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http://www.beatymuseum.ubc.ca/herbarium/prints/printPmunitum.jpg Propagation Protocol for munitum (western swordfern) ESRM 412 – Native Plant Production

TAXONOMY Family Names Family Scientific Name: Family Common Wood Family1 Name: Scientific Names : Polystichum Roth Species: P. munitum Species (Kaulf.) C. Presl Authority: Variety: NA Sub-species: curtum, imbricans, munitum, nudatum, imbricans, incisoserratum3 Cultivar: NA Authority for NA Variety/Sub- species: Common (Kaulfuss) K. Presl incisoserratum Synonym(s) (include full scientific names (e.g., Elymus glaucus Buckley), including variety or subspecies information) Common Western Swordfern1, Common Sword Fern1, Western Sword Fern3, Name(s): Pineland Swordfern6, Giant Holly Fern9

Species Code (as POMU per USDA database): GENERAL INFORMATION Geographical range (distribution maps for and Washington state)

http://plants.usda.gov/maps/large/PO/POMU.png Grows along the west coast from southeastern to Santa Barbara County, , and eastward through Washington and northern into northwest . Disjunct populations have been found in and on Guadalupe Island off Baja California. 1, 5

Ecological Terrestrial, coniferous forest floor, mixed evergreen forests, shrubby or distribution grass spaces, and at higher elevations growing among rocks. Also, (ecosystems it occasionally found in wetlands.6, 8 occurs in, etc): Climate and Moist forest, Western sword fern grows from sea level to mid-elevations in elevation range the mountains throughout its range, primarily below 2500 feet elevation.1, 2

Local habitat and Can be found extensively in Western Washington and one of the most abundance; may abundant in the western flora; It is also extensively used in include landscaping with the trade being mainly in wild-collected plants.1 commonly associated species Plant strategy Facultative Seral Species4 type / successional stage (stress- tolerator, competitor, weedy/colonizer , seral, late successional) Plant Fern characteristics (life form (shrub, grass, forb), longevity, key characteristics, etc) PROPAGATION DETAILS Ecotype (this is Muir Woods, California10 meant primarily for experimentally derived protocols, and is a description of where the seed that was tested came from): Propagation Goal Plants10 (Options: Plants, Cuttings, Seeds, Bulbs, Somatic Embryos, and/or Other Propagules): Propagation Seed (spores)10 Method Division: Sword ferns may be divided in spring if the rhizome is large and (Options: Seed the roots are well developed in the Spring.4 or Vegetative): Product Type Container (plug)10 (options: Container (plug), Bareroot (field grown), Plug + (container-field grown hybrids, and/or Propagules (seeds, cuttings, poles, etc.)) Stock Type: 4 inch pots10

Time to Grow 2 Years10 (from seeding until plants are ready to be outplanted): Target Height: N/A Specifications Caliper: N/A (size or Root System: Firm plug in container10 characteristics of target plants to be produced): Propagule Spores are collected between June 1st and August 30th. Collection Mature spores are brown and are collected when the indusium folds back.10 (how, when, etc): Propagule Spore Cleaning: All chaff must be carefully cleaned from the spores. Processing/Prop Storage Conditions: Spores are kept dry and stored in a refrigerator.10 agule Spore viability highly variable, usually low after 1 year.4 Characteristics Store spores in glassine envelopes or in packets or waxed paper. Store (including seed packets in 1-4 degrees C, in moisture-tight and air-tight containers.5 density (# per pound), seed longevity, etc): Pre-Planting No dormancy breaking requirements.5 Propagule Carefully sterilize growing media, flats and flat coverings before sowing Treatments spores.10 (cleaning, dormancy treatments, etc): Growing Area Greenhouse. Preparation / Spores are sown on petri plates with distilled water and sealed in a sterile Annual ziplock bag.10 Practices for Perennial Crops (growing media, type and size of containers, etc): Establishment After spores germinate, the prothalli are placed on sterile 3:1 Peat/Perlite Phase (from mix in rectangular flats. Flats are covered with glass to seal in moisture. seeding to Water only with distilled water.10 germination): Length of NA Establishment Phase: Active Growth Once fertilization occurs and the developing sporophytes at least 1 true leaf, Phase (from they are potted into individual containers (2 or 4 inch) containing standard germination potting mix of peat moss, fir bark, perlite, and sand. until plants are Plants should be grown in shade.10 no longer actively growing): Length of Active Evergreen, fast growing/ spreading.4 Growth Phase: Hardening Phase Plants grown from spores should be 1.5-2 years old before planting to (from end of ensure survival.5 active growth phase to end of growing season; primarily related to the development of cold-hardiness and preparation for winter): Length of NA Hardening Phase: Harvesting, NA Storage and Shipping (of seedlings): Length of Storage NA (of seedlings, between nursery and outplanting): Guidelines for Recommended planting density 18-24 in. Outplanting / Average Water Needs; Water regularly; do not overwater. Soil pH 6.1 to Performance on 6.5 (mildly acidic).5 Typical Sites (eg, percent survival, height or diameter growth, elapsed time before flowering): Other Comments NA (including collection restrictions or guidelines, if available): INFORMATION SOURCES References (full 1. Pojar, Jim, A. MacKinnon, and Paul B. Alaback. Plants of the citations): Pacific Northwest coast: Washington, Oregon, & Alaska. Redmond, Wash.: Lone Pine Pub., 1994. Print. 2. " Calflora: Polystichum munitum ." Calflora - Search for Plants. N.p., n.d. Web. 20 Apr. 2011. . 3. Houston, M., student in Geography 316, and Fall 2000. "Biogeography of swordfern (Polystichum munitum)."redirect. N.p., n.d. Web. 20 Apr. 2011.. 4. Jones, David Lloyd.Encyclopaedia of ferns: an introduction to ferns, their structure, biology, economic importance, cultivation, and propagation. Portland, Or.: Timber Press, 1987. Print. 5. Kruckeberg, Arthur R..Gardening with native plants of the Pacific Northwest: an illustrated guide. Seattle: University of Washington Press, 1982. Print. 6. "PLANTS Profile for Polystichum munitum (western swordfern) | USDA PLANTS."Welcome to the PLANTS Database | USDA PLANTS. N.p., n.d. Web. 20 Apr. 2011. . 7. "Polystichum munitum." US Forest Service - Caring for the land and serving people.. N.p., n.d. Web. 20 Apr. 2011. . 8. "Polystichum munitum (Western swordfern) | NPIN."Lady Bird Johnson Wildflower Center - The University of Texas at Austin. N.p., n.d. Web. 20 Apr. 2011. . 9. “Polystichum munitum in Flora of North America @ efloras.org." eFloras.org Home. N.p., n.d. Web. 20 Apr. 2011. . 10. "Untitled Page." Protocol Information. N.p., n.d. Web. 20 Apr. 2011. .

Other Sources Consulted (but that contained no pertinent information) (full citations): Protocol Author Jonathan Jue (First and last name): Date Protocol 04/19/2011 Created or Updated (MM/DD/YY):

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