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Vegetation as Site Indicator Vegetation as Site Indicator Presentation summary

• General ecology - why grow where they do • How plants indicate fertility • Tree and soil relationships with vegetation • Vegetation Identification examples

2 Vegetation as Site Indicator

Limiting Factors

• Every environmental factor has both minimum and maximum levels, beyond which a particular cannot survive.

• E.g. No humans permanently above 5 km

3 Vegetation as Site Indicator

Tolerance limits

4 Vegetation as Site Indicator

Vegetation Studies

• Scientists have studied plant species survival strategies, e.g. stress tolerance, competitiveness and speed of colonisation • Each species has been scored for environmental tolerances, e.g. Nitrogen, salt • Species have been grouped by habitat preferences, e.g. NVC classification

5 Vegetation as Site Indicator

Plant environmental limiting factors

• Temperature • Water • Nutrients • Grazing • Fire • People • Competition from other plants

6 Vegetation as Site Indicator

Plants as Fertility indicators

• Plant species indicate fertility (and wetness) • Tall herbs and annuals suggest RICH • Small woody perennials suggest POOR • Each species has a Soil Nutrient Regime (SNR) score • Some species have wider range than others

7 Vegetation as Site Indicator

Soil Nutrient Regime (SNR) Very Poor Poor Medium Rich Very Rich Moisture and Ellenberg SNR score 2 - 5.7 5.7 - 7.7 7.7 - 9.7 9.7 - 11.7 11.7 - 18 HUMUS MOR MOR - MODER MODER - MULL MULL MULL Slightly Dry wood sage burdock cowberry, bell- wavy hair-grass, raspberry, holly, bluebell (wild elder, yellow archangel, heather, heather common bent, greater stitchwort, hyacinth), hazel, ivy, wood spurge, common Nutrient grid bracken, common cow-wheat, hawthorn, false- hemp-nettle, spear thistle, violet, great chickweed, broom, brome, rosebay white clover, false oat- woodrush, slender gorse, bracken, barren willowherb, grass, hogweed St.John's wort, strawberry germander wavy hair-grass, tormentil, devil's bit speedwell, wood scabious sedge, pignut, common bent, Fresh primrose, cocksfoot, red fescue, yarrow bracken, common violet, great woodrush, blaeberry, heather, wood sorrel, scaly bramble, creeping tufted hair-grass, male dog's mercury, slender St.John's crowberry, green- male fern, hard fern, soft-grass, broad fern, herb robert goosegrass (sticky ribbed sedge heath bedstraw, heath buckler-fern, wood willies), wild garlic, woodrush anemone, foxglove, stinging nettle, hedge wort, tormentil, honeysuckle, wound wort, ground yorkshire fog, sweet ivy, wood avens, devil's bit scabious vernal-grass enchanter's nightshade, Moist lesser celandine, red campion, wood speedwell, common horsetail, creeping thistle, rough meadow-grass

mat grass, heath rush compact rush, lady fern, yellow bugle, wild angelica pimpernel, creeping buttercup, soft rush Soil Soil Moisture Regime (SMR) Very Moist

purple moor-grass, sharp flowered rush marsh thistle wood horsetail, harestail cotton- common valarien, grass, cross leaved meadow sweet, golden Wet heath, deer grass, saxifrage lousewort

common cotton- marsh marigold Very Wet grass, bog myrtle

ALL PLANTS MAYBE FOUND IN CONDITIONS ADJACENT TO THE CELL DISPLAYED 8 N.B.: Plant names in 'red type' are amendments by Scott Wilson, to the published Bulletin 124 _ 15-10-2007 Vegetation as Site Indicator

Soil Nutrient Regime (SNR) Very Poor Poor Medium Rich Very Rich Ellenberg SNR 2 - 5.7 5.7 - 7.7 7.7 - 9.7 9.7 - 11.7 11.7 - 18 HUMUS MOR MOR - MODER MODER - MULL MULL MULL

wood sage burdock Sl. Dry cowberry, bell- wavy hair- raspberry, bluebell (wild elder, yellow heather, grass, common holly, greater hyacinth), archangel, wood Moisture and heather bent, bracken, stitchwort, cow- hazel, ivy, spurge, common hemp- common violet, wheat, hawthorn, nettle, spear thistle, great chickweed, false-brome, white clover, false oat- woodrush, broom, gorse, rosebay grass, hogweed slender bracken, barren willowherb, St.John's wort, strawberry germander Nutrient grid tormentil, devil's speedwell, Fresh bit scabious wood sedge, pignut, primrose, cocksfoot, red fescue, • Each species has a yarrow

blaeberry, wood sorrel, bramble, tufted hair- dog's mercury, different range across heather, scaly male creeping soft- grass, male goosegrass (sticky crowberry, fern, hard fern, grass, broad fern, herb willies), wild garlic, green-ribbed heath bedstraw, buckler-fern, robert stinging nettle, nutrient and wetness sedge heath woodrush wood hedge wound wort, anemone, ground ivy, wood foxglove, avens, enchanter's

honeysuckle, nightshade, lesser Moist classes yorkshire fog, celandine, red sweet vernal- campion, wood grass speedwell, common

Bramble Soil Moisture (SMR) Regime horsetail, creeping thistle, rough meadow-grass

mat grass, compact rush, lady fern, bugle, wild heath rush yellow angelica pimpernel, creeping

buttercup, soft V. Moist • So the SNR value is an rush

purple moor- sharp flowered marsh thistle wood average grass, harestail rush horsetail, cotton-grass, common cross leaved valarien, Wet heath, deer meadow sweet, grass, golden saxifrage lousewort common cotton- marsh marigold grass, bog

myrtle Very Wet

ALL PLANTS MAYBE FOUND IN CONDITIONS ADJACENT TO THE CELL DISPLAYED 9 N.B.: Plant names in 'red type' are amendments by Scott Wilson, to the published Bulletin 124 _ 15-10-2007 Vegetation as Site Indicator

Soil Nutrient Regime (SNR) Very Poor Poor Medium Rich Very Rich Ellenberg SNR 2 - 5.7 5.7 - 7.7 7.7 - 9.7 9.7 - 11.7 11.7 - 18 HUMUS MOR MOR - MODER MODER - MULL MULL MULL

wood sage burdock Sl. Dry cowberry, bell- wavy hair- raspberry, bluebell (wild elder, yellow heather, grass, common holly, greater hyacinth), archangel, wood Moisture and heather bent, bracken, stitchwort, cow- hazel, ivy, spurge, common hemp- common violet, wheat, hawthorn, nettle, spear thistle, great chickweed, false-brome, white clover, false oat- woodrush, broom, gorse, rosebay grass, hogweed slender bracken, barren willowherb, St.John's wort, strawberry germander Nutrient grid tormentil, devil's speedwell, Fresh bit scabious wood sedge, pignut, primrose, cocksfoot, red fescue, • Each species has a yarrow

blaeberry, wood sorrel, bramble, tufted hair- dog's mercury, different range across heather, scaly male creeping soft- grass, male goosegrass (sticky crowberry, fern, hard fern, grass, broad fern, herb willies), wild garlic, green-ribbed heath bedstraw, buckler-fern, robert stinging nettle, nutrient and wetness sedge heath woodrush wood hedge wound wort, anemone, ground ivy, wood foxglove, avens, enchanter's

honeysuckle, nightshade, lesser Moist classes yorkshire fog, celandine, red sweet vernal- campion, wood grass speedwell, common

Heather Soil Moisture (SMR) Regime horsetail, creeping thistle, rough meadow-grass

mat grass, compact rush, lady fern, bugle, wild heath rush yellow angelica pimpernel, creeping

buttercup, soft V. Moist • So the SNR value is an rush

purple moor- sharp flowered marsh thistle wood average grass, harestail rush horsetail, cotton-grass, common cross leaved valarien, Wet heath, deer meadow sweet, grass, golden saxifrage lousewort common cotton- marsh marigold grass, bog

myrtle Very Wet

ALL PLANTS MAYBE FOUND IN CONDITIONS ADJACENT TO THE CELL DISPLAYED 1 N.B.: Plant names in 'red type' are amendments by Scott Wilson, to the published Bulletin 124 _ 15-10-2007 0 Vegetation as Site Indicator

Soil Nutrient Regime (SNR) Very Poor Poor Medium Rich Very Rich Ellenberg SNR 2 - 5.7 5.7 - 7.7 7.7 - 9.7 9.7 - 11.7 11.7 - 18 HUMUS MOR MOR - MODER MODER - MULL MULL MULL

wood sage burdock Sl. Dry cowberry, bell- wavy hair- raspberry, bluebell (wild elder, yellow heather, grass, common holly, greater hyacinth), archangel, wood Moisture and heather bent, bracken, stitchwort, cow- hazel, ivy, spurge, common hemp- common violet, wheat, hawthorn, nettle, spear thistle, great chickweed, false-brome, white clover, false oat- woodrush, broom, gorse, rosebay grass, hogweed slender bracken, barren willowherb, St.John's wort, strawberry germander Nutrient grid tormentil, devil's speedwell, Fresh bit scabious wood sedge, pignut, primrose, cocksfoot, red fescue, • Each species has a yarrow

blaeberry, wood sorrel, bramble, tufted hair- dog's mercury, different range across heather, scaly male creeping soft- grass, male goosegrass (sticky crowberry, fern, hard fern, grass, broad fern, herb willies), wild garlic, green-ribbed heath bedstraw, buckler-fern, robert stinging nettle, nutrient and wetness sedge heath woodrush wood hedge wound wort, anemone, ground ivy, wood foxglove, avens, enchanter's

honeysuckle, nightshade, lesser Moist classes yorkshire fog, celandine, red sweet vernal- campion, wood grass speedwell, common

Bell Soil Moisture (SMR) Regime horsetail, creeping thistle, rough meadow-grass

Heather mat grass, compact rush, lady fern, bugle, wild heath rush yellow angelica pimpernel, creeping

buttercup, soft V. Moist • So the SNR value is an rush

purple moor- sharp flowered marsh thistle wood average grass, harestail rush horsetail, cotton-grass, common cross leaved valarien, Wet heath, deer meadow sweet, grass, golden saxifrage lousewort common cotton- marsh marigold grass, bog

myrtle Very Wet

ALL PLANTS MAYBE FOUND IN CONDITIONS ADJACENT TO THE CELL DISPLAYED 1 N.B.: Plant names in 'red type' are amendments by Scott Wilson, to the published Bulletin 124 _ 15-10-2007 1 Vegetation as Site Indicator

Some species are more “specialist indicators” than others

1 2 Vegetation as Site Indicator

Soil Nutrient Regime (SNR) Very Poor Poor Medium Rich Very Rich Ellenberg SNR 2 - 5.7 5.7 - 7.7 7.7 - 9.7 9.7 - 11.7 11.7 - 18 HUMUS MOR MOR - MODER MODER - MULL MULL MULL wavy hair- wood sage burdock

grass, Sl. Dry cowberry, bell- wavy hair- raspberry, bluebell (wild elder, yellow common bent, heather, grass, common holly, greater hyacinth), archangel, wood heather bent, bracken, stitchwort, cow- hazel, ivy, spurge, common hemp- common violet, wheat, hawthorn, nettle, spear thistle, bracken, great chickweed, false-brome, white clover, false oat- woodrush, broom, gorse, rosebay grass, hogweed common slender bracken, barren willowherb, St.John's wort, strawberry germander violet, great tormentil, devil's speedwell, Fresh bit scabious wood sedge, woodrush, pignut, primrose, cocksfoot, slender red fescue, St.John's wort yarrow

blaeberry, wood sorrel, bramble, tufted hair- dog's mercury, tormentil, heather, scaly male creeping soft- grass, male goosegrass (sticky crowberry, fern, hard fern, grass, broad fern, herb willies), wild garlic, devil's bit green-ribbed heath bedstraw, buckler-fern, robert stinging nettle, sedge heath woodrush wood hedge wound wort, scabious, anemone, ground ivy, wood foxglove, avens, enchanter's

honeysuckle, nightshade, lesser Moist yorkshire fog, celandine, red • Each species has a different “footprint” sweet vernal- campion, wood grass speedwell, common

Soil Moisture (SMR) Regime horsetail, creeping thistle, • Range may vary on conditions rough meadow-grass mat grass, compact rush, lady fern, bugle, wild heath rush yellow angelica pimpernel, • Species overlap creeping

buttercup, soft V. Moist rush

purple moor- sharp flowered marsh thistle wood SNR must be estimated grass, harestail rush horsetail, cotton-grass, common cross leaved valarien, from presence and Wet heath, deer meadow sweet, grass, golden saxifrage lousewort abundance of several common cotton- marsh marigold grass, bog myrtle characteristic species Very Wet ALL PLANTS MAYBE FOUND IN CONDITIONS ADJACENT TO THE CELL DISPLAYED N.B.: Plant names in 'red type' are amendments by Scott Wilson, to the published Bulletin 124 _ 15-10-2007

1 3 Vegetation as Site Indicator Working out SNR - Numerical Method

species abundance [%] SNR value product

tufted hair grass 50 (5) 9 45 wavy hair grass 30 (3) x 5 = 15

soft rush 10 (1) 8 8

sum 90 (9) 68

use the 5 most abundant species

weighted average SNR = 68 / 9 = 7.5 poor - medium 1 4 Vegetation as Site Indicator Working out SNR - Shortcut Method Soil Nutrient Regime (SNR) Very Poor Poor Medium Rich Very Rich Ellenberg SNR 2 - 5.7 5.7 - 7.7 7.7 - 9.7 9.7 - 11.7 11.7 - 18 HUMUS MOR MOR - MODER MODER - MULL MULL MULL

wood sage burdock Sl. Dry cowberry, bell- wavy hair- raspberry, bluebell (wild elder, yellow heather, grass, common holly, greater hyacinth), archangel, wood heather bent, bracken, stitchwort, cow- hazel, ivy, spurge, common hemp- wavy hair grass common violet, wheat, hawthorn, nettle, spear thistle, great chickweed, false-brome, white clover, false oat- woodrush, broom, gorse, rosebay grass, hogweed slender bracken, barren willowherb, St.John's wort, strawberry germander tormentil, devil's speedwell, Fresh bit scabious wood sedge, tufted hair grass pignut, primrose, cocksfoot, red fescue, yarrow

blaeberry, wood sorrel, bramble, tufted hair- dog's mercury, soft rush heather, scaly male creeping soft- grass, male goosegrass (sticky crowberry, fern, hard fern, grass, broad fern, herb willies), wild garlic, green-ribbed heath bedstraw, buckler-fern, robert stinging nettle, sedge heath woodrush wood hedge wound wort, anemone, ground ivy, wood foxglove, avens, enchanter's

honeysuckle, nightshade, lesser Moist yorkshire fog, celandine, red sweet vernal- campion, wood grass speedwell, common

Soil Moisture (SMR) Regime horsetail, creeping thistle, rough meadow-grass

mat grass, compact rush, lady fern, bugle, wild heath rush yellow angelica pimpernel, creeping

buttercup, soft V. Moist medium rush purple moor- sharp flowered marsh thistle wood grass, harestail rush horsetail, cotton-grass, common cross leaved valarien,

moist Wet heath, deer meadow sweet, grass, golden saxifrage lousewort common cotton- marsh marigold grass, bog

use the 5 most abundant species myrtle Very Wet consider abundance of species ALL PLANTS MAYBE FOUND IN CONDITIONS ADJACENT TO THE CELL DISPLAYED N.B.: Plant names in 'red type' are amendments by Scott Wilson, to the published Bulletin 124 _ 15-10-2007 1 5 Rough peatland species distribution

flushed unflushed wet unflushed dry

8 9a 9b 9c 9d 9e 10 11b 11c 11a (11d)

Calluna vulgaris (heather) / Vaccinium myrtillus (blaeberry)

Erica tetralix (heath)

Molinia caerulea (rushes) (purple moor grass)

Trichophorum cespitosum

(deergrass) Juncussp. Narthecium and Rhacomitrum lichens Agrostis sp. Eriophorum sp. (bents) (cotton grass) ESC – Nutrient indicator plants

SNR Taxonomic name R N Rubus chamaemorus 1 1 2 Betula nana 1 1 Empetrum nigrum 2 1

Erica tetralix 2 1 3 Eriophorum vaginatum 2 1 cespitosum 2 1 Drosera intermedia & rotundifolia 2 1 Narthecium ossifragum 2 1 Arctostaphylos uva-ursi 2 2 Erica cinerea 2 2 Vaccinium vitis-idaea 2 2 4 vulgaris 2 2 Listera cordata 2 2 Vaccinium myrtillus 2 2 2 2

Purple - Heaths

1 7 ESC – Nutrient indicator plants

SNR Taxonomic name R N Moneses uniflora 4 1 4 1 Goodyera repens 3 2 Poa flexuosa 3 2 Salix herbacea 3 2 Agrostis vinealis 3 2 3 2 5 Festuca vivipara 3 2 Nardus stricta 3 2 erecta 3 2 caerulea 3 2 Pedicularis sylvatica 3 2 Myrica gale 3 2 flexuosa 2 3 Melampyrum pratense 2 3

Blue – Grasses (Phosphate)

1 8 Vegetation Identification

Vegetation identification - Families

Plants are grouped into families by general features, e.g.: • All grasses have hollow stems • All sedges have triangular stems • Rushes have solid or pithy stems

1 9 Vegetation Identification

Vegetation Identification - Species

• Every individual plant species has unique features • Some are easier to spot than others

2 0 Vegetation Identification

Deschampsia cespitosa - Tufted hair grass

• Very tall, loose seed head

2 1 Vegetation Identification

Deschampsia cespitosa - Tufted hair grass

• Very tall, loose seed head • Forms a large tussock

2 2 Vegetation Identification

Deschampsia cespitosa - Tufted hair grass

• Very tall, loose seed head • Forms a large tussock • are striped when held to the light (are rough in one direction only)

2 3 Vegetation Identification

Deschampsia cespitosa - Tufted hair grass SNR values for all indicator species can be found in FC bulletin 124, p. 22/23. Add columns N and R for total value!

Soil Nutrient Regime (SNR) Very Poor Poor Medium Rich Very Rich 2 - 5.7 5.7 - 7.7 7.7 - 9.7 9.7 - 11.7 11.7 - 18

2 4 Vegetation Identification

Molinia caerulea - Purple moor grass

• Thin seed head

• Forms a hard tussock

• Leaves persist over the winter

2 5 Vegetation Identification

Molinia caerulea - Purple moor grass

• Thin seed head

• Forms a hard tussock

• Leaves persist over the winter

• Root base is bulbous

2 6 Vegetation Identification

Molinia caerulea - Purple moor grass

Soil Nutrient Regime (SNR) Very Poor Poor Medium Rich Very Rich 2 - 5.7 5.7 - 7.7 7.7 - 9.7 9.7 - 11.7 11.7 - 18

2 7 Vegetation Identification

Erica tetralix - Cross-leaved heath

• Dwarf shrub • Woody stems • Waxy coating on leaves

2 8 Vegetation Identification

Erica tetralix - Cross-leaved heath • Dwarf shrub • Woody stems • Waxy coating on leaves • Leaflets form a cross shape on the stem

2 9 Vegetation Identification

Erica tetralix - Cross-leaved heath

Soil Nutrient Regime (SNR) Very Poor Poor Medium Rich Very Rich 2 - 5.7 5.7 - 7.7 7.7 - 9.7 9.7 - 11.7 11.7 - 18

3 0 Vegetation Identification Tree - Vegetation associations

• Purple moor grass - Downy birch, Sitka spruce • Heather - Scots pine, Silver birch • Nettle - Alder • Dogs Mercury - Ash • And many more….

3 1 Vegetation Identification

Mineral Soil - Vegetation associations

• Brown earths - Grasses, herbs and ferns • Podzols - Heather and fine grasses • Peaty gleys - Purple moor grass • Surface-water gley - Tufted hair-grass

3 2 Vegetation Identification

General Vegetation sampling

• Select an area of similar uniform vegetation • Pick some representative sampling areas • Split the vegetation into layers • Identify every species possible • Estimate the relative abundance (%)

3 3 Vegetation as Site Indicator Summary

• Plants indicate fertility • Trees and soils have relationships with other vegetation • Learn your plant ID…

• You must go and L K

3 4