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NAT. CROAT. VOL. 14 No 2 99¿120 ZAGREB June 30, 2005

original scientific paper / izvorni znanstveni rad

SURVEY OF THE GASTERAL (FUNGI) OF CROATIA

ZDENKO TKAL^EC,ARMIN ME[I] &OLEG ANTONI]

Laboratory of Biocoenotic Research, Ru|er Bo{kovi} Institute, Bijeni~ka cesta 54, 10000 Zagreb, Croatia (E-mails: [email protected], [email protected], [email protected])

Tkal~ec, Z., Me{i}, A. & Antoni}, O.: Survey of the gasteral Basidiomycota (Fungi) of Croatia. Nat. Croat., Vol. 14, No. 2., 99–120, 2005, Zagreb. A survey of the gasteral Basidiomycota of Croatia is given. 68 species belonging to 26 genera are presented. Five genera and 18 species are reported as new to Croatia. For each species, the pub- lished and unpublished sources of data are given, as well as the collections in which the material is deposited. Key words: Biodiversity, mycobiota, bibliography

Tkal~ec, Z., Me{i}, A. & Antoni}, O.: Pregled utrobnja~a (Basidiomycota, Fungi) Hrvatske. Nat. Croat., Vol. 14, No. 2., 99–120, 2005, Zagreb. Dat je pregled gljiva utrobnja~a Hrvatske. Sadr`i 68 vrsta iz 26 rodova. Pet rodova i 18 vrsta prvi je put publicirano za podru~je Hrvatske. Uz svaku vrstu navedeni su publicirani i nepub- licirani izvori podataka, kao i zbirke u kojima je pohranjen sakupljeni materijal. Klju~ne rije~i: biolo{ka raznolikost, mikobiota, bibliografija

INTRODUCTION

The mycobiota of Croatia is poorly explored. The gasteral Basidiomycota are no exception since few mycologists have researched the group. The first data on this fungal group in Croatia were published by Stjepan SCHULZER (1857), the only Cro- atian mycologist in the 19th century. In the second half of the 20th century more in- tensive research started again. Significant contributions to the research on gasteral Basidiomycota during that period were made by Milica Torti}, Ivan Focht, Zdenko Tkal~ec and Armin Me{i}. Although it has been a century and a half since the be- ginning of mycological research in Croatia, no survey of the species of gasteral Basidiomycota has hitherto been made. Accordingly, we have prepared a survey of species of this fungal group from Croatia, including all published and most of the unpublished sources of data known to us.

Croatian Natural History Museum, Demetrova 1, Zagreb, Croatia 100 Tkal~ec, Z., Me{i}, A. & Antoni}, O.: Survey of the gasteral Basidiomycota (Fungi) of Croatia

MATERIAL, METHODS AND PRESENTATION

In this paper we treated all species of the class Gasteromycetes in the classic con- cept (see JÜLICH, 1984). The survey includes all the published records of species, subspecies and varieties of gasteral Basidiomycota from Croatia known to us. Re- cords from daily and weekly newspapers have been excluded, as well as records from HAFNER (1997) due to false data. This survey also includes, except for a few cases, all the unpublished records known and available to us. Two manuscripts by Stjepan Schulzer have not been included (one is kept at the Library of the Hungar- ian Academy of Sciences in Budapest, and the other at the National and Univer- sity Library in Zagreb), as well as notes by Ivan Focht, because it is not possible to interpret their data in a satisfactory way without thorough revision. We have also excluded unpublished records the identification of which was doubtful to the author himself. The survey includes data published or collected by 31 December 2004. The material known to us is deposited in the following herbaria and fun- garia. BEO – Herbarium, Botany Department, Natural History Museum of Serbia, Beograd, Serbia and Montenegro. BP – Herbarium, Botanical Department, Hungarian Natural History Mu- seum, Budapest, Hungary. CNF – Croatian National Fungarium, Croatian Mycological Society, Zagreb, Croatia. IB – Herbarium, Innsbruck University, Innsbruck, Austria. POZM – Herbarium, Department of Plant Ecology and Environment Protec- tion, Adam Mickiewicz University, PoznaBn, Poland. PRM – Herbarium, Mycological Department, National Museum in Prague, Prague, Czech Republic. ZA – Herbarium Croaticum, University of Zagreb, Faculty of Natural Sci- ences and Mathematics, Botany Department, Zagreb, Croatia.

Along with each presented species we specify the published and unpublished sources of records for Croatia, as well as the collections in which the collected ma- terial is deposited. In this way we give a more or less complete survey of the pub- lished sources of data, as well as a good survey of the unpublished sources of data on species of gasteral Basidiomycota in Croatia. We also specify synonyms and mis- applied names which the authors of records have used, as well as those used in modern taxonomic literature. We have tried to interpret all records on the basis of taxa names and dates of recording in the sense of modern . The older the record, the greater the need for revision. So far we have revised only a part of the revisable records. We have revised all our entries in this survey, as well as a smaller number of records by other authors indicated as such in the text. Species and varieties revised by us or by authors of taxonomic literature which we have used for interpretation of records are marked with a black dot. At the end of the data specified for such species we cite the literature (in brackets) on the basis of which we have identified or revised the records. The name of each genus is fol- Nat. Croat. Vol. 14(2), 2005 101 lowed by the literature (given in brackets) used for the interpretation, nomencla- ture and synonyms of records in the genus. SCHULZER (1867) corrects one mistake from the paper by SCHULZER et al. (1866), so we give that quote as »SCHULZER et al. (1866) + SCHULZER (1867)«. We have cited abbreviations of authors of taxa accord- ing to KIRK &ANSELL (1992) and abbreviations of journals in chapter »Literature cited« according to BRIDSON &SMITH (1991). The chapter »Excluded records« comprises: (a) records of species and infra- specific taxa we were unable to find in modern taxonomic literature, (b) records we could not interpret in the sense of modern classification, (c) records which are a nomen nudum, and (d) records which have been incorrectly recorded for Croatia or have been recorded for the territory which does not now belong to Croatia. When we could not check whether the author of the record had given the correct author citation, we have given it herein according to that author.

Marks and abbreviations: * – taxon reported from Croatia for the first time in this paper; • – at least part of records of taxon revised; (1) – number in brackets indicates synonym or misapplied name under which the author of the record has quoted the taxon; ( ) – empty brackets indicate that the author of the record has also quoted the taxon under the name we use; ? – question mark instead of an author of combination means that we could not find the author in the literature available to us; = ? – taxon not found in modern taxonomic literature; Exs. – collections in which the dried material (exsiccatum) is deposited; HR? – taxon most probably found in the territory of Croatia (locality not precisely quoted); Incl. – including; Lit. – literature; Misappl. – misapplied name; Ref. – references (sources of records of taxon); rev. – revised (when we have revised records of other authors); unp. – unpublished data.

RESULTS

In this paper we present 68 species of gasteral Basidiomycota belonging to 26 gen- era found on the territory of Croatia, 52 species having been revised by us. Five genera and 18 species are reported here as new to Croatia. 102 Tkal~ec, Z., Me{i}, A. & Antoni}, O.: Survey of the gasteral Basidiomycota (Fungi) of Croatia

SURVEY OF GENERA AND SPECIES

Alpova C.W. Dodge (Lit.: GROß et al., 1980; JÜLICH, 1984; KREISEL, 2001; MONTECCHI &SARASINI, 2000)

Alpova rubescens (Vittad.) Trappe Ref.: VR[^AJ, 2002

Astraeus Morgan (Lit.: ARNOLDS et al., 1995; DÖRFELT, 1985; GROß et al., 1980; JÜLICH, 1984; KREI- SEL, 2001; MOYERSOEN &DEMOULIN, 1996; PEGLER et al., 1995; PILÁT, 1958)

• (Pers. : Pers.) Morgan = hygrometricum Pers. : Pers. (1) = Astraeus stellatus (Scop.) E. Fisch. (2) = Geastrum stellatum (Scop.) Wettst. (3) Ref.: BLAGAI], 1931 (1); BO@AC, 1984; COURTECUISSE &DUHEM, 2000; FOCHT, 1972; FOCHT, 1974; GJURA{IN, 1898 (1); LANGE, 1974; MOESZ, 1938 (2); PICBAUER, 1928 (2); [KORI], 1928 (3); TORTI], 1964; TORTI], 1993; TORTI], 2002; VR[^AJ, 2002; unp. KOTLABA (according to unp. TORTI]); unp. MATO^EC; unp. TKAL^EC &ME[I]; unp. TORTI] ( )(2); SACCARDO, 1888 HR? (1) Exs.: BEO, CNF, PRM (Lit.: JÜLICH, 1984; PEGLER et al., 1995)

Bovista Pers. : Pers. (Lit.: ARNOLDS et al., 1995; CALONGE, 1998; GMINDER et al., 2000; GROß et al., 1980; JÜLICH, 1984; KREISEL, 1967; KREISEL, 2001; MOYERSOEN &DEMOULIN, 1996; PEGLER et al., 1995; PILÁT, 1958)

• aestivalis (Bonord.) Demoulin = Bovista colorata (Peck) Kreisel = furfuraceum Schaeff. (1) = Bovista polymorpha (Vittad.) Kreisel (nom. illeg.) (2) = Bovista pusilliformis (Kreisel) Kreisel Ref.: GJURA[IN, 1898 (1); MOESZ, 1938 (1); TORTI], 1993 (2); unp. TKAL^EC &ME[I] Exs.: CNF (Lit.: CALONGE, 1998; KREISEL, 1967; PEGLER et al., 1995)

Bovista aestivalis (Bonord.) Demoulin agg. Incl.: (Bonord.) Demoulin Bovista dermoxantha (Vittad.) De Toni Bovista furfuracea (J.F. Gmel. : Pers.) Pers. Nat. Croat. Vol. 14(2), 2005 103

Bovista pusilla (Batsch : Pers.) Pers. (1) = Lycoperdon pusillum Batsch : Pers. (2) Ref.: FOCHT, 1972 (2); FOCHT, 1974 (2); VR[^AJ, 2002 (1) Annotation: Modern authors distinguish two or three species here, but dis- agreements exist about species concepts, synonymy and nomenclature (see AR- NOLDS et al., 1995; CALONGE, 1998; KREISEL, 2001; MOYERSOEN &DEMOULIN, 1996; PEGLER et al., 1995).

• Bovista nigrescens Pers. : Pers. Ref.: BAR^I], 1982; BAR^I], 1996; BLAGAI], 1931; FOCHT, 1974; MOESZ, 1938; TORTI], 1993; VR[^AJ, 2002; unp. TKAL^EC &ME[I] Exs.: CNF (Lit.: KREISEL, 1967; PEGLER et al., 1995)

• Pers. : Pers. = Bovista nuciformis Wallr. (1) Ref.: FOCHT, 1974; GJURA[IN, 1898; SCHULZER et al., 1866; [KORI], 1928; TORTI], 1966 a; TORTI], 1966 b; VOUK &PEVALEK, 1916 (1); unp. KOTLABA (according to unp. TORTI]); unp. TKAL^EC &ME[I]; unp. TORTI];SCHULZER, 1857 HR? Exs.: CNF, PRM (Lit.: KREISEL, 1967; PEGLER et al., 1995)

* Bovista tomentosa (Vittad.) Quél. Ref.: unp. TORTI] Exs.: CNF

Calvatia Fr. = Kreisel (Lit.: ARNOLDS et al., 1995; CALONGE, 1998; DEMOULIN, 1970; GMINDER et al., 2000; GROß et al., 1980; JÜLICH, 1984; KREISEL, 2001; MOYERSOEN &DEMOULIN, 1996; PEGLER et al., 1995; PILÁT, 1958)

• excipuliformis (Scop. : Pers.) Perdeck = (Scop. : Pers.) Kreisel = Lycoperdon saccatum Vahl (1) Ref.: BAR^I], 1982; BAR^I], 1996; FOCHT, 1972; FOCHT, 1974; FOCHT, 1979; KRANJ- ^EV, 1985 (1); LISIEWSKA &TORTI], 1990; TORTI], 1993; TORTI], 2002; VR[^AJ, 2002; unp. LINDTNER; unp. MATO^EC; unp. TKAL^EC &ME[I]; unp. TORTI] Exs.: BEO, CNF (Lit.: JÜLICH, 1984; PILÁT, 1958)

• Calvatia utriformis (Bull. : Pers.) Jaap = (Bull. : Pers.) Kreisel 104 Tkal~ec, Z., Me{i}, A. & Antoni}, O.: Survey of the gasteral Basidiomycota (Fungi) of Croatia

= Lycoperdon utriforme Bull. : Pers. (1) = Calvatia caelata (Bull.) Morgan (2) = Lycoperdon caelatum Bull. (3) Ref.: BAR^I], 1989 (2); BAR^I], 1996 (2); FOCHT, 1979 (2); KRANJ^EV, 1985 (3); KRIVI^I], 1999; MOESZ, 1938 (2); MU@IC &BO@AC, 1997 (3); RAGUZIN, 2001; [KORI], 1928 (3); TORTI], 1968 b (2); TORTI], 1973 (2); TORTI], 1993; VOUK & PEVALEK, 1916 (1)(3); VR[^AJ, 1990; VR[^AJ, 2002; unp. LINDTNER (2); unp. MATO^EC; unp. TKAL^EC &ME[I]; unp. TORTI] (rev.) (2) Exs.: BEO (2), CNF (Lit.: CALONGE, 1998; JÜLICH, 1984)

Clathrus P. Micheli ex L. : Pers. = Anthurus Kalchbr. et MacOwan

(Lit.: ARNOLDS et al., 1995; CALONGE, 1998; DRING, 1980; GROß et al., 1980; JÜLICH, 1984; KREISEL, 2001; MOYERSOEN &DEMOULIN, 1996; PEGLER et al., 1995; PILÁT, 1958) *• Clathrus archeri (Berk.) Dring = Anthurus archeri (Berk.) E. Fisch. Ref.: unp. TKAL^EC &ME[I] Exs.: CNF (Lit.: JÜLICH, 1984; PEGLER et al., 1995; PILÁT, 1958) • Clathrus ruber P. Micheli ex Battarra : Pers. = Clathrus cancellatus Tourn. ex L. (1) Ref.: BAR^I], 1982; BAR^I], 1984; BAR^I], 1996; BLAGAI], 1923 (1); BLAGAI], 1931 (1); BO@AC, 1984 (1); BO@AC, 1986 (1); BO@AC, 1989 (1); BO@AC, 1993 (1); FOCHT, 1974 (1); LOHWAG, 1935; MOESZ, 1938 (1); PICBAUER, 1928 (1); RAGUZIN, 2000; TORTI], 1993; VR[^AJ, 2002; unp. MATO^EC (1); unp. PEINTNER; unp. TKAL^EC & ME[I]; unp. TORTI] Exs.: CNF, IB, ZA (Lit.: CALONGE, 1998; JÜLICH, 1984; PILÁT, 1958)

Crucibulum Tul. et C. Tul. (Lit.: ARNOLDS et al., 1995; CALONGE, 1998; GMINDER et al., 2000; GROß et al., 1980; JÜLICH, 1984; KREISEL, 2001; MORNAND, 1985; MOYERSOEN &DEMOULIN, 1996; PEGLER et al., 1995; PILÁT, 1958) • Crucibulum crucibuliforme (Scop.) V.S. White = Cyathus crucibulum Pers. : Pers. (1) = Nidularia crucibulum (Pers. : Pers.) Fr. (2) = Crucibulum laeve (Huds.) Kambly (3) = Crucibulum vulgare Tul. et C. Tul. (4) Nat. Croat. Vol. 14(2), 2005 105

Ref.: COURTECUISSE &DUHEM, 2000 (3); FOCHT, 1972 (3); GJURA[IN, 1898 (1); JAAP, 1916; LISIEWSKA &TORTI], 1990; RAC, 2003 (3); SCHULZER et al., 1866 (1); TORTI], 1964 (4); TORTI], 1966 a (4); TORTI], 1966 b (4); TORTI], 2002 ( )(3); unp. MATO^EC (3); unp. TKAL^EC &ME[I]; unp. TORTI] (3)(4); SCHULZER, 1857 HR? (2) Exs.: CNF (Lit.: CALONGE, 1998; JÜLICH, 1984)

Cyathus Haller : Pers. (Lit.: ARNOLDS et al., 1995; CALONGE, 1998; GMINDER et al., 2000; GROß et al., 1980; JÜLICH, 1984; KREISEL, 2001; MORNAND, 1985; MOYERSOEN &DEMOULIN, 1996; PEGLER et al., 1995; PILÁT, 1958)

• Cyathus olla (Batsch : Pers.) Pers. = Cyathus olla (Batsch : Pers.) Pers. var. nitidus (Roth : Pers.) Pers. (1) = Nidularia campanulata With. (2) = Cyathus sericeus (Schaeff.) Quél. (3) Ref.: GJURA[IN, 1898 (3); JAAP, 1916; KRANJ^EV, 1985 a; MOESZ, 1938 (3); RANDI], 1998; SCHULZER et al., 1866 (1); TORTI], 1966 a; TORTI], 1966 b; TORTI], 1968 b; TORTI], 1993; unp. MATO^EC; unp. TKAL^EC &ME[I]; unp. TORTI] (rev.); SCHULZER, 1857 HR? (2) Exs.: CNF (Lit.: CALONGE, 1998; GROß et al., 1980; JÜLICH, 1984)

*• Cyathus stercoreus (Schwein.) De Toni Ref.: unp. TKAL^EC &ME[I]; unp. TORTI] (rev.) Exs.: CNF (Lit.: CALONGE, 1998; GROß et al., 1980; JÜLICH, 1984)

• Cyathus striatus (Huds. : Pers.) Willd. = Nidularia striata (Huds. : Pers.) With. (1) = Cyathus hirsutus (Schaeff.) Quél. (2) Ref.: GJURA[IN, 1898 (2); JELI] &TORTI], 1973; KRANJ^EV, 1985; MOESZ, 1938; RANDI], 1998; SCHULZER, 1882; SCHULZER et al., 1866; [KORI], 1928; TORTI], 1964; TORTI], 1966 a; TORTI], 1966 b; TORTI], 1989; TORTI], 1993; TORTI], 2002; TORTI] &LISIEWSKA, 1978; VOUK &PEVALEK, 1915; VOUK &PEVALEK, 1916; unp. MATO^EC; unp. TKAL^EC &ME[I]; unp. TORTI];SCHULZER, 1857 HR? (1) Exs.: CNF (Lit.: CALONGE, 1998; GROß et al., 1980; JÜLICH, 1984)

*Disciseda Czern. (Lit.: ARNOLDS et al., 1995; CALONGE, 1998; GMINDER et al., 2000; GROß et al., 1980; JÜLICH, 1984; KREISEL, 2001; PILÁT, 1958) 106 Tkal~ec, Z., Me{i}, A. & Antoni}, O.: Survey of the gasteral Basidiomycota (Fungi) of Croatia

*• Disciseda bovista (Klotzsch) Henn. Ref.: unp. TKAL^EC &ME[I] Exs.: CNF (Lit.: CALONGE, 1998; JÜLICH, 1984; PILÁT, 1958)

*• Disciseda candida (Schwein.) Lloyd = Disciseda calva (Z. Moravec) Z. Moravec Ref.: unp. TKAL^EC &ME[I] Exs.: CNF (Lit.: CALONGE, 1998; JÜLICH, 1984; PILÁT, 1958)

*Endoptychum Czern. (Lit.: JÜLICH, 1984; KREISEL, 2001; MONTECCHI &SARASINI, 2000; PILÁT, 1958)

*• Endoptychum agaricoides Czern. Ref.: unp. TKAL^EC &ME[I] Exs.: CNF (Lit.: JÜLICH, 1984; MONTECCHI &SARASINI, 2000; PILÁT, 1958)

Geastrum P. Micheli ex Pers. : Pers. (Lit.: ARNOLDS et al., 1995; CALONGE, 1998; DÖRFELT, 1985; GMINDER et al., 2000; GROß et al., 1980; JÜLICH, 1984; KREISEL, 2001; MOYERSOEN &DEMOULIN, 1996; PEGLER et al., 1995; PILÁT, 1958; SUNHEDE, 1989)

Geastrum corollinum (Batsch) Hollós = Geastrum mammosum Cheval. (1) = Geastrum recolligens (With.) Desv. Ref.: SACCARDO, 1888 (1); SCHULZER et al., 1866 (1)

Geastrum coronatum Pers. : Pers. = Plecostoma limbatum (Fr.) ? (1) Ref.: SCHULZER et al., 1866 (1)

• Geastrum fimbriatum Fr. = Geastrum djakovense Schulzer (1) = Geastrum sessile (Sowerby ®) Pouzar (2) Misappl.: Geastrum rufescens Pers. : Pers. sensu auct. p. p. non Pers. Ref.: BAR^I], 1996; BLAGAI], 1931; KRANJ^EV, 1985; KRANJ^EV, 1988; KRANJ^EV, 1995; LISIEWSKA &TORTI], 1990; SACCARDO, 1888 (1); SCHULZER, 1882 (1); TOR- TI], 1964; TORTI], 1966 a; TORTI], 1966 b; TORTI], 2002; VOUK &PEVALEK, 1916; unp. LISIEWSKA (2); unp. MATO^EC; unp. TKAL^EC &ME[I]; unp. TORTI] Exs.: CNF, POZM (2) (Lit.: SUNHEDE, 1989) Nat. Croat. Vol. 14(2), 2005 107

Geastrum fornicatum (Huds.) Hook. Ref.: STOITZNER, 1869

*• Schwein. Ref.: unp. TKAL^EC &ME[I] Exs.: CNF (Lit.: SUNHEDE, 1989)

• Geastrum pectinatum Pers. : Pers. Ref.: KRANJ^EV, 1985; KRANJ^EV, 1988; KRANJ^EV, 1995; unp. TKAL^EC &ME[I] Exs.: CNF (Lit.: SUNHEDE, 1989)

*• Pers. : Pers. Ref.: unp. TKAL^EC &ME[I] Exs.: CNF (Lit.: SUNHEDE, 1989)

• Geastrum rufescens Pers. : Pers. = Geastrum schaefferi Vittad. (1) = Geastrum vulgatum Vittad. Ref.: BLAGAI], 1921; BLAGAI], 1931; FOCHT, 1972; KRANJ^EV, 1979; KRANJ^EV, 1985; KRANJ^EV, 1988; KRANJ^EV, 1995; TORTI], 1968 b (1); TORTI], 2002; unp. MATO^EC; unp. TKAL^EC &ME[I]; unp. TORTI] Exs.: CNF (Lit.: SUNHEDE, 1989)

*• Fr. Ref.: unp. TKAL^EC &ME[I] Exs.: CNF (Lit.: SUNHEDE, 1989)

• Geastrum schmidelii Vittad. Misappl.: Geastrum nanum Pers. sensu auct. non Pers. (nom. illeg.) (1) = Geastrum nanum Pers. var. coniferarum V.J. Stanìk (nom. illeg.) (2) Ref.: KRANJ^EV, 1988 (1); KRANJ^EV, 1995 (1); unp. TKAL^EC &ME[I]; unp. TOR- TI] (2) Exs.: CNF, PRM (2) (Lit.: SUNHEDE, 1989)

Geastrum striatum DC. = Geastrum bryantii Berk. (1) Ref.: BLAGAI], 1931 (1) 108 Tkal~ec, Z., Me{i}, A. & Antoni}, O.: Survey of the gasteral Basidiomycota (Fungi) of Croatia

• Geastrum triplex Jungh. Ref.: BAR^I], 1989; BAR^I], 1996; BO@AC, 1982; BO@AC, 1984; BO@AC, 1986; BO- @AC, 1989; BO@AC, 1993; FOCHT, 1972; KRANJ^EV, 1985; KRANJ^EV, 1988; MOESZ, 1938; TORTI], 1968 b; TORTI], 1993; TORTI], 2002; unp. LINDTNER; unp. MATO - ^EC; unp. TKAL^EC &ME[I]; unp. TORTI] Exs.: BEO, CNF (Lit.: SUNHEDE, 1989)

Langermannia Rostk. (Lit.: ARNOLDS et al., 1995; CALONGE, 1998; GROß et al., 1980; JÜLICH, 1984; KREISEL, 2001; MOYERSOEN &DEMOULIN, 1996; PEGLER et al., 1995; PILÁT, 1958; SCHULZER, 1882)

• Langermannia gigantea (Batsch : Pers.) Rostk. = Bovista gigantea (Batsch : Pers.) Nees (1) = (Batsch : Pers.) Lloyd (2) = Lycoperdon giganteum Batsch : Pers. (3) = Globaria bovista (L.) Quél. (4) = Lycoperdon bovista L. (5) Ref.: GJURA[IN, 1898 (1); KRANJ^EV, 1985 (3); KRANJ^EV, 1994 (3); KRANJ^EV, 1995 (3); LANGE, 1974 (2); MU@IC &BO@AC, 1997 (3); SCHULZER, 1866 (5); SCHULZER, 1882 (4); SCHULZER et al., 1866 (5); TORTI], 1980; unp. TKAL^EC &ME[I];BO@AC, 1984 HR? (3); BO@AC, 1986 HR? (3); BO@AC, 1989 HR?; BO@AC, 1993 HR? Exs.: CNF (Lit.: JÜLICH, 1984)

*Leucogaster R. Hesse (Lit.: GROß et al., 1980; JÜLICH, 1984; KREISEL, 2001; MONTECCHI &SARASINI, 2000; PEGLER et al., 1993)

*• Leucogaster nudus (Hazsl.) Hollós Ref.: unp. TKAL^EC &ME[I] Exs.: CNF (Lit.: JÜLICH, 1984; MONTECCHI &SARASINI, 2000)

Lycoperdon Tourn. ex L. : Pers. (Lit.: ARNOLDS et al., 1995; CALONGE, 1998; DEMOULIN, 1970; DEMOULIN, 1983; GMINDER et al., 2000; GROß et al., 1980; JÜLICH, 1984; KREISEL, 2001; MOYERSOEN &DEMOULIN, 1996; PEGLER et al., 1995; PILÁT, 1958)

Lycoperdon caudatum J. Schröt. = Lycoperdon pedicellatum Peck (nom. illeg.), non L. pedicellatum Batsch Ref.: VOUK &PEVALEK, 1916 Nat. Croat. Vol. 14(2), 2005 109

• Pers. : Pers. = Lycoperdon constellatum Fr. (1) Ref.: BAR^I], 1982; BAR^I], 1996; BLAGAI], 1921 ( )(1); KRANJ^EV, 1985; SCHULZER et al., 1866 (1); TORTI], 1964; TORTI], 1993; TORTI], 2002; unp. LINDTNER; unp. MATO^EC; unp. TKAL^EC &ME[I]; unp. TORTI];SCHULZER, 1857 HR? (1) Exs.: BEO, CNF (Lit.: CALONGE, 1998; JÜLICH, 1984) *Lycoperdon ericaeum Bonord. = Lycoperdon muscorum Morgan (1) Ref.: unp. TORTI] (1) Exs.: CNF • Lycoperdon lividum Pers. = Lycoperdon fuscum Bonord. (1) Ref.: VOUK &PEVALEK, 1916 (1); unp. TKAL^EC &ME[I] Exs.: CNF (Lit.: CALONGE, 1998; GMINDER et al., 2000; JÜLICH, 1984) • Pers. : Pers. = Lycoperdon velatum Vittad. (1) Ref.: BO@AC, 1984; FOCHT, 1974 (1); GJURA[IN, 1898; VR[^AJ, 2003; unp. TKAL^EC &ME[I] Exs.: CNF (Lit.: CALONGE, 1998; JÜLICH, 1984) Vittad. ex Moris et De Not. Misappl.: Lycoperdon papillatum Schaeff. sensu auct. non Schaeff. (1) Ref.: GMINDER et al., 2000; MOESZ, 1938 (1); TORTI], 1993 • Lycoperdon molle Pers. : Pers. = Lycoperdon cupricum Bonord. (1) Ref.: VOUK &PEVALEK, 1916 (1); unp. TKAL^EC &ME[I]; unp. TORTI] Exs.: CNF (Lit.: DEMOULIN, 1983; GMINDER et al., 2000; JÜLICH, 1984) *• Lycoperdon nigrescens (Pers.) Lloyd = Lycoperdon foetidum Bonord. Ref.: unp. MATO^EC; unp. TKAL^EC &ME[I] Exs.: CNF (Lit.: DEMOULIN, 1983; GMINDER et al., 2000; JÜLICH, 1984) • Pers. : Pers. = Lycoperdon gemmatum Batsch var. perlatum (Pers. : Pers.) Fr. (1) 110 Tkal~ec, Z., Me{i}, A. & Antoni}, O.: Survey of the gasteral Basidiomycota (Fungi) of Croatia

= Lycoperdon gemmatum Batsch (2) Ref.: BAR^I], 1982; BAR^I], 1996; BLAGAI], 1921 (2); FOCHT, 1972; FOCHT, 1974; GJURA[IN, 1898 (2); KRANJ^EV, 1985; LISIEWSKA &TORTI], 1990; LORINSER-JANDA, 1877 (2); MOESZ, 1938 (2); RAGUZIN, 2001; SCHULZER et al., 1866 (1); [KORI], 1928 (2); TORTI], 1964; TORTI], 1966 a; TORTI], 1993; TORTI], 2002; TORTI] &LISIEWSKA, 1978; VOUK &PEVALEK, 1916 (2); VR[^AJ, 2003; unp. MATO^EC; unp. TKAL^EC & ME[I]; unp. TORTI];SACCARDO, 1888 HR? (2) Exs.: CNF (Lit.: CALONGE, 1998; DEMOULIN, 1983; JÜLICH, 1984)

• Lycoperdon pyriforme Schaeff. : Pers. = Bovista bresadolae (Schulzer) De Toni (1) = Globaria bresadolae Schulzer (2) Ref.: FOCHT, 1979; KRANJ^EV, 1985; MOESZ, 1938; RAGUZIN, 2001; SCHULZER, 1882; SCHULZER, 1885 (2); SCHULZER et al., 1866 + SCHULZER, 1867; [KORI], 1928; TOR- TI], 1966 a; TORTI], 1966 b; TORTI], 1968 b; TORTI], 1989; TORTI], 1993; TORTI], 2002; VOUK &PEVALEK, 1915; unp. MATO^EC; unp. TKAL^EC &ME[I]; unp. TORTI];SACCARDO, 1888 HR? (1); SCHULZER, 1857 HR? Exs.: CNF (Lit.: CALONGE, 1998; JÜLICH, 1984)

Lycoperdon umbrinum Pers. : Pers. Ref.: FOCHT, 1974; LISIEWSKA &TORTI], 1990; TORTI], 2002; unp. LISIEWSKA; unp. MATO^EC; unp. TORTI] Exs.: CNF, POZM

Melanogaster Corda (Lit.: ARNOLDS et al., 1995; GROß et al., 1980; JÜLICH, 1984; KREISEL, 2001; MON- TECCHI &SARASINI, 2000; PEGLER et al., 1993; PILÁT, 1958)

Melanogaster ambiguus (Vittad.) Tul. et C. Tul. Ref.: FOCHT, 1972

*• Melanogaster broomeanus Berk. Ref.: unp. TKAL^EC &ME[I] Exs.: CNF (Lit.: GROß et al., 1980; JÜLICH, 1984; PILÁT, 1958)

*• Melanogaster vittadinii Soehner et A. Knapp Ref.: unp. TKAL^EC &ME[I] Exs.: CNF (Lit.: GROß et al., 1980; JÜLICH, 1984) Nat. Croat. Vol. 14(2), 2005 111

Mutinus Fr. (Lit.: ARNOLDS et al., 1995; CALONGE, 1998; GROß et al., 1980; JÜLICH, 1984; KREI- SEL, 2001; PEGLER et al., 1995; PILÁT, 1958)

• Mutinus caninus (Huds. : Pers.) Fr. Ref.: BLAGAI], 1917; MOESZ, 1938; TORTI], 1964; TORTI], 1993; TORTI], 2002; VOUK &PEVALEK, 1916; VR[^AJ, 2003; unp. MATO^EC; unp. TKAL^EC &ME[I]; unp. TORTI] Exs.: CNF (Lit.: JÜLICH, 1984; PILÁT, 1958)

Mycenastrum Desv.

(Lit.: ARNOLDS et al., 1995; CALONGE, 1998; GROß et al., 1980; JÜLICH, 1984; KREI- SEL, 2001; PILÁT, 1958)

• Mycenastrum corium (Guers.) Desv. = Pachyderma strossmayeri Schulzer (1) Ref.: SACCARDO, 1895 (1); SCHULZER, 1876 (1); unp. TKAL^EC &ME[I] Exs.: CNF (Lit.: CALONGE, 1998; JÜLICH, 1984; PILÁT, 1958)

Myriostoma Desv. (Lit.: ARNOLDS et al., 1995; DÖRFELT, 1985; GROß et al., 1980; JÜLICH, 1984; KREI- SEL, 2001; PEGLER et al., 1995; PILÁT, 1958; SUNHEDE, 1989)

• Myriostoma coliforme (With. : Pers.) Corda Ref.: COURTECUISSE &DUHEM, 2000; FOCHT, 1974 • var. coliforme Ref.: unp. TORTI] (rev.) Exs.: CNF (Lit.: DÖRFELT, 1985; PILÁT, 1958; SUNHEDE, 1989)

*Nidularia Fr. (Lit.: ARNOLDS et al., 1995; CALONGE, 1998; GROß et al., 1980; JÜLICH, 1984; KREI- SEL, 2001; MORNAND, 1985; MOYERSOEN &DEMOULIN, 1996; PEGLER et al., 1995; PILÁT, 1958)

*Nidularia deformis (Willd. : Pers.) Fr. et Nordholm = Nidularia farcta (Roth : Pers.) Fr. (1) Ref.: unp. TORTI] (1) Exs.: CNF 112 Tkal~ec, Z., Me{i}, A. & Antoni}, O.: Survey of the gasteral Basidiomycota (Fungi) of Croatia

*Phallogaster Morgan (Lit.: CALONGE, 1998; GROß et al., 1980; JÜLICH, 1984; KREISEL, 2001; MONTECCHI &SARASINI, 2000; PILÁT, 1958)

*• Phallogaster saccatus Morgan Ref.: unp. TKAL^EC &ME[I] Exs.: CNF (Lit.: CALONGE, 1998; JÜLICH, 1984; MONTECCHI &SARASINI, 2000)

Phallus Junius ex L. : Pers. (Lit.: ARNOLDS et al., 1995; CALONGE, 1998; JÜLICH, 1984; KREISEL, 2001; MOYER- SOEN &DEMOULIN, 1996; PEGLER et al., 1995; PILÁT, 1958)

• Phallus hadriani Vent. : Pers. Ref.: BO@AC, 1984; KRANJ^EV, 1995; unp. TKAL^EC &ME[I] Exs.: CNF (Lit.: CALONGE, 1998; PEGLER et al., 1995; PILÁT, 1958)

• Phallus impudicus L. : Pers. = Ithyphallus impudicus (L. : Pers.) E. Fisch. (1) Ref.: BAR^I], 1989; BAR^I], 1996; BLAGAI], 1917 (1); BLAGAI], 1921; BLAGAI], 1931; FOCHT, 1974; KRANJ^EV, 1979; KRANJ^EV, 1980; LANGE, 1974; LISIEWSKA & TORTI], 1990; RAGUZIN, 2000; SCHULZER et al., 1866; [KORI], 1928; TORTI], 1966 a; TORTI], 1966 b; TORTI], 1968 a; TORTI], 1968 b; TORTI], 2002; VOUK &PEVALEK, 1916 (1); unp. MATO^EC, unp. TKAL^EC &ME[I]; unp. TORTI];SCHULZER, 1857 HR? Exs.: CNF (Lit.: CALONGE, 1998; PEGLER et al., 1995; PILÁT, 1958)

Pisolithus Alb. et Schwein. (Lit.: ARNOLDS et al., 1995; CALONGE, 1998; GROß et al., 1980; JÜLICH, 1984; KREI- SEL, 2001; MOYERSOEN &DEMOULIN, 1996; PEGLER et al., 1995; PILÁT, 1958)

• Pisolithus arhizus (Scop. : Pers.) Rauschert = Pisolithus tinctorius (P. Micheli ex Pers. : Pers.) Coker & Couch Ref.: RAGUZIN, 2001; unp. TKAL^EC &ME[I] Exs.: CNF (Lit.: JÜLICH, 1984; PEGLER et al., 1995; PILÁT, 1958)

Rhizopogon Fr. emend. Tul. et C. Tul. (Lit.: ARNOLDS et al., 1995; GROß et al., 1980; JÜLICH, 1984; KREISEL, 2001; MARTÍN, 1996; MONTECCHI &SARASINI, 2000; MOYERSOEN &DEMOULIN, 1996; PEGLER et al., 1993; PILÁT, 1958) Nat. Croat. Vol. 14(2), 2005 113

Rhizopogon obtextus (Spreng.) R. Rauschert = Rhizopogon luteolus Fr. emend. Tul. et C. Tul. (1) Ref.: TORTI], 2002; unp. TORTI] (1) Exs.: CNF Rhizopogon roseolus (Corda) Th. Fr. = Rhizopogon rubescens (Tul. et C. Tul.) Tul. et C. Tul. (1) Ref.: FOCHT, 1974 (1); JAAP, 1916 (1); MARTÍN, 1996; MU@IC &BO@AC, 1984; unp. POUZAR (according to unp. TORTI]) (1) Exs.: PRM (1) Rhizopogon roseolus (Corda) Th. Fr. agg. Incl.: Rhizopogon briardii Boud. Rhizopogon hymenogastrosporus Soehner Rhizopogon lapponicus P. Karst. Rhizopogon luteorubescens A.H. Sm. Rhizopogon provincialis Tul. et C. Tul. Rhizopogon rubescens (Tul. et C. Tul.) Tul. et C. Tul. Rhizopogon sardous Pacioni Rhizopogon ventricisporus A.H. Sm. Rhizopogon vulgaris (Vittad.) M. Lange Ref.: KREISEL, 2001

Scleroderma Pers. : Pers. (Lit.: ARNOLDS et al., 1995; CALONGE, 1998; GROß et al., 1980; JÜLICH, 1984; KREI- SEL, 2001; MOYERSOEN &DEMOULIN, 1996; PEGLER et al., 1995; PILÁT, 1958; SIMS et al., 1995) *• Ehrenb. Ref.: unp. KOTLABA (according to unp. TORTI]); unp. TKAL^EC &ME[I]; unp. TORTI] (rev.) Exs.: CNF, PRM (Lit.: JÜLICH, 1984; PEGLER et al., 1995; SIMS et al., 1995) • Scleroderma bovista Fr. = Scleroderma fuscum (Corda) E. Fisch. (1) Ref.: MOESZ, 1938; SCHULZER, 1882 a; SCHULZER et al., 1866; TORTI], 1993; TOR- TI], 2002; VOUK &PEVALEK, 1916 ( )(1); unp. TKAL^EC &ME[I]; unp. TORTI] Exs.: CNF (Lit.: JÜLICH, 1984; PEGLER et al., 1995) • Scleroderma cepa Pers. : Pers. Ref.: TORTI], 1968 b; unp. TKAL^EC &ME[I] Exs.: CNF (Lit.: JÜLICH, 1984; PEGLER et al., 1995) 114 Tkal~ec, Z., Me{i}, A. & Antoni}, O.: Survey of the gasteral Basidiomycota (Fungi) of Croatia

• Scleroderma citrinum Pers. : Pers. = Scleroderma vulgare Hornem. (1) Misappl.: Scleroderma aurantium (L. : Pers.) Pers. sensu auct. non L. (2) Ref.: BAR^I], 1982 (2); BAR^I], 1996 (2); BLAGAI], 1931 (1); GJURA[IN, 1898 (1); KRANJ^EV, 1979 (1); KRANJ^EV, 1985 a (2); SCHULZER, 1882 a (1); SCHULZER et al., 1866 (1); [KORI], 1928 (1); TORTI], 1964 (2); TORTI], 1968 b (2); TORTI], 2002 ( )(1); VOUK &PEVALEK, 1916 (1); unp. MATO^EC (2); unp. TKAL^EC &ME[I]; unp. TORTI] ( )(1)(2); SCHULZER, 1857 HR? (1) Exs.: CNF (Lit.: JÜLICH, 1984; PEGLER et al., 1995) • Scleroderma polyrhizum (J.F. Gmel. : Pers.) Pers. = Scleroderma geaster Fr. (1) Ref.: FOCHT, 1974 (1); unp. TKAL^EC &ME[I];BO@AC, 1984 HR? (1) Exs.: CNF (Lit.: JÜLICH, 1984; CALONGE, 1998) • Scleroderma verrucosum (Bull. : Pers.) Pers. = Scleroderma bresadolae Schulzer (1) Ref.: BAR^I], 1982; BAR^I], 1996; FOCHT, 1974; GJURA[IN, 1898; RAGUZIN, 1998; RAGUZIN, 2001; SACCARDO, 1888 b (1); SCHULZER, 1884 a (1); SCHULZER, 1884 b (1); SCHULZER, 1885 (1); TORTI], 1968 b; TORTI], 2002; TORTI] &LISIEWSKA, 1978; VR[^AJ, 2003; unp. MATO^EC; unp. TKAL^EC &ME[I]; unp. TORTI];FOCHT, 1979 HR? Exs.: CNF (Lit.: JÜLICH, 1984; PEGLER et al., 1995; SIMS et al., 1995)

Sphaerobolus Tode : Pers. (Lit.: ARNOLDS et al., 1995; CALONGE, 1998; GMINDER et al., 2000; GROß et al., 1980; JÜLICH, 1984; KREISEL, 2001; MOYERSOEN &DEMOULIN, 1996; PEGLER et al., 1995; PILÁT, 1958) • Sphaerobolus stellatus Tode : Pers. Ref.: FOCHT, 1972; GJURA[IN, 1898; unp. TKAL^EC &ME[I] Exs.: CNF (Lit.: CALONGE, 1998; JÜLICH, 1984)

Tulostoma Pers. : Pers. (Lit.: ARNOLDS et al., 1995; CALONGE, 1998; GMINDER et al., 2000; GROß et al., 1980; JÜLICH, 1984; KREISEL, 2001; MORNAND, 1989; MOYERSOEN &DEMOULIN, 1996; PEGLER et al., 1995; PILÁT, 1958; WRIGHT, 1987) *Tulostoma armillatum Bres. = Tulostoma fulvellum Bres. Nat. Croat. Vol. 14(2), 2005 115

Ref.: unp. PEINTNER Exs.: IB

• Tulostoma brumale Pers. : Pers. = Tulostoma mammosum (P. Micheli) ex Fr. (1) Ref.: JAAP, 1916 (1); KRANJ^EV, 1985; KRANJ^EV, 1995; RANDI], 1998; unp. TKAL- ^EC &ME[I]; unp. TORTI] Exs.: CNF (Lit.: GROß et al., 1980; JÜLICH, 1984; PILÁT, 1958)

• Tulostoma fimbriatum Fr. Ref.: RANDI], 1998; unp. TKAL^EC &ME[I] Exs.: CNF (Lit.: GROß et al., 1980; JÜLICH, 1984; PILÁT, 1958)

• Tulostoma melanocyclum Bres. Ref.: WRIGHT, 1987 Exs.: BP (Lit.: WRIGHT, 1987) Annotation: Author of the quote specified that locality ([ugarska Duliba, Vele- bit) belongs to Hungary, but the locality belongs to Croatia.

Tulostoma squamosum J.F. Gmel. : Pers. Ref.: FOCHT, 1974; MOESZ, 1938; TORTI], 1993

Vascellum F. [marda (Lit.: ARNOLDS et al., 1995; CALONGE, 1998; DEMOULIN, 1970; GMINER et al., 2000; JÜLICH, 1984; KREISEL, 2001; MOYERSOEN &DEMOULIN, 1996; PEGLER et al., 1995; PILÁT, 1958; SACCARDO, 1888)

• pratense (Pers. : Pers.) Kreisel = Pers. : Pers. (1) = Lycoperdon depressum Bonord. (2) = Vascellum depressum (Bonord.) F. [marda (3) = Lycoperdon hyemale Bull. (4) = Bovista queletii (Schulzer) De Toni (5) = Globaria queletii Schulzer (6) Ref.: FOCHT, 1972 (1); FOCHT, 1974 (1); FOCHT, 1979 (1); KRANJ^EV, 1985 (1); MOESZ, 1938 (4); SCHULZER, 1885 (6); TORTI], 1966 a (3); TORTI], 1966 b (3); TORTI], 1993; VOUK &PEVALEK, 1916 (2); VR[^AJ, 2003; unp. TKAL^EC &ME[I]; unp. TORTI] ( )(3); SACCARDO, 1888 HR? (5) Exs.: CNF (Lit.: JÜLICH, 1984) 116 Tkal~ec, Z., Me{i}, A. & Antoni}, O.: Survey of the gasteral Basidiomycota (Fungi) of Croatia

EXCLUDED RECORDS

Anixia minuta Schulzer = ? Ref.: SCHULZER, 1872 Dictyophora croatica Mu`ic et Bo`ac – nom. nud. Ref.: MU@IC &BO@AC, 1997 Endoptychum agaricoides Czern. = Secotium acuminatum Mont. (1) = Secotium thunii (Schulzer) Schulzer (2) Ref.: SACCARDO, 1888 (2); SCHULZER, 1883 a (1); SCHULZER, 1883 b (1) Annotation: Authors of the quotes specified Slavonia (part of Croatia today) as a locality, but Schulzer found the species in the part of former Slavonia that now belong to Serbia and Montenegro (SCHULZER, 1865). Geastrum striatum DC. =! Geastrum coronatum Pers. : Pers. Ref.: SACCARDO, 1888 HR? Annotation: We exclude this record due to incorrect synonymy. Lycoperdon involvatum Schulzer – nom. nud. Ref.: SCHULZER et al., 1866 Lycoperdon silvaticum Schulzer – nom. nud. Ref.: SCHULZER et al., 1866 Lycoperdon silvaticum Wettst. = ? Ref.: [KORI], 1928; VOUK &PEVALEK, 1916 Scleroderma vulgare Hornem. =!Scleroderma bovista Fr. =!Scleroderma verrucosum (Bull. : Pers.) Pers. Ref.: BLAGAI], 1921 Annotation: We exclude this record due to incorrect synonymy.

ACKNOWLEDGEMENTS

We are grateful to Dr. Milica Torti}, Mr. Neven Mato~ec, Prof. Dr. Maria Lisi- ewska and Dr. Ursula Peintner for generously allowing us to use the unpublished data from their records; to Dr. Milica Torti}, Mr. Andrej Piltaver, and Prof. Dr. Bogdan Cvjetkovi} for their assistance with literature and to Mrs. Maja Prezel for helping with the English version of the text. We are grateful to Ministry of Science, Education and Sports and Croatian Academy of Sciences and Arts for financial as- sistance. We also thank all collectors of material and all those who have in any way contributed to the writing of this paper. Received December 14, 2004 Nat. Croat. Vol. 14(2), 2005 117

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Unpublished:

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