1872): OShr; [NIO-IA]; [IP] PMBC Pseudocheilinus hexataenia (Bleeker, 1857): Cr; [IP]; Parapercis clathrata Ogilby, 1910: Cr, OShr; [WCP]; PMBC PMBC Pseudodax moluccanus (Valenciennes, 1840): Cr; [IP]; Parapercis cylindrica (Bloch, 1792): Cr: [EIO-WP], PMBC PMBC Parapercis hexophthalma (Cuvier, 1829): Cr, OShr; [IWP]; Pseudojuloides kaleidos Kuiter & Randall, 1995: Cr; [EIO]; PMBC PMBC Parapercis maculata (Bloch & Schneider, 1801): OShr; Pteragogus cryptus Randall, 1981: Cr; [IWP]; PMBC [IO] Stethojulis albovittata (Bonnaterre, 1788): Cr; [IO]; PMBC Parapercis millepunctata (Günther, 1860): Cr, OShr; [EIO- Stethojulis bandanensis (Bleeker, 1851): Cr; [WCP] WCP]; PMBC Stethojulis interrupta (Bleeker, 1851): Cr; [IWP]; PMBC Parapercis punctata (Cuvier, 1829): Mg, OShr; [IO]; Stethojulis strigiventer (Bennett, 1832): Cr, Sg; [IP] PMBC Stethojulis trilineata (Bloch & Schneider, 1801): Cr; [EIO- Parapercis quadrispinosa (Weber, 1913): OShr; [EIO]; WCP]; PMBC PMBC Suezichthys caudovittatus (Steindachner, 1898): OShr; [IO]; Parapercis schauinslandi (Steindachner, 1900): Cr; [IP] PMBC Parapercis snyderi Jordan & Starks, 1905: Cr, OShr; [WP] Thalassoma amblycephalum (Bleeker, 1856): Cr; [IP]; Parapercis tetracantha (Lacepède, 1800): Cr, OShr; [IWP]; PMBC PMBC Thalassoma hardwicke (Bennett, 1830): Cr; [IP]; PMBC Parapercis xanthozona (Bleeker, 1849): Cr, OShr; [IWP]; Thalassoma jansenii (Bleeker, 1856): Cr; [EIO-WP]; PMBC PMBC Trichonotidae Thalassoma lunare (Linnaeus, 1758): Cr; [IP]; PMBC Trichonotus setiger Bloch & Schneider, 1801: OShr; [WP] 64 Phuket mar. biol. Cent. Res. Bull.

Trichonotus sp.1: Cr, OShr Entomacrodus vermiculatus (Valenciennes, 1836): Cr; [IO]; Percophidae PMBC Bembrops caudimaculata Steindachner, 1876: OShr; [IWP] Exallias brevis (Kner, 1968): Cr; [IP] Bembrops platyrhynchus (Alcock, 1894): OShr; [IWP]; Glyptoparus delicatulus Smith, 1959: Cr; [IWP] PMBC Istiblennius bellus (Günther, 1861): Cr; [IP]; PMBC Ammodytidae Istiblennius dussumieri (Valenciennes, 1836): Cr; [IWP]; Bleekeria kallolepis Günther, 1862: OShr; [EIO] PMBC Uranoscopidae Istiblennius edentulus (Schneider, 1801): Cr; [IP]; PMBC Ichthyoscopus lebeck (Schneider, 1801): OShr; [EIO]; Istiblennius lineatus (Valenciennes, 1836): Cr; [EIO-WCP] PMBC Laiphognathus multimaculatus Smith, 1955: Cr; [IWP]; Uranoscopus affinis Cuvier, 1829: OShr; [IWP]; PMBC PMBC Uranoscopus bicinctus Temminck & Schlegel, 1843: OShr; Meiacanthus smithi Klausewitz, 1916: Cr; [EIO]; PMBC [EIO-WP] Meiacanthus urostigma Smith-Vaniz, Satapoomin & Allen, Uranoscopus cognatus Cantor, 1849: OShr; [EIO-WP] 2001: Cr; [AS]; PMBC Uranoscopus oligolepis Bleeker, 1878: OShr; [IO] Omobranchus elongatus (Peters, 1855): Cr; [IWP]; PMBC Tripterygiidae Omobranchus fasciolatus (Valenciennes, 1836): Mg; [IO]; Ennaepterygius fasciatus (Weber, 1909): Cr; [IP] PMBC Ennaepterygius nanus (Schultz, 1960): Cr; [WP] Omobranchus ferox (Herre, 1927): Mg; [IP]; PMBC Ennaepterygius philippinus (Peters, 1868): Cr; [IWP] Omobranchus obliquus (Garman, 1903): Cr; [WCP]; Enneapterygius sp.1: PMBC Enneapterygius tutuilae Jordan & Seale, 1906: Cr; [IP]; Omobranchus punctatus (Valenciennes, 1836): Mg; [IWP]; PMBC PMBC Helcogramma chica Rosenblatt, 1960: Cr; [EIO-WP] Omobranchus smithi (Rao, 1973): Mg; [EIO-IM]; PMBC Helcogramma lacuna Williams & Howe, 2003: Cr; [AS] Omobranchus zebra (Bleeker, 1868): Mg; [EIO-IM] Helcogramma obtusirostre Klunzinger, 1871: Cr; [IWP] Petroscirtes breviceps (Valenciennes, 1836): Cr, Mg, Sg; Helcogramma rosea Holleman, 2006: Cr; [EIO] [IWP]; PMBC Helcogramma sp.1: Cr Petroscirtes mitratus Rüppell, 1830: Cr, Sg; [IP]; PMBC Helcogramma springeri Hansen, 1986: Cr; [WP]; PMBC Petroscirtes thepassi Bleeker, 1853: Sg; [WP] Helcogramma striata Hansen, 1986: Cr; [EIO-WP]; PMBC Petroscirtes variabilis Cantor, 1850: Sg; [EIO-IA]; PMBC Norfolkia brachylepis (Schultz, 1960): Cr; [IP] Plagiotremus phenax Smith-Vaniz, 1976: Cr; [EIO]; PMBC Ucla xenogrammus Holleman, 1993: Cr; [IP]; PMBC Plagiotremus rhinorhynchos (Bleeker, 1852): Cr; [IP]; Blenniidae PMBC Alticus saliens (Lacepède, 1800): Cr; [IP]; PMBC Plagiotremus tapeinosoma (Bleeker, 1857): Cr; [IP]; Andamia reyi Sauvage, 1880: Cr; [IM]; PMBC PMBC Aspidontus dussumieri (Valenciennes, 1836): Cr; [IP] Plagiotremus townsendi (Regan, 1905): Cr; [IO] Aspidontus tractus Fowler, 1903: Cr; [IO]; PMBC Praealticus oortii (Bleeker, 1851): Cr; [IM] Atrosalarias fuscus (Rüppell, 1835): Cr; [IO]; PMBC Praealticus triangulus (Chapman, 1951): Cr; [EEIO] Blenniella chrysospilos (Bleeker, 1857): Cr; [IP] Salarias fasciatus (Bloch, 1786): Cr; [IWP]; PMBC Blenniella cyanostigma (Bleeker, 1849): Cr; [IO] Salarias guttatus Valenciennes, 1836: Cr; [IA]; PMBC Blenniella leopardus (Fowler, 1904): Cr; [AS]; PMBC Salarias sinuosus Snyder, 1908: Cr; [WP]; PMBC Blenniella periophthalmus (Valenciennes, 1836): Cr; [IWP] Xiphasia setifer Swainson, 1839: OShr; [IWP] Cirripectes auritus Carlson, 1980: Cr; [IP] Gobiesocidae Cirripectes castaneus (Valenciennes, 1836): Cr; [IWP] Diademichthys lineatus (Sauvage, 1883): Cr; [IWP] Cirripectes filamentosus (Alleyne & Macleay, 1877): Cr; Discotrema crinophilum Briggs, 1976: Cr; [WP] [IWP]; PMBC Lepadichthys lineatus Briggs, 1966: Cr; [IWP] Cirripectes stigmaticus Strasburg & Schultz, 1953: Cr; Callionymidae [IWP] Bathycallionymus kaianus (Günther, 1880): OShr; [IWP] Crossosalarias macrospilus Smith-Vaniz & Springer, Calliurichthys japonicus (Houttuyn, 1782): OShr; [WP]; 1971: Cr; [WP] PMBC Ecsenius bicolor (Day, 1888): Cr; [EIO-WP]; PMBC Dactylopus dactylopus (Valenciennes, 1837): Mg, Sg, OShr; Ecsenius lubbocki Springer, 1988: Cr; [EEIO]; PMBC [WP]; PMBC Ecsenius midas Starck, 1969: Cr; [IP]; PMBC Repomucenus belcheri recurvispinnis (Li, 1966): OShr; Ecsenius nalolo Smith, 1959: Cr; [IO]; PMBC [EIO-IM] Ecsenius paroculus Springer, 1988: Cr; [EEIO]; PMBC Repomucenus doryssus (Jordan & Fowler, 1903): OShr; Enchelyurus kraussi (Klunzinger, 1871): Cr; [IWP]; PMBC [WP] Entomacrodus striatus (Quoy & Gaimard, 1836): Cr; [IP]; Repomucenus filamentosus (Valenciennes, 1837): OShr; [IP] PMBC 65 The fishes of southwestern Thailand

Repomucenus hindsi (Richardson, 1844): Sg, OShr; [NIO- Amblyeleotris steinitzi (Klausewits, 1974): Cr; [IWP]; IM] PMBC Repomucenus octostigmatus (Fricke, 1981): OShr; [IM] Amblyeleotris triguttatus Randall, 1994: Cr; [IO] Repomucenus sagitta (Pallas, 1770): OShr; [EIO-IM] Amblyeleotris wheeleri (Polunin & Lubbock, 1977): Cr; Repomucenus schaapii (Bleeker, 1852): Mg, Sg, OShr; [EIO- [IWP] IM]; PMBC Amblygobius hectori (Smith, 1957): Cr; [IWP]; PMBC Paradiplogrammus enneactis (Bleeker, 1879): Mg, Sg; Amblygobius nocturnus (Herre, 1945): Cr; [NIO-WCP]; [WP]; PMBC PMBC Synchiropus lineolatus (Valenciennes, 1837): Cr; [IWP] Amblygobius semicinctus (Bennett, 1833): Cr; [IO]; PMBC Synchiropus stellatus Smith, 1963: Cr; [IO] Amblygobius stethophthalmus (Bleeker, 1851): Cr; [IM] Eleotridae Amoya gracilis (Bleeker, 1875): Mg; [IA]; PMBC Bostrychus sinensis (Lacepède, 1801): Mg; [EIO-WP] Amoya moloanus (Herre, 1927): Mg, Sg; [WP]; PMBC Butis butis (Hamilton, 1822): Mg, Sg; [IWP]; PMBC Apocryptodon madurensis (Bleeker, 1849): Mg, Sg; [EIO- Butis humeralis (Valenciennes, 1837): Mg; [EIO-WP]; WP]; PMBC PMBC Arcygobius baliurus (Valenciennes, 1837): Sg; [IWP]; Butis koilomatodon (Bleeker, 1849): Mg, Sg; [IWP]; PMBC PMBC Calumia godeffroyi (Günther, 1877): Cr; [IP] Asterropteryx dfh sp.3: Cr Eleotris melanosoma Bleeker, 1852): Mg; [IWP]; PMBC Asterropteryx ensiferus (Bleeker, 1874): Cr; [IP]; PMBC Odonteleotris macrodon (Bleeker, 1853): Mg; [EIO-IM]; Asterropteryx semipunctatus Rüppell, 1828: Cr; [IP]; PMBC PMBC Ophiocara aporos (Bleeker, 1854): Mg; [IWP]; PMBC Barbuligobius boehlkei Lachner & McKinney, 1974: Cr; Ophiocara porocephala (Valenciennes, 1837): Mg; [IWP]; [IWP] PMBC Bathygobius cocosensis (Bleeker, 1854): Cr; [IP]; PMBC Xenisthmidae Bathygobius crassiceps (Jordan & Seale, 1906): Cr; [IP]; Xenisthmus africanus Smith, 1958: Cr; [IO] PMBC Xenisthmus polyzonatus (Klunzinger, 1817): Cr; [IP] Bathygobius cyclopterus (Valenciennes, 1837): Cr; [IWP]; Gobiidae PMBC Acentrogobius audax Smith, 1959: Sg; [IWP]; PMBC Bathygobius fuscus (Rüppell, 1830): Cr, Sg; [IWP]; PMBC Acentrogobius caninus (Valenciennes,1837): Mg, Sg; [IWP]; Bathygobius karachiensis Hoda & Goren, 1990: Cr; [EIO] PMBC Bathygobius laddi (Fowler, 1931): Cr; [IP]; PMBC Acentrogobius chlorostigmatoides (Bleeker, 1849): OShr; Bathygobius meggitti (Hora & Mukuji, 1936): Cr; [WP] [WP]; PMBC Boleophthalmus boddarti (Pallas, 1770): Mg; [NIO-WP]; Acentrogobius cyanomos (Bleeker, 1849): Mg, Sg; [EIO- PMBC IM]; PMBC Brachygobius kabiliensis Inger, 1958: Mg; [IM] Acentrogobius janthinopterus (Bleeker, 1852): Mg; [WP]; Bryaninops amplus Larson, 1985: Cr; [IP] PMBC Bryaninops erythops (Jordan & Seale, 1906): Cr; [EIO- Acentrogobius madraspatensis (Day, 1868): Mg, Sg; [EIO- WP] WP]; PMBC Bryaninops loki Larson, 1985: Cr; [IWP] Acentrogobius multifasciatus (Herre, 1927): Cr, Sg; [WP]; Bryaninops natans Larson, 1985: Cr; [IWP]; PMBC PMBC Bryaninops ridens Smith, 1959: Cr; [IWP]; PMBC Acentrogobius sp.1: Sg Bryaninops youngei (Davis & Cohen, 1969): Cr; [IP] Acentrogobius viridipunctatus (Valenciennes, 1837): Mg; Cabillus tongarevae (Fowler, 1927): Cr; [EIO-WP]; PMBC [IWP]; PMBC Callogobius andamanensis Menon & Chatterjee, 1974: Mg; Amblyeleotris aurora (Polunin & Lubbock, 1977): Cr; [IO] [AS] Amblyeleotris diagonalis Polunin & Lubbock, 1979: Cr; Callogobius dfh sp.3: Cr [IP]; PMBC Callogobius dfh sp.6: Cr Amblyeleotris downingi Randall, 1994: Cr; OShr; [NIO]; Callogobius dfh sp.11: Cr PMBC Callogobius dfh sp.13: Cr Amblyeleotris fontanesii (Bleeker, 1852): Cr, OShr; [WP]; Callogobius dfh sp.15: Cr PMBC Callogobius dfh sp.16: Cr Amblyeleotris gymnocephala (Bleeker, 1853): Cr; [IA]; Callogobius flavobrunneus (Smith, 1958): Cr; [IO]; PMBC PMBC Callogobius maculipinnis (Fowler, 1918): Cr; [IWP] Amblyeleotris latifasciata Polunin & Lubbock, 1979: Cr; Callogobius plumatus (Smith, 1959): Cr; [IO] [IA]; PMBC Callogobius rw-th sp.2: Mg Amblyeleotris periophthalma (Bleeker, 1853): Cr; [IWP]; Callogobius sclateri (Steindachner, 1880): Cr; [IP]; PMBC PMBC Cristatogobius nonatoae (Ablan, 1940): Mg; [WP]; PMBC 66 Phuket mar. biol. Cent. Res. Bull.

Cryptocentrus caeruleomaculatus (Herre, 1933): Cr, Sg; Eviota zonura Jordan & Seale, 1906: Cr; [WCP] [IP]; PMBC Exyrias bellissimus (Smith, 1959): Cr; [IWP]; PMBC Cryptocentrus cyanotaenia (Bleeker, 1853): Cr, Mg, OShr; Exyrias puntang (Bleeker, 1851): Mg; [EIO-WP] [IA]; PMBC Favonigobius melanobranchus (Fowler, 1934): Cr, Mg, Cryptocentrus fasciatus (Playfair & Gunther, 1867): Cr; Sg; [IWP]; PMBC [IWP]; PMBC Favonigobius reichei (Bleeker, 1853): Cr, Mg, Sg; [IWP]; Cryptocentrus leucostictus (Günther, 1871): Cr; [WCP] PMBC Cryptocentrus maudae Fowler, 1937: Cr; [IA]; PMBC Flabelligobius latruncularius (Klausewitz, 1974): Cr; [IO] Cryptocentrus melanopus (Bleeker, 1860): Cr, Sg, OShr; Flabelligobius russus (Cantor, 1849): OShr; [IM]; PMBC [WP]; PMBC Fusigobius dfh sp.4: Cr Cryptocentrus pavoninoides (Bleeker, 1849): Cr, OShr; Fusigobius duospilus Hoese & Reader, 1985: Cr; [IP]; [IA]; PMBC PMBC Cryptocentrus sp.1 (dfh sp.27): Cr, Sg; [WP]; PMBC Fusigobius humeralis (Randall, 2001): Cr; [IP] Cryptocentrus strigilliceps (Jordan & Seale, 1906): Cr, Sg; Fusigobius inflamaculatus Randall, 1994: Cr; [IWP] [IP]; PMBC Fusigobius maximus (Randall, 2001): Cr; [IWP] Ctenogobiops aurocingulus (Herre, 1935): Cr; [WCP] Fusigobius melacron (Randall, 2001): Cr; [WP] Ctenogobiops crocineus Smith, 1959: Cr; [IWP]; PMBC Fusigobius neophytus (Günther, 1877): Cr; [IP]; PMBC Ctenogobiops feroculus Lubbock & Polunin, 1977: Cr; Fusigobius pallidus (Randall, 2001): Cr; [IWP] [IWP] Fusigobius signipinnis Hoese & Obika, 1988: Cr; [WP] Ctenogobiops pomastictus Lubbock & Polunin, 1977: Cr; Gladiogobius rex Shibukawa & Allen, 2007: Cr; [IO]; [WP]; PMBC PMBC Drombus globiceps (Hora, 1923): Mg; [EIO-WP]; PMBC Glossogobius bicirrhosus (Weber, 1894): Mg; [WP]; Drombus key (Smith, 1947): Cr, Mg, Sg; [IO]; PMBC PMBC Drombus ocyurus (Jordan & Seale, 1907): Mg, Sg; [WP]; Glossogobius circumspectus (Macleay, 1883): Mg; [WP]; PMBC PMBC Drombus triangularis (Weber, 1911): Mg, Sg; [IWP]; Glossogobius guiris (Hamilton, 1822): Mg; [IWP] PMBC Glossogobius sparsipapillus Akihito & Meguro, 1976: Mg; Eugnathogobius illotus (Larson, 1999): Mg; [IM]; PMBC [WP]; PMBC Eugnathogobius microps Smith, 1931: Mg; [IM (presently Gnatholepis anjerensis (Bleeker, 1850): Cr; [IP]; PMBC known from west coast of Thailand, Gulf of Thailand Gnatholepis cauerensis (Bleeker, 1853): Cr; [IP] and southern Vietnam)] Gobiodon citrinus (Rüppell, 1853): Cr; [IWP] Eugnathogobius mindora (Herre, 1945): Mg; WP Gobiodon dfh sp.11: Cr Eugnathogobius polylepis (Wu & Ni, 1985): Mg; [IA] Gobiodon erythrospilus Bleeker, 1857: Cr; [WP] Eugnathogobius variegatus (Peters, 1869): Mg; [IM]; Gobiodon histrio (Valenciennes, 1837): Cr; [IWP]; PMBC PMBC Gobiodon prolixus Winterbottom & Harold, 2005: Cr; Eviota afelei Jordan & Seale, 1906: Cr; [WCP] [EIO-WP] Eviota albolineata Jewett & Lachner, 1983: Cr; [IP]; PMBC Gobiodon quinquestrigatus (Cuvier & Valenciennes, 1837): Eviota herrei Jordan & Seale, 1906: Cr; [WCP] Cr; [WCP] Eviota indica Lachner & Karnella, 1980: Cr; [IO]; PMBC Gobiodon rivulatus (Rüppell, 1828): Cr; [IWP]; PMBC Eviota infulata (Smith, 1956): Cr; [IP] Gobiodon rw-th sp.1: Cr Eviota mikiae Allen, 2001: Cr; [EIO] Gobiodon rw-th sp.2: Cr Eviota nigriventris Giltay, 1933: Cr; [IA]; PMBC Gobiodon rw-th sp.3: Cr Eviota pellucida Larson, 1976: Cr; [WCP] Gobiopsis aporia Lachner & McKinney, 1978: Cr; [EIO- Eviota prasina (Klunzinger, 1871): Cr; [IWP]; PMBC WP]; PMBC Eviota prasites Jordan & Seale, 1906: Cr; [IP] Gobiopsis macrostoma Steindachner, 1861: Mg; [NIO- Eviota punctulata Jewett & Lachner, 1983: Cr; [WCP]; IM]; PMBC PMBC Gobiopsis quinquecincta (Smith, 1931): Cr; [EIO-WP]; Eviota queenslandica Whitley, 1932: Cr; [WP]; PMBC PMBC Eviota rw-th sp.1: Cr Gobiopterus brachypterus (Bleeker, 1855): Mg; [IA]; Eviota rw-th sp.2: Cr PMBC Eviota rw-th sp.4: Cr Gobiopterus panayensis (Herre, 1944): Mg; [WP]; PMBC Eviota sebreei Jordan & Seale, 1906: Cr; [IP]; PMBC Gobiopterus rw-th sp.1: Mg Eviota sigillata Jewett & Lachner, 1983: Cr; [IWP]; PMBC Hemigobius hoevenii (Bleeker, 1851): Mg; [IA] Eviota spilota Lachner & Karnella, 1980: Cr; [WP]; PMBC Hemigobius mingi Herre, 1936: Mg; [IM] Eviota zebrina Lachner & Karnella, 1978: Cr: [IWP]; Istigobius decoratus (Herre, 1927): Cr; [IWP]; PMBC PMBC Istigobius diadema (Steindachner, 1877): Cr, Mg; [EIO- IA]; PMBC 67 The fishes of southwestern Thailand

Istigobius goldmanni (Bleeker, 1852): Cr; [WP]; PMBC Periophthalmus walailakae Darumas & Tantochodok, Istigobius ornatus (Rüppell, 1830): Cr; [IP]; PMBC 2002: Mg; [IM (presently known from west coast of Istigobius rigilius (Herre, 1953): Cr; [WCP] Thailand and Singapore)]; PMBC Lophogobius bleekeri Popta, 1921: Mg; [IM]; PMBC Pleurosicya bilobata (Koumans, 1941): Sg; [IWP] Mahidolia mystacina (Valenciennes, 1837): Cr, Sg, OShr; Pleurosicya coerulea Larson, 1990: Cr; [IP] [IWP]; PMBC Pleurosicya fringilla Larson, 1990: Cr; [IWP]; PMBC Mangarinus seshaiyai (Jacop & Rangarajan, 1960): Mg; Pleurosicya labiata (Weber, 1913): Cr; [EIO-IA]; PMBC [EIO] Pleurosicya micheli Fourmanoir, 1971: Cr; [IP] Mangarinus waterousi Herre, 1943: Mg; [WP]; PMBC Pleurosicya mossambica Smith, 1959: Cr; [IP]; PMBC Mugilogobius chulae (Smith, 1932): Mg; [IM]; PMBC Pleurosicya plicata Larson, 1990: Cr; [IWP] Mugilogobius fasciatus Larson, 2001: Mg; [IM (presently Pleurosicya prognatha Goren, 1984: Cr; [IWP] known from west coast of Thailand and Singapore)]; Priolepis cinctus (Regan, 1908): Cr; [IWP]; PMBC PMBC Priolepis inhaca (Smith, 1949): Cr; [IWP] Mugilogobius mertoni (Weber, 1911): Mg; [IWP] Priolepis nuchifasciatus (Günther, 1873): Cr; [IA] Mugilogobius tigrinus Larson, 2001: Mg; [IM (presently Priolepis semidoliatus (Valenciennes, 1837): Cr; [IP]; known from west coast of Thailand, Singapore and Gulf PMBC of Thailand)]; PMBC Psammogobius biocellatus (Valenciennes, 1837): Mg, Sg; Myersina adonis Shibukawa & Satapoomin, 2006; Mg, Sg; [IWP]; PMBC [IM (presently known from west coast of Thailand and Pseudapocryptes lanceolatus (Bloch & Schneider, 1801): Singapore]; PMBC Mg; [EIO-IM] Myersina crocatus (Wongratana, 1975): Cr, OShr; [IM]; Pseudogobius javanicus (Bleeker, 1856): Mg; [IM]; PMBC PMBC Redigobius bikolanus Herre, 1927: Mg; [WP]; PMBC Myersina filifer (Cuvier & Valenciennes, 1837): OShr; [IWP]; Redigobius chrysosoma (Bleeker, 1857): Mg; [IA]; PMBC PMBC Scartelaos cantoris (Day, 1871): Mg, Sg; [AS]; PMBC Myersina yangii (Chen, 1960): Cr, OShr; [WP]; PMBC Scartelaos histophorus (Valenciennes, 1837): Mg, Sg; [NIO- Oligolepis acutipennis (Valenciennes, 1837): Mg; [IWP] WP]; PMBC Oplopomus caninoides (Bleeker, 1852): Cr, OShr; [EIO- Silhouettea sp.1:Cr,Sg;PMBC WP]; PMBC Stigmatogobius pleurostigma (Bleeker, 1849): Mg; [IM] Oplopomus oplopomus (Valenciennes, 1837): Cr, OShr; Stigmatogobius sandanundio (Hamilton, 1822): Mg; [EIO]; [IP]; PMBC PMBC Oxyurichthys microlepis (Bleeker, 1849): Mg, Sg; [IWP]; Stonogobiops nematodes Hoese & Randall, 1982: Cr; [IA] PMBC Sueviota lachneri Winterbottom & Hoese, 1988: Cr; [IWP]; Oxyurichthys ophthalmonema (Bleeker, 1856): Mg, Sg; PMBC [EIO-WP] Taenioides cirratus (Blyth, 1860): Mg, Sg, OShr; [IWP]; Oxyurichthys papuensis (Valenciennes, 1837): Cr, OShr; PMBC [IWP]; PMBC Taenioides gracilis (Valenciennes, 1837): OShr; [IWP]; Oxyurichthys tentacularis (Valenciennes, 1837): Mg, Sg; PMBC [WP]; PMBC Trimma emeryi Winterbottom, 1985: Cr; [IO] Palutrus scapulopunctatus (De Beaufort, 1912): Cr; [IWP]; Trimma flammeum (Smith, 1959): Cr; [IO]; PMBC PMBC Trimma fucatum Winterbottom & Southcott, 2007: Cr; Pandaka lidwilli (McCulloch, 1917): Mg; [WP]; PMBC [AS]; PMBC Pandaka pygmaea Herre, 1927: Mg; [IM] Trimma naudei Smith, 1956: Cr; [IWP]; PMBC Parachaeturichthys polynema (Bleeker, 1853): Mg; [IWP]; Trimma okinawae (Aoyaki, 1949): Cr; [WCP]; PMBC PMBC Trimma sanguinellus Winterbottom & Southcott, 2007: Paragobiodon modestus (Regan, 1908): Cr; [IP]; PMBC Cr; [AS]; PMBC Paratrypauchen microcephalus (Bleeker, 1860): Mg, Sg; Trimma striata (Herre, 1945): Cr; [EIO-WP]; PMBC [IWP] Trimma talori Lobel, 1979: Cr; [IP] Periophthalmus argentilineatus Valenciennes, 1837: Mg; Trimma winterbottomi Randall & Downing, 1994: Cr; [IO]; [IWP]; PMBC PMBC Periophthalmus chrysospilos Bleeker, 1852: Mg; [EIO- Trypauchen vagina (Bloch & Schneider, 1801): OShr; [NIO- IM]; PMBC IM]; PMBC Periophthalmus kalolo Lesson, 1830: Mg; [IWP]; PMBC Valenciennea immaculata (Ni, 1981): Sg; [WP] Periophthalmus minutus Eggert, 1935: Mg; [IA]; PMBC Valenciennea limicola Hoese & Larson, 1994: Cr; [WP]; Periophthalmus novemradiatus (Hamilton, 1822): Mg; PMBC [EIO-IA]; PMBC Valenciennea mularis (Valenciennes, 1837): Cr, Sg, OShr; Periophthalmus septemradiatus (Hamilton, 1822): Mg; [IA] [EIO-IM] Valenciennea parva Hoese & Larson, 1994: Cr; [IWP] 68 Phuket mar. biol. Cent. Res. Bull.

Valenciennea puellaris (Tomiyama, 1956): Cr, OShr; [IP]; Siganus magnificus (Burgess, 1977): Cr; [EEIO]; PMBC PMBC Siganus puelloides Woodland & Randall, 1979: Cr; [EIO]; Valenciennea randalli Hoese & Larson, 1994: Cr, OShr; PMBC [WP]; PMBC Siganus stellatus laqueus Von Bonde, 1934: Cr, OShr; [IO]; Valenciennea sexguttata (Valenciennes, 1837): Cr, OShr; PMBC [IP]; PMBC Siganus vermiculatus (Valenciennes, 1835): Cr, Mg, Sg, Valenciennea strigata (Broussonet, 1782): Cr; [IP]; PMBC OShr; [EIO-WP]; PMBC Valenciennea wardii (Playfair, 1867): Cr, OShr; [IWP]; Siganus virgatus (Valenciennes, 1835): Cr, Sg, OShr; [EIO- PMBC WP]; PMBC Vanderhorstia ambanoro (Fourmanoir, 1957): Cr, OShr; Zanclidae [IWP]; PMBC Zanclus cornutus (Linnaeus, 1758): Cr; [IP]; PMBC Vanderhorstia ornatissima Smith, 1959: Cr; [IP] Acanthuridae Yongeichthys nebulosus (Forsskål, 1775): Cr, Mg, Sg, OShr; Acanthurus bariene Lesson, 1830: Cr; [WP-EIO]; PMBC [IWP]; PMBC Acanthurus blochii Valenciennes, 1835: Cr; [IP] Microdesmidae Acanthurus guttatus Forster, 1801: Cr; [IP] Gunnellichthys curiosus Dawson, 1968: Cr; [IP] Acanthurus leucocheilus Herre, 1927: Cr; [IWP] Gunnellichthys viridescens Dawson, 1968: Cr; [IP] Acanthurus leucosternon Bennett, 1832: Cr; [IO]; PMBC Ptereleotridae Acanthurus lineatus (Linnaeus, 1758): Cr; [IP]; PMBC Nemateleotris decora Randall & Allen, 1973: Cr; [IWP] Acanthurus maculiceps (Ahl, 1923): Cr; [EIO-WCP] Nemateleotris magnifica Fowler, 1928: Cr; [IP] Acanthurus mata (Cuvier, 1829): Cr; [IP]; PMBC Oxymetopon compressus Chan, 1966: Cr, OShr; [WP]; Acanthurus nigricauda Duncker & Mohr, 1929: Cr; [IP]; PMBC PMBC Parioglossus formosus (Smith, 1931): Cr; [WP]; PMBC Acanthurus nigrofuscus (Forsskål, 1775): Cr; [IP] Parioglossus palustris (Herre, 1945): Mg; [WP] Acanthurus olivaceus Forster, 1801: Cr; [WCP] Parioglossus philippinus (Herre, 1940): Cr; [IWP]; PMBC Acanthurus tennenti Günther, 1861: Cr; [IO] Ptereleotris arabica Randall & Hoese, 1985: Cr, OShr; Acanthurus thompsoni (Fowler, 1923): Cr; [IP]; PMBC [IO]; PMBC Acanthurus triostegus (Linnaeus, 1758): Cr; [IP]; PMBC Ptereleotris evides (Jordan & Hubbs, 1925): Cr; [IP] Acanthurus tristis Tickell In Day, 1888: Cr; [IO]; PMBC Ptereleotris heteroptera (Bleeker, 1855): Cr; [IP]; PMBC Acanthurus xanthopterus Valenciennes, 1835: Cr; [IP]; Ptereleotris microlepis (Bleeker, 1856): Cr; [IP]; PMBC PMBC Ptereleotris monoptera (Bleeker, 1855): Cr; [IP] Ctenochaetus binotatus Randall, 1955: Cr; [IP]; PMBC Ptereleotris zebra (Fowler, 1938): Cr; [IP] Ctenochaetus striatus (Quoy & Gaimard, 1825): Cr; [IP]; Kurtidae PMBC Kurtus indicus Bloch, 1786: OShr; [EIO-IM]; PMBC Ctenochaetus truncatus Randall & Clements, 2001: Cr; Ephippidae [IO]; PMBC Ephippus orbis (Bloch, 1787): Mg, Sg, OShr; [IWP]; Naso brachycentron (Valenciennes, 1835): Cr; [IP] PMBC Naso brevirostris (Valenciennes, 1835): Cr; [IP]; PMBC Platax boersi Bleeker, 1852: Cr; [IP] Naso caeruleocauda Randall, 1994: Cr; [IA] Platax orbicularis (Forsskål, 1775): Cr, Mg, Sg, OShr; [IP]; Naso elegans (Rüppell, 1829): Cr; [IO]; PMBC PMBC Naso hexacanthus (Bleeker, 1855): Cr; [IP]; PMBC Platax pinnatus (Linnaeus, 1758): Cr, OShr; [WP]; PMBC Naso lopezi Herre, 1927: Cr; [WP] Platax teira (Forsskål, 1775): Cr, OShr; [IWP]; PMBC Naso mcdadei Johnson, 2002: Cr; [IWP] Scatophagidae Naso thynnoides (Valenciennes, 1835): Cr; [IP] Scatophagus argus (Linnaeus, 1758): Mg, Sg; [NIO-WP]; Naso unicornis (Forsskål, 1775): Cr; [IP]; PMBC PMBC Naso vlamingii (Valenciennes, 1835): Cr; [IP]; PMBC Siganidae Paracanthurus hepatus (Linnaeus, 1766): Cr; [IP] Siganus argenteus (Quoy & Gaimard, 1825): Cr; [IP]; Zebrasoma desjardinii (Bennett, 1835): Cr; [IO]; PMBC PMBC Zebrasoma scopas (Cuvier, 1829): Cr; [IP]; PMBC Siganus canaliculatus (Park, 1797): Cr, Mg, Sg, OShr; [NIO- Sphyraenidae IA]; PMBC Sphyraena baracuda (Walbaum, 1792): Cr, Sg, OShr; Siganus corallinus (Valenciennes, 1835): Cr; [IWP]; PMBC [COS]; PMBC Siganus fuscescens (Houttuyn, 1782): Cr; [WP]; PMBC Sphyraena forsteri Cuvier, 1829: Cr, OShr; [IP]; PMBC Siganus guttatus (Bloch, 1787): Cr, Mg, Sg, OShr; [IM] Sphyraena jello Cuvier, 1829: Cr, Mg, Sg, OShr; [IP]; Siganus javus (Linnaeus, 1766): Cr, Mg, Sg, OShr; [NIO- PMBC WP] Sphyraena obtusata Cuvier, 1829: Cr, Sg, OShr; [IWP]; Siganus lineatus (Valenciennes, 1835): Cr; [EIO-WP] PMBC 69 The fishes of southwestern Thailand

Sphyraena pinguis Günther, 1874: Cr; [IWP] Psenopsis obscura Headrich, 1967: OShr; [IM] Sphyraena putnamae Jordan & Seale, 1905: Mg, Sg, OShr; Nomeidae [IWP] Cubiceps pauciradiatus Günther, 1872: OShr; [CIR]; Sphyraena qenie Klunzinger, 1870: Cr; [IP] PMBC Gempylidae Cubiceps whiteleggii (Waite, 1894): OShr; [IWP] Gempylus serpens Cuvier, 1829: OShr; [COS] Ariommatidae Neoepinnula orientalis (Gilchrist & von Bonde, 1924): OShr; Ariomma indicum (Day, 1870): OShr; [IWP]; PMBC [IWP]; PMBC Stromateidae Nesiarchus nasutus Johnson, 1862: OShr; [COS]; PMBC Pampus argenteus (Euphrasen, 1788): OShr; [NIO-WP]; Promethichthys prometheus (Cuvier, 1832): OShr; [COS] PMBC Rexea bengalensis (Alcock, 1884): OShr; [IWP] Pampus chinensis (Euphrasen, 1788): OShr; [NIO-WP] Thyrsitoides marleyi Fowler, 1929: OShr; [IWP] Caproidae Trichiuridae Antigonia rubescens (Günther, 1860): OShr; [WP] Eupleurogrammus muticus (Gray, 1831): OShr; [NIO-IM] Order Pleuronectiformes Tentoriceps cristatus (Klunzinger, 1884): OShr; [IWP]; Psettodidae PMBC Psettodes erumei (Bloch & Schneider, 1801): Mg, OShr; Trichiurus gangeticus Gupta, 1966: OShr; [EIO]; PMBC [IWP]; PMBC Trichiurus lepturus Linnaeus, 1758: Mg, Sg, OShr; [COS]; Citharidae PMBC Brachypleura novaezeelandiae Günther, 1862: OShr; Scombridae [NIO-IA]; PMBC Acanthocybium solandi (Cuvier, 1831): Cr, OShr; [COS] Paralichthyidae Auxis rochei rochei (Risso, 1810): OShr; [COS]; PMBC Pseudorhombus argus Weber, 1913: OShr; [IA] Auxis thazard thazard (Lacepède, 1800): OShr; [COS]; Pseudorhombus arsius (Hamilton, 1822): Mg, Sg, OShr; PMBC [IWP]; PMBC Euthynnus affinis (Cantor, 1849): Cr, OShr; [IP]; PMBC Pseudorhombus dupliciocellatus Regan, 1905: OShr; [IA]; Grammatorcynos bilineatus (Rüppell, 1836): Cr, OShr; PMBC [IP]; PMBC Pseudorhombus elevatus Ogilby, 1912: Mg, OShr; [IWP]; Gymnosarda unicolor (Rüppell, 1838): Cr, OShr; [IP]; PMBC PMBC Pseudorhombus malayanus Bleeker, 1866: Mg, OShr; Katsuwonus pelamis (Linnaeus, 1758): OShr; [COS]; [NIO-IM] PMBC Pseudorhombus quinquocellatus Weber & de Beaufort, Rastrelliger brachysoma (Bleeker, 1851): OShr; [WP]; 1929: OShr; [EIO-IA]; PMBC PMBC Pseudorhombus triocellatus (Schneider, 1801): OShr; [EIO- Rastrelliger faughni Matsui, 1967: OShr; [EIO-WP]; IA]; PMBC PMBC Bothidae Rastrelliger kanagurta (Cuvier, 1817): Cr, Mg, Sg, OShr; Arnoglossus aspilos (Bleeker, 1851): Cr; [NIO-IA]; PMBC [IP]; PMBC Arnoglossus macrolophus Alcock, 1889: OShr; [IWP] Scomberomorus commerson (Lacepède, 1800): Cr, Mg, Arnoglossus tapeinosoma (Bleeker, 1865): OShr; [NIO- OShr; [IP]; PMBC WP] Scomberomorus guttatus (Bloch & Schneider, 1801): OShr; Arnoglossus tenuis Günther, 1880: OShr; [WP]; PMBC [NIO-WP]; PMBC Asterorhombus intermedius (Bleeker, 1866): OShr; [IWP] Scomberomorus lineolatus (Cuvier, 1831): OShr; [NIO- Bothus pantherinus (Rüppell, 1830): OShr; [IP] IM] Chascanopsetta lugubris Alcock, 1894: OShr; [IWP] Thunnus alalunga (Bonnaterre, 1788): OShr; [COS] Engyprosopon grandisquama (Temminck & Schlegel, Thunnus albacares (Bonnaterre, 1788): OShr; [COS]; 1846): OShr; [IWP]; PMBC PMBC Engyprosopon latifrons (Regan, 1908): OShr; [IWP]; Thunnus obesus (Lowe, 1839): OShr; [COS] PMBC Thunnus tonggol (Bleeker, 1851): OShr; [IWP]; PMBC Engyprosopon maldivensis (Regan, 1908): OShr; [EIO-WP] Xiphiidae Grammatobothus polyophthalmus (Bleeker, 1866): OShr; Xiphias gladius Linnaeus, 1758: OShr; [COS] [EIO-WP]; PMBC Istiophoridae Laeops guentheri Alcock, 1890: OShr; [NIO-IM] Istiophorus platypterus (Shaw & Nodder, 1792): OShr; Laeops kitaharae (Smith & Pope, 1906): OShr; [IWP] [IP]; PMBC Poecilopsettidae Makaira indica (Cuvier, 1832): OShr; [IP] Poecilopsetta colorata Günther, 1880: OShr; [EIO-IM] Tetrapturus audax (Philippi, 1887): OShr; [IP] Samaridae Centrolophidae Samaris cristatus Gray, 1831: OShr; [IWP]; PMBC 70 Phuket mar. biol. Cent. Res. Bull.

Soleidae Balistapus undulatus (Park, 1797): Cr, OShr; [IP]; PMBC Aesopia cornuta Kaup, 1858: OShr; [IWP]; PMBC Balistoides conspicillum (Bloch & Schneider, 1801): Cr, Aseraggodes dubius Weber, 1913: OShr; [WP] OShr; [IP]; PMBC Aseraggodes satapoomini Randall & Desoutter-Meniger, Balistoides viridescens (Bloch & Schneider, 1801): Cr, 2007: Cr; [AS] OShr; [IP]; PMBC Brachirus pan (Hamilton, 1822): OShr; [EIO-IM] Canthidermis maculatus (Bloch, 1786): OShr; [COS]; Heteromycteris oculus (Alcock, 1889): OShr; [IWP]; PMBC PMBC Canthidermis rotundatus (Marion de Procé, 1822): OShr; Liachirus melanospilos (Bleeker, 1854): OShr; [EIO-WP] [IWP]; PMBC Pardachirus pavoninus (Lacepède, 1802): Cr, Sg, OShr; Melichthys indicus Randall & Klausewitz, 1973: Cr, OShr; [EIO-WP]; PMBC [IO]; PMBC Solea ovata Richardson, 1846: Sg, OShr; [EIO-IM]; PMBC Melichthys vidua (Solander, 1844): Cr; [IP] Synaptura commersonii (Lacepède, 1802): Sg, OShr; [NIO- Odonus niger (Rüppell, 1837): Cr, OShr; [IP]; PMBC IM]; PMBC Pseudobalistes flavimarginatus (Rüppell, 1829): Cr, OShr; Zebrias altipinnis (Alcock, 1890): OShr; [EIO] [IP] Zebrias quagga Kaup, 1858: Mg, Sg, OShr; [IWP]; PMBC Pseudobalistes fuscus (Bloch & Schneider, 1801): Cr, OShr; Zebrias synapturoides (Jenkins, 1910): OShr; [NIO-WP] [IP]; PMBC Cynoglossidae Rhinecanthus aculeatus (Linnaeus, 1758): Cr, OShr; [IP] Cynoglossus arel (Schneider, 1801): Mg, OShr; [NIO-IM] Rhinecanthus rectangulus (Bloch & Schneider, 1801): Cr, Cynoglossus bilineatus (Lacepède, 1802): Mg, OShr; [NIO- OShr; [IP] WP]; PMBC Rhinecanthus verrucosus (Linnaeus, 1758): Cr, OShr; [IWP] Cynoglossus cynoglossus (Hamilton, 1822): OShr; [NIO- Sufflamen bursa (Bloch & Schneider, 1801): Cr; [IP] IM]; PMBC Sufflamen chrysopterum (Bloch & Schneider, 1801): Cr, Cynoglossus kopsi (Bleeker, 1851): OShr; [NIO-WP] OShr; [IWP]; PMBC Cynoglossus lida (Bleeker, 1851): Mg, Sg, OShr; [IWP]; Sufflamen fraenatum (Latreille, 1804): Cr, OShr; [IP]; PMBC PMBC Cynoglossus lingua Hamilton, 1822: Mg, OShr; [IWP] Xanthichthys auromarginatus (Bennett, 1831): Cr, OShr; Cynoglossus macrolepidotus (Bleeker, 1851): OShr; [WP]; [IP] PMBC Monacanthidae Cynoglossus monopus (Bleeker, 1849): OShr; [EIO-IM] Acreichthys tomentosus (Linnaeus, 1758): Sg; [IA] Cynoglossus oligolepis (Bleeker, 1854): OShr; [WP]; Aluterus monoceros (Linnaeus, 1758): OShr; [CIR]; PMBC PMBC Aluterus scriptus (Osbeck, 1765): Cr, OShr; [CIR]; PMBC Cynoglossus puncticeps (Richardson, 1846): Sg, OShr; Amanses scopas (Cuvier, 1829): Cr; [IP]; PMBC [NIO-IA]; PMBC Anacanthus barbatus Gray, 1831: Sg, OShr; [NIO-IA] Cynoglossus semifasciatus Day 1877: Mg, Sg, OShr; [EIO]; Cantherhines dumerilli (Hollard, 1854): Cr, OShr; [IP] PMBC Cantherhines multilineatus (Tanaka, 1918): OShr; [IWP; Cynoglossus sibogae Weber, 1913: OShr; [IM]; PMBC PMBC Paraplagusia bilineata (Bloch, 1787): OShr; [IWP] Cantherhines pardalis (Rüppell, 1837): Cr, OShr; [IP]; Paraplagusia blochii (Bleeker, 1851): OShr; [NIO-WP] PMBC Symphurus septemstriatus (Alcock, 1891): OShr; [EIO- Monacanthus chinensis Osbeck, 1765: Cr, Mg, Sg, OShr; IM] [WP]; PMBC Order Tetraodontiformes Oxymonacanthus longirostris (Bloch & Schneider, 1801): Triacanthodidae Cr; [IP]; PMBC Halimochirurgus alcocki Weber, 1913: OShr; [IWP] Paraluteres sp.1: Cr; [AS] Tydemania navigatoris Weber, 1913: OShr; [IWP]; PMBC Paramonacanthus japonicus (Tilesius, 1810): OShr; [EIO- Triacanthidae WP] Pseudotriacanthus strigilifer (Cantor, 1849): OShr; [NIO- Paramonacanthus tricuspis (Hollard, 1854): Mg, Sg, OShr; IA]; PMBC [EIO]; PMBC Triacanthus biaculeatus (Bloch, 1786): Mg, Sg, OShr; Pseudomonacanthus macrurus (Bleeker, 1857): Mg, Sg, [NIO-WP]; PMBC OShr; [IWP]; PMBC Triacanthus nieuhofi Bleeker, 1852: Sg, OShr; [EIO-IA] Pseudomonacanthus peroni (Hollard, 1854): OShr; [IA] Trixiphichthys weberi (Chaudhuri, 1910): OShr; [EIO-IA]; Ostraciidae PMBC Lactoria cornuta (Linnaeus, 1758): Cr, Mg, Sg, OShr; [IP]; Balistidae PMBC Abalistes stellatus (Anonymous, 1798): Cr, OShr; [IWP]; Lactoria diaphana (Bloch & Schneider, 1801): Mg, OShr; PMBC [IP] 71 The fishes of southwestern Thailand

Ostracion cubicus Linnaeus, 1758: Cr, Mg, Sg, OShr; [IP]; Chelonodon patoca (Hamilton, 1822): Mg, Sg, OShr; [IWP]; PMBC PMBC Ostracion meleagris Shaw, 1796: Cr; [IP]; PMBC Lagocephalus gloveri Abe & Tabeta, 1983: OShr; [IWP] Ostracion nasus Bloch, 1785: OShr; [EIO-WP]; PMBC Lagocephalus inermis (Temminck & Schlegel, 1844): OShr; Ostracion rhinorhynchus Bleeker, 1852: Cr, OShr; [IWP] [IWP]; PMBC Ostracion solorensis Bleeker, 1853: Cr; [IWP] Lagocephalus lunaris (Bloch & Schneider, 1801): Mg, Sg, Tetrosomus gibbosus (Linnaeus, 1758): Cr, OShr; [IWP]; OShr; [IWP]; PMBC PMBC Lagocephalus scleratus (Gmelin, 1789): OShr; [IWP]; Tetrosomus reipublicae (Whitley, 1930): OShr; [IWP] PMBC Tetraodontidae Lagocephalus spadiceus (Richardson, 1844): OShr; [IWP] Amblyrhynchotes honckenii (Bloch, 1795): OShr; [IWP] Takifugu oblongus (Bloch, 1786): Mg, Sg, OShr; [IWP]; Arothron caeruleopunctatus Matsuura, 1994: Cr, Mg, Sg; PMBC [EIO-WP] Tetraodon fluviatilis Hamilton, 1822: OShr; [EIO-IM]; Arothron hispidus (Linnaeus, 1758): Cr, Mg, Sg, OShr; PMBC [IP]; PMBC Tetraodon nigroviridis Procé, 1822: Mg, Sg, OShr; [EIO- Arothron immaculatus (Bloch & Schneider, 1801): Cr, OShr; WP]; PMBC [IWP]; PMBC Torquigener hypselogeneion (Bleeker, 1852): Cr; [IWP]; Arothron manilensis (Marion de Porcé, 1822): Cr, Sg, OShr; PMBC [WCP] Diodontidae Arothron mappa (Lesson, 1830): Cr, OShr; [IP]; PMBC Cyclichthys orbicularis (Bloch, 1785): Mg, Sg, OShr; Arothron nigropunctatus (Bloch & Schneider, 1801): Cr, [IWP]; PMBC OShr; [IP]; PMBC Diodon eydouxii Brisout de Barneville, 1846: OShr; [IWP] Arothron reticularis (Bloch & Schneider, 1801): Mg, OShr; Diodon holocanthus Linnaeus, 1758: OShr; [CIR]; PMBC [IWP]; PMBC Diodon hystrix Linnaeus, 1758: Cr, OShr; [CIR]; PMBC Arothron stellatus (Bloch & Schneider, 1801): Cr, OShr; Diodon liturosus Shaw, 1804: Cr, Sg, OShr; [IP]; PMBC [IP]; PMBC Molidae Canthigaster smithae Allen & Randall, 1977: Cr; [IO] Masturus lanceolatus (Lienard, 1840): OShr; [COS] Canthigaster solandri (Richardson, 1844): Cr, OShr; [IP]; Mola mola (Linnaeus, 1758): OShr; [COS]; PMBC PMBC 72 Phuket mar. biol. Cent. Res. Bull.

Appendix B

Selected taxonomic literature pertaining to the records of fishes from southwestern Thailand, the Andaman Sea

Class Myxini Wongratana, T. 1983. Eptatretus indrambaryai, a new species of hagfish (Myxinidae) from the Andaman Sea. Nat Hist. Bull. Siam Soc. 31(2): 139–150. Class Chondrichthyes Compagno, L.J.V.and P.R. Last. 2008. A new species of wedgefish, Rhynchobatus palpebratus sp. nov. (Rhynchobatoidei: Rhynchobatidae), from the Indo-West Pacific. In: P.R. Last, W.T. White and J.J. Pogonoski (eds.). Descriptions of New Australian Chondrichthyans. CISRO Marine and Atmospheric Research Paper No. 022. pp. 227–240. Krajangdara, T., S. Puteeka, R. Phoonsawat, S. Vibunpant, R. Sojittosakul and R. Chayakul. 2006. Sharks found in Thai waters and adjacent areas. Marine Fisheries Research and Development Bureau, Department of Fisheries. Technical Paper No. 17/2006.99p.(inThai) Monkolprasit, S. 1984. The cartilaginous fishes (Class Elasmobranchii) found in Thai waters and adjacent areas. Department of Fishery Biology, Faculty of Fisheries, Kasetsart University, Bangkok, Thailand. 175 p. Class Actinopterygii Order Albuliformes Hidaka, K., Y. Iwatsuki and J.E. Randall. 2008. A review of the Indo-Pacific bonefishes of the Albula argentea complex, with a description of a new species. Ichthyol. Res. 55(1): 53–64. Order Anguilliformes McCosker, J.E. 1972. Two new genera and two new species of western Pacific snake-eels (Apodes: Ophichthidae). Proc. Cal. Acad. Sci. 39(10): 111–119. McCosker, J.E. and J.E. Randall. 2001. Revision of the snake-eel genus Brachysomophis (Anguilliformes: Ophichthidae), with descriptions of two new species and comments on the species of Mystriophis. Indo-Pacific Fishes 33:1– 32, 2 pls. Böhlke, E.B. 2000. Notes on the identity of small, brown, unpatterned Indo-Pacific moray eels, with descriptions of three new species (Anguilliformes: Muraenidae). Pac. Sci. 54(4): 395–416. Böhlke, E.B. and J.E. Randall. 2000. A review of the moray eels (Anguilliformes; Muraenidae) of the Hawaiian Islands, with descriptions of two new species. Proc. Acad. nat. Sci. Philad. 150: 203–278, 9 pls. Order Clupeiformes Whitehead, P.J.P. 1985. FAO Species Catalogue. Vol. 7. Clupeoid fishes of the world (suborder Clupeoidei). An annotated and illustrated catalogue of the herrings, sardines, pilchards, sprats, shads, anchovies, and wolf-herrings. Part 1 – Chirocentridae, Clupeidae and Pristigasteridae. FAO Species Synopsis No. 125,Vol.7, Pt. 1: 1–303. Whitehead, P.J.P., G.J. Nelson and T. Wongratana. 1988. FAO Species Catalogue. Vol. 7. Clupeoid fishes of the world (suborder Clupeoidei). An annotated and illustrated catalogue of the herrings, sardines, pilchards, sprats, shads, anchovies, and wolf-herrings. Part 2 – Engraulididae. FAO Species Synopsis No. 125,Vol.7,Pt.2:305– 579. Order Ophidiiformes Markle, D.F. and J.E. Olney. 1990. Systematics of the pearlfishes (Pisces: Carapidae). Bull. Mar. Sci. 47(2): 269–410. Møller, P.R. and W. Schwarzhans. 2008. Review of the Dinematichthyini (Teleostei: Bythitidae) of the Indo-west Pacific. Part IV. Dinematichthys and two new genera with descriptions of nine new species. The Beagle, Records of the Museum of Art Galleries of the Northern Territory 24: 87–146. Nielsen, J.G. and Y. Machida. 1988. Revision of the Indo-Pacific bathyal fish genus Glyptophidium (Ophidiiformes, Ophidiidae). Jap. J. Ichthyol. 35(3): 289–319. Order Lophiiformes Bradbury, M.G. 1988. Rare fishes of the deep-sea genus Halieutopsis: a review with descriptions of four new species (Lophiiformes: Ogcocephalidae). Fieldiana, Zool. 44: 1–22. Order Atheriniformes Kimura, S., D. Golani, Y. Iwatsuki, M. Tabuchi and T. Yoshino. 2007. Redescriptions of the Indo-Pacific atherinid fishes Atherinomorus forskalii, Atherinomorus lacunosus,andAtherinomorus pinguis. Ichthyol. Res. 54(2): 145– 159. Parenti, L.R. 1989. A phylogenetic revision of the phallostethid fishes (Atherinomorpha, Phallostethidae). Proc. Cal. Acad. Sci. 46(11): 243–277. Order Beloniformes 73 The fishes of southwestern Thailand

Parenti, L.R. 2008. A phylogenetic analysis and taxonomic revision of ricefishes, Oryzias and relatives (Beloniformes, Andrianichthyidae). Zool. J. Linn. Soc. 154: 494–610. Order Beryciformes Randall, J.E. 1998. Revision of the Indo-Pacific squirrelfishes (Beryciformes: Holocentridae: Holocentrinae) of the genus Sargocentron, with descriptions of four new species. Indo-Pacific Fishes 27: 1–105, 11 pls. Order Gasterosteiformes Dawson, C.E. 1985. Indo-Pacific Pipefishes (Red Sea to the Americas). The Gulf Coast Research Laboratory, Ocean Springs, Mississippi. 230 p. Order Scorpaeniformes Imamura, H. 1996. Phylogeny of the family Platycephalidae and related taxa (Pisces: Scorpaeniformes). Species Diversity 1(2): 123–233. Knapp, L.W. and T. Wongratana. 1987. Sorsogona melanoptera, a new flathead fish from the northern Indian Ocean (Teleostei: Platycephalidae). Proc. Biol. Soc. Wash. 100(2): 381–385. Poss, S.G. and W.N. Eschmeyer. 1980. Xenaploactis, a new genus for Prosopodasys asperrimus Günther (Pisces: Aploactinidae), with descriptions of two new species. Proc. Cal. Acad. Sci. 42(8): 287–293. Randall, J.E. and W.N. Eschmeyer. 2001. Revision of the Indo-Pacific scorpionfish genus Scorpaenopsis, with descriptions of eight new species. Indo-Pacific Fishes 34: 1–79, 12 pls. Wongratana, T. 1975. A record of a flathead fish, Thysanophrys papillolabium Schultz, in the Andaman Sea, with a key to fourteen species of the Platycephalidae from Thai waters (Pisces: Platycephalidae). Phuket mar. biol. Cent. Res. Bull. 7:1–9. Order Perciformes Family Acropomatidae Okamoto, M. and H. Ida. 2002. Acropoma argentistigma, a new species from the Andaman Sea, off southern Thailand (Perciformes: Acropomatidae). Ichthyol. Res. 49(3): 281–285. Family Serranidae Randall, J.E. and C.C. Baldwin. 1997. Revision of the serranid fishes of the subtribe Pseudogrammina, with descriptions of five new species. Indo-Pacific Fishes 26: 1–56, 1 pl. Randall, J.E. and P.C. Heemstra. 1991. Revision of Indo-Pacific groupers (Perciformes: Serranidae: Epinephelinae), with descriptions of five new species. Indo-Pacific Fishes 20: 1–332, 41 pls. Randall, J.E. and P.C. Heemstra. 2007. Meganthias filiferus, a new species of anthiine fish (Perciformes: Serranidae), from the Andaman Sea off southwestern Thailand. Phuket mar. biol. Cent. Res. Bull. 68:5–9. Randall, J.E. and R. Lubbock. 1981. A revision of the serranid fishes of the subgenus Mirolabrichthys (Anthiinae: Anthias), with descriptions of five new species. Contr. Sci. 333: 1–27. Randall, J.E. and U. Satapoomin. 2000. Cephalopholis polyspila, a new species of grouper (Perciformes: Serranidae: Epinephelinae) from southwestern Thailand and Sumatra. Phuket mar. biol. Cent. Res. Bull. 63:1–7. Family Pseudochromidae Gill, A.C. 2004. Revision of the Indo-Pacific dottyback fish subfamily Pseudochrominae (Perciformes: Pseudochromidae). Smithiana Monograph 1. 213 p, 12 pls. Winterbottom, R. and J.E. Randall. 1994. Two new species of congrogadins (Teleostei: Pseudochromidae), with range extensions for four other species. Can. J. Zool. 72: 750–756. Family Plesiopidae Mooi, R.D. 1995. Revision, phylogeny, and discussion of biology and biogeography of the fish genus Plesiops (Perciformes: Plesiopidae). Royal Ontario Museum Life Sciences Contributions No. 159.iv+107p. Family Opistognathidae Smith-Vaniz, W.F. 2009. Three new species of Indo-Pacific jawfishes (Opistognathus: Opistognathidae), with the posterior end of the upper jaw produced as a thin flexible lamina. Aqua, Int. J. Ichthyol. 15(2): 69–108. Family Apogonidae Gon, O. 1996. Revision of the cardinalfish subgenus Jaydia (Perciformes, Apogonidae, Apogon). Trans. Roy. Soc. S. Afr. 51: 147–194. Gon, O. 2000. The taxonomic status of the cardinalfish species Apogon niger, A. nigripinnis, A. pharaonis, A. sialis,and related species (Perciformes: Apogonidae). J.L.B. Smith Inst. Ichthyol. Spec. Publ. 65: 1–20. Gon, O. and J.E. Randall. 2003. Revision of the Indo-Pacific cardinalfish genus Archamia (Perciformes: Apogonidae), with description of a new species. Indo-Pacific Fishes 35: 1–49, 3 pls. Fraser, T.H. 2005. A review of the species in the Apogon fasciatus group with a description of a new species of cardinalfish from the Indo-west Pacific (Perciformes: Apogonidae). Zootaxa 924: 1–30. Fraser, T.H. 2008. Cardinafishes of the genus Nectamia (Apogonidae, Perciformes) from the Indo-Pacific region with descriptions of four new species. Zootaxa 1691: 1–52. 74 Phuket mar. biol. Cent. Res. Bull.

Fraser, T.H. and E.A. Lachner. 1985. A revision of the cardinalfish subgenera Pristiapogon and Zoramia (Genus Apogon) of the Indo-Pacific region (Teleostei: Apogonidae). Smithson. Contr. Zool. 42.47p. Randall, J.E., T.H. Fraser and E.A. Lachner. 1990. On the validity of the Indo-Pacific cardinalfishes Apogon aureus (Lacepède) and A. fleurieu (Lacepède), with description of a related new species from the Red Sea. Proc. Biol. Soc. Wash. 103(1): 39–62. Family Sillaginidae McKay, R.J. 1985. A revision of the fishes of the family Sillaginidae. Mem. Qd. Mus. 22(1): 1–73. Family Leiognathidae Kimura, S., T. Ito, T. Peristiwady, Y. Iwatsuki and T. Yoshino. 2005. The Leiognathus splendens complex (Perciformes: Leiognathidae) with the description of a new species, Leiognathus kupanensis Kimura and Peristiwady. Ichthyol. Res. 52(3): 275–291. Kimura, S., R. Kimura and K. Ikejima. 2008. Revision of the genus Nuchequula with descriptions of three new species (Perciformes: Leiognathidae). Ichthyol. Res. 55(1): 22–42. Kimura, S., T. Yamashita and Y. Iwatsuki. 2000. A new species, Gazza rhombea, from the Indo-west Pacific, with a redescription of G.achlamys Jordan & Starks, 1917 (Perciformes: Leiognathidae). Ichthyol. Res. 47(1): 1–12. Mochizuki, K. and M. Hayashi. 1989. Revision of the leiognathid fishes of the genus Secutor, with two new species. Sci. Rept. Yokosuka City Mus. 37: 83–95. Yamashita, T., S. Kimura and Y.Iwatsuki. 1998. Validity of the leiognathid fish, Gazza dentex (Valenciennes in Cuvier & Valenciennes, 1835), with designation of a lectotype, and redescription of G. m in u t a (Bloch, 1795). Ichthyol. Res. 45(3): 271–280. Family Lutjanidae Allen, G.R. and F.H. Talbot. 1985. Review of the snappers of the genus Lutjanus (Pisces: Lutjanidae) from the Indo- Pacific, with description of a new species. Indo-Pacific Fishes 11: 1–87, 10 pls. Family Caesionidae Carpenter, K.E. 1987. Revision of the Indo-Pacific fish family Caesionidae (Lutjanoidea), with descriptions of five new species. Indo-Pacific Fishes 15: 1–56, 7 pls. Family Gerreidae Iwatsuki, Y. and P.C. Heemstra. 2001. Gerres phaiya: a new species of gerreid fish (Teleostei: Perciformes: Gerreidae) from India, with comments on Gerres poieti and the Gerres erythrourus complex. Copeia 2001(4): 1043– 1049. Iwatsuki, Y., S. Kimura, H. Kishimoto and T. Yoshino. 1996. Validity of the gerreid fish, Gerres macracanthus Bleeker, 1854, with designation of lectotype, and designation of a neotype for G. filamentosus Cuvier, 1829. Ichthyol. Res. 43(4): 417–429. Iwatsuki, Y., S. Kimura and T. Yoshino. 1999a. Description of Gerres chrysops sp. nov. from Thailand and redescription of Gerres setifer (Hamilton, 1822) and G. decacanthus (Bleeker, 1865) (Perciformes: Gerreidae). Ichthyol. Res. 46(1): 27–41. Iwatsuki, Y., S. Kimura and T. Yoshino. 1999b. Redescription of Gerres baconensis (Evermann & Seale, 1907), G. equulus Temminck & Schlegel, 1844 and G. o ye n a (Forsskål, 1775), included in the “G. o ye n a complex”, with notes on other related species (Perciformes: Gerreidae). Ichthyol. Res. 46(4): 377–395. Iwatsuki, Y., S. Kimura and T. Yoshino. 2001. Redescription of Gerres longirostris (Lacepède, 1801) and Gerres oblongus Cuvier in Cuvier and Valenciennes, 1830, included in the Gerres longirostris complex (Perciformes: Gerreidae). Copeia 2001(4): 954–965. Iwatsuki, Y., S. Kimura and T. Yoshino. 2007. A review of the Gerres subfasciatus complex from the Indo-West Pacific, with three new species (Perciformes: Gerreidae). Ichthyol. Res. 54(2): 168–185. Family Haemulidae Johnson, J.W., J.E. Randall and S.F. Chenoweth. 2001. Diagramma melanacrum, new species of haemulid fish from Indonesia, Borneo and the Philippines with a generic review. Mem. Qd. Mus. 46(2): 657–676. Iwatsuki, Y. and B.C. Russell. 2006. Revision of the genus Hapalogenys (Teleostei: Perciformes) with two new species from Indo-West Pacific. Mem. Mus. Vict. 63(1): 29–46. Iwatsuki, Y., U. Satapoomin and K. Amaoka. 2000. New species: Hapalogenys merguiensis (Teleostei; Perciformes) from Andaman Sea. Copeia 2000(1): 129–139. McKay, R.J. and U. Satapoomin. 1994. Pomadasys andamanensis, a new species of haemulid fish from Thailand. Phuket mar. biol. Cent. Res. Bull. 59:1–4. Satapoomin, U and J.E. Randall. 2000. Plectorhinchus macrospilus, a new species of sweetlip (Perciformes: Haemulidae) from the southwestern Thailand, Andaman Sea. Phuket mar. biol. Cent. Res. Bull. 63: 9–16. Family Nemipteridae 75 The fishes of southwestern Thailand

Russell, B.C. 1986. Review of the western Indian Ocean species of Nemipterus Swainson 1839, with description of a new species. Senckenberg. biol. 67(1/3): 19–35. Russell, B.C. 1991. On the validity of Nemipterus furcosus (Valenciennes) (Nemipteridae). Cybium 15(1): 35–41. Russell, B.C. and D. Golani. 1993. A review of the fish genus Parascolopsis (Nemipteridae) of the western Indian Ocean, with description of a new species from the northern Red Sea. Israel J. Zool. 39: 337–347. Family Polynemidae Feltes, R.M. 1991. Revision of the polynemid fish genus Filimanus, with description of two new species. Copeia 1991(2): 302–322. Motomura, H. 2002. Revision of the Indo-Pacific threadfin genus Polydactylus (Perciformes, Polynemidae) with a key to the species. Bull. Natl. Sci. Mus., Ser. A. 28(3): 171–194. Motomura, H., Y. Iwatsuki, S. Kimura and T. Yoshino. 2001. A new species, Polydactylus siamensis, from Thailand and redescription of P. plebeius (Broussonet, 1782) with designation of a neotype (Perciformes: Polynemidae). Ichthyol. Res. 48(2): 117–126. Motomura, H., Y. Iwatsuki and T. Yoshino. 2002. Revision of the Indo-Pacific polynemid fish genus Eleutheronema (Teleostei: Perciformes). Ichthyol. Res. 49(1): 47–61. Motomura, H., U. Satapoomin and Y. Iwatsuki. 2000. A new record of the threadfin, Filimanus perplexa Feltes, 1991 (Perciformes: Polynemidae), from the Andaman Sea, Thailand. Phuket mar. biol. Cent. Res. Bull. 63: 17–20. Family Sciaenidae Sasaki, K. 1992. Two new and two resurrected species of sciaenid genus Johnius (Johnius) from the west Pacific. Jap. J. Ichthyol. 39(3): 191–199. Sasaki, K. 1996. Sciaenid fishes of the Indian Ocean (Teleostei, Perciformes). Mem. Fac. Sci. Kochi Univ. Ser. D (Biol.) 16/17: 83–95. Family Mullidae Randall, J.E. 2004. Revision of the goatfish genus Parupeneus (Perciformes: Mullidae), with descriptions of two new species. Indo-Pacific Fishes 36: 1–64, 16 pls. Family Kuhliidae Randall, J.E. and H.A. Randall. 2001. Review of the fishes of the genus Kuhlia (Perciformes: Kuhliidae) of the central Pacific. Pac. Sci. 55(3): 227–256. Family Pomacentridae Allen, G.R. 1991. Damselfishes of the World. Mergus Publishers, Melle, Germany. 271 p. Allen, G.R. and A.R. Emery. 1985. A review of the pomacentrid fishes of the genus Stegastes from the Indo-Pacific, with descriptions of two new species. Indo-Pacific Fishes 3: 1–31, 3 pls. Allen, G.R. and M.V. Erdmann. 2005. Chromis xouthos, a new species of damselfish (Pomacentridae) from the east Andaman Sea and central Indian Ocean. Aqua, J. Ichthyol. Aquat. Biol. 10(3): 89–94. Allen, G.R. and J.E. Randall. 2002. A review of the leucogaster species complex of the Indo-Pacific pomacentrid genus Amblyglyphidodon, with descriptions of two new species. Aqua, J. Ichthyol. Aquat. Biol. 5(4): 139–152. Randall, J.E. and G.R. Allen. 2005. Neopomacentrus sororius, a new species of damselfish from the Indian Ocean, with description of a neotype for its sister species, N. azysron (Bleeker). Aqua, J. Ichthyol. Aquat. Biol. 10(2): 73– 80. Family Labridae Gomon, M.F. 2006. A revision of the labrid fish genus Bodianus with descriptions of eight new species. Rec. Aust. Mus. Suppl. 30: 1–133. Randall, J.E. 1996. Second revision of the labrid fish genus Leptojulis, with descriptions of two new species. Indo- Pacific Fishes. 24: 1–20, 3 pls. Randall, J.E. 1999. Revision of the Indo-Pacific labrid fishes of the genus Pseudocheilinus, with descriptions of three new species. Indo-Pacific Fishes. 28: 1–34, 2 pls. Randall, J.E. 1999. Revision of the Indo-Pacific labrid fishes of the genus Coris, with descriptions of five new species. Indo-Pacific Fishes. 29: 1–74, 22 pls. Randall, J.E. 2000. Revision of the Indo-Pacific labrid fishes of the genus Stethojulis, with descriptions of two new species. Indo-Pacific Fishes. 31: 1–42, 6 pls. Randall, J.E. and Lubbock, R. 1981. Labrid fishes of the genus Paracheilinus, with descriptions of three new species from the Philippines. Jap. J. Ichthyol. 28(1): 19–30, 2 pls. Randall, J.E. and M.M. Smith. 1982. A review of the labrid fishes of the genus Halichoeres of the western Indian Ocean, with descriptions of six new species. Ichthyol. Bull. J.L.B. Smith Inst. Ichthyol. 45: 1–26, 6 pls. Family Scaridae Satapoomin, U., R.H. Kuiter and J.E. Randall. 1994. First Record of the parrotfish Scarus viridifucatus from Thailand (the Andaman Sea) and Indonesia. Phuket mar. biol. Cent. Res. Bull. 59:5–9. 76 Phuket mar. biol. Cent. Res. Bull.

Westneat, M., U. Satapoomin and J.E. Randall. 2007. Scarus maculipinna, a new species of parrotfish (Perciformes, Scaridae) from eastern Indian Ocean. Zootaxa. 1628: 59–68. Family Ammodytidae Ida, H., P. Sirimontaporn and S. Monkolprasit. 1994. Comparative morphology of the fishes of the family Ammodytidae, with description of two new genera and two new species. Zool. Stud. 33(4): 251–277. Family Tripterygiidae Holleman, W. 2005. A review of the triple fish genus Enneapterygius (Blennioidei: Tripterygiidae) in the western Indain Ocean, with descriptions of four new species. Smithiana Bulletin. 5: 1–25, 2 pls. Holleman, W. 2006. Fishes of the Helcogramma steinitzi species group (Blennioidei: Tripterygiidae) from the Indain Ocean, with descriptions of two new species. Aqua, J. Ichthyol. Aquat. Biol. 11(3): 89–104. Williams, J.T. and J.C. Howe. 2003. Seven new species of the triplefin genus Helcogramma (Tripterygiidae) from the Indo-Pacific. Aqua, J. Ichthyol. Aquat. Biol. 7(4): 151–176. Family Blenniidae Bath, H. 1992. Revision der gattung Praealticus Schultz & Chapman 1960 (Pisces: Blenniidae). Senckenberg. biol. 72(4/ 6): 237–316. Carlson, B.A. 1980. A new Indo-Pacific fish of the genus Cirripectes (Blenniidae, Salariini). Pac. Sci. 34(4): 407–414. Smith-Vaniz, W.F. 1976. The saber-toothed blennies, tribe Nemophini (Pisces: Blenniidae). The Academy of Natural Sciences of Philadelphia. Monograph 19.vii+196p. Smith-Vaniz,W.F., U. Satapoomin and G.R.Allen. 2001. Meiacanthus urostigma, a new fangblenny from the northeastern Indian Ocean, with discussion and examples of mimicry in species of Meiacanthus (Teleostei: Blenniidae: Nemophini). Aqua, J. Ichthyol. Aquat. Biol. 5(1): 25–43. Springer, V.G. 1988. The Indo-Pacific blenniid fish genus Ecsenius. Smithson. Contr. Zool. 465.134p.,14pls. Springer, V.G. and M.F. Gomon. 1975. Revision of the blenniid fish genus Omobranchus with descriptions of three new species and notes on other species of the tribe Omobranchini. Smithson. Contr. Zool. 177.135p. Springer, V.G. and J.E. Randall. 1999. Ecsenius polystictus, new species of blenniid fish from Mentawai Islands, Indonesia, with notes on other species of Ecsenius. Revue fr. Aquariol. 26(1–2): 39–48. Springer, V.G. and J.T. Williams. 1994. The Indo-west Pacific blenniid fish genus Istiblennius reappraised: A revision of Istiblennius, Blenniella,andParalticus, new genus. Smithson. Contr. Zool. 565.193p. Williams, J.T. 1988. Revision and phylogenetic relationships of the blenniid fish genus Cirripectes. Indo-Pacific Fishes. 17: 1–78, 7 pls. Family Callionymidae Fricke, R. 1983. Revision of the Indo-Pacific genera and species of the dragonet family Callionymidae. Theses Zoologicae Vo l. 3.774p. Family Gobiidae Akihito and Meguro, K. 2000. Review of the Gobiid genus Cristatogobius found in Japan with description of a new species. Ichthyol. Res. 47(3): 249–261. Darumas, U. and P. Tantichodok. 2002. A new species of mudskipper (Gobiidae: Oxucercinae) from southern Thailand. Phuket mar. biol. Cent. Res. Bull. 64: 101–107. Hoese, D.F. and H.K. Larson. 1994. Revision of the Indo-Pacific gobiid fish genus Valenciennea, with descriptions of seven new species. Indo-Pacific Fishes. 23: 1–71, 6 pls. Lachner, E.A. and S.J. Karnella. 1980. Fishes of the Indo-Pacific fish genus Eviota with descriptions of eight new species (Teleostei: Gobiidae). Smithson. Contr. Zool. 315. 127 p. Lachner, E.A. and J.F. McKinney. 1978. A revision of the Indo-Pacific fish genus Gobiopsis with descriptions of four new species (Pisces: Gobiidae). Smithson. Contr. Zool. 262. 52 p. Larson, H.K. 1999. A review of the mangrove goby genus Hemigobius (Gobioidei, Gobiidae, Gobionellinae). The Beagle, Records of the Museums and Art Galleries of the Northern Territory. 15: 23–42. Larson, H.K. 1999. Allocation to Calamiana and redescription of the fish species Apocryptes variegates and Vaimosa mindora (Gobioidei: Gobiidae: Gobionellinae), with description of a new species. Raffles Bull. Zool. 47(1): 257–281. Larson, H.K. 2001. A revision of the gobiid fish genus Mugilogobius (Teleostei: Gobioidei). Rec. West. Aust. Mus. Suppl. No. 62: 1–233. Larson, H.K. 2005. A revision of the gobiid genus Stigmatogobius (Teleostei: Gobiidae), with descriptions of two new species. Ichthyol. Explor. Freshwaters. 16(4): 347–370. Larson, H.K. 2009. Review of the gobiid fish genera Eugnathogobius and Pseudogobiopsis (Gobioidei: Gobiidae: Gobionellinae), with descriptions of three new species. Raffles Bull. Zool. 57(1): 127–181. Murdy, E.O. 1985. A review of the gobiid fish genera Exyrias and Macrodontogobius, with description of a new species of Exyrias. Indo-Pacific Fishes. 10: 1–14, 2 pls. 77 The fishes of southwestern Thailand

Murdy, E.O. 1989. A taxonomic revision and cladistic analysis of the oxudercine gobies (Gobiidae: Oxudercinae). Rec. Aust.Mus.Suppl.11: 1–93. Murdy, E.O. and D.F. Hoese. 1985. Revision of the gobiid fish genus Istigobius. Indo-Pacific Fishes 4: 1–41, 3 pls. Polunin, N.V.C. and R. Lubbock. 1979. Five new prawn-associated gobies (Teleostei: Gobiidae) of the genus Amblyeleotris. Bull. Br. Mus. Nat. Hist. (Zool.). 36(4): 239–249. Randall, J.E. 2001. Five new Indo-Pacific gobiid fishes of the genus Coryphopterus.Zool.Stud.40(3): 206–225. Randall, J.E. and D.W. Greenfield. 2001. A preliminary review of the Indo-Pacific gobiid fishes of the genus Gnatholepis. Ichthyol. Bull. J.L.B. Smith Inst. Ichthyol. 69: 1–17, 2 pls. Randall, J.E. and D.F. Hoese. 1985. Revision of the Indo-Pacific dartfishes, genus Ptereleotris (Perciformes: Goioidei) Indo-Pacific Fishes. 7: 1–36, 4 pls. Shibukawa, K. and G.R.Allen. 2007. Review of the cheek-spine goby genus Gladiogobius (Actinopterigii, Perciformes, Gobiidae), with descriptions of two new species from the Indo-Pacific. Bull. Natl. Mus. Nat. Sci., Ser. A, 33(4): 193–206. Shibukawa, K. and U. Satapoomin. 2006. Myersina adonis, a new species of shrimp-associated goby (Pisces: Perciformes: Gobiidae) from the Andaman Sea. Bull. Natl. Sci. Mus., Ser. A, 32(1): 29–37. Shibukawa, K., T. Suzuki, H. Senou and K. Yano. 2005. Records of three shrimp-goby species (Teleostei, Perciformes, Gobiidae) from the Ryukyu Archipelago, Japan. Bull. Natl. Sci. Mus., Ser. A, 31(4): 191–204. Satapoomin, U. and R. Winterbottom. 2002. Redescription of the gobioid fish Cryptocentrus pavoninoides (Bleeker, 1849), with notes on sexual dichromatism in shrimp gobies. Aqua, J. Ichthyol. Aquat. Biol. 5(2): 53–64. Winterbottom, R. 2002. A redescription of Cryptocentrus crocatus Wongratana, a redefinition of Myersina Herre (Acanthopterygii; Gobiidae), a key to the species, and comments on relationships. Ichthyol. Res. 49(1): 69– 75. Winterbottom, R. and M. Burridge. 1993. Revision of the species of Priolepis possessing a reduced transverse pattern of cheek papillae and no predorsal scales (Teleostei: Gobiidae). Can. J. Zool. 71: 494–514. Winterbottom, R. and A.S. Harold. 2005. Gobiodon prolixus, a new species of gobiid fish (Teleostei: Perciformes: Gobiidae) from the Indo-west Pacific. Proc. Biol. Soc. Wash. 118(3): 582–589. Winterbottom, R. and L. Southcott. 2007. Two new species of the genus Trimma (Percomorpha: Gobiidae) from western Thailand. Aqua, Int. J. Ichthyol. 13(2): 69–76. Family Siganidae Woodland, D.J. 1990. Revision of the fish family Siganidae with descriptions of two new species and comments on distribution and biology. Indo-Pacific Fishes. 19: 1–136, 11 pls. Family Acanthuridae Randall, J.E. 1993. Acanthurus tristis,avalidIndianOceansurgeonfish(Perciformes: Acanthuridae). J.L.B. Smith Inst. Ichthyol. Spec. Publ. 54:1–8. Randall, J.E. and K.D. Clements. 2001. Second revision of the surgeonfish genus Ctenochaetus (Perciformes:Acanthuridae), with descriptions of two new species. Indo-Pacific Fishes. 32: 1–33, 6 pls. Family Sphyraenidae Doiuchi, R. and T. Nakabo. 2005. The Sphyraena obtusata group (Perciformes: Sphyraenidae) with a description of a new species from southern Japan. Ichthyol. Res. 52(2): 132–151 Order Pleuronectiformes Menon, A.G.K. 1977. A systematic monograph of the tongue soles of the genus Cynoglossus Hamilton-Buchanan (Pisces: Cynoglossidae). Smithson. Contr. Zool. 238.129p. Randall, J.E. and M. Desoutter-Meniger. 2007. Review of the soles of the genus Aseraggodes (Pleuronectiformes: Soleidae) from the Indo-Malayan region, with descriptions of nine new species. Cybium. 31(3): 301–331. Randall, J.E. and J.W. Johnson. 2007. Revision of the soleid fish genus Pardachirus. Indo-Pacific Fishes. 39: 1–22, 4 pls. Order Tetraodontiformes Hutchins, J.B. 1997. Review of the monacanthid fish genus Paramonacanthus, with descriptions of three new species. Rec. West. Aust. Mus. Suppl. No. 54.57p. Leis, J.M. 2006. Nomenclature and distribution of the species of the porcupinefish family Diodontidae. Mem. Mus. Vict. 63(1): 77–90. Randall, J.E. and W. Klausewitz. 1973. A review of the triggerfish genus Melichthys, with description of a new species from the Indian Ocean. Senckenberg. biol. 54(1/3): 57–69. 78 Phuket mar. biol. Cent. Res. Bull.