WTU Herbarium Specimen Label Data Generated from the WTU Herbarium Database October 10, 2021 at 2:22 am http://biology.burke.washington.edu/herbarium/collections/search.php

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Label Query: Genus = "Polystichum" Dryopteridaceae Dryopteridaceae Polystichum munitum (Kaulf.) C. Presl Polystichum lonchitis (L.) Roth U.S.A., WASHINGTON, WHATCOM COUNTY: U.S.A., OREGON, WALLOWA COUNTY: Little Beaver Trail, about 1 mile east of Perry Creek; North Cascades Wallowa-Whitman National Forest. Lostine National Park. Canyon, Bowman Trail, where trail first crosses Elev. 2034 ft. a feeder creek. 48° 55.3' N, 121° 8.1' W; UTM Zone 10, 636625.715E, Elev. 6689 ft. 5420205.349N; Source: Field notes, UTM calculated from Lat-Lon. 45° 17.289' N, 117° 24.645' W Dry gneiss talus, adjacent Douglas-fir forest, and adjacent cold Dry, north-facing outcrop; open area, break in stream bank just below talus; southern aspect. Growing on mossy, trees. Granite cliff and talus. Growing in rock shaded rock. Phenology: Fertile. Origin: Native. crevices. Altimeter reading 6450 ft. Phenology: Fertile. Origin: Native. Ben Legler 22A 13 Jul 2002 Sarah Gage 7343 Herbarium: NOCA, NPS accession 632, catalog 22826 with Don Knoke, Dale Blum, Richard Robohm 21 Jul 2002 Dryopteridaceae WTU-360645, WTU-V-021059 Polystichum lonchitis (L.) Roth U.S.A., WASHINGTON, SKAGIT COUNTY: North Cascades National Park. Quarter mile Dryopteridaceae southeast of wetland complex. Fisher Basin. North facing wall. Polystichum lonchitis (L.) Roth 48° 33' 17.85241" N, 120° 50' 39.95755" W; UTM Zone 10, 659064E, 5380009N; Source: U.S.A., WASHINGTON, PIERCE COUNTY: Calc. from UTM, UTM from field notes. Mount Rainier National Park, on northeast side at The Palisades, E Wet, mossy cliffs with small waterfalls. Partial end below cliffs just NW of Upper Palisades Lake. sun. Plants restricted to soil pockets and Elev. 6100 ft. hanging gardens among cracks. Common. 46° 57' 7" N, 121° 35' 45" W; UTM Zone 10, 606901E, 5200692N; Large clumps with roots stretching deep among NAD 27, uncertainty: 9 m., Source: GPS talus. Full sun. Phenology: Fertile. Origin: Steep, east-facing rocky slope below small cliffs, with Lutkea Native. pectinata, Carex spectabilis, Poa sp. Uncommon, in deep shady cliff crevices. Phenology: Fertile. Origin: Native. Steve Hahn 1041 with Walt Lockwood Ben Legler 2230 18 Aug 2004 with Pete Del Zotto 23 Aug 2003 Herbarium: MORA, NPS accession 448, catalog 15103 WTU-357990, WTU-V-021087, NPS accession 656, catalog 24463 Dryopteridaceae Dryopteridaceae Polystichum andersonii M. Hopkins Polystichum braunii (Spenner) Fée U.S.A., WASHINGTON, SNOHOMISH RUSSIAN FEDERATION, SAKHALIN REGION: COUNTY: Sakhalin Island, central part of island; along Cascade Mountains. Along Perry Creek Trail. west side of the Orlovka River circa 25 Elev. 3000-3500 ft. kilometers west of town of Smirnykh and 4 48.063294°, -121.513675°; WGS 84, kilometers southeast of mouth of Ognannaya uncertainty: 10000 m., Source: GeoLocate, River. Georef'd by WTU Staff Elev. 394 ft. Deep wooded slopes. Phenology: Spores. 49° 43' 46.62" N, 142° 31' 27.36" E Origin: Native. Northeast facing slope above river, with mix of Abies-Betula forest, open Rubus-Epilobium J. William Thompson 17483 meadows and small brushy stream gullies. Growing in shade in gully. Phenology: Fertile. 30 Jun 1953 Origin: Native. WTU-173350, WTU-V-016083 Ben Legler 908 26 Jul 2003 WTU-357637 Dryopteridaceae Dryopteridaceae Polystichum andersonii M. Hopkins Polystichum imbricans (D.C. Eaton) D.H. Wagner ssp. imbricans U.S.A., WASHINGTON, WHATCOM COUNTY: Along ruth Creek, 6 miles above Shuksan U.S.A., WASHINGTON, SKAGIT COUNTY: Townsite. Cliffs on south side of Bald Mtn. Elev. 3500 ft. Elev. 1600 ft. 48.894272°, -121.662696°; WGS 84, T33N R6E S9; NAD 27, uncertainty: 805 m., uncertainty: 500 m., Source: GeoLocate, Source: TRS2LL, Georef'd by Ben Legler Georef'd by WTU Staff Origin: Native. Viviparous. Growing under Acer circinatum. Origin: Native. Elroy Burnett s.n. with M. Arnot Carl S. English 2225 with Edith Hardin English 31 Mar 1990 6 Aug 1934 WTU-318194, WTU-V-016185 WTU-315831, WTU-V-016085 Dryopteridaceae Dryopteridaceae Polystichum imbricans (D.C. Eaton) D.H. Wagner ssp. imbricans Polystichum californicum (D.C. Eaton) Diels U.S.A., WASHINGTON, CHELAN COUNTY: U.S.A., WASHINGTON, THURSTON COUNTY: On Rock Mtn Trail 1.5 to 2 miles from Hwy 2. [locality and coordinates withheld; status in Elev. 4400 ft. Washington: G4/S1 (Threatened; WANHP)] T26N R15E S3 NE 1/4; WGS84, uncertainty: 1000 m., Source: South side of cliff rock. Nos. 6 to 8 with normal Georeferenced, Georef'd by WTU Staff spores. Phenology: Spores. Origin: Native. With Ceanothus velutinus, Acer glabrum, and Sorbus scopulina. A few small Pseudotsuga nearby. Origin: Native. Ed Alverson s.n. Elroy Burnett s.n. 20 Jul 1991 21 Mar 1980 WTU-328845 WTU-316893, WTU-V-016113 Dryopteridaceae Polystichum kruckebergii W.H. Wagner Dryopteridaceae U.S.A., WASHINGTON, CLALLAM COUNTY: Polystichum munitum (Kaulf.) C. Presl Obstruction Point, Olympic National Park. 48.26667°, -124.675°; WGS 84, uncertainty: U.S.A., WASHINGTON, SAN JUAN COUNTY: 10000 m., Source: GeoLocate, Georef'd by Mount Constitution, Moran State Park, Orcas WTU Staff Island. Along road to summit ca. 1 mile north of Slate crevices of cliffs. With Aster Mountain Lake turnoff. paucicapitatus, Viola flettii, Oxyria digyna, 48.664472°, -122.81167°; WGS 84, Campanula piperi, and Arnica cordifolia. Also uncertainty: 500 m., Source: GeoLocate, one lone plant of Polystichum lonchitis nearby. Georef'd by WTU Staff Infrequent at this site. n = 82 (normal pairing). Shaded, steep rocky slope, moss covered and Origin: Native. westerly exposure, with Polystichum munitum and Heuchera micrantha. Infrequent perennials. A. R. Kruckeberg 5666 Origin: Native. 1 Aug 1963 Scott R. Atkinson 370 WTU-218786, WTU-V-016148 10 May 1992 WTU-325095, WTU-V-18285 Dryopteridaceae Polystichum lemmonii Underw. U.S.A., WASHINGTON, WHATCOM COUNTY: Twin Sisters Range (west side), ca. 13 miles N of Hamilton. Elev. 4700 ft. 48.70833°, -121.99167°; WGS 84, uncertainty: 10000 m., Source: GeoLocate, Georef'd by WTU Staff On serpentine (ultramafic) soil; dry SW talus slope. Origin: Native. G. W. Douglas 1836 7 Aug 1969 WTU-267329, WTU-V-016222 Dryopteridaceae Dryopteridaceae Polystichum lemmonii Underw. Polystichum lonchitis (L.) Roth U.S.A., WASHINGTON, KITTITAS COUNTY: U.S.A., WASHINGTON, KITTITAS COUNTY: Upper Eldorado Creek, North Fork Teanaway Thorp Mountain, Knox Creek Trail. River area. 47.370833°, -121.203333°; NAD 27, 47.413046°, -120.936314°; WGS 84, uncertainty: 1000 m., Source: Georeferenced, uncertainty: 10000 m., Source: GeoLocate, Georef'd by WTU Staff Georef'd by WTU Staff Origin: Native. In deep shade, on peridotite rock. Origin: Native. Maxine Hagen s.n. A. R. Kruckeberg 5789 10 Jul 1983 6 Jul 1964 WTU-335199, WTU-V-021136 WTU-313557, WTU-V-016216

Dryopteridaceae Dryopteridaceae Polystichum munitum (Kaulf.) C. Presl Polystichum lemmonii Underw. U.S.A., WASHINGTON, PIERCE COUNTY: U.S.A., WASHINGTON, CHELAN COUNTY: Sparsely vegetated till ridge between forks of Approx. 13 miles northwest of Coles Corner. Paradise River at source. Take FS Road 6503 to near its end, then cross Elev. 6000 ft. country to ultramafic rocks. 46.785509°, -121.729663°; WGS 84, Elev. 4950 ft. uncertainty: 10000 m., Source: GeoLocate, T28N R15E S35 SW; WGS84, uncertainty: Georef'd by WTU Staff 1000 m., Source: GeoLocate, Georef'd by Origin: Native. WTU Staff With Adiantum pedatum ssp. Calderi and very Harold W. Smith 2216 abundant Aspidotus densa. Also Juncus sp. And Castilleja elmeri. Origin: Native. 13 Aug 1938 Elroy Burnett 589 WTU-48753, WTU-V-18256 30 Jun 1992 WTU-328731, WTU-V-016209 Dryopteridaceae Polystichum munitum (Kaulf.) C. Presl Dryopteridaceae U.S.A., WASHINGTON, JEFFERSON Polystichum lonchitis (L.) Roth COUNTY: Along Quinalt River, near Graves Creek U.S.A., WASHINGTON, SNOHOMISH Campground, Olympic National Park. COUNTY: 47.573819°, -123.571128°; WGS 84, Perry Creek, near Mt. Pilchuck, south fork of the uncertainty: 5000 m., Source: GeoLocate, Stillaguamish River. Georef'd by WTU Staff Elev. 3000 ft. Origin: Native. 48.063294°, -121.513675°; WGS 84, uncertainty: 10000 m., Source: GeoLocate, H. T. Rogers 818 Georef'd by WTU Staff Talus slopes in Tsuga 23 Jun 1941 heterophylla-Chamaecyparis nootkatensis-Alnus rubra zone. Common. Origin: Native. WTU-97233, WTU-V-18269 Melinda F. Denton 2106 Dryopteridaceae 7 Oct 1972 Polystichum munitum (Kaulf.) C. Presl WTU-258282, WTU-V-021064 U.S.A., WASHINGTON, MASON COUNTY: Schafer State Park. 47.09583°, -123.46389°; WGS 84, uncertainty: 10000 m., Source: GeoLocate, Georef'd by WTU Staff Collected in moist woods. Specimen represents small individuals of species to facilitate mounting. Fronds grow to 2 meters in length. Origin: Native. Phillips B. Freer 87 23 Aug 1949 WTU-127100, WTU-V-18400 Dryopteridaceae Dryopteridaceae Polystichum munitum (Kaulf.) C. Presl Polystichum munitum (Kaulf.) C. Presl U.S.A., WASHINGTON, KITSAP COUNTY: U.S.A., WASHINGTON, SAN JUAN COUNTY: Poulsbo. Yellow Island. Elev. 60 ft. 48.592222°, -123.031944°; WGS 84, uncertainty: 250 m., Source: 47.73611°, -122.64528°; WGS 84, Georeferenced, Georef'd by WTU Staff uncertainty: 10000 m., Source: GeoLocate, Center of island, in Pseudotsuga menziesii woodland. Origin: Native. Georef'd by WTU Staff Deep woods. Origin: Native. Eliza Habegger s.n. 17 Apr 1996 S. R. Wellings 35 WTU-333131-A 14 Jul 1948 Dryopteridaceae WTU-183071, WTU-V-18271 Polystichum munitum (Kaulf.) C. Presl U.S.A., WASHINGTON, SKAMANIA COUNTY: Dryopteridaceae Pumice Plain north of crater breach, Mt. St. Helens. Polystichum munitum (Kaulf.) C. Presl 46.24583°, -122.18333°; WGS 84, uncertainty: 10000 m., Source: GeoLocate, U.S.A., WASHINGTON, CLALLAM COUNTY: Georef'd by WTU Staff Steep west facing slope of Mt. Angeles, Olympic Upland, log pile. Origin: Native. National Park. Halfway to summit, in rocks. 48.26667°, -124.675°; WGS 84, uncertainty: Jon Titus s.n. 10000 m., Source: GeoLocate, Georef'd by WTU Staff 9 Aug 1994 Lone plant. Origin: Native. WTU-334431, WTU-V-18403 A. R. Kruckeberg 5676 1 Aug 1963 Dryopteridaceae WTU-214837, WTU-V-18260 Polystichum munitum (Kaulf.) C. Presl U.S.A., WASHINGTON, CHELAN COUNTY: Dryopteridaceae Wenatchee Lake Quad. 1.5 to 2 miles up Rock Mountain trail. Polystichum munitum (Kaulf.) C. Presl Elev. 4400 ft. T26N R15E S3 NE; WGS84, uncertainty: 500 U.S.A., WASHINGTON, SNOHOMISH m., Source: Georeferenced, Georef'd by WTU COUNTY: Staff Perry Creek, near Mt. Pilchuck, south fork of the Growing with Ceanothus velutinus, Acer Stillaguamish River. glabrum and Sorbus scopulina. A few Elev. 3000 ft. Pseudotsuga menziesii nearby. Origin: Native. 48.063294°, -121.513675°; WGS 84, uncertainty: 10000 m., Source: GeoLocate, Elroy Burnett 290 Georef'd by WTU Staff Talus slopes in Tsuga 12 Nov 1990 heterophylla-Chamaecyparis nootkatensis-Alnus rubra zone. Common. Origin: Native. WTU-339784, WTU-V-18255 Melinda F. Denton 2110 Dryopteridaceae 7 Oct 1972 Polystichum munitum (Kaulf.) C. Presl WTU-258280, WTU-V-18258 U.S.A., WASHINGTON, THURSTON COUNTY: Priest Point Park, Olympia. East of East Bay Dryopteridaceae Drive, southern part of park. Elev. 100-120 ft. Polystichum munitum (Kaulf.) C. Presl 47° 4' N, 122° 53' W; T18N R2W S2 or 11 Last season's sori. Phenology: Fertile. Origin: U.S.A., WASHINGTON, KING COUNTY: Native. Cougar Mountain. 47.54056°, -122.09806°; WGS 84, Sarah Gage G&R 035 uncertainty: 10000 m., Source: GeoLocate, with Sharon Rodman Georef'd by WTU Staff Origin: Native. 22 May 1992 David M. Chapin s.n. WTU-345188, WTU-V-18328 9 Oct 1982 WTU-314230, WTU-V-18329 Dryopteridaceae Dryopteridaceae Polystichum scopulinum (D.C. Eaton) Maxon Polystichum munitum (Kaulf.) C. Presl U.S.A., WASHINGTON, CHELAN COUNTY: U.S.A., WASHINGTON, KING COUNTY: SE facing slopes above Wenatchee River, Cascade Mountains of Western Washington: directly across from Tumwater Camp, along Rattlesnake Ledge trail. Hatchery Creek Road. Elev. 1491 ft. 47.672041°, -120.742415°; WGS84, 47° 25.979' N, 121° 46.79' W; T23N R8E S33 uncertainty: 1000 m., Source: GeoLocate, Open area along trail. Very common on slope. Georef'd by WTU Staff Phenology: Fertile. Origin: Native. In boulder strewn talus of massive metamorphic outcrops. Origin: Native. Sarah Gage 7223 with Cindy Spurgeon, Janet Sailer A. R. Kruckeberg 2804 23 Jun 2001 23 Aug 1951 WTU-350880, WTU-V-18327 WTU-156703, WTU-V-021737

Dryopteridaceae Dryopteridaceae Polystichum munitum (Kaulf.) C. Presl Polystichum munitum (Kaulf.) C. Presl U.S.A., WASHINGTON, KING COUNTY: U.S.A., WASHINGTON, ISLAND COUNTY: Cascade Mountains of Western Washington: Hoypus Point, Deception Pass State Park, Whidbey Island. Cedar River bridge at Barneston. 48.39389°, -122.6475°; WGS 84, uncertainty: 10000 m., Source: Elev. 1000 ft. GeoLocate, Georef'd by WTU Staff 47° 23.33' 33" N, 121° 51.68' 51" W; T22N Roadside. Phenology: Flowers. Origin: Native. R7E S14 Woods, riparian area, and roadside near bridge. Harold W. Smith 1607 7 Aug 1936 1 meter tall. Phenology: Fertile. Origin: Native. WTU-38650 P. F. Zika 16002 with Jeff Walker, David Kiefer, Kari Stices, Bill Winije Dryopteridaceae 26 May 2001 Polystichum setigerum (C. Presl) C. Presl WTU-350667, WTU-V-18323 U.S.A., ALASKA COUNTY: [locality and coordinates withheld; status in Alaska: S3 (Rare; AKNHP)] Mountain slope in coniferous woods. Indefinite Dryopteridaceae loan by Tacoma High School to University of Washington. In pencil on sheet: " Also from Polystichum munitum (Kaulf.) C. Presl Olympus - Isotype." [First number (1) on collection number is hidden under specimen on U.S.A., WASHINGTON, KING COUNTY: sheet]. Phenology: Spores. Origin: Native. Type Cascade Mountains of Western Washington: Specimen: Isotype. Wetlands along road 30; near headwaters of Williams Creek. J. B. Flett 917 Elev. 1000 ft. T22N R7E S2 NE 1/4; NAD 27, uncertainty: 19 Jul 1901 805 m., Source: TRS2LL, Georef'd by Ben Legler WTU-36715, WTU-V-000011 Other plants in vicinity: Thuja plicata, Tsuga heterophylla, Psuedotsuga menziesii, Acer circinatum, Galium sp., Glycyrrhiza sp. Phenology: Fertile. Origin: Native. Dryopteridaceae Alan C. Yen 01-47 Polystichum munitum (Kaulf.) C. Presl 26 May 2001 U.S.A., OREGON, CURRY COUNTY: Siskiyou Mountains: Siskiyou Mountains. WTU-350950, WTU-V-18401 T38S R10W S35 SW; NAD 27, uncertainty: 805 m., Source: TRS2LL, Georef'd by Ben Legler Dry serpentine, Calocedrus, Jeffrey pine woodland with Quercus vaccinifolia, Garrya laxifolia. Phenology: Fertile. Origin: Native. Sasa Stefanovic 98-26 with Alison Colwell, John Myers, Randall Hitchin 26 Jun 1998 WTU-349158, WTU-V-18288 Dryopteridaceae Dryopteridaceae Polystichum munitum (Kaulf.) C. Presl Polystichum imbricans (D.C. Eaton) D.H. Wagner ssp. imbricans U.S.A., WASHINGTON, KING COUNTY: Cascade Mountains of Western Washington: U.S.A., WASHINGTON, WHATCOM COUNTY: Law's Ledge. North Cascades National Park; Thunder Creek Elev. 3400 ft. drainage; the first large rocky bald encountered T22N R8E S10 NE; NAD 27, uncertainty: 805 on the trail to 4th of July Pass starting at the m., Source: TRS2LL, Georef'd by Ben Legler junction with the thunder Creek Trail. A small Old clear cut, rock, gravel. Some firs. way trail goes north onto the bald. Phenology: Fertile. Origin: Native. Elev. 1950 ft. 48° 40' 22.7599" N, 121° 3' 44.95907" W; David Kiefer 01-037 UTM Zone 10, 642641E, 5392697N; Source: with Sharon Brown, Bob Antieau, Rena Calc. from UTM, UTM from field notes. Hamburger Very dry, exposed rock outcrops covered with moss. Uncommon; occurs in large clumps of 23 Jun 2001 20-60 fronds; in full sunlight. Phenology: Fertile. Origin: Native. WTU-351145, WTU-V-18319 Steve Hahn 550 15 Jun 2002 WTU-353797, WTU-V-016187, NPS accession 632, catalog 23076 Dryopteridaceae Polystichum lonchitis (L.) Roth Dryopteridaceae U.S.A., WASHINGTON, CHELAN COUNTY: Polystichum munitum (Kaulf.) C. Presl North Cascades National Park; Pelton Basin east of Cascade Pass; hillside opposite camp. U.S.A., WASHINGTON, WHATCOM COUNTY: Elev. 4670 ft. North Cascades National Park; Stetattle Creek 48° 27.6' N, 121° 2.78' W; UTM Zone 10, Trail, at an unnamed tributary to Stetattle Creek. 644435.733E, 5369053.888N; Source: Field Elev. 1329 ft. notes, UTM calculated from Lat-Lon. 48° 43.783' N, 121° 9.342' W; UTM Zone 10, Wet scree slope, coarse rock with some sandy 635626.483E, 5398832.286N; Source: Field soil; scattered Salix, Epilobium latifolium, notes, UTM calculated from Lat-Lon. Athyrium distentifolium; much of slope under Riparian mixed forest, with Acer macrophyllum, snow. Phenology: Flowers. Origin: Native. Thuja plicata, Tsuga heterophylla; understory of Polystichum munitum, Acer circinatum; rock Don Knoke 251 faces at edge of waterfall with Asplenium viride, Adiantum aleuticum, Montia sp., Valeriana 10 Aug 2002 sitchensis, Cryptogramma crispa. Scattered on dry mossy slope. Phenology: Fertile. Origin: WTU-353766, WTU-V-021085, NPS accession Native. 632, catalog 23198 Jeff Walker 3 15 Jun 2002 with Sarah Gage, Nancy Job, Amy Dearborn Dryopteridaceae WTU-354034, WTU-V-18438, NPS accession 632, catalog 23573 Polystichum munitum (Kaulf.) C. Presl U.S.A., WASHINGTON, SKAGIT COUNTY: Dryopteridaceae Lake Cavanaugh Road where it crosses Bear Creek, 2 miles NW of Lake Cavanaugh. Polystichum munitum (Kaulf.) C. Presl Elev. 875 ft. 48° 20' 41" N, 122° 4' 3" W U.S.A., WASHINGTON, WHATCOM COUNTY: Moist woods along creek, with Thuja plicata, North Cascades National Park; Perry Creek Tsuga heterophylla, Alnus rubra, Rubus Shelter, near confluence of Perry Creek and parviflorus, Maianthemum dilatatum. Common Little Beaver Creek. in understory; fronds clustered. Phenology: Elev. 2350 ft. Fertile. Origin: Native. 48° 55.2' N, 121° 8.9' W; UTM Zone 10, 635653.508E, 5419996.201N; Source: Field Ben Legler 284 notes, UTM calculated from Lat-Lon. Shady forest of Thuja plicata and Pseudotsuga 29 Apr 2003 menziesii; thick duff, loose soil. Phenology: Fertile. Origin: Native. WTU-353652, WTU-V-18290 Justin Hohn 26 13 Jul 2002 WTU-353987, WTU-V-18302, NPS accession 632, catalog 23954 Dryopteridaceae Dryopteridaceae Polystichum munitum (Kaulf.) C. Presl Polystichum munitum (Kaulf.) C. Presl U.S.A., WASHINGTON, WHATCOM COUNTY: U.S.A., WASHINGTON, WHATCOM COUNTY: Ross Lake National Recreation Area; Thunder Ross Lake National Recreation Area: Little Creek Trail. Beaver Creek trail. Along trail on east-facing 48° 40.724' N, 121° 4.858' W; UTM Zone 10, slope, within 0.5 miles of trailhead (before 641264.629E, 5393301.111N; Source: Field switchbacks that take trail northwest and finally notes, UTM calculated from Lat-Lon. west). East-facing slope with Tsuga heterophylla, Elev. 1589-2037 ft. Pseudotsuga menziesii, Acer circinatum, 48° 55.014' N, 121° 4.642' W; UTM Zone 10, Pachistima myrsinites, Rubus pedatus, Rosa 640860.997E, 5419780.74N; Source: Field gymnocarpa, Linnaea borealis. Phenology: notes, UTM calculated from Lat-Lon. Fertile. Origin: Native. Furthest point west of lake. Douglas fir forest; along edge of trail. Common; scales rusty-red. Cindy Spurgeon CHS2 Phenology: Fertile. Origin: Native. 15 Jun 2002 Sarah Gage 7270 WTU-353972, WTU-V-18301, NPS accession 13 Jul 2002 632, catalog 23893 WTU-354267, WTU-V-18300, NPS accession 632, catalog 22343 Dryopteridaceae Polystichum lonchitis (L.) Roth Dryopteridaceae U.S.A., WASHINGTON, SKAGIT COUNTY: Polystichum munitum (Kaulf.) C. Presl North Cascades National Park; Boston Basin, 2 miles NW of Cascade Pass; basin between U.S.A., WASHINGTON, WAHKIAKUM glacier and lower climber’s camp. COUNTY: Elev. 5700 ft. 4 miles north of Oneida, along Deep River 48° 29' 48.01041" N, 121° 3' 47.35594" W; Valley Road ca. 1.5 miles north of junction with UTM Zone 10, 643089E, 5373098N; Source: Anderson road. Calc. from UTM, UTM from field notes. Elev. 50 ft. Rocky stream-filled basin; with Salix, Carex, 46° 22' 43.5" N, 123° 40' 8.9" W Phyllodoce. Growing in patches of forest. Wet deciduous forest along small stream, Phenology: Fertile. Origin: Native. dominated by Alnus rubra, Rubus spectabilis, Lysichiton americanum. Fronds clustered from Gerald Schneeweiss 72 rootstalk. Phenology: Fertile. Origin: Native. with Hanna Weiss, Ben Legler Ben Legler 539 10 Aug 2002 1 Jun 2003 WTU-353953, WTU-V-021128, NPS accession 632, catalog 23811 Herbarium: WTU, WTU-V-18305 Dryopteridaceae Dryopteridaceae Polystichum munitum (Kaulf.) C. Presl Polystichum imbricans (D.C. Eaton) D.H. Wagner ssp. imbricans U.S.A., WASHINGTON, ISLAND COUNTY: Whidbey Island, Ebey's Landing; Hill Road c. 1 U.S.A., WASHINGTON, WHATCOM COUNTY: km South of main parking lot, just below summit Ross Lake National Recreation Area: Little of road. Bluff to west of road. Beaver Creek trail. Within 0.5 miles of trailhead Elev. 171 ft. (before switchbacks that take trail northwest and 48° 11' 17" N, 122° 42' 5" W; NAD 27 finally west). Steep bluff dominated by native and exotic Elev. 2035 ft. grasses with Lupinus, Eriophyllum, and Achillea; 48° 55.014' N, 121° 4.642' W; UTM Zone 10, backed by conifer woods. In woods behind bluff. 640860.997E, 5419780.74N; Source: Field Phenology: Fruits. Origin: Native. notes, UTM calculated from Lat-Lon. Furthest point west of lake. South-facing, rocky Peter W. Dunwiddie PWD 65 slope along trail. Growing in tufts out of rock outcrop. Phenology: Fertile. Origin: Native. 30 Apr 2004 Sarah Gage 7282 WTU-355516, WTU-V-18304 13 Jul 2002 WTU-354294, WTU-V-016192, NPS accession 632, catalog 22325 Dryopteridaceae Dryopteridaceae Polystichum braunii (Spenner) Fée Polystichum lonchitis (L.) Roth RUSSIAN FEDERATION, SAKHALIN REGION: U.S.A., WASHINGTON, CHELAN COUNTY: Sakhalin Island; on southern part of island along Okanogan National Forest. Lake Ann Trail, 1 the Nayba River circa 5 kilometers km. north of lake. north-northwest of town of Bykov. Elev. 5477 ft. Elev. 98 ft. 48.5105°, -120.746283°; WGS 84, 47° 59' 54.48" N, 142° 53' 56.52" E uncertainty: 500 m., Source: GeoLocate, Ecotone between uncut field along river and Georef'd by WTU Staff dense Abies Sachalinensis forest on slope Trailside. Forest edge and sloping subalpine above. Growing in shade of Alnus in moist soil. meadow with occasional pockets of talus. Phenology: Fertile. Origin: Native. Clump forming; growing underneath boulders. Phenology: Spores. Origin: Native. Ben Legler 681 David Tank 03-57 17 Jul 2003 with Jessie Johanson, Cindy Spurgeon, Marianne Powell, David Giblin, Kara Ardern, WTU-357049 Suzanne Bagshaw 27 Jul 2003 Dryopteridaceae WTU-362528, WTU-V-021055 Polystichum munitum (Kaulf.) C. Presl U.S.A., WASHINGTON, PACIFIC COUNTY: Dryopteridaceae The Nature Conservancy's Ellsworth Creek Preserve. Old logging road 0.5 km north of first Polystichum andersonii M. Hopkins gate on main road, west side of estuary. Elev. 70 ft. U.S.A., WASHINGTON, GRAYS HARBOR 46° 24.424' N, 123° 53.376' W COUNTY: Edge of clearcut and riparian strip; Alnus and On mountain north alder avalanche slope, Tsuga overstory; Rubus and Sambucus shrub USFS Road 2375, west fork Humptulips River, layer. Trillium along main road. Abundant; new headwaters, 6 miles above Campbell Grove fronds emerging. Phenology: Fertile. Origin: Campground. Native. 47.22956°, -123.956466°; WGS 84, uncertainty: 10000 m., Source: GeoLocate, Jessie Johanson 03-34 Georef'd by WTU Staff Phenology: Spores. Origin: Native. 31 May 2003 A. R. Kruckeberg s.n. WTU-357491, WTU-V-18325 with R. L. Taylor 30 Jun 1978 Dryopteridaceae WTU-280831, WTU-V-016086 Polystichum munitum (Kaulf.) C. Presl Dryopteridaceae U.S.A., WASHINGTON, PACIFIC COUNTY: The Nature Conservancy's Ellsworth Creek Polystichum andersonii M. Hopkins Preserve. Waypoint #36. Along trail in forest tract just above logging spur road. CANADA, BRITISH COLUMBIA: Elev. 161 ft. Queen Charlotte Islands. Moresby Island. About 46° 23.68' N, 123° 53.054' W 3 miles west of head of Cumshewa Inlet below Cedar/spruce dominants with Gaultheria, north face of the Mount Moresby. Vaccinium, and Polystichum. Phenology: Fertile. Elev. 800 ft. Origin: Native. 53.029103°, -132.083496°; WGS 84, uncertainty: 1500 m., Source: Digital Map, Richard G. Olmstead 2003-34 Georef'd by Joseph Stormer In Alder thickets at base of steep rocky-gravelly, 31 May 2003 north-facing runnel. Common in thickets. Phenology: Spores. Origin: Native. WTU-357517, WTU-V-18303 J. A. Calder 36397 with R. L. Taylor 31 Jul 1964 WTU-242452, WTU-V-016076 Dryopteridaceae Dryopteridaceae Polystichum andersonii M. Hopkins Polystichum andersonii M. Hopkins U.S.A., WASHINGTON, SNOHOMISH U.S.A., WASHINGTON, PIERCE COUNTY: COUNTY: Mount Rainier. By trail to Van Trump Park. Cascade Range. One mile east Monte Cristo. Elev. 3000 ft. Elev. 4000 ft. 46.79806°, -121.77806°; WGS 84, 47.98583°, -121.39278°; WGS 84, uncertainty: 10000 m., Source: GeoLocate, uncertainty: 10000 m., Source: GeoLocate, Georef'd by WTU Staff Georef'd by WTU Staff Rocky alder slopes. Phenology: Spores. Origin: Phenology: Spores. Origin: Native. Native. T. Darling Jr. s.n. J. William Thompson 12555 11 Aug 1962 14 Aug 1935 WTU-234934, WTU-V-016060 WTU-150JWT, WTU-V-016069

Dryopteridaceae Dryopteridaceae Polystichum andersonii M. Hopkins Polystichum andersonii M. Hopkins U.S.A., WASHINGTON, SNOHOMISH COUNTY: U.S.A., IDAHO COUNTY: Mt. Dickerman; talus above Perry Creek. Roundtop. Elev. 3000 ft. Elev. 6000 ft. 48.063294°, -121.513675°; WGS 84, T42N R3E S3; NAD 27, uncertainty: 805 m., uncertainty: 10000 m., Source: GeoLocate, Source: TRS2LL, Georef'd by Ben Legler Georef'd by WTU Staff Found in rock crevasses. Phenology: Spores. Shaded mossy talus; with P. munitum and Origin: Native. hybrid. Locally abundant. N=82. Phenology: Spores. Origin: Native. R. C. Stillinger 623 W. H. Wagner 63116-7 3 Aug 1941 with A. R. Kruckeberg WTU-299380, WTU-V-016088 28 Jun 1963 WTU-213555, WTU-V-016073 Dryopteridaceae Dryopteridaceae Polystichum andersonii M. Hopkins Polystichum andersonii M. Hopkins U.S.A., WASHINGTON, SNOHOMISH COUNTY: U.S.A., WASHINGTON, SNOHOMISH West slope of Cascade Mountains, Mount COUNTY: Baker-Snoqualmie National Forest. Northwest Mt. Dickerman; talus above Perry Creek. slope of Dickerman Mountain. Perry Creek Elev. 3000 ft. drainage, c. [circa] 0.5 miles above trailhead 48.063294°, -121.513675°; WGS 84, and northeast end of Forest Service Road 4063. uncertainty: 10000 m., Source: GeoLocate, Elev. 2197 ft. Georef'd by WTU Staff 48° 4.4' N, 121° 29.5' W; T30N R10E S23 Shaded mossy talus; with P. munitum and Shaded trailside, northwest aspect, steep mossy hybrid. Locally abundant. N=82. Voucher. and rocky slope, with Blechnum spicant, Acer Origin: Native. circinatum, Athyrium filix-femina, Botrychium multifidum, Polystichum munitum. Caespitose; W. H. Wagner 63116-2 mid-green; common. Phenology: Spores. Origin: with A. R. Kruckeberg Native. 28 Jun 1963 P. F. Zika 19083 20 Sep 2003 WTU-213556, WTU-V-016057 WTU-354255, WTU-V-016067 Dryopteridaceae Polystichum andersonii M. Hopkins U.S.A., ALASKA COUNTY: About 10 miles north of Juneau. East side of Mendenhall Glacier. About 1/3 mile south of Nugget Creek along East Glacier Trail. In forested area. Phenology: Spores. Origin: Native. Mary Clay Muller 5697 5 Aug 1992 WTU-331493, WTU-V-016087 Dryopteridaceae Dryopteridaceae Polystichum andersonii M. Hopkins Polystichum andersonii M. Hopkins U.S.A., ALASKA COUNTY: U.S.A., WASHINGTON, PIERCE COUNTY: About 10 miles north of Juneau. East side of Mount Rainier. By trail to Van Trump Park. Mendenhall Glacier. About 1/3 mile south of Elev. 3000 ft. Nugget Creek along East Glacier Trail. 46.79806°, -121.77806°; WGS 84, In forested area. Phenology: Spores. Origin: uncertainty: 10000 m., Source: GeoLocate, Native. Georef'd by WTU Staff Rocky alder slopes. Phenology: Spores. Origin: Mary Clay Muller 5696 Native. 5 Aug 1992 J. William Thompson 12555 WTU-331489, WTU-V-016068 14 Aug 1935 WTU-33055, WTU-V-016059 Dryopteridaceae Polystichum andersonii M. Hopkins Dryopteridaceae U.S.A., MONTANA, LINCOLN COUNTY: Polystichum andersonii M. Hopkins Cabinet Mountains. North-facing mixed shrubfield snowchute. [Transect 63]. U.S.A., WASHINGTON, SNOHOMISH Elev. 4400 ft. COUNTY: T28N R31W S18 SE; NAD 27, uncertainty: Perry Creek, near Mt. Pilchuck, south fork of the 805 m., Source: TRS2LL, Georef'd by Ben Stillaguamish River. Legler Elev. 3000 ft. North-facing mixed shrubfield snowchute. Under 48.063294°, -121.513675°; WGS 84, canopy of Alnus 2+ meters high, with uncertainty: 10000 m., Source: GeoLocate, Oplopanax horridum, Aconitum, etc. Phenology: Georef'd by WTU Staff Spores. Origin: Native. Talus slopes in Tsuga heterophylla- Chamaecyparis nootkatensis- Alnus rubra zone. Christopher Clampitt 124 Locally common. Phenology: Spores. Origin: Native. 25 Aug 1985 Melinda F. Denton 2108 WTU-313278, WTU-V-016065 7 Oct 1972 Dryopteridaceae WTU-258281, WTU-V-016075 Polystichum setigerum (C. Presl) C. Presl Dryopteridaceae U.S.A., ALASKA COUNTY: [locality and coordinates withheld; status in Polystichum andersonii M. Hopkins Alaska: S3 (Rare; AKNHP)] Elev. 250 ft. U.S.A., WASHINGTON, SNOHOMISH Herbaceous slopes and rocky places. COUNTY: Phenology: Spores. Origin: Native. Perry Creek from garden of A. R. Kruckeberg. 48.063294°, -121.513675°; WGS 84, Robert M. Hardy 404 uncertainty: 10000 m., Source: GeoLocate, Georef'd by WTU Staff 23 Sep 1945 Garden. Phenology: Spores. Origin: Native. WTU-97295, WTU-V-021040 A. R. Kruckeberg K-1 24 Jun 1963 Dryopteridaceae WTU-213566, WTU-V-016080 Polystichum andersonii M. Hopkins U.S.A., WASHINGTON, LEWIS COUNTY: Headwaters Cold Creek, Ladd Rock, in the "Little Rockies" (Deschutes Mountains), between Elbe and Morton. 46.55861°, -122.27389°; WGS 84, uncertainty: 10000 m., Source: GeoLocate, Georef'd by WTU Staff Phenology: Spores. Origin: Native. A. R. Kruckeberg s.n. 4 Jul 1978 WTU-280832, WTU-V-016090 Dryopteridaceae Dryopteridaceae Polystichum andersonii M. Hopkins Polystichum andersonii M. Hopkins U.S.A., WASHINGTON, SNOHOMISH U.S.A., WASHINGTON, GRAYS HARBOR COUNTY: COUNTY: Cascade Mountains. Mount Dickerman. Olympic Mountains. Trail to Mount Colonel Bob. Elev. 3000 ft. Elev. 4000 ft. 48.06829°, -121.47068°; WGS 84, 47.48889°, -123.74556°; WGS 84, uncertainty: 3000 m., Source: GeoLocate, uncertainty: 10000 m., Source: GeoLocate, Georef'd by WTU Staff Georef'd by WTU Staff Wooded talus slopes of mountain. Phenology: Wooded slopes. Phenology: Spores. Origin: Spores. Origin: Native. Native. J. William Thompson 8779 J. William Thompson 9964 17 Jul 1932 22 Aug 1933 WTU-10216JWT, WTU-V-016062 WTU-10219JWT, WTU-V-016053

Dryopteridaceae Dryopteridaceae Polystichum andersonii M. Hopkins Polystichum andersonii M. Hopkins U.S.A., WASHINGTON, LEWIS COUNTY: U.S.A., WASHINGTON, GRAYS HARBOR Stevens Canyon, Mt. Rainier National Park. COUNTY: 46.71583°, -121.60139°; WGS 84, By trail to Mount Colonel Bob Lookout, Olympic uncertainty: 10000 m., Source: GeoLocate, Mountains. Georef'd by WTU Staff Elev. 4000 ft. Rocky slope. Phenology: Spores. Origin: Native. 47.487222°, -123.745556°; NAD 27, uncertainty: 1000 m., Source: Georeferenced, F. A. Warren 1790 Georef'd by WTU Staff Rocky talus slopes. Phenology: Spores. Origin: 31 Jul 1932 Native. WTU-25207, WTU-V-016078 J. William Thompson 7276 9 Jul 1931 Dryopteridaceae WTU-10214JWT, WTU-V-016056 Polystichum andersonii M. Hopkins Dryopteridaceae U.S.A., MONTANA, RAVALLI COUNTY: Bitterroot Mountains. East slope of the ridge Polystichum andersonii M. Hopkins between South Kootenail and the Heavenly Twins. U.S.A., WASHINGTON, SNOHOMISH Elev. 6400 ft. COUNTY: East slope of the ridge. Extensive colonies (with Cascade Mountains. Mount Dickerman. Dryopteris felix-mas and D. austriaca) beneath Elev. 3000 ft. Alnus sinuata. Origin: Native. 48.06829°, -121.47068°; WGS 84, uncertainty: 3000 m., Source: GeoLocate, Klaus H. Lackschewitz 3264 Georef'd by WTU Staff Wooded talus slopes. Phenology: Spores. 15 Aug 1971 Origin: Native. WTU-257671, WTU-V-016061 J. William Thompson 8779 17 Jul 1932 Dryopteridaceae WTU-10220JWT, WTU-V-016055 Polystichum andersonii M. Hopkins Dryopteridaceae U.S.A., WASHINGTON, PIERCE COUNTY: Mount Rainier. Old Moraines. Carbon Glacier. Polystichum andersonii M. Hopkins Elev. 4000 ft. 46.940083°, -121.789677°; WGS 84, U.S.A., WASHINGTON, CLALLAM COUNTY: uncertainty: 2030 m., Source: Digital Map, Olympic Mountains. Elwha Canyon. Georef'd by Joseph Stormer Elev. 4000 ft. In shade of Alders. Indefinite loan by Tacoma 48.06083°, -123.59306°; Source: Inferred High School to University of Washington. from duplicate specimen, Georef'd by Ed Phenology: Spores. Origin: Native. Alverson On wet slope. Phenology: Spores. Origin: J. B. Flett 2020 Native. 28 Aug 1902 J. B. Flett s.n. WTU-37539, WTU-V-016064 9 Aug 1907 WTU-29781, WTU-V-016046 Dryopteridaceae Dryopteridaceae Polystichum andersonii M. Hopkins Polystichum andersonii M. Hopkins U.S.A., WASHINGTON, GRAYS HARBOR U.S.A., WASHINGTON, SNOHOMISH COUNTY: COUNTY: By trail to Mount Colonel Bob Lookout, Olympic Cascade Mountains. By Perry Creek Trail. Mountains. Elev. 3000 ft. Elev. 4000 ft. 48.063294°, -121.513675°; WGS 84, 47.487222°, -123.745556°; NAD 27, uncertainty: 10000 m., Source: GeoLocate, uncertainty: 1000 m., Source: Georeferenced, Georef'd by WTU Staff Georef'd by WTU Staff Deep woods. Phenology: Spores. Origin: Native. Rocky talus slopes. Phenology: Spores. Origin: Native. J. William Thompson 14745 J. William Thompson 7276 15 Jun 1940 9 Jul 1931 WTU-291584, WTU-V-016039 WTU-10213JWT, WTU-V-016091 Dryopteridaceae Dryopteridaceae Polystichum andersonii M. Hopkins Polystichum andersonii M. Hopkins U.S.A., IDAHO, SHOSHONE COUNTY: Lost Lake. U.S.A., ALASKA COUNTY: 47.07361°, -115.95694°; WGS 84, Left side of Mendenhall glacier, Juneau. Trail uncertainty: 10000 m., Source: GeoLocate, around Mount McGinnis. Georef'd by WTU Staff In alder and willow. Abundant. Phenology: Steep talus at head of cirque, with Alnus Spores. Origin: Native. sinuata, Mertensia, Senecio tri. Spruce-fir vegetation zone; seral open scrub. Phenology: Maxcine Williams 2103 Spores. Origin: Native. 6 Nov 1968 R. F. Daubenmire 45179 WTU-240058, WTU-V-016077 10 Sep 1945 WTU-271406, WTU-V-016070

Dryopteridaceae Dryopteridaceae Polystichum andersonii M. Hopkins Polystichum andersonii M. Hopkins U.S.A., ALASKA COUNTY: Tongass N.F. Mainland. About 28 miles N of U.S.A., ALASKA COUNTY: Juneau. About 4 miles up the Eagle Glacier trail. Tongass National Forest. West side of Baranof Growing in the forest near the trail. Phenology: Island at Sandy Cove about 4 miles south of Spores. Origin: Native. Sitka. Elev. 0 ft. Mary Clay Muller 4066 Slightly vegetated river bar near stream mouth. Several small Polystichum andersonii seen next 21 Aug 1980 to large plant. Polystichum braunii also present. Phenology: Spores. Origin: Native. WTU-289237, WTU-V-016092 Mary Clay Muller 4705 26 Nov 1981 Dryopteridaceae WTU-282129, WTU-V-016081 Polystichum andersonii M. Hopkins U.S.A., ALASKA COUNTY: Dryopteridaceae Juneau area. About 10 miles north of town at Mendenhall Glacier. Collected along East Polystichum andersonii M. Hopkins Glacier Trail. About 0.25 mile south of Nugget Creek. U.S.A., WASHINGTON, PIERCE COUNTY: Elev. 200 ft. Mount Rainier National Park. In Ipsut Pass. Growing with herbaceous plants in an open area 46.95083°, -121.86639°; WGS 84, near a rock cliff. Phenology: Spores. Origin: uncertainty: 10000 m., Source: GeoLocate, Native. Georef'd by WTU Staff Rocky slopes. Phenology: Spores. Origin: Mary Clay Muller 4312 Native. 9 Jul 1981 F. A. Warren s.n. WTU-282680, WTU-V-016089 25 Jul 1930 WTU-10215JWT, WTU-V-016048 Dryopteridaceae Dryopteridaceae Polystichum andersonii M. Hopkins Polystichum andersonii M. Hopkins U.S.A., WASHINGTON, SNOHOMISH U.S.A., IDAHO, SHOSHONE COUNTY: COUNTY: Freezout. Fourteen miles east of Clarkia. Cascade Mountains. By Perry Creek Trail. 47.01083°, -116.25194°; WGS 84, Elev. 3000 ft. uncertainty: 10000 m., Source: GeoLocate, 48.063294°, -121.513675°; WGS 84, Georef'd by WTU Staff uncertainty: 10000 m., Source: GeoLocate, Open granite slope, glacial cirque. Phenology: Georef'd by WTU Staff Spores. Origin: Native. Deep woods. Phenology: Spores. Origin: Native. William H. Baker 16233 J. William Thompson 14745 21 Jul 1961 15 Jun 1940 WTU-261402, WTU-V-016066 WTU-21920JWT, WTU-V-016082

Dryopteridaceae Dryopteridaceae Polystichum andersonii M. Hopkins Polystichum andersonii M. Hopkins U.S.A., ALASKA COUNTY: U.S.A., ALASKA COUNTY: Juneau area. About 10 miles north of town at Juneau area. About 10 miles north of town at Mendenhall Glacier. Collected along East Mendenhall Glacier. Collected along East Glacier Trail. Glacier Trail. Collected just north of AJ Creek. Elev. 200 ft. Elev. 200 ft. West-southwest facing open brushy area. Open brushy area. Phenology: Spores. Origin: Collected about 100 north of AJ Creek. Native. Phenology: Spores. Origin: Native. Mary Clay Muller 4320 Mary Clay Muller 4319 9 Jul 1981 9 Jul 1981 WTU-282620, WTU-V-016084 WTU-282621, WTU-V-016079

Dryopteridaceae Dryopteridaceae Polystichum andersonii M. Hopkins Polystichum andersonii M. Hopkins U.S.A., ALASKA COUNTY: U.S.A., OREGON, HOOD RIVER COUNTY: Tongass National Forest. Near Mendenhall Mount Hood region. Lost Lake. Glacier, about 10 miles northwest of Juneau. Elev. 3000 ft. Collected along East Glacier Trail. 45.489391°, -121.831552°; WGS 84, Open area; this area was recently deglaciated. uncertainty: 1777 m., Source: Digital Map, Near guardrail overlooking ice. Growing with Georef'd by Sara Legler Polystichum lonchitis. Common. Phenology: Timbered rock slides at lake. First Oregon Spores. Origin: Native. record. Phenology: Spores. Origin: Native. Mary Clay Muller 4185 J. William Thompson 11192 30 Oct 1980 31 Jul 1934 WTU-289205, WTU-V-016072 WTU-25533, WTU-V-016050

Dryopteridaceae Dryopteridaceae Polystichum braunii (Spenner) Fée Polystichum andersonii M. Hopkins U.S.A., ALASKA, SITKA COUNTY: U.S.A., OREGON, HOOD RIVER COUNTY: Chichagof Island, Klag Bay, western shore Mount Hood region. Lost Lake. about 1/4 mile from mine. Elev. 3000 ft. Beach forest ecotene, hemlock and spruce 45.489391°, -121.831552°; WGS 84, forest on moss. Locally common. Origin: Native. uncertainty: 1777 m., Source: Digital Map, Georef'd by Sara Legler Mary Clay Muller 5431 Timbered rock slides at lake. Phenology: Spores. Origin: Native. 6 Jun 1984 J. William Thompson 11192 WTU-330851, WTU-V-016114 31 Jul 1934 WTU-140JWT, WTU-V-016047 Dryopteridaceae Dryopteridaceae Polystichum setigerum (C. Presl) C. Presl Polystichum imbricans (D.C. Eaton) D.H. Wagner ssp. imbricans U.S.A., ALASKA, PRINCE WALES KETCHIKAN COUNTY: U.S.A., WASHINGTON, CHELAN COUNTY: [locality and coordinates withheld; status in Wenatchee lake Quadrangle; 1.5 to 2 miles up Alaska: S3 (Rare; AKNHP)] Rock Mountian Trail. Cedar-hemlock forest, bank of dry side channel Elev. 4400 ft. of cobbles. Origin: Native. T26N R15E S3 NE 1/4; WGS84, uncertainty: 10000 m., Source: GeoLocate, Georef'd by M. C. Stensvold 6195 WTU Staff Origin: Native. 11 Aug 1993 E. Burnett 289 WTU-332940, WTU-V-021031 12 Nov 1990 WTU-339785, WTU-V-016190 Dryopteridaceae Polystichum braunii (Spenner) Fée Dryopteridaceae U.S.A., ALASKA, YAKUTAT CITY AND BOROUGH COUNTY: Polystichum setigerum (C. Presl) C. Presl 21 miles east of the town, 1 mile north of Forest Highway 10, Pike Lakes area, between the large U.S.A., ALASKA, WRANGELL PETERSBURG muskeg north of the road and the Lakes. COUNTY: 59° 29' 45" N, 139° 9' 30" W [locality and coordinates withheld; status in Mixed coniferous forest, well drained area, near Alaska: S3 (Rare; AKNHP)] highwater river channel. Phenology: Fertile. Rock crevice in stream bank. Origin: Native. Origin: Native. W. J. Eyerdam 5451 M. C. Stensvold 5606 20 Aug 1947 16 Jul 1991 WTU-293245, WTU-V-021032 WTU-333161, WTU-V-016094

Dryopteridaceae Dryopteridaceae Polystichum braunii (Spenner) Fée Polystichum braunii (Spenner) Fée U.S.A., ALASKA, KENAI PENINSULA COUNTY: U.S.A., ALASKA COUNTY: Base of Mount Marathon, Seward. Tongass National Forest. Mainland. About 28 60° 6' N, 149° 27' W miles north of Juneau. About 4 miles up the Moist lush wooded area, base of mountain. Eagle Glacier trail. Rare, only 2 noted. Origin: Native. Forest near trail. Origin: Native. J. A. Calder 6630 Mary Clay Muller 4080 10 Aug 1951 21 Aug 1980 WTU-173797, WTU-V-016097 WTU-289243, WTU-V-016110

Dryopteridaceae Dryopteridaceae Polystichum braunii (Spenner) Fée Polystichum braunii (Spenner) Fée U.S.A., ALASKA COUNTY: Mountain near Juneau. CANADA, BRITISH COLUMBIA: 58.339447°, -134.342818°; uncertainty: 50000 Head of McClinton Bay, Masset Inlet, Graham m. Island, along McClinton Creek and below cliffs. Origin: Native. Elev. 1700 ft. Coniferous woods. Common. Origin: Native. Emma A. Shumway s.n. J. A. Calder 21595 6 Aug 1891 with D. B. O. Savile, R. L. Taylor WTU-90311, WTU-V-016104 18 Jun 1957 WTU-242455, WTU-V-016109 Dryopteridaceae Dryopteridaceae Polystichum setigerum (C. Presl) C. Presl Polystichum braunii (Spenner) Fée U.S.A., ALASKA, SKAGWAY YAKUTAT CANADA, BRITISH COLUMBIA: ANGOON COUNTY: Along Yakoun River near junction with Ghost [locality and coordinates withheld; status in Creek, 12 miles south of Juskatla, Graham Alaska: S3 (Rare; AKNHP)] Island. Just above tide line. Phenology: Fertile. Origin: 53.433249°, -132.283643°; WGS 84, Native. uncertainty: 2500 m., Source: Digital Map, Georef'd by Joseph Stormer Mary Clay Muller 3300 Grassy banks of river, open coniferous forest. Common. Origin: Native. 25 Aug 1979 J. A. Calder 35500 WTU-281759, WTU-V-021041 with R. L. Taylor 27 Jun 1964 Dryopteridaceae WTU-251436, WTU-V-016106 Polystichum imbricans (D.C. Eaton) D.H. Wagner ssp. imbricans Dryopteridaceae U.S.A., OREGON, JOSEPHINE COUNTY: Siskiyou National Forest, 20-23 miles west and Polystichum imbricans (D.C. Eaton) D.H. Wagner ssp. north of Selma on Illinois River Road (3504) or imbricans 8-11 miles north of intersection of 3504 and 374 on 3504. U.S.A., OREGON, JOSEPHINE COUNTY: Elev. 1180 ft. Along Big Tree Trail at Cave Creek. Oregon 42.35201°, -123.779622°; WGS 84, National Mounument. uncertainty: 4000 m., Source: Digital Map, Georef'd by Joseph Stormer William H. Baker 555 Origin: Native. 2 Aug 1949 Melinda F. Denton 2466 WTU-298181, WTU-V-016173 21 Jun 1973 WTU-259372, WTU-V-016182

Dryopteridaceae Dryopteridaceae Polystichum imbricans (D.C. Eaton) D.H. Wagner ssp. Polystichum imbricans (D.C. Eaton) D.H. Wagner ssp. imbricans imbricans U.S.A., OREGON, KLAMATH COUNTY: U.S.A., OREGON, JOSEPHINE COUNTY: Southwest corner, north wall, redblanket Along the Merline Galice road 2.8 miles canyon, Crater Lake National Park. upstream from the junction with lower Graves creek road. W. H. Baker 7104 Elev. 751 ft. T34S R8W S13; NAD 27, uncertainty: 805 m., 29 Jul 1950 Source: TRS2LL, Georef'd by Ben Legler WTU-299575, WTU-V-016189 Richard R. Halse 4977 8 Aug 1995 WTU-333632, WTU-V-016172

Dryopteridaceae Polystichum braunii (Spenner) Fée U.S.A., ALASKA, SOUTHEAST FAIRBANKS COUNTY: Tongass National Forest, Stikine River at Shakes Hot Springs, north side of Ketili Slough, 30 miles northeast of Wrangell. Forest. Origin: Native. Mary Clay Muller 3777 4 Jul 1980 WTU-289133, WTU-V-016108 Dryopteridaceae Dryopteridaceae Polystichum imbricans (D.C. Eaton) D.H. Wagner ssp. Polystichum braunii (Spenner) Fée imbricans U.S.A., ALASKA, SKAGWAY YAKUTAT U.S.A., OREGON, JOSEPHINE COUNTY: ANGOON COUNTY: Along Forest Road 4105, 2.8 miles from junction Southeast, Chichagof Island, Icy Strait, Mud with Forest Road 4103. Siskiyou National Bay, Slope 0-5% . Forest. Elev. 260 ft. Elev. 2400 ft. T43S R59E S15 1/4NE 42° 18' 55" N, 123° 42' 58" W; T37S R9W Flood plain, sandy undeveloped soil, growing S35 or 36 with Alnus rubra, Ribes bracteosum, Picea Origin: Native. sitchensis, Tiarella trifoliata. Infrequent. Origin: Native. Richard Robohm 67 Kitty LaBounty 207 26 Jun 1998 26 Aug 1981 WTU-345749, WTU-V-016181 WTU-282148, WTU-V-016105 Dryopteridaceae Dryopteridaceae Polystichum imbricans (D.C. Eaton) D.H. Wagner ssp. Polystichum imbricans (D.C. Eaton) D.H. Wagner ssp. imbricans imbricans U.S.A., OREGON, JOSEPHINE COUNTY: CANADA, BRITISH COLUMBIA: No Name Trail. Little Saanich Mountain, V.I. Origin: Native. Phenology: Fertile. Origin: Native. William H. Baker 456 J. P. Anderson s.n. with George C. Ruhle 20 Oct 1917 13 Jul 1949 WTU-123981, WTU-V-016188 WTU-298180, WTU-V-016176

Dryopteridaceae Dryopteridaceae Polystichum setigerum (C. Presl) C. Presl Polystichum imbricans (D.C. Eaton) D.H. Wagner ssp. imbricans U.S.A., ALASKA, WRANGELL PETERSBURG COUNTY: U.S.A., WASHINGTON, CHELAN COUNTY: [locality and coordinates withheld; status in On cliffs along trail on noth side of lake Chelan Alaska: S3 (Rare; AKNHP)] about one mile south of stehekin. Shady spruce forest. Origin: Native. 47.84111°, -120.01528°; WGS84, uncertainty: 10000 m., Source: GeoLocate, Georef'd by W. J. Eyerdam 5431 WTU Staff 15 Aug 1947 B. O. Mulligan s.n. WTU-293244, WTU-V-021035 12 Jul 1969 WTU-260104, WTU-V-016126

Dryopteridaceae Dryopteridaceae Polystichum imbricans (D.C. Eaton) D.H. Wagner ssp. imbricans Polystichum braunii (Spenner) Fée U.S.A., WASHINGTON, CHELAN COUNTY: U.S.A., ALASKA, SKAGWAY YAKUTAT Rock Mountain Trail 1.5 to 2 miles from ANGOON COUNTY: Highway 2. Chichagof Island, Icy Strait, Gallagher Creek Elev. 4400 ft. Drainage, aspect south/southeast slope 60%. T26N R15E S3 NE 1/4; WGS84, uncertainty: Elev. 1300 ft. 500 m., Source: Georeferenced, Georef'd by T43S R60E S1 WTU Staff Ribes bracteosum thicket on an old rock slide, Origin: Native. growing with Oplapanax horridum, Rubus spectabilis, Galium kamtschaticum. Uncommon. E. Burnett s.n. Origin: Native. 20 Jul 1991 Kitty LaBounty 193 WTU-321914, WTU-V-030981 17 Aug 1981 WTU-282210, WTU-V-016099 Dryopteridaceae Dryopteridaceae Polystichum imbricans (D.C. Eaton) D.H. Wagner ssp. Polystichum imbricans (D.C. Eaton) D.H. Wagner ssp. imbricans imbricans U.S.A., WASHINGTON, CHELAN COUNTY: U.S.A., WASHINGTON, SKAMANIA COUNTY: Tumwater canyon. Dry rock mountain slopes, Wind river. Elev. 1350 ft. 45.79707°, -121.91456°; WGS 84, 47.58472°, -120.67722°; WGS84, uncertainty: uncertainty: 3000 m., Source: GeoLocate, 10000 m., Source: GeoLocate, Georef'd by Georef'd by WTU Staff WTU Staff Origin: Native. I. C. Otis 1052 J. B. Flett s.n. 15 Sep 1920 8 Aug 1899 WTU-28930, WTU-V-016123 WTU-37538, WTU-V-016127

Dryopteridaceae Dryopteridaceae Polystichum imbricans (D.C. Eaton) D.H. Wagner ssp. Polystichum braunii (Spenner) Fée imbricans U.S.A., ALASKA, JUNEAU COUNTY: U.S.A., WASHINGTON, JEFFERSON Juneau. COUNTY: 58.301944°, -134.419722°; uncertainty: 10000 Olympic Mountains. m. Georef'd by Ed Alverson, locality too vague to Woods. Another number written at top of label georeference #5591. Origin: Native. Origin: Native. J. P. Anderson 6159 A. D. E. Elmer 2815 8 Aug 1940 Jul 1900 WTU-292961, WTU-V-016101 WTU-8114JWT, WTU-V-016125

Dryopteridaceae Dryopteridaceae Polystichum imbricans (D.C. Eaton) D.H. Wagner ssp. Polystichum imbricans (D.C. Eaton) D.H. Wagner ssp. imbricans imbricans U.S.A., WASHINGTON, CHELAN COUNTY: U.S.A., WASHINGTON, CLALLAM COUNTY: Power house between Leavenworth and Steep rocky cliff in shade, Lake crescent. Tumwater. 48.09472°, -123.80583°; WGS 84, Elev. 1350 ft. uncertainty: 10000 m., Source: GeoLocate, 47.59639°, -120.66028°; WGS84, uncertainty: Georef'd by WTU Staff 10000 m., Source: GeoLocate, Georef'd by Origin: Native. WTU Staff Phenology: Fertile. Origin: Native. Carl S. English 2209 I. C. Otis 1058 4 Jul 1932 28 Sep 1920 WTU-34943, WTU-V-016128 WTU-2975JWT, WTU-V-016184

Dryopteridaceae Dryopteridaceae Polystichum braunii (Spenner) Fée Polystichum braunii (Spenner) Fée U.S.A., ALASKA, SKAGWAY YAKUTAT ANGOON COUNTY: U.S.A., ALASKA, JUNEAU COUNTY: Southeast, Chichagof Island, Icy Strait, Point Side of Mount Juneau. Adolphus area, aspect east slope 25%, bottom Origin: Native. of a large, shrubby avalanche track. Elev. 1460 ft. Frank Beals s.n. T42S R59E S6 In thick cover of Ribes bracteosum, Rubus 13 Aug 1943 spectabilis. Locally common. Origin: Native. WTU-110860, WTU-V-016098 Kitty LaBounty 162 12 Aug 1981 WTU-282097, WTU-V-016093 Dryopteridaceae Dryopteridaceae Polystichum imbricans (D.C. Eaton) D.H. Wagner ssp. Polystichum imbricans (D.C. Eaton) D.H. Wagner ssp. imbricans imbricans U.S.A., OREGON, JOSEPHINE COUNTY: U.S.A., OREGON, DOUGLAS COUNTY: Ridge between north and south forks of cave Cow Creek near Nichols Station. creek. 42.888727°, -123.538683°; WGS 84, Origin: Native. uncertainty: 1500 m., Source: GeoLocate, Georef'd by John Haskins Kent Beattie 5384 Chiefly on rocks. Phenology: Spores. Origin: Native. 28 Aug 1922 L. F. Ward s.n. WTU-108081, WTU-V-016186 17 Sep 1899 WTU-239518, WTU-V-016122

Dryopteridaceae Dryopteridaceae Polystichum braunii (Spenner) Fée Polystichum imbricans (D.C. Eaton) D.H. Wagner ssp. U.S.A., ALASKA, WRANGELL PETERSBURG imbricans COUNTY: Washington Bay, Kuiu Island. U.S.A., OREGON, LAKE COUNTY: 56.71885°, -134.364815°; WGS 84, Dry cleft west slope of crane mountains. uncertainty: 2000 m., Source: Digital Map, Elev. 7000 ft. Georef'd by Joseph Stormer Origin: Native. Rock crevices in stream bank. Origin: Native. M. E. Peck 15565a W. J. Eyerdam 5451 11 Jul 1927 20 Aug 1947 WTU-8115JWT, WTU-V-016180 WTU-118032, WTU-V-016102

Dryopteridaceae Polystichum braunii (Spenner) Fée Dryopteridaceae U.S.A., ALASKA COUNTY: Polystichum californicum (D.C. Eaton) Diels × Juneau area. About 10 miles north of town at Polystichum munitum (Kaulf.) C. Presl Mendenhall Glacier. Collected along East Glacier Trail. U.S.A., WASHINGTON, THURSTON COUNTY: Elev. 200 ft. Bald Hills Lake. Open area with deciduous shrubs and herbs. 46.82027°, -122.44414°; WGS 84, Common. Origin: Native. uncertainty: 500 m., Source: GeoLocate, Georef'd by WTU Staff Mary Clay Muller 4316 Shady cliff. Sporangia abortive. Phenology: Spores. Origin: Native. 9 Jul 1981 Ed Alverson s.n. WTU-282681, WTU-V-016107 15 Mar 1980 WTU-316897, WTU-V-030989 Dryopteridaceae Polystichum braunii (Spenner) Fée Dryopteridaceae U.S.A., ALASKA COUNTY: Mainland 30 miles north of Juneau, 1 mile south Polystichum imbricans (D.C. Eaton) D.H. Wagner ssp. of the head of Echo Cove, north shore of Cowee imbricans Creek, 1/4mile above he confluence with Davies Creek. U.S.A., OREGON, WASCO COUNTY: Elev. 50 ft. Rocky slopes at Siwash flats near Mosier. Flat area of alluvial deposits, spruce-hemlock Origin: Native. forest, upper river bank. Origin: Native. J. William Thompson 5152 Mary Clay Muller 4097 6 Jul 1929 11 Sep 1980 WTU-5367JWT, WTU-V-016120 WTU-289190, WTU-V-016096 Dryopteridaceae Dryopteridaceae Polystichum imbricans (D.C. Eaton) D.H. Wagner ssp. Polystichum californicum (D.C. Eaton) Diels imbricans U.S.A., WASHINGTON, THURSTON COUNTY: U.S.A., OREGON, HOOD RIVER COUNTY: [locality and coordinates withheld; status in Margins of Lost Lake, Mount Hood National Washington: G4/S1 (Threatened; WANHP)] Forest. Elev. 800 ft. Elev. 3000 ft. Shaded cliff crevices. Rare. Origin: Native. 45.489349°, -121.821695°; uncertainty: 2000 m. Ed Alverson 250 Origin: Native. 6 Jan 1980 J. William Thompson 3668 WTU-277099, WTU-V-016116 21 Aug 1927 WTU-5364JWT, WTU-V-016179 Dryopteridaceae Dryopteridaceae Polystichum imbricans (D.C. Eaton) D.H. Wagner ssp. Polystichum imbricans (D.C. Eaton) D.H. Wagner ssp. imbricans imbricans U.S.A., WASHINGTON, CHELAN COUNTY: U.S.A., OREGON, CLACKAMAS COUNTY: Tumwater. Rocky embankment above Oswego Lake. Elev. 1400 ft. Origin: Native. 47.678278°, -120.734379°; WGS84, uncertainty: 500 m., Source: Georeferenced, J. William Thompson 4153 Georef'd by WTU Staff Origin: Native. 6 May 1928 I. C. Otis 1063 WTU-5346JWT, WTU-V-016130 3 Oct 1920 WTU-28932, WTU-V-016177

Dryopteridaceae Dryopteridaceae Polystichum californicum (D.C. Eaton) Diels Polystichum californicum (D.C. Eaton) Diels U.S.A., WASHINGTON, PIERCE COUNTY: U.S.A., WASHINGTON, PIERCE COUNTY: [locality and coordinates withheld; status in [locality and coordinates withheld; status in Washington: G4/S1 (Threatened; WANHP)] Washington: G4/S1 (Threatened; WANHP)] Crevices. Specimen used for illustration in Elev. 1200 ft. Revision of Flora of the Pacific Northwest, WTU Shaded cliff crevice. Common. W/Polystichum Herbarium, 2015. Origin: Native. munitum and hybirds. Origin: Native. D. Wagner 730 Ed Alverson 242 23 Jun 1976 11 Nov 1979 WTU-267218, WTU-V-021966 WTU-277096, WTU-V-016115

Dryopteridaceae Dryopteridaceae Polystichum imbricans (D.C. Eaton) D.H. Wagner ssp. Polystichum imbricans (D.C. Eaton) D.H. Wagner ssp. imbricans imbricans U.S.A., OREGON COUNTY: U.S.A., WASHINGTON, CHELAN COUNTY: 1 mile east of Dalles. Tumwater canyon, Wenatchee mountains. Origin: Native. Elev. 1400 ft. 47.58472°, -120.67722°; WGS84, uncertainty: Morton E. Peck 14871a 10000 m., Source: GeoLocate, Georef'd by WTU Staff WTU-8127JWT, WTU-V-016191 Origin: Native. I. C. Otis 1063 3 Oct 1920 WTU-8117JWT, WTU-V-016178 Dryopteridaceae Dryopteridaceae Polystichum imbricans (D.C. Eaton) D.H. Wagner ssp. Polystichum californicum (D.C. Eaton) Diels imbricans U.S.A., WASHINGTON, PIERCE COUNTY: U.S.A., OREGON, JOSEPHINE COUNTY: [locality and coordinates withheld; status in Summit of Lake Mountain, Oregon Caves Washington: G4/S1 (Threatened; WANHP)] National Monuments. Crevices of rocky ledges. Rare. Origin: Native. 42.090139°, -123.359861°; WGS 84, uncertainty: 800 m., Source: Digital map, J. B. Flett 1701 Georef'd by Joseph Stormer Origin: Native. 29 Sep 1900 W. H. Baker 352 WTU-174854, WTU-V-016118 3 Jul 1949 WTU-298179, WTU-V-016175 Dryopteridaceae Dryopteridaceae Polystichum californicum (D.C. Eaton) Diels Polystichum californicum (D.C. Eaton) Diels U.S.A., WASHINGTON, THURSTON COUNTY: [locality and coordinates withheld; status in U.S.A., WASHINGTON, PIERCE COUNTY: Washington: G4/S1 (Threatened; WANHP)] [locality and coordinates withheld; status in Cliff. #4 less dissected large frond: P. Washington: G4/S1 (Threatened; WANHP)] californicum X P. munitum,sporangia abortive. Elev. 1200 ft. #5 more dissected large frond: P. Shady cliff crevice. Common. With Polystichum californicum,spores normal. Phenology: Spores. munitum and hybrids. Specimen used for Origin: Native. illustration in Revision of Flora of the Pacific Northwest, WTU Herbarium, 2015. Origin: Ed Alverson s.n. Native. 21 Mar 1980 Ed Alverson 241B WTU-316894, WTU-V-030988 11 Nov 1979 WTU-277097, WTU-V-021967 Dryopteridaceae Dryopteridaceae Polystichum kruckebergii W.H. Wagner Polystichum kruckebergii W.H. Wagner U.S.A., WASHINGTON, CLALLAM COUNTY: In crevices of rock above talus. East slope of U.S.A., WASHINGTON COUNTY: obstruction point, Hurricane ridge. Olympic Mountains, without definite locality. 47.93333°, -123.40833°; WGS 84, Phenology: Spores. Origin: Native. uncertainty: 10000 m., Source: GeoLocate, Georef'd by WTU Staff J. Nieman s.n. Origin: Native. 1932 A. R. Kruckeberg s.n. WTU-23689, WTU-V-016142 5 Sep 1971 WTU-274738, WTU-V-016145

Dryopteridaceae Dryopteridaceae Polystichum scopulinum (D.C. Eaton) Maxon Polystichum californicum (D.C. Eaton) Diels CANADA, BRITISH COLUMBIA: U.S.A., WASHINGTON, THURSTON COUNTY: [locality and coordinates withheld; status in [locality and coordinates withheld; status in British Columbia: S2 (Imperiled, Red List; Washington: G4/S1 (Threatened; WANHP)] BCCDC)] #3P. californicum, more dissected, spores Elev. 5200 ft. normal;#2 P. californicum X P. munitum, less dissected frond. Phenology: Spores. Origin: A. R. Kruckeberg 5738 Native. 10 Sep 1963 Ed Alverson s.n. WTU-217647, WTU-V-016138 21 Mar 1980 WTU-316895, WTU-V-030987 Dryopteridaceae Dryopteridaceae Polystichum lonchitis (L.) Roth Polystichum kruckebergii W.H. Wagner U.S.A., MONTANA, FLATHEAD COUNTY: U.S.A., WASHINGTON, MASON COUNTY: Jewel Basin, Swan Range, 6 miles northeast of Lake Cushman. Echo Lake. 47.42167°, -123.22167°; WGS 84, Elev. 6500 ft. uncertainty: 10000 m., Source: GeoLocate, 48.170236°, -113.924276°; WGS 84, Georef'd by WTU Staff uncertainty: 722 m., Source: GeoLocate, Georef'd by Joseph Stormer C. V. Piper s.n. Rocky slope. Origin: Native. Aug 1890 J. M. Rogers 1285 WTU-2044, WTU-V-016141 28 Aug 1942 WTU-243327, WTU-V-016245 Dryopteridaceae Dryopteridaceae Polystichum lemmonii Underw. Polystichum lemmonii Underw. U.S.A., WASHINGTON, KITTITAS COUNTY: Along Beverly Creek, Wenatchee Mts. U.S.A., WASHINGTON, CHELAN COUNTY: Elev. 4000 ft. Mt. Stuart region, Wenatchee Mts. 47.372454°, -120.877017°; WGS 84, Elev. 4500-8000 ft. uncertainty: 10000 m., Source: GeoLocate, 47.47024°, -120.78503°; WGS84, uncertainty: Georef'd by WTU Staff 13000 m., Source: Georeferenced, Georef'd Underside of boulders near timberline. by WTU Staff Phenology: Fertile. Origin: Native. Abundant. Collected: 27-31 July 1931. Phenology: Fertile. Origin: Native. J. William Thompson 10036 J. William Thompson 7686 2 Sep 1933 27 Jul 1931 WTU-11135JWT, WTU-V-016161 WTU-11131JWT, WTU-V-016215 Dryopteridaceae Dryopteridaceae Polystichum lonchitis (L.) Roth Polystichum kruckebergii W.H. Wagner U.S.A., WASHINGTON, KING COUNTY: Guy Peak, near Snoqualmie Pass. U.S.A., MONTANA, DEER LODGE COUNTY: Elev. 3500 ft. [locality and coordinates withheld; status in 47.52889°, -121.82417°; WGS 84, Montana: S2S3 (Rare; MTNHP)] uncertainty: 5000 m., Source: GeoLocate, Elev. 9300 ft. Georef'd by WTU Staff Origin: Native. Rocky crevices. Origin: Native. Klaus H. Lackschewitz 5793 J. William Thompson 9685 4 Sep 1974 7 Aug 1933 WTU-267355, WTU-V-016129 WTU-11534JWT, WTU-V-021137

Dryopteridaceae Dryopteridaceae Polystichum kruckebergii W.H. Wagner Polystichum kruckebergii W.H. Wagner U.S.A., WASHINGTON, YAKIMA COUNTY: U.S.A., MONTANA, DEER LODGE COUNTY: Crevices of rocks, Mount Adams. [locality and coordinates withheld; status in Elev. 6000-7000 ft. Montana: S2S3 (Rare; MTNHP)] 46.20255°, -121.4908°; WGS 84, uncertainty: Elev. 9275 ft. 8000 m., Source: GeoLocate, Georef'd by Origin: Native. WTU Staff Origin: Native. Klaus H. Lackschewitz 4111 W. N. Suksdorf 297 10 Sep 1972 1 Oct 1881 WTU-257668, WTU-V-016135 WTU-270513, WTU-V-016143 Dryopteridaceae Dryopteridaceae Polystichum lonchitis (L.) Roth Polystichum lonchitis (L.) Roth U.S.A., WASHINGTON, KITTITAS COUNTY: U.S.A., ALASKA, ALEUTIANS WEST COUNTY: Slopes above Hyas Lake, 37 miles northwest of Unalaska. Cle Elum. 53.583944°, -166.796107°; uncertainty: 68000 Elev. 3600 ft. m. 47.56722°, -121.12114°; WGS 84, Origin: Native. uncertainty: 1000 m., Source: GeoLocate, Georef'd by WTU Staff J. B. Flett 1523 Under spruce forest. Origin: Native. 19 Jun 1900 C. L. Hitchcock 8049 WTU-37541, WTU-V-016244 14 Jul 1942 WTU-75330, WTU-V-016246 Dryopteridaceae Dryopteridaceae Polystichum lemmonii Underw. Polystichum lemmonii Underw. U.S.A., WASHINGTON, KING COUNTY: Slopes bordering Eldorado Creek, about De U.S.A., WASHINGTON, KITTITAS COUNTY: Roux Forest Camp, north fork of the Teanaway Wenatchee Mountains. North fork Teanaway river area. River. 47.559138°, -120.005888°; WGS 84, Elev. 4500 ft. uncertainty: 10000 m., Source: GeoLocate, 47.251427°, -120.87694°; WGS 84, Georef'd by David Giblin, Invalid coordinates uncertainty: 10000 m., Source: GeoLocate, (located outside county bounds). Georef'd by WTU Staff Open wooded serpentine sloes boarding creek. Phenology: Fertile. Origin: Native. With P. Lemmonii and P. scopulinum. Mass collection of single fronds in population mix of P. C. S. English s.n. Lemmonii and P. scopulinum. Phenology: Fertile. Jul 1932 A. R. Kruckeberg 5787 WTU-315830, WTU-V-016162 6 Jul 1964 WTU-313567, WTU-V-016221 Dryopteridaceae Polystichum lonchitis (L.) Roth Dryopteridaceae U.S.A., WASHINGTON, PIERCE COUNTY: Polystichum lonchitis (L.) Roth Near Owyhigh Lakes, Mount Rainier National Park. U.S.A., MONTANA, PARK COUNTY: Elev. 5500 ft. Crazy Mountain, Blue Lake Trail. 46.75667°, -122.05472°; WGS 84, Elev. 7400 ft. uncertainty: 10000 m., Source: GeoLocate, Ledge, granite. Origin: Native. Georef'd by WTU Staff Rocky crevices of cliffs. Origin: Native. F. B. Cotner s.n. J. William Thompson 8030 20 Aug 1938 27 Aug 1931 WTU-174491, WTU-V-016243 WTU-11136JWT, WTU-V-021135

Dryopteridaceae Dryopteridaceae Polystichum lemmonii Underw. Polystichum lonchitis (L.) Roth U.S.A., WASHINGTON, CHELAN COUNTY: U.S.A., OREGON, KLAMATH COUNTY: Mt. Stuart region. Kerr Notch, Crater Lake National Park. Elev. 4000-6000 ft. Origin: Native. 47.47024°, -120.78503°; WGS84, uncertainty: 13000 m., Source: Georeferenced, Georef'd W. H. Baker 7139 by WTU Staff By edges of granite boulders. Phenology: 3 Aug 1950 Fertile. Origin: Native. WTU-294306, WTU-V-021133 J. William Thompson 5870 20 Aug 1930 WTU-23232, WTU-V-016159 Dryopteridaceae Dryopteridaceae Polystichum lonchitis (L.) Roth Polystichum lemmonii Underw. U.S.A., IDAHO, BONNER COUNTY: U.S.A., WASHINGTON, KITTITAS COUNTY: Granite Pass, 16 miles north of Nordman. Scatter Creek Trail, 2 miles from junction with Rocky cliffs. Origin: Native. Fish Creek Road. Elev. 4200 ft. William H. Baker 16914 47.507151°, -121.059494°; WGS 84, uncertainty: 10000 m., Source: GeoLocate, 3 Jul 1965 Georef'd by WTU Staff Rock crevices. Phenology: Fertile. Origin: WTU-263936, WTU-V-021131 Native. C. L. Hitchcock 7994 6 Jul 1942 WTU-75321, WTU-V-016160 Dryopteridaceae Polystichum lemmonii Underw. Dryopteridaceae U.S.A., WASHINGTON, KITTITAS COUNTY: Polystichum lonchitis (L.) Roth Along Boulder Creek. Elev. 3500 ft. U.S.A., MONTANA, GALLATIN COUNTY: 47.198258°, -120.4389°; WGS 84, Hyalite Canyon, Bozeman. uncertainty: 10000 m., Source: GeoLocate, 45.490246°, -110.993154°; WGS 84, Georef'd by WTU Staff uncertainty: 10000 m., Source: Digital Map, Rock slides along creek. Phenology: Spores. Georef'd by Joseph Stormer Origin: Native. Deep shade, lodge pole pine, moist few scattered plants. Origin: Native. J. William Thompson 10496 W. E. Booth s.n. 12 May 1934 10 Aug 1946 WTU-24599, WTU-V-016218 WTU-116386, WTU-V-016241

Dryopteridaceae Dryopteridaceae Polystichum lonchitis (L.) Roth Polystichum lemmonii Underw. U.S.A., WASHINGTON, GRAYS HARBOR COUNTY: U.S.A., OREGON, GRANT COUNTY: Bob Mountain [Colonel Bob Mountain]. North rim of summit, Baldy Mountain, Malheur Elev. 4000 ft. National Forest. 47.48852°, -123.74628°; WGS 84, Peridotite talus. Steep talus and rock crevaices. uncertainty: 500 m., Source: GeoLocate, Open subalpine vegitation is very catted and Georef'd by WTU Staff stunted Drouglas fir and limber pine. Phenology: Crevices of shaded cliff. Origin: Native. Fertile. Origin: Native. Harold E. Helmrich 227 A. R. Kruckeberg 5484 12 Jul 1933 21 Jul 1962 WTU-306231, WTU-V-021129 WTU-215168, WTU-V-016157

Dryopteridaceae Dryopteridaceae Polystichum lonchitis (L.) Roth Polystichum lemmonii Underw. U.S.A., ALASKA COUNTY: Tongass National Forest. Near Mendenhall U.S.A., WASHINGTON, CHELAN COUNTY: Glacier, about 10 miles northwest of Juneau. Mt. Stuart region, Wenatchee Mts. Collected along East Glacier Trail. Elev. 4500-8000 ft. Elev. 100 ft. 47.47024°, -120.78503°; WGS84, uncertainty: Moraine, area recently deglaciated, growing with 13000 m., Source: Georeferenced, Georef'd P. Braunii, P. Andersonii. Origin: Native. by WTU Staff Abundant. Collected: 27-31 July 1931. Mary Clay Muller 4192 Phenology: Fertile. Origin: Native. 30 Oct 1980 J. William Thompson 7686 WTU-289220, WTU-V-021127 27 Jul 1931 WTU-11130JWT, WTU-V-016158 Dryopteridaceae Dryopteridaceae Polystichum lonchitis (L.) Roth Polystichum lemmonii Underw. U.S.A., WASHINGTON, JEFFERSON U.S.A., WASHINGTON, KITTITAS COUNTY: COUNTY: Just below west face of Mt. Hawkins on truck South of Marmot Pass. trail from Huckleberry Mt., ca 4.5 mi above and 47.81833°, -123.13222°; WGS 84, east of Camp Creek road, upper Cle Elum River uncertainty: 10000 m., Source: GeoLocate, drainage, Wenatchee Nat Forest. Georef'd by WTU Staff Elev. 5000 ft. Alpine. Origin: Native. 47.459731°, -121.049059°; WGS 84, uncertainty: 10000 m., Source: GeoLocate, Harold E. Helmrich 360 Georef'd by WTU Staff Serpentine outcrop. Phenology: Fertile. Origin: 18 Aug 1933 Native. WTU-306257, WTU-V-021047 A. R. Kruckeberg 5656 13 Jul 1963 Dryopteridaceae WTU-217983, WTU-V-016155 Polystichum lonchitis (L.) Roth U.S.A., IDAHO, IDAHO COUNTY: Above Canteen Meadows, just below Fenn Dryopteridaceae Mountains, Crags Mountains, Middle fork Selway river ranger district. Nez Perce National Polystichum lemmonii Underw. Forest. Elev. 7000 ft. U.S.A., WASHINGTON, CHELAN COUNTY: 46.082289°, -115.425324°; WGS 84, Mt. Ingalls, Mt. Stuart District. uncertainty: 5000 m., Source: GeoLocate, Elev. 7216 ft. Georef'd by WTU Staff 47.473451°, -120.946478°; WGS84, Steep subalpine granitic slopes. Origin: Native. uncertainty: 1400 m., Source: Georeferenced, Georef'd by WTU Staff A. R. Kruckeberg 4117 Rock crevices. Rare. Range: Alaska to California. Zone: Arctic. Phenology: Fertile. 8 Jul 1957 Origin: Native. WTU-192055, WTU-V-016242 W. J. Eyerdam 1002 4 Jul 1937 Dryopteridaceae WTU-291652, WTU-V-016217 Polystichum lonchitis (L.) Roth U.S.A., IDAHO, LATAH COUNTY: Dryopteridaceae Southern slope of Bald Mountain. 47.03167°, -116.57083°; WGS 84, Polystichum lonchitis (L.) Roth uncertainty: 10000 m., Source: GeoLocate, Georef'd by WTU Staff U.S.A., IDAHO, SHOSHONE COUNTY: Along streamside. Commonly associated with Along Bullion Creek, 2 miles below summit of Rubus. Origin: Native. Bullion Pass. 47.377453°, -115.764825°; WGS 84, E. L. Richards 4 uncertainty: 10000 m., Source: GeoLocate, Georef'd by WTU Staff 25 Jun 1956 Rocky cliffs. Origin: Native. WTU-299587, WTU-V-021072 William H. Baker 13476 15 Jul 1955 WTU-299573, WTU-V-021089

Dryopteridaceae Polystichum lonchitis (L.) Roth U.S.A., IDAHO, BONNER COUNTY: Priest River. 48.200171°, -116.90305°; WGS 84, uncertainty: 10000 m., Source: GeoLocate, Georef'd by WTU Staff Forest. Origin: Native. R. F. Daubenmire 43362 29 Jul 1943 WTU-252038, WTU-V-016233 Dryopteridaceae Dryopteridaceae Polystichum lonchitis (L.) Roth Polystichum lonchitis (L.) Roth U.S.A., ALASKA, PRINCE WALES KETCHIKAN CANADA, BRITISH COLUMBIA: COUNTY: Moat Lake. Prince of Wales Island. Elev. 4000 ft. 55.536393°, -132.733415°; WGS 84, 49° 41' N, 125° 23' W uncertainty: 112649 m., Source: Digital Map, Open scree. Origin: Native. Georef'd by Joseph Stormer Alpine summits, snowline. Origin: Native. J. E. Underhill 494 J. B. Flett 2015 25 Jul 1957 4 Aug 1901 WTU-214802, WTU-V-016229 WTU-37540, WTU-V-016234

Dryopteridaceae Dryopteridaceae Polystichum lonchitis (L.) Roth Polystichum lemmonii Underw. CANADA, BRITISH COLUMBIA: U.S.A., WASHINGTON, KITTITAS COUNTY: Queen Charlotte Islands, 20 miles south of Just above De Roux Forest Camp, North Fork Moresby Camp, Moresby Island. Teanaway River. Elev. 2250 ft. 47.251427°, -120.87694°; WGS 84, 52° 55' N, 132° 3' W uncertainty: 10000 m., Source: GeoLocate, Large rock colony in runnel at base of rock cliffs. Georef'd by WTU Staff Rare. Collection dates: Jul29-Aug2/1957. Origin: Steep barren serpentine talus. Phenology: Native. Fertile. Origin: Native. J. A. Calder 23115 A. R. Kruckeberg 5560 with R. L. Taylor, D. B. O. Savile 5 Jun 1963 29 Jul 1957 WTU-216016, WTU-V-016153 WTU-239382, WTU-V-016232

Dryopteridaceae Dryopteridaceae Polystichum lonchitis (L.) Roth Polystichum lemmonii Underw. U.S.A., IDAHO, BOUNDARY COUNTY: U.S.A., WASHINGTON, KITTITAS COUNTY: Ledge of east facing cliff Mount Roothaan. Ridge south of highest peak northeast of Fish 48.61326°, -116.707987°; WGS 84, lake; 30 miles nothwest of Cle Elum. uncertainty: 10000 m., Source: GeoLocate, Elev. 5600 ft. Georef'd by WTU Staff 47.19556°, -120.93806°; WGS 84, Spruce and fir forest. Phenology: Fertile. Origin: uncertainty: 10000 m., Source: GeoLocate, Native. Georef'd by WTU Staff Crevices of serpentine rocks. Phenology: R. F. Daubenmire 44318 Fertile. Origin: Native. 3 Jul 1944 C. L. Hitchcock 7962 WTU-271589, WTU-V-021088 6 Jul 1942 WTU-76205, WTU-V-016156 Dryopteridaceae Polystichum lemmonii Underw. Dryopteridaceae U.S.A., WASHINGTON, CHELAN COUNTY: Polystichum lonchitis (L.) Roth Mount Stuart. 47.47503°, -120.90223°; WGS 84, CANADA, BRITISH COLUMBIA: uncertainty: 1500 m., Source: GeoLocate, Haines Road. Mile 60. Georef'd by WTU Staff Elev. 4100 ft. Phenology: Fertile. Origin: Native. 59° 30' N, 136° 30' W Grassy alpine slope. Origin: Native. A. D. E. Elmer 1114 T. M.C. Taylor 1243 Aug 1898 with A. F. Szczawinski, M. Bell WTU-33581, WTU-V-016149 9 Jul 1956 WTU-182064, WTU-V-016231 Dryopteridaceae Dryopteridaceae Polystichum lonchitis (L.) Roth Polystichum lonchitis (L.) Roth U.S.A., WASHINGTON, JEFFERSON U.S.A., IDAHO, IDAHO COUNTY: COUNTY: Southeast of Burnt Knob Lookout. Foot of damp rocks, Elwha Basin, Olympic Elev. 8100 ft. Mountains. 45.46833°, -115.28194°; WGS 84, 47.75583°, -123.56611°; WGS 84, uncertainty: 10000 m., Source: GeoLocate, uncertainty: 10000 m., Source: GeoLocate, Georef'd by WTU Staff Georef'd by WTU Staff Rock cliffs. Origin: Native. Damp rocks. Origin: Native. William H. Baker 12644 Myrtle Culmer s.n. 17 Aug 1954 8 Aug 1913 WTU-299366, WTU-V-021084 WTU-11392JWT, WTU-V-016230

Dryopteridaceae Dryopteridaceae Polystichum lemmonii Underw. Polystichum lonchitis (L.) Roth U.S.A., WASHINGTON, CHELAN COUNTY: U.S.A., WASHINGTON COUNTY: Mt. Stuart region. Olympic Mountains. Elev. 4000-6000 ft. Elev. 5000 ft. 47.47024°, -120.78503°; WGS84, uncertainty: 47.811111°, -123.626944°; Source: Inferred 13000 m., Source: Georeferenced, Georef'd from duplicate specimen, Georef'd by Ed by WTU Staff Alverson By side of granite boulders. Very abundant. Rock crevices. Origin: Native. Phenology: Fertile. Origin: Native. C. V. Piper s.n. J. William Thompson 5870 Sep 1890 20 Aug 1930 WTU-2039, WTU-V-021124 WTU-1299JWT, WTU-V-016150 Dryopteridaceae Dryopteridaceae Polystichum lemmonii Underw. Polystichum lonchitis (L.) Roth U.S.A., WASHINGTON, CHELAN COUNTY: Mt. Stuart. U.S.A., WASHINGTON COUNTY: 47.47522°, -120.90238°; Source: Inferred Goat Mountains. from duplicate specimen, Georef'd by Ed Origin: Native. Alverson Phenology: Fertile. Origin: Native. J. B. Flett s.n. J. H. Sandberg s.n. 19 Aug 1903 with J. B. Leiberg WTU-42912, WTU-V-016227 Aug 1893 WTU-1800, WTU-V-016219

Dryopteridaceae Dryopteridaceae Polystichum lonchitis (L.) Roth Polystichum lemmonii Underw. U.S.A., OREGON, HARNEY COUNTY: Southeast canyon hillside, Steens Mount U.S.A., WASHINGTON, KITTITAS COUNTY: Region, 18 miles due southeast of Frenchglen. De Roux Forest Camp, upper north fork Elev. 8400 ft. Teanaway River, Wenatchee National Forest. T33S R33E S2; NAD 27, uncertainty: 805 m., Above camp. Source: TRS2LL, Georef'd by Ben Legler 47.251427°, -120.87694°; WGS 84, Grass-sedge zone, dry, rocky, in rock cleft. uncertainty: 10000 m., Source: GeoLocate, Rare. Origin: Native. Georef'd by WTU Staff Steep sterile, barren serpentinized slopes. Chas. G. Hansen 633 Phenology: Fertile. Origin: Native. 16 Aug 1953 A. R. Kruckeberg 3940 WTU-294146, WTU-V-021082 14 Aug 1955 WTU-192063, WTU-V-016204 Dryopteridaceae Dryopteridaceae Polystichum lemmonii Underw. Polystichum lonchitis (L.) Roth U.S.A., WASHINGTON, CHELAN COUNTY: U.S.A., OREGON, HARNEY COUNTY: Mt. Stuart region. Southeast canyon hillside, Steens Mount Elev. 5000 ft. Region, 18 miles due southeast of Frenchglen. 47.47503°, -120.90223°; WGS 84, Elev. 8400 ft. uncertainty: 1500 m., Source: GeoLocate, T33S R33E S2; NAD 27, uncertainty: 805 m., Georef'd by WTU Staff Source: TRS2LL, Georef'd by Ben Legler Under boulders, near timberline. Phenology: Grassy-sedge zone, dry, rocky, in rock cleft. Fertile. Origin: Native. Rare. Origin: Native. J. William Thompson 10036 Chas. G. Hansen 633 3 Sep 1933 16 Aug 1953 WTU-111199, WTU-V-016154 WTU-163101, WTU-V-021057

Dryopteridaceae Dryopteridaceae Polystichum lonchitis (L.) Roth Polystichum lonchitis (L.) Roth CANADA, BRITISH COLUMBIA: U.S.A., ALASKA COUNTY: Murray Range at north end of Azouzetta Lake Tongass National Forest. Near Mendenhall along Hart Highway, Pine Pass. Glacier, about 10 miles northwest of Juneau. Elev. 4500 ft. Collected along East Glacier Trail. Among stunted balsam on slope. Origin: Native. Elev. 100 ft. Plants growing on moraine, this area recently J. A. Calder 12615 deglaciated. Growing with P. Braunii, P. with D. B. O. Savile, J. M. Ferguson andersonii. Origin: Native. 7 Jul 1954 Mary Clay Muller 4193 WTU-173798, WTU-V-016228 30 Oct 1980 WTU-289202, WTU-V-021080 Dryopteridaceae Polystichum lemmonii Underw. Dryopteridaceae U.S.A., WASHINGTON, SNOHOMISH Polystichum lemmonii Underw. COUNTY: Just above and east of Dougle Eagle lakes, U.S.A., WASHINGTON, WHATCOM COUNTY: upper drainage of west fork of Coal Creek, ca. 4 Near summit of Yellow Aster Butte. miles east of Silverton, south fork of the 48.95556°, -121.67833°; WGS 84, Stillaguamish River. uncertainty: 10000 m., Source: GeoLocate, 48.114685°, -121.520223°; WGS 84, Georef'd by WTU Staff uncertainty: 1000 m., Source: GeoLocate, Clefts of schist. Phenology: Fertile. Origin: Georef'd by WTU Staff Native. Massive peridotite rocks. Grassy areas. Mtn Hemlock Alaska ceader and Pacific silver fir W. C. Muenscher 7559 forest nearby. Common at base fo peridotite rocks and in grassy areas. Phenology: Fertile. 3 Jul 1937 Origin: Native. WTU-55860, WTU-V-016151 A. R. Kruckeberg 6638 20 Sep 1979 Dryopteridaceae WTU-305250, WTU-V-016208 Polystichum lonchitis (L.) Roth U.S.A., ALASKA, ALEUTIANS WEST COUNTY: Holtz Bay Region. Elev. 150 ft. 52.938533°, 173.172031°; uncertainty: 5000 m. Loam. Origin: Native. M.D. Gilmore W. Soule 325 5 Aug 1945 WTU-155634, WTU-V-016226 Dryopteridaceae Dryopteridaceae Polystichum lemmonii Underw. Polystichum lonchitis (L.) Roth U.S.A., WASHINGTON, WHATCOM COUNTY: U.S.A., MONTANA, LEWIS AND CLARK Twin Sister Range. COUNTY: 48.68259°, -121.97036°; WGS 84, Lewis and Clark National Forest. Sun River uncertainty: 4100 m., Source: Nat. Geog. Primitive Area. East slope of China [Chinese] TOPO!, Georef'd by John Bassett Wall near north end. Rocky slope. Phenology: Spores. Origin: Native. 47.714251°, -113.150509°; WGS 84, uncertainty: 3000 m., Source: Digital Map, W. C. Muenscher 10230 Georef'd by Joseph Stormer Origin: Native. 13 Aug 1939 C. L. Hitchcock 18926 WTU-56757, WTU-V-016210 27 Jul 1948 WTU-126011, WTU-V-021058 Dryopteridaceae Polystichum lonchitis (L.) Roth Dryopteridaceae U.S.A., ALASKA, SKAGWAY YAKUTAT Polystichum lonchitis (L.) Roth ANGOON COUNTY: Tongass National Forest, Chichagof Island U.S.A., MONTANA, LAKE COUNTY: northwest part, west side of Lisianski Inlet, Mission Range, high on canyon walls south and across from Pelican, along Cann Creek. west of McDonald Lake. Origin: Native. 47.416371°, -113.978498°; WGS 84, uncertainty: 1675 m., Source: Digital Map, Mary Clay Muller 3291 Georef'd by Joseph Stormer Origin: Native. 24 Aug 1979 C. L. Hitchcock 18265 WTU-280511, WTU-V-021078 12 Jul 1948 WTU-126012, WTU-V-021060 Dryopteridaceae Polystichum lonchitis (L.) Roth Dryopteridaceae U.S.A., OREGON, CLACKAMAS COUNTY: Trail from Twin Bridges to Paradise Park, Mount Polystichum lonchitis (L.) Roth Hood. 45.331459°, -121.779923°; uncertainty: 3000 U.S.A., ALASKA COUNTY: m., Georef'd by Joseph Stormer Attu Island; Engineer Canyon. Origin: Native. Elev. 200 ft. 52.884154°, 173.193026°; uncertainty: 400 m. J. William Thompson 3437 Moist slopes among rocks. Origin: Native. 5 Aug 1927 Robert M. Hardy 54 WTU-5374JWT, WTU-V-016223 26 Apr 1945 WTU-97619, WTU-V-021056 Dryopteridaceae Polystichum lemmonii Underw. Dryopteridaceae U.S.A., WASHINGTON, CHELAN COUNTY: Mount Stuart. Polystichum lemmonii Underw. Elev. 3800-7000 ft. 47.47522°, -120.90238°; Georef'd by Ed U.S.A., WASHINGTON, KITTITAS COUNTY: Alverson Slides along Beverly Creek, Wenatchee Mts. Phenology: Fertile. Origin: Native. Elev. 4500 ft. 47.372454°, -120.877017°; WGS 84, J. H. Sandberg 812 uncertainty: 10000 m., Source: GeoLocate, Georef'd by WTU Staff 28 Aug 1893 Serpentine slides. Phenology: Fertile. Origin: Native. WTU-37584, WTU-V-016152 J. William Thompson 9584 25 Jul 1933 WTU-11129JWT, WTU-V-016220 Dryopteridaceae Dryopteridaceae Polystichum lonchitis (L.) Roth Polystichum lonchitis (L.) Roth U.S.A., WASHINGTON, WHATCOM COUNTY: U.S.A., WASHINGTON, CLALLAM COUNTY: Mount Hermann, Mount Baker National Forest. Mount Angeles, Olympic Mountains. Elev. 4500 ft. Elev. 5500 ft. 48.86361°, -121.68111°; Source: Inferred 47.993333°, -123.47275°; Source: Inferred from duplicate specimen, Georef'd by Ed from duplicate specimen, Georef'd by Ed Alverson Alverson Rocky Cliffs. Collection date: jul22-25/1930. Rocky Crevices. Origin: Native. Origin: Native. J. William Thompson 5526 J. William Thompson 5253 2 Aug 1930 22 Jul 1930 WTU-11512JWT, WTU-V-021012 WTU-11139JWT, WTU-V-021140 Dryopteridaceae Dryopteridaceae Polystichum lemmonii Underw. Polystichum lemmonii Underw. U.S.A., WASHINGTON, CHELAN COUNTY: U.S.A., WASHINGTON, CHELAN COUNTY: Mount Stuart. Base of Three Brothers. 47.47503°, -120.90223°; WGS 84, Elev. 4000 ft. uncertainty: 1500 m., Source: GeoLocate, 47.43891°, -120.74198°; WGS84, uncertainty: Georef'd by WTU Staff 2000 m., Source: Georeferenced, Georef'd by Phenology: Fertile. Origin: Native. WTU Staff Rocky slopes. Phenology: Fertile. Origin: Native. A. D. E. Elmer 1114 J. William Thompson 10753 Aug 1898 21 Jun 1934 WTU-37537, WTU-V-016166 WTU-25868, WTU-V-016147 Dryopteridaceae Dryopteridaceae Polystichum lonchitis (L.) Roth Polystichum lemmonii Underw. U.S.A., WASHINGTON COUNTY: Olympic Mountains. U.S.A., WASHINGTON, KITTITAS COUNTY: Elev. 5500 ft. Edge of serpentine outcrop of east-facing slope 47.811111°, -123.626944°; Source: Inferred of Koppen Mt. ridge, ca. 2 miles above De Roux from duplicate specimen, Georef'd by Ed Cr., North Fork Teanaway River drainage. Alverson Elev. 5000 ft. Rocky places. Origin: Native. 47.251427°, -120.87694°; WGS 84, uncertainty: 10000 m., Source: GeoLocate, J. B. Flett s.n. Georef'd by WTU Staff In shade of Tsuga mertensiana forest, east of 28 Aug 1898 serpentine outcrop. Phenology: Fertile. Origin: Native. WTU-37542, WTU-V-021142 A. R. Kruckeberg 5763 Dryopteridaceae 20 Sep 1963 Polystichum lonchitis (L.) Roth WTU-217659, WTU-V-016164 U.S.A., ALASKA COUNTY: Southeast; Chichagof Island; Icy Strait; Gallagher Creek drainage. Dryopteridaceae Elev. 1590 ft. T43S R60E S2; WGS 84, uncertainty: 8376 Polystichum lemmonii Underw. m., Source: Digital Map, Georef'd by Joseph Stormer U.S.A., WASHINGTON, WHATCOM COUNTY: Aspect South; slope 70%; open volcanic rock Winchester Pass, Cascade Mts. outcrop in Western Hemlock-Spruce-Picea Elev. 5500 ft. forest; growing with Cryptogramma crispa, Rocky slopes. Phenology: Fertile. Origin: Native. Heuchera glabra, Galium kamtschaticm. Locally common. Phenology: Spores. Origin: Native. Harry B. Higman s.n. Kitty LaBounty 180 Aug 1931 16 Aug 1981 WTU-11134JWT, WTU-V-016163 WTU-282072, WTU-V-021126 Dryopteridaceae Dryopteridaceae Polystichum lemmonii Underw. Polystichum lemmonii Underw. U.S.A., WASHINGTON, KITTITAS COUNTY: U.S.A., WASHINGTON, CHELAN COUNTY: Head of Beverly Creek, Wenatchee Mountains. At 6 mile mark on trail to 8-Mile Lake, just above Elev. 6000 ft. Little 8-Mile Lake, Icicle Creek Drainage. 47.372454°, -120.877017°; WGS 84, 47.557948°, -120.672115°; WGS84, uncertainty: 10000 m., Source: GeoLocate, uncertainty: 10000 m., Source: GeoLocate, Georef'd by WTU Staff Georef'd by WTU Staff Crevices of cliffs. Collected: 27-31 July 1931. Shallow stony soil of massive peridotite talus. In Phenology: Fertile. Origin: Native. deep shade of rocks less dwarfed and stenophyllous. Some plts. Phenology: Fertile. J. William Thompson 7667 Origin: Native. 27 Jul 1931 A. R. Kruckeberg 5425 WTU-11133JWT, WTU-V-016165 7 Jul 1962 WTU-218816, WTU-V-016168 Dryopteridaceae Polystichum lonchitis (L.) Roth Dryopteridaceae U.S.A., WASHINGTON, JEFFERSON Polystichum lonchitis (L.) Roth COUNTY: Marmot Pass, Olympic Mountains. U.S.A., OREGON, CLACKAMAS COUNTY: Elev. 5500 ft. Along base of high cliffs at Mirror Lake, 47.81833°, -123.13222°; WGS 84, Mountain Hood National Forest. uncertainty: 10000 m., Source: GeoLocate, Elev. 4000 ft. Georef'd by WTU Staff 45.294893°, -121.794802°; WGS 84, Crevices of cliffs. Origin: Native. uncertainty: 1000 m., Source: Digital Map, Georef'd by Joseph Stormer J. William Thompson 9924 Phenology: Spores. Origin: Native. 18 Aug 1933 J. William Thompson 3572 WTU-11510JWT, WTU-V-021010 17 Aug 1927 WTU-11509JWT, WTU-V-021065 Dryopteridaceae Polystichum lonchitis (L.) Roth Dryopteridaceae U.S.A., WASHINGTON, CLALLAM COUNTY: Obstruction Point, Olympic National Park. Polystichum lonchitis (L.) Roth 48.26667°, -124.675°; WGS 84, uncertainty: 10000 m., Source: GeoLocate, Georef'd by U.S.A., MONTANA, SWEET GRASS COUNTY: WTU Staff Under granite boulders, below Blue Lake, above Slate crevices of cliffs. Rare "only plant seen", Big Timber Creek, Crazy Mountains. with Aster paucicapitatus, Viola flettii, Oxyria Moist slope. Origin: Native. digyna, Campanula piperi, Arnica cordifolia. N=41. Origin: Native. C. L. Hitchcock 13240 with C. V. Muhlick A. R. Kruckeberg 5667 5 Aug 1945 13 Jul 1963 WTU-100803, WTU-V-021011 WTU-216605, WTU-V-021013

Dryopteridaceae Polystichum lemmonii Underw. U.S.A., WASHINGTON, KITTITAS COUNTY: Along Beverly Creek, Wenatchee Mts. Elev. 4000-9000 ft. 47.372454°, -120.877017°; WGS 84, uncertainty: 10000 m., Source: GeoLocate, Georef'd by WTU Staff Rocky open slopes. Phenology: Fertile. Origin: Native. J. William Thompson 6391 16 May 1931 WTU-11132JWT, WTU-V-016203 Dryopteridaceae Dryopteridaceae Polystichum lemmonii Underw. Polystichum lonchitis (L.) Roth U.S.A., WASHINGTON, SNOHOMISH U.S.A., OREGON, JOSEPHINE COUNTY: COUNTY: Ledges on Bolan Peak, Siskiyou Mountains. Narrow cirque above Double Eagle Lakes, Elev. 6000 ft. upper West Fork Coal Creek drainage. Crevices. Origin: Native. 48.086015°, -121.54013°; WGS 84, uncertainty: 10000 m., Source: GeoLocate, J. William Thompson 12483 Georef'd by WTU Staff Subalpine talus and fellfields in narrow cirque. 5 Aug 1935 Infrequent. on dunite-paridotite. Phenology: Fertile. Origin: Native. WTU-34059, WTU-V-021070 A. R. Kruckeberg 5327 3 Aug 1961 WTU-218778, WTU-V-016206 Dryopteridaceae Polystichum lemmonii Underw. Dryopteridaceae U.S.A., WASHINGTON, KITTITAS COUNTY: Polystichum lonchitis (L.) Roth Ca. 1 mils (southwest) below Teanaway Turnpike on ridge. Wenatchee Mts. U.S.A., OREGON, JOSEPHINE COUNTY: Elev. 6500 ft. Ledges on Bolan Peak, Siskiyou Mountains. 47.175°, -120.85639°; WGS 84, uncertainty: Elev. 6000 ft. 10000 m., Source: GeoLocate, Georef'd by Crevices. Origin: Native. WTU Staff Open serpentine slopes with scattered Pinus J. William Thompson 12483 albicaulis, Abies lasiocarpa. Phenology: Fertile. Origin: Native. 5 Aug 1935 A. R. Kruckeberg 2600 WTU-5128JWT, WTU-V-021009 8 Aug 1950 WTU-169971, WTU-V-016167

Dryopteridaceae Dryopteridaceae Polystichum lemmonii Underw. Polystichum lonchitis (L.) Roth U.S.A., WASHINGTON, CHELAN COUNTY: Southeast-facing upper slopes of Three U.S.A., WASHINGTON, SNOHOMISH Brothers Peak, just below lookout, on trail to COUNTY: Etienne Creek. Mount Pugh, Cascade Mountains. 47.428889°, -120.7625°; NAD 27, uncertainty: Elev. 7000 ft. 500 m., Source: Georeferenced, Georef'd by 48.143056°, -121.372778°; NAD 27, WTU Staff uncertainty: 1000 m., Source: Georeferenced, Whitebark pine ("krummholz") forest type. Georef'd by WTU Staff Serpentine, but with a mixture of sandstone, Alpine slopes. Origin: Native. gabbroic rock. Talus of slope. Phenology: Fertile. Origin: Native. J. William Thompson 14329 A. R. Kruckeberg 5282 18 Aug 1938 20 Jul 1961 WTU-17671JWT, WTU-V-021069 WTU-215167, WTU-V-016205 Dryopteridaceae Polystichum lonchitis (L.) Roth Dryopteridaceae U.S.A., WASHINGTON, CHELAN COUNTY: Polystichum lonchitis (L.) Roth Trail to Snow Lakes, Wenatchee Mountains. Elev. 3500 ft. U.S.A., MONTANA, RAVALLI COUNTY: 47.48556°, -120.74222°; WGS84, uncertainty: Above Blodgett Lake; Bitterroot-Selway Divide. 10000 m., Source: GeoLocate, Georef'd by Elev. 7800 ft. WTU Staff East-slope, north-exposure; crevice with Moist rock crevices. Origin: Native. seeping moisture. Phenology: Spores. Origin: Native. J. William Thompson 17140 Klaus H. Lackschewitz 1846 11 Aug 1952 with Tor Fageraas WTU-153936, WTU-V-021141 17 Aug 1969 WTU-257667, WTU-V-021054 Dryopteridaceae Dryopteridaceae Polystichum lemmonii Underw. Polystichum lemmonii Underw. U.S.A., WASHINGTON, KITTITAS COUNTY: U.S.A., WASHINGTON, WHATCOM COUNTY: Slopes at Paddy-Go-Easy Pass, Wenatchee Steep SW-facing slope of Twin Sisters Mtn. National Forest. (north peak), above Orsina Creek. 47.545°, -121.05889°; WGS 84, uncertainty: 48.70833°, -121.99167°; WGS 84, 10000 m., Source: GeoLocate, Georef'd by uncertainty: 10000 m., Source: GeoLocate, WTU Staff, Invalid coordinates (located Georef'd by WTU Staff outside county bounds). Olivine talus. Subalpine fir-alaska cedar and Steep rocky slopes. Phenology: Fertile. Origin: lodgepole pine. Phenology: Fertile. Origin: Native. Native. A. R. Kruckeberg 2947 A. R. Kruckeberg 5231 22 Jun 1952 12 Jul 1961 WTU-154496, WTU-V-016170 WTU-217548, WTU-V-016169

Dryopteridaceae Dryopteridaceae Polystichum lonchitis (L.) Roth Polystichum lonchitis (L.) Roth U.S.A., OREGON, JOSEPHINE COUNTY: CANADA, BRITISH COLUMBIA: Ledges above Upper Biglow Lake, Oregon Mountain near Kennedy River along Caves National Monument. TofinoUcluelet road. Rock ledges. Origin: Native. Elev. 3000-3500 ft. 49° 13' N, 125° 23' W W. H. Baker 706 Occasional on talus slope and along margin of with G. C. Ruhle rock runnels. Phenology: Spores. 3 Sep 1949 J. A. Calder 32003 with K. T. MacKay WTU-298242, WTU-V-021139 1 Aug 1961 WTU-215722, WTU-V-021063 Dryopteridaceae Polystichum imbricans (D.C. Eaton) D.H. Wagner ssp. Dryopteridaceae imbricans Polystichum lonchitis (L.) Roth U.S.A., OREGON, HOOD RIVER COUNTY: At Viento, Columbia Gorge. U.S.A., MONTANA, FLATHEAD COUNTY: Rocky cliffs. Phenology: Spores. Origin: Native. Meadow, Logan Pass, Glacier National Park. 48.69953°, -113.718857°; uncertainty: 800 m. J. William Thompson 4041 Origin: Native. 7 Apr 1928 Fred A. Barkley 1727 WTU-5345JWT, WTU-V-016199 18 Aug 1937 WTU-41643, WTU-V-021068

Dryopteridaceae Dryopteridaceae Polystichum lonchitis (L.) Roth Polystichum munitum (Kaulf.) C. Presl U.S.A., OREGON, KLAMATH COUNTY: Kerr Notch, Crater Lake National Park. U.S.A., WASHINGTON, SAN JUAN COUNTY: Origin: Native. Yellow Island; center of island. 48.592222°, -123.031944°; WGS 84, W. H. Baker 7139 uncertainty: 250 m., Source: Georeferenced, Georef'd by WTU Staff 3 Aug 1950 Pseudotsuga menziesii woodland. Phenology: Spores. Origin: Native. WTU-299570, WTU-V-021073 E. Habegger s.n. 17 Apr 1996 WTU-333131-B, WTU-V-18318 Dryopteridaceae Dryopteridaceae Polystichum lonchitis (L.) Roth Polystichum lonchitis (L.) Roth U.S.A., OREGON, BAKER COUNTY: U.S.A., ALASKA, JUNEAU COUNTY: Elkhorn Ridge, Rocky northwest slopes above Mountain near Juneau. Pine Creek Reservoir. Elev. 3000 ft. Elev. 6750 ft. 58.339447°, -134.342818°; uncertainty: 50000 Occassionally associated with Pinus albicaulis, m. heuchera cylindrica alpina, Pellaea breweri, Phenology: Spores. Origin: Native. Ribes montigenum. Phenology: Spores. Origin: Native. Emma A. Shumway s.n. Brenda Bafus 379 6 Aug 1891 27 Jul 1979 WTU-90308, WTU-V-021062 WTU-278618, WTU-V-021130 Dryopteridaceae Dryopteridaceae Polystichum lonchitis (L.) Roth Polystichum lonchitis (L.) Roth U.S.A., ALASKA COUNTY: Attu Island; Engineer Canyon; Sidden's Valley. U.S.A., ALASKA, JUNEAU COUNTY: Elev. 250 ft. Nugget Creek Trail. 52.884154°, 173.193026°; WGS 84, Holly fern. Origin: Native. uncertainty: 432 m., Source: Digital Map, Georef'd by Joseph Stormer Lucille Stonehouse s.n. Herbaceous slopes. Origin: Native. 20 Oct 1943 Robert M. Hardy 405 WTU-110852, WTU-V-016240 23 Sep 1945 WTU-97615, WTU-V-016239

Dryopteridaceae Dryopteridaceae Polystichum lonchitis (L.) Roth Polystichum munitum (Kaulf.) C. Presl U.S.A., OREGON, WALLOWA COUNTY: U.S.A., WASHINGTON, SNOHOMISH Razz Lake, Wallowa Mountains. COUNTY: 45.212444°, -117.277325°; WGS 84, Perry Creek drainage, ca. .5 miles above uncertainty: 473 m., Source: GeoLocate, trailhead and northeast end of Forest Service Georef'd by Joseph Stormer Road 4063, northwest slope of Dickerman Origin: Native. Mountian. Cascade Mountains, Mt. Baker-Snoqualmie National Forest. Lilla Leach 4217 Elev. 2197 ft. 48° 4.4' N, 121° 29.5' W; T30N R10E S23 1 Aug 1931 Shaded trailside, northwest aspect, steep mossy and rocky slope, with Blechnum spicant, Acer WTU-255902, WTU-V-016238 circinatum, Athyrium filix-femina, Botrychium multifidum, Polystichum andersonii. Cespitose, evergreen, dark green, common. Phenology: Fertile. Origin: Native. Dryopteridaceae P. F. Zika 19082 20 Sep 2003 Polystichum munitum (Kaulf.) C. Presl WTU-354254, WTU-V-18335 U.S.A., IDAHO, LATAH COUNTY: Idler's Rest north of Moscow near Moscow Dryopteridaceae Mountain. 46.7325°, -116.99917°; WGS 84, uncertainty: Polystichum munitum (Kaulf.) C. Presl 5000 m., Source: GeoLocate, Georef'd by WTU Staff U.S.A., IDAHO COUNTY: East aspect in dense forest canopy near a Coeur D'Alene Mountains. Forks on St. Mary's creek, little sunlight. Moist soils associated wtih River. Cupressaceae, Trillium ovatum, and some Elev. 4756 ft. grasses. 5-15 dm tall, reproduction by spores on Phenology: Spores. Origin: Native. underside of leaflets. common christmans-fern, Sword-fern. Phenology: Fertile. Origin: Native. John B. Leiberg 1147 J. D. Robertson 859 3 Jul 1895 13 Apr 2003 WTU-56196, WTU-V-18429 WTU-354406, WTU-V-18437 Dryopteridaceae Dryopteridaceae Polystichum lonchitis (L.) Roth Polystichum imbricans (D.C. Eaton) D.H. Wagner ssp. imbricans U.S.A., ALASKA, VALDEZ CORDOVA COUNTY: U.S.A., WASHINGTON, CHELAN COUNTY: Evans Island. Port San Juan. Between Drury and Chiwaukum, near G. N. 60.051325°, -148.071491°; uncertainty: 1200 snow shed 1680.3. m., Georef'd by Joseph Stormer Elev. 1600 ft. Cliff crevice. Very rare. Origin: Native. 47.656962°, -120.729561°; WGS84, uncertainty: 4600 m., Source: Georeferenced, Walter J. Eyerdam 7192 Georef'd by WTU Staff Phenology: Spores. Origin: Native. 30 Aug 1948 I. C. Otis 1053 WTU-293242, WTU-V-021122 16 Sep 1920 WTU-8126JWT, WTU-V-016202 Dryopteridaceae Polystichum lonchitis (L.) Roth Dryopteridaceae U.S.A., ALASKA, ALEUTIANS WEST COUNTY: Unalaska. Polystichum lonchitis (L.) Roth 53.583944°, -166.796107°; uncertainty: 68000 m. U.S.A., MONTANA, FERGUS COUNTY: Phenology: Spores. Origin: Native. On limestone cliff, Half Moon Canyon. 3.5miles from mouth, Big Snowy Mountains. Trevor Kincaid 101 Phenology: Spores. Origin: Native. 18 Sep 1897 C. L. Hitchcock 12000 with C. V. Muhlick WTU-101409, WTU-V-021061 5 Jul 1945 WTU-98358, WTU-V-021050 Dryopteridaceae Polystichum munitum (Kaulf.) C. Presl Dryopteridaceae U.S.A., OREGON COUNTY: Salem. Polystichum lonchitis (L.) Roth Phenology: Fertile. Origin: Native. U.S.A., MONTANA, FLATHEAD COUNTY: O. B. Johnson s.n. Logan Pass, St. Mary Lake, Hudsonian zone, Glacier National Park, slope north. Oct 1899 Elev. 7000 ft. 48.69953°, -113.718857°; uncertainty: 800 m. WTU-1989, WTU-V-18326 Origin: Native. Lyman Benson 5240 17 Jul 1933 WTU-56186, WTU-V-021125 Dryopteridaceae Polystichum lonchitis (L.) Roth Dryopteridaceae U.S.A., WASHINGTON, WHATCOM COUNTY: Near summit, Church Mountain. Polystichum imbricans (D.C. Eaton) D.H. Wagner ssp. 48.92694°, -121.88083°; WGS 84, imbricans uncertainty: 10000 m., Source: GeoLocate, Georef'd by WTU Staff U.S.A., OREGON, CLACKAMAS COUNTY: Exposed cliff. A dwarfed form with general Clackamas River above Estacada. appearance of P. lonchites. Phenology: Spores. Growing in rocky places along river. Phenology: Origin: Native. Spores. Origin: Native. W. C. Muenscher 9838 J. William Thompson 2642 9 Jun 1939 11 Jun 1927 WTU-56759, WTU-V-016237 WTU-5368JWT, WTU-V-016201 Dryopteridaceae Dryopteridaceae Polystichum munitum (Kaulf.) C. Presl Polystichum munitum (Kaulf.) C. Presl U.S.A., IDAHO COUNTY: U.S.A., IDAHO, IDAHO COUNTY: Chatcolet. Slim's Camp near mouth of Meadow Creek, Phenology: Fertile. Origin: Native. South of Selway River. 45.481973°, -115.931284°; WGS 84, Carl Cozier s.n. uncertainty: 10000 m., Source: GeoLocate, Georef'd by WTU Staff 25 Jun 1904 Rocky bar. Phenology: Fertile. Origin: Native. WTU-139299, WTU-V-18257 W. H. Baker 14255 6 Jul 1956 WTU-299590, WTU-V-18436

Dryopteridaceae Dryopteridaceae Polystichum lonchitis (L.) Roth Polystichum lonchitis (L.) Roth U.S.A., WASHINGTON, CHELAN COUNTY: Iron Mountain, Wenatchee Mountains. U.S.A., WASHINGTON, WHATCOM COUNTY: Elev. 5000 ft. Mount Baker, Minning District. 47.41222°, -120.71417°; WGS 84, 48.9665°, -121.62758°; WGS 84, uncertainty: uncertainty: 10000 m., Source: GeoLocate, 3500 m., Source: Nat. Geog. TOPO!, Georef'd Georef'd by WTU Staff by John Bassett Crevices of rocky outcrops. Origin: Native. Origin: Native. J. William Thompson 6651 J. B. Flett s.n. 13 Jun 1931 20 Jul 1898 WTU-11137JWT, WTU-V-016235 WTU-37450, WTU-V-021121

Dryopteridaceae Dryopteridaceae Polystichum lonchitis (L.) Roth Polystichum munitum (Kaulf.) C. Presl U.S.A., OREGON, WALLOWA COUNTY: U.S.A., OREGON, UMATILLA COUNTY: Source of Wallowa River, Wallowa Mountains. Bingham Springs. Elev. 10000 ft. 45.740935°, -118.233709°; uncertainty: 2000 Cliffs. Phenology: Spores. Origin: Native. m., Georef'd by Joseph Stormer Phenology: Fertile. Origin: Native. W. C. Cusick 3390a Lilla Leach 2643 16 Aug 1909 13 May 1930 WTU-147049, WTU-V-021052 WTU-266271, WTU-V-18435

Dryopteridaceae Dryopteridaceae Polystichum imbricans (D.C. Eaton) D.H. Wagner ssp. Polystichum munitum (Kaulf.) C. Presl imbricans U.S.A., ALASKA COUNTY: U.S.A., WASHINGTON, CHELAN COUNTY: Baranof island, southeast side. Northeast part of Chelan National Forest, Valley of Purple Creek. Port Armstrong just west of the site of old whale Sawtooth Ridge. station. Tongass National Forest. Southeastern Elev. 2000 ft. Alaska. 48.320776°, -120.606623°; WGS84, Spruce-hemlock forest just back from beach. uncertainty: 10000 m., Source: GeoLocate, Well drained rocky soil. Mossy forest floor. Georef'd by WTU Staff Common. Phenology: Fertile. Origin: Native. Granite ledges. Phenology: Spores. Origin: Native. Mary Clay Muller 4623 Harold St. John 3756 16 Aug 1981 with W. D. Courtney, Charles S. Parker WTU-282675, WTU-V-18402 24 Jul 1921 WTU-107920, WTU-V-016119 Dryopteridaceae Dryopteridaceae Polystichum lonchitis (L.) Roth Polystichum munitum (Kaulf.) C. Presl U.S.A., WASHINGTON COUNTY: U.S.A., ALASKA COUNTY: Goat Mountains. Baranof Island, east side. Port Walter, about 2 Origin: Native. miles in from the mouth. North side, west end of narrows, west edge of rock slide. Tongrass J. B. Flett s.n. National Forest. Southeastern Alaska. Elev. 30 ft. 19 Aug 1903 Well drained mossy area, overlying jumbled rocky unstable soil. With P. braunii. Other WTU-42792, WTU-V-016236 associated plants: Aluns rubra, Sambucus ravemosa, Oplopanax harridum, and misc. herbs. Phenology: Fertile. Origin: Native. Mary Clay Muller 4630 17 Aug 1981 Dryopteridaceae WTU-282631, WTU-V-18434 Polystichum imbricans (D.C. Eaton) D.H. Wagner ssp. imbricans U.S.A., WASHINGTON, CHELAN COUNTY: Dryopteridaceae Lake Wenatchee, near Ranger Station. 47.42361°, -120.30917°; WGS84, uncertainty: Polystichum munitum (Kaulf.) C. Presl 10000 m., Source: GeoLocate, Georef'd by WTU Staff U.S.A., ALASKA COUNTY: Rocky slopes. Phenology: Spores. Origin: Washington Bay, Kuiu Island. Native. 56.721111°, -134.356907°; WGS 84, uncertainty: 2400 m., Source: Digital Map, J. William Thompson 6798 Georef'd by Joseph Stormer In shady spruce forest. Phenology: Spores. 15 Jun 1931 Origin: Native. WTU-5347JWT, WTU-V-016171 W. J. Eyerdam 5371 20 Jul 1947 Dryopteridaceae WTU-293243, WTU-V-18286 Polystichum lonchitis (L.) Roth U.S.A., WASHINGTON, SKAMANIA COUNTY: Little White Salmon River. Dryopteridaceae 45.720881°, -121.640615°; WGS 84, uncertainty: 10000 m., Source: GeoLocate, Polystichum lonchitis (L.) Roth Georef'd by WTU Staff Phenology: Spores. Origin: Native. U.S.A., IDAHO, BEAR LAKE COUNTY: West point of cirque wall on east facing slope, Wilhelm N. Suksdorf 5433 Bloomington Lake. 42.19194°, -111.40056°; WGS 84, 11 Aug 1886 uncertainty: 10000 m., Source: GeoLocate, Georef'd by WTU Staff WTU-268968, WTU-V-021132 Phenology: Spores. Origin: Native. E. J. Larrison s.n. with M. Jollie Dryopteridaceae 10 Jul 1949 Polystichum lonchitis (L.) Roth WTU-297558, WTU-V-021134 U.S.A., ALASKA, KENAI PENINSULA COUNTY: Head of Rocky Creek Valey near Kenai Lake. Elev. 3000 ft. 60° 22' N, 149° 21' W Quite common on alpine slopes above and just below tree line. Phenology: Spores. Origin: Native. J. A. Calder 5189 11 Jun 1951 WTU-173335, WTU-V-021048 Dryopteridaceae Dryopteridaceae Polystichum munitum (Kaulf.) C. Presl Polystichum munitum (Kaulf.) C. Presl U.S.A., OREGON, CLACKAMAS COUNTY: U.S.A., WASHINGTON, GRAYS HARBOR Slopes near Paradise Park, Mt. Hood. COUNTY: Elev. 5000 ft. Near Montesano. 45.3369°, -121.77358°; WGS 84, uncertainty: Elev. 200 ft. 3676 m., Source: Digital Map, Georef'd by 46.98139°, -123.60139°; Source: Inferred Joseph Stormer from duplicate specimen, Georef'd by Ed Rocky Slopes. Phenology: Fertile. Origin: Alverson Native. Phenology: Spores. Origin: Native. J. William Thompson 3410 A. A. Heller 4035 with E. Gertrude Heller 4 Aug 1927 11 Jul 1898 WTU-5363JWT, WTU-V-18259 WTU-82932, WTU-V-18306 Dryopteridaceae Dryopteridaceae Polystichum munitum (Kaulf.) C. Presl Polystichum munitum (Kaulf.) C. Presl U.S.A., OREGON, UMATILLA COUNTY: U.S.A., OREGON, TILLAMOOK COUNTY: Umatilla River, Bingham Springs; western Blue Along Sand Creek, 4-5 miles northwest of Mountains. Hemlock. Elev. 760 ft. T3S R10W S4 S; NAD 27, uncertainty: 805 45.740935°, -118.233709°; uncertainty: 2000 m., Source: TRS2LL, Georef'd by Ben Legler m., Georef'd by Joseph Stormer With Deer Fern under Tsuga and Thuja forest. Basaltic soil, thick forests. Phenology: Spores. Very common. Phenology: Fertile. Origin: Origin: Native. Native. W. C. Cusick 3285 Albert N. Steward 6622 with John E. Smith 16 Jul 1908 2 Jan 1954 WTU-147051, WTU-V-18428 WTU-155456, WTU-V-18263

Dryopteridaceae Dryopteridaceae Polystichum munitum (Kaulf.) C. Presl Polystichum scopulinum (D.C. Eaton) Maxon CANADA, BRITISH COLUMBIA: U.S.A., IDAHO, BUTTE COUNTY: Kennedy river area near Ucluelet. Drainage Walls inside the Great Rift, just north of the Kennedy river. Watchman, Craters of the Moon National Elev. 600 ft. Monument. Forest floor of ceadar and hemlock. Exp.: 43.41667°, -113.51667°; WGS 84, southwest. Slope: .5%. Phenology: Fertile. uncertainty: 10000 m., Source: GeoLocate, Georef'd by WTU Staff A. Young 65 Phenology: Spores. Origin: Native. with W. Hubbard William H. Baker 14131 28 May 1963 24 Jun 1956 WTU-221384, WTU-V-18261 WTU-299576, WTU-V-021113

Dryopteridaceae Dryopteridaceae Polystichum munitum (Kaulf.) C. Presl Polystichum lonchitis (L.) Roth CANADA, BRITISH COLUMBIA: Chert and phyllite slate talus and outcrop above U.S.A., WASHINGTON, MASON COUNTY: Fraser River, at Hicks, 2 miles below Boston Mt. Ellinor. Bar. Elev. 4920 ft. Patchy Douglas Fir forest and talus/outcrop site 47.52205°, -123.26061°; WGS 84, (treeless); west-facing, rather dry. Phenology: uncertainty: 500 m., Source: GeoLocate, Spores. Georef'd by WTU Staff Amongst boulders in a mountain meadow. A. R. Kruckeberg 5749 Phenology: Spores. Origin: Native. 10 Sep 1963 W. J. Eyerdam 1266 WTU-313561, WTU-V-18287 14 Aug 1937 WTU-291623, WTU-V-016248 Dryopteridaceae Dryopteridaceae Polystichum munitum (Kaulf.) C. Presl Polystichum munitum (Kaulf.) C. Presl CANADA, BRITISH COLUMBIA: U.S.A., WASHINGTON, JEFFERSON One mile east of Bella Coola. COUNTY: Steep slope in coniferous woods. Very common. Hoh River bottoms near Nolan Creek. Phenology: Fertile. Elev. 100 ft. 47.750312°, -124.340519°; WGS 84, J. A. Calder 18444 uncertainty: 5000 m., Source: GeoLocate, with J. A. Parmelee, R. L. Taylor Georef'd by WTU Staff Phenology: Spores. Origin: Native. 6 Jul 1946 I. C. Otis 1315 WTU-173799, WTU-V-18411 19 Jul 1924 WTU-8119JWT, WTU-V-18424 Dryopteridaceae Dryopteridaceae Polystichum munitum (Kaulf.) C. Presl Polystichum munitum (Kaulf.) C. Presl U.S.A., WASHINGTON, JEFFERSON COUNTY: U.S.A., OREGON, TILLAMOOK COUNTY: Hoh River. bottoms below Big Bend. Slopes along the Wilson River, 22 miles above Elev. 200 ft. Tillamook. T26N R12W S4; Source: Inferred from Rocky slopes along river. Phenology: Fertile. duplicate specimen, Georef'd by Ed Alverson Origin: Native. Phenology: Spores. Origin: Native. J. William Thompson 3125 I. C. Otis 1294 22 Jul 1927 29 Jun 1924 WTU-5361JWT, WTU-V-18407 WTU-28641, WTU-V-18426

Dryopteridaceae Dryopteridaceae Polystichum scopulinum (D.C. Eaton) Maxon Polystichum lonchitis (L.) Roth U.S.A., IDAHO, CLEARWATER COUNTY: Powerhouse, north of Pierce. U.S.A., IDAHO, VALLEY COUNTY: 46.49111°, -115.79806°; WGS 84, Trail to Snowslide Lake from Lick Cr Road, 10 uncertainty: 10000 m., Source: GeoLocate, airmiles northeast of McCall. Georef'd by WTU Staff Elev. 7000 ft. Phenology: Vegetative. Origin: Native. T19N R4E S14 NW; WGS 84, uncertainty: 10000 m., Source: GeoLocate, Georef'd by R. M. Joslyn 246-249 WTU Staff Under granite rocks and shrubs of dry streamlet 2 Jul 1946 near lake; local in deep shade. Phenology: Spores. WTU-299381, WTU-V-021111 Barbara Ertter 7779 with Susan D’Alcamo, Cristopher Davidson Dryopteridaceae 21 Jul 1988 Polystichum munitum (Kaulf.) C. Presl WTU-318439, WTU-V-021138 CANADA, BRITISH COLUMBIA: Vancouver Island; Kennedy River area near Dryopteridaceae Ucluelet. Kennedy River drainage. Elev. 600 ft. Polystichum munitum (Kaulf.) C. Presl Forest floor of ceadar and hemlock. Exp.: southwest. Slope: .5%. Phenology: Fertile. CANADA, BRITISH COLUMBIA: Just southf of Skidegate Mission on road to A. Young 65 Queen Charlotte City, Graham Island, Queen with W. Hubbard Charlotte Islands. Slopes and bluffs in coniferous woods back from 28 May 1963 shoreline. Common. Phenology: Fertile. WTU-218838, WTU-V-18409 J. A. Calder 21405 with D. B. O. Savile, R. L. Taylor 11 Jun 1957 WTU-239367, WTU-V-18420 Dryopteridaceae Dryopteridaceae Polystichum munitum (Kaulf.) C. Presl Polystichum munitum (Kaulf.) C. Presl U.S.A., WASHINGTON, SAN JUAN COUNTY: U.S.A., WASHINGTON, SNOHOMISH Friday Harbor. COUNTY: 48.541389°, -123.0225°; WGS 84, 205th and Alaska Way, Snohomish. uncertainty: 2000 m., Source: Georeferenced, 47.91822°, -122.08872°; WGS 84, Georef'd by WTU Staff uncertainty: 2000 m., Source: GeoLocate, Phenology: Spores. Origin: Native. Georef'd by WTU Staff In the transition from deciduous forest with A. S. Pope s.n. Rubus undergrowth to evergreen forest with mostly fern and small herb undergrowth. 25 Jun 1904 Phenology: Spores. Origin: Native. WTU-2024, WTU-V-18422 Christie S. Isler 34-5-98 16 May 1998 Dryopteridaceae WTU-335310, WTU-V-18324 Polystichum munitum (Kaulf.) C. Presl Dryopteridaceae CANADA, BRITISH COLUMBIA: Boat Basin, Vancouver Island. Polystichum munitum (Kaulf.) C. Presl 49° N, 126° W; Misleading coordinates given only to nearest degree. U.S.A., OREGON, MULTNOMAH COUNTY: Coniferous forest. Phenology: Fertile. Portland. 45.52361°, -122.675°; WGS 84, uncertainty: A. F. Szczawinski s.n. 12756 m., Source: GeoLocate, Georef'd by Maria Yousoufian 27 Jun 1961 Rocky Banks. Phenology: Fertile. Origin: Native. WTU-203798, WTU-V-18405 B.S.S. s.n. 18 Aug 1889 WTU-2000, WTU-V-18421 Dryopteridaceae Polystichum munitum (Kaulf.) C. Presl Dryopteridaceae U.S.A., WASHINGTON, JEFFERSON COUNTY: Polystichum lonchitis (L.) Roth Hoh River bottoms near Nolan Creek. Elev. 100 ft. U.S.A., WASHINGTON, SNOHOMISH 47.750312°, -124.340519°; WGS 84, COUNTY: uncertainty: 5000 m., Source: GeoLocate, Mount Dickerman, Cascade Mountains. Georef'd by WTU Staff Elev. 4000 ft. Phenology: Spores. Origin: Native. 48.06829°, -121.47068°; WGS 84, uncertainty: 3000 m., Source: GeoLocate, I. C. Otis 1315 Georef'd by WTU Staff Crevices of cliffs. Phenology: Spores. 19 Jul 1924 J. William Thompson 8800 WTU-28642, WTU-V-18406 17 Jul 1932 Dryopteridaceae WTU-11515JWT, WTU-V-021083 Polystichum scopulinum (D.C. Eaton) Maxon Dryopteridaceae U.S.A., MONTANA, SANDERS COUNTY: [locality and coordinates withheld; status in Polystichum lonchitis (L.) Roth Montana: S1S2 (Imperiled; MTNHP)] Elev. 6500 ft. U.S.A., WASHINGTON, CLALLAM COUNTY: Rock crevice; with Cryptogramma crispa, East face of Mount Angeles, Olympic National Parnassia, Tofieldia, Polemonium pulcherimum. Park. Phenology: Spores. 48.26667°, -124.675°; WGS 84, uncertainty: 10000 m., Source: GeoLocate, Georef'd by Christopher Clampitt 150 WTU Staff Steep, cool, shaded draw. Associated with 18 Aug 1985 Polypodium vulgare and Cryptogramma achrostichoides. Phenology: Spores. Origin: WTU-313257, WTU-V-021109 Native. A. R. Kruckeberg 5675 1 Aug 1963 WTU-214836, WTU-V-016259 Dryopteridaceae Dryopteridaceae Polystichum lonchitis (L.) Roth Polystichum lonchitis (L.) Roth U.S.A., WASHINGTON COUNTY: U.S.A., WASHINGTON, CHELAN COUNTY: Olympics. Mount Stuart, Wenatchee Mountains. 47.811111°, -123.626944°; Source: Inferred Elev. 6000 ft. from duplicate specimen, Georef'd by Ed 47.47522°, -120.90238°; WGS84, uncertainty: Alverson 2000 m., Source: Georeferenced, Georef'd by Phenology: Spores. WTU Staff Most crevices of cliffs. The collection date is J. B. Flett 915 from 27 to 31 of July. Phenology: Spores. with C.R.P. J. William Thompson 77673 WTU-2040, WTU-V-021076 27 Jul 1931 WTU-11511JWT, WTU-V-016260 Dryopteridaceae Polystichum lonchitis (L.) Roth Dryopteridaceae U.S.A., WASHINGTON, WHATCOM COUNTY: Polystichum lonchitis (L.) Roth Below Austin Pass, Mount Baker National Forest. U.S.A., WASHINGTON, CHELAN COUNTY: Elev. 3000 ft. Upper Jack Creek, 3 miles Northwest of Stuart 48.85194°, -121.68389°; WGS 84, Pass. uncertainty: 10000 m., Source: GeoLocate, 47.515562°, -120.950439°; WGS84, Georef'd by WTU Staff uncertainty: 1000 m., Source: GeoLocate, Crevices and rocky slopes. Phenology: Spores. Georef'd by WTU Staff At base of "granitoid" rock talus and outcropping J. William Thompson 8008 in otherwise massive peridotite area. Phenology: Spores. Origin: Native. 19 Aug 1931 A. R. Kruckeberg 5135 WTU-11535JWT, WTU-V-021067 29 Jul 1960 Dryopteridaceae WTU-218779, WTU-V-016257 Polystichum lonchitis (L.) Roth U.S.A., WASHINGTON, JEFFERSON Dryopteridaceae COUNTY: Above Lake Constance; Olympic Mountains. Polystichum lonchitis (L.) Roth Elev. 6000 ft. 47.87222°, -123.66667°; WGS 84, U.S.A., WASHINGTON, GRAYS HARBOR uncertainty: 10000 m., Source: GeoLocate, COUNTY: Georef'd by WTU Staff Near summit of Mount Colonel Bob, Olympic Crevices of cliffs. Phenology: Spores. Origin: Mountains. Native. Elev. 4700 ft. 47.48889°, -123.74556°; WGS 84, J. William Thompson 7895 uncertainty: 10000 m., Source: GeoLocate, Georef'd by WTU Staff 11 Aug 1931 Crevices of cliffs. Phenology: Spores. WTU-11138JWT, WTU-V-021066 J. William Thompson 7254 9 Jul 1931 Dryopteridaceae WTU-11536JWT, WTU-V-016258 Polystichum lonchitis (L.) Roth U.S.A., WASHINGTON, YAKIMA COUNTY: Dryopteridaceae Mount Adams. 46.20255°, -121.4908°; WGS 84, uncertainty: Polystichum lonchitis (L.) Roth 8000 m., Source: GeoLocate, Georef'd by WTU Staff U.S.A., WASHINGTON, CHELAN COUNTY: Damp, shady rocks, "wiesehalde". Phenology: Mount Stuart, Wenatchee Mountains. Spores. Origin: Native. Elev. 8000 ft. 47.47522°, -120.90238°; WGS84, uncertainty: Wilhelm N. Suksdorf 4568 2000 m., Source: Georeferenced, Georef'd by WTU Staff 5 Oct 1902 Crevices of cliffs. Phenology: Spores. WTU-268971, WTU-V-021081 J. William Thompson 5782 20 Aug 1930 WTU-11513JWT, WTU-V-016255 Dryopteridaceae Dryopteridaceae Polystichum lonchitis (L.) Roth Polystichum munitum (Kaulf.) C. Presl U.S.A., WASHINGTON, WHATCOM COUNTY: U.S.A., WASHINGTON, CLALLAM COUNTY: South slope of Mount Hermann, mount Baker Olympic Mountains. Mount Angeles. Lodge. Elev. 5000 ft. Elev. 4500 ft. 47.995278°, -123.474444°; NAD 27, 48.86361°, -121.68111°; WGS 84, uncertainty: 1000 m., Source: Georeferenced, uncertainty: 10000 m., Source: GeoLocate, Georef'd by WTU Staff Georef'd by WTU Staff Phenology: Spores. Origin: Native. Crevices of cliffs. The collection date should be from 22 to 25 of July. Phenology: Spores. J. William Thompson 7503 Origin: Native. 18 Jul 1931 J. William Thompson 5253 WTU-5372JWT, WTU-V-18418 22 Jul 1930 WTU-11514JWT, WTU-V-021079 Dryopteridaceae Polystichum munitum (Kaulf.) C. Presl Dryopteridaceae U.S.A., WASHINGTON, SNOHOMISH Polystichum lonchitis (L.) Roth COUNTY: Mt. Dickerman; talus above Perry Creek. U.S.A., WASHINGTON, WHATCOM COUNTY: Elev. 3000 ft. Alpine slopes around Twin Lakes, Cascade 48.063294°, -121.513675°; WGS 84, Mountains. uncertainty: 10000 m., Source: GeoLocate, Elev. 5000 ft. Georef'd by WTU Staff 48.95194°, -121.63444°; WGS 84, Shaded mossy talus. Abundant with P. uncertainty: 5000 m., Source: GeoLocate, andersonii and hybrid. Phenology: Spores. Georef'd by WTU Staff Origin: Native. Rock crevices. Phenology: Spores. W. H. Wagner 63114-2 J. William Thompson 17212 with A. R. Kruckeberg 21 Aug 1952 28 Jun 1963 WTU-154967, WTU-V-016256 WTU-213565, WTU-V-18416 Dryopteridaceae Dryopteridaceae Polystichum munitum (Kaulf.) C. Presl Polystichum munitum (Kaulf.) C. Presl U.S.A., WASHINGTON, CLALLAM COUNTY: Mount Angeles. U.S.A., WASHINGTON, SKAMANIA COUNTY: Elev. 2500 ft. Below Flattop Mountain; 25 miles west of 47.99538°, -123.46784°; WGS 84, Glenwood. uncertainty: 2500 m., Source: GeoLocate, Elev. 3148 ft. Georef'd by WTU Staff 46.02852°, -121.61425°; WGS 84, Phenology: Spores. Origin: Native. uncertainty: 500 m., Source: GeoLocate, Georef'd by WTU Staff G. B. Rigg s.n. Phenology: Spores. Origin: Native. 22 Jul 1929 E. Hunn 651 WTU-39451, WTU-V-18414 2 Oct 1977 WTU-271158, WTU-V-18289 Dryopteridaceae Polystichum munitum (Kaulf.) C. Presl Dryopteridaceae U.S.A., WASHINGTON, MASON COUNTY: Polystichum munitum (Kaulf.) C. Presl Twanoh State Park, Hood Canal. 47.37345°, -122.97428°; WGS 84, U.S.A., WASHINGTON, JEFFERSON uncertainty: 1000 m., Source: GeoLocate, COUNTY: Georef'd by WTU Staff Along Hoh River near Jackson Ranger Station. Phenology: Spores. Origin: Native. 47.814011°, -124.176698°; WGS 84, uncertainty: 10000 m., Source: GeoLocate, Harold W. Smith 182 Georef'd by WTU Staff Phenology: Spores. Origin: Native. 21 Sep 1936 I. C. Otis 1363 WTU-38849, WTU-V-18410 6 Aug 1924 WTU-8124JWT, WTU-V-18316 Dryopteridaceae Dryopteridaceae Polystichum munitum (Kaulf.) C. Presl Polystichum munitum (Kaulf.) C. Presl U.S.A., WASHINGTON, KING COUNTY: U.S.A., WASHINGTON COUNTY: 3 miles east of Kirkland. Garden of C. L. Hitchcock. 47.681652°, -122.143002°; WGS 84, Narrow form. Phenology: Spores. Origin: Native. uncertainty: 1000 m., Source: GeoLocate, Georef'd by WTU Staff W. H. Wagner 63124 Western Sword Fern. Phenology: Spores. Origin: Native. 27 Jun 1963 Anita Watson 2 WTU-213551, WTU-V-18408 28 Jul 1949 WTU-165220, WTU-V-18412

Dryopteridaceae Dryopteridaceae Polystichum munitum (Kaulf.) C. Presl Polystichum imbricans (D.C. Eaton) D.H. Wagner × Polystichum munitum (Kaulf.) Presl U.S.A., WASHINGTON, KING COUNTY: South Seattle. U.S.A., WASHINGTON, CHELAN COUNTY: 47.60639°, -122.33083°; WGS 84, Wenatchee lake Quadrangle; 1.5 to 2 miles up uncertainty: 10000 m., Source: GeoLocate, Rock Mountian Trail. Georef'd by WTU Staff Elev. 4400 ft. Moist hillsides. Phenology: Spores. Origin: T26N R15E S3 NE 1/4; WGS84, uncertainty: Native. 10000 m., Source: GeoLocate, Georef'd by WTU Staff J. William Thompson 5894 Growing with Ceanothus velutinus, Acer glabrum and Sorbus scopulina. A few 1 Feb 1931 Pseudotsuga menziesii nearby. Phenology: Spores. Origin: Native. WTU-5373JWT, WTU-V-18419 E. Burnett 291 12 Nov 1990 Dryopteridaceae WTU-339781, WTU-V-030980 Polystichum munitum (Kaulf.) C. Presl U.S.A., WASHINGTON, KING COUNTY: South Seattle. Dryopteridaceae 47.60639°, -122.33083°; WGS 84, uncertainty: 10000 m., Source: GeoLocate, Polystichum imbricans (D.C. Eaton) D.H. Wagner × Georef'd by WTU Staff Polystichum munitum (Kaulf.) Presl Wooded hillsides. Phenology: Spores. Origin: Native. U.S.A., WASHINGTON, CHELAN COUNTY: Wenatchee lake Quadrangle; 1.5 to 2 miles up J. William Thompson 5180 Rock Mountian Trail. Elev. 4400 ft. Oct 1929 T26N R15E S3 NE 1/4; WGS84, uncertainty: 10000 m., Source: GeoLocate, Georef'd by WTU-5348JWT, WTU-V-18417 WTU Staff Growing with Ceanothus velutinus, Acer glabrum and Sorbus scopulina. A few Pseudotsuga menziesii nearby. Phenology: Dryopteridaceae Spores. Origin: Native. Polystichum munitum (Kaulf.) C. Presl E. Burnett 292 U.S.A., WASHINGTON, KING COUNTY: 12 Nov 1990 Seattle. 47.613617°, -122.310098°; WGS 84, WTU-339786, WTU-V-030983 uncertainty: 15750 m., Source: Digital Map, Georef'd by Joseph Stormer Phenology: Spores. Origin: Native. Emma J. Cole 49155 14 Jul 1905 WTU-266664, WTU-V-18315 Dryopteridaceae Dryopteridaceae Polystichum munitum (Kaulf.) C. Presl Polystichum munitum (Kaulf.) C. Presl U.S.A., WASHINGTON, KING COUNTY: U.S.A., WASHINGTON, CLALLAM COUNTY: Seattle. Lake Crescent. 47.613617°, -122.310098°; WGS 84, 48.09472°, -123.80583°; WGS 84, uncertainty: 10000 m., Source: uncertainty: 15750 m., Source: Digital Map, GeoLocate, Georef'd by WTU Staff Georef'd by Joseph Stormer Phenology: Spores. Origin: Native. Young plant. Phenology: Spores. Origin: Native. G. N. Jones 3483 Jul 1931 Eleanor Blodgett s.n. WTU-21393 20 Jun 1904 WTU-2034, WTU-V-18415 Dryopteridaceae Polystichum munitum (Kaulf.) C. Presl Dryopteridaceae U.S.A., IDAHO COUNTY: Coeur D'Alene Mountains; Forks of St. Mary's River. Polystichum munitum (Kaulf.) C. Presl Elev. 4756 ft. Phenology: Spores. Origin: Native. U.S.A., WASHINGTON, KING COUNTY: Seattle. John B. Leiberg 1147 3 Jul 1895 47.613617°, -122.310098°; WGS 84, uncertainty: 15750 m., Source: Digital Map, WTU-8123JWT Georef'd by Joseph Stormer Phenology: Spores. Origin: Native. Dryopteridaceae Emma A. Shumway s.n. Polystichum munitum (Kaulf.) C. Presl Oct 1889 U.S.A., WASHINGTON, KING COUNTY: WTU-90305, WTU-V-18413 Lake Forest Park, Seattle. 47.75694°, -122.27972°; WGS 84, uncertainty: 10000 m., Source: GeoLocate, Georef'd by WTU Staff Dryopteridaceae Phenology: Spores. Origin: Native. Polystichum munitum (Kaulf.) C. Presl G. L. Wittrock s.n. U.S.A., WASHINGTON, JEFFERSON 12 Oct 1927 COUNTY: Olympic National Park: Royal Creek Trail; circa WTU-31285, WTU-V-18297 2 miles below Royal Basin & Lake. 47.854841°, -123.214107°; Source: Inferred from duplicate specimen, Georef'd by Ed Alverson Dryopteridaceae Seep in coniferous woods, with Polystichum lonchitis. Possible hybridity?. Phenology: Polystichum munitum (Kaulf.) C. Presl Spores. Origin: Native. U.S.A., WASHINGTON, PIERCE COUNTY: A. R. Kruckeberg s.n. Tacoma. 47.25306°, -122.44306°; Source: Inferred from duplicate 8 Oct 1979 specimen, Georef'd by Ed Alverson Moist woods. Phenology: Spores. Origin: Native. WTU-305224, WTU-V-18299 J. B. Flett s.n. 1897 Dryopteridaceae WTU-39472 Polystichum munitum (Kaulf.) C. Presl U.S.A., WASHINGTON, JEFFERSON Dryopteridaceae COUNTY: Hoh River valley near Nolan Creek. Polystichum munitum (Kaulf.) C. Presl Elev. 100 ft. 47.750312°, -124.340519°; WGS 84, U.S.A., WASHINGTON, KING COUNTY: uncertainty: 5000 m., Source: GeoLocate, Seattle and Western Washington. Georef'd by WTU Staff 47.60639°, -122.33083°; WGS 84, uncertainty: 10000 m., Source: Phenology: Spores. Origin: Native. GeoLocate, Georef'd by WTU Staff Moist dark woods. Common; Wood Sheild-Fern. Phenology: Spores. I. C. Otis 1371 Origin: Native. 14 Aug 1924 Louis F. Henderson s.n. 22 Jul 1892 WTU-8122JWT, WTU-V-18298 WTU-2030 Dryopteridaceae Dryopteridaceae Polystichum munitum (Kaulf.) C. Presl Polystichum munitum (Kaulf.) C. Presl U.S.A., WASHINGTON, JEFFERSON COUNTY: U.S.A., ALASKA, KETCHIKAN GATEWAY COUNTY: Port Discovery. Ketchikan. 48.01556°, -122.86056°; WGS 84, uncertainty: 10000 m., Source: 55.342222°, -131.646111°; WGS 84, uncertainty: 7400 m., GeoLocate, Georef'd by WTU Staff Source: GeoLocate, Georef'd by Joseph Stormer Evergreen woods. Phenology: Spores. Origin: Native. Rocky seashore-shade. Phenology: Spores. Origin: Native. Harold St. John 5849 12 Sep 1921 J. B. Flett 1912 8 Aug 1901 WTU-108000 WTU-37583

Dryopteridaceae Dryopteridaceae Polystichum imbricans (D.C. Eaton) D.H. Wagner ssp. Polystichum munitum (Kaulf.) C. Presl imbricans U.S.A., WASHINGTON, JEFFERSON CANADA, BRITISH COLUMBIA: COUNTY: Little Saanich Mountain, Vancouver Island. Olympic Peninsula; Duckabush River. Phenology: Spores. Origin: Native. Elev. 1000 ft. 47.648433°, -122.933459°; WGS 84, J. R. Anderson s.n. uncertainty: 10000 m., Source: GeoLocate, Georef'd by WTU Staff 20 Oct 1917 Open woods. Phenology: Spores. Origin: Native. WTU-86707, WTU-V-016195 Geo. N. Jones 3067 Jun 1931 WTU-21390, WTU-V-18296 Dryopteridaceae Dryopteridaceae Polystichum munitum (Kaulf.) C. Presl Polystichum imbricans (D.C. Eaton) D.H. Wagner ssp. imbricans U.S.A., IDAHO, CLEARWATER COUNTY: Near east fork of Potlatch Creek; 10 miles east of Bovill. U.S.A., WASHINGTON, CHELAN COUNTY: Elev. 3300 ft. Power house between Leavenworth and T41N R2E S20; WGS 84, uncertainty: 10000 m., Source: Tumwater. GeoLocate, Georef'd by WTU Staff Elev. 1350 ft. On north slopes in mixed coniferous woods. In large clumps from a 47.59639°, -120.66028°; WGS84, uncertainty: compact caudex. Phenology: Spores. Origin: Native. 10000 m., Source: GeoLocate, Georef'd by WTU Staff Arthur Cronquist 5991 24 Jul 1949 Phenology: Spores. Origin: Native. with Quentin Jones I. C. Otis 1058 WTU-168988 28 Sep 1920 Dryopteridaceae WTU-8125JWT, WTU-V-016198 Polystichum munitum (Kaulf.) C. Presl U.S.A., OREGON, DOUGLAS-COOS COUNTY: Dryopteridaceae Coast Mountains. Coniferous forests. Very common. Phenology: Spores. Origin: Polystichum munitum (Kaulf.) C. Presl Native. U.S.A., WASHINGTON, JEFFERSON W. C. Cusick 4302 8 Jul 1914 COUNTY: Along Hoh River near Jackson Ranger Station. WTU-147050 47.814011°, -124.176698°; WGS 84, uncertainty: 10000 m., Source: GeoLocate, Dryopteridaceae Georef'd by WTU Staff Phenology: Spores. Origin: Native. Polystichum munitum (Kaulf.) C. Presl I. C. Otis 1363 U.S.A., WASHINGTON, KING COUNTY: Seattle. 6 Aug 1924 47.613617°, -122.310098°; WGS 84, uncertainty: 15750 m., Source: Digital Map, Georef'd by Joseph Stormer WTU-28650, WTU-V-18295 Common name: Sheild Fern. Phenology: Spores. Origin: Native. C. V. Piper 336 16 Sep 1888 WTU-2025 Dryopteridaceae Dryopteridaceae Polystichum munitum (Kaulf.) C. Presl Polystichum munitum (Kaulf.) C. Presl U.S.A., IDAHO COUNTY: U.S.A., OREGON, MARION COUNTY: Above Ahsahka. [No locality given on label] Pine woods. Phenology: Spores. Origin: Native. Phenology: Spores. Origin: Native. F. L. Pickett 588 18 Aug 1916 O. B. Johnson s.n. Jul 1880 WTU-67163 WTU-1990

Dryopteridaceae Dryopteridaceae Polystichum munitum (Kaulf.) C. Presl Polystichum munitum (Kaulf.) C. Presl U.S.A., WASHINGTON, PIERCE COUNTY: U.S.A., IDAHO, CLEARWATER COUNTY: Near Puyallup. Little North Fork of Clearwater River, Clearwater Forest Reserve. 47.18556°, -122.29167°; WGS 84, uncertainty: 5000 m., Source: In dense white pine forest. Common and abundant. Phenology: GeoLocate, Georef'd by WTU Staff Spores. Origin: Native. Wooded areas in shade. Forming dense clumps. Phenology: Spores. Origin: Native. F. L. Pickett 1128 11 Aug 1917 C. L. Porter 4214 13 Jun 1947 WTU-66023 WTU-122117 Dryopteridaceae Dryopteridaceae Polystichum munitum (Kaulf.) C. Presl Polystichum munitum (Kaulf.) C. Presl CANADA, BRITISH COLUMBIA: Queen Charlotte Islands; Masset, Graham U.S.A., OREGON, MULTNOMAH COUNTY: Island. Back of Crown Point, near Corbett. 54.016667°, -132.15°; WGS 84, uncertainty: Woods. Phenology: Spores. Origin: Native. 6029 m., Source: GeoLocate, Georef'd by Joseph Stormer J. William Thompson 3009 Phenology: Spores. Origin: Native. 17 Jul 1927 Edward Woolman s.n. WTU-5365JWT, WTU-V-18404 Aug 1930 WTU-56830, WTU-V-18395

Dryopteridaceae Dryopteridaceae Polystichum munitum (Kaulf.) C. Presl Polystichum imbricans (D.C. Eaton) D.H. Wagner ssp. U.S.A., IDAHO, IDAHO COUNTY: imbricans Lowell-Kooskia road, 3 miles west of Lowell. T32N R6E S1; WGS 84, uncertainty: 1000 m., Source: U.S.A., OREGON, KLAMATH COUNTY: GeoLocate, Georef'd by WTU Staff Southwest corner, north wall, Red Blanket In seepage from spring, in dense Thuja stand. Phenology: Spores. Canyon. Crater Lake National Park. Origin: Native. 42.784045°, -122.367427°; WGS 84, uncertainty: 2500 m., Source: Digital Map, Quentin Jones 46 16 Jun 1950 Georef'd by Joseph Stormer, Invalid coordinates (located outside county bounds). WTU-142736 Phenology: Spores. Origin: Native. William H. Baker 7104 Dryopteridaceae 29 Jul 1950 Polystichum munitum (Kaulf.) C. Presl WTU-294304, WTU-V-016197 U.S.A., IDAHO, IDAHO COUNTY: Lochsa Valley. No collection date given. Phenology: Spores. Origin: Native. Dryopteridaceae Fred A. Barkley 1859 Polystichum munitum (Kaulf.) C. Presl WTU-41522, WTU-V-18396 U.S.A., ALASKA COUNTY: Southeastern Alaska. Coastal areas. Phenology: Spores. Origin: Native. Mr. Ernest P. Walker 1083 Jun 1916 with Mrs. Ernest P. Walker WTU-74627 Dryopteridaceae Dryopteridaceae Polystichum munitum (Kaulf.) C. Presl Polystichum andersonii M. Hopkins × Polystichum munitum (Kaulf.) C. Presl U.S.A., OREGON, HOOD RIVER COUNTY: Cascades; along the Loop Highway in the Mount Hood National U.S.A., WASHINGTON, SNOHOMISH Forest. COUNTY: 45.353641°, -121.590584°; WGS 84, uncertainty: 11873 m., Perry Creek drainage; circa 0.5 miles above Source: Digital Map, Georef'd by Joseph Stormer trailhead and northeast end of Forest Service West slope. Spores collected by A. R. C. October 7, 1980. Road 4063; northwest slope of Dickerman Phenology: Spores. Origin: Native. Mountain. West slope of Cascade Mountains, Mt. Baker-Snoqualmie National Forest. William A. Weber 1973 7 Feb 1941 Elev. 2197 ft. 48° 4.4' N, 121° 29.5' W; T30N R10E S23 WTU-67643 Shaded trailside; northwest aspect; steep mossy and rocky slope. With Blechnum spicant, Acer circinatum, Athyrium filix-femina, Botrychium Dryopteridaceae multifidum, Polystichum andersonii, P. munitum. Cepitose, rare, one plant seen. Fronds dark Polystichum munitum (Kaulf.) C. Presl green as in P. munitum, and more leathery than P. andersonii at this site. Lower pinnae less U.S.A., IDAHO, BONNER COUNTY: divided than P. andersonii, much more divided Preist River Exptl. Forest. than P. munitum. Sheet 1 of 3. Phenology: 48.35783°, -116.79602°; WGS 84, Spores. Origin: Native. uncertainty: 5000 m., Source: GeoLocate, Georef'd by WTU Staff Peter F. Zika 19084 20 Sep 2003 Douglas Fir vegetation zone; Community near-climax, open forest with dominant WTU-354256, WTU-V-021028 Pseudotsuga, Pinus monticola, Pinus ponderosa. Phenology: Spores. Origin: Native. Dryopteridaceae R. F. Daubenmire 44270 Polystichum andersonii M. Hopkins × Polystichum munitum (Kaulf.) C. Presl 10 Jun 1944 U.S.A., WASHINGTON, SNOHOMISH WTU-241062, WTU-V-18398 COUNTY: Perry Creek drainage; circa 0.5 miles above trailhead and northeast end of Forest Service Road 4063; northwest slope of Dickerman Dryopteridaceae Mountain. West slope of Cascade Mountains, Mt. Baker-Snoqualmie National Forest. Polystichum munitum (Kaulf.) C. Presl Elev. 2197 ft. 48° 4.4' N, 121° 29.5' W; T30N R10E S23 CANADA, BRITISH COLUMBIA: Shaded trailside; northwest aspect; steep mossy Vancouver Island; Elk River drainage area. and rocky slope. With Blechnum spicant, Acer Elev. 1100 ft. circinatum, Athyrium filix-femina, Botrychium In moss under Hemlock; Exp.-E, Slope-2. Sword Fern. Phenology: multifidum, Polystichum andersonii, P. munitum. Spores. Origin: Native. Cepitose, rare, one plant seen. Fronds dark green as in P. munitum, and more leathery than A. Young 610 10 Jul 1963 P. andersonii at this site. Lower pinnae less with W. Hubbard divided than P. andersonii, much more divided than P. munitum. Sheet 2 of 3. Phenology: WTU-219317 Spores. Origin: Native. Peter F. Zika 19084 20 Sep 2003 Dryopteridaceae WTU-354257, WTU-V-021029 Polystichum munitum (Kaulf.) C. Presl U.S.A., OREGON, TILLAMOOK COUNTY: Neskowin Creek. Phenology: Spores. Origin: Native. G. E. Morrill 221 25 Jul 1933 WTU-27689 Dryopteridaceae Dryopteridaceae Polystichum andersonii M. Hopkins × Polystichum Polystichum andersonii M. Hopkins × Polystichum munitum (Kaulf.) C. Presl munitum (Kaulf.) C. Presl U.S.A., WASHINGTON, SNOHOMISH U.S.A., WASHINGTON, SNOHOMISH COUNTY: COUNTY: Perry Creek drainage; circa 0.5 miles above Mt. Dickerman; talus above Perry Creek. trailhead and northeast end of Forest Service Elev. 3000 ft. Road 4063; northwest slope of Dickerman 48.063294°, -121.513675°; WGS 84, Mountain. West slope of Cascade Mountains, uncertainty: 10000 m., Source: GeoLocate, Mt. Baker-Snoqualmie National Forest. Georef'd by WTU Staff Elev. 2197 ft. Shaded mossy talus. About 8 plants, with 48° 4.4' N, 121° 29.5' W; T30N R10E S23 parents. Phenology: Spores. Origin: Native. Shaded trailside; northwest aspect; steep mossy and rocky slope. With Blechnum spicant, Acer W. H. Wagner 63115-13 circinatum, Athyrium filix-femina, Botrychium with A. R. Kruckeberg multifidum, Polystichum andersonii, P. munitum. Cepitose, rare, one plant seen. Fronds dark 28 Jun 1963 green as in P. munitum, and more leathery than P. andersonii at this site. Lower pinnae less WTU-213562, WTU-V-021022 divided than P. andersonii, much more divided than P. munitum. Sheet 3 of 3. Phenology: Spores. Origin: Native. Dryopteridaceae Peter F. Zika 19084 20 Sep 2003 Polystichum andersonii M. Hopkins × Polystichum munitum (Kaulf.) C. Presl WTU-354258, WTU-V-021027 U.S.A., WASHINGTON, SNOHOMISH Dryopteridaceae COUNTY: Mt. Dickerman; talus above Perry Creek. Polystichum andersonii M. Hopkins × Polystichum Elev. 3000 ft. munitum (Kaulf.) C. Presl 48.063294°, -121.513675°; WGS 84, uncertainty: 10000 m., Source: GeoLocate, U.S.A., WASHINGTON, SNOHOMISH Georef'd by WTU Staff COUNTY: Shaded mossy talus. About 8 plants, with Mt. Dickerman; talus above Perry Creek. parents. Phenology: Spores. Origin: Native. Elev. 3000 ft. 48.063294°, -121.513675°; WGS 84, W. H. Wagner 63115-18 uncertainty: 10000 m., Source: GeoLocate, with A. R. Kruckeberg Georef'd by WTU Staff Shaded mossy talus. About 8 plants, with 28 Jun 1963 parents. Phenology: Spores. Origin: Native. WTU-213559, WTU-V-021025 W. H. Wagner 63115-14 with A. R. Kruckeberg Dryopteridaceae 28 Jun 1963 Polystichum andersonii M. Hopkins × Polystichum WTU-213561, WTU-V-021026 munitum (Kaulf.) C. Presl U.S.A., WASHINGTON, SNOHOMISH Dryopteridaceae COUNTY: Mt. Dickerman; talus above Perry Creek. Polystichum andersonii M. Hopkins × Polystichum Elev. 3000 ft. munitum (Kaulf.) C. Presl 48.063294°, -121.513675°; WGS 84, uncertainty: 10000 m., Source: GeoLocate, U.S.A., WASHINGTON, SNOHOMISH Georef'd by WTU Staff COUNTY: Shaded mossy talus. About 8 plants, with Mt. Dickerman; talus above Perry Creek. parents. Phenology: Spores. Origin: Native. Elev. 3000 ft. 48.063294°, -121.513675°; WGS 84, W. H. Wagner 63115-12 uncertainty: 10000 m., Source: GeoLocate, with A. R. Kruckeberg Georef'd by WTU Staff Shaded mossy talus. About 8 plants, with 28 Jun 1963 parents. Phenology: Spores. Origin: Native. WTU-213563, WTU-V-021023 W. H. Wagner 63115-21 with A. R. Kruckeberg 28 Jun 1963 WTU-213558, WTU-V-021024 Dryopteridaceae Dryopteridaceae Polystichum andersonii M. Hopkins × Polystichum Polystichum andersonii M. Hopkins × Polystichum munitum (Kaulf.) C. Presl munitum (Kaulf.) C. Presl U.S.A., WASHINGTON, SNOHOMISH U.S.A., WASHINGTON, SNOHOMISH COUNTY: COUNTY: Mt. Dickerman; talus above Perry Creek. Mt. Dickerman; talus above Perry Creek. Elev. 3000 ft. Elev. 3000 ft. 48.063294°, -121.513675°; WGS 84, 48.063294°, -121.513675°; WGS 84, uncertainty: 10000 m., Source: GeoLocate, uncertainty: 10000 m., Source: GeoLocate, Georef'd by WTU Staff Georef'd by WTU Staff Shaded mossy talus. About 8 plants, with Shaded mossy talus. About 8 plants, with parents. Phenology: Spores. Origin: Native. parents. Phenology: Spores. Origin: Native. W. H. Wagner 63115-15 W. H. Wagner 63115-8 with A. R. Kruckeberg with A. R. Kruckeberg 28 Jun 1963 28 Jun 1963 WTU-213560, WTU-V-021021 WTU-213522, WTU-V-021018

Dryopteridaceae Dryopteridaceae Polystichum andersonii M. Hopkins × Polystichum Polystichum andersonii M. Hopkins × Polystichum munitum (Kaulf.) C. Presl munitum (Kaulf.) C. Presl U.S.A., WASHINGTON, SNOHOMISH U.S.A., WASHINGTON, SNOHOMISH COUNTY: COUNTY: Mt. Dickerman; talus above Perry Creek. Mt. Dickerman; talus above Perry Creek. Elev. 3000 ft. Elev. 3000 ft. 48.063294°, -121.513675°; WGS 84, 48.063294°, -121.513675°; WGS 84, uncertainty: 10000 m., Source: GeoLocate, uncertainty: 10000 m., Source: GeoLocate, Georef'd by WTU Staff Georef'd by WTU Staff Shaded mossy talus. About 8 plants, with Shaded mossy talus. About 8 plants, with parents. Phenology: Spores. Origin: Native. parents. Phenology: Spores. Origin: Native. W. H. Wagner 63115-3 W. H. Wagner 63115-22 with A. R. Kruckeberg with A. R. Kruckeberg 28 Jun 1963 28 Jun 1963 WTU-213564, WTU-V-021020 WTU-213557, WTU-V-021017

Dryopteridaceae Dryopteridaceae Polystichum andersonii M. Hopkins × Polystichum Polystichum scopulinum (D.C. Eaton) Maxon munitum (Kaulf.) C. Presl U.S.A., OREGON, KLAMATH COUNTY: U.S.A., WASHINGTON, SNOHOMISH Pete's Point, Wallowa Mountains. COUNTY: Elev. 2700 ft. Mt. Dickerman; talus above Perry Creek. Origin: Native. Elev. 3000 ft. 48.063294°, -121.513675°; WGS 84, Morton E. Peck 18059 uncertainty: 10000 m., Source: GeoLocate, Georef'd by WTU Staff 29 Jul 1933 Shaded mossy talus. About 8 plants, with parents. Phenology: Spores. Origin: Native. WTU-8440JWT, WTU-V-021736 W. H. Wagner 63115-4 with A. R. Kruckeberg 28 Jun 1963 WTU-213520, WTU-V-021019 Dryopteridaceae Dryopteridaceae Polystichum andersonii M. Hopkins × Polystichum Polystichum andersonii M. Hopkins × Polystichum munitum (Kaulf.) C. Presl munitum (Kaulf.) C. Presl U.S.A., WASHINGTON, SNOHOMISH U.S.A., WASHINGTON, SNOHOMISH COUNTY: COUNTY: Mt. Dickerman; talus above Perry Creek. Perry Creek, near Mt. Pilchuck, south fork of the Elev. 3000 ft. Stillaguamish River. 48.063294°, -121.513675°; WGS 84, Elev. 3000 ft. uncertainty: 10000 m., Source: GeoLocate, 48.063294°, -121.513675°; WGS 84, Georef'd by WTU Staff uncertainty: 10000 m., Source: GeoLocate, Shaded mossy talus. About 8 plants, with Georef'd by WTU Staff parents. Phenology: Spores. Origin: Native. Talus slopes in Tsuga heterophylla-Chamacyparis nootkatensis-Alnus W. H. Wagner 63115-11 rubra zone. Not common. Phenology: Spores. with A. R. Kruckeberg Origin: Native. 28 Jun 1963 Melinda F. Denton 2109 WTU-213521, WTU-V-021016 7 Oct 1972 WTU-258279, WTU-V-021102 Dryopteridaceae Polystichum andersonii M. Hopkins × Polystichum Dryopteridaceae munitum (Kaulf.) C. Presl Polystichum scopulinum (D.C. Eaton) Maxon U.S.A., WASHINGTON, SNOHOMISH COUNTY: U.S.A., IDAHO, IDAHO COUNTY: Perry Creek Trail; Stillaguamish River drainage. Clearwater National Forest, south east slope off 48.063294°, -121.513675°; WGS 84, the Walton Lakes Trail. uncertainty: 10000 m., Source: GeoLocate, Elev. 6400 ft. Georef'd by WTU Staff 46.0225°, -115.89°; WGS 84, uncertainty: In shade of vine maple, rocky slopes along trail. 10000 m., Source: GeoLocate, Georef'd by With Polystichum munitum, Athyrium WTU Staff filix-foemina, and Douglas Fir. Phenology: Crevices in bedrock. Origin: Native. Spores. Origin: Native. Klaus H. Lackschewitz 3899 A. R. Kruckeberg 2773 6 Aug 1979 21 Jul 1951 WTU-257660, WTU-V-021117 WTU-154495, WTU-V-021015 Dryopteridaceae Dryopteridaceae Polystichum scopulinum (D.C. Eaton) Maxon Polystichum kruckebergii W.H. Wagner U.S.A., WASHINGTON, CHELAN COUNTY: U.S.A., OREGON, KLAMATH COUNTY: Slope along G.N.Ry. near Drury. Pete's Point, Wallowa Mountains. Elev. 1600 ft. Elev. 2700 ft. 47.656962°, -120.729561°; WGS84, Origin: Native. uncertainty: 4600 m., Source: Georeferenced, Georef'd by WTU Staff Morton E. Peck 18059 29 Jul 1933 Dry slope. Origin: Native. WTU-8440JWT I. C. Otis 860 9 Sep 1918 WTU-28928, WTU-V-021096 Dryopteridaceae Dryopteridaceae Polystichum scopulinum (D.C. Eaton) Maxon Polystichum californicum (D.C. Eaton) Diels × Polystichum munitum (Kaulf.) C. Presl U.S.A., IDAHO, CASSIA COUNTY: Cache Peak Range (Albion Mountains), Silent U.S.A., WASHINGTON, PIERCE COUNTY: City of Rocks, near the summit of road, 13 air Eatonville Rim, 2 mile northeast of Eatonville. miles southeast of Oakley. Elev. 1200 ft. Elev. 6850 ft. 46.887969°, -122.235335°; WGS 84, T15S R23E S26; WGS 84, uncertainty: 10000 uncertainty: 1000 m., Source: GeoLocate, m., Source: GeoLocate, Georef'd by WTU Georef'd by WTU Staff Staff Rocky woods. Spores abortive (many sporangia Shady cracks of rock. Origin: Native. unopened and empty). A single plant with both parents nearby. Phenology: Vegetative. Origin: Noel H. Holmgren 5884 Native. with Patricia K. Holmgren Ed Alverson 240A 13 Jun 1972 11 Nov 1979 WTU-275974, WTU-V-021115 WTU-277095, WTU-V-030986 Dryopteridaceae Polystichum scopulinum (D.C. Eaton) Maxon Dryopteridaceae U.S.A., WASHINGTON, KITTITAS COUNTY: Polystichum scopulinum (D.C. Eaton) Maxon Crags along Beverly Creek, Wenatchee Mountains. U.S.A., WASHINGTON, KITTITAS COUNTY: Elev. 5000 ft. Along Beverly Creek, upper north fork of 47.372454°, -120.877017°; WGS 84, Teanaway River, Wenatchee Mountains, uncertainty: 10000 m., Source: GeoLocate, Wenatchee National Forest. Georef'd by WTU Staff 47.372454°, -120.877017°; WGS 84, Rocky crags. Origin: Native. uncertainty: 10000 m., Source: GeoLocate, Georef'd by WTU Staff J. William Thompson 9590 Growing with P. mohrioides, P. scopulinum and backcross. Origin: Native. 25 Jul 1933 W. H. Wagner 63099 WTU-5360JWT, WTU-V-021095 with A. R. Kruckeberg 25 Jul 1963 Dryopteridaceae WTU-213526, WTU-V-021094 Polystichum braunii (Spenner) Fee × Polystichum setigerum (K. Presl) K. Presl Dryopteridaceae CANADA, BRITISH COLUMBIA: Lower Squamish River Valley. Polystichum californicum (D.C. Eaton) Diels × On floodplain with second-growth mixed Polystichum munitum (Kaulf.) C. Presl ? hardwood-conifer forest. Site visited with Prof. T. M. C. Taylor, BC pteridologist. Phenology: U.S.A., WASHINGTON, PIERCE COUNTY: Spores. Origin: Native. Eatonville Rim, 2 mile northeast of Eatonville. Elev. 1200 ft. A. R. Kruckeberg 5767a 46.887969°, -122.235335°; WGS 84, uncertainty: 1000 m., Source: GeoLocate, 10 Nov 1963 Georef'd by WTU Staff Dry, rocky woods. Some spores seem to be WTU-313376, WTU-V-030990 viable. Both parents nearby. Resembles P. munitum except for the two lowest pairs of pinnae, which are bi-pinnatifid, as in P. californicum. Phenology: Spores. Origin: Native. Ed Alverson 239 11 Nov 1979 WTU-277101, WTU-V-030985 Dryopteridaceae Dryopteridaceae Polystichum scopulinum (D.C. Eaton) Maxon Polystichum scopulinum (D.C. Eaton) Maxon U.S.A., WASHINGTON, KITTITAS COUNTY: U.S.A., IDAHO, VALLEY COUNTY: Along Beverly Creek, at 2.5 mile mark, upper Gold Fork Lookout, Payette National Forest, north fork of Teanaway, R. Wenatchee Sawtooth Mountains. Mountains. Wenachee national forests. Elev. 8000 ft. 47.372454°, -120.877017°; WGS 84, 44.67194°, -115.83667°; WGS 84, uncertainty: 10000 m., Source: GeoLocate, uncertainty: 10000 m., Source: GeoLocate, Georef'd by WTU Staff Georef'd by WTU Staff Creek bed on rocks, growing with P. mohrioides Granite ledges. Origin: Native. and hybrid. Origin: Native. J. William Thompson 13797 W. H. Wagner 63092 with A. R. Kruckeberg 11 Jul 1937 25 Jun 1963 WTU-10416JWT, WTU-V-021092 WTU-213542, WTU-V-021093 Dryopteridaceae Dryopteridaceae Polystichum scopulinum (D.C. Eaton) Maxon Polystichum californicum (D.C. Eaton) Diels × U.S.A., WASHINGTON, KITTITAS COUNTY: Polystichum munitum (Kaulf.) C. Presl High steep open slope above Cle Elum river, Between Paris Creek and Little Boulder Creek. U.S.A., WASHINGTON, PIERCE COUNTY: 2 miles below little Boulder Creek. Near Eatonville, 35 miles from Tacoma. 47.407818°, -121.060366°; WGS 84, 46.8675°, -122.26528°; WGS 84, uncertainty: uncertainty: 1000 m., Source: GeoLocate, 5000 m., Source: GeoLocate, Georef'd by Georef'd by WTU Staff WTU Staff Plants somewhat intermediate; approaching P. Base of rocky crags. Phenology: Spores. Origin: scopulinum. Origin: Native. Native. A. R. Kruckeberg 5640 J. B. Flett 1926 12 Jul 1963 28 Aug 1901 WTU-217658, WTU-V-021091 WTU-174855, WTU-V-030984

Dryopteridaceae Dryopteridaceae Polystichum munitum (Kaulf.) C. Presl Polystichum scopulinum (D.C. Eaton) Maxon U.S.A., WASHINGTON, CLALLAM COUNTY: U.S.A., IDAHO, IDAHO COUNTY: Lower slopes of Mount Angeles, Olympic Above Sheep Creek Lakes #2, Dry Diggins Mountains. Area, Seven Devils Mountains, Nez Perce Elev. 3000 ft. National Forest. 47.995278°, -123.4575°; Source: Inferred Elev. 7500 ft. from duplicate specimen, Georef'd by Ed 45.483916°, -115.814891°; WGS 84, Alverson uncertainty: 10000 m., Source: GeoLocate, Wooded slopes. Phenology: Fertile. Origin: Georef'd by WTU Staff Native. Talus and in crevices. Origin: Native. J. William Thompson 7564 Kruckeberg 3192 20 Jul 1931 30 Jul 1952 WTU-5371JWT, WTU-V-18272 WTU-154486, WTU-V-021114 Dryopteridaceae Dryopteridaceae Polystichum scopulinum (D.C. Eaton) Maxon Polystichum scopulinum (D.C. Eaton) Maxon U.S.A., WASHINGTON, CHELAN COUNTY: U.S.A., WASHINGTON, CHELAN COUNTY: Slope on Bills Peak, head of Beverly Creek, Mount Stuart. Wenatchee Mountains. 47.47503°, -120.90223°; WGS 84, Elev. 5500 ft. uncertainty: 1500 m., Source: GeoLocate, 47.424°, -120.8977°; WGS84, uncertainty: Georef'd by WTU Staff 1000 m., Source: Georeferenced, Georef'd by Origin: Native. WTU Staff Rocky open slopes. Origin: Native. A. D. E. Elmer 1113 J. William Thompson 6395 Aug 1898 16 May 1931 WTU-37384, WTU-V-021086 WTU-5369JWT, WTU-V-021090 Dryopteridaceae Dryopteridaceae Polystichum munitum (Kaulf.) C. Presl Polystichum munitum (Kaulf.) C. Presl U.S.A., WASHINGTON, JEFFERSON COUNTY: U.S.A., WASHINGTON, JEFFERSON Trail to Constance Ridge. COUNTY: 47.77561°, -123.12014°; WGS 84, Hoh River valley near Nolan Creek. uncertainty: 1300 m., Source: GeoLocate, Elev. 100 ft. Georef'd by WTU Staff 47.750312°, -124.340519°; WGS 84, Dry rocky mountainside. Phenology: Fertile. uncertainty: 5000 m., Source: GeoLocate, Origin: Native. Georef'd by WTU Staff Phenology: Fertile. Origin: Native. G. N. Jones 5856 I. C. Otis 1371 8 Aug 1934 14 Aug 1924 WTU-39452, WTU-V-18268 WTU-28651, WTU-V-18273 Dryopteridaceae Dryopteridaceae Polystichum munitum (Kaulf.) C. Presl Polystichum munitum (Kaulf.) C. Presl U.S.A., WASHINGTON, CLALLAM COUNTY: Along Elwha River, vicinity of Altaire Public U.S.A., WASHINGTON, SKAMANIA COUNTY: Camp, Olympic Mountains. Mount St. Helens; First creek east of Timberline 48.060047°, -123.581458°; WGS 84, Camp. uncertainty: 10000 m., Source: GeoLocate, Elev. 4400 ft. Georef'd by WTU Staff 46.19893°, -122.18888°; WGS 84, Pseudotsuga forest along river. Phenology: uncertainty: 5000 m., Source: GeoLocate, Fertile. Origin: Native. Georef'd by WTU Staff Phenology: Vegetative. Origin: Native. Ira L. Wiggins 9419 Donald B. Lawrence s.n. 22 Jun 1940 6 Aug 1937 WTU-77913, WTU-V-18266 WTU-324193, WTU-V-18433 Dryopteridaceae Dryopteridaceae Polystichum munitum (Kaulf.) C. Presl Polystichum munitum (Kaulf.) C. Presl U.S.A., WASHINGTON, SKAMANIA COUNTY: Along the footpath which climbs Beacon Rock, U.S.A., WASHINGTON, JEFFERSON Beacon Rock State Park (along U.S. 830). COUNTY: T2N R6E S25; Source: Inferred from duplicate Hoh River valley near Nolan Creek. specimen, Georef'd by Ed Alverson Elev. 100 ft. Crevices of rock. Phenology: Fertile. Origin: 47.750312°, -124.340519°; WGS 84, Native. uncertainty: 5000 m., Source: GeoLocate, Georef'd by WTU Staff D. R. Simpson 322 Phenology: Fertile. Origin: Native. 20 Feb 1966 I. C. Otis 1370 WTU-288498, WTU-V-18317 14 Aug 1924 WTU-8118JWT, WTU-V-18270 Dryopteridaceae Dryopteridaceae Polystichum munitum (Kaulf.) C. Presl Polystichum scopulinum (D.C. Eaton) Maxon U.S.A., WASHINGTON, SAN JUAN COUNTY: U.S.A., WASHINGTON, KITTITAS COUNTY: San Juan Island; west side of Mt. Dallas. On Trailhead of Boulder Creek trail, 6.7 miles by FR west side of the road. 4330 and FR 138 from Salmon la Sac. 30 48.54111°, -123.08472°; WGS 84, degrees south-facing slope. uncertainty: 10000 m., Source: GeoLocate, Elev. 3500 ft. Georef'd by WTU Staff 47° 28.812' N, 121° 2.577' W; T23N R14E Open and dry rocky slople above shoreline. S36 Common among the rocks. Phenology: Fertile. Soil with some serpentine rocks, growing with Origin: Native. Pseudotsuga menziesii, Phlox diffusa, Aspidotus densa. Origin: Native. Melinda F. Denton 3506 Don Knoke 429 2 Aug 1974 25 May 2004 WTU-259545, WTU-V-18264 WTU-357773, WTU-V-021044

Dryopteridaceae Dryopteridaceae Polystichum scopulinum (D.C. Eaton) Maxon Polystichum scopulinum (D.C. Eaton) Maxon U.S.A., OREGON, HARNEY COUNTY: U.S.A., WASHINGTON, CHELAN COUNTY: Cliffs along the south tributary to Alvord Creek, Crevices of talus slopes, Mount stuart, about a mile above the falls; east side of the Wenatchee mountains. Steens Mountains, 100 miles southeast of Elev. 7000 ft. Burns. 47.47522°, -120.90238°; WGS84, uncertainty: Elev. 7500 ft. 2000 m., Source: Georeferenced, Georef'd by T33S R34E S32; NAD 27, uncertainty: 805 WTU Staff m., Source: TRS2LL, Georef'd by Ben Legler Crevices. Origin: Native. Crevices in basaltic cliffs. Plants apparently evergreen, these being las year's fronds. Origin: J. William Thompson 7674 Native. 27 Jul 1931 Arthur Cronquist 8256 WTU-5358JWT, WTU-V-021043 20 May 1959 WTU-211687, WTU-V-021105 Dryopteridaceae Dryopteridaceae Polystichum scopulinum (D.C. Eaton) Maxon Polystichum munitum (Kaulf.) C. Presl U.S.A., WASHINGTON, CHELAN COUNTY: Between Chiwaukum and Drury. U.S.A., WASHINGTON, SAN JUAN COUNTY: Elev. 1700 ft. Stuart Island. 47.656962°, -120.729561°; WGS84, 48.674722°, -123.210556°; WGS 84, uncertainty: 4600 m., Source: Georeferenced, uncertainty: 3700 m., Source: Georeferenced, Georef'd by WTU Staff Georef'd by WTU Staff Origin: Native. Deep moist woods. Phenology: Fertile. Origin: Native. I. C. Otis 1041 W. H. Lawrence 31 18 Aug 1920 18 Jun 1904 WTU-115953, WTU-V-021738 WTU-95451, WTU-V-18262 Dryopteridaceae Dryopteridaceae Polystichum munitum (Kaulf.) C. Presl Polystichum munitum (Kaulf.) C. Presl U.S.A., WASHINGTON, KING COUNTY: Snoqualmie National Forest. U.S.A., WASHINGTON, PIERCE COUNTY: Elev. 450 ft. Ranger camp, Mount Rainier. 47.52889°, -121.82417°; WGS 84, Phenology: Fertile. Origin: Native. uncertainty: 10000 m., Source: GeoLocate, Georef'd by WTU Staff J. M. Grant s.n. Damp bottom woods. Very large trees. Phenology: Fertile. Origin: Native. Aug 1925 Delzie Demaree 60901 WTU-22265, WTU-V-18333 21 Sep 1969 WTU-274606, WTU-V-18332 Dryopteridaceae Dryopteridaceae Polystichum scopulinum (D.C. Eaton) Maxon Polystichum munitum (Kaulf.) C. Presl U.S.A., WASHINGTON, CHELAN COUNTY: U.S.A., IDAHO, CLEARWATER COUNTY: Mount Stuart. Wenatchee Mountains. Little North Fork of Clearwater River, Clearwater Elev. 8000 ft. Forest Reserve. 47.474167°, -120.915°; NAD 27, uncertainty: Phenology: Fertile. Origin: Native. 3000 m., Source: Georeferenced, Georef'd by WTU Staff F. L. Pickett 1128 Crevices of high cliffs. Origin: Native. 11 Aug 1917 J. William Thompson 5837 WTU-66617, WTU-V-18276 20 Aug 1930 WTU-5357JWT, WTU-V-021104

Dryopteridaceae Dryopteridaceae Polystichum scopulinum (D.C. Eaton) Maxon Polystichum scopulinum (D.C. Eaton) Maxon U.S.A., WASHINGTON, CHELAN COUNTY: U.S.A., WASHINGTON, CHELAN COUNTY: Head of Ingall's Creek, Wenatchee Mountains. Wenatchee Forest, in cleft in rocks on Icicle Elev. 4000 ft. River, slope 0. 47.42361°, -120.30917°; WGS84, uncertainty: Elev. 6600 ft. 10000 m., Source: GeoLocate, Georef'd by 47.718591°, -121.054146°; WGS 84, WTU Staff uncertainty: 10000 m., Source: Digital Map, Open rocky slopes. Collection date:July27-31 Georef'd by Dan Post 1931. Origin: Native. Barren Rocks, no other vegetation present, clefts. Origin: Native. J. William Thompson 7747 Frank B. Lenzie 313 27 Jul 1931 29 Sep 1917 WTU-10209JWT, WTU-V-021108 WTU-323928, WTU-V-021739 Dryopteridaceae Dryopteridaceae Polystichum munitum (Kaulf.) C. Presl Polystichum munitum (Kaulf.) C. Presl U.S.A., OREGON COUNTY: Near Cortland. U.S.A., MONTANA COUNTY: Phenology: Vegetative. Origin: Native. Within the wester half of section 7 and the eastern half of section 8 along the Bear creek William Palmer s.n. trail in the Bitterroot Mtns. just south and west of Victor. 22 May 1905 Shaded alcove. Only a few representatives of this species were noted. Phenology: Fertile. WTU-243698, WTU-V-18275 Origin: Native. Clark schaack 1033 2 Oct 1976 WTU-272506, WTU-V-18294 Dryopteridaceae Polystichum munitum (Kaulf.) C. Presl Dryopteridaceae U.S.A., ALASKA COUNTY: Baranof island, southeast side. Northeast part of Polystichum scopulinum (D.C. Eaton) Maxon Port Armstrong just west of the site of old whale station. Tongass National Forest. Southeastern CANADA, BRITISH COLUMBIA: Alaska. [locality and coordinates withheld; status in Grassy south facing. Rock cliff just above British Columbia: S2 (Imperiled, Red List; narrow beach meadow and garden spot. BCCDC)] Phenology: Fertile. Origin: Native. Outcrop of peridotite and serpentine. Origin: Native. Mary Clay Muller 4625 A. R. Kruckeberg 5728 16 Aug 1981 7 Sep 1963 WTU-282674, WTU-V-18293 WTU-217656, WTU-V-021103 Dryopteridaceae Dryopteridaceae Polystichum scopulinum (D.C. Eaton) Maxon Polystichum munitum (Kaulf.) C. Presl U.S.A., OREGON, KLAMATH/LAKE COUNTY: U.S.A., ALASKA, KETCHIKAN GATEWAY Summit of Gearhart Mountain, Sprague River COUNTY: region. Southeastern Alaska, Revillagigedo Island, Elev. 8000 ft. south side. About 10 miles west of Ketchikan Origin: Native. near Totem Bight. North side of the highway. Soil on an ustable rock cliff. Last year's fronds. Elmer I. Applegate 4528 Phenology: Fertile. Origin: Native. 27 Aug 1925 Mary Clay Muller 7414 WTU-8487JWT, WTU-V-021112 6 Mar 1982 WTU-282143, WTU-V-18292

Dryopteridaceae Dryopteridaceae Polystichum munitum (Kaulf.) C. Presl Polystichum munitum (Kaulf.) C. Presl U.S.A., OREGON, JOSEPHINE COUNTY: U.S.A., OREGON, CLACKAMAS COUNTY: Along Big Tree Trail at Cave Creek. Oregon Upper Clackamas River. National Mounument. Three Links. Phenology: Fertile. Origin: Native. Phenology: Fertile. Origin: Native. Emily M. Anderson s.n. William H. Baker 519 with George C. Ruhle 31 Mar 1928 2 Aug 1949 WTU-8121JWT, WTU-V-18274 WTU-298182, WTU-V-18291

Dryopteridaceae Dryopteridaceae Polystichum munitum (Kaulf.) C. Presl Polystichum scopulinum (D.C. Eaton) Maxon U.S.A., OREGON, CLACKAMAS COUNTY: U.S.A., OREGON, BAKER COUNTY: Near Mirror Lake, Mt. Hood region. Anthony Lakes Region, Blue Mountains. Elev. 4000 ft. Elev. 7100 ft. 45.297067°, -121.793085°; uncertainty: 3500 44.959998°, -118.228984°; WGS 84, m. uncertainty: 1912 m., Source: Digital Map, Rocky crevices. Phenology: Fertile. Origin: Georef'd by Joseph Stormer Native. Origin: Native. J. William Thompson 3570 J. William Thompson 13452 17 Aug 1927 23 Jul 1936 WTU-5375JWT, WTU-V-18277 WTU-5099JWT, WTU-V-021734

Dryopteridaceae Dryopteridaceae Polystichum scopulinum (D.C. Eaton) Maxon Polystichum munitum (Kaulf.) C. Presl U.S.A., WASHINGTON, CHELAN COUNTY: Hillside east of G.N. bridge across the U.S.A., IDAHO, IDAHO COUNTY: Wenatchee River, between Drury and Near Selway River, 2 miles up from Selway Chiwaukum. Falls. Elev. 1700 ft. 46.053515°, -115.309684°; WGS 84, 47.656962°, -120.729561°; WGS84, uncertainty: 1000 m., Source: GeoLocate, uncertainty: 4600 m., Source: Georeferenced, Georef'd by WTU Staff Georef'd by WTU Staff Comon. Phenology: Fertile. Origin: Native. Rocky hillside. Origin: Native. J. L. McMullen 1485 I. C. Otis 1061 3 Jun 1954 2 Oct 1920 WTU-299574, WTU-V-18314 WTU-8504JWT, WTU-V-021735 Dryopteridaceae Dryopteridaceae Polystichum scopulinum (D.C. Eaton) Maxon Polystichum munitum (Kaulf.) C. Presl U.S.A., OREGON COUNTY: U.S.A., OREGON, CURRY COUNTY: China Cap Peak, Wallowa Mountains. Rogue River Canyon. Along a stream at Cherry Elev. 2894 ft. Flat 4 miles north of Agness. Extra label of Polystichum lonchitis (L.) Roth. Woods along a stream. Phenology: Fertile. Origin: Native. Origin: Native. W. C. Cusick 3050 W. H. Baker 6115 Jun 1906 13 Jun 1949 WTU-8481JWT, WTU-V-021110 WTU-299364, WTU-V-18312

Dryopteridaceae Dryopteridaceae Polystichum munitum (Kaulf.) C. Presl Polystichum lonchitis (L.) Roth U.S.A., ALASKA COUNTY: U.S.A., WASHINGTON, CLALLAM COUNTY: Baranof island, southeast side. Northeast part of Hudsonian zone, Crystal Creek, Mount Olympus Port Armstrong just west of the site of old whale National Monument. station. Tongass National Forest. Southeastern 47.977296°, -123.684108°; WGS 84, Alaska. uncertainty: 10000 m., Source: GeoLocate, Narrow steeply sloping upper beach meadow. Georef'd by WTU Staff Phenology: Fertile. Origin: Native. Rocky slope. Phenology: Spores. Origin: Native. Mary Clay Muller 4626 George Neville Jones 8433 16 Aug 1981 7 Sep 1935 WTU-282676, WTU-V-18313 WTU-30206, WTU-V-016253

Dryopteridaceae Dryopteridaceae Polystichum imbricans (D.C. Eaton) D.H. Wagner ssp. Polystichum munitum (Kaulf.) C. Presl imbricans U.S.A., IDAHO COUNTY: U.S.A., OREGON, JOSEPHINE COUNTY: Big Creek, Kaniksu National Forest. 1 mile west bridge, Hellgate, Rouge River T57N R3W S6; NAD 27, uncertainty: 805 m., Canyon. Source: TRS2LL, Georef'd by Ben Legler Phenology: Fertile. Origin: Native. Phenology: Fertile. Origin: Native. Lilla Leach s.n. R. C. Stillinger 749 30 May 1928 7 Jul 1941 WTU-11457JWT, WTU-V-016200 WTU-299379, WTU-V-18311

Dryopteridaceae Dryopteridaceae Polystichum scopulinum (D.C. Eaton) Maxon Polystichum scopulinum (D.C. Eaton) Maxon U.S.A., OREGON COUNTY: U.S.A., OREGON, HARNEY COUNTY: Crater Lake National Park, Wizard Island, rock Cliff crevices on the northeast side of crevices on the south shore. Felsite-porphyry outcrop atop the Trout Creek 42.93619°, -122.149742°; WGS 84, Mountains. uncertainty: 750 m., Source: Digital Map, Elev. 9000 ft. Georef'd by Joseph Stormer T41S R38E S9; NAD 27, uncertainty: 805 m., Rock crevices. Origin: Native. Source: TRS2LL, Georef'd by Ben Legler Origin: Native. William H. Baker 6357 Arthur Cronquist 8664 21 Aug 1949 9 Jul 1959 WTU-129665, WTU-V-021101 WTU-211754, WTU-V-021107 Dryopteridaceae Dryopteridaceae Polystichum munitum (Kaulf.) C. Presl Polystichum lonchitis (L.) Roth U.S.A., IDAHO, CLEARWATER COUNTY: U.S.A., WASHINGTON, SNOHOMISH 4 miles east of Pierce. COUNTY: 46.491079°, -115.713946°; WGS 84, Mount Dickerman, shaded mossy talus above uncertainty: 1000 m., Source: GeoLocate, Perry Creek. Georef'd by WTU Staff Elev. 3000 ft. Woods. Phenology: Fertile. Origin: Native. 48.063294°, -121.513675°; WGS 84, uncertainty: 10000 m., Source: GeoLocate, William H. Baker 12779 Georef'd by WTU Staff Frequent on rocks. Phenology: Spores. Origin: 23 Aug 1954 Native. WTU-299571, WTU-V-18310 W. H. Wagner 63117 with A. R. Kruckeberg 28 Jun 1963 Dryopteridaceae WTU-213554, WTU-V-016254 Polystichum munitum (Kaulf.) C. Presl U.S.A., OREGON, KLAMATH COUNTY: Dryopteridaceae North wall of Redblanket Canyon, southwest corner of Crater Lake National Park. Polystichum munitum (Kaulf.) C. Presl 42.790111°, -122.309642°; WGS 84, uncertainty: 9000 m., Source: Digital Map, U.S.A., ALASKA COUNTY: Georef'd by Joseph Stormer, Invalid Dixon Entrance Quad.: Dall I., Crace Mountain. coordinates (located outside county bounds). C.a. 224 m south-southwest of peak 2443. Phenology: Fertile. Origin: Native. 54° 55' N, 132° 57' W Limestone substrate. Mixed conifer subalpine William H. Baker 7018 forest. Open understory, mossy west-facing slope. 640 m.s.m. Phenology: Fertile. Origin: 29 Jul 1950 Native. WTU-299572, WTU-V-18309 M. C. Muller 5596 17 Aug 1990 Dryopteridaceae WTU-333156, WTU-V-18307, 3013 Polystichum munitum (Kaulf.) C. Presl U.S.A., IDAHO, CLEARWATER COUNTY: Dryopteridaceae On north slopes near east fork of Potlatch Creek, 10 miles east of Boville, Twp. Polystichum scopulinum (D.C. Eaton) Maxon Elev. 3300 ft. T14N R2E S20; WGS 84, uncertainty: 10000 U.S.A., WASHINGTON, CHELAN COUNTY: m., Source: GeoLocate, Georef'd by WTU Ridge back of Ingalls Creek Ranger Station. Staff Elev. 4000 ft. Mixed coniferous woods. In large clumps from a 47.462899°, -120.661069°; WGS84, compact caudex. Phenology: Fertile. Origin: uncertainty: 10000 m., Source: GeoLocate, Native. Georef'd by WTU Staff Rocky ridge. Origin: Native. Quentin Jones 5991 Harry B. Higman s.n. 24 Jul 1949 11 Jun 1933 WTU-292830, WTU-V-18308 WTU-10244JWT, WTU-V-021099 Dryopteridaceae Polystichum scopulinum (D.C. Eaton) Maxon Dryopteridaceae U.S.A., OREGON COUNTY: Polystichum scopulinum (D.C. Eaton) Maxon Ledge of mountain north of Tumalo Ranger Station. U.S.A., WASHINGTON, CHELAN COUNTY: Origin: Native. Hillside, west of Drury. Elev. 1700 ft. K. Whited s.n. 47.61975°, -120.72665°; WGS84, uncertainty: 500 m., Source: Georeferenced, Georef'd by 1 Aug 1922 WTU Staff Rocks on hillside. Origin: Native. WTU-8336JWT, WTU-V-021100 I. C. Otis 1041 18 Aug 1920 WTU-8503JWT, WTU-V-021733 Dryopteridaceae Dryopteridaceae Polystichum lonchitis (L.) Roth Polystichum scopulinum (D.C. Eaton) Maxon U.S.A., WASHINGTON, CHELAN COUNTY: U.S.A., WASHINGTON, KLICKITAT COUNTY: Near Ice Creek. Cliffs near Columbia River. 48.105535°, -120.737582°; WGS84, 45.71167°, -120.72611°; WGS 84, uncertainty: 5000 m., Source: GeoLocate, uncertainty: 10000 m., Source: GeoLocate, Georef'd by WTU Staff Georef'd by WTU Staff On cliffs. Phenology: Spores. Origin: Native. Cliffs. Origin: Native. G. E. Morrill 302 W. N. Suksdorf 2084 22 Aug 1933 11 Aug 1892 WTU-27765, WTU-V-016251 WTU-270514, WTU-V-021042

Dryopteridaceae Dryopteridaceae Polystichum scopulinum (D.C. Eaton) Maxon Polystichum lonchitis (L.) Roth U.S.A., WASHINGTON, KITTITAS COUNTY: U.S.A., WASHINGTON, GRAYS HARBOR Wenatchee Mountains. North fork Teanaway COUNTY: River. Upper reaches of west fork Humptulips River, Elev. 4500 ft. above Campbell Grove campground. 47.251427°, -120.87694°; WGS 84, 47.22956°, -123.956466°; WGS 84, uncertainty: 10000 m., Source: GeoLocate, uncertainty: 10000 m., Source: GeoLocate, Georef'd by WTU Staff Georef'd by WTU Staff Origin: Native. Phenology: Spores. Origin: Native. C. S. English s.n. A. R. Kruckeberg s.n. Jul 1932 30 Jun 1978 WTU-23865, WTU-V-021118 WTU-280834, WTU-V-021077

Dryopteridaceae Dryopteridaceae Polystichum scopulinum (D.C. Eaton) Maxon Polystichum scopulinum (D.C. Eaton) Maxon U.S.A., WASHINGTON, CHELAN COUNTY: U.S.A., WASHINGTON, KITTITAS COUNTY: Slopes of Three Brothers Peak, Wenatchee Slopes along Beverly Creek, Wenatchee Mountains. Mountains. Elev. 5500 ft. Elev. 4000 ft. 47.43891°, -120.74198°; WGS84, uncertainty: 47.372454°, -120.877017°; WGS 84, 2000 m., Source: Georeferenced, Georef'd by uncertainty: 10000 m., Source: GeoLocate, WTU Staff Georef'd by WTU Staff Origin: Native. Rocky slopes. Origin: Native. J. William Thompson 8560 J. William Thompson 8755 25 Jun 1932 13 Jul 1932 WTU-5366JWT, WTU-V-021098 WTU-5362JWT, WTU-V-021116

Dryopteridaceae Dryopteridaceae Polystichum lonchitis (L.) Roth Polystichum lonchitis (L.) Roth U.S.A., WASHINGTON, JEFFERSON U.S.A., WASHINGTON, OKANOGAN COUNTY: COUNTY: Lower crags of Billy Goat Mountain. Olympic National Park: Royal Creek trail, 2 Elev. 6000 ft. miles below Royal Basin and Lake. 48.81278°, -120.34222°; WGS 84, 47.854841°, -123.214107°; Source: Inferred uncertainty: 1000 m., Source: GeoLocate, from duplicate specimen, Georef'd by Ed Georef'd by WTU Staff Alverson Phenology: Spores. Seep in coniferous woods, with P. munitum. Phenology: Spores. Origin: Native. J. William Thompson 10889 A. R. Kruckeberg s.n. 1 Jul 1934 8 Oct 1979 WTU-2910JWT, WTU-V-016252 WTU-305223, WTU-V-021075 Dryopteridaceae Dryopteridaceae Polystichum scopulinum (D.C. Eaton) Maxon Polystichum setigerum (C. Presl) C. Presl U.S.A., OREGON COUNTY: U.S.A., ALASKA COUNTY: China Cap Peak, Wallowa Mountains. [locality and coordinates withheld; status in Elev. 2894 ft. Alaska: S3 (Rare; AKNHP)] Origin: Native. Elev. 150 ft. Moist slope among herbaceous plants. Origin: W. C. Cusick 3050 Native. Jun 1906 Robert M. Hardy 52 WTU-139193, WTU-V-021097 26 Apr 1945 WTU-97626, WTU-V-021038

Dryopteridaceae Dryopteridaceae Polystichum setigerum (C. Presl) C. Presl Polystichum setigerum (C. Presl) C. Presl U.S.A., ALASKA COUNTY: U.S.A., ALASKA, WRANGELL PETERSBURG [locality and coordinates withheld; status in COUNTY: Alaska: S3 (Rare; AKNHP)] [locality and coordinates withheld; status in Origin: Native. Alaska: S3 (Rare; AKNHP)] In devil climb thicket. Origin: Native. Mr. Ernest P. Walker 1084 with Mrs. Ernest P. Walker W. J. Eyerdam 5452 Jun 1916 20 Aug 1947 WTU-74626, WTU-V-021039 WTU-118037, WTU-V-021036

Dryopteridaceae Dryopteridaceae Polystichum lonchitis (L.) Roth Polystichum setigerum (C. Presl) C. Presl U.S.A., WASHINGTON, WHATCOM COUNTY: U.S.A., ALASKA COUNTY: At base of slate ledges above Twin Lakes; [locality and coordinates withheld; status in Mount Baker National Forest. Alaska: S3 (Rare; AKNHP)] Elev. 5500 ft. Phenology: Spores. Origin: Native. 48.95194°, -121.63444°; WGS 84, uncertainty: 10000 m., Source: GeoLocate, Mr. Ernest P. Walker 1084 Georef'd by WTU Staff with Mrs. Ernest P. Walker Tsuga mertensiana- Vaccinium association. Phenology: Spores. Origin: Native. Jun 1916 A. R. Kruckeberg 3799 WTU-130366, WTU-V-021037 26 Sep 1954 WTU-190451, WTU-V-016249 Dryopteridaceae Polystichum setigerum (C. Presl) C. Presl Dryopteridaceae U.S.A., ALASKA COUNTY: Polystichum lonchitis (L.) Roth [locality and coordinates withheld; status in Alaska: S3 (Rare; AKNHP)] U.S.A., WASHINGTON, KITTITAS COUNTY: Elev. 250 ft. Mount Daniel, 4 miles west of Hyas Lake. Herbarceous slopes and rocky rocky places. Elev. 6000 ft. Origin: Native. 47.56256°, -121.17189°; WGS 84, uncertainty: 2000 m., Source: GeoLocate, Robert M. Hardy 404 Georef'd by WTU Staff Phenology: Spores. Origin: Native. 23 Sep 1945 C. L. Hitchcock 8024 WTU-97563, WTU-V-021033 12 Jul 1942 WTU-75661, WTU-V-016250 Dryopteridaceae Dryopteridaceae Polystichum microchlamys (H. Christ) Matsum. Polystichum lemmonii Underw. U.S.A., ALASKA, ALEUTIANS WEST COUNTY: U.S.A., OREGON, JACKSON COUNTY: [locality and coordinates withheld; status in Observation Gap, about 1 1/2 miless south of Alaska: S1 (Critically Imperiled; AKNHP)] Dutchmans Peak. Elev. 250 ft. Elev. 7000 ft. Steep slope among ferns and tall herbs. 42° 1' N, 122° 53' W Phenology: Spores. Origin: Native. Parent material peridotite. Phenology: Spores. Origin: Native. Robert M. Hardy 316 Richard Waring 320 8 Aug 1945 22 Aug 1963 WTU-97296, WTU-V-021014 WTU-312946, WTU-V-016207

Dryopteridaceae Dryopteridaceae Polystichum setigerum (C. Presl) C. Presl Polystichum imbricans (D.C. Eaton) D.H. Wagner × U.S.A., ALASKA COUNTY: Polystichum munitum (Kaulf.) Presl [locality and coordinates withheld; status in Alaska: S3 (Rare; AKNHP)] U.S.A., WASHINGTON, CHELAN COUNTY: Elev. 150 ft. Rock Mountain Trail 1.5 to 2 miles from Western hemlock-Sitka spruce forest, growing Highway 2. on a steep grassy slope under the canopy. Elev. 4400 ft. Origin: Native. T26N R15E S3 NE 1/4; WGS84, uncertainty: 500 m., Source: Georeferenced, Georef'd by Mary Clay Stensvold 5618 WTU Staff With Ceanothus velutinus, Acer glabrum, and 16 Jul 1991 Sorbus scopulina. A few small Pseudotsuga menziesii nearby. Phenology: Spores. Origin: WTU-332864, WTU-V-021034 Native. E. Burnett s.n. Dryopteridaceae 20 Jul 1991 Polystichum microchlamys (H. Christ) Matsum. WTU-323847, WTU-V-030982 U.S.A., ALASKA, ALEUTIANS WEST COUNTY: [locality and coordinates withheld; status in Alaska: S1 (Critically Imperiled; AKNHP)] Dryopteridaceae Elev. 250 ft. Steep slope among ferns and tall herbs. Polystichum imbricans (D.C. Eaton) D.H. Wagner ssp. Phenology: Fertile. Origin: Native. imbricans Robert M. Hardy 316 U.S.A., WASHINGTON, CHELAN COUNTY: Rock Mountain Trail 1.5 to 2 miles from 8 Aug 1945 Highway 2. Elev. 4400 ft. WTU-97562, WTU-V-021071 T26N R15E S3 NE 1/4; WGS84, uncertainty: 500 m., Source: Georeferenced, Georef'd by WTU Staff With Ceanothus velutinus, Acer glabrum, and Dryopteridaceae Sorbus scopulina. A few small Pseudotsuga menziesii nearby. Phenology: Spores. Origin: Polystichum setigerum (C. Presl) C. Presl Native. U.S.A., ALASKA, SOUTHEAST FAIRBANKS E. Burnett s.n. COUNTY: [locality and coordinates withheld; status in 20 Jul 1991 Alaska: S3 (Rare; AKNHP)] Grassy upper beach meadow, about 20 feet WTU-323846, WTU-V-030979 east of P. Munitum. Origin: Native. Mary Clay Muller 4627 16 Aug 1981 WTU-282126, WTU-V-021030 Dryopteridaceae Dryopteridaceae Polystichum imbricans (D.C. Eaton) D.H. Wagner × Polystichum imbricans (D.C. Eaton) D.H. Wagner × Polystichum scopulinum (D.C. Eaton) Maxon Polystichum scopulinum (D.C. Eaton) Maxon U.S.A., WASHINGTON, CHELAN COUNTY: U.S.A., WASHINGTON, KITTITAS COUNTY: Between Drury and Chiwaukum. Along Beverly Creek. Elev. 1700 ft. Elev. 3000 ft. 47.656962°, -120.729561°; WGS84, 47.372454°, -120.877017°; WGS 84, uncertainty: 4600 m., Source: Georeferenced, uncertainty: 10000 m., Source: GeoLocate, Georef'd by WTU Staff Georef'd by WTU Staff Phenology: Spores. Origin: Native. Rocky creek beds. Phenology: Spores. Origin: Native. I. C. Otis 1062 J. William Thompson 9475 2 Oct 1920 19 Jul 1933 WTU-115952, WTU-V-030977 WTU-10206JWT, WTU-V-030978

Dryopteridaceae Dryopteridaceae Polystichum imbricans (D.C. Eaton) D.H. Wagner × Polystichum scopulinum (D.C. Eaton) Maxon Polystichum imbricans (D.C. Eaton) D.H. Wagner × Polystichum scopulinum (D.C. Eaton) Maxon U.S.A., WASHINGTON, CHELAN COUNTY: Between Drury and Chiwaukum. U.S.A., WASHINGTON, CHELAN COUNTY: Elev. 1700 ft. Hillside east of G.N. bridge across the 47.656962°, -120.729561°; WGS84, Wenatchee River, between Drury and uncertainty: 4600 m., Source: Georeferenced, Chiwaukum. Georef'd by WTU Staff Elev. 1700 ft. Phenology: Spores. Origin: Native. 47.656962°, -120.729561°; WGS84, uncertainty: 4600 m., Source: Georeferenced, I. C. Otis 1062 Georef'd by WTU Staff On rocky hillside. Phenology: Spores. Origin: 2 Oct 1920 Native. WTU-2913JWT, WTU-V-030976 I. C. Otis 1062 2 Oct 1920 Dryopteridaceae WTU-8505JWT, WTU-V-030974 Polystichum imbricans (D.C. Eaton) D.H. Wagner × Polystichum scopulinum (D.C. Eaton) Maxon Dryopteridaceae U.S.A., WASHINGTON, KITTITAS COUNTY: Along Beverly Creek, upper north fork of Polystichum lemmonii Underw. × Polystichum Teanaway River; Wenatchee Mountains, scopulinum (D.C. Eaton) Maxon Wenatchee National Forest. 47.372454°, -120.877017°; WGS 84, U.S.A., WASHINGTON, KITTITAS COUNTY: uncertainty: 10000 m., Source: GeoLocate, Along Beverly Creek, at 2.5 mile mark, upper Georef'd by WTU Staff north fork of Teanaway, R. Wenatchee One plant with parents. Phenology: Spores. Mountains. Wenachee national forests. Origin: Native. 47.372454°, -120.877017°; WGS 84, uncertainty: 10000 m., Source: GeoLocate, W. H. Wagner 63089 Georef'd by WTU Staff with A. R. Kruckeberg Along creek on rocks. Specimen #2. With parents. Phenology: Spores. Origin: Native. 25 Jun 1963 W. H. Wagner 63094 WTU-213543, WTU-V-030975 with A. R. Kruckeberg 25 Jun 1963 WTU-213529, WTU-V-030969 Dryopteridaceae Dryopteridaceae Polystichum lemmonii Underw. × Polystichum Polystichum lemmonii Underw. × Polystichum scopulinum (D.C. Eaton) Maxon scopulinum (D.C. Eaton) Maxon U.S.A., WASHINGTON, KITTITAS COUNTY: U.S.A., WASHINGTON, KITTITAS COUNTY: Along Beverly Creek, upper north fork of Along Beverly Creek, upper north fork of Teanaway River; Wenatchee Mountians, Teanaway River; Wenatchee Mountians, Wenatchee National Forest. Wenatchee National Forest. 47.372454°, -120.877017°; WGS 84, 47.372454°, -120.877017°; WGS 84, uncertainty: 10000 m., Source: GeoLocate, uncertainty: 10000 m., Source: GeoLocate, Georef'd by WTU Staff Georef'd by WTU Staff With parents. Phenology: Spores. Origin: With Polystichum mohrioides, Polystichum Native. scopulinum, and Polystichum mohrioides backcross. Specimen #3. Phenology: Spores. W. H. Wagner 63100 Origin: Native. with A. R. Kruckeberg W. H. Wagner 63098 25 Jun 1963 with A. R. Kruckeberg WTU-213524, WTU-V-030973 25 Jun 1963 WTU-213528, WTU-V-030968 Dryopteridaceae Polystichum lemmonii Underw. × Polystichum scopulinum (D.C. Eaton) Maxon Dryopteridaceae U.S.A., WASHINGTON, KITTITAS COUNTY: Polystichum imbricans (D.C. Eaton) D.H. Wagner × Along Beverly Creek, at 2 miles mark, upper Polystichum lemmonii Underw. north fork of Teanaway River; Wenatchee Mountains, Wenatchee National Forest. U.S.A., WASHINGTON, KITTITAS COUNTY: 47.372454°, -120.877017°; WGS 84, Miller Peak. Talus of northeast slope. uncertainty: 10000 m., Source: GeoLocate, Elev. 6420 ft. Georef'd by WTU Staff 47.39472°, -120.74222°; WGS 84, Deep in wooded valley on rocks. With parents. uncertainty: 10000 m., Source: GeoLocate, Phenology: Spores. Origin: Native. Georef'd by WTU Staff Shale-sandstone-conglomerate; subalpine W. H. Wagner 63093 fir-mountain hemlock-whitebark pine forest type. with A. R. Kruckeberg Phenology: Spores. Origin: Native. 25 Jun 1963 A. R. Kruckeberg 3957 WTU-213530, WTU-V-030971 23 Aug 1955 WTU-192106, WTU-V-030966 Dryopteridaceae Polystichum lemmonii Underw. × Polystichum Dryopteridaceae scopulinum (D.C. Eaton) Maxon Polystichum lemmonii Underw. × Polystichum U.S.A., WASHINGTON, KITTITAS COUNTY: scopulinum (D.C. Eaton) Maxon Along Beverly Creek, half way up slope above 3 mile mark, upper north fork of Teanaway River; U.S.A., WASHINGTON, CHELAN COUNTY: Wenatchee Mountains, Wenatchee National Base of Three Brothers. Forest. Near specimen #5. Elev. 4000 ft. 47.372454°, -120.877017°; WGS 84, 47.43891°, -120.74198°; WGS84, uncertainty: uncertainty: 10000 m., Source: GeoLocate, 2000 m., Source: Georeferenced, Georef'd by Georef'd by WTU Staff WTU Staff With Polystichum mohrioides. Specimen #4. Rocky slopes. Phenology: Spores. Origin: With parents. Phenology: Spores. Origin: Native. Native. J. William Thompson 10753 W. H. Wagner 63097 with A. R. Kruckeberg 21 Jun 1934 25 Jun 1963 WTU-138JWT, WTU-V-030964 WTU-213527, WTU-V-030970 Dryopteridaceae Dryopteridaceae Polystichum lemmonii Underw. × Polystichum Polystichum lonchitis (L.) Roth × Polystichum scopulinum (D.C. Eaton) Maxon scopulinum (D.C. Eaton) Maxon U.S.A., OREGON, JACKSON COUNTY: U.S.A., OREGON, JOSEPHINE COUNTY: Serpentine slopes of Red Mountain, Siskiyou Alpine slopes back of the Oregon Caves. Mountains. Phenology: Spores. Origin: Native. Rocky slopes. First Oregon record. Phenology: Spores. Origin: Native. J. William Thompson 12948 J. William Thompson 12376 26 Jun 1936 29 Jul 1935 WTU-5134JWT, WTU-V-030962 WTU-34971, WTU-V-030967

Dryopteridaceae Dryopteridaceae Polystichum mohrioides (Bory) Presl × Polystichum Polystichum munitum (Kaulf.) C. Presl × Polystichum scopulinum (D.C. Eaton) Maxon scopulinum (D.C. Eaton) Maxon U.S.A., OREGON, JACKSON COUNTY: U.S.A., WASHINGTON, CHELAN COUNTY: Serpentine slopes of Red Mountain, Siskiyou Mountains. Between Drury and Chiwaukum. Rocky slopes. First Oregon record. Phenology: Spores. Origin: Elev. 1700 ft. Native. 47.656962°, -120.729561°; WGS84, uncertainty: 4600 m., Source: Georeferenced, J. William Thompson 12376 29 Jul 1935 Georef'd by WTU Staff Phenology: Spores. Origin: Native. WTU-594JWTa I. C. Otis 1062 Dryopteridaceae 2 Oct 1920 Polystichum lemmonii Underw. WTU-28931, WTU-V-030958 U.S.A., OREGON, JACKSON COUNTY: Serpentine slopes of Red Mountain, Siskiyou Mountains. Rocky slopes. First Oregon record. Phenology: Spores. Origin: Native. Dryopteridaceae J. William Thompson 12376 29 Jul 1935 Polystichum munitum (Kaulf.) C. Presl × Polystichum scopulinum (D.C. Eaton) Maxon WTU-594JWTb U.S.A., WASHINGTON, KITTITAS COUNTY: Dryopteridaceae By trail up Beverly Creek, Wenatchee Mountains. Polystichum lemmonii Underw. × Polystichum Elev. 4000 ft. scopulinum (D.C. Eaton) Maxon 47.372454°, -120.877017°; WGS 84, uncertainty: 10000 m., Source: GeoLocate, U.S.A., WASHINGTON, KITTITAS COUNTY: Georef'd by WTU Staff Along Boulder Creek. Open slope. Phenology: Spores. Origin: Native. Elev. 3500 ft. 47.198258°, -120.4389°; WGS 84, J. William Thompson 6387 uncertainty: 10000 m., Source: GeoLocate, Georef'd by WTU Staff 16 May 1931 Rock slides. Phenology: Spores. Origin: Native. WTU-10211JWT, WTU-V-030960 J. William Thompson 10496 12 May 1934 Dryopteridaceae WTU-139JWT, WTU-V-030963 Polystichum californicum (D.C. Eaton) Diels U.S.A., WASHINGTON, PIERCE COUNTY: [locality and coordinates withheld; status in Washington: G4/S1 (Threatened; WANHP)] Crevices of rocky mountain slopes. No collection date given. Phenology: Spores. Origin: Native. J. B. Flett 1770 WTU-33580, WTU-V-030961 Dryopteridaceae Dryopteridaceae Polystichum munitum (Kaulf.) C. Presl × Polystichum Polystichum munitum (Kaulf.) C. Presl × Polystichum scopulinum (D.C. Eaton) Maxon scopulinum (D.C. Eaton) Maxon U.S.A., WASHINGTON, CHELAN COUNTY: U.S.A., WASHINGTON, KITTITAS COUNTY: Head of Ingall's Creek, Wenatchee Mountains. Between Paris Creek and Boulder Creek, upper Elev. 4000 ft. Cle Elum River valley. On benches above river 47.42361°, -120.30917°; WGS84, uncertainty: and road. 10000 m., Source: GeoLocate, Georef'd by 47.198258°, -120.4389°; WGS 84, WTU Staff uncertainty: 10000 m., Source: GeoLocate, In open rocky slopes. Occasional. Verbatium Georef'd by WTU Staff collection dates: July 27-31, 1931. Phenology: On wooded alluvial benches; mixed parent Spores. Origin: Native. material, thouh predominantly ultramafic rock. Hybrid population. 5639a: P. munitum, vel aff. P. J. William Thompson 7747 scopulinum (meiosis irregular and non-staining spores). 5639b: P. munitum - meiosis 27 Jul 1931 apparently regular; n - 41. 5639c: P. scopulinum. 5639d: P. scopulinum - P. munitum WTU-10208JWT, WTU-V-030959 possible hybrid. Phenology: Spores. Origin: Native. A. R. Kruckeberg 5639 12 Jul 1963 Dryopteridaceae WTU-217982, WTU-V-030952 Polystichum munitum (Kaulf.) C. Presl × Polystichum scopulinum (D.C. Eaton) Maxon Dryopteridaceae U.S.A., WASHINGTON, KITTITAS COUNTY: Polystichum munitum (Kaulf.) C. Presl × Polystichum Slopes along Beverly Creek, Wenatchee scopulinum (D.C. Eaton) Maxon Mountains. Elev. 4000 ft. U.S.A., WASHINGTON, KITTITAS COUNTY: 47.372454°, -120.877017°; WGS 84, Between Paris Creek and Boulder Creek, upper uncertainty: 10000 m., Source: GeoLocate, Cle Elum River valley. On benches above river Georef'd by WTU Staff and road. Rocky slopes. Phenology: Spores. Origin: 47.198258°, -120.4389°; WGS 84, Native. uncertainty: 10000 m., Source: GeoLocate, Georef'd by WTU Staff J. William Thompson 8756 On wooded alluvial benches; mixed parent material, thouh predominantly ultramafic rock. 13 Jul 1932 Hybrid population. Specimen also mounted with 5639 b-d. P. munitum, vel aff. P. scopulinum WTU-10207JWT, WTU-V-030954 (meiosis irregular and non-staining spores). Phenology: Spores. Origin: Native. Dryopteridaceae A. R. Kruckeberg 5639a Polystichum munitum (Kaulf.) C. Presl × Polystichum 12 Jul 1963 scopulinum (D.C. Eaton) Maxon WTU-214838A, WTU-V-030955 U.S.A., WASHINGTON, KITTITAS COUNTY: By trail up Beverly Creek; Wenatchee Mountains. Elev. 4000 ft. Dryopteridaceae 47.372454°, -120.877017°; WGS 84, uncertainty: 10000 m., Source: GeoLocate, Polystichum munitum (Kaulf.) C. Presl Georef'd by WTU Staff Phenology: Spores. Origin: Native. U.S.A., WASHINGTON, KITTITAS COUNTY: Between Paris Creek and Boulder Creek, upper Cle Elum River J. William Thompson 5871 valley. On benches above river and road. 47.198258°, -120.4389°; WGS 84, uncertainty: 10000 m., Source: 20 Aug 1930 GeoLocate, Georef'd by WTU Staff On wooded alluvial benches; mixed parent material, thouh WTU-10210JWT, WTU-V-030956 predominantly ultramafic rock. Hybrid population. Specimen also mounted with 5639 a,c,d. 5639b: P. munitum - meiosis apparently regular; n - 41. Phenology: Spores. Origin: Native. A. R. Kruckeberg 5639b 12 Jul 1963 WTU-214838B Dryopteridaceae Dryopteridaceae Polystichum scopulinum (D.C. Eaton) Maxon Polystichum munitum (Kaulf.) C. Presl × Polystichum scopulinum (D.C. Eaton) Maxon U.S.A., WASHINGTON, KITTITAS COUNTY: Between Paris Creek and Boulder Creek, upper Cle Elum River U.S.A., WASHINGTON, KITTITAS COUNTY: valley. On benches above river and road. Just above serpentine outcrop at the base of 47.198258°, -120.4389°; WGS 84, uncertainty: 10000 m., Source: Esmerelda Peaks in sight of Forest Service GeoLocate, Georef'd by WTU Staff Road 9737. On wooded alluvial benches; mixed parent material, thouh Phenology: Spores. Origin: Native. predominantly ultramafic rock. Hybrid population. Also mounted with 5639 a, b,d. Phenology: Spores. Origin: Native. Elroy Burnett 26 A. R. Kruckeberg 5639c 12 Jul 1963 12 Jun 1988 WTU-214838C WTU-315427, WTU-V-030957

Dryopteridaceae Polystichum kruckebergii W.H. Wagner Dryopteridaceae U.S.A., IDAHO, CUSTER COUNTY: Polystichum kruckebergii W.H. Wagner [locality and coordinates withheld; status in Idaho: S2 (Imperiled; IDNHP)] U.S.A., WASHINGTON, OKANOGAN COUNTY: Elev. 8500 ft. Okanogan National Forest, 27 miles west of Origin: Native. Winthrop, 3 miles from Harts Pass along Pacific Crest Trail between Tatie Peak and Glacier J. William Thompson 14039 Pass. Elev. 6000 ft. 29 Jul 1937 T37N R17E S4; WGS 84, uncertainty: 1000 m., Source: GeoLocate, Georef'd by WTU WTU-10506JWT, WTU-V-016137 Staff Origin: Native. L. Standley 372 Dryopteridaceae 27 Jul 1979 Polystichum imbricans (D.C. Eaton) D.H. Wagner × WTU-281188, WTU-V-016131 Polystichum scopulinum (D.C. Eaton) Maxon Dryopteridaceae U.S.A., WASHINGTON, KITTITAS COUNTY: Between Paris Creek and Boulder Creek, upper Cle Elum River Polystichum kruckebergii W.H. Wagner valley. On benches above river and road. 47.198258°, -120.4389°; WGS 84, uncertainty: 10000 m., Source: U.S.A., WASHINGTON, CLALLAM COUNTY: GeoLocate, Georef'd by WTU Staff Hurricane Ridge, Olympic Mountains. On wooded alluvial benches; mixed parent material, thouh Elev. 6500 ft. predominantly ultramafic rock. Hybrid population. Also mounted with 47.961667°, -123.4575°; NAD 27, uncertainty: 5639 a-c. 5639d: P. scopulinum - P. munitum possible hybrid. 10000 m., Source: Georeferenced, Georef'd Phenology: Spores. Origin: Native. by WTU Staff Origin: Native. A. R. Kruckeberg 5639d 12 Jul 1963 J. William Thompson 14178 WTU-214838D 6 Sep 1937 Dryopteridaceae WTU-10364JWT, WTU-V-016146 Polystichum kruckebergii W.H. Wagner U.S.A., WASHINGTON, JEFFERSON Dryopteridaceae COUNTY: Olympic native monument, Hayden Pass. Polystichum kruckebergii W.H. Wagner Elev. 5000 ft. 47.78278°, -123.34889°; WGS 84, U.S.A., WASHINGTON, OKANOGAN COUNTY: uncertainty: 10000 m., Source: GeoLocate, Between Chopaka Lake and summit of road to Georef'd by WTU Staff Chopaka Mountain. Origin: Native. 48.99944°, -119.72361°; WGS 84, uncertainty: 10000 m., Source: GeoLocate, Harold E. Helmrich 409 Georef'd by WTU Staff Origin: Native. 16 Jul 1934 A. R. Kruckeberg 6450 WTU-306101, WTU-V-016132 17 Jul 1975 WTU-305222, WTU-V-016134 Dryopteridaceae Dryopteridaceae Polystichum kruckebergii W.H. Wagner Polystichum kruckebergii W.H. Wagner U.S.A., WASHINGTON, KLICKITAT COUNTY: CANADA, BRITISH COLUMBIA: Bird creek Meadows, Mount Adams area. [locality and coordinates withheld; status in 45.981988°, -121.263985°; WGS 84, British Columbia: S2S3 (Rare, Blue List; uncertainty: 10000 m., Source: GeoLocate, BCCDC)] Georef'd by WTU Staff [Mixed collection. Also in packet: Kruckeberg Origin: Native. 5738, Polystichum kruckebergii, WTU-V-016133; Kruckeberg 5763, Polystichum A. R. Kruckeberg 5541 lemmonii, WTU-V-016286. Filed under Polystichum kruckebergii.] Phenology: Spores. 1 Sep 1962 Origin: Native. WTU-217549, WTU-V-016139 A. R. Kruckeberg 5728 7 Sep 1963 Dryopteridaceae WTU-316043C, WTU-V-016287 Polystichum kruckebergii W.H. Wagner Dryopteridaceae CANADA, BRITISH COLUMBIA: Polystichum kruckebergii W.H. Wagner [locality and coordinates withheld; status in British Columbia: S2S3 (Rare, Blue List; U.S.A., ALASKA COUNTY: BCCDC)] [locality and coordinates withheld; status in [Mixed collection. Also in packet: Kruckeberg Alaska: S1 (Critically Imperiled; AKNHP)] 5763, Polystichum lemmonii, WTU-V-016286; Elev. 2410 ft. Kruckeberg 5728, Polystichum kruckebergii, Origin: Native. WTU-V-016287. Filed under Polystichum kruckebergii.] Phenology: Spores. Origin: M. C. Stensvold 7259 Native. 9 Jul 1998 A. R. Kruckeberg 5738 WTU-340591, WTU-V-016136 10 Sep 1963 WTU-316043A, WTU-V-016133 Dryopteridaceae Dryopteridaceae Polystichum lemmonii Underw. Polystichum munitum (Kaulf.) C. Presl U.S.A., WASHINGTON, KITTITAS COUNTY: U.S.A., WASHINGTON, PIERCE COUNTY: Edge of serpentine outcrop on east-facing slope South Prairie. of Koppen Mountain ridge, circa 2 miles above 47.13944°, -122.09694°; WGS 84, De Roux Creek, North Fork Teanaway River uncertainty: 10000 m., Source: GeoLocate, drainage. Georef'd by WTU Staff Elev. 5000 ft. Collector's initials may be on sheet, but 47.251427°, -120.87694°; WGS 84, handwriting in unreadable. Phenology: Fertile. uncertainty: 10000 m., Source: GeoLocate, Origin: Native. Georef'd by WTU Staff In shade of mountain hemlock. [Mixed unknown s.n. collection. Also in packet: Kruckeberg 5738, Polystichum kruckebergii, WTU-V-016133; 8 Aug 1898 Kruckeberg 5728, Polystichum kruckebergii, WTU-V-016287. Filed under Polystichum WTU-22840, WTU-V-18265 kruckebergii.] Phenology: Spores. Origin: Native. A. R. Kruckeberg 5763 20 Sep 1963 Dryopteridaceae WTU-316043B, WTU-V-016286 Polystichum kruckebergii W.H. Wagner U.S.A., ALASKA COUNTY: [locality and coordinates withheld; status in Alaska: S1 (Critically Imperiled; AKNHP)] Elev. 2525 ft. Bare ultramafic rock. With few alpine vascular plants grwoing in cracks in otherwise bare ultramafics. Two plants growing on organic material accumulated in grack of bare ultramafic rock. Phenology: Spores. Origin: Native. Tatiana G. Sapozhnikova 027 11 Aug 1997 WTU-358332 Dryopteridaceae Dryopteridaceae Polystichum lonchitis (L.) Roth Polystichum braunii (Spenner) Fée ? U.S.A., WASHINGTON, SKAMANIA COUNTY: U.S.A., ALASKA COUNTY: Davenport, in litt. Near Butterfly Lake. Washington Bay, Kuiu Island. Elev. 4000-5000 ft. 56.721111°, -134.356907°; WGS 84, 46.02331°, -121.84207°; WGS 84, uncertainty: 2400 m., Source: Digital Map, uncertainty: 50000 m., Source: GeoLocate, Georef'd by Joseph Stormer Georef'd by WTU Staff Spruce and hemlock forest. Phenology: Fertile. Steep rocks. Phenology: Spores. Origin: Native. Origin: Native. W. N. Suksdorf 2056 W. J. Eyerdam 8368 24 Aug 1891 25 Aug 1956 WTU-358334, WTU-V-021123 WTU-292743, WTU-V-016103

Dryopteridaceae Dryopteridaceae Polystichum andersonii M. Hopkins Polystichum andersonii M. Hopkins U.S.A., IDAHO, BONNER COUNTY: CANADA, BRITISH COLUMBIA: Priest River Experimental Forest. Queen Charlotte Islands, head of Pallant Creek 48.355678°, -116.787586°; WGS 84, below Mount Moreby west of Cumshewa Inlet, uncertainty: 4000 m., Source: Digital Map, Moresby Island. Georef'd by Melissa Bowers Open alder-coniferous woods near margin of Arborvitae-hemlock veg. zone. open forest. spring run-off creek on stabilized talus below Phenology: Vegetative. Origin: Native. mountain. Common. Phenology: Fertile. R. F. Daubenmire 43360 J. A. Calder 35316 with R. L. Taylor 29 Jul 1943 21 Jun 1964 WTU-358337, WTU-V-016058 WTU-242441, WTU-V-016063

Dryopteridaceae Dryopteridaceae Polystichum munitum (Kaulf.) C. Presl Polystichum andersonii M. Hopkins CANADA, BRITISH COLUMBIA: Sooke River, Victoria. U.S.A., ALASKA COUNTY: Phenology: Vegetative. Origin: Native. Southeast, Baranof Island (west side), Mount Verstovia about 2 miles east of Sitka. F. Comte 5580 Elev. 300 ft. Rocky gully on the lower parts of the mountain 10 Mar 1970 in a blowdown area. Last year's fronds. Phenology: Fertile. Origin: Native. WTU-329881, WTU-V-18267 Mary Clay Muller 3457 13 May 1980 WTU-289176, WTU-V-016051 Dryopteridaceae Polystichum munitum (Kaulf.) C. Presl ? Dryopteridaceae U.S.A., WASHINGTON, KING COUNTY: Polystichum andersonii M. Hopkins Cedar River Watershed. North facing felsenmeer south of Little Mountain (south of U.S.A., ALASKA COUNTY: 100-300 and 390 Roads junction). Southeastern, Baranof Island, Sitka, about 1/3 Elev. 1800 ft. mile up the Indian River Trail. T22N R9E S26 NW, SW; Source: Inferred Elev. 75 ft. from duplicate specimen, Georef'd by Ed Open forest about 100 feet from the river. Alverson Phenology: Fertile. Origin: Native. Margin at edge of shade(?) area. North facing felsenmeer. Phenology: Vegetative. Origin: Mary Clay Muller 4197 Native. 18 Jan 1981 Carla Culver 1-02 WTU-289143, WTU-V-016052 13 Jul 2002 WTU-364396, WTU-V-18284 Dryopteridaceae Dryopteridaceae Polystichum munitum (Kaulf.) C. Presl Polystichum kruckebergii W.H. Wagner U.S.A., WASHINGTON, SAN JUAN COUNTY: CANADA, BRITISH COLUMBIA: San Juan Islands National Wildlife Refuge and [locality and coordinates withheld; status in Wilderness Area. Charles Island. British Columbia: S2S3 (Rare, Blue List; Elev. 25 ft. BCCDC)] 48° 26.583' N, 122° 54.486' W; Source: Elev. 5100 ft. Specimen label Open grassy bank in opening in coniferous Vicinity of Caretaker's house. Occasional on woods. Occasional locally. In pencil on sheet: mossy slopes, north side. Phenology: Spores. "loneh. x mohr. spore slide photographed Origin: Native. indusih[?] slide 13." Phenology: Spores. Origin: Native. Type Specimen: Holotype. Steve Hahn 05-80 J. A. Calder 15550 27 Apr 2005 with D. B. O. Savile, J. M. Ferguson WTU-385143, WTU-V-18334 6 Sep 1954 WTU-173800, WTU-V-000012 Dryopteridaceae Dryopteridaceae Polystichum lonchitis (L.) Roth Polystichum lonchitis (L.) Roth U.S.A., WASHINGTON, CHELAN COUNTY: North Cascades National Park. Base of cliffs on U.S.A., MONTANA COUNTY: northwest side of Reynolds Peak. Just above Glacier National Park; Two Medicine Lake. talus at base of seepy cliffs. . Herb MBG 1,011,462 1,161,629 (mich.) Gray's Elev. 7708 ft. Bot. 37-1. Phenology: Fertile. Origin: Native. 48° 22.504' N, 120° 34.062' W; UTM Zone 10, 680119.009E, 5360628.409N; T33N F. Comte 1765 R18E S11; Source: GPS, UTM calculated from Lat-Lon. 14 Aug 1957 Base of wet cliffs. Very steep. Small micro-benches with lush hanging gardens. WTU-330063, WTU-V-021074 Vegetation very scattered. Loose, piles of rock stacked on slopes. Aspect: north. Slope: 20-90 degrees. Full shade, rock crevice, cool and moist site. Phenology: Fertile. Origin: Native. Dorothy Wallace-Senft 23 23 Jul 2005 Dryopteridaceae with Steve Hahn, Emily Sloane, Peter Zika, Lindsey Hamilton, Laura Potash Polystichum scopulinum (D.C. Eaton) Maxon WTU-366289, WTU-V-016225, NPS accession 764, catalog 32694 U.S.A., WASHINGTON, CHELAN COUNTY: Tumwater Canyon, west of Leavenworth. Dryopteridaceae 47.58472°, -120.67722°; WGS84, uncertainty: 10000 m., Source: GeoLocate, Georef'd by Polystichum munitum (Kaulf.) C. Presl WTU Staff Above climbing rock; growing in a clump of U.S.A. COUNTY: Polystichum imbricans. Phenology: Fertile. [No locality given on label] Origin: Native. Photo of type specimen is stored in type collection. Sheffield Scientific School Daniel C. Eaton Collection. Cited first by D. C. Elroy Burnett s.n. Easton in original description. HOLOTYPE! Aspidium munitum var incisoserratum D. C. Eaton Investigation of North American 15 May 1991 Polystichum, D. H. Wagner, 1976. Washington State University. Phenology: Spores. WTU-340736, WTU-V-021106 Dr. Lyall s.n. 1858 WTU-111322, WTU-V-000010 Dryopteridaceae Dryopteridaceae Polystichum scopulinum (D.C. Eaton) Maxon Polystichum munitum (Kaulf.) C. Presl U.S.A., OREGON, LAKE COUNTY: U.S.A., WASHINGTON, SAN JUAN COUNTY: Fremont National Forest. At the Palisade Rocks San Juan Islands. Sentinel Island. on southeast side of Gearhart Mountain. 48° 38.4' N, 123° 9.139' W Elev. 6478 ft. West end of island. Rare in Douglas fir forest. 42° 27.904' N, 120° 48.557' W Two plants. Phenology: Spores. Origin: Native. Eroding rock pinnacles with loose, mostly bare rocky soil. Scattered Pinus. Uncommon. In Eliza Habegger EH-1074 narrow, moist, shaded rock crevices on east-facing outcrop. Phenology: Fertile. Origin: 27 Apr 2006 Native. WTU-386173, WTU-V-18279 Ben Legler 1932 with Sarah Gage, Wendy Gibble, Robert Goff 3 Jul 2004 Dryopteridaceae WTU-369519, WTU-V-021045 Polystichum imbricans (D.C. Eaton) D.H. Wagner ssp. imbricans Dryopteridaceae U.S.A., WASHINGTON, CHELAN COUNTY: Lake Chelan National Recreation Area. Polystichum munitum (Kaulf.) C. Presl Outcrops on south side of Coon Lake. One kilometer northeast of High Bridge. Stehekin U.S.A., WASHINGTON, LEWIS COUNTY: River Valley. Logging road on west side of Highway 97 Elev. 2200 ft. between Carlson and Divide, circa 10 road miles 48° 23' 13" N, 120° 49' 52.1" W; UTM Zone north of Morton. 10, 660569.263E, 5361581.615N; WGS 84, 46° 40' 55" N, 122° 11' 54" W; WGS 84, uncertainty: 8 m., Source: GPS, UTM uncertainty: 1000 m., Source: Digital map calculated from Lat-Lon. Understory. Fronds in large clumps; common. Open dry rocky outcrops in open Pseudotsuga Phenology: Spores. Origin: Native. forest. With Archtostaphylos nevadensis, Calamagrostis rubescens and Stipa. Uncommon Jessica Ni 33 on shaded outcrop. In clumps. Phenology: with Ben Legler Spores. Origin: Native. 16 Jun 2006 Steve Hahn 06-210 29 Jun 2006 WTU-362942, WTU-V-18282 WTU-366771, WTU-V-016194 Dryopteridaceae Dryopteridaceae Polystichum imbricans (D.C. Eaton) D.H. Wagner ssp. imbricans Polystichum imbricans (D.C. Eaton) D.H. Wagner ssp. imbricans U.S.A., WASHINGTON, CHELAN COUNTY: Lake Chelan National Recreational Area. Base U.S.A., WASHINGTON, CHELAN COUNTY: of Purple Creek Pass Trail where trail passes Lake Chelan National Recreational Area. waterfall. 300 meters east of Stehekin. Stehekin State Airstrip. Elev. 1280 ft. Elev. 1196 ft. UTM Zone 10, 673977E, 5353360N; WGS 84, 48.34332° N, 120.71854° W; UTM Zone 10, Source: Georeferenced, Georef'd by WTU 669049.268E, 5356975.232N; WGS 83, Staff uncertainty: 5 m., Source: GPS, UTM Open mixed conifer forest with shady mesic calculated from Lat-Lon. rocky understory along perennial stream. In Flat mowed area approximately 3,000 feet long clumps in rock crevices and rocky soil. by 300 feet wide. Public airstrip cut out of valley Phenology: Spores. Origin: Native. floor forest. Abies grandis, Alnus rubra, Acer macrophylla, Pseudotsuga menziesii dominate Jennette Timmer 406 forest edge. Airstrip dominated by non-native grasses. Uncommon among rocks along 28 Jun 2006 roadside. Phenology: Spores. Origin: Native. WTU-366867, WTU-V-016193 Dale Blum 06-36 27 Jun 2006 WTU-366662, WTU-V-016196 Dryopteridaceae Dryopteridaceae Polystichum lonchitis (L.) Roth Polystichum imbricans (D.C. Eaton) D.H. Wagner ssp. imbricans U.S.A., WASHINGTON, PIERCE COUNTY: Mount Rainier National Park. Ipsut Pass circa 1 U.S.A., WASHINGTON, PIERCE COUNTY: mile north of Mowich Lake. Mount Rainier National Park; rock outcrop just west of pass south of Elev. 5150 ft. Owyhigh Lakes. 46° 57.039' N, 121° 52.09' W; UTM Zone 10, Elev. 5490 ft. 586127.103E, 5200301.74N; WGS 84, 46° 51.774' N, 121° 35.499' W; UTM Zone 10, 607343.509E, uncertainty: 7 m., Source: GPS, UTM 5190891.659N; WGS 84, uncertainty: 16 m., Source: GPS, UTM calculated from Lat-Lon. calculated from Lat-Lon. Steep northeast-facing brushy cirque with Small east-facing rock outcrop in mix of subalpine meadows and Rubus, Sorbus, Valeriana, Veratrum, topped by small Chamaecyparis forest. Fronds in tufts in rock crevices. Single steep dry rock faces and subalpine conifer plant found. Phenology: Spores. Origin: Native. forest. Scattered on rocky areas under shrubs. Phenology: Spores. Origin: Native. Pete Del Zotto 50 20 Jul 2006 with Ben Legler, David Giblin Lucia Harrison 06-23 Herbarium: WTU 8 Aug 2006 WTU-366687, WTU-V-016224 Dryopteridaceae Polystichum lonchitis (L.) Roth Dryopteridaceae U.S.A., IDAHO, SHOSHONE COUNTY: Clearwater National Forest. Hoodoo Pass, Polystichum andersonii M. Hopkins South Trailhead parking lot. Elev. 5993 ft. U.S.A., WASHINGTON, PIERCE COUNTY: 46.97483°, -115.02681°; WGS 84, Mount Rainier National Park. Along the uncertainty: 500 m., Source: Nat. Geog. Wonderland Trail circa 0.25 mile northeast of TOPO!, Georef'd by John Bassett Ipsut Pass. Rocky meadow surrounded by forest of Tsuga, Elev. 4600 ft. Abies and Picea. Artemisia species and 46° 57.19' N, 121° 51.981' W; UTM Zone 10, Penstemon dominant. In darkest, dampest rock 586261.3E, 5200583.399N; WGS 84, crevices beneath overhanging rock. Phenology: uncertainty: 10 m., Source: GPS, UTM Vegetative. Origin: Native. calculated from Lat-Lon. Steep, brushy, rocky northeast-facing cirque Mary Johanson 05-37 bordered by cliffs, with Alnus sinuata, Aruncus with Robert Goff, Doug Williams, Joe Johanson, dioicus, Rubus parviflorus, Sorbus sitchensis. Jessie Johanson, Nomi Odano, Julia Odano, Fronds tufted from large rhizome; scattered and Ben Legler, David Giblin uncommon under Alnus sinuata. Phenology: Spores. Origin: Native. 8 Jul 2005 Lara Massey 06-21 8 Aug 2006 WTU-369594, WTU-V-021051 WTU-366664, WTU-V-016054 Dryopteridaceae Dryopteridaceae Polystichum munitum (Kaulf.) C. Presl Polystichum lonchitis (L.) Roth U.S.A., WASHINGTON, SAN JUAN COUNTY: U.S.A., WASHINGTON, PIERCE COUNTY: San Juan Islands. Flattop Island. Weed Point on Mount Rainier National Park; Steep rock face about 1/8 mile due northeast end of Flattop Island; northern part of west of Owyhigh Lakes. San Juan Archipelago. Elev. 5600 ft. Elev. 0-10 ft. 46° 52.072' N, 121° 35.422' W; UTM Zone 10, 607431.41E, 48° 38.871' N, 123° 4.707' W; WGS 84, 5191445.319N; WGS 84, uncertainty: 3 m., Source: GPS, UTM uncertainty: 2 m. calculated from Lat-Lon. Rocky open point, treeless, with thin soil, with Steep east-facing rocky slope with Phlox diffusa, Delphinium Rosa nutkana, Grindelia, Lathyrus japonicus, glareosum, Aster ledophyllus, Vaccinium, Chamaecyparis grasses. Scattered on forest edge. Leaf material nootkatensis. Scattered in clumps in rock crevices and stable scree. removed for RADseq analysis by Megan Ruffley Phenology: Spores. Origin: Native. at WTU, February 2016. Phenology: Spores. Origin: Native. Lindsey Koepke 06-2 18 Jul 2006 with Ben Legler, David Giblin, Diane Doss Carrie Marschner 06-55 Herbarium: WTU 31 May 2006 WTU-388045, WTU-V-019144 Dryopteridaceae Dryopteridaceae Polystichum munitum (Kaulf.) C. Presl Polystichum lemmonii Underw. U.S.A., WASHINGTON, SAN JUAN COUNTY: U.S.A., WASHINGTON, SNOHOMISH San Juan Islands. McConnell Island, middle of COUNTY: island; southwest of Orcas Island. East side of Devil's Thumb, in rocky bowl below Elev. 40 ft. southern point on ridge, above Windom Lake. 48° 35.8' N, 123° 1.45' W; WGS 84, Elev. 4590 ft. uncertainty: 100 m., Source: GPS 48° 7' 29" N, 121° 32' 34" W; NAD 27, Shady Pseudotsuga forest; most trees less than Source: Digital map ca. [circa] 50 years old; grassy or duff Open rocky east-facing subalpine slope understory; grazed in past. Scattered in forest. composed of dunite, with large talus below; soil Phenology: Spores. Origin: Native. thin, rocky; with Adiantum aleuticum, Polystichum lemmonii, grasses. Common in soil David Giblin 489 derived from dunite; stems clumped. Phenology: Spores. Origin: Native. 30 May 2006 Ben Legler 2368 WTU-385133, WTU-V-18278 26 Sep 2004 WTU-382517, WTU-V-016214 Dryopteridaceae Polystichum munitum (Kaulf.) C. Presl Dryopteridaceae U.S.A., WASHINGTON, JEFFERSON Polystichum munitum (Kaulf.) C. Presl COUNTY: Protection Island. U.S.A., WASHINGTON, KITTITAS COUNTY: Elev. 178 ft. Near stream along Forest Road 46, ca. [circa] 2 48° 7.733' N, 122° 55.692' W miles west of Cooper Pass. A tiny remnant native prairie near the forest. Elev. 2765 ft. Collections from both habitats. Occasional 47° 25.544' N, 121° 13.942' W under shrubs in Pseudotsuga forest. Phenology: Roadside in large rocks with some soil; other Spores. Origin: Native. plants: Pteridium aquilinum, Taxus brevifolia, Linnaea borealis, Vaccinium membranaceum. Fred Weinmann 06-1000 Uncommon. Phenology: Spores. Origin: Native. 3 May 2006 Don Knoke 766 WTU-371089, WTU-V-18280 16 Jun 2005 WTU-364833, WTU-V-18337 Dryopteridaceae Polystichum munitum (Kaulf.) C. Presl Dryopteridaceae U.S.A., WASHINGTON, SKAGIT COUNTY: Padilla Bay National Estuarine Research Polystichum andersonii M. Hopkins Reserve. Saddlebag Island. Elev. 17 ft. CANADA, BRITISH COLUMBIA: 48° 32.117' N, 122° 33.333' W; WGS 84, Northeast shore of Princess Royal Island, on Source: GPS southwest side of McKay Reach, abandoned High island with rocky shoreline cliffs; Douglas grounds of Butedale Cannery. fir/Madrone forests with Snowberry, Mahonia; Elev. 49 ft. forest and understory. Frequent in understory. 53° 9.7' N, 128° 41.7' W Leaf material removed for RADseq analysis by Partial shade of Rubus spectablilis, Megan Ruffley at WTU, February 2016. shrub-invaded mossy clearing, abandoned Phenology: Spores. Origin: Native. grounds. Rare, cespitose, semi-evergreen. Phenology: Spores. Origin: Native. Ben Legler 3378 Peter F. Zika 22741 23 May 2006 27 May 2006 WTU-371020-A, WTU-V-019145 WTU-365137, WTU-V-016071 Dryopteridaceae Dryopteridaceae Polystichum braunii (Spenner) Fée Polystichum lemmonii Underw. U.S.A., ALASKA COUNTY: U.S.A., WASHINGTON, KITTITAS COUNTY: In town of Sitka, near junction of Sawmill Creek Beverly Creek Trail. Road and Raptor Way, Baranof Island. Elev. 4920 ft. Elev. 49 ft. 47° 24.44' N, 120° 52.53' W 57° 3.1' N, 135° 18.9' W Serpentine, rocky moderate slope. Dry Partly shaded thickets, with Alnus rubra, Tsuga south-facing sunny slope. Pseudotsuga and heterophylla, Ribes laxiflorum, Athyrium Abies lasiocarpa forest. Growing at base of rock filix-femina, Aruncus. Cespitose, uncommon, outcrops. common fern. Phenology: Spores. last years leaves beaten down by snow and still Origin: Native. slightly green. SHEET 2 OF 2. Phenology: Spores. Origin: Native. Valerie Soza 1896 with Sheila Olmstead, Keri Cardon, Richard G. Peter F. Zika 22743 Olmstead, Maria Gerace, Ryan Miller, Linda Brooking, Cole Brooking 31 May 2006 7 Jul 2006 WTU-365133, WTU-V-016095 WTU-371614, WTU-V-016212

Dryopteridaceae Dryopteridaceae Polystichum braunii (Spenner) Fée Polystichum munitum (Kaulf.) C. Presl U.S.A., ALASKA COUNTY: In town of Sitka, near junction of Sawmill Creek U.S.A., WASHINGTON, SAN JUAN COUNTY: Road and Raptor Way, Baranof Island. Reef Island. Elev. 49 ft. 48° 36.179' N, 123° 1.021' W 57° 3.1' N, 135° 18.9' W Large island dominated by Pseudotsuga Partly shaded thickets, with Alnus rubra, Tsuga menziesii; open understory with Elymus and heterophylla, Ribes laxiflorum, Athyrium Holodiscus. Occasional in forest understory. filix-femina, Aruncus. Cespitose, uncommon, Leaf material removed for RADseq analysis by last years leaves beaten down by snow and still Megan Ruffley at WTU, February 2016. slightly green. SHEET 1 OF 2. Phenology: Phenology: Spores. Origin: Native. Spores. Origin: Native. Peter Dunwiddie 07-94 Peter F. Zika 22743 with Phil Green, Megan Jensen, David Giblin, Ted Thomas 31 May 2006 19 Sep 2007 WTU-365132, WTU-V-016112 WTU-375789, WTU-V-019143

Dryopteridaceae Dryopteridaceae Polystichum lonchitis (L.) Roth Polystichum munitum (Kaulf.) C. Presl U.S.A., WASHINGTON, PIERCE COUNTY: Mount Rainier National Park. Crystal Lakes area, just inside Park U.S.A., WASHINGTON, SAN JUAN COUNTY: boundary, about 0.75 kilometer west from junction with Pacific Crest Castle Island. Trail. 48.421° N, 122.823° W; WGS 84, Source: Elev. 6271 ft. Digital map 46.90381° N, 121.50206° W; WGS 84, uncertainty: 8 m. West side is high quality Festuca rubra Southwest facing subalpine meadow dominated by Lupinus latifolius, grassland on rocky benches with a diverse array and adjacent steep talus slope and rock cliffs. Growing at base of of forbs, Koeleria, and Elymus. Top is thickets of rock face in crevices. Phenology: Spores. Origin: Native. Arctostaphylos, Symphoricarpos, Rosa nutkana, Prunus emarginata, and Pseudotsuga Vicky Weafer 07-19 13 Aug 2007 menziesii. A few plants on north-facing slope. with Dan Paquette, David Giblin, Megan A. Jensen, Richard Leaf material removed for RADseq analysis by Ramsden Megan Ruffley at WTU, February 2016. Phenology: Spores. Origin: Native. Herbarium: WTU Barbara French 07-22 with David Giblin, Megan A. Jensen, Peter Dunwiddie, Ted Thomas, Phil Green 21 Sep 2007 WTU-375578, WTU-V-019146 Dryopteridaceae Dryopteridaceae Polystichum munitum (Kaulf.) C. Presl Polystichum braunii (Spenner) Fée U.S.A., WASHINGTON, SKAGIT COUNTY: U.S.A., ALASKA, HAINES COUNTY: Padilla Bay National Estuarine Research Tongass National Forest. Reserve. Dot Island. 58° 59' 23.609" N, 135° 24' 2.84" W Elev. 69 ft. Rocky floodplain dominated by Alnus and 48° 32.0412' N, 122° 33.2112' W; Source: Populus. Understory of Pyrola secunda, Digital map Dryopteris expansa, and Calamagrostis Sloping island with high end at north end. Rich canadensis. Rare; growing in rocky floodplain slopes of Festuca rubra, Plectritis, Cerastium, under deciduous canopy. Phenology: Spores. Camassia, and Zygadenus. Juniperus are Origin: Native. common, very shaped and stunted by salt spray, little deer browse. No geese nesting; otter Jeff Walker 210 use on southwest side. A couple of small plants with Kristin Marsh under shrubs. Phenology: Fertile. Origin: Native. 30 Jul 2004 Ben Legler 3785 WTU-370032, WTU-V-016111 1 May 2007 WTU-377330, WTU-V-18341 Dryopteridaceae Polystichum sp. Dryopteridaceae U.S.A., ALASKA, HAINES COUNTY: Polystichum munitum (Kaulf.) C. Presl Tongass National Forest. 58° 59' 23.609" N, 135° 24' 2.84" W U.S.A., WASHINGTON, SKAGIT COUNTY: Rocky floodplain dominated by Alnus and Padilla Bay National Estuarine Research Populus. Understory of Pyrola secunda, Reserve. Southwest side of Hat Island. Dryopteris expansa, and Calamagrostis Elev. 32 ft. canadensis. Rare; growing in rocky floodplain 48° 31.37' N, 122° 33.098' W; WGS 84 under deciduous canopy. Phenology: Spores. Steep grassy slopes with cliffs and rock outcrops. Abundant Elymus glaucus, Koeleria Jeff Walker 209 macrantha, Festuca rubra, and Zigadenus with Kristin Marsh venenosus. Common in Pseudotsuga menziesii forest understory. Phenology: Spores. Origin: 30 Jul 2004 Native. WTU-370086, WTU-V-030953 Peter Dunwiddie 07-4 30 Apr 2007 Dryopteridaceae WTU-377281, WTU-V-18281 Polystichum lonchitis (L.) Roth Dryopteridaceae U.S.A., ALASKA, HAINES COUNTY: Tongass National Forest. Polystichum munitum (Kaulf.) C. Presl 58° 59' 23.609" N, 135° 24' 2.84" W Rocky floodplain dominated by Alnus and U.S.A., WASHINGTON, SNOHOMISH Populus. Understory of Pyrola secunda, COUNTY: Dryopteris expansa, and Calamagrostis Southwest of Bryant. canadensis. Infrequent; growing in mossy rocks 48.23917°, -122.15667°; WGS 84, in old rocky floodplain dominated by deciduous uncertainty: 10000 m., Source: GeoLocate, trees. Phenology: Spores. Origin: Native. Georef'd by WTU Staff Deep conifer woods. Abundant. [Located in Jeff Walker 211 teaching collection]. Phenology: Spores. Origin: with Kristin Marsh Native. 30 Jul 2004 M. Arnot 110 WTU-370085, WTU-V-021120 20 May 1988 WTU-314697, WTU-V-005722 Dryopteridaceae Dryopteridaceae Polystichum munitum (Kaulf.) C. Presl Polystichum andersonii M. Hopkins U.S.A., WASHINGTON, SAN JUAN COUNTY: U.S.A., OREGON, WALLOWA COUNTY: Little Sucia Island, off the southwest corner of West side of Lostine River, Lostine River Sucia Island. canyon, Wallowa Mountains. 48° 45.283' N, 122° 55.198' W; Source: GPS Elev. 5260 ft. Includes forested uplands (circa 80 feet high) T3S R43E S25 NW 1/4 of NW 1/4; NAD 27, with shrub understory, steep sandstone cliffs, uncertainty: 805 m., Source: TRS2LL, eroding dirt bluffs. Linked by a low, Georef'd by Ben Legler shrub-covered (Rosa nutkana) shell-midden Full shade, among granitic rocks in dry berm. Uncommon in forest understory. creekbed, with Alnus sinuata, Galium triflorum. Phenology: Spores. Origin: Native. All fronds with buds in axils of upper pinnae, one large evergreen plant (old fronds collected). Erica Nees 08-24 Phenology: Spores. Origin: Native. 23 Apr 2008 P. F. Zika 11793 with E. R. Alverson WTU-393526, WTU-V-18340 29 Jul 1992 Dryopteridaceae WTU-369795, WTU-V-016074 Polystichum munitum (Kaulf.) C. Presl U.S.A., WASHINGTON, SAN JUAN COUNTY: Dryopteridaceae Matia Island. 48.74791° N, 122.84196° W; Source: GPS Polystichum munitum (Kaulf.) C. Presl Vicinity of campsites at west end of island. Large mixed conifer forest, with several open, U.S.A., WASHINGTON, SAN JUAN COUNTY: disturbed campsites. Common and widespread Ewing Island. Long, narrow island off NE end of in Pseudotsuga menziesii understory. Leaf Sucia Island. Steep sandstone ridges run length material removed for RADseq analysis by of island. Megan Ruffley at WTU, February 2016. 48.76455° N, 122.88093° W; WGS 84, Phenology: Spores. Origin: Native. Source: Digital map Mostly forested with PSME, ARME, and JUMA. Peter Dunwiddie 08-16 Grassy balds along south shore and at east end of island. A few plants scattered through forest. 24 Apr 2008 Phenology: Spores. Origin: Native. WTU-377297, WTU-V-019142 Janneke Hille-Ris Lambers JHRL 08-72 28 May 2008 Dryopteridaceae WTU-396167, WTU-V-18339 Polystichum munitum (Kaulf.) C. Presl U.S.A., WASHINGTON, SAN JUAN COUNTY: Dryopteridaceae San Juan Islands. West Matia #1. Large island at north side of harbor, west end of Matia Island. Polystichum munitum (Kaulf.) C. Presl 48.74946° N, 122.84378° W; WGS 84, Source: Digital map U.S.A., WASHINGTON, SAN JUAN COUNTY: Cliffy sandstone island rising steeply out of cove San Juan Islands National Wildlife Refuge and at west end of Matia Island. Approximately 50 Wilderness Area. Turn Island. Large island 0.25 feet high, with Pseudotsuga menziesii and miles off the east shore of San Juan Island. Arbutus menziesii forest across top. 1 plant. 48° 31.98' N, 122° 58.22' W; WGS 84, Phenology: Spores. Origin: Native. uncertainty: 10 m., Source: GPS On dry rocks. Occasional under Pseudotsuga. Megan A. Jensen 567 Phenology: Spores. Origin: Native. 24 Apr 2008 Eliza Habegger EH1094 WTU-376226, WTU-V-18283 29 May 2008 WTU-395102, WTU-V-18338 Dryopteridaceae Dryopteridaceae Polystichum munitum (Kaulf.) C. Presl Polystichum munitum (Kaulf.) C. Presl U.S.A., WASHINGTON, SAN JUAN COUNTY: U.S.A., WASHINGTON, SAN JUAN COUNTY: San Juan Islands National Wildlife Refuge and North Finger Island. Wilderness Area. Aleck Rock. South of Lopez Elev. 0-65 ft. Island. 48° 45' 24.18" N, 122° 53' 27.64" W Elev. 0-32 ft. 0.8 mile-long island dominated by Pseudotsuga 48° 25.399' N, 122° 51.038' W menziesii overstory. Coastal bluff habitat on Island includes two islets separated by a gravel south and extreme eastern edge of island. berm which probably is submerged at very high Common in forest understory. Phenology: tides. Rare on ledge near west end of island. Spores. Origin: Native. Phenology: Spores. Origin: Native. David Giblin 1507 Phil Green 08-54 13 May 2008 30 May 2008 WTU-385873, WTU-V-18330 WTU-394513, WTU-V-18342

Dryopteridaceae Dryopteridaceae Polystichum munitum (Kaulf.) C. Presl Polystichum braunii (Spenner) Fée U.S.A., WASHINGTON, KING COUNTY: CANADA, BRITISH COLUMBIA: Corner of 22nd Avenue Northeast and East Cheekeye, 7 miles north off river road. Louisa Street, Montlake, Seattle. 49° N, 123° E; Misleading coordinates given Elev. 65 ft. only to nearest degree. 47° 38.5' N, 122° 18.2' W; T25N R4E S21 Rare in rich woods, Oplopanax community. No Shaded fenceline, damp irrigated soil, with bulblets. Long: 123 NE. Phenology: Spores. Athyrium filix-femina, Ranunculus repens. North Origin: Native. aspect. Cespitose, evergreen. Phenology: Spores. Origin: Native. T. M. C. Taylor 6557 Peter F. Zika 18932 24 May 1963 2 Sep 2003 WTU-371161, WTU-V-016100 WTU-371533, WTU-V-18331

Dryopteridaceae Dryopteridaceae Polystichum munitum (Kaulf.) C. Presl Polystichum munitum (Kaulf.) C. Presl CANADA, BRITISH COLUMBIA: Burnett Bay, south third of beach. U.S.A., WASHINGTON, SNOHOMISH Elev. 0 ft. COUNTY: 51° 6.8' N, 127° 40.2' W; WGS 84 Along Mountain Loop Highway, north side, at In sand on rotted driftwood mound at head of Verlot Ranger Station, Robe, west side of highly sea-exposed beach; under small Picea Cascade Mountains. sitchensis. Sea level, flat. Phenology: Spores. Elev. 951 ft. Origin: Native. 48° 5.5' N, 121° 48.8' W; T30N R8E S8 Deep shade of Pseudotsuga menziesii, Thuja Eric Lamb 2002-262 plicata, Tsuga heterophylla, second growth with Sarah Martz conifer stand. Cespitose, evergreen, common. Phenology: Spores. Origin: Native. 28 Jun 2002 Peter F. Zika 19081 WTU-371215, WTU-V-18432 20 Sep 2003 WTU-371518, WTU-V-18427 Dryopteridaceae Polystichum andersonii M. Hopkins CANADA, BRITISH COLUMBIA: 2 miles west on highway #16. Elev. 0 ft. 54° 15' N, 129° 35' W Rocky slope. Phenology: Spores. Origin: Native. T. M. C. Taylor 6975 14 Jun 1966 WTU-371204 Dryopteridaceae Dryopteridaceae Polystichum lonchitis (L.) Roth Polystichum lemmonii Underw. U.S.A., WASHINGTON, SNOHOMISH U.S.A., WASHINGTON, KITTITAS COUNTY: COUNTY: Iron Peak Trail, 1 air mile from trailhead, Perry Creek drainage, circa 1 mile above Wenatchee National Forest, Wenatchee trailhead and northeast end of Forest Service Mountains. Road 4063, northwest slope of Dickerman Elev. 5838 ft. Mountain, west slope of Cascade Mountains, 47° 25.3' N, 120° 54.5' W Mount Baker-Snoqualmie National Forest. Barren serpentine slope, full sun, with Anemone Elev. 2296 ft. drummondii, Thlaspi montanum, Erysimum 48° 4.5' N, 121° 29.2' W; T30N R10E S23 arenicola, dwarf Abies lasiocarpa; west aspect. Mossy shaded talus, north aspect, with Acer Cespitose. Phenology: Spores. Origin: Native. circinatum, Athyrium filix-femina, Polystichum munitum, P. andersonii. Cespitose, dark green, Peter F. Zika 17041 evergreen, occasional. Phenology: Spores. Origin: Native. 17 Jul 2002 Peter F. Zika 19085 WTU-371961, WTU-V-016213 20 Sep 2003 WTU-371519, WTU-V-021053 Dryopteridaceae Polystichum lemmonii Underw. Dryopteridaceae U.S.A., WASHINGTON, KITTITAS COUNTY: Polystichum munitum (Kaulf.) C. Presl Circa 2 miles up Iron Peak Trail, Wenatchee National Forest, Wenatchee Mountains. U.S.A., WASHINGTON, PACIFIC COUNTY: Elev. 5543 ft. Bruceport County Park campground. 47° 25.3' N, 120° 54.8' W Elev. 131 ft. Crevice in serpentine ledge, full sun, with 46° 40.9' N, 123° 53.7' W; T14N R10W S22 Anemone drummondii, Poa curtifolia, Forested bluff over the ocean, shade of Tsuga heterophylla, Picea Eriogonum pyrolifolium, adjacent forest an open sitchensis, Blechnum spicant, Gaultheria shallon, Rubus parviflorus, stand of Abies lasiocarpa, Pinus contorta, P. R. spectablis, Sambucus racemosa. Cespitose, evergreen, common. albicaulis, Tsuga mertensiana; northwest Phenology: Spores. Origin: Native. Cultivated. aspect. Cespitose. Phenology: Spores. Origin: Native. Peter F. Zika 18934 3 Sep 2003 Peter F. Zika 17040 WTU-371782 17 Jul 2002 Dryopteridaceae WTU-371962, WTU-V-016211 Polystichum munitum (Kaulf.) C. Presl Dryopteridaceae U.S.A., WASHINGTON, PACIFIC COUNTY: Bruceport County Park campground. Polystichum munitum (Kaulf.) C. Presl Elev. 131 ft. 46° 40.9' N, 123° 53.7' W; T14N R10W S22 U.S.A., WASHINGTON, KING COUNTY: Forested bluff over the ocean, shade of Tsuga Southeast side of Tinkham Peak are near heterophylla, Picea sitchensis, with Blechnum Twilight Lake, at end of 565 Road. spicant, Gaultheria shallon, Rubus parviflorus, Elev. 3600 ft. R. spectablis, Sambucus racemosa. Cepitose, 47.33° N, 121.43° W; T21N R11E S5 evergreen, common. Phenology: Spores. Origin: Phenology: Spores. Origin: Native. Native. Julia Bent 3-2 Peter F. Zika 18934 with Jeannette Banobi, Tyler Marschall, Helmut with D. Knoke, David Giblin Isenmann, Dorte Isenmann 3 Sep 2003 6 Sep 2008 WTU-371770, WTU-V-18423 WTU-372387, WTU-V-18336 Dryopteridaceae Dryopteridaceae Polystichum lonchitis (L.) Roth Polystichum imbricans (D.C. Eaton) D.H. Wagner U.S.A., WASHINGTON, CHELAN COUNTY: U.S.A., WASHINGTON, KLICKITAT COUNTY: North Cascades National Park. 1 km west of Fisher Hill Rd. Top of U.S. BLM property and Heather Pass. southeast facing, open slope below. Elev. 6366 ft. Elev. 728 ft. UTM Zone 10, 664210E, 5375047N; WGS 84, 45.72258° N, 121.25898° W; WGS 84, uncertainty: 500 m., Source: GeoLocate, uncertainty: 5 m., Source: GPS Georef'd by WTU Staff, Invalid coordinates Largely open, probably overgrazed slope with (located outside county bounds). mineral soil, rock outcroppings, and Quercus Subalpine meadow comprising low wet area fed garryana forest at base. Rhus toxicodendron by snowmelt and higher areas with steep, rocky abundant. Uncommon in cool rock crevice. outcroppings. Scattered in moist pockets of Phenology: Spores. Origin: Native. rocky outcropping. Phenology: Spores. Origin: Native. David Giblin 2753 with Steve Hahn Mignonne Bivin 08-12 with Cheryl Lowe, Maria Gerace, David Giblin 13 May 2009 16 Aug 2008 WTU-378817, WTU-V-016174 WTU-374312, WTU-V-021119 Dryopteridaceae Dryopteridaceae Polystichum munitum (Kaulf.) C. Presl Polystichum munitum (Kaulf.) C. Presl U.S.A., OREGON, COLUMBIA COUNTY: U.S.A., WASHINGTON, SAN JUAN COUNTY: Southwest shore of Columbia River, Diblee San Juan Islands National Wildlife Refuge and Point, north of Rinearson Slough. Wilderness Area. Willow Island. A large, 100' Elev. 16 ft. high rocky island 1/3 mile west of Blakely Island. 46° 6.5' N, 122° 58.95' W; T7N R2W S6 Elev. 100 ft. In dry sandy floodplain forest, shade of 48.5402°, -122.82242°; WGS 84, Source: Popopulus balsamifera. Cespitose, infrequent. Georeferenced, Georef'd by WTU Staff Phenology: Spores. Origin: Native. Partially treed with Pseudotsuga and Arbutus. Open understory dominated by Elymus glaucus Peter F. Zika 19065 and Symphoricarpos albus. East side has extensive cliffs and ledges and is particularly 18 Sep 2003 rich with ferns. Considerable deer browse. Rock appears to be serpentine. Occasional, scattered WTU-379141, WTU-V-18399 under Pseudotsuga, across island. Phenology: Spores. Origin: Native. Steve Hahn 08-79 25 Sep 2008 Dryopteridaceae WTU-392569, WTU-V-18322 Polystichum munitum (Kaulf.) C. Presl U.S.A., WASHINGTON, COWLITZ COUNTY: Dryopteridaceae Solo Slough, Mount Solo Road, 100 meters north of junction with Route 432. Polystichum munitum (Kaulf.) C. Presl Elev. 32 ft. 46° 9.8' N, 123° 1.5' W; T8N R3W S23 U.S.A., WASHINGTON, SAN JUAN COUNTY: Shore. Cespitose, evergreen. Phenology: San Juan Islands. Gossip Island. Spores. Origin: Native. Elev. 25 ft. 48° 39.789' N, 123° 10.374' W Peter F. Zika 19016 Low conglomerate and sandstone bedrock island with gently sloping bedrock on all sides. 10 Sep 2003 Scattered madrone, Douglas fir, and junipers across island with lush camas on swales WTU-379197, WTU-V-18397 between outcrop. One plant under shrubs. Phenology: Spores. Origin: Native. Brianne Cohen 09-73 Dryopteridaceae 23 Sep 2009 Polystichum munitum (Kaulf.) C. Presl WTU-403759, WTU-V-012789 U.S.A., WASHINGTON, LEWIS COUNTY: West side of I-5, east side of Knowles Road, 1 road kilometer south of Winlock-Toledo Road, Drews Prairie. Elev. 295 ft. 46° 28' N, 122° 53.3' W; T11N R2W S2 In shade of Amelanchier alnifolia, Rhamnus purshiana, Quercus garryana, Fraxinus latifolia, wet prairie remnant. Cespitose, evergreen, occasional. Phenology: Spores. Origin: Native. Peter F. Zika 18994 10 Sep 2003 WTU-379190 Dryopteridaceae Dryopteridaceae Polystichum scopulinum (D.C. Eaton) Maxon Polystichum imbricans (D.C. Eaton) D.H. Wagner ssp. imbricans U.S.A., IDAHO, IDAHO COUNTY: Nez Perce National Forest, Gospel Peak region, U.S.A., WASHINGTON, WHATCOM COUNTY: 1 1/4 mile southwest of Middle Knob Lake. Ross Lake National Recreation Area. Area T26N R4E SSW; WGS 84, uncertainty: 1000 between Ponderosa and Lightning Creek m., Source: GeoLocate, Georef'd by WTU campgrounds. Staff Elev. 485 m. Rocks west end of meadow. Occasional in 48.8682° N, 121.0174° W; WGS 84, protected areas. Phenology: Spores. Origin: uncertainty: 100 m., Source: Digital map Native. Mixture of grassy balds, light forest overstory, and seasonally inundated lake bottom. Douglass Henderson 3394 Uncommon; single plant on moist, west facing outcropping in forest understory. Phenology: 27 Jul 1976 Spores. Origin: Native. WTU-376099, WTU-V-021046 David Giblin 3376 12 May 2010 Dryopteridaceae WTU-381877, WTU-V-016124 Polystichum imbricans (D.C. Eaton) D.H. Wagner ssp. imbricans U.S.A., WASHINGTON, KLICKITAT COUNTY: Dryopteridaceae U. S. Bureau of Land Management property opposite Dillacort Canyon on west side of Polystichum munitum (Kaulf.) C. Presl Klickitat River. Elev. 393 ft. U.S.A., WASHINGTON, SKAGIT COUNTY: 45.741° N, 121.22549° W; WGS 84, North side of Skagit River and Route 20, 1 air mile northwest of uncertainty: 10 m., Source: GPS Rockport. Vernally moist, steep southeast-facing slope Elev. 492 ft. dominated by Lomatium grayi. Uncommon; in 48° 29.4' N, 121° 36.7' W; T35N R9E S26 east-facing rocky outcropping. Phenology: Deep shade of Acer circinatum, Rubus spectabilis, old growth forest Remnants. Origin: Native. of Tsuga heterophylla, Pseudotsuga menziesii, Thuja plicata. Cespitose, fronds to 166 centimeters; dominant in understory. David Giblin 3330 SHEET 1 OF 2. Phenology: Vegetative. Origin: Native. with Steve Hahn Peter F. Zika 18930 31 Aug 2003 8 Apr 2010 WTU-381217 WTU-379985, WTU-V-016121 Dryopteridaceae Dryopteridaceae Polystichum munitum (Kaulf.) C. Presl Polystichum munitum (Kaulf.) C. Presl U.S.A., WASHINGTON, KING COUNTY: Seattle, Discovery Park: Loop trail entry. U.S.A., WASHINGTON, WHATCOM COUNTY: 47.659817°, -122.417069°; WGS 84, uncertainty: 1000 m., Ross Lake National Recreation Area; east of trail; approximately 1.0 Source: Digital Map, Georef'd by Ibro Polovina kilometer northeast of Lodgepole Campground. FN #58. Phenology: Vegetative. Origin: Native. Elev. 1600 ft. 48.85658° N, 121.01725° W; WGS 84, uncertainty: 5 m., Source: Amy Galperin s.n. Apr 1977 GPS Pseudotsuga menziesii and Pinus ponderosa forest with Acer WTU-381726 circinatum in understory; occasional relictual balds. Scattered in moist area of forest understory. Phenology: Fertile. Origin: Native. Dryopteridaceae Ben Legler 11738 12 May 2010 Polystichum andersonii M. Hopkins WTU-381904 U.S.A., WASHINGTON, SNOHOMISH COUNTY: Dryopteridaceae Mt. Baker-Snoqualmie National Forest: slope at base of northeast face of Baring Mountain, just Polystichum munitum (Kaulf.) C. Presl south of Barclay Lake. Elev. 2700 ft. U.S.A., WASHINGTON, KITTITAS COUNTY: 47.781447° N, 121.425598° W; WGS 84, Along the west fork of Caseknife Creek about 1 mile from Taneum uncertainty: 100 m., Source: Digital Map, Creek. Georef'd by Ben Legler Elev. 3599 ft. North-facing talus and gravel deposits at base of 47° 5.322' N, 120° 58.76' W; T18N R16E S4; WGS 84 a narrow avalanche chute and stream gully; cool Dry, gravelly soil; 20° southeast-facing slope. Other plants: and shady; vegetated with Chamerion Pseudotsuga menziesii, Spiraea betulifolia, Calamagrostis angustifolium. Single plant with fronds erect to rubescens. Occasional. Phenology: Spores. Origin: Native. arching, found at base of large boulder at edge of forest. Phenology: Spores. Origin: Native. Don Knoke 1868 23 Jun 2009 Ben Legler 11866 3 Oct 2010 WTU-381092 WTU-407111, WTU-V-020313 Dryopteridaceae Dryopteridaceae Polystichum imbricans (D.C. Eaton) D.H. Wagner ssp. Polystichum braunii (Spenner) Fée imbricans RUSSIAN FEDERATION, SAKHALIN REGION: U.S.A., WASHINGTON, MASON COUNTY: Southeast Moneron Island; environs of creek Olympic National Forest: along Forest Service and drainage basin; near Poyasnoy Islands. Road 2353 ca. 1 road mile NW of its junction Elev. 371 ft. with Road 2355; north side of Skokomish River 46° 14.728' N, 141° 15.344' E valley. Bank next to river; dense Petasites grove with Elev. 464 m. sparse herb understory; on north-facing slope; 47.443384° N, 123.375469° W; WGS 84, scattered Angelica, 2 meters tall. Plants uncertainty: 17 m., Source: GPS, Georef'd by scattered under 1.5 meter Petasites groves. Ben Legler Phenology: Spores. Origin: Native. Dry, south-facing opening among young Pseudotsuga forest; thin soil over rock; Kari Stiles 2001-387 dominated by Arctostaphylos columbiana. Fronds in large clumps; pinnae ascending, 24 Aug 2001 oriented vertically; plants uncommon. Phenology: Spores. Origin: Native. WTU-391756 Ben Legler 12081 24 Jul 2011 WTU-409964, WTU-V-023974 Dryopteridaceae Dryopteridaceae Polystichum munitum (Kaulf.) C. Presl Polystichum kruckebergii W.H. Wagner U.S.A., WASHINGTON, STEVENS COUNTY: Western Colville National Forest and vicinity: U.S.A., WASHINGTON, CHELAN COUNTY: Little Roundtop Mountain: Middle Fork Mill East wall of cirque on south side of Cutthroat Creek, circa 8 air miles northeast of Colville Peak, ca. 2.2 air miles west of Washington along Irish Mountain Road from Aladdin Route. Pass. Elev. 2700-3842 ft. Elev. 1988 m. T36N R40E S9 SE; T36N R40E S10 W 1/2; 48.522304° N, 120.700168° W; WGS 84, Source: Inferred from duplicate specimen, uncertainty: 9 m., Source: Digital Map, Georef'd by Ed Alverson Georef'd by Ben Legler East face of mountain with scattered large Sloping subalpine meadows and vegetated ponderosa pine and Douglas fir regeneration scree, rocky, withPhyllodoce empetriformis, near top; ninebark and ocean spray; mixed Antennaria lanata, Anemone occidentalis, forest, hemlock/cedar at base. Phenology: Vaccinium. Several large tufts of fronds in Vegetative. Origin: Native. crevice of rock face above scree; uncommon. Phenology: Spores. Origin: Native. Jean Wood 3775 20 Aug 1995 Ben Legler 12254 WTU-387091, WTU-V-18431 21 Aug 2011 Dryopteridaceae WTU-407038, WTU-V-019833 Polystichum kruckebergii W.H. Wagner Dryopteridaceae U.S.A., WASHINGTON, OKANOGAN COUNTY: South-facing slope below summit of Hurley Polystichum lonchitis (L.) Roth Peak. Elev. 7600 ft. U.S.A., ALASKA, JUNEAU COUNTY: 48.966999°, -119.776598°; WGS 84, Admiralty Island, Alexander Archipelago, north uncertainty: 500 m., Source: Digital Map, end of island, ridge just north of the Greens Georef'd by Crystal Shin Creek mine. Cliff crevices and rocks. Little soil development. Elev. 3063 ft. Associated with Oxyria digyna. Phenology: 58.09732° N, 134.65434° W Spores. Origin: Native. Alpine ridge, rock outcrop with little vegetation, southwest-facing scree and boulder slope, 70 Steve Wagstaff 21 percent unvegetated, growing with Salix reticulata, Dryas sp., Silene acaulis, and 28 Aug 1981 Parnassia fimbriata. Phenology: Spores. Origin: Native. WTU-387088, WTU-V-016140 Mary Clay Stensvold 8209 9 Jul 2004 WTU-384746, WTU-V-021049 Dryopteridaceae Dryopteridaceae Polystichum imbricans (D.C. Eaton) D.H. Wagner Polystichum munitum (Kaulf.) C. Presl U.S.A., WASHINGTON, CHELAN COUNTY: U.S.A., WASHINGTON, KITSAP COUNTY: Chiwaukum Creek Trail, approximately 1.0 Circa 2 miles south of Lofall above Hood Canal. kilometers from trailhead. 47.786576°, -122.676186°; uncertainty: 1500 Elev. 2197 ft. m., Georef'd by Joseph Stormer 47.6912° N, 120.7639° W; WGS 84, In moist, shady area of dense woods. Shrub uncertainty: 500 m., Source: Digital map circa 9-10 decimeters tall. Phenology: Spores. Rocky outcroppings and dry, open slopes along Origin: Native. side of trail. Scattered in south-facing rocky outcropping. Phenology: Spores. Origin: Native. Carl A. Linde Jr. s.n. David Giblin 4182 27 Apr 1957 with Julia Monahan WTU-390376, WTU-V-18430 3 Jun 2012 WTU-389329, WTU-V-016117 Dryopteridaceae Polystichum munitum (Kaulf.) C. Presl Dryopteridaceae U.S.A., WASHINGTON, KING COUNTY: Polystichum andersonii M. Hopkins Woods, 25 meters east of Riverton Hospital. 47.487109°, -122.295051°; WGS 84, U.S.A., WASHINGTON, WHATCOM COUNTY: uncertainty: 200 m., Source: Digital Map, Ross Lake National Recreation Area: along Big Georef'd by Ibro Polovina Beaver Trail on north side of Big Beaver Creek Forest, loam soil, well shaded. Phenology: ca. 3 miles northeast of Ross Lake. Spores. Origin: Native. Elev. 536 m. 48.798174° N, 121.102014° W; WGS 84, Jerry Sheldon 19 uncertainty: 17 m., Source: GPS, Georef'd by Ben Legler 3 Mar 1973 Old growth Thuja plicata with brushy understory of Acer circinarum. Fronds arching from a basal WTU-391168, WTU-V-18320 clump; vegetative buds present on fronds; plants common under shrubs. Phenology: Spores. Origin: Native. Dryopteridaceae Ben Legler 12106 Polystichum munitum (Kaulf.) C. Presl 27 Jul 2011 U.S.A., WASHINGTON, KITSAP COUNTY: WTU-393092, WTU-V-016049 Hood Canal circa 2 miles south of Lofall. 47.786279°, -122.677466°; uncertainty: 3000 m., Georef'd by Marina DeMarco Forest. Phenology: Spores. Origin: Native. Dryopteridaceae Nancy Karns 58 Polystichum munitum (Kaulf.) C. Presl 11 Apr 1959 U.S.A., WASHINGTON, SKAGIT COUNTY: Vendovi Island. Near cove on north side of WTU-391175, WTU-V-18321 island. In general vicinity of house. Elev. 16-65 ft. 48° 36' 46.44" N, 122° 36' 50.4" W; WGS 84, uncertainty: 50 m., Source: Digital map Open, mowed area around house and outbuildings. Abundant in forests throughout island. Phenology: Spores. Origin: Native. Peter Dunwiddie 2011-56 1 Jun 2011 WTU-404341, WTU-V-012405 Dryopteridaceae Dryopteridaceae Polystichum lonchitis (L.) Roth Polystichum lonchitis (L.) Roth U.S.A., WASHINGTON, PEND OREILLE U.S.A., WASHINGTON, PEND OREILLE COUNTY: COUNTY: Colville National Forest. Along old road bed Colville National Forest. United States Forest converted to trail between trailhead and start of Service Road 2220. trail to Hall Mountain at end of Forest Service Elev. 1360 m. Road 500 off Sullivan Creek Road. 48.9197° N, 117.0884° W; WGS 84, Elev. 1670 m. uncertainty: 500 m., Source: GPS, Georef'd 48.80119° N, 117.231037° W; WGS 84, by Maria Yousoufian uncertainty: 18 m., Source: GPS, Georef'd by Roadside edge below rocky outcropping; Maria Yousoufian gravelly soil, open. Uncommon; scattered Moist mossy soil of road bank and bed under across hillside. Phenology: Spores. Origin: Alnus sinuata and conifer forest. Locally Native. common on shaded road bank under shrubs. Phenology: Spores. Origin: Native. David Giblin 4227 Don Knoke 2300 21 Jul 2012 21 Jul 2012 WTU-408697, WTU-V-022475 WTU-406838, WTU-V-019560 Dryopteridaceae Polystichum lonchitis (L.) Roth Dryopteridaceae U.S.A., WASHINGTON, WHATCOM COUNTY: Polystichum lonchitis (L.) Roth Ross Lake National Recreation Area: along Panther Creek Trail ca. 1.3 miles from trailhead. U.S.A., WASHINGTON, PEND OREILLE Elev. 1242 m. COUNTY: 48.693909° N, 120.982717° W; WGS 84, Colville National Forest. Along Round Top uncertainty: 33 m., Source: GPS, Georef'd by Mountain Trail #512 (Shedroof Divide trail). Ben Legler Elev. 1830 m. Rocky forest understory, with Ribes lacustre, 48.8061° N, 117.1298° W; WGS 84, Athyrium filix-femina, Oplopanax horridus. uncertainty: 1000 m., Source: Digital map, Fronds grass-green; petiole very short and the Georef'd by Maria Yousoufian lower pinnae much reduced; teeth few, short, Mixed coniferous open forest along trail. Some mostly incurved; single plant, growing with P. rocky outcrops and seeps. Phenology: Spores. munitum (other P. lonchitis seen elsewhere Origin: Native. along trail). Photographed. Phenology: Spores. Susan Kephart 777 Ben Legler 12735 7 Jul 2013 21 Jul 2012 WTU-403922, WTU-V-012133 WTU-407928, WTU-V-021156 Dryopteridaceae Dryopteridaceae Polystichum lonchitis (L.) Roth Polystichum lonchitis (L.) Roth U.S.A., MONTANA, GRANITE COUNTY: Beaverhead-Deerlodge National Forest: along U.S.A., WASHINGTON, PEND OREILLE North Fork Flint Creek 2.4 air km due south of COUNTY: Red Lion Mountain. Colville National Forest. Bear Pasture trailhead Elev. 2219 m. and along road near trailhead. 46.265633° N, 113.198902° W; WGS 84, Elev. 5620 ft. uncertainty: 6 m., Source: GPS, Georef'd by 48.91724° N, 117.13914° W Ben Legler Phenology: Spores. Origin: Native. Open Picea engelmanii forest with lush herbaceous ground layer; adjacent to stream. Doug Williams 2012-18 Uncommon in shade by small flowing seep. Phenology: Spores. 22 Jul 2012 Ben Legler 12857 WTU-407539, WTU-V-020596 18 Aug 2013 WTU-403552, WTU-V-011958 Dryopteridaceae Dryopteridaceae Polystichum lonchitis (L.) Roth Polystichum munitum (Kaulf.) C. Presl U.S.A., MONTANA, MINERAL COUNTY: U.S.A., WASHINGTON, SNOHOMISH Lolo National Forest: along Forest Service Road COUNTY: 342 on south side of Fourth of July Gulch, ca 7 East Warm Beach Senior Community, air km ENE of Little Joe Mountain. Stanwood. Elev. 1510 m. 48.181907°, -122.348357°; WGS 84, 47.175222° N, 115.159927° W; WGS 84, uncertainty: 224 m., Source: Digital Map, uncertainty: 13 m., Source: GPS, Georef'd by Georef'd by Maria Yousoufian Ben Legler Mesic, mixed woods. Phenology: Spores. Mossy road cut on north-facing slope, with Origin: Native. Polystichum lonchitis, Bromus. Fronds in rosettes; plants common. Phenology: Spores. Clayton Haberman s.n. Ben Legler 12875 4 Aug 2010 19 Aug 2013 WTU-402773, WTU-V-010541 WTU-403522, WTU-V-011928 Dryopteridaceae Polystichum imbricans (D.C. Eaton) D.H. Wagner ssp. imbricans Dryopteridaceae U.S.A., WASHINGTON, SKAMANIA COUNTY: Polystichum munitum (Kaulf.) C. Presl Southwest side of Mount Adams. Salt Creek Trail #75. Elev. 1166 m. U.S.A., WASHINGTON, PIERCE COUNTY: 46° 8.958' N, 121° 34.489' W; Source: Field notes North Shore Henderson Bay, Gig Harbor. Only a few fronds with spores. Phenology: Vegetative. Origin: 47.372° N, 122.663° W; Source: Specimen Native. Label Woods. Leaf material removed for RADseq David Biek 05-143 25 Jul 2005 analysis by Megan Ruffley at WTU, February with Susan McDougall 2016. Phenology: Spores. Origin: Native. Herbarium: WTU Bill Harms 7151 Dryopteridaceae 24 Nov 2011 Polystichum munitum (Kaulf.) C. Presl × Polystichum WTU-402600, WTU-V-010508 scopulinum (D.C. Eaton) Maxon U.S.A., WASHINGTON, KITTITAS COUNTY: Dryopteridaceae North side of bridge over mouth of Beverly Creek, 5.4 air kilometers east of Malcolm Polystichum munitum (Kaulf.) C. Presl Mountain. Elev. 955 m. U.S.A., WASHINGTON, YAKIMA COUNTY: 47.375° N, 120.877° W; Source: Specimen Southeast side of Mount Adams. Forest Road 071. Label Elev. 1199 m. Partial shade, dry rocky northeast slope above 46° 5.588' N, 121° 27.745' W; Source: Field notes road, over serpentine, with Aspidotis, Achillea, Calypso spot. Phenology: Spores. Origin: Native. Polystichum scopulinum (which is only half as tall). Cespitose, rare. Phenology: Spores. Origin: David Biek 04-83 3 Jul 2004 Native. with Susan McDougall Peter F. Zika 27147 Herbarium: WTU 23 Jul 2015 Dryopteridaceae WTU-404212, WTU-V-012341 Polystichum munitum (Kaulf.) C. Presl Dryopteridaceae U.S.A., WASHINGTON, JEFFERSON Polystichum scopulinum (D.C. Eaton) Maxon COUNTY: Kalaloch. U.S.A., WASHINGTON, KITTITAS COUNTY: 47.60472°, -124.37111°; WGS 84, North side of bridge over mouth of Beverly uncertainty: 5000 m., Source: GeoLocate, Creek, 5.4 air kilometers east of Malcolm Georef'd by WTU Staff Mountain. [Label says Clallam County but this location is Elev. 955 m. actually in Jefferson County.] Phenology: 47.375° N, 120.877° W; Source: Specimen Vegetative. Origin: Native. Label Partial shade, dry rocky northeast slope above Clayton Haberman s.n. road, over serpentine, with Aspidotis, Achillea, Polystichum munitum x scopulinum. Cespitose, 28 Sep 1991 common. Phenology: Spores. Origin: Native. WTU-402770, WTU-V-010537 Peter F. Zika 27146 23 Jul 2015 WTU-404213, WTU-V-012340 Dryopteridaceae Dryopteridaceae Polystichum munitum (Kaulf.) C. Presl Polystichum lonchitis (L.) Roth U.S.A., WASHINGTON, KITTITAS COUNTY: U.S.A., WASHINGTON, PEND OREILLE South side of Snoqualmie Pass, 1 air kilometer COUNTY: north-northeast of Lodge Lake, Cascade Range. Road 2220, 2.5 air kilometers south-southeast Elev. 1050 m. of Salmo Mountain. 47.419° N, 121.423° W; Source: Specimen Elev. 1560 m. Label 48.946° N, 117.097° W; Source: Specimen Full sun, east aspect, ski slope, with Label Lycopodium sitchense, Tsuga mertensiana. On dry slope, west aspect, full sun, with Cespitose, evergreen. Phenology: Spores. Athyrium filix-femina, Dryopteris filix-mas, Origin: Native. Montia, Anaphalis, Alnus, Picea. Cespitose; common. Phenology: Spores. Origin: Native. Peter F. Zika 27130 Peter F. Zika 27245 21 Jul 2015 18 Aug 2015 WTU-404248, WTU-V-012317 WTU-404398, WTU-V-012489 Dryopteridaceae Dryopteridaceae Polystichum munitum (Kaulf.) C. Presl Polystichum munitum (Kaulf.) C. Presl U.S.A., WASHINGTON, PEND OREILLE COUNTY: U.S.A., WASHINGTON, KITTITAS COUNTY: Road 57, North Fork Granite Creek drainage, Approximately 2 kilometers east of junction of 2.4 air kilometers northwest of Little Grass Leisure Land Lane and Iron Horse Trail; just Mountain (in Idaho). southeast of I-90 Exit 80 interchange. Elev. 1060 m. Elev. 606 m. 48.771° N, 117.053° W; Source: Specimen 47.1779° N, 121.0054° W; WGS 84, Label uncertainty: 20 m., Source: Digital map In partial shade, steep vegetated talus, with Impounded ponds and riparian zone ephemeral Thuja, Polystichum lonchitis, Dryopteris pond along edge of Pinus ponderosa and carthusiana, D. expansa, D. filix-mas, Athyrium Populus trichocarpa forest; water level filix-femina, Gymnocarpium disjunctum. fluctuations associated with nearby Yakima Cespitose, evergreen, uncommon. Phenology: River. Uncommon on north-facing gravel berm Spores. Origin: Native. of Iron Horse Trail. Phenology: Spores. Origin: Native. Peter F. Zika 27228 David Giblin 4240 18 Aug 2015 with Don Knoke WTU-404410, WTU-V-012501 24 Sep 2012 Dryopteridaceae WTU-415272, WTU-V-032559 Polystichum lonchitis (L.) Roth Dryopteridaceae U.S.A., WASHINGTON, PEND OREILLE COUNTY: Polystichum scopulinum (D.C. Eaton) Maxon Road 57, North Fork Granite Creek drainage, 2.4 air kilometers northwest of Little Grass U.S.A., OREGON, GRANT COUNTY: Mountain (in Idaho). Malheur/Umatilla National Forest: slope on west side of cirque in Elev. 1060 m. headwaters of East Fork Clear Creek ca. 0.5 air miles NW of 48.771° N, 117.053° W; Source: Specimen Vinegar Hill. Label Elev. 2157 m. In partial shade, steep vegetated talus, with 44.720651° N, 118.563084° W; WGS 84, uncertainty: 25 m., Thuja, Polystichum munitum, Dryopteris Source: GPS, Georef'd by Ben Legler carthusiana, D. expansa, D. filix-mas, Athyrium East facing, dry slope with ultramaphic outcrops; sparsely vegetated filix-femina, Gymnocarpium disjunctum. with Eriophyllum, Artemisia, Agastache, Melica, Aspidotis densa. Cespitose, evergreen, common. Phenology: Tufted in rock crevices. Phenology: Spores. Origin: Native. Spores. Origin: Native. Ben Legler 12204 16 Aug 2011 Peter F. Zika 27229 with Nick Otting, Richard Brainerd 18 Aug 2015 Herbarium: WTU WTU-404402, WTU-V-012493 Dryopteridaceae Dryopteridaceae Polystichum andersonii M. Hopkins Polystichum andersonii M. Hopkins U.S.A., WASHINGTON, SKAGIT COUNTY: U.S.A., WASHINGTON, CLARK COUNTY: Mt. Baker-Snoqualmie National Forest: along Trail #769 ca. 1/8 mile Yacolt Burn State Forest: just below Tarbell Trail past end of Forest Service Road 1590; South Fork Cascade River. on east-trending ridge of Larch Mountain ca 1.3 Elev. 511 m. air km ENE of summit. 48.449431° N, 121.1375° W; WGS 84, uncertainty: 14 m., Elev. 894 m. Source: GPS, Georef'd by Ben Legler 45.719813° N, 122.279475° W; WGS 84, Open old growth forest of Thuja plicata and Tsuga heterophylla, with uncertainty: 5 m., Source: GPS, Georef'd by understory of Athyrium filix-femina, Polystichum. Fronds in basal Ben Legler rosettes; vegetative bud present near frond tip; ca. 10 plants here. Top edge of a large, north-facing talus slope, Phenology: Spores. shaded by Acer circinatum and Holodiscus discolor; volcanic rock covered in moss. Fronds Ben Legler 12697 23 Jun 2013 yellow-green, many with small vegetative bud near tip, arching from a cespitose base; plants Herbarium: WTU frequent in shaded mossy talus. Phenology: Spores. Dryopteridaceae Ben Legler 13863 12 Sep 2015 Polystichum kruckebergii W.H. Wagner WTU-409256, WTU-V-023136 U.S.A., WASHINGTON, SKAGIT COUNTY: Mt. Baker-Snoqualmie National Forest: on west side of saddle along Dryopteridaceae Hidden Lake Ridge; ca. 2.2 air km northeast of Hidden lake Lookout. Elev. 1861 m. Polystichum imbricans (D.C. Eaton) D.H. Wagner ssp. 48.512696° N, 121.191756° W; WGS 84, uncertainty: 10 m., imbricans Source: GPS, Georef'd by Ben Legler Steep, rocky, southwest-facing slope with small sloping benches of U.S.A., WASHINGTON, CLARK COUNTY: gravelly soil, and low cover of perennial herbs and shrubs including Yacolt Burn State Forest: just below Tarbell Trail Solidago, Fragaria virginiana, Sibbaldia, Dasiphora fruticosa, on east-trending ridge of Larch Mountain ca 1.3 Artemisia, Vaccinium cespitosum, and Juniperus communis. air km ENE of summit. Scattered in mossy soil in crevices of steep, broken rock face. Elev. 894 m. Phenology: Spores. 45.719813° N, 122.279475° W; WGS 84, uncertainty: 5 m., Source: GPS, Georef'd by Ben Legler 12915 25 Aug 2013 Ben Legler Top edge of a large, north-facing talus slope, Herbarium: WTU shaded by Acer circinatum and Holodiscus discolor; volcanic rock covered in moss. Fronds tufted, erect, with pinnae folded inwards or Dryopteridaceae spreading, mostly imbricate; plants common in mossy talus. Phenology: Spores. Polystichum andersonii M. Hopkins Ben Legler 13865 12 Sep 2015 U.S.A., WASHINGTON, SNOHOMISH COUNTY: Mt Baker-Snoqualmie National Forest: Northeast side of Goat Lake WTU-409268, WTU-V-023124 ca. 0.4 km from outlet; east of Barlow Pass. Elev. 942 m. 48.02031° N, 121.350525° W; WGS 84, uncertainty: 7 m., Dryopteridaceae Source: GPS, Georef'd by Ben Legler Brushy slope above lake shore; south-facing, with Acer circinatum, Polystichum imbricans (D.C. Eaton) D.H. Wagner ssp. Cornus sericea, Aruncus dioicus, Rubus parviflorus. Fronds arching, imbricans tufted from a cespitose base; vegetative bud present near frond tip; plants scattered under shrubs, growing with Polystichum munitum U.S.A., WASHINGTON, LEWIS COUNTY: (no hybrids seen). Phenology: Spores. Gifford Pinchot National Forest: along Forest Service Road 21 at bend in road 2.7 air km Ben Legler 13830 7 Sep 2015 southeast of junction of Road 21 with Highway 12, south of Packwood. Herbarium: WTU Elev. 578 m. 46.564037° N, 121.664242° W; WGS 84, uncertainty: 22 m., Source: Digital Map, Georef'd by Ben Legler Dry, rocky openings and road cuts on southwest-facing slope among Pseudotsuga menziesii forest, with Arctostaphylos columbiana, Collinsia, Selaginella wallacei, and moss. Fronds erect, pinnae imbricate; forming large tufts in thin dry soil over rock. Phenology: Spores. Ben Legler 13508 9 May 2015 WTU-409973, WTU-V-023966 Dryopteridaceae Dryopteridaceae Polystichum imbricans (D.C. Eaton) D.H. Wagner ssp. Polystichum munitum (Kaulf.) C. Presl imbricans U.S.A., WASHINGTON, ISLAND COUNTY: U.S.A., WASHINGTON, LEWIS COUNTY: West Troxell Road, 0.5 road miles east of Mt. Baker-Snoqualmie National Forest: bald on junction with Monkey Hill Road, 3.2 air Point 2103 just northwest of where Cortright kilometers southwest of Hoypus Hill, Whidbey Creek joins the Clear Fork of the Cowlitz River; Island. 12 air km northeast of Packwood. Elev. 80 m. Elev. 624 m. 48.3764° N, 122.6254° W; Source: Specimen 46.690016° N, 121.566714° W; WGS 84, Label uncertainty: 60 m., Source: Digital Map, Partly shaded edge of conifer forest, in dense Georef'd by Ben Legler understory, with Gaultheria shallon, Vaccinium Large, south-facing bald on edge of hill above parvifolium, Rubus spectabilis, Holodiscus stream; with large areas of dry, mossy soil over discolor, Tsuga heterophylla, Pseudotsuga rock, large patches of Arctostaphylos menziesii. Cespitose, evergreen, common. columbiana, scattered Penstemon fruticosus, SHEET 2 OF 2. Phenology: Spores. Origin: and Festuca idahoensis. Fronds erect, pinnae Native. imbricate; plants in tufts in moss over rock, variable in size, some with fronds only 10 cm Peter F. Zika 27583 17 Oct 2015 long; scattered across bald. Phenology: Spores. WTU-405833, WTU-V-18078 Ben Legler 13677 28 Jun 2015 Dryopteridaceae WTU-409212, WTU-V-023188 Polystichum munitum (Kaulf.) C. Presl Dryopteridaceae U.S.A., WASHINGTON, CLALLAM COUNTY: 1 air kilometer east of Lake Sutherland, south of Polystichum kruckebergii W.H. Wagner and above Route 101, Elwha River drainage, Olympic Peninsula. U.S.A., WASHINGTON, PIERCE COUNTY: Elev. 155 m. Mount Rainier. 48.074° N, 123.674° W; Source: Specimen 46.852827°, -121.760441°; WGS 84, Label uncertainty: 17500 m., Source: GeoLocate, In full or partial shade at forest edge, with Acer Georef'd by Joseph Stormer macrophyllum, A. circinatum, Rubus spectabilis. Phenology: Spores. Origin: Native. Cespitose, common. SHEET 2 OF 2. Phenology: Spores. Origin: Native. E. C. Smith 876 Peter F. Zika 27266 Aug 1890 27 Aug 2015 WTU-2038, WTU-V-016144 WTU-405784, WTU-V-18048

Dryopteridaceae Dryopteridaceae Polystichum imbricans (D.C. Eaton) D.H. Wagner ssp. Polystichum acrostichoides (Michx.) Schott imbricans U.S.A., WISCONSIN, WAUKESHA COUNTY: U.S.A., WASHINGTON, CHELAN COUNTY: Village of Menomonee Falls. Rock Mountain Trail 1.5 to 2 miles from 43.178897°, -88.117313°; WGS 84, Highway 2. uncertainty: 8365 m., Source: GeoLocate, Elev. 4400 ft. Georef'd by Maria Yousoufian T26N R15E S3 NE 1/4; WGS84, uncertainty: Mixed hardwood stand. Phenology: Vegetative. 500 m., Source: Georeferenced, Georef'd by Origin: Native. WTU Staff With Ceanothus velutinus, Acer glabrum and Clayton Haberman s.n. Sorbus scopulina. A few small Pseudotsuga menziesii nearby. Phenology: Spores. Origin: 26 Jun 2003 Native. WTU-407229, WTU-V-020950 E. Burnett s.n. 20 Jul 1991 WTU-323845, WTU-V-016183 Dryopteridaceae Dryopteridaceae Polystichum lonchitis (L.) Roth Polystichum andersonii M. Hopkins U.S.A., WASHINGTON, WHATCOM COUNTY: U.S.A., WASHINGTON, SNOHOMISH COUNTY: Mt. Baker-Snoqualmie National Forest: talus slope and bordering Mt Baker-Snoqualmie National Forest: along Sauk River Trail ca rock faces on west slope of South Fork Divide ca 0.3 km south of 03.8 air km southeast of trailhead. Bell Creek. Elev. 830 m. Elev. 1170 m. 48.044293° N, 121.241528° W; WGS 84, uncertainty: 8 m., 48.700986° N, 121.862323° W; WGS 84, uncertainty: 100 m., Source: GPS, Georef'd by Ben Legler Source: Digital Map, Georef'd by Ben Legler Buried, vegetated talus slope dominated by Acer macrophyllum, with Exposed band of marble associated with rocks of the Chilliwack Acer circinatum, Athyrium filix-femina, Rubus parviflorus. Fronds Group, bordered below by blocky marble talus; full sun, dry; with ascending, tufted from a cespitose crown; single vegetative bud Rubus parviflorus, Pteridium aquilinum, and Artemisia sp. Fronds present below frond tip; plants uncommon on edge of brush. arching, tufted; pinnae thick glossy green; plants scattered on talus Phenology: Spores. slope in pockets of soil. Phenology: Spores. Ben Legler 14352 28 Aug 2016 Ben Legler 14226 29 Jul 2016 with Ed Alverson, Peter Zika Herbarium: WTU Herbarium: WTU Dryopteridaceae Dryopteridaceae Polystichum munitum (Kaulf.) C. Presl ? Polystichum lemmonii Underw. U.S.A., WASHINGTON, CLALLAM COUNTY: Near Ozette Lake. U.S.A., WASHINGTON, WHATCOM COUNTY: 48.14585° N, 124.5927° W; Source: GPS Mt. Baker-Snoqualmie National Forest: On west Forested wetland with large amounts of downed side of ridge along Sisters Divide 0.1 air km wood as growing substrate between areas of southeast of outlet of Lake Wiseman. inundated and/or saturated soil. Growing with Elev. 1315 m. Lysichiton americanus, Rubus spectabilis under 48.708547° N, 121.937441° W; WGS 84, overstory of Thuja plicata, Picea sitchensis, uncertainty: 6 m., Source: GPS, Georef'd by Alnus rubra. Abundance: Uncommon. Growth Ben Legler Habit: Erect fern (young?) Site ID: Exposures of dunite outcrops on west-facing NWCA16-4861. Visit #1. Collection #U1 slope, among subalpine woodland of Tsuga (Unknown Species). Phenology: Vegetative. mertensiana, Chamaecyparis nootkatensis, and Origin: Native. Pinus contorta, with Cassiope mertensiana, Adiantum aleuticum, Aspidotis densa, and Patricia Johnson NWCA16-4861-V1-U1 Polystichum lemmonii. Fronds erect to spreading, from a cespitose caudex, varying 28 Jun 2016 from 3 cm long in exposed plants to ca 30 cm long in shaded plants; plants common around WTU-410806, WTU-V-024858 bases of boulders. Phenology: Spores. Ben Legler 14251 6 Aug 2016 Dryopteridaceae WTU-418965, WTU-V-041870 Polystichum munitum (Kaulf.) C. Presl U.S.A., WASHINGTON, SKAGIT COUNTY: Dryopteridaceae Near Anacortes on refinery property. 48.46472° N, 122.54925° W; Source: GPS Polystichum lonchitis (L.) Roth Forested wetland. With Populus balsamifera, P. tremuloides, Cornus sericea, Alnus rubra, U.S.A., WASHINGTON, WHATCOM COUNTY: Symphoricarpos albus, Tellima grandiflora. Mt. Baker-Snoqualmie National Forest: southeast slope of Liumchen Abundance: Sparse. Growth Habit: Perennial Mountain ca 0.6 km south of Canada and just north of Canyon fern growing on ground surface. Site ID: Ridge. NWCA16-4858. Visit #1. Collection #Q2 (QA Elev. 1569 m. Voucher). Phenology: Vegetative. Origin: 48.993068° N, 121.856542° W; WGS 84, uncertainty: 4 m., Native. Source: GPS, Georef'd by Ben Legler Steep, exposed limestone outcrops on a dry southeast facing Patricia Johnson NWCA16-4858-V1-Q2 subalpine slope, bordered below by blocky limestone talus; with Phlox diffusa, Phacelia hastata var. leptosepala, Fragaria virginiana, 3 May 2016 Saxifraga austromontana, Dasiphora fruticosa, Juniperus communis, Polystichum lonchitis. Fronds tufted, dark glossy green; plants WTU-411361, WTU-V-025769 scattered in soil over talus, and in limestone crevices. Phenology: Spores. Ben Legler 14306 13 Aug 2016 with Sara Legler Herbarium: WTU Dryopteridaceae Dryopteridaceae Polystichum braunii (Spenner) Fée Polystichum braunii (Spenner) Fée U.S.A., ALASKA, KETCHIKAN GATEWAY U.S.A., ALASKA, PETERSBURG COUNTY: COUNTY: Kupreanof Island, west of Duncan Canal. Revilla Island, Swan Lake northeast of Elev. 20 ft. Ketchikan. UTM Zone 8, 602432E, 6289376N Elev. 335 ft. Plants growing on moist mossy cliff wall UTM Zone 9, 358569E, 6168289N; T72S adjacent large shallow creek. Associated R92E S12 NW 1/4 of SW 1/4 species: Adiantum pedatum, Cystopteris fragilis, Moist, undulating, moss, duff, and soil on Thelypteris phegopteris, Athyrium filix-femina, hummocks, under tall shrub layer, in riparian Actaea rubra, Saxifraga mertensiana, terrace. Aspect: 335 degrees. Slope: 1%. Adenocaulon bicolor, Heuchera glabra, Tiarella Associated species: Picea engelmannii, Alnus trifoliata, Streptopus amplexicaulis. rubra, Ribes bracteosum, Rubus spectabilis, Oplopanax horridum, Vaccinium parviflorum, Kathryn A. Beck 201166 Gymnocarpium dryopteris, Dryopteris expansa, Athyrium filix-femina, Tiarella trifoliata, 22 Jul 2011 Streptopus amplexicaulis, Trisetum cernuum. Phenology: Vegetative. WTU-414226, WTU-V-031285 Kathryn A. Beck 201201 7 Aug 2012 WTU-412632, WTU-V-027184 Dryopteridaceae Dryopteridaceae Polystichum munitum (Kaulf.) C. Presl Polystichum munitum (Kaulf.) C. Presl U.S.A., WASHINGTON, ISLAND COUNTY: U.S.A., WASHINGTON, CLALLAM COUNTY: West Troxell Road, 0.5 road miles east of 1 air km E of Lake Sutherland, S of and above junction with Monkey Hill Road, 3.2 air Rout 101, Elwha River drainage, Olympic kilometers southwest of Hoypus Hill, Whidbey Peninsula. Island. Elev. 155 m. Elev. 80 m. 48.074° N, 123.674° W; Source: Specimen 48.3764° N, 122.6254° W; Source: Specimen Label Label Cespitose, common in full or partial shade at Partly shaded edge of conifer forest, in dense forest edge, with Acer macrophyllum, A. understory, with Gaultheria shallon, Vaccinium circinatum, Rubus spectabilis. Phenology: parvifolia, Rubus spectabilis, Holodiscus Spores. Origin: Native. discolor, Tsuga heterophylla, Pseudotsuga menziesii. Cespitose, evergreen, common. Peter F. Zika 27266 Phenology: Vegetative. 27 Aug 2015 Peter F. Zika 27583 WTU-405788, WTU-V-18113 17 Oct 2015 WTU-405826, WTU-V-18127 Dryopteridaceae Polystichum imbricans (D.C. Eaton) D.H. Wagner ssp. Dryopteridaceae imbricans Polystichum andersonii M. Hopkins × Polystichum U.S.A., OREGON, JOSEPHINE COUNTY: munitum (Kaulf.) C. Presl SE side of Horseshoe bend, above E bank of Rogue River. U.S.A., OREGON, LANE COUNTY: Elev. 150 m. Cultivated at 501 Irving Road, Eugene. A 42.6799° N, 123.7344° W; Source: Specimen division from a plant originally collected from Label Clackamas County, Oregon, near Williams Cespitose, common, dry mossy ledges in full Lake, headwaters of South Fork Clackamas sun or partial shade, over sandstone, with River, circa 13 miles (21 kilometers) Quercus chrysolepis, Selaginella wallacei. south-southeast of Estacada along BLM Road 5 Phenology: Spores. Origin: Native. 4E-26. Elev. 3600 ft. P.F. Zika 27300 T5S R4E S26 with R. E. Brainerd A division from a cespitose plant originally collected on 16 July 1991 by David H. Wagner. 7 Sep 2015 Phenology: Vegetative. Cultivated. WTU-405863, WTU-V-18119 Edward R. Alverson 2015-14 15 Jun 2015 WTU-405823, WTU-V-18130 Dryopteridaceae Dryopteridaceae Polystichum munitum (Kaulf.) C. Presl Polystichum munitum (Kaulf.) C. Presl U.S.A., WASHINGTON, SKAGIT COUNTY: U.S.A., IDAHO, SHOSHONE COUNTY: Thunder Lane, 0.1 road miles east of junction Coeur d'Alene National Forest. Shadow Falls, with Cottage Lane, 1.2 air kilometers southwest along and above trail at Yellow Dog Creek. of Ginnett Hill, Fidalgo Island. [Elevation approximate, from digital map.] Elev. 105 m. Elev. 3200 ft. 48.4317° N, 122.6553° W; Source: Specimen 47.7596° N, 116.1049° W; Source: GPS, Label Digital map In partly shaded thickets, with Rubus parviflorus, West-facing slope along and above trail. Sambucus racemosa, Alnus rubra, Thuja Western hemlock dominant, ericaceous plicata. Cespitose, evergreen, common. understory. (Vaccinium, etc.) Phenology: Phenology: Vegetative. Origin: Native. Spores. Origin: Native. Peter F. Zika 27576 Ashley Powell 2016-40 17 Oct 2015 9 Jul 2016 WTU-405877, WTU-V-18157 WTU-420985, WTU-V-50113

Dryopteridaceae Dryopteridaceae Polystichum munitum (Kaulf.) C. Presl Polystichum andersonii M. Hopkins U.S.A., WASHINGTON, SKAGIT COUNTY: U.S.A., ALASKA COUNTY: Thunder Lane, 0.1 road miles east of junction Southeast Alaska. Tongass National Forest, with Cottage Lane, 1.2 air kilometers southwest Yakobi Island (north end). About 1/2 mile south of Ginnett Hill, Fidalgo Island. of Soapstone Point. Between Soapstone Harbor Elev. 105 m. and sandy beach on P. Ocean. 48.4317° N, 122.6553° W; Source: Specimen T43S R54S S36 Label South face of rock cliff of north side of sandy In partly shaded thickets, with Rubus parviflorus, ocean beach. U.S. Forest Service Sitka, Alaska, Sambucus racemosa, Alnus rubra, Thuja Herbarium, 00141. Origin: Native. plicata. Cespitose, evergreen, common. Phenology: Vegetative. Origin: Native. Mary Clay Muller 3073 Peter F. Zika 27576 24 Jun 1979 17 Oct 2015 WTU-408560, WTU-V-022698 WTU-405870, WTU-V-18164 Dryopteridaceae Dryopteridaceae Polystichum braunii (Spenner) Fée Polystichum lonchitis (L.) Roth U.S.A., ALASKA COUNTY: Juneau area about 10 miles north of downtown U.S.A., IDAHO, SHOSHONE COUNTY: at Mendenhall Glacier. Collected along East Coeur d'Alene National Forest. Shoulder of Glacier Trail. About 1/4 mile south of the Nugget Forest Service Road 208. Creek waterfall. Elev. 739 m. T40S R66E S9 NE 1/4 NW 1/4 47.74572° N, 116.020736° W; WGS 84, Growing along the trail in somewhat open areas. uncertainty: 25 m., Source: GPS Forest Service Alaska Region, TNFS Vertical rock face, northeast facing, with Herbarium, v. 5631. Origin: Native. Saxifraga mertensiana, Prenanthus sagittata, Campanula rotundifolia, moss. Fronds tufted; Mary Stensvold 6808 uncommon in shaded moist areas at base of rock face. Phenology: Spores. Origin: Native. 22 Jul 1996 Gabriel Wisswaesser 2016-68 WTU-408557, WTU-V-022710 9 Jul 2016 WTU-419199, WTU-V-041060 Dryopteridaceae Dryopteridaceae Polystichum andersonii M. Hopkins Polystichum lonchitis (L.) Roth U.S.A., ALASKA COUNTY: U.S.A., ALASKA COUNTY: Tongass National Forest. Near Mendenhall Chichagof Island, west side of Lisianski Inlet, Glacier, about 10 miles northwest of Juneau. Cann Creek drainage. About 1 1/4 mile Collected along East Glacier Trail, open area. southwest of the mouth of Cann Creek. Apex Elev. 100 ft. mine portal. Grassy cliff just west of small waterfall, along T45S R56E S23 NE 1/4 SW 1/4 trail. U.S. Forest Service Sitka, Alaska, Growing on north side of waste rock pile in front Herbarium, 01474. Origin: Native. of portal. Forest Service Alaska Region, TNFS Herbarium, v.3802. Origin: Native. Mary Clay Muller 4189 Mary Clay Stensvold 6311 30 Oct 1980 23 Aug 1993 WTU-408326, WTU-V-022695 WTU-408374, WTU-V-022705

Dryopteridaceae Dryopteridaceae Polystichum andersonii M. Hopkins Polystichum lonchitis (L.) Roth U.S.A., ALASKA COUNTY: U.S.A., ALASKA COUNTY: Tongass National Forest. Near Mendenhall Alexander Archipelago. Heceta Island. Bald Glacier, about 10 miles northwest of Juneau. mountain. Coordinates for Bald mountain, Collected along East Glacier Trail. specific location unknown. Elev. 100 ft. Elev. 2000 ft. Open area. This area was recently deglaciated. 55.7237° N, 133.5622° W; WGS 84, Source: Common. U.S. Forest Service Sitka, Alaska, Specimen Label Herbarium, 01475. Origin: Native. Subalpine/alpine talus slope. Forest Service Alaska Region, TNFS Herbarium, v. 6761. Mary Clay Muller 4186 Origin: Native. 30 Oct 1980 Phyllis Woolwine 565 WTU-408322, WTU-V-022696 13 Jul 1999 WTU-408373, WTU-V-022706 Dryopteridaceae Polystichum andersonii M. Hopkins Dryopteridaceae U.S.A., ALASKA COUNTY: Polystichum lonchitis (L.) Roth Southeastern Alaska. Baranof Island, southeast part. North end of Patterson Bay. Between head U.S.A., ALASKA COUNTY: of Bay and small lake just south of Parry Lake. Admiralty Island, north end of the Island. About Near head of bay. 15 kilometers east southeast of Hawk Inlet on T60S R69E S27 the north-facing ridgeface south of Greens Edge of dry rocky stream bed, under forest Creek. canopy. U.S. Forest Service Sitka, Alaska, Elev. 930 m. Herbarium, 02723. Origin: Native. 58.0674° N, 134.585° W; Source: Specimen Label Mary Clay Muller 3609 Among boulders and rocks on a scree slope. Occasional. Pinna margins deeply lacerate. 3 20 Jun 1980 morphotypes found in an area of 15 meters square. Forest Service Alaska Region, TNFS WTU-408321, WTU-V-022697 Herbarium, v. 7241. Origin: Native. Mary Clay Stensvold 8189 8 Jul 2004 WTU-408372, WTU-V-022707 Dryopteridaceae Dryopteridaceae Polystichum kruckebergii W.H. Wagner Polystichum braunii (Spenner) Fée U.S.A., ALASKA COUNTY: U.S.A., ALASKA COUNTY: [locality and coordinates withheld; status in Juneau. Alaska: S1 (Critically Imperiled; AKNHP)] 58.301944°, -134.419722°; uncertainty: 10000 Elev. 286 m. m. Rare on 10? slope on northeast aspect, Woods. Forest Service Alaska Region, TNFS ultramafic rock outcrop, bluff with overhang. Herbarium, v. 6408. Origin: Native. Forest Service Alaska Region, TNFS Herbarium, v. 7186. Origin: Native. J. P. Anderson 6159 K. Calhoun 18 18 Aug 1940 with B. Krieckhaus WTU-408367, WTU-V-022713 18 Jun 2004 WTU-408371, WTU-V-022708 Dryopteridaceae Polystichum braunii (Spenner) Fée Dryopteridaceae U.S.A., ALASKA COUNTY: Polystichum braunii (Spenner) Fée Chichagof Island, west side of Lisianski Inlet, Cann Creek drainage. About 1/2 mile southwest U.S.A., ALASKA COUNTY: of the mouth of Cann Creek. Southeastern Alaska. About 30 miles north of 57.957° N, 136.276° W; T45S R56E S24 Juneau at Berners Bay. West side of Echo Cove SW1/4 NW 1/4; Source: Specimen Label along the beach. Hemlock-spruce forest along trail above lower Forest ecotone. Growing about 30 feet back bog. Forest Service Alaska Region, TNFS from the high tide line at the forest/beach Herbarium, v. 4768. Origin: Native. ecotone among deciduous shrubs. U.S. Forest Service Sitka, Alaska, Herbarium, 01423. Origin: Mary Clay Stensvold 6307 Native. 23 Aug 1993 Mary Clay Muller 4149 WTU-408366, WTU-V-022714 14 Sep 1980 WTU-408370, WTU-V-022709 Dryopteridaceae Dryopteridaceae Polystichum braunii (Spenner) Fée Polystichum braunii (Spenner) Fée U.S.A., ALASKA COUNTY: Tongass National Forest. Mainland. About 28 U.S.A., ALASKA COUNTY: miles north of Juneau. About 4 miles up the Tongass National Forest, Chatham Area. Indian Eagle Glacier trail. River (Tenakee). Growing in the forest near the trail. U.S. Forest U.S. Forest Service Sitka, Alaska, Herbarium, Service Sitka, Alaska, Herbarium, 01447. Origin: 00145. Origin: Native. Native. James Downs s.n. Mary Clay Muller 4081 30 Sep 1976 21 Aug 1980 WTU-408369, WTU-V-022711 WTU-408363, WTU-V-022717

Dryopteridaceae Dryopteridaceae Polystichum setigerum (C. Presl) C. Presl Polystichum braunii (Spenn) Fu var. alaskana (Maxon) Huttin U.S.A., ALASKA COUNTY: [locality and coordinates withheld; status in U.S.A., ALASKA COUNTY: Alaska: S3 (Rare; AKNHP)] Washington Bay, Kuiu Island. Elev. 30 ft. 56.721111°, -134.356907°; WGS 84, Riverbank about a quarter mile upriver from uncertainty: 2400 m., Source: Digital Map, estuary. Associated plants: Streptopus Georef'd by Joseph Stormer amplexifolia, Osmorhiza sp., Trisetum spicatum, In spruce and hemlock forest. Forest Service Tiarella trifoliate, Arnica latifolia, Viola sp., Alaska Region, TNFS Herbarium, v. 7517. Rubus specatabilis, Alnus sinuate. Forest Origin: Native. Service Alaska Region, TNFS Herbarium, v. 7304. Origin: Native. W. J. Eyerdam 8368 Brad Krieckhaus 342 25 Aug 1956 28 Jun 2005 WTU-408368, WTU-V-022712 WTU-408362, WTU-V-022718 Dryopteridaceae Dryopteridaceae Polystichum braunii (Spenner) Fée Polystichum munitum (Kaulf.) C. Presl U.S.A., ALASKA COUNTY: U.S.A., ALASKA COUNTY: Tongass National Forest. Baranof Island, east Mouth of Fillmore Inlet, point of land south of side. Big Port Walter. About 2 miles in from the Just Island, south of mooring cove. Prince mouth, north side, west end of narrows. Rupert D-2. Elev. 20 ft. Elev. 15 m. West edge of rock slide. Well drained wet area 54° 47' 38" N, 130° 38' 14" W; T82S R99E with mossy substrate which overlies jumbled S4 NW4NW4; Source: Specimen Label rocky unstable soil. Growing with P. munitum. On steep slope next to wet rock face in closed Other associated plants: Alnus rubra, Sambucus Sitka spruce western red cedar forest. Forest racemosa, Oplopanax horridum, miscellaneous Service Alaska Region, TNFS Herbarium, v. herbs. U.S. Forest Service Sitka, Alaska, 4802. Origin: Native. Herbarium, 01886. Origin: Native. J. DeLapp 93-385 Mary Clay Muller 4632 with M. Duffy 17 Aug 1981 9 Jul 1993 WTU-408361, WTU-V-022719 WTU-408358, WTU-V-022722

Dryopteridaceae Dryopteridaceae Polystichum lonchitis (L.) Roth Polystichum braunii (Spenner) Fée U.S.A., ALASKA COUNTY: U.S.A., ALASKA COUNTY: Skagway Quad.: Goat Lake, 7 miles northeast Southeast Alaska. Baranof Island. Mt. Verstovia of Skagway, south end of lake, south side of about 2 miles east of Sitka. outlet stream. Elev. 300 ft. Elev. 884 m. Growing in a rocky gully on the lower parts of 59° 31' 36" N, 135° 11' 11" W; Source: the mountain in a blowdown area. This year's Specimen Label frond. From same plant as 3457. U.S. Forest Herbaceous subalpine meadow adjacent to Service Sitka, Alaska, Herbarium, 01301. Origin: wide pool-stream area. Forest Service Alaska Native. Region, TNFS Herbarium, v. 5105. Origin: Native. Mary Clay Muller 3893 M. C. Stensvold 6106 25 Jul 1980 27 Jul 1993 WTU-408360, WTU-V-022720 WTU-408357, WTU-V-022723 Dryopteridaceae Dryopteridaceae Polystichum setigerum (C. Presl) C. Presl Polystichum lonchitis (L.) Roth U.S.A., ALASKA COUNTY: [locality and coordinates withheld; status in U.S.A., ALASKA COUNTY: Alaska: S3 (Rare; AKNHP)] Tongass National Forest. Near Mendenhall Mixed conifer subalpine forest. Open Glacier, about 10 miles northwest of Juneau. understory, mossy west-facing slope. Circa 244 Collected along East Glacier Trail. meters south southwest of peak 2443. Open area. Plants growing on moraine, this Limestone substrate. ALA V107042. U.S. Forest area recently deglaciated. Growing with P. Service Sitka, Alaska, Herbarium, 2898. Origin: braunii and P. andersonii. U.S. Forest Service Native. Sitka, Alaska, Herbarium, 01473. Origin: Native. M. C. Muller 5593 Mary Clay Muller 4194 with P. Alaback 30 Oct 1980 17 Aug 1990 WTU-408356, WTU-V-022724 WTU-408359, WTU-V-022721 Dryopteridaceae Dryopteridaceae Polystichum munitum (Kaulf.) C. Presl Polystichum lonchitis (L.) Roth U.S.A., IDAHO, SHOSHONE COUNTY: CANADA, BRITISH COLUMBIA: Coeur d'Alene National Forest: West side of Alamo Trail, 0.9 air km E of Idaho Peak, Central Eagle Creek just downstream of where Forest Kootenay Regional District. Service Road 805 crosses creek. Elev. 2095 m. Elev. 884 m. 49.9718° N, 117.2888° W; Source: Specimen 47.684871° N, 115.876866° W; WGS 84, Label uncertainty: 40 m., Source: GPS On steep w-facing rocky subalpine meadow, Mature forest of Thuja plicata, Abies grandis, partial shade or full sun, with Abies lasiocarpa, and Tsuga heterophylla, with Coptis Rhododendron albiflorum, Valeriana, Fragaria occidentalis, Asarum caudatum, Dryopteris spp. virginiana, Gentianella amarella, Epilobium Fronds evergreen, arching, tufted from a stout angustifolium. Leaves shiny, evergreen; root crown; common. Phenology: Spores. occasional on steep w-facing rocky subalpine Origin: Native. meadow. Origin: Native. Don Estberg 2016-09 Peter F. Zika 28349 9 Jul 2016 28 Aug 2016 WTU-420657, WTU-V-048549 WTU-412150, WTU-V-026598

Dryopteridaceae Dryopteridaceae Polystichum munitum (Kaulf.) C. Presl Polystichum munitum (Kaulf.) C. Presl U.S.A., IDAHO, SHOSHONE COUNTY: U.S.A., WASHINGTON, KING COUNTY: Coeur d'Alene National Forest. Forest Service Near S end of 32nd Avenue, Leschi, Seattle. Road 900. Area between road and Elev. 60 m. Independence Creek. 47.6° N, 122.291° W; Source: Specimen Elev. 2980 ft. Label 47° 52.996' N, 116° 12.663' W; WGS 84, Shade of deciduous forest, gentle E slope, with uncertainty: 500 m., Source: GPS Corylus cornuta, Oemleria cerasiformis, Acer Highly disturbed meadow with Hypericum macrophyllum. Cespitose, evergreen, common. perforatum, adjacent riparian corridor, dry forest Phenology: Spores. Origin: Native. understory and forested riparian zone. Uncommon on edge of forested riparian zone. Peter F. Zika 27584 Phenology: Spores. Origin: Native. 18 Oct 2015 Jim Duemmel 2016-10 WTU-406025, WTU-V-17985 9 Jul 2016 WTU-420665, WTU-V-048670 Dryopteridaceae Polystichum munitum (Kaulf.) C. Presl Dryopteridaceae U.S.A., WASHINGTON, KING COUNTY: Near S end of 32nd Avenue, Leschi, Seattle. Polystichum munitum (Kaulf.) C. Presl Elev. 60 m. 47.6° N, 122.291° W; Source: Specimen U.S.A., WASHINGTON, PIERCE COUNTY: Label No locality given. Shade of deciduous forest, gentle E slope, with Elev. 250 ft. Corylus cornuta, Oemleria cerasiformis, Acer T19N R2E S5; WGS 84, uncertainty: 62000 macrophyllum. Cespitose, evergreen, common. m., Source: Digital Map, Georef'd by Ibro Phenology: Vegetative. Polovina Moist conifer woods, open or deep shade, sea Peter F. Zika 27584 level to mid-mountain. Highly palatable fiddlehead in sprnig, otherwise low palatability. 18 Oct 2015 Origin: Native. WTU-406026, WTU-V-17986 Mengert 27 1 May 1977 WTU-405723, WTU-V-016769 Dryopteridaceae Dryopteridaceae Polystichum lonchitis (L.) Roth Polystichum lonchitis (L.) Roth U.S.A., IDAHO, BOUNDARY COUNTY: U.S.A., WASHINGTON, PEND OREILLE Colville N.F. Little Snowy Top. COUNTY: Elev. 6800 ft. Sullivan Lake Dist. Colville National Forest. 48.973542°, -117.003017°; WGS 84, T39N R45E S2; WGS 84, uncertainty: 2000 uncertainty: 1500 m., Source: GeoLocate, m., Source: GeoLocate, Georef'd by Crystal Georef'd by Dan Post Shin Growing from under edge of boulders and cliffs Moist areas above creek in cedar and hemlock on east side of Little Snowy Top. Phenology: forest. Phenology: Spores. Origin: Native. Vegetative. Origin: Native. N. Althauser s.n. T. E. Burke 175 with S, Althauser 23 Aug 1974 12 Aug 1979 WTU-410933, WTU-V-025247 WTU-410936, WTU-V-025258 Dryopteridaceae Dryopteridaceae Polystichum lonchitis (L.) Roth Polystichum lonchitis (L.) Roth U.S.A., WASHINGTON, STEVENS COUNTY: Washington State land, Trombetta Canyon, U.S.A., IDAHO, BOUNDARY COUNTY: about 2.5 mi. S of Northport. Take County R. Kaniksu [National Forest] administered by #9435 SE of Northport and then private roads to Colville. Little Snowy Top the SE corner of section 16. Walk NW about Elev. 6400-6800 ft. 0.5 mi. into Trombetta Canyon. T65N R5W S20 NW¼; WGS 84, uncertainty: Elev. 2500 ft. 569 m., Source: Calc. from TRS, Georef'd by T39N R40E S16 W½; WGS 84, uncertainty: Dan Post 3908 m., Source: GeoLocate, Georef'd by AF/Luhi Origin: Native. Crystal Shin Plants were growing on limestone cliffs with RCP 620 Dryas drummondii, Pellaea gastonyi, Asplenium trichomanes, Anemone cylindrica, Arenaria 23 Aug 1974 serpyllifolia and Alyssum desertorum. Origin: Native. WTU-410934, WTU-V-025248 Kathy Ahlenslager 540 4 Jul 1998 with Jean Wood Dryopteridaceae WTU-410935, WTU-V-025259 Polystichum andersonii M. Hopkins Dryopteridaceae U.S.A., WASHINGTON, STEVENS COUNTY: Polystichum braunii (Spenn) Fu var. alaskana (Maxon) Colville National Forest. Interrupted unmapped Huttin tributary of Thomason Creek, below the Flowery Trail Rd. east of Chewelah. U.S.A., ALASKA COUNTY: Elev. 3200 ft. Washington Bay, Kuiu Island. T32N R41W S3 NWNW; WGS 84, 56.721111°, -134.356907°; WGS 84, uncertainty: 1000 m., Source: Digital Map, uncertainty: 2400 m., Source: Digital Map, Georef'd by Crystal Shin Georef'd by Joseph Stormer Eight plants at spring/seep slump in short In shady ravine in alder woods. Origin: Native. trib[utary] to seasonal creek channel, slope 45%, aspect 260°. Rather open tree canopy. W. J. Eyerdam 8138 Associated spp: Larix occidentalis, Abies grandis, Thuja plicata, Pseudotsuga menziesii, 10 Aug 1955 Alnus viridis, Acer glabrum, Rubus parviflorus, Symphoricarpos albus, Linnaea borealis, WTU-414054, WTU-V-029526 Chimaphila umbellata, Rosa gymnocarpa, Ribes lacustre, Mahonia aquifolius, Pyrola secunda, Polystichum munitum, Gymnocarpum dryopteris, Osmorhiza chilensis, Galium triflorum, Disporum hoodii, Veronica officinalis, Maiathemum stellatum, Goodyear oblongifolia, Clintonia uniflora, Athyrium filix-femina, Dryopteris filix-mas, Asarum caudatum, moss. Phenology: Spores. Origin: Native. Linda Swartz 540 11 Aug 1999 WTU-410937, WTU-V-025257 Dryopteridaceae Dryopteridaceae Polystichum munitum (Kaulf.) C. Presl Polystichum squarrosum (D. Don) Fée U.S.A., ALASKA COUNTY: BHUTAN: Dixon Entrance Quad.: Dall Island, Grace On road from Haa to Thimpu, (Chhudzom-Haa Mountain. Ca. [about] 244 meters road), 23 air km SSE of Paro airstrip, foothills of south-southwest peak 2443. Himalayas. Elev. 640 m. Elev. 2555 54° 55' N, 132° 57' W; Source: Specimen 27.1995° 89.5057°; Source: Specimen Label Label S slope, occasional in shade of young stand of Mixed conifer subalpine forest. Open Pinus wallichiana, with grasses and other ferns. understory, mossy west-facing slope. Cespitose, large plants, evergreen, fronds Limestone substrate. Origin: Native. somewhat shiny. Sheet 1 of 2. Origin: Native. M. C. Muller 5595 Peter F. Zika 26958 with Alaback, P. 23 Mar 2015 17 Aug 1990 WTU-406196, WTU-V-17133 WTU-414071, WTU-V-029543

Dryopteridaceae Dryopteridaceae Polystichum imbricans (D.C. Eaton) D.H. Wagner ssp. Polystichum andersonii M. Hopkins imbricans U.S.A., WASHINGTON, KING COUNTY: U.S.A., CALIFORNIA, DEL NORTE COUNTY: Along West Fork Foss Lakes Trail ca 0.75 km north of Trout Lake. Above Road 427, 5.1 air km NW of Canthook Elev. 553 m. Mountain, Klamath Range (KR). 47.625271° N, 121.314799° W; WGS 84, uncertainty: 14 m., Elev. 45 m. Source: GPS, Georef'd by Ben Legler 41.801° N, 124.054° W; Source: Specimen Old, shaded, gravelly debris flow on side slope beneath canopy of Label young Acer macrophyllum; moist duff over gravel with Polystichum Steep ledges, NE aspect, full sun, with andersonii, P. munitum, Tolmeia menziesii, Dicentra formosa. Pentagramma, Eriogonum, Briza, Montia, Fronds tufted, arching, deciduous, each with a vegetative bud near Pseudotsuga, Alnus. Cespitose. Phenology: tip; pinnae divided nearly to midrib; plants locally common. Spores. Origin: Native. Phenology: Spores. Peter F. Zika 26984 Ben Legler 14677 7 Jul 2018 16 May 2015 Herbarium: WTU WTU-406002, WTU-V-17962 Dryopteridaceae Polystichum andersonii M. Hopkins × Polystichum Dryopteridaceae munitum (Kaulf.) C. Presl Polystichum squarrosum (D. Don) Fée U.S.A., WASHINGTON, KING COUNTY: Along West Fork Foss Lakes Trail ca 0.75 km north of Trout Lake. BHUTAN: Elev. 553 m. On road from Haa to Thimpu, (Chhudzom-Haa 47.625271° N, 121.314799° W; WGS 84, uncertainty: 14 m., road), 23 air km SSE of Paro airstrip, foothills of Source: GPS, Georef'd by Ben Legler Himalayas Old, shaded, gravelly debris flow on side slope beneath canopy of Elev. 2555 m. young Acer macrophyllum; moist duff over gravel with Polystichum 27.1995° N, 89.5057° E; Source: Specimen andersonii, P. munitum, Tolmeia menziesii, Dicentra formosa. Single Label plant among both parents; fronds arching, each with a vegetative S slope, occasional in shade of young stand of bud near tip; pinnae divided ca halfway to midrib. Phenology: Pinus wallichiana, with grasses and other ferns. Spores. Cespitose, large plants, evergreen, fronds somewhat shiny. Sheet 2 of 2. Origin: Native. Ben Legler 14678 7 Jul 2018 Peter F. Zika 26958 Herbarium: WTU 23 Mar 2015 WTU-406195, WTU-V-17132 Dryopteridaceae Dryopteridaceae Polystichum imbricans (D.C. Eaton) D.H. Wagner × Polystichum munitum (Kaulf.) C. Presl Polystichum scopulinum (D.C. Eaton) Maxon U.S.A., WASHINGTON, KING COUNTY: U.S.A., CALIFORNIA, SISKIYOU COUNTY: 0.8 air kilometer south of summit of Kelly Butte, 2 miles east White Mt. Siskiyou Mts. Cascade Range. Elev. 6300 ft. Elev. 1515 m. 41.948468°, -123.021357°; WGS 84, 47.162° N, 121.4889° W; Source: Specimen uncertainty: 1500 m., Source: Digital Map, Label Georef'd by Dan Post Subalpine talus slope, south aspect, with Shady north rock crevices. Origin: Native. Epilobium angustifolium, Lupinus latifolius, Vaccinium membranaceum, Paxistima Louis C. Wheeler 3172 myrsinites. Cespitose, evergreen, basal scales to 4 millimeters wide and with blackish central 16 Aug 1934 stripe, dwarf plants, uncommon. Phenology: Spores. Origin: Native. WTU-181JWT, WTU-V-030972 Peter F. Zika 29374 6 Aug 2018 Dryopteridaceae WTU-419524, WTU-V-042587 Polystichum chilense (Christ) Diels ARGENTINA, RíO NEGRO: Cerro Llao Llao, Peninsula Llao Llao, Lago Dryopteridaceae Nahuèl Huapi, 23 air km NW of San Carlos de Bariloche. Polystichum munitum (Kaulf.) C. Presl Elev. 990 m. 41.0443° S, 71.5562° W; Source: Specimen U.S.A., WASHINGTON, PIERCE COUNTY: Label JBLM, TA 10. Occasional on dry NW-facing forested slope, Elev. 115 m. shaded with Aristotelia chilensis, Austrocedrus 47.05692° N, 122.47941° W; WGS 84, Source: GPS chilensis, Berberis darwinii, Colletia hystrix, Common throughout understory of Pseudotsuga menziesii - Acer Fragaria chiloensis, Mutisia spinosa, macrophyllum forest. Origin: Native. Nothofagus dombeyi, Ribes magellanicum. Loosely cespitose. Origin: Native. Peter Dunwiddie PWD-16-229 13 Jun 2016 with Rod Gilbert, Sarah Krock, Peter Zika P. F. Zika 27616 Herbarium: WTU 29 Mar 2016 WTU-407915, WTU-V-021682 Dryopteridaceae Polystichum imbricans (D.C. Eaton) D.H. Wagner ssp. imbricans Dryopteridaceae U.S.A., CALIFORNIA, TEHAMA COUNTY: Polystichum imbricans (D.C. Eaton) D.H. Wagner Klamath Ranges/High North Coast Ranges; at the Tehama/Trinity County line; Stuart Gap, 2.7 U.S.A., OREGON, GILLIAM COUNTY: air kilometers north of North Yolla Bolly. Philippi Canyon Road Exit off I-84. Elev. 1525 m. 45.686299° N, 120.490138° W; Source: GoogleEarth, Georef'd by 40° 13' 44" N, 122° 58' 31" W; T28N R10W Sarah Winkowski NW 1/4 of SE 1/4 of Sec. 34; WGS 84, At foot of grassy slopes with some talus. Source: Specimen Label North-facing rock outcrop, +/- shaded by Peter Dunwiddie PWD-17-287 28 May 2017 Douglas-fir, Ponderosa pine, incense cedar, with Peter Zika, Stacy Kinsell white fir; also Corylus cornuta, Rubus virginiana. 7.5'x15' quad; pinnae rotated about 90 degrees Herbarium: WTU to the frond axis. Phenology: Spores. Origin: Native. Dryopteridaceae L. P. Janeway 12162 5 Apr 2016 with B. Castro Polystichum scopulinum (D.C. Eaton) Maxon Herbarium: WTU, WTU-V-043151 CANADA, BRITISH COLUMBIA: [locality and coordinates withheld; status in British Columbia: S2 (Imperiled, Red List; BCCDC)] Elev. 955 m. Base of shaded outcrop. Phenology: Spores. Origin: Native. Terry McIntosh 2011_99 with K. Sadler, M. Weston, A. Robinson 22 Jun 2011 WTU-417953, WTU-V-039955 Dryopteridaceae Dryopteridaceae Polystichum munitum (Kaulf.) C. Presl Polystichum acrostichoides (Michx.) Schott U.S.A., WASHINGTON, THURSTON COUNTY: U.S.A., ILLINOIS, TAZEWELL COUNTY: Scatter Creek North Unit Groveland Tp; Near East Peoria. Elev. 200 m. 40.592539°, -89.534542°; WGS 84, 46.84° N, 123.00293° W; WGS 84, Source: GPS uncertainty: 3036 m., Source: GeoLocate, Common in understory of Quercus garryana forest, with Georef'd by John Haskins Symphoricarpos albus Phenology: Fruits. Origin: Native. Wooded hillside. Peter Dunwiddie PWD-17-503 30 Jun 2017 Virginius H. Chase 3107 with Etsuko Reistroffer 10 Jun 1919 Herbarium: WTU WTU-201548, WTU-V-057820 Dryopteridaceae Rumohra adiantiformis (G. Forst.) Ching Dryopteridaceae AUSTRALIA, VICTORIA: Polystichum acrostichoides (Michx.) Schott Coleraine. -37.60142°, 141.69808°; WGS 84, U.S.A., PENNSYLVANIA, BERKS COUNTY: uncertainty: 3000 m., Source: GeoLocate, 2 miles north of Friedensburg. Georef'd by John Haskins Wooded hillside. Poisoned. Phenology: Spores. Hans Wilkens 991 A. MacCaskill s.n. 21 Jun 1931 1927 WTU-10194JWT, WTU-V-057819 WTU-10180JWT, WTU-V-057072

Dryopteridaceae Polystichum acrostichoides (Michx.) Schott Dryopteridaceae U.S.A., NEW YORK, CORTLAND COUNTY: Polystichum acrostichoides (Michx.) Schott [locality and coordinates withheld; status in New York: EV (Exploitably Vulnerable; USDA Plants)] U.S.A., MICHIGAN, OAKLAND COUNTY: Moist Acer-Fagus woods. Proud Lake. 42.565634°, -83.527814°; WGS 84, S. J. Smith 1133 uncertainty: 826 m., Source: Digital Map, with A. V. Leuchs, E. & M. Palmatier Georef'd by Ghislaine Ng In beech-maple woods. Fronds fertile only at 26 Jun 1939 the tips. WTU-94423, WTU-V-057826 Ira L. Wiggins 11949 31 Jul 1948 WTU-178956, WTU-V-057105 Dryopteridaceae Polystichum acrostichoides (Michx.) Schott Dryopteridaceae U.S.A., NORTH CAROLINA, VANCE COUNTY: Ruin Creek, 5.2 miles southwest of Bearpond. Polystichum acrostichoides (Michx.) Schott 36.230425°, -78.485277°; WGS 84, uncertainty: 3000 m., Source: GeoLocate, U.S.A., MASSACHUSETTS, FRANKLIN Georef'd by John Haskins COUNTY: Wooded slope. Deerfield River, Power Plant #2, Buckland. Along upper slope, side of railroad tracks. Harry E. Ahles 17395 with R. S. Leisner Stephen Garanin 7241 21 Jul 1956 5 Sep 1972 WTU-194358, WTU-V-057823 WTU-277912, WTU-V-057098 Dryopteridaceae Dryopteridaceae Polystichum acrostichoides (Michx.) Schott Polystichum californicum (D.C. Eaton) Diels U.S.A., NORTH CAROLINA, MECKLENBURG U.S.A., CALIFORNIA, SANTA CLARA COUNTY: COUNTY: East side of Mary Alexander Drive, UNC-C Foothills west of Los Gatos. campus, Charlotte. 37.234538°, -122.004206°; WGS 84, 35.308666°, -80.727427°; WGS 84, uncertainty: 5000 m., Source: Digital Map, uncertainty: 159 m., Source: Digital Map, Georef'd by Erik Petersen Georef'd by Ghislaine Ng Phenology: Spores. Beside creek. A. A. Heller 7225 Wayne C. Williams s.n. 11 Jan 1904 4 Jun 1966 WTU-155JWT, WTU-V-057827 WTU-227484, WTU-V-057088 Dryopteridaceae Dryopteridaceae Polystichum imbricans (D.C. Eaton) D.H. Wagner ssp. Polystichum munitum (Kaulf.) C. Presl imbricans U.S.A., WASHINGTON, YAKIMA COUNTY: U.S.A., OREGON, GILLIAM COUNTY: Forest Service Road 1902 approximately 2 kilometers west of Cliffs above Highway I-84, 3 miles east of John Day River. junction with Forest Road 19. 45.735807° N, 120.586609° W; WGS 84, Source: GPS Elev. 855 m. On steep, grassy slope at base of north-facing basalt cliffs. 1 plant, 47.01638° N, 121.14439° W; WGS 84, uncertainty: 100 m., fertile pinnae imbricate. With Pseudoroegneria spicata. Phenology: Source: GPS Immature. Origin: Native. South-facing basalt outcropping and adjacent seasonally moist open slope with loose soil; Penstemon richardsonii and Bromus carinatus Peter Dunwiddie PWD-18-100 10 May 2018 common. Uncommon in north-facing rocky outcropping. Phenology: with Peter Zika Spores. Origin: Native. Herbarium: WTU David Giblin 4998 20 Jun 2013 Herbarium: WTU Dryopteridaceae Polystichum braunii (Spenner) Fée Dryopteridaceae U.S.A., IOWA, VAN BUREN COUNTY: Polystichum aculeatum (L.) Roth. Bentonsport. 40.725586°, -91.85378°; WGS 84, PHILIPPINES, BENGUET: uncertainty: 1000 m., Source: GeoLocate, Island Luzon Mt. Pulog. Georef'd by Ghislaine Ng Elev. 2700 ft. From plant growing in my fernery. But plant Phenology: Spores. came originally from Superior, Wis. E. B. Copeland 162 E. W. Graves s.n. May 1909 1931 WTU-10186JWT, WTU-V-057142 WTU-10212JWT, WTU-V-056994

Dryopteridaceae Dryopteridaceae Polystichum braunii (Spenner) Fée Polystichum aculeatum (L.) Roth. CANADA, ONTARIO: Smith Lake. BOLIVIA: 47° N, 84° 45' W; WGS 84, uncertainty: 1276 Insel Coaty (Mondiusel) in Titicacasee. m., Source: GeoLocate, Georef'd by Ghislaine Elev. 3840 m. Ng Poisoned. Phenology: Spores. Woods among rooted lava talus. Buchtien 2802 T. M. C Taylor 131 with R. C. Hosie, R. E. Fitzpatrick, S. T. Losee, Mar 1910 A. Leslie. WTU-10181JWT, WTU-V-057143 15 Jul 1935 WTU-121586, WTU-V-056991 Dryopteridaceae Dryopteridaceae Polystichum braunii (Spenner) Fée Polystichum braunii (Spenner) Fée U.S.A., VERMONT, ORLEANS COUNTY: CANADA, ONTARIO: Lower east slope of Mt. Hor. Batchawana Falls. 44.708956°, -72.046769°; WGS 84, 46.98775°, -84.525413°; WGS 84, uncertainty: 677 m., Source: Digital Map, uncertainty: 619 m., Source: Digital Map, Georef'd by Ghislaine Ng Georef'd by Ghislaine Ng Moist rocky woods. Damp rocky woods. George R. Proctor 1590 T. M. C Taylor 7418 6 Jul 1945 27 Jun 1967 WTU-97426A, WTU-V-056988 WTU-371006, WTU-V-056968

Dryopteridaceae Dryopteridaceae Polystichum braunii (Spenner) Fée Polystichum imbricans (D.C. Eaton) D.H. Wagner CANADA, ONTARIO: U.S.A., CALIFORNIA, SISKIYOU COUNTY: Near top of east ridge; Havilland Bay. Above McBride Springs. 47° N, 84° 45' W; WGS 84, uncertainty: 647 Elev. 5000 ft. m., Source: Digital Map, Georef'd by 41.353969°, -122.281678°; WGS 84, Ghislaine Ng uncertainty: 90 m., Source: Digital Map, Damp rocky woods. Georef'd by Ghislaine Ng In cracks between lava boulders on the lava T. M. C Taylor 137 flow. with R. C. Hosie, R. E. Fitzpatrick, S. T. Losee, A. Leslie. W. B. Cooke 17732 13 Aug 1935 25 Jul 1947 WTU-137151, WTU-V-056985 WTU-134863, WTU-V-056926

Dryopteridaceae Dryopteridaceae Polystichum braunii (Spenner) Fée Polystichum imbricans (D.C. Eaton) D.H. Wagner U.S.A., WISCONSIN, IRON COUNTY: U.S.A., CALIFORNIA, TRINITY COUNTY: [locality and coordinates withheld; status in Canyon Creek(tributary of Trinity River). Wisconsin: T (Threatened; USDA Plants)] Elev. 5500 ft. Moist shady mossy rocks. Abundant. 40.685347°, -123.508597°; WGS 84, uncertainty: 406 m., Source: Digital Map, A.M.Fuller 4497 Georef'd by Ghislaine Ng Under edge of rock, granite slope. 11 Aug 1931 Annie M. Alexander 5478 WTU-85241, WTU-V-056977 with Louise Kellogg 11 Aug 1948 WTU-184301, WTU-V-056922 Dryopteridaceae Polystichum braunii (Spenner) Fée Dryopteridaceae CANADA, ONTARIO: Polystichum imbricans (D.C. Eaton) D.H. Wagner Batchawana Falls. 46.986548°, -84.52301°; WGS 84, U.S.A., CALIFORNIA, TULARE COUNTY: uncertainty: 589 m., Source: Digital Map, Along ridge of Mule Peak, 100 yds. Southwest Georef'd by Ghislaine Ng of lookout. Sequoia N.F. Damp rocky woods. Elev. 8100 ft. 35.993146°, -118.61687°; WGS 84, T. M. C Taylor 7418 uncertainty: 2317 m., Source: Digital Map, Georef'd by Ghislaine Ng 27 Jun 1967 In crevices of decomposing granite. WTU-371010, WTU-V-056969 A. R. Kruckeberg 3377 2 Jul 1954 WTU-189773, WTU-V-056921 Dryopteridaceae Dryopteridaceae Polystichum imbricans (D.C. Eaton) D.H. Wagner Polystichum imbricans (D.C. Eaton) D.H. Wagner ssp. imbricans U.S.A., CALIFORNIA, PLACER COUNTY: [no locality given on label]. U.S.A., CALIFORNIA, TEHAMA COUNTY: 39.055597°, -120.806169°; WGS 84, On the North side of Deer Creek Trail, between uncertainty: 66545 m., Source: Digital Map, the fish ladder and Big Leaf Pond, about 1 3/4 Georef'd by Ghislaine Ng miles by trail to Highway 32. Elev. 2900 ft. Matt Herbrand(of Cisco) s.n. 40° 10' 11" N, 121° 34' 13.3" W; T27N, R3E, near center of Section 25. ; WGS 84, 20 Dec 1917 uncertainty: 90 m., Source: GeoLocate, Georef'd by Ghislaine Ng WTU-214193, WTU-V-056920 On damp soil; mixed Evergreen Forest. From a many fronded plant; normal size fronds; common. Lowell Ahart 20,698-A Dryopteridaceae with John Dittes, Josephine Guardino. Polystichum imbricans (D.C. Eaton) D.H. Wagner 11 Feb 2016 U.S.A., CALIFORNIA, TUOLUMNE COUNTY: WTU-412073, WTU-V-056939 Old Colterville road, 1 mile from main Big Oak Flat road. 37.823539°, -120.258246°; WGS 84, Dryopteridaceae uncertainty: 301 m., Source: GeoLocate, Georef'd by Ghislaine Ng Polystichum imbricans (D.C. Eaton) D.H. Wagner ssp. imbricans Ira L. Wiggins 9243 U.S.A., CALIFORNIA, PLUMAS COUNTY: 20 Jun 1939 On the west side of Slate Creek Reservoir, about 100 yards northwest of the intake WTU-62958, WTU-V-056919 building, about 2 1/4 miles(air) northeast of La Porte Road, about 6 miles (air) southwest of La Porte. Elev. 3701 ft. Dryopteridaceae 39° 37' 0.2" N, 121° 3' 5.9" W; T20N, R8E, southwest 1/4 Section 2. ; WGS 84, Polystichum imbricans (D.C. Eaton) D.H. Wagner uncertainty: 969 m., Source: GeoLocate, Georef'd by Ghislaine Ng U.S.A., CALIFORNIA, DEL NORTE COUNTY: On damp soil, rock outcrops, filtered shade, Siskiyou National Forest; on hwy. 40S03, 14.1 steep mountain side, mixed Evergreen Forest. m S of intersection of 40S03 and Waldo Rd. A few fronds from several small clumps. Fronds (intersection 4 m E of O'Brien); above Whiskey upright, twisted. Common, scattered. Lake. Elev. 5160 ft. Lowell Ahart 20,578 23 Oct 2015 41.924597°, -123.645407°; WGS 84, uncertainty: 2000 m., Source: Digital Map, WTU-413563, WTU-V-056938 Georef'd by Ghislaine Ng Serpentine slopes. With Pinus attenuata, Dryopteridaceae Pseudotsugam Ceanothus pumilus, and Arctostaphylos; frequent. Polystichum imbricans (D.C. Eaton) D.H. Wagner ssp. imbricans Melinda F. Denton 2911 U.S.A., CALIFORNIA, TEHAMA COUNTY: 30 Jun 1973 Near the fish ladder on Deer Creek, about 2 miles by trail to Highway 32. Foothill Woodland. WTU-258969, WTU-V-056918 Elev. 2900 ft. 40° 10' 10.1" N, 121° 34' 52" W; T27N, R3E, near center of Section 25. ; WGS 84, uncertainty: 90 m., Source: GeoLocate, Dryopteridaceae Georef'd by Ghislaine Ng On damp soil. Normal size fronds, from a many Polystichum lemmonii Underw. fronded plant. Common. U.S.A., CALIFORNIA COUNTY: Lowell Ahart 20,698 Scott Mt., N. side of summit. with John Dittes, Josephine Guardino. 41.238819°, -122.730835°; WGS 84, uncertainty: 4756 m., Source: Digital Map, 11 Feb 2016 Georef'd by Ghislaine Ng Growing in crevices between rocks. WTU-412068, WTU-V-056937 J. Tucker 1572 17 May 1947 WTU-213587, WTU-V-056949 Dryopteridaceae Dryopteridaceae Polystichum imbricans (D.C. Eaton) D.H. Wagner ssp. Polystichum imbricans (D.C. Eaton) D.H. Wagner ssp. curtum (Ewan) D.H. Wagner curtum (Ewan) D.H. Wagner U.S.A., CALIFORNIA, COLUSA COUNTY: U.S.A., CALIFORNIA, BUTTE COUNTY: On the east side of the gravel road, about 1/2 Macabes Creek Canyon ca. 100 ft E of Lumpkin mile north of Letts Lake, about 1/2 mile south of Rd, ca. 2.5 mi N of its jct with Forbestown Road, the large campground, about 13 miles (air) ca. 18 mi NE of Oroville. southwest of Stonyford. Elev. 1300 ft. Elev. 3170 ft. T19N R6E sw sec 8. ; WGS 84, uncertainty: 39° 19' 37.3" N, 122° 43' 29.5" W; T17N, 90 m., Source: GeoLocate, Georef'd by R8W, southeast 1/4 Section 24. ; WGS 84, Ghislaine Ng uncertainty: 969 m., Source: GeoLocate, Cool, wet Ripar ian Woodland. Georef'd by Ghislaine Ng On damp soil; mixed Evergreen Forest. Normal M. S. Taylor 2388 size fronds, from a many fronded plant; common. 19 Mar 1980 Lowell Ahart 20,721 WTU-325180, WTU-V-056933 with John Dittes, Josephine Guardino. 14 Feb 2016 Dryopteridaceae WTU-412076, WTU-V-056936 Polystichum imbricans (D.C. Eaton) D.H. Wagner ssp. curtum (Ewan) D.H. Wagner Dryopteridaceae U.S.A., CALIFORNIA, YUBA COUNTY: On the west side of the road of Slate Creek, Polystichum imbricans (D.C. Eaton) D.H. Wagner ssp. about 1/4 mile southwest of Slate Creek Dam, imbricans about 2 miles (air) northwest of La Porte Road, about 6 1/4 miles (air) southwest of La Porte. U.S.A., CALIFORNIA, YUBA COUNTY: Elev. 3673 ft. On the west side of the road to Slate Creek, 39° 36' 35.5" N, 121° 3' 19.1" W; T20N, R8E, about 1/8 mile southwest of the Slate Creek northwest 1/4 Section 11. ; WGS 84, Dam, about 2 1/8 miles (air) northeast of La uncertainty: 969 m., Source: GeoLocate, Porte Road, about 6 miles (air) southwest of La Georef'd by Ghislaine Ng Porte. On damp soil, in shade, steep mountain side, Elev. 3818 ft. mixed Evergreen Forest. Fronds from one large 39° 36' 18" N, 121° 3' 35.5" W; T20N, R8E, clump. Common, scattered. Fronds flat, arched northwest 1/4 Section 11.; WGS 84, downward. uncertainty: 969 m., Source: GeoLocate, Georef'd by Ghislaine Ng Lowell Ahart 20,580 23 Oct 2015 On damp soil, rock outcrops, some sun, steep mountain side, mixed Evergreen Forest. Fronds WTU-413569, WTU-V-056931 from one large clump. Common, scattered. Fronds upright, twisted. Dryopteridaceae Lowell Ahart 20,579 23 Oct 2015 Polystichum californicum (D.C. Eaton) Diels WTU-413570, WTU-V-056935 U.S.A., CALIFORNIA, BUTTE COUNTY: Dryopteridaceae On the east side of Big Chico Creej, about 40 yards west of Ten Mile House Road, about 3/4 Polystichum imbricans (D.C. Eaton) D.H. Wagner ssp. mile (air) west of Highway 32, about 9 miles (air) curtum (Ewan) D.H. Wagner northeast of Chico. Elev. 649 ft. U.S.A., CALIFORNIA, PLUMAS COUNTY: 39° 48' 29.5" N, 121° 43' 27.7" W; T22N, On the west side of Slate Creek Reservoir, R2E, southwest 1/4 Section 35.; WGS 84, about 100 yards northwest of the intake uncertainty: 969 m., Source: GeoLocate, building, about 2 1/4 miles (air) northeast of La Georef'd by Ghislaine Ng Porte Road, about 6 miles (air) southwest of La On damp soil, in deep shade from trees and Porte. rocks, riparian woodland. Fronds from one Elev. 3701 ft. clump, uncommon, fronds flat, arched 39° 37' 0.2" N, 121° 3' 5.9" W; T20N, R8E, downward. southwest 1/4 Section 2. ; WGS 84, uncertainty: 969 m., Source: GeoLocate, Lowell Ahart 20,585 Georef'd by Ghislaine Ng On damp soil, filtered shade, steep mountain 24 Oct 2015 side, mixed Evergreen Forest. A few fronds from several small clumps. Common, scattered. WTU-413566, WTU-V-057829 Fronds flat, arched downward. Lowell Ahart 20,577 23 Oct 2015 WTU-413564, WTU-V-056934 Dryopteridaceae Dryopteridaceae Polystichum acrostichoides (Michx.) Schott Polystichum californicum (D.C. Eaton) Diels U.S.A., INDIANA, MORGAN COUNTY: U.S.A., CALIFORNIA, SONOMA COUNTY: Along Syacamore Creek, 2 miles west Sonoma Canyon, near Kenwood. Centerton. 38.4°, -122.5333°; WGS 84, uncertainty: 301 39.514777°, -86.440297°; WGS 84, m., Source: GeoLocate, Georef'd by Ghislaine uncertainty: 3649 m., Source: Digital Map, Ng Georef'd by Ghislaine Ng Wooded ravine. W. H. Wagner s.n. Ray C. Friesner 8805 30 Oct 1948 29 Jun 1935 WTU-213588, WTU-V-057831 WTU-62483, WTU-V-057117

Dryopteridaceae Dryopteridaceae Polystichum acrostichoides (Michx.) Schott Polystichum craspedosorum (Maxim.) Diels U.S.A., MISSOURI, JEFFERSON COUNTY: JAPAN, OMI: Along McNamee Road, ca. 3/4 mi N of its jct. Hondo: Mt. Tbuki. with State Road F, ca, 4 air mi SW of Eureka. Phenology: Spores. Very wet, overgrown road embankment. Shaded fern on moist soil; upper parts of fronds M. Togasi 450 dimorphically fertile. with M. Matuoka. M. S. Taylor 6017 3 Nov 1952 26 May 1990 WTU-180981, WTU-V-056962 WTU-331985, WTU-V-057103

Dryopteridaceae Dryopteridaceae Polystichum acrostichoides (Michx.) Schott Polystichum scopulinum (D.C. Eaton) Maxon U.S.A., OKLAHOMA, MCCURTAIN COUNTY: U.S.A., CALIFORNIA, SISKIYOU COUNTY: 1.5 miles east of State Highway 21 on road to Above Panther Creek Meadows, Mt. Shasta. Beavers Bend State Park. Elev. 9000 ft. 34.143°, -94.7008°; WGS 84, uncertainty: 41.282879°, -122.141274°; WGS 84, 2000 m., Source: GeoLocate, Georef'd by uncertainty: 2278 m., Source: Digital Map, Ghislaine Ng Georef'd by Ghislaine Ng Seepage area in oak-hickory-pine woods. Under boulders on ridge in basin. G. T. Robbins 2833 W. B. Cooke 17611 with V. G. Cooke 25 Oct 1947 29 Aug 1946 WTU-184355, WTU-V-057091 WTU-135673, WTU-V-056765 Dryopteridaceae Polystichum lonchitis (L.) Roth Dryopteridaceae CANADA, ALBERTA: Polystichum scopulinum (D.C. Eaton) Maxon Picea-Pseudotsuga forest; near Cameron Lake, Waterton Lakes Nat'l. Park. U.S.A., CALIFORNIA, SISKIYOU COUNTY: Elev. 5900 ft. On s.w. -facing brushy talus between Cook and 49.045884°, -114.022575°; WGS 84, Green Pass and Red Butte. uncertainty: 2366 m., Source: Digital Map, 41.466343°, -123.020927°; WGS 84, Georef'd by Ghislaine Ng uncertainty: 5432 m., Source: Digital Map, Georef'd by Ghislaine Ng C. L. Hitchcock 7925 Massive peridotite talus w/ Quercus vaccinifolia, with J.S.Martin Pinus monticola, Arctostaphylos sp. 7 Aug 1941 Arthur R. Kruckeberg 6212 WTU-68936, WTU-V-056724 10 Jun 1972 WTU-256933, WTU-V-056764 Dryopteridaceae Dryopteridaceae Polystichum platyphyllum (Willd.) C. Presl Polystichum munitum (Kaulf.) C. Presl ECUADOR, CHIMBORAZO: U.S.A., CALIFORNIA, SONOMA COUNTY: Canon of the rio Chanchan, about 5 km. north of Rancho del Paradiso, Russian River. Huigra. 38.901932°, -123.061445°; WGS 84, Elev. 5000-6500 ft. uncertainty: 6943 m., Source: Digital Map, -2.243834°, -78.983293°; WGS 84, Georef'd by Ghislaine Ng uncertainty: 6536 m., Source: GeoLocate, A dry form with imbricate pinnae. Growing Georef'd by Ghislaine Ng wherever the woods have been cleared, or Moist forested valleys in the afternoon fogbelt. where the plants are otherwise exposed. In Lvs deep green above, pale below. Sori naked. shaddier places the plants are 3 -5 times as (See #3417A). large. W. H. Camp E-3417 W. H. Wagner Jr. 4650 19 May 1945 18 Aug 1946 WTU-143899, WTU-V-056761 WTU-213586, WTU-V-056742

Dryopteridaceae Dryopteridaceae Polystichum munitum (Kaulf.) C. Presl Polystichum rhizophyllum (Sw.) C. Presl U.S.A., CALIFORNIA, TRINITY COUNTY: U.S.A., PUERTO RICO COUNTY: .5 mile above Ripstein Camp, Canyon Creek. Between Cayey and Guayama. Elev. 2600 ft. 18.050736°, -66.141243°; WGS 84, 40.952176°, -123.030235°; WGS 84, uncertainty: 4034 m., Source: Digital Map, uncertainty: 8047 m., Source: Digital Map, Georef'd by Ghislaine Ng Georef'd by Ghislaine Ng On moist soil, wooded slope. Semi-shade on rock, with humus Mixed forest among Philadelphus, Corylus. H.L. Blomquist 13095 E. K. Balls 13678 12 Apr 1942 30 Jun 1949 WTU-93982, WTU-V-056756 WTU-132598, WTU-V-056740

Dryopteridaceae Dryopteridaceae Polystichum munitum (Kaulf.) C. Presl Polystichum munitum (Kaulf.) C. Presl U.S.A., CALIFORNIA, SAN FRANCISCO COUNTY: U.S.A., CALIFORNIA, COLUSA COUNTY: Mt. Davidson. On the east side of the gravel road, about 1/2 37.738259°, -122.45463°; WGS 84, mile north of Letts Lake, about 1.2 mile south of uncertainty: 462 m., Source: GeoLocate, the large campground, about 13 miles (air) Georef'd by Ghislaine Ng southwest of Stonyford. Damp wooded slopes. Elev. 3170 ft. 39° 19' 48.1" N, 122° 43' 6.4" W; T17N, R8W, Lewis S. Rose 34007 southwest 1/4 Section 24. ; WGS 84, uncertainty: 969 m., Source: GeoLocate, 28 Jan 1934 Georef'd by Ghislaine Ng On wet soil, on the edge of a small pond, deep WTU-41365, WTU-V-056748 shade, mixed Evergreen Forest. Normal size frond, from a many fronded large plant, uncommon. Dryopteridaceae Lowell Ahart 20,720 14 Feb 2016 Polystichum munitum (Kaulf.) C. Presl with John Dittes, Josephine Guardino. U.S.A., CALIFORNIA, SAN MATEO COUNTY: WTU-412071, WTU-V-056734 La Honda road, east slope of Coast Range above Searsville Lake. Elev. 1000 ft. 37.40954°, -122.245487°; WGS 84, uncertainty: 4673 m., Source: Digital Map, Georef'd by Ghislaine Ng With Scoliopus bigelovii, Aspidium rigidum argutum, and Corylus rostrata californica, in wet shade of Sequoia sempervirens. Common. David D. Keck 1360 12 Feb 1932 WTU-22931, WTU-V-056746 Dryopteridaceae Dryopteridaceae Polystichum lonchitis (L.) Roth Polystichum acrostichoides (Michx.) Schott U.S.A., IDAHO, BONNEVILLE COUNTY: U.S.A., MISSOURI, MARION COUNTY: Caribou Range, southeastern end of Big Elk South of Oakwood. Mountain, 9 airline miles south of Palisades 39.683474°, -91.362485°; WGS 84, Dam. uncertainty: 1169 m., Source: Digital Map, Elev. 8400 ft. Georef'd by Ghislaine Ng T.2 S., R.45E., Sec. 31. ; WGS 84, uncertainty: 301 m., Source: GeoLocate, Rev. John Davis s.n. Georef'd by Ghislaine Ng Common on open rock outcrop surrounded by 10 Jul 1913 firs. WTU-56193, WTU-V-057085 Noel H. Holmgren 5571 with Vernon Marttala 22 Jul 1971 Dryopteridaceae WTU-263348, WTU-V-056723 Polystichum acrostichoides (Michx.) Schott U.S.A., MAINE, PENOBSCOT COUNTY: Dryopteridaceae Bangor. 44.801182°, -68.777814°; WGS 84, Polystichum lonchitis (L.) Roth uncertainty: 1000 m., Source: GeoLocate, Georef'd by Ghislaine Ng U.S.A., MONTANA, MISSOULA COUNTY: Deep woods. One mi. S.E. of Upper Holland Lake; Flathead Range; Flathead National Forest. O. W. Knight s.n. 47.450537°, -113.508592°; WGS 84, uncertainty: 1561 m., Source: Digital Map, 22 Jul 1905 Georef'd by Ghislaine Ng WTU-124873, WTU-V-057084 CLHitchcock 18450 15 Jul 1948 Dryopteridaceae WTU-126015, WTU-V-056722 Polystichum acrostichoides (Michx.) Schott U.S.A., TENNESSEE, SEVIER COUNTY: Dryopteridaceae Dupont Community, N. slope of Sugar Loaf Mt. 35.8493°, -83.681518°; WGS 84, uncertainty: Polystichum lonchitis (L.) Roth 2068 m., Source: Digital Map, Georef'd by Ghislaine Ng U.S.A., NEVADA, ELKO COUNTY: To Island Lake Ruby Range. Ken Whitman 80 40.615248°, -115.368846°; WGS 84, with Bot. 205, et. al. uncertainty: 1305 m., Source: Digital Map, Georef'd by Ghislaine Ng 25 Apr 1968 Moist protected places in crevices of rocks, cascades. Frequent. WTU-242007, WTU-V-057082 Bassett Maguire 22635 with A. H. Holmgren Dryopteridaceae 18 Aug 1943 Polystichum acrostichoides (Michx.) Schott WTU-72436, WTU-V-056718 U.S.A., IOWA, VAN BUREN COUNTY: Bentonsport. 40.725586°, -91.85378°; WGS 84, Dryopteridaceae uncertainty: 1000 m., Source: GeoLocate, Georef'd by Ghislaine Ng Polystichum acrostichoides (Michx.) Schott Damp shady banks. U.S.A., MASSACHUSETTS, HAMPSHIRE E. W. Graves s.n. COUNTY: North of Amherst. 1928 42.381382°, -72.520225°; WGS 84, uncertainty: 851 m., Source: Digital Map, WTU-10217JWT, WTU-V-057081 Georef'd by Ghislaine Ng Mixed woods. H. E. Ahles 64696 18 Sep 1966 WTU-253703, WTU-V-057086 Dryopteridaceae Dryopteridaceae Polystichum acrostichoides (Michx.) Schott Polystichum U.S.A., LOUISIANA, CALDWELL COUNTY: U.S.A., VIRGINIA COUNTY: 8 mi. N. of Columbia on Hwy. 546. Cass. W. Va.. 32.221261°, -92.077913°; WGS 84, uncertainty: 11293 m., Source: GeoLocate, F.W. Gray s.n. Georef'd by Ghislaine Ng 1927 M. Morris 14 with Bot. 205. WTU-316711, WTU-V-056678 6 Mar 1968 WTU-242006, WTU-V-057080

Dryopteridaceae Dryopteridaceae Polystichum acrostichoides (Michx.) Schott Polystichum munitum (Kaulf.) C. Presl U.S.A., SOUTH CAROLINA, NEWBERRY U.S.A., OREGON, WASCO COUNTY: COUNTY: 6.8 miles east of The Dalles Dam on south side of I-84. [Original 0.4 miles nw. of Brown's Crossroad. (ne. of label mistakenly says Sherman]. Newberry). 45.645341° N, 120.99547° W; WGS 84, Source: GPS 34.310454°, -81.570126°; WGS 84, One plant in wet crevice on north-facing basalt cliff, with Anaphalis uncertainty: 1799 m., Source: Digital Map, margaritacea, Solidago lepida. Origin: Native. Georef'd by Ghislaine Ng Pine woodland with exposed granite. Peter F. Zika PWD-19-407 2 Aug 2019 with Peter Dunwiddie C. Ritchie Bell 9105 Herbarium: WTU 24 Jun 1957 WTU-230005, WTU-V-057079 Dryopteridaceae Polystichum acrostichoides (Michx.) Schott Dryopteridaceae U.S.A., VIRGINIA COUNTY: Polystichum imbricans (D.C. Eaton) D.H. Wagner ssp. Shenandoah National Park. N. of Skyland. imbricans 38.491772°, -78.469102°; WGS 84, uncertainty: 2289 m., Source: GeoLocate, U.S.A., OREGON, SHERMAN COUNTY: Georef'd by Ghislaine Ng South side of highway I-84 one mile west of LePage Park. Scattered in open woods. 45.71943° N, 120.673527° W; WGS 84, Source: GPS One plant at base of basalt cliff along roadside, with Secale cereale, P. M. Mazzeo 1699 Phacelia hastata, Asclepia fasciculatus. Origin: Native. 27 Aug 1966 Peter Dunwiddie PWD-19-369 31 Jul 2019 with Peter Zika WTU-228494, WTU-V-057087 Herbarium: WTU Dryopteridaceae Dryopteridaceae Polystichum acrostichoides (Michx.) Schott Polystichum U.S.A., VERMONT COUNTY: U.S.A., VIRGINIA COUNTY: N. H. line, 80 mi. N. of Dartmouth. [No locality given on label]. Moist woods. 37.570962°, -80.027549°; WGS 84, uncertainty: 394289 m., Source: GeoLocate, C. L. Hitchcock 2757 Georef'd by Joseph Stormer Phenology: Spores. 16 Jun 1935 Elva Lawton s.n. WTU-38535, WTU-V-057092 18 Sep 1926 WTU-325503, WTU-V-056677 Dryopteridaceae Dryopteridaceae Polystichum acrostichoides (Michx.) Schott Polystichum acrostichoides (Michx.) Schott U.S.A., MASSACHUSETTS COUNTY: U.S.A., MASSACHUSETTS COUNTY: Rich woods, Stoughton. Woodland Spencer. 42.122172°, -71.107936°; WGS 84, 42.243981°, -71.992297°; WGS 84, uncertainty: 1222 m., Source: Digital Map, uncertainty: 2229 m., Source: GeoLocate, Georef'd by Ghislaine Ng Georef'd by Ghislaine Ng S. F. Blake s.n. G. E. Stone s.n. 3 Sep 1932 1886 WTU-10187JWT, WTU-V-057093 WTU-146055, WTU-V-057108

Dryopteridaceae Dryopteridaceae Polystichum acrostichoides (Michx.) Schott Polystichum acrostichoides (Michx.) Schott CANADA, NOVA SCOTIA, INVERNESS: U.S.A., NEW YORK COUNTY: Cape Breton Is. N side of Anse River, 3 km SW [locality and coordinates withheld; status in New of Sammys Barren. York: EV (Exploitably Vulnerable; USDA Plants)] Elev. 75 m. Woods. Common. 46° 49' N, 60° 44' W; WGS 84, uncertainty: 1911 m., Source: GeoLocate, Georef'd by Stewart H. Burnham 1387 Ghislaine Ng Occasional in 300 year-old hardwood forest, 23 Nov 1914 with Acer saccharum, Fagus grandifolius, Trientalis borealis. WTU-124793, WTU-V-057112 P. F. Zika 13970 29 Jul 1999 WTU-339893, WTU-V-057101 Dryopteridaceae Polystichum acrostichoides (Michx.) Schott Dryopteridaceae U.S.A., NEW YORK, SUFFOLK COUNTY: Polystichum acrostichoides (Michx.) Schott [locality and coordinates withheld; status in New York: EV (Exploitably Vulnerable; USDA Plants)] U.S.A., MAINE, SOMERSET COUNTY: Northeast corner of Detroit township. E. L. s.n. 44° 45' N, 69° 15' W; WGS 84, uncertainty: 986 m., Source: Digital Map, Georef'd by 31 Jul 1922 Ghislaine Ng Secondary forest of Betula, Tsuga, Populus. WTU-316708, WTU-V-057113 Sori brown, leaves in rosettes, evergreen, dark green. James F. Smith 2719 23 Dec 1993 Dryopteridaceae WTU-328844, WTU-V-057104 Polystichum acrostichoides (Michx.) Schott Dryopteridaceae U.S.A., PENNSYLVANIA, WESTMORELAND Polystichum acrostichoides (Michx.) Schott COUNTY: Trafford City. U.S.A., INDIANA, PARKE COUNTY: 40.384934°, -79.757381°; WGS 84, [no locality given on label]. uncertainty: 701 m., Source: Digital Map, 39.789136°, -87.219238°; WGS 84, Georef'd by Ghislaine Ng uncertainty: 20313 m., Source: Digital Map, Georef'd by Ghislaine Ng Elva Lawton s.n. R. F. Daubenmire s.n. 15 Jul 1924 22 Aug 1930 WTU-316709, WTU-V-057114 WTU-175700, WTU-V-057107 Dryopteridaceae Dryopteridaceae Polystichum californicum (D.C. Eaton) Diels Polystichum munitum (Kaulf.) C. Presl U.S.A., CALIFORNIA, HUMBOLDT COUNTY: U.S.A., CALIFORNIA, SIERRA COUNTY: Halfway between Miranda and Garberville. On the north side of the Canyon Creek Trail, Vicinity of Gaberville. about 50 yards southeast of the foot bridge Elev. 500 ft. across Boyce Ravine, trail on the north side of 40.170887°, -123.793504°; WGS 84, the North Fork of the Yuba River, about one uncertainty: 8676 m., Source: Digital Map, mile west of highway 49, about 2 miles west of Georef'd by Ghislaine Ng Indian Valley. Stream bank in shade. Phenology: Spores. Elev. 2352 ft. 39° 30' 57.8" N, 121° 2' 5" W; T19N, R8E, Harold E. Parks 11547 northwest 1/4 Section 13. ; WGS 84, with Joseph P. Tracy. uncertainty: 4652 m., Source: GeoLocate, Georef'd by Ghislaine Ng 16 May 1942 On good dry rocky soil, road to trail cut. Mixed Evergreen Forest. Small size frond, some WTU-207304, WTU-V-057828 plants four feet tall. Common. Lowell Ahart 20,154 9 May 2015 Dryopteridaceae WTU-413559, WTU-V-056735 Polystichum lonchitis (L.) Roth Dryopteridaceae CANADA, ONTARIO: Carp Lake. Polystichum muricatum (L.) Fée 45° N, 84° 45' W; WGS 84, uncertainty: 1189 m., Source: Digital Map, Georef'd by MEXICO, GUERRERO: Ghislaine Ng Teotepec, Galeana. Elev. 2450 m. T. M. C Taylor 2445 By stream in oak and pine forest. with R. C. Hosie, R. E. Fitzpatrick, S. T. Losee, A. Leslie. Geo. B. Hinton 14288 with Et al. 6 Sep 1935 19 May 1939 WTU-121585, WTU-V-056729 WTU-71255, WTU-V-056737 Dryopteridaceae Polystichum lonchitis (L.) Roth Dryopteridaceae CANADA, ONTARIO: Kemble, Owen Sound. Polystichum munitum (Kaulf.) C. Presl 44.568501°, -80.938921°; WGS 84, uncertainty: 1376 m., Source: Digital Map, U.S.A., CALIFORNIA, MONTEREY COUNTY: Georef'd by Ghislaine Ng Carmel Highlands above Yankee Pt. Rocky woods. Elev. 600 ft. 36.504129°, -121.931621°; WGS 84, C. C. Stewart 1740 uncertainty: 332 m., Source: GeoLocate, Georef'd by Ghislaine Ng 9 Jul 1900 In shade, on steep canyon slopes, in iduff. Closed-cone pine forest. Hgt. to 2 1/2 ft, WTU-146796, WTU-V-056732 common. E. K. Balls 23607 23 Sep 1958 WTU-197429, WTU-V-056743 Dryopteridaceae Dryopteridaceae Polystichum munitum (Kaulf.) C. Presl Polystichum polyblepharum (Roem. & Kunze) C. Presl U.S.A., CALIFORNIA, SAN MATEO COUNTY: JAPAN: 4 mi. west of Woodside,. 1 mi. east of Skyline Hondo: Chihayamura in Kawachi. Blvd. On Old LaHonda Rd. Santa Cruz mt. San TSN. Francisco Bay drainage area. Slope east. Elev. 1600 M. Togasi 894 37.429916°, -122.326896°; WGS 84, uncertainty: 8812 m., Source: GeoLocate, 10 Mar 1953 Georef'd by Ghislaine Ng Sandy loam. RedWood forest. WTU-157525, WTU-V-056759 Patrick Elvander 221 10 Jun 1970 WTU-266152, WTU-V-056744 Dryopteridaceae Polystichum platyphyllum (Willd.) C. Presl Dryopteridaceae COLOMBIA : Polystichum obtusum J.Sm. Boca Toma Forest, Finca El Recuerdo. Elev. 2500 ft. PHILIPPINES , RIZAL : 5.616389°, -73.653889°; WGS 84, Island Luzon. Rizal(Montalban). uncertainty: 3036 m., Source: GeoLocate, 14.726148°, 121.146253°; WGS 84, Georef'd by Ghislaine Ng uncertainty: 2591 m., Source: Digital Map, Georef'd by Ghislaine Ng Dora S. Bennett 19 In steep woods. Mar 1936 E. B. Copeland 97 WTU-55903, WTU-V-056762 Nov 1908 WTU-8264JWT, WTU-V-056751 Dryopteridaceae Polystichum dudleyi Maxon Dryopteridaceae U.S.A., CALIFORNIA, SANTA CLARA Polystichum ×potteri Barrington COUNTY: Foothills west of Los Gatos. CANADA, NOVA SCOTIA, INVERNESS: 37.230985°, -122.035105°; WGS 84, N side of Nase River, 3 km SW of Sammys uncertainty: 5281 m., Source: Digital Map, Barren. Georef'd by Ghislaine Ng Elev. 75 m. 46° 49' N, 60° 44' W; WGS 84, uncertainty: A. A. Heller 7226 1911 m., Source: GeoLocate, Georef'd by Ghislaine Ng 11 Jan 1904 Rare in 300 year-old hardwood forest, with Acer saccharum, Fagus grandifolius, Trientalis WTU-124882, WTU-V-056905 borealis, and both parent taxa. P. F. Zika 13971 Dryopteridaceae 29 Jul 1999 Polystichum dudleyi Maxon WTU-339900, WTU-V-056752 U.S.A., CALIFORNIA COUNTY: Santa Cruz Mts. Dryopteridaceae 37.122678°, -121.85123°; WGS 84, uncertainty: 1177 m., Source: Digital Map, Polystichum retroso-paleaceum (Kodama) Tagawa Georef'd by Ghislaine Ng JAPAN: unknown s.n. Honshu: Mt. Zaozan in Echigo. 38.152115°, 140.444768°; WGS 84, No collection date given uncertainty: 1000 m., Source: Digital Map, Georef'd by Ghislaine Ng WTU-11128JWT, WTU-V-056906 TNS. M. Togasi 1756 13 Jun 1958 WTU-198696, WTU-V-056757 Dryopteridaceae Dryopteridaceae Polystichum amabile (Blume) J.Sm. Polystichum horizontale C.Presl CHINA , KWONG TUNG : PHILIPPINES : Teng Woo Mt. Mt. Banahao, near San Pablo City, Laguna, Luzon Is. Levine 51 Elev. 1100 m. with Groff 14° 5' N, 121° 31' E; WGS 84, uncertainty: 3036 m., Source: Digital Map, Georef'd by 18 Nov 1916 Ghislaine Ng Terrestrial and at various heights on Cyathea WTU-213130, WTU-V-057074 trunks. Possibly starts life epiphytic. C. J. Marchant 78.267 with B.Hernaez. Apr 1978 Dryopteridaceae WTU-376017, WTU-V-056915 Polystichum acrostichoides (Michx.) Schott Dryopteridaceae U.S.A., MICHIGAN, LAPEER COUNTY: Deerfield Twp. S side of M-90, E of M-24; Polystichum imbricans (D.C. Eaton) D.H. Wagner floodplain of North Branch of Flint River. Sec. 4/90, T9N, R10E. ; WGS 84, uncertainty: U.S.A., CALIFORNIA, TRINITY COUNTY: 31667 m., Source: GeoLocate, Georef'd by 4.5 mi NE of Forest Glen along Post Creek or Ghislaine Ng 1.5 mi NE of intersection of highway 36 and Damp, low woods; moderate shade of Ostrya Post Creek Road. virginiana, Acer rubrum; w/ Botrychium Elev. 3000 ft. virginianum, Juglans cinerea, Acer 40.419302°, -123.264772°; WGS 84, saccharinum. Clumped fern w/ leathery lvs. uncertainty: 2950 m., Source: GeoLocate, Georef'd by Ghislaine Ng Ruth B. Alford 3981 Narrow canyon with small creek and serpentine slopes. Zone of Pinus sabianiana, Quercus 6 Jun 1976 kelloggii, and Quercus chrysolepis. Locally frequent in rock crevices. WTU-348269, WTU-V-057078 Melinda F. Denton 3960 23 Jun 1976 Dryopteridaceae WTU-268018, WTU-V-056917 Polystichum acrostichoides (Michx.) Schott U.S.A., MASSACHUSETTS, HAMPSHIRE COUNTY: Dryopteridaceae Amherst. 42.37352°, -72.520053°; WGS 84, Polystichum imbricans (D.C. Eaton) D.H. Wagner ssp. uncertainty: 1318 m., Source: Digital Map, curtum (Ewan) D.H. Wagner Georef'd by Ghislaine Ng Open woodland. U.S.A., CALIFORNIA, SAN BERNARDINO COUNTY: S. J. Ewer s.n. San Bernardino Mts., 8 mi. below Lake Arrowhead, on Lake Arrowhead Road. Aug 1940 34.132276°, -117.189208°; WGS 84, uncertainty: 13800 m., Source: GeoLocate, WTU-182393, WTU-V-057115 Georef'd by Ghislaine Ng Shade, cracks in granite rock. Life Zone: Arid transition. Height: 1 ft. Spread: 1 1/2 ft. Dryopteridaceae Carl B. Wolf 4397 Polystichum hillebrandii Carruth. 9 Nov 1932 U.S.A., HAWAII, MAUI COUNTY: Paliku, Haleakala. WTU-107108, WTU-V-056928 20.723363°, -156.142839°; WGS 84, uncertainty: 1566 m., Source: Digital Map, Georef'd by Ghislaine Ng Grassy, open forest. Otto Degener 17196 5 Aug 1939 WTU-152981, WTU-V-056913 Dryopteridaceae Dryopteridaceae Polystichum imbricans (D.C. Eaton) D.H. Wagner ssp. Polystichum kruckebergii W.H. Wagner imbricans U.S.A., UTAH, BOX ELDER COUNTY: U.S.A., CALIFORNIA, BUTTE COUNTY: Head of Middle Fork of Dunn Canyon, Raft About 100 yards southwest o the bridge across River Mts. Butte Creek, on the north side of Doe Mill Road, Elev. 9500 ft. about 3 miles (air) north of De Sabla Reservoir. 41.871708°, -113.3658°; WGS 84, Elev. 2166 ft. uncertainty: 2000 m., Source: Digital Map, 39° 54' 31.6" N, 121° 37' 3.8" W; T24N, R3E, Georef'd by Ghislaine Ng northeast 1/4 Section 27. ; WGS 84, Dry places in crevices of rocks in cliffs, in uncertainty: 90 m., Source: GeoLocate, contact of quartzite and serpentine. Infrequent. Georef'd by Ghislaine Ng On damp soil, in sun, cliffs. Mixed Evergreen Bassett Maguire 22205 Forest. Frond from one clump, fronds upright, with A.H. Holmgren some twisted. Common. 31 Jul 1943 Lowell Ahart 20,584 WTU-72435, WTU-V-056943 24 Oct 2015 WTU-143565, WTU-V-056940 Dryopteridaceae Polystichum lemmonii Underw. Dryopteridaceae U.S.A., CALIFORNIA, TRINITY COUNTY: Polystichum imbricans (D.C. Eaton) D.H. Wagner ssp. Ca. 200 feet above and to south of Mosquito imbricans Lake. Salmon-Trinity alps Wilderness Area, Shasta National Forest. U.S.A., CALIFORNIA, BUTTE COUNTY: Elev. 6600 ft. On the south side of Forbestown Reservoir, 41.019599°, -123.107929°; WGS 84, about 400 yards southeast of the Forbestown uncertainty: 1927 m., Source: GeoLocate, Reservoir Dam, about 4 miles (air) northeast of Georef'd by Ghislaine Ng Forbestown. Dry slope in forest of red fir, white fir, mountain Elev. 1853 ft. hemlock, western white pine, Jeffrey pine. 39° 33' 4" N, 121° 12' 30.3" W; T20N, R7E, Rhizomes deeply embedded under boulder in northeast 1/4 Section 32. ; WGS 84, granite scree. uncertainty: 969 m., Source: GeoLocate, Georef'd by Ghislaine Ng Helen K. Sharsmith 4830 On damp soil, in shade, serpentine rock outcrops. Mixed Evergreen Forest. Fronds from 27 Aug 1960 two or three clumps, fronds upright, twisted. Common, scattered. WTU-212473, WTU-V-056944 Lowell Ahart 20,581 23 Oct 2015 WTU-413568, WTU-V-056941 Dryopteridaceae Polystichum braunii (Spenner) Fée Dryopteridaceae U.S.A., VERMONT, CALEDONIA COUNTY: Polystichum keysserianum Rosenst SW of Mt. Hor, Sutton. 44.710639°, -72.062767°; WGS 84, PAPUA NEW GUINEA , MOROBE : uncertainty: 1807 m., Source: Digital Map, Ekuti Divide, nr. Bulolo. Georef'd by Ghislaine Ng 7° 10' S, 146° 40' E; WGS 84, uncertainty: Rocky ravine. 3036 m., Source: GeoLocate, Georef'd by Ghislaine Ng Peter F. Zika 3392 C. J. Marchant 78.340 3 Jun 1981 Feb 1978 WTU-378190, WTU-V-056966 WTU-375991, WTU-V-056942 Dryopteridaceae Dryopteridaceae Polystichum braunii (Spenner) Fée Polystichum lonchitis (L.) Roth CANADA, NOVA SCOTIA, INVERNESS: CANADA, ONTARIO: N side of Anse River, 3 km SW of Sammys Owen Sound, comte de Bruce: sur les rochers Barren. calcaires. Elev. 75 m. 46° 49' N, 60° 44' W; WGS 84, uncertainty: Frere Marie-Victorin 45925 1911 m., Source: GeoLocate, Georef'd by with F. Rolland-Germain, Emile Jacques. Ghislaine Ng In 300 year-old hardwood forest, with Acer 9 Jun 1932 saccharum, Fagus grandifolius, Trientalis borealis. Uncommon. WTU-60273, WTU-V-056716 P. F. Zika 13972 29 Jul 1999 WTU-339899, WTU-V-056970 Dryopteridaceae Polystichum lonchitis (L.) Roth Dryopteridaceae CANADA, QUEBEC: Polystichum braunii (Spenner) Fée Mont St-Pierre, Cte Gaspe-Nord, P. Que.. Phenology: Spores. RUSSIA : Kuril Archipelgo: Iturup, about 4 km east of Père Louis-Marie s.n. Kitovyi by road; environs of Podoshevka River; with FF. Fabius & Adonis M. Raymond & J. near abandoned fish hatchery. Paquin. 45° 15' 54" N, 147° 55' 35" E Collected along stream on lichen covered trees 12 Aug 1934 (Salix sp.); moss covered rocks; under Sasa and other herbaceous vegetation (Petasites). WTU-239994, WTU-V-056717 Fronds very scaly; Fide Ohwi, Flora of Japan. Birgit Semsrott BS 1024 29 Jul 1997 Dryopteridaceae WTU-348754, WTU-V-056972 Polystichum lonchitis (L.) Roth U.S.A., MAINE, KENNEBEC COUNTY: Augusta. Dryopteridaceae 44.310624°, -69.77949°; WGS 84, uncertainty: 1000 m., Source: GeoLocate, Polystichum braunii (Spenner) Fée Georef'd by Ghislaine Ng Phenology: Spores. U.S.A., VERMONT, ORLEANS COUNTY: W of Jobs Pond, town of Westmore. E. C. Smith s.n. 44.772622°, -72.038181°; WGS 84, uncertainty: 1772 m., Source: Digital Map, 1 Jun 1886 Georef'd by Ghislaine Ng Among boulders, base of cliffs. WTU-124786, WTU-V-056719 Peter F. Zika 3146 18 May 1981 Dryopteridaceae WTU-369029, WTU-V-056982 Polystichum lonchitis (L.) Roth U.S.A., CALIFORNIA, TRINITY COUNTY: Above Whites Creek Lake. Devils Canyon Dryopteridaceae Mountains at head of White Creek. Elev. 6500 ft. Polystichum lonchitis (L.) Roth 40.965758°, -123.224197°; WGS 84, uncertainty: 1000 m., Source: Digital Map, CANADA, ONTARIO, BRUCE.: Georef'd by Ghislaine Ng 1.5 miles north of Kemble, Bruce County. Crevices of rock in cirque. Phenology: Spores. 44° 44' N, 80° 56' W; Source: Specimen Label Joseph P. Tracy 14711 Common in woods on very shallow rich soil over limestone on top of escarpment. Phenology: 8 Aug 1935 Spores. WTU-207301, WTU-V-056725 W. J. Cody 12390 with L.C. Sherk 6 Sep 1962 WTU-356518, WTU-V-056713 Dryopteridaceae Dryopteridaceae Polystichum lonchitis (L.) Roth Polystichum munitum (Kaulf.) C. Presl RUSSIA, KAMCHATSKIY: U.S.A., CALIFORNIA, SONOMA COUNTY: Kamchatskiy Region, Yelizovskiy district, head 6/10 miles inland from stewarts point. water of river Avacha, research station 38.727923°, -123.46376°; WGS 84, "Tymonovskiye sources". uncertainty: 1000 m., Source: Digital Map, Betula ermanii timberline, common. Phenology: Georef'd by Ghislaine Ng Spores. Humid transition. Redwood forest floor, shade. Plant type: Perennial. Height: 3 feet. abundant. S. Kharkevich 36 Phenology: Spores. with T.Buch. Carl B. Wolf 5798 26 Sep 1975 29 Jun 1934 WTU-343257, WTU-V-056726 WTU-109336, WTU-V-056747

Dryopteridaceae Dryopteridaceae Polystichum lonchitis (L.) Roth Polystichum munitum (Kaulf.) C. Presl SWEDEN: Lule lappmark: Jokkmokk parish, Mt. Rittok. U.S.A., CALIFORNIA, SAN DIEGO COUNTY: North peak Cuyamaca mountains. Tor Ake Tengwall s.n. Elev. 4000 ft. 32.952947°, -116.568032°; WGS 84, 15 Jul 1911 uncertainty: 2000 m., Source: Digital Map, Georef'd by Ghislaine Ng WTU-57190, WTU-V-056728 Semi-shade. Roots deeply buried in leaf mould. Phenology: Spores. F. M. Cota 89 1930 WTU-8479JWT, WTU-V-056749 Dryopteridaceae Polystichum munitum (Kaulf.) C. Presl Dryopteridaceae U.S.A., CALIFORNIA, GLENN COUNTY: Coast Range near Bennett Spring, on the Polystichum mohrioides (Bory) C. Presl Newville Covelo road in the yellow pine belt. Elev. 3000 ft. ARGENTINA, SANTA CRUZ: On a timbered north slope, growing in a damp Depto. Deseado, 3 kilometers west of Puerto ravine, the common under shrubs Ceanothus Deseado, road to Comodoro Rivadavia. and Toxicodendron. Phenology: Spores. Feldspathic granite outcrop, rock crevices. Fern to 0.25 meters. Associated with Nos. A. A. Heller 11546 23863-868. Phenology: Spores. 8 Jul 1914 W. J. Eyerdam 23862 with A.A. Beetle, E. Grondona WTU-81390, WTU-V-056739 29 Dec 1938 WTU-26923JWT, WTU-V-056750 Dryopteridaceae Polystichum munitum (Kaulf.) C. Presl Dryopteridaceae U.S.A., CALIFORNIA, SAN FRANCISCO COUNTY: Polystichum platyphyllum (Willd.) C. Presl Southeast slope of the island. Red Rock, San Francisco Bay. BOLIVIA: 37.928642°, -122.430233°; WGS 84, Lambramani. uncertainty: 146 m., Source: Digital Map, Very common in wet soil. Poisoned. Phenology: Georef'd by Ghislaine Ng Spores. On steep Franciscan chert bluffs. Forming very dense stools. Leaves erect, stiff. Phenology: M. Cardenas 1236 Spores. Nov 1933 Edward Lee 1983 WTU-23342JWT, WTU-V-056760 19 Jun 1936 WTU-26989JWT, WTU-V-056745 Dryopteridaceae Dryopteridaceae Polystichum scopulinum (D.C. Eaton) Maxon Polystichum lemmonii Underw. U.S.A., CALIFORNIA, SISKIYOU COUNTY: U.S.A., CALIFORNIA, TRINITY COUNTY: On the Red Ridge to the west of and above Just below outlet of Deer Lake, at contact Panther Creek Meadows. Hudsonian Zone. between peridotite and granodiorite. Elev. 9000 ft. Salmon-Trinity Primitive Area. Trinity Alps. 41.759444°, -122.248052°; WGS 84, 40.942708°, -122.89279°; WGS 84, uncertainty: 2000 m., Source: Digital Map, uncertainty: 250 m., Source: Digital Map, Georef'd by Ghislaine Ng Georef'd by Ghislaine Ng Under shaded portions of lava boulders and Open rocky slopes among scattered mountain strata. Poisoned. Phenology: Spores. hemlock and white pine. Phenology: Spores. unknown 13910 A. R. Kruckeberg 3765 24 Aug 1939 11 Aug 1954 WTU-23864JWT, WTU-V-056763 WTU-189772, WTU-V-056948

Dryopteridaceae Dryopteridaceae Polystichum setiferum (Forssk.) Moore ex Woynar var. Polystichum sp. subtripinnatum Moore RUSSIA: FRANCE: Kuril Archipelaho: Simushir, inland coastal Cher - Paro du Chateau de la Beuvriere et rives margin of Milna Cove in Kitoboynaua Bay; near de l'Arnon en amont du pont de bois, pres du unnamed waterfall. moulin du meme nom, commune de Elev. 15 m. Saint-Hilaire-de-Court. 46° 51' 14" N, 151° 47' 50" E Phenology: Spores. Coastal upper beach zone; surrounding wooded area and rock wall with shrubby Alnus and J. Prudhomme 784 Petasites as the dominant vegetation. Growing under Alnus, among boulders. 30 Aug 1949 Sarah Gage SG 1287 WTU-136647, WTU-V-056771 10 Aug 1995 WTU-337769, WTU-V-056676 Dryopteridaceae Polystichum japonicum Diels Dryopteridaceae JAPAN, KANAGAWA: Hondo: Taura in Sagami. Polystichum tridens (T.Moore ex Hook.) Fée Phenology: Spores. JAMAICA: M. Togasi 873 Arntully. 18.016667°, -76.6°; WGS 84, uncertainty: Dec 1951 3036 m., Source: GeoLocate, Georef'd by Ghislaine Ng WTU-160545, WTU-V-056916 Orcutt 2702 1927 WTU-8513JWT, WTU-V-056680 Dryopteridaceae Polystichum munitum (Kaulf.) C. Presl U.S.A., CALIFORNIA, GLENN COUNTY: Slap Jack camp west of Alder Springs. Elev. 5000 ft. 39.650132°, -122.738072°; WGS 84, uncertainty: 2000 m., Source: Digital Map, Georef'd by Ghislaine Ng In moist places in the yellow pine belt. Sugar pine is also plentiful. Ceanothus cordulatus and Grossularia Lobbii are common shrubs. Phenology: Spores. A. A. Heller 12805 5 Jul 1917 WTU-87483, WTU-V-056924 Dryopteridaceae Dryopteridaceae Polystichum vestitum (G.Forst.) C.Presl Polystichum braunii (Spenner) Fée CHILE : RUSSIA : Juan Fernandez Islands, Isla Mas Atierra. Along Kuril Archipelago: Simushir, inland coastal road to Mr. Weber's house. margin of Milna Cove in Kitoboynaya Bay; near Elev. 150 m. unnamed waterfall. -33.800399°, -79.766597°; WGS 84, Elev. 15 m. uncertainty: 101382 m., Source: GeoLocate, 46° 51' 14" N, 151° 47' 50" E Georef'd by Ghislaine Ng Coastal upper beach zone; surrounding wooded Among bushes. Fern 0.4m. Directior, T. H. area and rock wall with shrubby Alnus and Goodspeed. Petasites as the dominant vegetation. Growing under Alnus, among boulders. R. Wagenknecht 18514 Sarah Gage SG 1286 21 Mar 1940 10 Aug 1995 WTU-290025, WTU-V-056681 WTU-337658, WTU-V-056979

Dryopteridaceae Dryopteridaceae Polystichum distans Fourn. Polystichum braunii (Spenner) Fée MEXICO, MéXICO: Temascaltepex, Mex. Cumbre de Tejupilco. RUSSIA: 18.906122°, -100.149459°; WGS 84, Kuril Archipelago: Ekarma, northwest corner of uncertainty: 1277 m., Source: Digital Map, island; unnamed bay just east of Cape Georef'd by Ghislaine Ng Shpilevoy. 48° 57' 57" N, 153° 55' 20" E Geo. B. Hinton 7134 Understory with Parasenecio auriculata with Et al. dominant. Alnus grove with trees 12-15 feet tall, with trunks up to 5 inch diameter. Growing 20 Dec 1934 under Parasenecio auricalata. WTU-123500, WTU-V-056908 Sarah Gage SG 2234 10 Aug 1996 Dryopteridaceae WTU-342886, WTU-V-056980 Polystichum braunii (Spenner) Fée RUSSIA: Dryopteridaceae Kuril Archipelago: Shiashkotan, inland from Zakatnaya bay on east side of Cape Grotovii, Polystichum braunii (Spenner) Fée top of plateau. Elev. 200 ft. RUSSIA : 48° 46' 51" N, 154° 2' 19" E; WGS 84, Kuril Archipelago: Iturup, konservnaya Bay. uncertainty: 6573 m., Source: GeoLocate, 45° 20' 7" N, 147° 59' 32" E; WGS 84, Georef'd by Ghislaine Ng uncertainty: 56728 m., Source: GeoLocate, Meadow at top of plateau, dominated by Georef'd by Ghislaine Ng Calamagrostis, other grasses and forbs. Coastal forest with much evidence of human impact; moist area along stream dominated by Sarah Gage SG 2267 Cacalia, Filipendula, Sasa. Overstory includes Salix, Alnus, Betula. Growing on stream bank. 11 Aug 1996 Sarah Gage SG 2483 WTU-342897, WTU-V-056978 18 Aug 1996 WTU-342913, WTU-V-056981 Dryopteridaceae Dryopteridaceae Polystichum braunii (Spenner) Fée Polystichum braunii (Spenner) Fée RUSSIA : RUSSIA: Kuril Archipelago: Urup, Ukromnaya River, small Kuril Archipelago: Ushishir Group, Yankicha tributary ca. 3 km inland from Ukromnaya Bay. Island; inland environs of Kraternaya Bay; along 45° 35' 31" 149° 32' 12"; WGS 84, south side of northern slope of bay, just below uncertainty: 6905 m., Source: GeoLocate, ridge. Georef'd by Ghislaine Ng 47° 31' 10" N, 152° 49' 8" E; WGS 84, Riverbanks and sandbars of river, huge basalt uncertainty: 487 m., Source: Digital Map, block at edge of river and muddy riverbanks. Georef'd by Ghislaine Ng Leaves deep green above and almost leathery. Ridge tundra vegetation turns into lush tall Growing in clump at base of basalt rock race. herbaceous meadow. Sarah Gage SG 2560 Birgit Semsrott BS 1111 20 Aug 1996 1 Aug 1997 WTU-342687, WTU-V-056983 WTU-348676, WTU-V-056989

Dryopteridaceae Dryopteridaceae Polystichum braunii (Spenner) Fée Polystichum braunii (Spenner) Fée RUSSIA: RUSSIA: Kuril Archipelago: Alaid, inland from Alaidskaya Kuril Archipelago: Makanrushi, inland from Bay, near abandoned settlement of Atlasova. Zakat Bay. Elev. 325 ft. 49° 44' 16" N, 154° 24' 52" E; WGS 84, 50° 50' 11" N, 155° 39' 31" E; WGS 84, uncertainty: 796 m., Source: Digital Map, uncertainty: 3036 m., Source: GeoLocate, Georef'd by Ghislaine Ng Georef'd by Ghislaine Ng Moist meadow with patches of lichen covered North facing. Growing under Alnus on an Alnus, rocks. Fronds glossy, dark green above. Elve: Filipendula, Artemisia dominated slope. Upper near sea level. side of leaves glossy green. Sarah Gage SG 4581 Sarah Gage SG 4380 18 Aug 1997 12 Aug 1997 WTU-348699, WTU-V-056990 WTU-348667, WTU-V-056986

Dryopteridaceae Dryopteridaceae Polystichum sp. Polystichum braunii (Spenner) Fée RUSSIA: RUSSIA: Kuril Archipelago: Urup, inland coastal margin of Kuril Archipelago: Iturup, eastern side of Chirip Natalie Bay; environs of Vesolaya River. Peninsula; inland coastal margin of Elev. 15 m. Konservnaya Bay. 46° 5' 7" N, 150° 8' 27" E 45° 19' 53" N, 147° 59' 41" E; WGS 84, Along steep bank of river at edge of uncertainty: 39646 m., Source: GeoLocate, Betula-Alnus-Sasa forest. Growing on soil. Georef'd by Ghislaine Ng With sori; under deciduous tree on vertical Sarah Gage SG 1194 root/soil wall. Sasa dominant under deciduous trees (Acer, Alnus, Quercus). 6 Aug 1995 Birgit Semsrott BS 1067 WTU-337657, WTU-V-056674 30 Jul 1997 WTU-348645, WTU-V-056987 Dryopteridaceae Polystichum tripteron (Kunze) C.Presl JAPAN: Hondo: Jummuji in Sagami. J. Ohwi 282 Apr 1951 WTU-148278, WTU-V-056683 Dryopteridaceae Dryopteridaceae Polystichum tripteron (Kunze) C.Presl Polystichum munitum (Kaulf.) C. Presl JAPAN: U.S.A., CALIFORNIA, HUMBOLDT COUNTY: Hondo: Minomo in Settsu. South Fork Mountain; openings in fir forest near McKay Camp. M. Togasi 484 Elev. 5500 ft. 40.525093°, -123.549285°; WGS 84, 11 Jun 1952 uncertainty: 3397 m., Source: Digital Map, Georef'd by Ghislaine Ng WTU-180988, WTU-V-056685 rocky point in woods. Phenology: Spores. Joseph P. Tracy 9089 July 16 to 19, 1930 WTU-56192, WTU-V-056927 Dryopteridaceae Cyrtomium falcatum (L. f.) C. Presl Dryopteridaceae CHINA, KWONG TUNG: Polystichum imbricans (D.C. Eaton) D.H. Wagner ssp. Loh Fau Shan (Loh Fau Mountain). curtum (Ewan) D.H. Wagner C. O. Levine 509 U.S.A., CALIFORNIA, SAN BERNARDINO COUNTY: Oct. 27-30 1916. San Bernardino mountains. Elev. 3000 ft. WTU-213131, WTU-V-056912 34.060458°, -116.868175°; WGS 84, uncertainty: 1000 m., Source: Digital Map, Georef'd by Ghislaine Ng Phenology: Spores. S. B. Parish 11390 14 Jun 1917 Dryopteridaceae WTU-124789, WTU-V-056930 Polystichum japonicum Diels JAPAN, KANAGAWA: Dryopteridaceae Manazuru. Phenology: Spores. Polystichum imbricans (D.C. Eaton) D.H. Wagner Shigetaka Suzuki s.n. U.S.A., CALIFORNIA, HUMBOLDT COUNTY: Trinity River Valley, near the south fork. 16 Oct 1951 Elev. 600 ft. 41.184849°, -123.708672°; WGS 84, WTU-149668, WTU-V-056914 uncertainty: 1000 m., Source: GeoLocate, Georef'd by Ghislaine Ng Rocky hillside in woods. Common. Phenology: Spores. Joseph P. Tracy 7270 Dryopteridaceae 20 Sep 1925 Polystichum imbricans (D.C. Eaton) D.H. Wagner WTU-207302, WTU-V-056932 U.S.A., CALIFORNIA, DEL NORTE COUNTY: Siskiyou mountains, low divide. Elev. 1800 ft. Dryopteridaceae 41.834552°, -123.667807°; WGS 84, uncertainty: 2000 m., Source: Digital Map, Polystichum lemmonii Underw. Georef'd by Ghislaine Ng Arid transition:- scrub tan oak. sun, dry open U.S.A., CALIFORNIA, SISKIYOU COUNTY: flats, red clay over serpentine. Plant type: East fork house creek. Siskiyou mountains. Perennila. Height: 1'-2'. Abundant. Phenology: Elev. 6500 ft. Spores. About rocks, bare north slope. Phenology: Spores. Carl B. Wolf 9141 Louis C. Wheeler 3114 10 Aug 1937 13 Aug 1934 WTU-116639, WTU-V-056925 WTU-2934JWT, WTU-V-056945 Dryopteridaceae Dryopteridaceae Polystichum lemmonii Underw. Polystichum braunii (Spenner) Fée U.S.A., CALIFORNIA, SISKIYOU COUNTY: CANADA, QUEBEC: Mt. Eddy. Riviere A Claude: pres de l'embouchure. Elev. 7700 ft. Phenology: Spores. 41.319871°, -122.479184°; WGS 84, uncertainty: 1000 m., Source: GeoLocate, Père Louis-Marie 34328 Georef'd by Ghislaine Ng with FF. Fabius et Adonis, M. Raymond et J. Phenology: Spores. Paquin. A. A. Heller 11739 3-9 aout 1934 28 Aug 1914 WTU-146JWT, WTU-V-056975 WTU-84696, WTU-V-056950

Dryopteridaceae Dryopteridaceae Polystichum braunii (Spenner) Fée Polystichum lobatum (Huds.) Pr. U.S.A., VERMONT, WINDHAM COUNTY: MONTENEGRO: Haystack mountain, Wilmington. Crna Gora; 10 kilometers northeast of Kolasin. Shady woods near brook. Only plant seen. Lake of Biogradsko Jezero. Phenology: Spores. Elev. 1200 m. In Fagus-forest near the lake. Phenology: F. G. F. 2573 Spores. 25 Sep 1908 unknown 68-1927 with Stud. biol. Rheno-Trai. in itinere. WTU-213132, WTU-V-056984 16 Jul 1968 WTU-313156, WTU-V-056951 Dryopteridaceae Dryopteridaceae Polystichum braunii (Spenner) Fée Polystichum lobatum (Huds.) Pr. CANADA, QUEBEC: POLAND, SUBCARPATHIA, DEBICA: Stoneham, comte de Quebec. Southern Poland Poisoned. Phenology: Spores. In the complex Fagetum. Phenology: Spores. Fr. Anselme s.n. E.Pancer s.n. 18 Jul 1931 27 Apr 1947 WTU-11866JWT, WTU-V-056992 WTU-279421, WTU-V-056958

Dryopteridaceae Dryopteridaceae Polystichum braunii (Spenner) Fée Polystichum braunii (Spenner) Fée CANADA, QUEBEC: U.S.A., VERMONT, RUTLAND COUNTY: Ruisseau Castor, Cte Gaspe-Nord, P. Que. Sur Region of Mt. Killington. Sherburne. les falaises. Phenology: Spores. Phenology: Spores. D. Lewis Dutton s.n. Adrien Gosselin s.n. 11 Sep 1910 23 Jul 1936 WTU-44437, WTU-V-056974 WTU-243708, WTU-V-056997 Dryopteridaceae Dryopteridaceae Polystichum californicum (D.C. Eaton) Diels Polystichum braunii (Spenner) Fée U.S.A., CALIFORNIA, SANTA CLARA CANADA, QUEBEC: COUNTY: Riviere-a-Claude, a 1.5 mille de la mer. Comte Foothills west of Los Gatos. Gaspe-Nord. 37.234265°, -122.024118°; WGS 84, 49° 12' N, 65° 54' W; Source: Specimen uncertainty: 1000 m., Source: Digital Map, Label Georef'd by Ghislaine Ng Populetum boreale a Pteretis pensylvanica. Phenology: Spores. Phenology: Spores. A. A. Heller 7226 V. Gerardin 4775 with M. Grandtner, G. Lemieux, L. Rousseau. 11 Jan 1904 4 Jul 1963 WTU-156JWT, WTU-V-056999 WTU-229365, WTU-V-056998 Dryopteridaceae Dryopteridaceae Polystichum lobatum (Huds.) Pr. Polystichum aculeatum (L.) Roth. FRANCE, ORLéANS: Cher - Avec le type, le long de la route de U.S.A., NEW HAMPSHIRE, CARROLL Mennetou-Salon aux Crots-de-Veaux. COUNTY: Phenology: Spores. Dolloffs Cliff, Jackson. Rich, shady, moist, rocky slope. Fairly J. Prudhomme 782 abundant. Phenology: Spores. with Vidit J. Calle F. G. F. 2482 22 Sep 1949 27 Sep 1907 WTU-136646, WTU-V-056959 WTU-210280, WTU-V-054604

Dryopteridaceae Dryopteridaceae Polystichum lobatum (Huds.) Pr. Polystichum aculeatum (L.) Roth. var. aristatum (Christ) FRANCE, DIJON: Askerov Haute-Saone - Colonies eparses dans les Gorges du Rahin en amont de la Vieille Hutte, FRANCE: sur les pentes au Ballon d'Alsace. Col de Bussang (France, chaine des Vosges). Phenology: Spores. Elev. 700 m. Pentes boisees. Phenology: Spores. E.Walter 1540 with J. Prudhomme, Vidit J. Calle. Bernard Retz 12241 30 Aug 1951 13 Jun 1936 WTU-166911, WTU-V-056960 WTU-168619, WTU-V-054607

Dryopteridaceae Polystichum braunii (Spenner) Fée Dryopteridaceae CANADA, QUEBEC: Polystichum acrostichoides (Michx.) Schott Riviere-a-Claude, a 1.5 mille de la mer. Comte Gaspe-Nord. CANADA, QUEBEC, 49° 12' N, 65° 54' W; Source: Specimen VAUDREUIL-SOULANGES: Label Erablière laurentienne. St-Polycarpe. Populetum boreale a Pteretis pensylvanica. 45.302663°, -74.304079°; WGS 84, Phenology: Spores. uncertainty: 898 m., Source: Digital Map, Georef'd by Ghislaine Ng V. Gerardin 4775 with M. Grandtner, G. Lemieux, L. Rousseau. Gilles Lemieux 3196 4 Jul 1963 7 Jul 1962 WTU-219566, WTU-V-056996 WTU-220147, WTU-V-057089 Dryopteridaceae Dryopteridaceae Polystichum acrostichoides (Michx.) Schott Polystichum imbricans (D.C. Eaton) D.H. Wagner U.S.A., VERMONT, RUTLAND COUNTY: U.S.A., CALIFORNIA, HUMBOLDT COUNTY: Middletown Springs. Trinity river valley, near the south fork. 43.485231°, -73.118331°; WGS 84, Elev. 600 ft. uncertainty: 400 m., Source: Digital Map, 41.166243°, -123.696141°; WGS 84, Georef'd by Ghislaine Ng uncertainty: 742 m., Source: Digital Map, Christmans Fern. Common. Georef'd by Ghislaine Ng Accompanying P. munitum but less common. Dana S. Carpenter 199 Rocky hillside in woods. Phenology: Spores. Jul Joseph P. Tracy 7271 WTU-26933JWT, WTU-V-057100 20 Sep 1925 WTU-207303, WTU-V-056923 Dryopteridaceae Polystichum ? Dryopteridaceae U.S.A., ARKANSAS COUNTY: Polystichum lepidocaulon (Hook.) J.Sm. [no locality given on label]. 34.858171°, -92.423095°; WGS 84, JAPAN, CHIBA: uncertainty: 297729 m., Source: Digital Map, Mt. Kiyosumi. Georef'd by Joseph Stormer Phenology: Spores. Elva Lawton s.n. Shigetaka Suzuki s.n. Sep 1927 4 Nov 1951 WTU-325505, WTU-V-056672 WTU-149100, WTU-V-056953

Dryopteridaceae Polystichum Dryopteridaceae U.S.A., WEST VIRGINIA COUNTY: Polystichum acrostichoides (Michx.) Schott Cass. 38.396785°, -79.914783°; WGS 84, U.S.A., NEW YORK, DELAWARE COUNTY: uncertainty: 450 m., Source: GeoLocate, [locality and coordinates withheld; status in New Georef'd by Ghislaine Ng York: EV (Exploitably Vulnerable; USDA Plants)] F.W. Gray s.n. D. Leroy Topping 179 Nov 1927 24 Jun 1906 WTU-316710, WTU-V-056675 WTU-10188JWT, WTU-V-057083

Dryopteridaceae Polystichum lonchitis (L.) Roth Dryopteridaceae CANADA, ONTARIO, BRUCE: Polystichum dudleyi Maxon 1 1/2 miles north of Kemble. 44° 44' N, 80° 56' W; WGS 84, uncertainty: U.S.A., CALIFORNIA, SAN LUIS OBISPO 3219 m., Source: Digital Map, Georef'd by COUNTY: Ghislaine Ng Coast just south of Monterey Co. Common in woods on very shallow rich soil over 35.757353°, -121.307751°; WGS 84, limestone on top of escarpment. uncertainty: 4000 m., Source: Digital Map, Georef'd by John Haskins W. J. Cody 12390 Line, on moist rocky canyon-wall. Phenology: with L.C.Sherk. Spores. 6 Sep 1962 R. F. Hoover 6672 WTU-396631, WTU-V-056714 16 Mar 1947 WTU-238113, WTU-V-056904 Dryopteridaceae Dryopteridaceae Polystichum dudleyi Maxon Polystichum tripteron (Kunze) C.Presl U.S.A., CALIFORNIA, SANTA CLARA JAPAN: COUNTY: Sendai. Home Canon, Saratoga. Elev. 900 ft. A. Yasuda s.n. Poisoned. Phenology: Spores. 4 Jun 1912 R. L. Pendleton s.n. WTU-124787, WTU-V-056682 7 May 1907 WTU-13578JWT, WTU-V-056907

Dryopteridaceae Dryopteridaceae Polystichum echinatum (J.F. Gmel.) C. Chr. Polystichum tripteron (Kunze) C.Presl CUBA, MATANZAS: JAPAN, YAMANASHI: Bellavista. Mt. Kuradake Dry rocky bank. Phenology: Spores. Shigetaka Suzuki s.n. W. R. Maxon 4459 26 May 1952 4 May 1907 WTU-184507, WTU-V-056684 WTU-2684JWT, WTU-V-056910

Dryopteridaceae Dryopteridaceae Polystichum falcatum (L. f.) Diels var. genuinum (Makino) Polystichum lonchitis (L.) Roth Matsum NORWAY: JAPAN, SHOBUTA: [Indecipherable] Rikugen, Japan. Phenology: Spores. Origin: Native. Phenology: Spores. T. Lillefosse s.n. A. Yasuda s.n. 28 Jun 1933 17 Jun 1917 WTU-124807, WTU-V-056720 WTU-213133, WTU-V-056911

Dryopteridaceae Dryopteridaceae Polystichum lonchitis (L.) Roth Polystichum lobatum (Huds.) Pr. CANADA, ONTARIO, GREY : CZECH REPUBLIC: Woodford School Site. Moravia Centralis: in declivibus silvaticis ad p. Elev. 913 ft. Smilov prope urbem Olomouc. Rock crevices. Exchanged with J.W. Thompson. Phenology: Spores. var. marginale - sori on edge. Ex. Rare, one plant only. J. Otruba 111 U.R.W. Coll s.n. 5 Sep 1926 Sep 1924 Herbarium: WTU, WTU-V-056954 Herbarium: WTU, WTU-V-056721 Dryopteridaceae Dryopteridaceae Polystichum lonchitis (L.) Roth Polystichum muricatum (L.) Fée CANADA, ONTARIO, GREY: COLUMBIA: Woodford school (orchard) Dept. Cundinamarca; Macizo de Bogotá; Elev. 923 ft. Quebrodo de Chico. Rock crevice in sun. Exchanged with J.W. Elev. 2800-3000 m. Thompson. Rare. J. Cuatrecasas 5489 U.R.W. Coll s.n. 14 Jun 1939 Sep 1924 WTU-23375JWT, WTU-V-056738 WTU-11459JWT, WTU-V-056727

Dryopteridaceae Dryopteridaceae Polystichum lonchitis (L.) Roth Polystichum munitum (Kaulf.) C. Presl CANADA, ONTARIO, GREY: U.S.A., CALIFORNIA, SAN MATEO COUNTY: Woodford [indecipherable text] Pescadero Creek Cañon 6 mi above Elev. 923 ft. Pescadero. In rock crevices. Exchanged with J.W. 37.337005°, -122.3667°; WGS 84, Thompson. Largest found here. uncertainty: 1000 m., Source: GeoLocate, Georef'd by Ghislaine Ng U.R.W. Coll s.n. Roxana S. Ferris 2111 1931 20 Mar 1921 WTU-11458JWT, WTU-V-056730 WTU-71593, WTU-V-056741

Dryopteridaceae Dryopteridaceae Polystichum lonchitis (L.) Roth Polystichum richardii (Hook.) J.Sm. U.S.A., UTAH, UTAH COUNTY: NEW ZEALAND: Mt. Timpanogos. Palmerston North. A. O. Garrett 3441 V. D. Zotav s.n. 13 Aug 1925 [no collection date given] WTU-146797, WTU-V-056731 WTU-123416, WTU-V-056753

Dryopteridaceae Dryopteridaceae Polystichum munitum (Kaulf.) C. Presl Polystichum richardii (Hook.) J.Sm. U.S.A., CALIFORNIA, LOS ANGELES NEW ZEALAND: COUNTY: Lake Rotoiti. Big Tujunga Canyon. San Gabriel Mts. Elev. 4500 P. H. Buddle s.n. 34.27278°, -118.3148°; WGS 84, uncertainty: 2000 m., Source: GeoLocate, Georef'd by Jan 1941 Ghislaine Ng Open hot ridge, in granite fissures. WTU-124563, WTU-V-056754 J. Ewan 4751 5 Jul 1931 WTU-8488JWT-A, WTU-V-056736 Dryopteridaceae Dryopteridaceae Polystichum richardii (Hook.) J.Sm. Polystichum setiferum (Forssk.) Moore ex Woynar NEW ZEALAND: JERSEY : Lake Rotoiti. Ile de Jersey. Clairiere du Collège N.D. de Bonsecours, Highlands. P. H. Buddle s.n. Haies, chemins couverts. Jan 1941 Louis-Arsène 1541 WTU-124562, WTU-V-056755 10 Jul 1951 WTU-166912, WTU-V-056773

Dryopteridaceae Dryopteridaceae Polystichum pseudomakinoi Tagawa Polystichum seemannii (Hook.) J.Sm. JAPAN, TOKYO: COLOMBIA: Mt. Takao Santa Marta. Shigetaka Suzuki s.n. H.H. Smith s.n. 5 Oct 1952 Feb 1900 WTU-161128, WTU-V-056758 WTU-36624, WTU-V-056774

Dryopteridaceae Dryopteridaceae Polystichum semicordatum (Sw.) T.Moore Polystichum vestitum (G.Forst.) C.Presl COLOMBIA: NEW ZEALAND: Santa Marta. Rimutaka Range. H. H. Smith s.n. Doris W. Filmer s.n. Jul 1899 [no collection date given] WTU-37576, WTU-V-056766 WTU-123418, WTU-V-056775

Dryopteridaceae Dryopteridaceae Polystichum setiferum (Forssk.) Moore ex Woynar Polystichum varium (L.) C.Presl GERMANY: JAPAN: Mittel-Mosel: Senhals bei Cochem. Berg-wald Sendai. an der ehemaligen Brauerei. Elev. 120 m. A. Yasuda s.n. Laubwald. 3 Nov 1916 H. Andres 302 WTU-124788, WTU-V-056776 [no collection date given] WTU-135541, WTU-V-056772 Dryopteridaceae Dryopteridaceae Polystichum lemmonii Underw. Polystichum craspedosorum (Maxim.) Diels U.S.A., CALIFORNIA, SISKIYOU COUNTY: JAPAN, TOCHIGI: North Side of White Mt. Mt. Izuru-yama. Tochigi, Honshu. Elev. 5800 ft. 41.970461°, -123.059043°; WGS 84, Shigetaka Suzuki s.n. uncertainty: 2000 m., Source: Digital Map, Georef'd by Ghislaine Ng 19 May 1947 Dry slope. WTU-187846, WTU-V-056964 L. C. Wheeler 3137 14 Aug 1934 WTU-56188, WTU-V-056952

Dryopteridaceae Dryopteridaceae Polystichum craspedosorum (Maxim.) Diels Polystichum lobatum (Huds.) Pr. JAPAN: NORWAY, ASKER: Sendai. Skaugum [illegible] A. Yasuda s.n. Fr. Werenskiold s.n. 9 Sep 1917 12 Oct 1895 WTU-212217, WTU-V-056965 WTU-124790, WTU-V-056956

Dryopteridaceae Dryopteridaceae Polystichum braunii (Spenner) Fée Polystichum lobatum (Huds.) Pr. USSR: INDIA: Rokitno Wielkie in districtu Leopoliensi-URSS Pulney Hills Locis humidis, umbrosis, declivibus in fageto, Elev. 6200 ft. prope saltum "Mutynka." Parce. A. Sauliere 597 J. Madalski s.n. with S. Radwanski 3 Mar 1914 25 Aug 1939 Herbarium: WTU, WTU-V-056957 WTU-253305, WTU-V-056976

Dryopteridaceae Dryopteridaceae Polystichum braunii (Spenner) Fée Polystichum caruifolium Diels USSR: INDIA: Pulney Hills E I. Kikodse s.n. A. Sauliere 784 7 Jul 1913 [no collection date given] WTU-291304, WTU-V-056993 WTU-10128JWT, WTU-V-056963 Dryopteridaceae Dryopteridaceae Polystichum braunii (Spenner) Fée Polystichum angulare (Kit. ex Willd.) C.Presl NORWAY, ASKER: : Skougumaasen. [no locality given on label] Fr. Werenskiold s.n. Elsass s.n. 12 Oct 1895 [no collection date given] WTU-124791, WTU-V-056995 WTU-10178JWT, WTU-V-057076

Dryopteridaceae Dryopteridaceae Polystichum aculeatum (L.) Roth. var. aristatum (Christ) Polystichum angulare (Kit. ex Willd.) C.Presl Askerov ENGLAND: AUSTALIA, VICTORIA: Croughton. [no locality given on label] Steep Wood [illegible text]. Forests, Gullies [additional illegible notes] E. Waterfall 2892 unknown s.n. May 1930 1928 WTU-31152, WTU-V-057077 WTU-10184JWT, WTU-V-057070

Dryopteridaceae Dryopteridaceae Polystichum acrostichoides (Michx.) Schott Rumohra adiantiformis (G. Forst.) Ching U.S.A., INDIANA COUNTY: URUGUAY, CERRO LARGO.: Mt. Tom. [no locality given on label] 41.662642°, -87.056251°; WGS 84, uncertainty: 1000 m., Source: GeoLocate, Rosengurtt PE-4653 1/2 Georef'd by Ghislaine Ng Shaded dunes. 30 May 1942 L. M. Umbach 6236 WTU-122158, WTU-V-057073 29 Aug 1917 WTU-43049, WTU-V-057094

Dryopteridaceae Dryopteridaceae Polystichum acrostichoides (Michx.) Schott Polystichum angulare (Kit. ex Willd.) C.Presl U.S.A., NORTH CAROLINA, JACKSON ENGLAND: COUNTY: [no locality given on label] Near [what appears to read "Luckasinger" or Polydactylum (Homozygous). "Luckasingen"] Falls. I. Andersson s.n. Vida M. Lovett 55 1 Dec 1927 4 Jul 1934 WTU-316840, WTU-V-057075 WTU-56387, WTU-V-057095 Dryopteridaceae Dryopteridaceae Polystichum acrostichoides (Michx.) Schott Cyrtomium falcatum (L. f.) C. Presl U.S.A., IOWA, VAN BUREN COUNTY: JAPAN, FUKUOKA: Lacey Keosauqua State Park. Najima, Chikuzen [now a part of Fukuoka 40.715147°, -91.998888°; WGS 84, prefecture]. uncertainty: 2000 m., Source: Digital Map, Georef'd by Ghislaine Ng M. Takenouchi 152 Jess Sults s.n. 8 Apr 1922 May 1934 WTU-137167, WTU-V-057125 WTU-73126, WTU-V-057096

Dryopteridaceae Dryopteridaceae Polystichum acrostichoides (Michx.) Schott Polystichum aculeatum (L.) Roth. U.S.A., NORTH CAROLINA, MACON COUNTY: SWITZERLAND ?: Along Grover Creek near its junction with the [no locality given on label] Cullasaja River, 10 or so miles N. W. of Highlands. unknown s.n. Hardwood Forest. [no collection date given on label] R. L. Wilbur 1201 WTU-18974, WTU-V-057139 12 Aug 1948 WTU-141870, WTU-V-057106

Dryopteridaceae Dryopteridaceae Polystichum acrostichoides (Michx.) Schott Polystichum aculeatum (L.) Roth. U.S.A., VIRGINIA, ARLINGTON COUNTY: CANARY ISLANDS (SPAIN): Near Roslyn [Rosslyn]. Las Mercedes, Tenerlife. D. Leroy Topping s.n. Alice Carter Cook 1010 11 Aug 1895 Aug 1897 WTU-61419, WTU-V-057110 WTU-124795, WTU-V-057141

Dryopteridaceae Dryopteridaceae Polystichum acrostichoides (Michx.) Schott Polystichum aculeatum (L.) Roth. U.S.A., MARYLAND, MONTGOMERY JAPAN: COUNTY: Prov. Iwashiro. Garrett Park. 39.038165°, -77.093034°; WGS 84, A. Yasuda s.n. uncertainty: 1000 m., Source: GeoLocate, Georef'd by Ghislaine Ng 20 May 1914 D. Leroy Topping s.n. WTU-124796, WTU-V-057145 20 Oct 1895 WTU-61405, WTU-V-057116 Dryopteridaceae Dryopteridaceae Polystichum aculeatum (L.) Roth. Polystichum lemmonii Underw. FRANCE, HAUT-RHIN: U.S.A., CALIFORNIA, TRINITY COUNTY: Vosges, Alsace Near Siskiyou Co. line, Shasta National Forest; west and south slopes of Mt. Eddy. unknown s.n. Elev. 7800-9000 ft. 41.319533°, -122.480346°; WGS 84, [no collection date given] uncertainty: 1000 m., Source: Digital Map, Georef'd by Ghislaine Ng WTU-10183JWT, WTU-V-057147 Large clumps formed around rocks. Melinda F. Denton 4113 29 Jul 1977 WTU-271915, WTU-V-056946 Dryopteridaceae Polystichum aculeatum (L.) Roth. var. japonicum Franch. Dryopteridaceae et Sav. Polystichum acrostichoides (Michx.) Schott JAPAN, FUKUOKA: Shugakoe, Chikuzen [now Fukuoka]. U.S.A., NORTH CAROLINA, POLK COUNTY: var. japonicum. Mt. Side. Phenology: Spores. Origin: Native. M. Takenouchi 161 unknown s.n. 5 Mar 1922 8 Aug 1997 WTU-134141, WTU-V-057148 WTU-124794, WTU-V-057817

Dryopteridaceae Dryopteridaceae Polystichum aculeatum (L.) Roth. Polystichum acrostichoides (Michx.) Schott FRANCE, HAUTE-CORSE: Corse [Haute-Corse], Vizzavona (la Forêt). U.S.A., NEW YORK, SUFFOLK COUNTY: Elev. 950 m. [locality and coordinates withheld; status in New Bois peu humides. York: EV (Exploitably Vulnerable; USDA Plants)] Phenology: Spores. Origin: Native. B. de Retz s.n. unknown s.n. 1 Aug 1932 19 Jul 1919 WTU-10182JWT, WTU-V-057149 WTU-213111, WTU-V-057822

Dryopteridaceae Dryopteridaceae Polystichum acrostichoides (Michx.) Schott Polystichum fragrans (L.) Roth U.S.A., ARKANSAS, BRADLEY COUNTY: P.O. Johnsonville. CANADA, QUEBEC: Elev. 200 ft. Saguenay, Canada East, 18 Wooded bluff. 48.3771°, -71.694013°; WGS 84, uncertainty: 10000 m., Source: Digital map, Georef'd by Delzie Demaree 21025 Dan Post In clefts of rocks, on mossy cliffs, and 5 May 1940 sometimes on stony hillsides. Phenology: Spores. Origin: Native. WTU-185005, WTU-V-057090 D. A. Watt 25 [no collection date given] WTU-3230JWT, WTU-V-055702 Dryopteridaceae Dryopteridaceae Rumohra adiantiformis (G. Forst.) Ching Polystichum varium (L.) C.Presl AUSTRALIA, VICTORIA: JAPAN, MIYAGI: East Villt. Sendai. -36.479716°, 145.726294°; WGS 84, 38.291671°, 140.836974°; WGS 84, uncertainty: 300000 m., Source: Digital Map, uncertainty: 10000 m., Source: Digital Map, Georef'd by Dan Post Georef'd by Dan Post [undecipherable notes] Phenology: Spores. Phenology: Spores. Origin: Native. Origin: Native. A. Yasuda s.n. Allan Maebassui s.n. 23 Oct 1917 6 Dec 1929 WTU-213114, WTU-V-056777 WTU-10179JWT, WTU-V-057071

Dryopteridaceae Dryopteridaceae Polystichum lemmonii Underw. Polystichum acrostichoides (Michx.) Schott U.S.A., CALIFORNIA, SISKIYOU COUNTY: U.S.A., MISSOURI COUNTY: Scott Mt. near Shasta. Windsor Forest. 41.286458°, -122.679515°; WGS 84, 38.53224°, -93.522154°; WGS 84, uncertainty: 5000 m., Source: Digital Map, uncertainty: 10000 m., Source: GeoLocate, Georef'd by Marina DeMarco Georef'd by Dan Post Origin: Native. Christmas Fern. Origin: Native. J. G. Lemmon s.n. F. Comte 240 with and wife, Oakland California 15 May 1956 12 Aug 1878 WTU-329882, WTU-V-057102 WTU-1917, WTU-V-056947

Dryopteridaceae Dryopteridaceae Polystichum acrostichoides (Michx.) Schott Polystichum munitum (Kaulf.) C. Presl U.S.A., MAINE, KNOX COUNTY: U.S.A., WASHINGTON, CHELAN COUNTY: Camden, Flanks Mt.; Megunticook. Okanogan-Wenatchee National Forest. Forest 44.2413°, -69.06768°; WGS 84, uncertainty: Service Road 65 at Soda Springs campground 2500 m., Source: GeoLocate, Georef'd by area. Dan Post Elev. 682 m. Phenology: Spores. Origin: Native. 47.85821° N, 120.96986° W; WGS 84, uncertainty: 100 m., Source: GPS A. H. Norton s.n. Forested wetland and adjacent upland; Thuja plicata dominated in wet areas, Pseudotsuga 13 Aug 1913 menziesii where dry. Locally common in upland adjacent to wet area. Phenology: Spores. WTU-208514, WTU-V-057821 Origin: Native. David Giblin 7044 Dryopteridaceae 19 Jul 2019 Polystichum californicum (D.C. Eaton) Diels WTU-425226, WTU-V-068795 U.S.A., CALIFORNIA, MARIN COUNTY: Forest between Phoenix Lake and Lake Lagunitas. 37.949952°, -122.585961°; WGS 84, uncertainty: 1500 m., Source: Digital Map, Georef'd by Dan Post Phenology: Spores. Origin: Native. Fred Hein s.n. 12 Nov 1931 WTU-214194, WTU-V-057830 Dryopteridaceae Dryopteridaceae Polystichum scopulinum (D.C. Eaton) Maxon Polystichum kruckebergii W.H. Wagner U.S.A., WASHINGTON, CHELAN COUNTY: U.S.A., WASHINGTON, OKANOGAN COUNTY: Okanogan-Wenatchee National Forest. White River Road. Pacific Crest Trail approximately 4 km south from Brown Bear Approximately 2 km east of Grasshopper Meadow Campground. trailhead from Saddle below Tatie Peak to near top of ridgeline to the Elev. 596 m. south with unnamed peak. GPS reading from highest point. 47.92873° N, 120.90678° W Elev. 2285 m. South-facing boulder and talus slope and adjacent rocky 48.68962° N, 120.71127° W; WGS 84, uncertainty: 500 m., outcroppings in broken canopy, Pseudotsuga menziesii and Pinus Source: GPS ponderosa pine forest. Pteridium aquilinum and Moehringia Mix of east and northern exposures with scree slopes dry fellfields macrophyllum common. Rock brownish-orange, possibly ultramafic. and rocky outcroppings; Silene acaulis, Saxifraga austormontana, Uncommon in rock crevice; lower pinnae deeply incised. Phenology: and Antenarria lanata common. Uncommon in rock crevices; last Spores. Origin: Native. year's fronds absent. Phenology: Spores. Origin: Native. David Giblin 6882 16 Jul 2019 David Giblin 7780 19 Aug 2020 Herbarium: WTU Herbarium: WTU

Dryopteridaceae Dryopteridaceae Polystichum munitum (Kaulf.) C. Presl Polystichum lonchitis (L.) Roth U.S.A., WASHINGTON, CHELAN COUNTY: U.S.A., WASHINGTON, OKANOGAN COUNTY: Okanogan-Wenatchee National Forest. White River Road. Pacific Crest Trail approximately 4 km south from Brown Bear Approximately 2 km east of Grasshopper Meadow Campground. trailhead from Saddle below Tatie Peak to near top of ridgeline to the Elev. 596 m. south with unnamed peak. GPS reading from highest point. 47.92873° N, 120.90678° W Elev. 2285 m. South-facing boulder and talus slope and adjacent rocky 48.68962° N, 120.71127° W; WGS 84, uncertainty: 500 m., outcroppings in broken canopy, Pseudotsuga menziesii and Pinus Source: GPS ponderosa pine forest. Pteridium aquilinum and Moehringia Mix of east and northern exposures with scree slopes dry fellfields macrophyllum common. Rock brownish-orange, possibly ultramafic. and rocky outcroppings; Silene acaulis, Saxifraga austormontana, Common and abundant throughout area. Phenology: Spores. Origin: and Antenarria lanata common. Scattered in rock crevices; last Native. year;s fronds present. Phenology: Spores. Origin: Native. David Giblin 6889 16 Jul 2019 David Giblin 7779 19 Aug 2020 Herbarium: WTU Herbarium: WTU Dryopteridaceae Dryopteridaceae Polystichum lonchitis (L.) Roth Polystichum kruckebergii W.H. Wagner U.S.A., WASHINGTON, CHELAN COUNTY: Okanogan-Wenatchee National Forest. Mountain David trail from U.S.A., WASHINGTON, OKANOGAN COUNTY: where it emerges from forest into historic burn to approximately 0.5 Okanogan-Wenatchee National Forest. Pacific Crest Trail km from summit. approximately 4.5 km south of Brown Bear Trailhead. Elev. 200 m. Elev. 2054 m. 47.9592° N, 120.9866° W; WGS 84, uncertainty: 1000 m., 48.69103° N, 120.70537° W; WGS 84, uncertainty: 100 m., Source: Digital map Source: GPS Mix of north and south-facing slopes along ridgeline; Penstemon Southeast-facing scree and talus slope with outcroppings that may davidsonii and Lomatium brandegeei common and abundant. be ultramafic rock; Lomatium brandegeei, Eucephalus engelmannii, Uncommon on north-facing rocky outcropping in crevices. and Arnica common. Uncommon in rock crevices and rock Phenology: Spores. Origin: Native. overhangs; last year's fronds absent. Phenology: Spores. Origin: Native. David Giblin 6948 17 Jul 2019 David Giblin 7785 19 Aug 2020 Herbarium: WTU Herbarium: WTU Dryopteridaceae Dryopteridaceae Polystichum lonchitis (L.) Roth Polystichum munitum (Kaulf.) C. Presl U.S.A., WASHINGTON, KING COUNTY: U.S.A., WASHINGTON, CLALLAM COUNTY: Above FS Road NF-7034, 4.6 air km ESE of Adjacent to an unnamed tributary to the Soleduck River, west of the Kelly Butte, W slope of Cascade Range. community of Shuwah. Elev. 1445 m. 48.031666° N, 124.397378° W 47.1523° N, 121.4352° W Moist on hummocks; part shade; Forested wetland hummocks. Steep NW-facing talus, with Vaccinium Common, forb, erect, Phenology: Spores. Origin: Native. membranaceum, Sedul divergens, Arnica latifolia. Cespitose, occasional. Patricia Johnson NWC21-WA-10039-Q22 29 Jun 2021 P. F. Zika 29999 Herbarium: WTU 5 Jul 2020 WTU-424403, WTU-V-067171