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Conspectus Familiae Pannariaceae (Ascomycetes Lichenosae). by Per

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Conspectus familiae Pannariaceae (Ascomycetes lichenosae). By Per M. Jørgensen Botanisk institutt Universitetet i Bergen Revised version 2006 1 ISBN-82-7460-014-2 ISSN-0807-2116 2 Jørgensen, P.M. 2002. Conspectus familiae Pannariaceae (Ascomycetes lichenosae). – Ilicifolia 4: 1-78. ISBN-82-7460-014-2. ISSN -0807-2116. (Revised version 2006) Abstract: This conspectus lists all the names for lichens in the family Pannariaceae. No nomenclatural novelities are presented except for some necessary typifications. The list is available on the net and will be updated yearly. A key to the accepted genera is included in the introduction. Per Magnus Jørgensen, Department of Natural history,Bergen Museum, University of Bergen, Allégaten 41, N-5007 Bergen, Norway. CONTENTS INTRODUCTION …………………………………………….…… 4 KEY …………………………………………………………….….. 5 LIST …………………………………………………………………7 CONCLUSIVE REMARKS ……………………………………… 81 ACKNOWLEDGEMENTS ………………………………………. 81 REFERENCES …………………………………………………….82 3 INTRODUCTION difficulties in tracing the high number of such, often unimportant It is now 28 years since I published my thesis on the European Pannariaceae (Jørgensen 1978), and in the meantime I names, and their types. All epithets which have published a number of papers on its have been used on specific level are in different genera from all parts of the principle included, as well as such which world. I had originally intended to make a have been clarified with readily available full world-monograph, but particularly in types. the later years I have found it more amenable to publish smaller, regional Surely further additions and adjustments treatments. In the following I try to give a will be necessary in the future, but I hope nomenclatural conspectus of the family the following will prove a useful basis for with references to the relevant papers further research. The list of names in this where a fuller taxonomic treatment can be conspectus will also be available on the found. My intention is to present an net, and will be yearly updated there, with updated and improved "Zahlbruckner" for printed versions appearing when the names in the Pannariaceae. I follow necessary. Ekman & Jørgensen (2002) in the circumscription of the family. The names are given in alphabetical order from generic level down to form. Names in bold are the correct ones for taxa presently recognized by me, and full citation including citation of types are found there. If names originally were published in another genus of the Pannariaceae, they are cited from their original publication as well, with reference to their present position. Names of species which I recognize, but which are of uncertain taxonomic position at the moment, are marked by an asterisk. Uncertainties are otherwise indicated by a question mark. I have not presented any new taxa, or proposed any new combinations or names in this list, though I have cited some names which are in ed. In order not to create any doubt I have omitted the page references where new combinations in ed. are cited. Accordingly no nomenclature novelities are presented, except that in a few cases where new lectotypes had to be designated. The coverage of names in lower ranks are incomplete due to 4 KEY TO GENERA OF PANNARIACEAE Though this is basically a nomenclatural conspectus, it may be useful also to present an introductory key to the accepted genera: 1 Thallus gelatinous, homoiomerous, PD- ………….………………..…………..…2 1 Thallus mostly not gelatinous, or if so with distinct upper cortex and/or PD+ orange ………………………………………………………………….……….………...3 2(1) Thallus usually fertile, without gymnidia and terpenes; neotropical …………………………………………………...……………… 8 Lepidocollema 2 Thallus usually sterile, with bluish gymnidia and terpenes; paleotropical ………………………………………………………………...………... 7 Kroswia 3(1) Thallus totally dissolved into blue-grey grains, without lichen substances …………………………………………...…………..…………... 10 Moelleropsis 3 Thallus not totally dissolved into granules, usually containing lichen substances ………………………………………………………………..………………….. 4 4(3) Cyanobacteria penetrating into the subhymenial parts ………...…………….…... 5 4 No cyanobacteria in the subhymenial parts …………...…………………….…….7 5(4) Thallus foliose; upper surface with stiff hairs, mostly PD+ orange …………………………………………………………………………4 Erioderma 5 Thallus mostly of squamules; upper surface glabrous, or if hairy, arachnoid, usually PD- …………………………………………………………………………..……….. 6 6(5) Thallus subfoliose, bluish, upper surface cobwebby, asci with I+ blue cap ..….……………………………………………………...…………… 8 Leioderma 6 Thallus squamulose, brownish, upper surface at most scabrid, asci without I+ blue cap …………………………………………………………….…… 5 Fuscoderma 7(4) Apothecia without thalline margin ………………….………...…………..….….. 8 7 Apothecia normally with thalline margin …………………………………...….. 14 8(7) Hymenium hemiamyloid, finally red-brown ……………..…………………..….. 9 8 Hymenium amyloid, constantly blue ………………….…..……………….…… 12 9(8) Thallus gelatinous with cortex on both surfaces ……………….. 16 Santessoniella 9 Thallus not gelatinous, cortex only on upper surface …………………………... 10 10(9) Thallus PD+; cortex sclerenchymatous ………………………...…17 Siphulastrum 5 10 Thallus PD-; cortex paraplectenchymatous …………………………………..… 11 11(10) Thallus small-squamulose; apothecia surrounded by a weft of hyphae ………………………………...……………………………………….. 1 Austrella 11 Thallus rosettiform-placodioid; apothecia without such a weft …...…. 3 Degeliella 12(8) Thallus placodioid, with dense medulla of parallel hyphae; ascus with sheet-like amyloid apical structures ……………………………………..………... 2 Degelia 12 Thallus small-squamulose, with medulla of unorientated hyphae; asci with tubular, amyloid apical structures ……...……………………………………………….. 13 13(12) Thallus with green photobiont and cephalodia ……………..…….15 Psoromidium 13 Thallus with cyanobacteria, no cephalodia ......................………… 12 Parmeliella 14(7) Prothallus feltet, prominent, thallus a closely attached rosette; hymenium I+ blue, asci with tubular amyloid apical structures; subtropical ……………………………………………….…...……Parmeliella mariana group 14 Prothallus not prominent or felted, thallus loosely attached; hymenium hemiamyloid, or when I+ blue without amyloid structures in asci …………….. 15 15(14) Thallus squamulose to subfoliose; hymenium I+ blue, asci without amyloid apical structures ………………………………………………………………………...…………16 15 Thallus crustose to small-squamulose; hymenial reaction otherwise, asci with amyloid apical structures ……………………………………………………….. 17 16(15) Thallus small-squamulose, PD-, brownish; perispore warted; bipolar ………………………………………………………...…………13 Protopannaria 16 Thallus subfoliose, usually PD+, bluish gray; perispore at most uneven; subtropical to warm-temperate ……………………...………………. 11 Pannaria 17(15) Apothecia with distinct thalline margin; spores markedly warted; hymenium I+ dirty blackish blue; bipolar ……………………………………..…… 14 Psoroma 17 Apothecia with variable thalline margin, sometimes poorly developed, spores at most uneven; hymenium I+ blue-green, turning red-brown; temperate ……………………………………………………………...…….6 Fuscopannaria The , Lepidocollema and Moelleropsis in this key are mainly based on thallus characters and may eventually need to be included in other genera. On the other, hand some genera, particularly Psoroma s. lat., need to be split in further genera. 6 LIST OF NAMES 1. AUSTRELLA P.M. Jørg., Bibl. Lich.88: 230 (2004). - Generitype: Austrella arachnoidea P.M. Jørg. Small, recently recognized Southern Hemisphere genus, as yet only known from SE Australia and New Zealand. Austrella arachnoidea P.M. Jørg., Bibl. Lich.88: 230 (2004). - Type: Australia, A. C. T., Condor Creek, 700 m, 31. Aug. 1981, P. M. Jørgensen 8200 (BG, holotype!). Austrella brunnea (P.M. Jørg.) P.M. Jørg., Bibl. Lich.88: 231 (2004); Santessoniella brunnea P.M. Jørg., New Zeal. Journ. Bot. 37 (1999). - Type: New Zealand, South Isl., Canterbury, Cass Hill, P.M. Jørgensen 8469 (BG, holotype!). 2. DEGELIA Arv. & D.J. Galloway, Lichenologist 13: 28 (1981). - Generitype: Degelia gayana (Mont.) Arv. & D.J. Galloway This genus was from its description regarded as part of the Pannariaceae, and has been treated as such by me in several papers. However, its inclusion in the family is uncertain (Ekman & Jørgensen 2002), but it is in spite of this treated here since the names are so closely linked with those of Parmeliella, and its position needs to be studied more in detail before the final decision is taken. In a recent analysis Wiklund & Wedin (2003) showed that Degelia groups with Erioderma and Leioderma, which appears reasonable. However, the ranking of this group is still unclear. The genus is heterogeneous and has been split in three sections: sect. Amphiloma (Fr.) P.M. Jørg. & P. James (cold temperate, N. Hemisphere), sect. Degelia (warm temperate S. Hemisphere), and sect. Frigidae P.M. Jørg. (subantarctic-alpine, predominantly S. Hemisphere.). Degelia atlantica (Degel.) P.M. Jørg. & P. James, Bibl. Lich. 38: 264 (1990). Parmeliella atlantica Degel., Acta Phyt. Suec. 7: 131-133 (1935). - Type: Ireland. Killarney, near Muckross Lake, 1933, G. Degelius (UPS, holotype!). Degelia calcicola P.M. Jørg. & Kantvilas in Jørgensen, Kantvilas & Galloway, Lichenologist 32: 257 (2000). - Type: Australia, Tasmania, Lake Sydney, on limestone, 680 m, G. Kantvilas 44/98 (HO, holotype!). Degelia crustacea P.M. Jørg.& D.J. Galloway, Lichenologist 32: 259 (2000). -Type: New Zealand, North Island, Auckland, Wattakere Range, Piha stream, 17. March 1979, J.K. Bartlett 32436a (AK). Degelia duplomarginata (P. James
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  • BLS Bulletin 111 Winter 2012.Pdf

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    1 BRITISH LICHEN SOCIETY OFFICERS AND CONTACTS 2012 PRESIDENT B.P. Hilton, Beauregard, 5 Alscott Gardens, Alverdiscott, Barnstaple, Devon EX31 3QJ; e-mail [email protected] VICE-PRESIDENT J. Simkin, 41 North Road, Ponteland, Newcastle upon Tyne NE20 9UN, email [email protected] SECRETARY C. Ellis, Royal Botanic Garden, 20A Inverleith Row, Edinburgh EH3 5LR; email [email protected] TREASURER J.F. Skinner, 28 Parkanaur Avenue, Southend-on-Sea, Essex SS1 3HY, email [email protected] ASSISTANT TREASURER AND MEMBERSHIP SECRETARY H. Döring, Mycology Section, Royal Botanic Gardens, Kew, Richmond, Surrey TW9 3AB, email [email protected] REGIONAL TREASURER (Americas) J.W. Hinds, 254 Forest Avenue, Orono, Maine 04473-3202, USA; email [email protected]. CHAIR OF THE DATA COMMITTEE D.J. Hill, Yew Tree Cottage, Yew Tree Lane, Compton Martin, Bristol BS40 6JS, email [email protected] MAPPING RECORDER AND ARCHIVIST M.R.D. Seaward, Department of Archaeological, Geographical & Environmental Sciences, University of Bradford, West Yorkshire BD7 1DP, email [email protected] DATA MANAGER J. Simkin, 41 North Road, Ponteland, Newcastle upon Tyne NE20 9UN, email [email protected] SENIOR EDITOR (LICHENOLOGIST) P.D. Crittenden, School of Life Science, The University, Nottingham NG7 2RD, email [email protected] BULLETIN EDITOR P.F. Cannon, CABI and Royal Botanic Gardens Kew; postal address Royal Botanic Gardens, Kew, Richmond, Surrey TW9 3AB, email [email protected] CHAIR OF CONSERVATION COMMITTEE & CONSERVATION OFFICER B.W. Edwards, DERC, Library Headquarters, Colliton Park, Dorchester, Dorset DT1 1XJ, email [email protected] CHAIR OF THE EDUCATION AND PROMOTION COMMITTEE: S.