The Palaeontological Association 45th Annual Meeting 15th-19th December, 2001 Geological Museum University of Copenhagen ABSTRACTS Abstracts 2001 2 TALKS Abstracts of oral presentations Compiled and edited by David A.T. Harper The tristichopterid fishes: Almost, but not quite, tetrapods Per Erik Ahlberg Department of Palaeontology, The Natural History Museum, Cromwell Road, London SW7 5BD, UK <
[email protected]> The clade Tristichopteridae occupies a relatively high position within the tetrapod stem group; it is the most advanced “osteolepiform” group, and forms the sister taxon to Panderichthys + tetrapods. The best known tristichopterid is the primitive early Frasnian genus Eusthenopteron, but the clade persisted to the end of the Devonian. The phase of rapid morphological change traditionally identified as the “fish-tetrapod transition” begins with the late Givetian Panderichthys and continues through to Famennian stem-group tetrapods such as Acanthostega and Ichthyostega. By contrast, the less crownward “osteolepiform fish” part of the stem group shows limited morphological change and no obvious overall trend towards terrestriality. However, within the Tristichopteridae there is independent acquisition of a Panderichthys-like body morphology. Recently, the advanced late Frasnian tristichopterid Mandageria has been found to possess a true neck joint, previously thought to be a unique tetrapod character. The extent, nature, TALKS and likely ecological significance of the convergence between tristichopterids and tetrapods will be discussed in the light of these findings. Phylogenetic analysis of the tetrapod stem group has shown that the lower, “fish” part consists of (in ascending sequence) the clade Rhizodontida, a paraphyletic array of osteolepiforms, the clade Tristichopteridae, and Panderichthys. In morphological terms, the “fish-tetrapod transition” begins at the level of Panderichthys.