DOCSLIB.ORG

Zoologische Mededelingen

Total Page：16

File Type：pdf, Size：1020Kb

Download full-text PDF Read full-text
Zoologische Mededelingen MINISTERIE VAN ONDERWIJS, KUNSTEN EN WETENSCHAPPEN ZOOLOGISCHE MEDEDELINGEN UITGEGEVEN DOOR HET RIJKSMUSEUM VAN NATUURLIJKE HISTORIE TE LEIDEN DEEL XXXVIII, No. 15 18 sept. 1963 CLAUSILIIDAE (GASTROPODA, PULMONATA) COLLECTED BY THE NETHERLANDS BIOLOGICAL EXPEDITION TO TURKEY IN 1959 by F. E. LOOSJES The route of the Netherlands Biological Expedition to Turkey in 1959 is published elsewhere (Hennipman a.o., 1961; Anonymus, 1963). As is mentioned there also, the zoological material collected is preserved in the Rijksmuseum van Natuurlijke Historie at Leiden. I am greatly indebted to Dr. C. O. van Regteren Altena, curator of that Museum, for placing the Clausiliidae collected by the expedition at my disposal. Moreover my thanks are due to Mrs. W. S. S. van der Feen-van Benthem Jutting, Amsterdam, and Dr. A. Zilch, Frankfurt am Main, for specimens lent for the purpose of comparison. The Clausiliidae were all collected by Mr. W. J. M. Vader, one of the members of the expedition. Only one sample is from Greece (Porto Lagos), the others are from Istanbul, near the Bosporus, and from Asiatic Turkey. All the samples are preserved dry, the soft parts having hardened in the top whorls of the shells by loss of moisture. In the systematic account the arrangement given in the „Handbuch der Palaozoologie" vol. 6 by Wenz-Zilch (1959/60) is followed. Some of the specimens are in the author's collection and in the collection of Mr. W. J. M. Vader. PHAEDUSINAE Tribus Serrulineae Serrulina Mousson, 1873 Serrulina (Serrulina) serrulata (L. Pfeiffer, 1847) Material examined. — Turkey, Vilayet of Trabzon, 3 km S. of Hamsikoy (pi. XX), about 45 km SSW. of the city of Trabson, altitude ca. 1900 m, 244 F. E. LOOSJES in a wet forest with firs and beech trees; a small number of specimens was found on the trunks of living beech trees, the majority, however, were living in a decaying beech trunk, lying on a small mountain pasture, 5-VI-1959 (29 specimens). Length of shells 12.2-15.5 mm, diameter 2.8-3.4 mm; whorls: 10-12. The columellar side of the peristome is folded as usual in the species, but in most specimens the outer peristome is not or only faintly folded. The set thus shows features tending towards the var. semiserrata Lindholm (1913) that was described from Wasrya (Gouvernement of Batum). Distribution. — Type locality "Tauria". The species is known from the coastal areas of the Black Sea, on the east side (O. Boettger, 1879, 1880, 1883, 1889; Von Martens, 1874; Mousson, 1863; Retowski, 1889; Rosen, 1907), on the south side (Gotting, 1961; Retowski, 1889) and on the west side (Hesse, 1913; Jaeckel, 1957; Urbanski, 1960a). Washed ashore on the Crimea peninsula (Retowski, 1883, 1887). The locality given is thus within the known area of distribution. Biotope. — Shady forests. O. Boettger (1880) recorded (according to Leder) that the species lives: "in alten, faulenden, durch und durch nassen und audi bei der grossten Trockenheit doch stets reichlich Wasser haltenden Stammen einer grossblatterigen Acer Art, deren Holz man Stuck fiir Stuck zerfasern muss". Retowski (1889): "in einem faulenden Baumstamme". Urbanski (1960a) : "unter der Rinde alter, faulender Stubben und Baum• stamme, falls diese nicht ganz trocken sind". Gotting (1961) mentioned: "an und unter Baumen". These data are in accordance with the habitat mentioned by Vader. Serrulina (Pravispira) semilamellata (Mousson, 1863) Material examined. — Turkey, Vilayet of Trabzon, 3 km S. of Hamsikoy (pi. XX), about 45 km SSW. of the city of Trabzon, alt. ca. 1900 m, in a wet forest with firs and beech trees; collected together with S. serrulrta Pfeiffer, thus on trunks of living beech trees or in a decaying beech trunk, lying on a small mountain pasture. It is likely that the present two specimens lived in the decaying trunk and not on the living trees as most of the Serrulina material came from the trunk and as the species is especially known from such a biotope. 5-VI-1959 (2 specimens). Length of shells 12.4 and 13.4 mm, diameter 2.9 and 3.1 mm; whorls: 9 and 9. The specimens have no third (middle) plica palatalis and the upper (principalis) and lower plicae are less strong than in typical specimens. Retowski (1889) described this condition for his variety serrulosa from Sephanos near Trebisond (= Trabzon). The small folds along the right CLAUSILIIDAE OF THE TURKEY EXPEDITION 245 side of the peristome, however, are as in typical S. semilamellata and not as numerous as was described for var. serrulosa Retowski. The specimens are large as compared with the original set from Reduktaleh (length 10 mm, diam. 2.4 mm, Mousson, 1863) and with var. serrulosa Retowski (length 9 mm, diam. 2 mm). Specimens from other localities, however, may have larger dimensions (among others Swanetien: length 11J4-14 mm> diam. 2J4-3 mm; O. Boettger, 1883). According to O. Boettger (1880) shape, dimensions and sculpture of the species are very variable. Distribution. — S. semilamellata is only known from the coast of the Black Sea (O. Boettger, 1879, 1880, 1883, 1889; Clessin, 1881; Von Mar• tens, 1876; Mousson, 1863; Retowski, 1883, 1889), from the SE coast (Retowski, 1889) and washed ashore on the Crimea peninsula (Retowski, 1887). The locality Hamsikoy is within the known area of distribution. Biotope. — The biotope is the same as that of S. serrulata. According to O. Boettger (1880) both species were found together in old, wet, decaying tree trunks. Retowski (1889) collected the species in completely decayed wood. Rosen (1907) recorded: "sie lebt sehr vereinzelt und nie in Colonien wie andere Arten, an Bergabhangen im Walde unter faulenden Baum- stammen. Unter einen dicken Baumstumpf fand ich eine Colonie von Clausilia serrulata Midd. in 80 Exemplaren und darunter bloss zwei Exemplare semilamellata. Sie fiihrt wahrscheinlich ein Einsiedlerleben, denn ich konnte nie mehr wie ein oder hochstens zwei Exemplare an einem Ort finden". This is in accordance with the situation at Hamsikoy where 2 specimens of 5*. semilamellata were collected among more than 20 S. serrulata. CLAUSILIINAE Laciniaria Hartmann, 1844 Laciniaria (Denticularia) thessalonica thasia (O. Boettger, 1907 ex recte Urbanski, i960) Material examined. — Greece, Macedonia, road from Komotini to Porto Lagos, under stones in poor grassland, 28-VI-1959 (10 specimens). Length of shells 11.2-12.6 mm, diameter 2.9-3.1 mm; whorls: 11-12. The specimens are short and rather ventricose with somewhat concave lateral outlines, the striae are occasionally whitish only on the upper parts of a whorl. Most of the specimens (7) have only small folds on the peristome, one is also provided with some interlamellar plicae. Plica principalis and lower palatal plica long, visible in the aperture; the middle palatal plica is short in all except two specimens, where it is clearly visible in the aperture. 246 F. E. LOOSJES Distribution. — The polytypic species is known from Central Greece (Euboea), Thessalia, Macedonia and SE. Bulgaria, from the coasts of the Sea of Marmara and from W. Turkey (O. Boettger, 1885; Fuchs & Kaufel, 1936; Urbanski, 1960b). L. thessalonica thasia is known from the island of Thasos (O. Boettger, 1907; Urbanski, 1960b), from the island of Thasopoula (Urbanski, 1960b) and according to Urbanski (1960b) presumably also from eastern Macedonia. Our locality is in accordance with Urbanski's expectations. Biotope. — Under decaying wood (O. Boettger, 1885; Gotting, 1961) under shrubs (Hesse, 1914; Urbanski, 1960b). Vader found the specimens under stones. Euxina O. Boettger, 1877 Euxina (Euxina) circumdata (L. Pfeiffer, 1848) (pi. XVIII figs. 16, 17) Material examined. — Turkey, Istanbul, in a small park near the Aya Sophia, clinging to the bark of the only tree present (it was a very small park indeed!), 21-III-1959 (3 specimens). Length of shells 12.2, 12.7 and 13.4 mm, diameter 3.1, 3.1 and 3.2 mm; whorls: 11. Turkey, Rumeli Hisar, along the Bosporus, along the road under stones, 1-IV-1959 (4 specimens and 3 juv.). Length of shells 12.2-13.2 mm, diameter 2.9-3.1 mm; whorls: Turkey, Rumeli Hisar, along the Bosporus, on a stony grass-slope, under a stone, 25-III-1959 (1 specimen). Length of shell 11.7 mm, diameter 2.9 mm; whorls: 11. The specimens are typical. The var. byzantina Nagele (1910) from the same district is insufficiently characterized. Distribution. — The typical set was collected near Brussa in Natolia; the species occurs in NW. Asia Minor, E. Thracia and Bulgaria (A. J. Wagner, 1920; Urbanski, 1960a; Likharev, 1962), and was found washed ashore on the coast of the Crimea peninsula (Retowski, 1887). The three localities mentioned above are all west of the Bosporus. The species is already known from this area: Constantinople, on walls of the Serail (Mousson, 1863), Bujukdere (Retowski, 1889), Constantinople, garden of the Jezuits (Nagele, 1910). Biotope. — On walls (Mousson, 1863), common in shady woods, under fallen leaves, dead wood, more rarely under stones; sometimes in light oak woods and even in Paliurus shrubs (Urbanski, 1960a). The habitats where Vader collected his specimens were indeed hardly wooded at all. CLAUSILIIDAE OF THE TURKEY EXPEDITION 247 Euxina (Multiplicaria) duboisi (Charpentier, 1852) Material examined. — Turkey, Trabzon, on old city walls in dry places, under ferns, etc., 2-VI-1959 (39 specimens). Length of shells 10.9-13.7 mm, diameter 2.6-3.0 mm; whorls: ioj£-i2j4. The specimens are typical, among 39 shells most have smooth peristomes, only a few have the peristome faintly folded; both forms occur together in the same locality as was already recorded by Retowski (1889). Distribution. — The species was described from "Tauria" (Charpentier, 1852). According to O. Boettger (1879) the species is: "im ganzen Gebiet des Kaukasus verbreitet und entschieden eine der hauf igeren dortigen Clausi- lienformen".
Load more

Recommended publications
  • Guidelines for the Capture and Management of Digital Zoological Names Information Francisco W
    Guidelines for the Capture and Management of Digital Zoological Names Information Francisco W. Welter-Schultes Version 1.1 March 2013 Suggested citation: Welter-Schultes, F.W. (2012). Guidelines for the capture and management of digital zoological names information. Version 1.1 released on March 2013. Copenhagen: Global Biodiversity Information Facility, 126 pp, ISBN: 87-92020-44-5, accessible online at http://www.gbif.org/orc/?doc_id=2784. ISBN: 87-92020-44-5 (10 digits), 978-87-92020-44-4 (13 digits). Persistent URI: http://www.gbif.org/orc/?doc_id=2784. Language: English. Copyright © F. W. Welter-Schultes & Global Biodiversity Information Facility, 2012. Disclaimer: The information, ideas, and opinions presented in this publication are those of the author and do not represent those of GBIF. License: This document is licensed under Creative Commons Attribution 3.0. Document Control: Version Description Date of release Author(s) 0.1 First complete draft. January 2012 F. W. Welter- Schultes 0.2 Document re-structured to improve February 2012 F. W. Welter- usability. Available for public Schultes & A. review. González-Talaván 1.0 First public version of the June 2012 F. W. Welter- document. Schultes 1.1 Minor editions March 2013 F. W. Welter- Schultes Cover Credit: GBIF Secretariat, 2012. Image by Levi Szekeres (Romania), obtained by stock.xchng (http://www.sxc.hu/photo/1389360). March 2013 ii Guidelines for the management of digital zoological names information Version 1.1 Table of Contents How to use this book ......................................................................... 1 SECTION I 1. Introduction ................................................................................ 2 1.1. Identifiers and the role of Linnean names ......................................... 2 1.1.1 Identifiers ..................................................................................
    [Show full text]
  • Magyar Malakológiai Bibliográfia (1727–2004)
    01_cikkek.qxd 2005.11.21. 10:00 Page 5 MALAKOLÓGIAI TÁJÉKOZTATÓ MALACOLOGICAL NEWSLETTER 2005 23: 5–129 Magyar Malakológiai Bibliográfia (1727–2004) Varga András, Fűköh Levente & Krolopp Endre Az első Magyar Malakológiai Bibliográfia 1983-ban a Budapesten megrendezett Unitas Malacologica Kongresszus alkalmával jelent meg, Merényi László szerkesztésében. A ki- advány – Magyarországi pleisztocén és recens malakológiai bibliográfia címen – igyekezett áttekintést adni mindazokról a munkákról, melyek Magyarország területén végzett kutatá- sok eredményeit tartalmazták. Az összeállítás Magyarország alatt az adott munka megjele- nési idejének megfelelő ország területét vette alapul. Jelen összeállítás is követi ezt az alap- elvet, de a fentieken túl tartalmaz minden olyan munkát is, mely magyar szerzők nevéhez fűződően hazánkon kívüli területek malakológiai anyagának feldolgozását tartalmazza, vagy valamilyen formában kapcsolatba került a magyarországi malakológiai kutatásokkal (pl. VERDCOURT 2004). Az 1983-ban megjelent kiadványt követően egyre gyakrabban jelentkezett az igény, hogy az időközben fellelt, s a kötetből kimaradt régebbi irodalmi adatok, illetve az eltelt időszak- ban elvégzett munkák összegzése is jelenjen meg. Ennek az igénynek igyekezett eleget tenni az 1991-ben PERJÉSI György és HORVÁTH Csaba által összeállított munka, „Magyarországi pleisztocén és recens malakológiai bibliográfia (Pótlás)” a Malakológiai Tájékoztató 10. (1991) kötetében, majd ugyancsak ez a szerzőpáros készítette el az időközben megjelent újabb dolgozatok bibliográfiáját: „Magyarországi pleisztocén, quarter és recens malakológiai bibliográfia 1980-1990.” (sic!) címmel a Malakológiai Tájékoztató 11. (1992) kötetében. A fentiekben megjelent munkák óta eltelt közel negyed század ismét felvetette az igényt, hogy készüljön egy új, részben eltérő szerkesztési elveket követő munka, melynek céljai közt szerepel, hogy egyrészt összevonja a MERÉNYI, PERJÉSI & HORVÁTH féle össze- állításokat, másrészt az anyagot 2004-ig bezáróan kiegészítse az időközben megjelent dol- gozatokkal.
    [Show full text]
  • Of Strandzha Mountains and the Black Sea Coast
    Historia naturalis bulgarica, 21: 13-48, 2015 Terrestrial gastropods (Mollusca, Gastropoda) of Strandzha Mountain and the Black Sea coast (Bulgaria and Turkey) Atanas Irikov, Ivelin Mollov Abstract: The current synopsis presents an overview of the terrestrial malacofauna of Strandzha Mountain in Bulgaria and Turkey, based on previously published and new data. As a result of the research we recorded 101 species and subspecies of ter- restrial molluscs belonging to 27 families. The data on the terrestrial malacofauna from the Turkish part of Strandzha is entirely new and presented here for the first time. The synopsis includes a list of synonymous species and subspecies concerning the area of research, all known localities, new localities reported for 50 taxa, systematic and environmental data. For the first time a zoogeographical and conservation analysis of the terrestrial snails is made. Key words: terrestrial gastropods, snails, slugs, Strandzha Mountain, Bulgaria, Turkey. Introduction 50 publications contain data on the terrestrial Original data on species distribution, as well as malacofauna of Strandzha Mountain in Bulgaria. For taxonomic, systematic, zoogeographical and ecologi- Strandzha Mnt. in Turkey there are limited data only cal data resulting from studies in the years 2002-2010 on the presence of a few species of the genus Monacha are included. (Hausdorf, 2000). Most publications report spe- cies composition and localities or describe new taxa. Material and methods Information on the malacofauna is fragmentary and Names of species and subspecies, with a few ex- mostly referring to the Bulgarian part of Strandzha. ceptions, follow Irikov & Mollov (2006). The list of With almost no data on the Turkish part of Strandzha, synonyms applies only to Strandzha Mnt.
    [Show full text]
  • Strasbourg, 19 April 2013
    Strasbourg, 25 October 2013 T-PVS (2013) 17 [tpvs17e_2013.doc] CONVENTION ON THE CONSERVATION OF EUROPEAN WILDLIFE AND NATURAL HABITATS Group of Experts on the Conservation of Invertebrates Tirana, Albania 23-24 September 2013 ---ooOoo--- REPORT Document prepared by the Directorate of Democratic Governance This document will not be distributed at the meeting. Please bring this copy. Ce document ne sera plus distribué en réunion. Prière de vous munir de cet exemplaire. T-PVS (2013) 17 - 2 - CONTENTS 1. Meeting report ................................................................................................................................... 3 2. Appendix 1: Agenda .......................................................................................................................... 6 3. Appendix 2: List of participants ........................................................................................................ 9 4. Appendix 3: Compilation of National Reports .................................................................................. 10 5. Appendix 4: Draft Recommendation on threats by neurotoxic insecticides to pollinators ................ 75 * * * The Standing Committee is invited to: 1. Take note of the report of the meeting; 2. Thank the Albanian government for the efficient preparation of the meeting and the excellent hospitality; 3. Continue with Bern Convention engagement with invertebrate conservation issues by further encouraging and monitoring national implementation of European Strategy for the Conservation
    [Show full text]
  • Abbreviation Kiel S. 2005, New and Little Known Gastropods from the Albian of the Mahajanga Basin, Northwestern Madagaskar
    1 Reference (Explanations see mollusca-database.eu) Abbreviation Kiel S. 2005, New and little known gastropods from the Albian of the Mahajanga Basin, Northwestern Madagaskar. AF01 http://www.geowiss.uni-hamburg.de/i-geolo/Palaeontologie/ForschungImadagaskar.htm (11.03.2007, abstract) Bandel K. 2003, Cretaceous volutid Neogastropoda from the Western Desert of Egypt and their place within the noegastropoda AF02 (Mollusca). Mitt. Geol.-Paläont. Inst. Univ. Hamburg, Heft 87, p 73-98, 49 figs., Hamburg (abstract). www.geowiss.uni-hamburg.de/i-geolo/Palaeontologie/Forschung/publications.htm (29.10.2007) Kiel S. & Bandel K. 2003, New taxonomic data for the gastropod fauna of the Uzamba Formation (Santonian-Campanian, South AF03 Africa) based on newly collected material. Cretaceous research 24, p. 449-475, 10 figs., Elsevier (abstract). www.geowiss.uni-hamburg.de/i-geolo/Palaeontologie/Forschung/publications.htm (29.10.2007) Emberton K.C. 2002, Owengriffithsius , a new genus of cyclophorid land snails endemic to northern Madagascar. The Veliger 45 (3) : AF04 203-217. http://www.theveliger.org/index.html Emberton K.C. 2002, Ankoravaratra , a new genus of landsnails endemic to northern Madagascar (Cyclophoroidea: Maizaniidae?). AF05 The Veliger 45 (4) : 278-289. http://www.theveliger.org/volume45(4).html Blaison & Bourquin 1966, Révision des "Collotia sensu lato": un nouveau sous-genre "Tintanticeras". Ann. sci. univ. Besancon, 3ème AF06 série, geologie. fasc.2 :69-77 (Abstract). www.fossile.org/pages-web/bibliographie_consacree_au_ammon.htp (20.7.2005) Bensalah M., Adaci M., Mahboubi M. & Kazi-Tani O., 2005, Les sediments continentaux d'age tertiaire dans les Hautes Plaines AF07 Oranaises et le Tell Tlemcenien (Algerie occidentale).
    [Show full text]
  • Die Arten in Gilan Und Mazandaran (Mit Beschreibung Neuer Taxa) (Gastropoda: Stylommatophora)*)
    download Biodiversity Heritage Library, http://www.biodiversitylibrary.org/ Stuttgarter Beiträge zur Naturkunde Serie A (Biologie) Herausgeber: Staatliches Museum für Naturkunde, Rosenstein 1, D Stuttgarter Beitr. Naturk. Ser. A Nr. 527 27 S. Stuttgart, 31. 7. 1995 Iranische Clausiliidae: Die Arten in Gilan und Mazandaran (mit Beschreibung neuer Taxa) (Gastropoda: Stylommatophora)*) Iranian Clausiliidae: The Species in Gilan and Mazandaran (with the Description of new Taxa) (Gastropoda: Stylommatophora) Von Hartmut Nordsieck, Villingen-Schwenningen Mit 3 Tafeln und 5 Abbildungen Summary Faumstic and systematic data on Iranian clausiliid-species from Gilän and Mazandaran are presented. The available informations on the localities in which most of the material was col- lected are reported (chapters 4. and 5.). Species not yet revised are diagnosed, and their ranges, as far as known, are specified (6.). The following taxa are described as new: Euxina acbrafensis n. sp., E. forcarti n. sp., E. gastron n. sp., Likharevia gilanensis n. sp., Euxina talyschana asta- rana n. subsp., and Likharevia gilanensis costulata n. subsp. The zoogeographical position of Hyrcania, the distribution and the ecology of the species, and the phylogeny of the Hyrcanian Euxina species are discussed (7.). Zusammenfassung Es werden faunistische und systematische Angaben über iranische Clausihen-Arten von Gilän und Mazandaran vorgelegt und die verfügbaren Informationen zu den Fundorten, an denen das meiste Material gesammelt wurde, gegeben (Kapitel 4. und 5.). Die bislang noch nicht revidierten Arten werden diagnostiziert und ihre bisher festgestellten Verbreitungsge- biete angegeben (6.). Die folgenden neuen Taxa werden beschrieben: Euxina acbrafensis n. sp., E. forcarti n. sp., E. gastron n.sp., Likharevia gilanensis n.
    [Show full text]
  • Identification Keys to the Clausiliidae J. E. Gray, 1855 (Mollusca: Stylommatophora) from Bulgaria
    Systematics and Phylogenetics ACTA ZOOLOGICA BULGARICA Research Article Acta zool. bulg., 67 (2), 2015: 167-177 Identification Keys to the Clausiliidae J. E. Gray, 1855 (Mollusca: Stylommatophora) from Bulgaria Atanas Irikov*, Dilian Georgiev Department of Ecology and Nature Conservation, Faculty of Biology, University of Plovdiv “Paisii Hilendarski”, 24 Tsar Assen Street, 4000 Plovdiv, Bulgaria, E-mails: [email protected]; [email protected] Abstract: The paper presents the currently known diversity of the family Clausiliidae J. E. Gray, 1855 on the territory of Bulgaria. All new systematic changes are noted and all new taxa described during the last years are listed. The main key characters for species identification are presented, with additional descriptions of some characters. Keywords: identification, genera, species, subspecies, Clausiliidae, Bulgaria Introduction The terrestrial malacofauna of Bulgaria is char- the different groups of Clausiliidae require expertise. acterised by high taxonomic diversity of the family According to the same authors, the genital system Clausiliidae Gray, 1855. The Europaen representa- structure also has an important taxonomical value in tives of this family are distributed mostly on the identifying some of the species. Balkan Peninsula. On the territory of Bulgaria, there Specific for the species and subspecies of is an extremely high diversity of species and sub- this family is that most of them have local distri- species of clausiliids. Most of these taxa are sum- butions on isolated rock complexes in Bulgaria of marised in the monograph of DAMJANOV , LIKHAREV the so-called ‘island-type’. Some of them occur (1975). Following the latter publication, many no- on very small territories as Macedonica pirinensis menclatural changes were made, new distributional Jaeckel, 1954, Macedonica zilchi Urbański, 1972, data were published and some additional species and Macedonica hartmuti Irikov, 2003, Macedonica subspecies were described.
    [Show full text]
  • Annotated List of Invertebrates of the Georgian Karst Caves
    ANNOTATED LIST OF INVERTEBRATES OF THE GEORGIAN KARST CAVES Shalva Barjadze, Maka Murvanidze, Tea Arabuli, Le- van Mumladze, Vera Pkhakadze, Revaz Djanashvili, Meri Salakaia Georgian Academic Book Tbilisi, 2015 Project: “Biodiversity of the Invertebrate Animals in Georgian Karst Caves” This monograph is published by financial support of Shota Rustaveli National Science Foundation (project №11/27) Non-comercial publishing Shalva Barjadze, Maka Murvanidze, Tea Arabuli, Levan Mumladze, Vera Pkhakadze, Revaz Djanashvili, Meri Salakaia Annotated List of Invertebrates of the Georgian Karst Caves Technical editor - Maia Gambashidze Cover design - Vakhtang Inanishvili Design - Giorgi Kakabadze © Shalva Barjadze, 2015 ISBN 978- 9941-9371-2-5 Georgian Academic Book Ir. Abashidze Str. 16, 0179 Tbilisi, Georgia Tel.: 222 7137, 551 227137, 551 960904 www.acad.ge; [email protected] Contents Introduction 5 List of caves with more than one name, mentioned in biospeleological and speleological literature 10 Part I. Annotated List of Invertebrates 12 Part II. Caves and their invertebrates’ communities 65 Acknowledgements 103 Literature 105 Alphabetical index of caves 118 DEDICATION Dedicated to the memory of Dr. Eristo Kvavadze, eminent Georgian zoologist INTRODUCTION According to the great species and ecosystem diversity, high endemism and high risks of destroying of biodiversity, 34 hot spots of wild life are re- vealed in the world, among which the Caucasus ecoregion has appropriated place (Mayrs et al., 2000; Zazanashvili et al., 2004; Zazanashvili, 2009). Western Georgia’s karstic outcrops lie along the southern slope of the Great- er Caucasus mountain range, extending some 325km from the Psou River to the Ertso Lake area. The total area of karstic rock outcrop amounts to about 4,475km2, or 6.4% of the total area of Georgia (Tsikarishvili et al., 2010).
    [Show full text]
  • Gastropoda Pulmonata)
    BASTERIA, 62: 169-173, 1998 New and little-known Clausiliidae (Gastropoda Pulmonata) from eastern Turkey M. Szekeres Alfoldi u. 16, H-6725 Szeged, Hungary Cristataria colbeauiana inversa and Roseniella nemethi from n. ssp. n. sp. are introduced as new taxa eastern Turkey and their systematic position is discussed. The genital morphology of Pontophaedusella ofensis Nordsieck is presented, with comments on itsrelevance to the phylogenetic this clausiliid of the Serrulina status of group. words: Key Gastropoda, Pulmonata, Clausiliidae, Cristataria, Roseniella, taxonomy, Turkey. In the of about the Clausiliidae fauna of has past couple years our knowledge Turkey increased significantly. Since the publication of a comprehensive overview by Bank & Menkhorst (1994), more than 45 new species taxa have been described and considerable to efforts have been made improve our understanding of the phylogenetic relationships and biogeographical significance ofTurkish Clausiliidae. This paper is a contribution to this ongoing work by providing the description ofnew taxa which were collected during in a field trip made by Mr. L. Nemeth and myself 1996. Alopiinae Cristataria colbeauiana inversa n. ssp. (fig. 1) — 1 Material. Limestone cliffs at 370 m alt. in a gorge km S. of Harbiye (S. of Antakya), BA41. The holotype is in the collection of the Hungarian Natural History Museum (Budapest); paratypes (471) are in the collections of the Hungarian Natural History Museum (2), the Nationaal Natuurhistorisch Museum, Leiden (NNM 57265/2), the Forschungsinstitut und Naturmuseum Senckenberg, Frankfurt am Main(SMF 311774/ 2), and those of L Nemeth and the author. Diagnosis. — A sinistral subspecies of Cristataria colbeauiana (L. Pfeiffer, 1861) with a and the lunellar close the more prominent sculpture region positioned relatively to ap- erture.
    [Show full text]
  • Gastropoda: Pulmonata: Clausiliidae) from Northern Turkey
    Ecologica Montenegrina 10: 26-30 (2017) This journal is available online at: www.biotaxa.org/em https://zoobank.org/urn:lsid:zoobank.org:pub:476BC71C-E055-47A4-8280-205F37A6A981 Cotyorica nemethi n. gen. n. sp., a remarkable Tertiary relict of the subfamily Phaedusinae (Gastropoda: Pulmonata: Clausiliidae) from northern Turkey JOZEF GREGO1 & MIKLÓS SZEKERES1 1 Horná Mičiná 219, SK-97401 Banská Bystrica, Slovakia, e-mail: [email protected] 2 Institute of Plant Biology, Biological Research Centre of the Hungarian Academy of Sciences, Temesvári krt. 62, H- 6726 Szeged, Hungary, e-mail: [email protected] Received: 10 February 2017 │ Accepted by V. Pešić: 15 March 2017 │ Published online: 17 March 2017. Abstract Cotyorica nemethi n. sp., one of the Tertiary relict western members of the geographically disjunct subfamily Phaedusinae, is described from Ordu Province of northern Turkey. It shows similarity to the species of the subterranean genera Graecophaedusa Rähle, 1982 and Nothoserrulina Németh & Szekeres, 1995 having high positioned lamellae inferior and subcolumellaris, but differs from these to an extent that justifies distinct generic status. Key words: Phaedusinae, endemism, Tertiary relict, subterranean habitat. Introduction Climatically well balanced coastal regions of the Mediterranean, Black and Caspian Seas with mild winters harbour a remarkably rich and diverse fauna of the land snail family Clausiliidae. Among its unique endemic elements are a few species that represent Tertiary relicts of the mainly Asiatic Phaedusinae subfamily (Nordsieck 1978). In addition to the favourable climate, the survival of most of these clausiliids was ensured by their subterranean habitats capable of providing frost-free and permanent high humidity environment.
    [Show full text]
  • Annotated Check-List of the Bulgarian Terrestrial Snails (Mollusca, Gastropoda)
    © Biologiezentrum Linz/Austria; download unter www.biologiezentrum.at Linzer biol. Beitr. 30/2 745-765 31.12.1998 Annotated check-list of the Bulgarian terrestrial snails (Mollusca, Gastropoda) I.K. DEDOV Abstract: In the present work 236 species and 42 subspecies (excluding the nominal ones) of terrestrial snails are listed. 13 of the species and 22 of the subspecies are considered as being with vague systematic status. The occurence of 7 species [Pagodulina pagodula (DESMOULINS), Chondrina spelta (BECK), Bulgaria* rugicollis (ROSSMASSLER), Oxyloma sarsii (ESMARK & HOYER), Tandonia rustica (MILLET), Umax subalpinus LESSONA, Deroceras pageti GROSSU] in Bulgaria is doubtfull. The Bulgarian and Balkan peninsula endemics as well as the rare species/subspecies are indicated. The taxa so far known as being tertiary relicts are marked. Key words: terrestrial snails, Bulgaria, checklist. Introduction The investigations on Bulgarian terrestrial snails were generally carried out by foreign authors. The works until 1974 were summarized by DAMAJANOV & LlKHAREV (1975). Subsequently, many authors have published on Bulgarian land snails. Numerous new species were described, other were firstly recorded for the Bulgarian fauna or synony- mised. This work sets the task to summarize the names of all Bulgarian terrestrial snails, to present the new tendencies of the systematics and to provide some critical remarks on some species. According to DELTCHEV et al. (1993) about 80 percent of the Bulgarian mollusc fauna is known. Despite of presence of a monograph on terrestrial snails several systematical, zoogeographical and ecological problems of this group are not solved. There are many described subspecies, forms and varieties of a vague systematic status.
    [Show full text]
  • Systematic and Geographical Notes on Clausiliidae.*) by Caesar R
    download www.zobodat.at Systematic and Geographical Notes on Clausiliidae.*) By Caesar R. Boettger. In a presidential address the Rev. A. H. Cooke has presented to the Malacological Society of London an admirable and clear study of this widely distributed family of land shells1), including a review of the geographical distribution of species and a most valuable comprehensive account showing how far classification based on the structure of the shell alone was able to increase our knowledge of this group. The landshells composing it have always been a most attractive object of study to systematists. Especially the peculiarly adapted clausal apparatus of the shell has been a promising field for research. Even more than in most of the other families of landshells the clausal apparatus of the shell of the Clausiliidae had been used as a leading feature, apparently well adapted to serve as a definite characteristic for classification. Not only the different species could thus be distinguished, but the characteristics made out were also used for a subdivision into well defined sections of the genus Clausilia itself. In this way the system of Clausiliidae became a well founded building. K iister, A. Schm idt, v. V est and O. B o ettg er were the most experienced workers on this subject. *) This paper was completed early in 1923, but, on the advice of the editor of the Proceedings of the Malacological Society of London was kept back to avoid repetition, as two other papers on a somewhat similar subject were about to appear (A. S. K ennard and B.
    [Show full text]