sign in sign up
<<
Home , Aciculidae, Aegopinella minor, Borysthenia naticina, Brephulopsis cylindrica, Bulgarica cana, Carychium tridentatum

Folia Malacol. 21(3): 175–181

http://dx.doi.org/10.12657/folmal.021.021

AN UPDATED ANNOTATED CHECKLIST OF THE MOLLUSCS OF THE REPUBLIC OF

1 2 3 4 IGOR A. BALASHOV , MIKHAIL O. SON , VIORICA COADÃ , FRANCISCO WELTER-SCHULTES

1Schmalhausen Institute of Zoology, National Academy of Sciences of , B. Khmelnytsky 15, Kiev, 01601, Ukraine (e-mail: [email protected]) 2Odessa Branch Institute of Biology of the Southern Seas, National Academy of Sciences of Ukraine, Pushkinskaya 37, Odessa, 65011, Ukraine (e-mail: [email protected]) 3Tiraspol State University, Gh. Iablocikin 5, MD 2069, Chiºinãu, Republic of Moldova (e-mail: [email protected]) 4Zoological Institute of Göttingen University, Berliner Strasse 28, 37073, Göttingen, (e-mail: [email protected]) ABSTRACT: An updated annotated checklist of the molluscs of the Republic of Moldova is provided. It includes 155 : 26 bivalves, 45 freshwater gastropods and 84 terrestrial gastropods. Some species such as claustralis, frumentum, petronella and Tandonia kusceri are reported for the first time from Moldova by original data. A number of unclear species reports are discussed. KEY WORDS: molluscs, , , , checklist, Moldova

INTRODUCTION

The first annotated checklist of the molluscs of the original authors (for most complete list see: STARO- Republic of Moldova, which included 112 species, was BOGATOV et al. 2004), we could not just list these addi- published recently (COADÃ &WELTER-SCHULTES tional names as synonyms of recognised species. Many 2011), but unfortunately some literature data were of these names are not synonyms in the taxonomic overlooked and this list was incomplete. Moreover, sense (which would require studying the type mate- some unpublished data from collection material be- rial), but only synonyms in the sense of these authors. came available in the last few years. This created a We can only suggest from our experience which spe- need to update that previous checklist. Our new cies can more likely be understood or determined un- checklist includes 155 species: 26 bivalves, 45 fresh- der these names. Names that are marked by water gastropods and 84 terrestrial gastropods. “auctorum non...” refer to a in the sense of Since views on of the freshwater some authors (here mainly STAROBOGATOV and his molluscs are quite controversial, we follow here followers) but not the author of the species name. mainly a recent guide of European non-marine However many of these names are nomina dubia with- molluscs (WELTER-SCHULTES 2012) to make the list out types and we did not study this problem in detail, better available to the European reader. A list of the so it is also possible that some of these names are actu- freshwater molluscs of the Republic of Moldova in the ally correct synomyms in the sense of their authors. classification applied by the Russian malacologists of For this reason we mark such names with “auct. non Starobogatov’s school was published recently and in- [?] ...”. cluded 100 species (MUNJIU 2012). We did not try to Species that were not given in the previous check- make any nomenclatural revision here, but intended list (COADÃ &WELTER-SCHULTES 2011) are high- only to adapt this list of 100 species for the European lighted in bold. We mention only those synonyms reader. Since STAROBOGATOV and his followers fre- which have been in use in Moldova over the last sev- quently use specific names not in the sense of their eral years. 176 Igor A. Balashov, Mikhail O. Son, Viorica Coadã, Francisco Welter-Schultes

MOLLUSCS OF MOLDOVA – CHECKLIST

Gastropoda Gray, 1850 Neritidae Rafinesque, 1815 polita (Hartmann, 1840) Theodoxus cf. danubialis (Pfeiffer, 1828) Bithyniidae Gray, 1857 In past publications T. danubialis has been re- Bithynia leachii (Sheppard, 1823) ported many times from Moldova (YAROSHENKO Syn. B. troschelii (Paasch, 1842), B. inflata (Hansen, 1957, GONTYA 1984, 1985, MUNJIU 2009, 2012, 1845) MUNJIU &SHUBERNETSKI 2010, COADÃ &WELTER- Occurring in the and SCHULTES 2011, WELTER-SCHULTES 2012), but in (YAROSHENKO 1957, GONTYA 1984, 1985, MUNJIU the opinion of some authors (ANISTRATENKO et al. 2012). 1999, SON 2010) an endemic species, Theodoxus Bithynia tentaculata (Linnaeus, 1758) donasteri (Lindholm, 1908) lives in the northern basin of the . According to these authors, Hydrobiidae Stimpson, 1865 reports of T. danubialis and maybe of some other Lithoglyphus naticoides (Pfeiffer, 1828) species refer to this endemic. The last revision of Potamopyrgus antipodarum (Gray, 1843) T. danubialis provided no data about its occurrence Syn. P. jenkinsi (Smith, 1889) in the Dniester and lower sections of other This non-native species was recorded in the Ku- Ponto-Caspian rivers (BUNJE 2007). Some recent churhan Liman (SON 2008, MUNJIU &SHUBER- preliminary molecular data partly supported a sep- NETSKI 2010, MUNJIU 2012). aration of T. donasteri (see BUNJE 2005, SEREDA et Turricaspia lincta (Milaschevich, 1908) al. 2007). More research is needed to clarify the Turricaspia triton (Eichwald, 1838) taxonomic status of this species. A recent report of Thiaridae Troschel, 1857 the marine species Theodoxus pallasi Lindholm, esperi (Férussac, 1823) 1924 (see MUNJIU 2012) requires confirmation for Microcolpia daudebartii (Prevost, 1821) Moldova, where no marine ecosystems exist (see Syn. M. acicularis (Férussac, 1823) discussion below). The Theodoxus species of Moldova require a revision. Physidae Fitzinger, 1833 Theodoxus euxinus (Clessin, 1886) Aplexa hypnorum (Linnaeus, 1758) This brackish water species is present in the ma- Physa acuta Draparnaud, 1805 rine part of the Dniester Liman, but its occurrence Physa fontinalis (Linnaeus, 1758) in the Moldavian part of the Dniester Syn. Physa taslei auct. non [?] Bourguignat, 1860 (MUNJIU &SHUBERNETSKI 2010, MUNJIU 2012) re- Occurring in the Prut River, Kuchurhan Liman quires confirmation. It is possible that fossil shells and other water bodies (YAROSHENKO 1957, were involved or that this occurrence was a result MUNJIU 2009, 2012). of casual transfer by birds or man. Theodoxus fluviatilis (Linnaeus, 1758) Lymnaeidae Rafinesque, 1815 Theodoxus transversalis (Pfeiffer, 1828) Galba truncatula (Müller, 1774) Lymnaea stagnalis (Linnaeus, 1758) Viviparidae Gray, 1847 Myxas glutinosa (Müller, 1774) Viviparus contectus (Millet, 1813) Occurs in the Dniester River (GÎLCÃ et al. 2010, Viviparus viviparus (Linnaeus, 1758) MUNJIU 2012). Gray, 1840 Radix auricularia (Linnaeus, 1758) naticina (Menke, 1845) Radix balthica (Linnaeus, 1758) Reported from the Dniester Basin (YAROSHENKO Syn. R. ovata (Draparnaud, 1805) 1957, MUNJIU 2012). Perhaps extinct in Moldova. It occurs in the Dniester and Prut rivers, Kuchur- Valvata cristata Müller, 1774 han Liman and other water bodies (YAROSHENKO Valvata macrostoma Mörch, 1864 1957, MUNJIU 2012). Syn. V. pulchella auct. non Studer, 1789 Radix labiata (Rossmässler, 1835) Valvata piscinalis (Müller, 1774) Syn. R. peregra (Müller, 1774) Syn. V. profunda auct. non [?] Clessin, 1887, V. Stagnicola palustris (Müller, 1774) ambigua auct. non [?] Westerlund, 1878 Occurring in the Dniester Basin (YAROSHENKO 1957, GONTYA 1984, MUNJIU 2009, 2012). It was Pomatiidae Newton, 1891 probably not determined anatomically, further in- Pomatias rivularis (Eichwald, 1829) vestigation of Moldavian Stagnicola is needed. Checklist of the molluscs of the Republic of Moldova 177

Acroloxidae Thiele, 1931 Turton, 1831 Acroloxus lacustris (Linnaeus, 1758) muscorum (Linnaeus, 1758) Rafinesque, 1815 Pilsbry, 1918 Ancylus fluviatilis Müller, 1774 Columella edentula (Draparnaud, 1805) Anisus septemgyratus (Rossmässler, 1835) It occurs in the Codry Nature Reserve (BAIDASHNI- Anisus spirorbis (Linnaeus, 1758) KOV 1993, COADÃ 2000). Syn. A. dazuri auct. non [?] (Mörch, 1868) Truncatellina claustralis (Gredler, 1856) Anisus vorticulus (Troschel, 1834) It has not been reported in the literature. There Anisus vortex (Linnaeus, 1758) are about 20 shells in the material of A. Bathyomphalus contortus (Linnaeus, 1758) BAIDASHNIKOV from the old forest in the Ferrissia fragilis (Tryon, 1863) Codry Nature Reserve (collection of Non-native species, occurs in the Dniester Basin Schmalhausen Institute of Zoology, Kiev). It was (SON 2007, MUNJIU 2012). also found by V. COADÃ near city (northern acronicus (Férussac, 1807) Moldova, material in private collection). It occurs in the Dniester River and Kuchurhan Truncatellina costulata (Nilsson, 1823) Liman (MUNJIU 2012). It occurs in the Codry Nature Reserve (BAIDASHNI- Gyraulus albus (Müller, 1774) KOV, 1993). Gyraulus crista (Linnaeus, 1758) Truncatellina cylindrica (Férussac, 1807) Syn. G. bielzi auct. non [?] (Kimakowicz, 1884) It occurs in the Codry Nature Reserve (BAIDASHNI- Gyraulus laevis (Alder, 1838) KOV 1993, COADÃ 2000). Reported from the Dniester Basin (YAROSHENKO Vertigo angustior Jeffreys, 1830 1957, MUNJIU 2012). Vertigo antivertigo (Draparnaud, 1801) complanatus (Linnaeus, 1758) Vertigo moulinsiana (Dupuy, 1849) Syn. fontana (Lightfoot, 1786), Hippeutis dia- Vertigo pusilla Müller, 1774 phanella auct. non [?] (Bourguignat, 1864) Vertigo pygmaea (Draparnaud, 1801) H. complanatus occurs in the Kuchurhan Liman Steenberg, 1925 and some other water bodies (YAROSHENKO 1957, MUNJIU 2012). (Draparnaud, 1801) Planorbarius corneus (Linnaeus, 1758) It has not been reported in the literature, but M. Syn. P. purpura auct. non [?] (Müller, 1774), P. SON found this species near Saharna village in grandis auct. non [?] (Dunker, 1850), P. banaticus 2005 (material in private collection). auct. non [?] (Lang, 1856) Woodward, 1903 carinatus Müller, 1774 Brephulopsis cylindrica (Menke, 1828) Planorbis planorbis (Linnaeus, 1758) Probably a non-native species (GONTYA 1984, SY- It occurs in the Dniester and Prut rivers SOEV &SCHILEYKO 2009). Numerous population AROSHENKO ONTYA UNJIU (Y 1957, G 1984, 1985, M was recently observed by V. COADÃ in Chiºinãu. 2009, 2012). tridens (Müller, 1774) Segmentina nitida (Müller, 1774) Merdigera obscura (Müller, 1774) Carychiidae Jeffreys, 1830 Mörch, 1864 minimum Müller, 1774 Balea biplicata (Montagu, 1803) Carychium tridentatum (Risso, 1826) cana (Held, 1836) Cochlicopidae Pilsbry, 1900 Occurring in the Codry Nature Reserve (BAIDASH- Cochlicopa lubrica (Müller, 1774) NIKOV 1993). Cochlicopa lubricella (Porro, 1838) Bulgarica vetusta (Rossmässler, 1836) Reported from northern Moldova (SYSOEV &SCHI- Orculidae Steenberg, 1925 LEYKO 2009). Sphyradium doliolum (Bruguière, 1792) Cochlodina laminata (Montagu, 1803) Cochlodina orthostoma (Menke, 1830) Morse, 1864 plicata (Draparnaud, 1801) Acanthinula aculeata (Müller, 1774) Macrogastra borealis (Boettger, 1878) costata (Müller, 1774) Syn. M. latestriata (Schmidt, 1856) Vallonia enniensis (Gredler, 1856) The species occurs in the Codry Nature Reserve It occurs in the Codry Nature Reserve (BAIDASHNI- (BAIDASHNIKOV 1993). It was also found by V. KOV 1993). COADÃ on the Varatic rock in district (ma- Vallonia excentrica Sterki, 1893 terial in private collection). Vallonia pulchella (Müller, 1774) 178 Igor A. Balashov, Mikhail O. Son, Viorica Coadã, Francisco Welter-Schultes

Ruthenica filograna (Rossmässler, 1836) Deroceras turcicum (Simroth, 1894) Serrulina serrulata (Pfeiffer, 1847) Occurring in the Codry Nature Reserve (BAIDA- SHNIKOV 1993). Ferussaciidae Bourguignat, 1883 Cecilioides (Müller, 1774) Arionidae Gray, 1840 Arion circumscriptus Johnston, 1828 Punctidae Morse, 1864 Arion silvaticus Lohmander, 1937 Punctum pygmaeum (Draparnaud, 1801) Arion subfuscus (Draparnaud, 1805) [sensu lato] Thiele, 1931 Bradybaenidae Pilsbry, 1939 perspectivus (Megerle von Mühlfeld, 1816) Fruticicola fruticum (Müller, 1774) Euconulidae Baker, 1928 Helicidae Rafinesque, 1815 Euconulus fulvus (Müller, 1774) Arianta arbustorum (Linnaeus, 1758) Tryon, 1866 Cepaea vindobonensis (Férussac, 1821) Zonitoides nitidus (Müller, 1774) Helicigona faustina (Rossmässler, 1835) It was found near Saharna village in the canyon of Zonitidae Mörch, 1864 [sensu RIEDEL 2000 and Saharna River by M. SON in 2005, material in the SHILEYKO 2003, non HAUSDORF 1998] collection of the State Natural History Museum in minor (Stabile, 1864) Lvov (GURAL-SVERLOVA &GURAL 2012). Aegopinella pura (Alder, 1830) Helix lutescens Rossmässler, 1837 petronella (Pfeiffer, 1853) Helix pomatia Linnaeus, 1758 It has not been reported in the literature. There is Tryon, 1866 one shell in the material of A. BAIDASHNIKOV from the hornbeam-oak forest in the Codry Nature Re- strigella (Draparnaud, 1801) serve (collection of Schmalhausen Institute of Zo- Helicopsis instabilis (Rossmässler, 1838) ology, Kiev). It was also found by V. COADÃ near Occurring in the Dniester Basin (GONTYA 1984, Soroca city (northern Moldova, material in private SYSOEV &SCHILEYKO 2009). collection). Helicopsis striata (Müller, 1774) Oxychilus glaber (Rossmässler, 1835) Lindholmiola girva (Frivaldszky, 1835) Vitrea contracta (Westerlund, 1871) Monacha cartusiana (Müller, 1774) Vitrea crystallina (Müller, 1774) Perhaps a non-native species (COADÃ &WELTER- Vitrea diaphana (Studer, 1820) SCHULTES 2011). incarnatus (Müller, 1774) Milacidae Ellis, 1926 Monachoides vicinus (Rossmässler, 1842) Tandonia kusceri (Wagner, 1931) Perforatella dibotrion (Bielz, 1860) It was found by V. COADÃ in 2011 in “Valea Far- Pseudotrichia rubiginosa (Rossmässler, 1838) mecilor” city park of Chiºinãu and in Ciorescu vil- Occurs in the Codry Nature Reserve (BAIDASHNI- lage near Chiºinãu (material in private collection, KOV 1993, COADÃ 2000). anatomically determined). Probably a non-native Trochulus hispidus (Linnaeus, 1758) species. Xeropicta derbentina (Krynicki, 1836) Vitrinidae Fitzinger, 1833 Perhaps a non-native species (COADÃ &WELTER- SCHULTES 2011). Vitrina pellucida (Müller, 1774) Xeropicta krynickii (Krynicki, 1833) Limacidae Refinesque, 1815 Perhaps a non-native species (COADÃ &WELTER- Lehmannia jaroslaviae Grossu, 1967 SCHULTES 2011). A single finding near “Kipriani” (Capriana) village Xerolenta obvia (Menke, 1828) (SYSOEV &SCHILEYKO 2009). Syn. X. candicans (Pfeiffer, 1841) Lehmannia marginata (Müller, 1774) Perhaps a non-native species (COADÃ &WELTER- Limax cinereoniger Wolf, 1803 SCHULTES 2011). Limax maximus Linnaeus, 1758 Beck, 1837 Probably a non-native species. Oxyloma elegans (Risso, 1826) Agriolimacidae Wagner, 1935 Oxyloma sarsii (Esmark, 1886) Deroceras agreste (Linnaeus, 1758) Occurs in the Codry Nature Reserve (COADÃ Deroceras laeve (Müller, 1774) 2000). Deroceras reticulatum (Müller, 1774) Succinea putris (Linnaeus, 1758) Deroceras sturanyi (Simroth, 1894) oblonga (Draparnaud, 1801) Checklist of the molluscs of the Republic of Moldova 179

Bivalvia rivicola (Lamarck, 1818) Sphaerium solidum (Normand, 1844) Unionidae Fleming, 1828 Reported from the Dniester River (YAROSHENKO, Anodonta anatina (Linnaeus, 1758) 1957; MUNJIU, 2012). Syn. A. zellensis (Gmelin, 1791), A. piscinalis Nilsson, 1823 Dreissenidae Gray, 1840 Anodonta cygnea (Linnaeus, 1758) Dreissena polymorpha (Pallas, 1771) Pseudanodonta complanata (Rossmässler, 1835) Dreissena rostriformis (Deshayes, 1838) Syn. P. rossmaessleri (Bourguignat, 1870) Represented by its pontic subspecies – D. rostri- Sinanodonta woodiana (Lea, 1834) formis bugensis Andrusov, 1897. Non-native species, Non-native species, occurs in the Prut Basin occurs in Dniester and Prut rivers (MUNJIU 2009, (MUNJIU 2009, MUNJIU &SHUBERNETSKI 2010). 2012, MUNJIU &SHUBERNETSKI 2010). Unio crassus Philipsson, 1788 Corbiculidae Gray, 1847 Unio pictorum (Linnaeus, 1758) Syn. U. longirostris (Rossmässler, 1836) Corbicula fluminea (Müller, 1774) Non-native species, first record in 2009, Prut River Unio tumidus Philipsson, 1788 (MUNJIU &SHUBERNETSKI 2010, MUNJIU 2012). Occurs in the Dniester and Prut rivers (YARO- SHENKO 1957, MUNJIU 2009, 2012). Cardiidae Lamarck, 1809 Deshayes, 1855 Hypanis colorata (Eichwald, 1829) Musculium lacustre (Müller, 1774) Reported from the Kuchurhan Liman (VLADIMI- amnicum (Müller, 1774) ROV 1984, GONTYA 1985, MUNJIU 2012). Listed in Pisidium casertanum (Poli, 1791) the Red Book of Moldova (official national Red List). Pisidium henslowanum (Sheppard, 1823) Hypanis laeviuscula (Martens, 1874) Reported from the Dniester River (YAROSHENKO Reported from the Dniester River and Kuchurhan 1957, MUNJIU 2009, 2012). Liman (YAROSHENKO 1957, VLADIMIROV 1984, Pisidium milium Held, 1836 GONTYA 1985, MUNJIU 2012). Represented by its Syn. P. tetragonum Normand, 1854 pontic subspecies – H. laeviuscula fragilis (Mila- Reported from the Dniester River (MUNJIU 2009, schewitsch, 1908). Listed in the Red Book of Mol- MUNJIU 2012). dova. Pisidium nitidum Jenyns, 1832 Hypanis plicata (Eichwald, 1829) Reported from the Dniester River (YAROSHENKO Reported from the Kuchurhan Liman (GONTYA 1957, MUNJIU 2012). 1985, MUNJIU 2012). Represented by its pontic Pisidium pulchellum Jenyns, 1832 subspecies – H. plicata relicta (Milachevitch, 1916). Reported from the Dniester River (YAROSHENKO Hypanis pontica (Eichwald, 1838) 1957, MUNJIU 2012). Reported from the Dniester River and Kuchurhan Pisidium subtruncatum Malm, 1855 Liman (YAROSHENKO 1957, VLADIMIROV 1984, Reported from the Dniester Basin (YAROSHENKO GONTYA 1985, MUNJIU 2012). Listed in the Red 1957, MUNJIU 2012). Book of Moldova. Pisidium supinum Schmidt, 1851 Sphaerium corneum (Linnaeus, 1758)

DISCUSSION

Besides the listed species several names of marine these reports may refer to the Dniester Liman in and brackish water forms such as Turricaspia variabilis Ukraine or could be based on fossil shells or shells (Eichwald, 1838), T. pseudotriton (Golikov et that were occasionally transferred to Moldova by birds Starobogatov, 1966), T. limanica (Golikov et or man. Moreover the taxonomic status of some of Starobogatov, 1966), Caspia gmelinii Clessin et these forms is unclear and we did not verify their de- Dybowski, 1888, C. knipowitchi Makarov, 1938, C. terminations. Marine ecosystems are absent in makarovi (Golikov et Starobogatov, 1966), Moldova. The only brackish water body in Moldova is Caspiohydrobia eichwaldiana (Golikov et Starobogatov, the Kuchurhan Liman. The Moldavian border is adja- 1966), Ecrobia ventrosa (Montagu, 1803), Hydrobia cent to the Dniester Liman in Ukraine, but it does not acuta (Draparnaud, 1805) and Cerastoderma lamarcki touch it. Some brackish water species invaded the (Reeve, 1845) were reported from the Republic of Dniester River in Moldova. In 1964 the Kuchurhan Moldova (GONTYA 1985, MUNJIU 2012). However Liman was turned into a cooling reservoir of a 180 Igor A. Balashov, Mikhail O. Son, Viorica Coadã, Francisco Welter-Schultes

Moldavian power station, so it is regulated and pol- in Moldova, since these species are quite common in luted. Currently Ponto-Caspian molluscs of adjacent regions of and Ukraine (GROSSU Hydrobiidae and Cardiidae are receding and rare in 1983, BALASHOV &GURAL-SVERLOVA 2012). Kuchurhan, some species are probably extinct, others Several more mollusc species were listed for the are threatened in Moldova. Republic of Moldova in some respectable official Reports of Ena montana (Draparnaud, 1801) and websites. However no references were established. It Helix albescens Rossmässler, 1839 (as H. vulgaris is possible that these reports were based on the confu- Rossmässler, 1839) from the Codry Nature Reserve sion with Moldova, a historical region in adjacent Ro- (GONTYA 1984) and of Daudebardia cavicola Soós, 1927 mania (today officially called the “Principality of from the Dniester valley (VITION 2004) should be Moldavia”). “Moldova” was used in the titles of several confirmed. Reports of Vertigo substriata (Jeffreys, papers about molluscs of Romania only. 1833) and Perforatella bidentata (Gmelin, 1791) from From our observations at least some terrestrial spe- the Codry Nature Reserve (COADÃ 2000) are not con- cies such as Pomatias rivularis, , Vertigo firmed here. moulinsiana, Truncatellina claustralis, Granaria Some species like Pisidium globulare Westerlund, frumentum, Serrulina serrulata, filograna, 1873, P. hibernicum Westerlund, 1894, P. moitessieria- Macrogastra borealis and Discus perspectivus are threat- num Paladilhe, 1866, P. obtusale (Lamarck, 1818), P. ened in the Republic of Moldova and should be in- personatum Malm, 1855, Sphaerium ovale (Férussac, cluded in the official national Red List, the Red Book 1807), S. nucleus (Studer, 1820), Viviparus acerosus of Moldova. The problems of mollusc conservation in Bourguignat, 1862, Cochlicopa nitens (Gallenstein, Moldova should be studied separately and in detail, 1848), Pupilla bigranata (Rossmässler, 1839), this has not been done yet. Chondrina arcadica (Reinhardt, 1881), Discus ruderatus ø (Férussac, 1821), (Str m, 1765), ACKNOWLEDGEMENTS Malacolimax tenellus (Müller, 1774), Deroceras moldavicum (Grossu et Lupu, 1961) and Arion fasciatus We are grateful to Dr. OXANA V. MUNJIU (Institute (Nilsson, 1823) were only suggested to live in the Re- of Zoology, Academy of Sciences of Moldova) for her public of Moldova (WELTER- SCHULTES 2012), but comments regarding a previous manuscript of this pa- there are no known findings so far. At least D. per. ruderatus and N. hammonis are very likely to be found

REFERENCES

ANISTRATENKO O. YU., STAROBOGATOV YA. I., ANISTRATENKO COADÃ V., WELTER-SCHULTES F. 2011. Checklist of land and V. V. 1999. Mollyuski roda Theodoxus (Gastropoda, Pecti- freshwater mollusc species in Moldova. Schrift. Mala- nibranchia, Neritidae) Azovo-Chernomorskogo kozool. 26: 41–46. basseyna. Vestnik Zoologii 3(33): 11–19. GÎLCÃ G., ªTIRBU S., LUCHIANOVA V., GUDZI T., ÞURCANU V., BAIDASHNIKOV A. A. 1993. Nazemnye mollyuski (Gastro- IALOVIÞCAIA N., RUSU E. 2010. Starea ecologicã a apei poda, ) zapovednika Kodry (Moldova). fluviului nistru în baza elementelor hidrobiologice Vestnik Zoologii 4: 10–15. pentru anul de referinþã 2009. Proceedings of the Inter- BALASHOV I., GURAL-SVERLOVA N. 2012. An annotated national Scientific and Practical Conference: Dniester checklist of the terrestrial molluscs of Ukraine. J. Conch. river basin: environmental issues and management trans- 41: 91–109. boundary natural resources. 15–16 October 2010. NGO «Ecospectr», : 60–63. BUNJE P. M. E. 2005. Pan-European phylogeography of the aquatic Theodoxus fluviatilis (Gastropoda: GONTYA F. A. 1984. Klass Bryuhonogiye. In: Mshanki, Neritidae). Mol. Ecol. 14: 4323–4340. http://dx.doi. mollyuski, chlenistonogiye. Seriya Zhivotnyj mir org/10.1111/j.1365-294X.2005.02703.x Moldovy. World of Moldova, Shtinica, Kishinev, pp. 13–30. BUNJE P. M. E. 2007. Fluvial range expansion, allopatry, and parallel evolution in a Danubian snail lineage. Biol. J. GONTYA F. A. 1985. Malakofauna vodoemov basseyna Linn. Soc. 90: 603–617. http://dx.doi.org/10.1111/ Dnestra. Abstract of a thesis for the scientific degree of j.1095-8312.2007.00750.x candidate of biological sciences, Chisinau. COADÃ V. 2000. Malakofauna lesnogo zapovednika “Kodry” GROSSU A. V. 1983. Gastropoda Romaniae 4. Ordo Respubliki Moldova. Fourth (Thirteenth) Conference . Suprafam.: Arionacea, Zonitacea, on the Study of Molluscs: Molluscs: taxonomy, ecology Ariophantacea ºi Helicacea. Editura Litera, Bucureºti. and phylogeny. St. Petersburg, Russia, 27–29 October, GURAL-SVERLOVA N. V., GURAL R. I. 2012. Naukovy kolekcyi 1998. Abstracts, Zoological Institute RAS, St. Petersburg: Derzhavnogo pryrodoznavchogo muzeyu. Vyp. 4. 19–20. Malakolog³chnyj fond. Derzhavnyj pryrodoznachij muzey, Lvov. Checklist of the molluscs of the Republic of Moldova 181

HAUSDORF B. 1998. Phylogeny of the Limacoidea sensu lato SON M. O. 2008. Rapid expansion of the New Zealand mud (Gastropoda, Stylommatophora). J. Mollus. Stud. 64: snail Potamopyrgus antipodarum (Gray, 1843) in the 35–66. http://dx.doi.org/10.1093/mollus/64.1.35 Azov-Black Sea Region. Aquatic Invasions 3: 335–340. MUNJIU O. 2009. The biodiversity of freshwater mollusc in http://dx.doi.org/10.3391/ai.2008.3.3.9 Moldova in the places with different degree of anthro- SON M. O. 2010. Alien mollusks within the territory of pogenic activity. Proceedings of the International Con- Ukraine: sources and directions of invasions. Russian J. ference: Transboundary River Basin Management and Biol. Invasions 2: 37–48. http://dx.doi.org/10.1134/ International Cooperation for Healthy Dniester River. S207511171001008X Odessa, September 30–October 1, 2009: 193–197. STAROBOGATOV YA. I., PROZOROVA L. A., BOGATOV V. V., MUNJIU O. 2012. Sovremennyi vidovoy sostav presnovod- SAYENKO E. M. 2004. Mollyuski. In: Opredilitel presno- nykh mollyuskov Moldovy. Proceeding of the interna- vodnykh bespozvonochnykh Rossii i sopredelnikh terri- tional conference: Geoecological and bioecological toriy. T. 6. Mollyuski, Polikhety, Nemertiny. Nauka, Saint problems of the north Black Sea coast.. Tiraspol, Novem- Petersburg, pp. 9–492. ber 9–10, 2012: 205–207. SYSOEV A., SCHILEYKO A. 2009. Land and slugs of Rus- MUNJIU O., SHUBERNETSKI I. 2010. First record of Asian sia and adjacent countries. Pensoft, Sofia. clam Corbicula fluminea (Müller, 1774) in the Republic of VITION P. 2004. Studiul zoocomplexuluiedafic din lunca Moldova. Aquatic Invasions 5(suppl. 1): 67–70. riului Nistru. Proceedings of the International Confer- http://dx.doi.org/10.3391/ai.2010.5.S1.015 ence: Integrated management of natural resources in RIEDEL A. 2000. Die Sammlung der Paläarktischen Zoni- the transboundary Dniester River Basin. Chisinau, Sep- tidae sensu lato (Gastropoda, Stylommatophora) in dem tember 16–17, Eco-TIRAS: 77–79. Museum und Institut für Zoologie der PAdW in War- VLADIMIROV M. Z. 1984. Klass Dvustvorchatye. In: Mshanki, szawa. Folia Malacol. 8: 37–85. mollyuski, chlenistonogiye. Serija Zhivotnyj mir Moldo- SEREDA S. V., ALBRECHT C., ANISTRATENKO V. V., WILKE T. vy. Animal World of Moldova series, Shtinica, Kishinev. 2007. Diversification of the Theodoxus (Neritidae) pp. 31–45. in the Black Sea Basin. Abstracts of World Congress of YAROSHENKO M. F. 1957. Gidrofauna Dnestra. Izdatelstvo Malacology, Antwerp, , 15–20 July 2007: Akademii Nauk SSSR, Moscow. 203–204. WELTER-SCHULTES F. W. 2012. European non-marine SCHILEYKO A. A. 2003. Treatise on recent terrestrial pulmo- molluscs, a guide for species identification. Planet Poster nate molluscs. Part 10: Ariophantidae, Ostracolethidae, Editions, Göttingen. Ryssotidae, Milacidae, Dyakiidae, Staffordiidea, Gastro- dontidae, Zonitidae, Daudebardiidae, Parmacellidae. Received: May 25th, 2013 Ruthenica (Suppl. 2) 10: 1309–1466. Revised: June 23rd, 2013 SON M. O. 2007. North American freshwater limpet Ferrissia Accepted: June 25th, 2013 fragilis (Tryon, 1863) – the cryptic invader in the North- ern Black Sea Region. Aquatic Invasions 2: 55–58. http://dx.doi.org/10.3391/ai.2007.2.1.6