The First Fish Field Guide Ever Published Dedicated to the Tubbataha Reefs
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The first fish field guide ever published dedicated to the Tubbataha Reefs Compiled for the use of guests, visitors, and marine scientists With over 200 pictures of different fish species Documented >30% of the total resident population (633) TUBBATAHA REEFS FISH GUIDE TUBBATAHA REEFS FISH GUIDE Acknowledgements Author’s note Foreword Acanthuridae Surgeonfishes, Unicornfishes (p. 1) Aulostomidae Trumpetfishes (p. 5) Balistidae Triggerfishes (p. 6) Caesionidae Fusiliers (p. 8) Carangidae Jacks, Trevallies (p. 9) Chaetodontidae Butterflyfishes (p. 10) Cirrhitidae Hawkfishes (p. 14) Ephippidae Batfishes (p. 15) Fistularidae Cornetfishes (p. 16) Gobiidae Gobies (p. 17) Sweetlips (p. 18) Haemulidae Holocentridae Squirrelfishes (p. 19) Kyphosidae Sea chubs (p. 20) Labridae Wrasses (p. 21) Lethrinidae Emperors (p. 26) Lutjanidae Snappers (p. 29) Mullidae Goatfishes (p. 32) Muraenidae Moray eels (p. 33) Nemipteridae Whiptail breams (p. 34) Ostraciidae Boxfishes (p. 35) Pinguipedidae Sand perches (p. 36) Pomacanthidae Angelfishes (p. 37) Pomacentridae Damselfishes (p. 38) Priacanthidae Bigeyes (p. 44) Ptereleotridae Dartfishes (p. 45) Scaridae Parrotfishes (p. 46) Scorpaenidae Scorpionfishes, Firefishes (p. 49) Serranidae Groupers, Sea basses (p. 50) Siganidae Rabbitfishes (p. 54) Sphyraenidae Barracudas (p. 55) Synodontidae Lizardfishes (p. 56) Tetraodontidae Pufferfishes (p. 57) Zanclidae Moorish idol (p. 59) FAMILY ACANTHURIDAE Surgeonfishes, Tangs, Unicornfishes from Greek Akantha Class ACTINOPTERYGII (thorn) + Oura (tail) Subclass Order PERCIFORMES Suborder ACANTHUROIDEI REMARKS - All species marine, circumtropical distribution, 82 species recorded under 6 genera - Small and terminal mouth, ctenoid scales, single row close-set teeth, many are brightly colored - Feed on benthic algae, zooplankton, and detritus. Pelagic spawners © Toppx2 © Segundo Conales Jr. 2013 JAPANESE SURGEONFISH LINED SURGEONFISH Acanthurus japonicus Acanthurus lineatus Max. length: Max. length: © Segundo Conales Jr. 2013 © Art Faburada ELONGATE SURGEONFISH WHITECHEEK SURGEONFISH Acanthurus mata Acanthurus nigricans Max. length: Max. length: 1 Tubbataha Reefs Fish Book Volume 1 © Reef Fish Identification: Tropical Pacific. p. 48 © Reef Fish Identification: Tropical Pacific. p. 50 BLACKSTREAK SURGEONFISH BROWN SURGEONFISH Acanthurus nigricaudus Acanthurus nigrofuscus Max. length: Max. length: © Reef Fish Identification: Tropical Pacific. p. 45 © Art Faburada ORANGESPOT SURGEONFISH CHOCOLATE SURGEONFISH Acanthurus olivaceus Acanthurus pyroferus Max. length: Max. length: © Art Faburada © Segundo Conales Jr. 2013 STRIATED SURGEONFISH WHITEMARGIN UNICORNFISH Ctenochaetus striatus Naso annulatus Max. length: Max. length: Tubbataha Reefs Fish Book 2 Volume 1 © Art Faburada © Segundo Conales Jr. 2013 SPOTTED UNICORNFISH SLEEK UNICORNFISH Naso brevirostris Naso hexacanthus Max. length: Max. length: © Segundo Conales Jr. 2013 © Segundo Conales Jr. 2013 ORANGESPINE UNICORNFISH ONEKNIFE UNICORNFISH Naso lituratus Naso thynnoides Max. length: Max. length: © Art Faburada © Art Faburada BLUESPINE UNICORNFISH BIGNOSE UNICORNFISH Naso unicornis Naso vlamingii Max. length: Max. length: Tubbataha Reefs Fish Book 3 Volume 1 © Art Faburada © Segundo Conales Jr. 2013 TWOTONE TANG SAILFIN TANG Zebrasoma scopas Zebrasoma velifer Max. length: Max. length: Tubbataha Reefs Fish Book 4 Volume 1 FAMILY AULOSTOMIDAE Trumpetfishes from Greek Aulos Class ACTINOPTERYGII (flute) + Stoma (mouth) Subclass Order SYNGNATHIFORMES Suborder REMARKS - All species marine, found in tropical atlantic and indo-pacific regions, 3 species recorded under 6 genera - Compressed and elongated scaly body, fleshy barbels at the tip of lower jaw, rounded caudal fin - Ambush predator, feeds on small fishes and crustaceans. Spawners © Art Faburada CHINESE TRUMPETFISH Aulostomus chinensis Max. length: 5 Tubbataha Reefs Fish Book Volume 1 FAMILY BALISTIDAE Triggerfishes from Greek Balein Class ACTINOPTERYGII (to throw) Subclass once considered as “trash fish” Order TETRAODONTIFORMES Suborder REMARKS - All species marine, found in Atlantic and Indo-Pacific regions, 42 species recorded under 12 genera - Odd-shaped compressed body, swim with their 2nd dorsal (adipose) and anal fins - Most species are carnivorous, some feed on algae. Lay demersal eggs, aggresive guarders © Art Faburada © Toppx2 ORANGE-LINED TRIGGERFISH CLOWN TRIGGERFISH Balistapus undulatus Balistoides conspicillum Max. length: Max. length: © Art Faburada © Segundo Conales Jr. 2013 TITAN TRIGGERFISH BLACK TRIGGERFISH Balistoides viridescens Melichthys niger Max. length: Max. length: Tubbataha Reefs Fish Book 6 Volume 1 © Art Faburada © Segundo Conales Jr. (2013) PINKTAIL TRIGGERFISH RED-TOOTHED TRIGGERFISH Melichthys vidua Odonus niger Max. length Max. length © Art Faburada © Segundo Conales Jr. (2013) BOOMERANG TRIGGERFISH HALFMOON TRIGGERFISH Sufflamen bursa Sufflamen chrysopterum Max. length Max. length © Art Faburada GILDED TRIGGERFISH Xanthichthys auromarginatus Max. length Tubbataha Reefs Fish Book 7 Volume 1 FAMILY CAESIONIDAE Fusiliers from Greek Caesius Class ACTINOPTERYGII (bluish-grey) Subclass Order PERCIFORMES Suborder PERCOIDEI REMARKS - All species marine, thrive in Indo-west Pacific, 23 species recorded under 4 genera - Elongate and fusiform body, small mouth, deeply-forked caudal fin - Active swimmers, feed on zooplankton. Spawners, nonguarders © Art Faburada © Art Faburada BLUE AND GOLD FUSILIER DARK-BANDED FUSILIER Caesio caerulaurea Pterocaesio tile Max. length: Max. length: © Segundo Conales Jr. (2013) BANANA FUSILIER Pterocaesio pisang Max. length: Tubbataha Reefs Fish Book 8 Volume 1 FAMILY CARANGIDAE CHAETODONTIDAE Jacks, Trevallies from French Carangue Class ACTINOPTERYGII (name of a Caribbean Subclass fish) Order PERCIFORMES Suborder REMARKS - Chiefly marine. rare in brackish environment, Atlantic and Indo-Pacific, 148 species under 30 genera - Shape varies from very deep to fusiform, spiny scutes along lateral line, widely-forked caudal fin - Fast swimmers, feed on small fishes and invertebrates. Spawners, nonguarders © Segundo Conales Jr. (2013) © Segundo Conales Jr. (2013) COACHWHIP TREVALLY GIANT TREVALLY Carangoides oblongus Caranx ignobilis Max. length: Max. length: © Segundo Conales Jr. (2013) © Art Faburada BLUEFIN TREVALLY BIGEYE TREVALLY Caranx melampygus Caranx sexfasciatus Max. length: Max. length: 9 Tubbataha Reefs Fish Book Volume 1 FAMILY CHAETODONTIDAE Butterflyfishes from Greek Chaite Class ACTINOPTERYGII (hair) + Odontos (teeth) Subclass NEOPTERYGII Order PERCIFORMES Suborder PERCOIDEI REMARKS - Marine-brackish, found in tropical Atlantic and Indo-west Pacific, 129 species recorded under 12 genera - Highly compressed body, mouth protrusible with small brushlike teeth, most have “eyespot” - Diet includes coelenterate polyps, inverts, fish eggs, filamentous algae, others planktivore. Spawners © Segundo Conales Jr. (2013) © Segundo Conales Jr. (2013) PHILIPPINE BUTTERFLYFISH THREADFIN BUTTERFLYFISH Chaetodon adiergastos Chaetodon auriga Max. length: Max. length: © Art Faburada © Segundo Conales Jr. (2013) EASTERN TRIANGULAR BUTTERFLYFISH BLUELASHED BUTTERFLYFISH Chaetodon baronessa Chaetodon bennetti Max. length: Max. length: Tubbataha Reefs Fish Book 10 Volume 1 © Segundo Conales Jr. (2013) © Segundo Conales Jr. (2013) SADDLE BUTTERFLYFISH SUNBURST BUTTERFLYFISH Chaetodon ephippium Chaetodon kleinii Max. length: Max. length: © Segundo Conales Jr. (2013) © Toppx2 BLACKBACK BUTTERFLYFISH ORNATE BUTTERFLYFISH Chaetodon melannotus Chaetodon ornatissimus Max. length: Max. length: © Segundo Conales Jr. (2013) © Art Faburada SPOT-NAPE BUTTERFLYFISH SPOTBAND BUTTERFLYFISH Chaetodon oxycephalus Chaetodon punctatofasciatus Max. length: Max. length: Tubbataha Reefs Fish Book 11 Volume 1 © Art Faburada © Segundo Conales Jr. (2013) LATTICED BUTTERFLYFISH CHEVRON BUTTERFLYFISH Chaetodon rafflesi Chaetodon trifascialis Max. length: Max. length: © Segundo Conales Jr. (2013) © Segundo Conales Jr. (2013) MELON BUTTERFLYFISH PACIFIC DOUBLE-SADDLE BUTTERFLYFISH Chaetodon trifasciatus Chaetodon ulietensis Max. length: Max. length: © Segundo Conales Jr. (2013) © Art Faburada VAGABOND BUTTERFLYFISH LONGNOSE BUTTERFLY FISH Chaetodon vagabundus Forcipiger flavissimus Max. length: Max. length: Tubbataha Reefs Fish Book 12 Volume 1 CIRRHITIDAE © Segundo Conales Jr. (2013) © Toppx2 LONGNOSE BUTTERFLYFISH PENNANT CORALFISH Forcipiger longirostris Heniochus acuminatus Max. length: Max. length: © Art Faburada HORNED BANNERFISH Heniochus varius Max. length: Tubbataha Reefs Fish Book 13 Volume 1 FAMILY CIRRHITIDAE Hawkfishes from Latin Cirrus Class ACTINOPTERYGII (curl, fringe) Subclass Order PERCIFORMES Suborder REMARKS - All species marine, found in Eastern and Western Atlantic and Indo-Pacific, 33 species from 12 genera - Usually small and colorful. Protogynous hermaphrodites - Feed on small crustaceans and fishes. Pelagic spawners, nonguarders © Art Faburada © Art Faburada DWARF HAWKFISH ARC-EYE HAWKFISH Cirrhitichthys falco Paracirrhites arcatus Max. length: Max. length: © Art Faburada BLACKSIDE HAWKFISH Paracirrhites forsteri Max. length: Tubbataha Reefs Fish Book 14 Volume 1 FAMILY EPHIPPIDAE FISTULARIIDAE Batfishes, Spadefishes from Latin Ephippium Class ACTINOPTERYGII (horse saddle) Subclass Order PERCIFORMES Suborder REMARKS - Rarely found in estuaries, present in Atlantic, Indian, and Pacific regions, 15 species under 8 genera - Compressed body, small mouth, wing-like dorsal and anal fins - Omnivorous species. Spawners © Art Faburada