DOCSLIB.ORG

Status and Palaeobiology of the Late Cretaceous Indian Theropods with Description of a New Theropod Eggshell Oogenus and Oospeci

Total Page：16

File Type：pdf, Size：1020Kb

Download full-text PDF Read full-text

Load more

GAIA N' 15, LlSBOAlLISBON, DEZEMBRO/DECEMBER 1998, pp. 379-387 (ISSN: 0871-5424) STATUS AND PALAEOBIOLOGY OF THE LATE CRETACEOUS INDIAN THEROPODS WITH DESCRIPTION OF A NEW THEROPOD EGGSHELL OOGENUS AND OOSPECIES, ELLIPSOOLITHUS KHEDAENSIS, FROM THE LAMETA FORMATION, DISTRICT KHEDA, GUJARAT, WESTERN INDIA Raminder S. LOYAL Centre of Advanced Study in Geology, Panjab University. CHANDIGARH· 160014. INDIA Dhananjay M. MOHABEY Palaeontology Division, Geological Survey of India, Seminary Hills. NAGPUR - 440 006. INDIA Ashu KHOSLA Centre of Advanced Study in Geology, Panjab University. CHANDIGARH - 160014. INDIA Ashok SAHNI Centre of Advanced Study in Geology, Panjab University. CHANDIGARH -160014. INDIA ABSTRACT: This paper highlights the Cretaceous theropod record in India along with its af­ finities, and palaeobiologic aspects including description of a new theropod eggshell from the Upper Cretaceous Lameta Formation of Gujarat, India. Recent discoveries and phyloge­ netic reassessment of various theropod families suggest the presence of the poorly known family Abelisauridae; recent reports confirm the presence of Majungasaurus crenatissimus (DEPRET, 1896), Indosuchus HUENE & MATLEY, 1933 possible Carnotaurus BONAPARTE, 1985 and ?Allosaurus MARSH, 1877. An updated list of various Cretaceous theropod genera in­ cludes abelisaurids (Indosuchus raptorius HUENE & MATLEY, 1932, Indosaurus matleyi HUENE & MATLEY, 1933, Majungasaurus crenatissimus (DEPRET, 1896) Carnotaurus sp., allo­ saurids (Compsosuchus solus HUENE, 1932, ?Allosaurus) and coelurosaurs (Jubbulpuria tenuis HUENE & MATLEY, 1933, Laevisuchus indicus HUENE & MATLEY, 1933). Other theropods, such as Ornithomimoides mobilis HUENE & MATLEY, 1933, O. (?)barasim/ensis HUENE & MAT­ LEY, 1933 and Dryptosauroidesgrandis HUENE, 1932 regarded as indeterminate, need further assessment for proper affiliation. In the ongoing discussion, we however, present signifi­ cant new data on the first ever report of dinosaur eggshell material referable to a new ooge­ nus and oospecies Ellipsoolithus khedaensis, (oofamily Elongatoolithidae) from near Rahioli, district Kheda, Gujarat. This material has been presently assigned to the Theropoda and can be easily differentiated from the other well known megaloolithid oospecies (broadly referable to Sauropoda due to associated titanosaurid bones; known from Madhya Pradesh, Gujarat, Maharashtra and Ariyalur in South India) on the basis of megascopic, microscopic and ultrastructural characteristics. Megascopically, the eggshell shape varies from elon­ gated ellipsoid, having an elongation ratio range of 1.5 to 1.7. The eggshell thickness is 1.2- 1.6 mm and the mammillary layer is 1/4-1/7 of the total thickness. At the microstructural level, an angusticanaliculate pore system is observed. As the present find constitutes the first record of elongatoolithid eggshells apart from well known Chinese and Mongolian lo­ calities, comments on possible sauropod-theropod palaeobiology are also discussed. 379 artigos/papers R.S. LOYAL; O.M. MOHABEY; A. KHOSLA & A. SAHNI THEROPOD RECORD IN INDIA AND gascar) including cross-sectional and serrational EGGSHELL STATUS morphologies are strongly similar to that of the In­ dian theropod Indosuchus raptorius HUENE & MAT­ Although Indian sauropod eggshell taxonomy lEY, 1933; for instance, both forms exhibit marked and skeletal discoveries have made considerable pattern of vertical ridges on the interdental plates. As advancements overthe lastfew decades, the thero­ such, the shared synapomorphies between these pods unfortunately have received scant attention theropods have led SAMPSON et al. (1996) to consid­ except for notable exceptions (HUENE & MATlEY, erthem as sister taxa within the clade Abelisauridae. 1933; CHATTERJEE, 1978; CHATTERJEE & RUDRA, This viewpoint is also corroborated by CHATTERJEE 1996). Except for the presence of a Triassic small & RUDRA (1996); in a detailed discourse on Gon­ theropod Alwalkeria maleriensis CHATTERJEE & dwana theropod relationships, these workers asso­ CREISlER, 1994 from the Maleri Formation ciated a more primitive phylogenetic position to the (Pranhita-Godawari Valley, district Adilabad, abelisaurids in comparison to the allosaurs and ty­ Andhra Pradesh), the dominant discoveries of rannosaurs. Further, the remains of Majungasaurus theropod skeletal remains are associated with the crenatissimus (Mahajanga Basin, Upper Creta­ Upper Cretaceous Lameta and Intertrappean se­ ceous) represented by one premaxilla has been quences outcropping in central and southern parts confirmed to belong to an abelisaurid with no "mega­ of peninsular India. Although no Jurassic theropod losaurian" ortyrannosaurid characteristics (KRAUSE record exists, the later theropods are stratigraphi­ & DODSON , 1994; KRAUSE & HARTMAN, 1996). More cally associated with the Upper Cretaceous Lameta recently, CHATTERJEE & RUDRA (1996: 517) believe and Intertrappean sequences, outcropping in cen­ that "Indosuchus is not a tyrannosaurid as was sup­ tral and southern parts of Peninsular India. We pro­ posed earlier, but possibly an abelisaurid". These vide below a brief overview of theropod groups workers discovered a complete skeleton of Indo­ prevailing in the Indian subcontinent during the Cre­ suchus from near Rahioli, Gujarat and referred it to taceous times. allosaurids on the basis of similar lengths of fore­ The Indian Cretaceous theropods are broadly limbs in these two taxa; however, hindlimb elements represented by skeletal remains of abelisaurids, al­ resemble those of Camotaurus, a well-established losaurids, coelurosaurs and some indeterminate abelisaurid. forms (HUENE & MATlEY, 1933; BUFFETAUT, 1987; Further, the other theropod known from Bara KRAUSE & HARTMAN, 1996; CHATTERJEE & RUDRA, Simla Hill, Indosaurus mat/eyi, bears thickened fron­ 1996). Amongst these forms, the taxonomic affini­ tals and elevated parietals; these features are also ties of larger theropods have remained in doubt, observed in Camotaurus sastrei, a well established awaiting further assessment; in particular "megalo­ and a diagnostic abelisaurid known from Goro Frigio saurids" from Kallamedu, south India (BLANFORD, Formation (Albanian-Cenomanian) of Chubut, Ar­ 1862; LYDEKKER , 1879), Takli (Maharashtra, gentina (BONAPARTE, 1985). Moreover, vertebral VIANEY-LiAUD, JAIN & SAHNI, 1987) and Rahioli elements closely similar to that of Camotaurus are (MATHUR & SRIVASTAVA, 1987). However, in com­ also known from Rahioli, Gujarat (CHATTERJEE & parison, the family Abelisauridae is a relatively bet­ RUDRA, 1996). ter known group from the Carnosaur Bed of Lameta Formation, Jabalpur and is represented by Indo­ The remaining forms from Bara Simla Hill locality suchus raptorius HUENE & MATlEY, 1933, Indosau­ by HUENE & MATlEY (1933)were referred to coeluro­ rus mat/eyi HUENE & MAllEY, 1933 and saurs, which included Compsosuchus solus HUENE, Camotasaurus BONAPARTE , 1985. Of these, the first 1932, Laevisuchus indicus HUENE & MATlEY, 1933, two forms, Indosuchus HUENE & MATlEY, 1933 and Jubbulpuria tenuis HUENE & MATlEY, 1933, Coelu­ Indosaurus HUENE & MATlEY, 1933 originally re­ roides largus HUENE, 1932, Oryptosauroides gran­ ferred to Allosauridae (HUENE & MATlEY, 1933), dis HUENE, 1932, Omithomimoides mobilis HUENE & have been variously assigned, for instance Indo­ MATlEY, 1933 and O. (?) barasimlensis HUENE & suchus as a tyrannosaur and Indosaurus as a mega­ MATlEY, 1933. Of these forms, Compsosuchus so­ losaur (CHATTERJEE, 1978; BUFFETAUT, 1987); and Ius was suggested by MOLNAR, KURZANOV & DONG more recently to abelisaurids (BONAPARTE & NOVAS, (1990) and MOLNAR & FARLOW (1990) to be an allo­ 1985; MOLNAR, 1990; MOLNAR, KURZANOV & DONG , saurid and particularly close to Allosaurus fragilis, 1990 and SAMPSON et al., 1996). on the basis of shared apomorphies. Recently, however, morphologic and phyloge­ From Bara Simla Hill locality, the only forms to netic comparison of various theropods including have retained their assignment to coelurosaurs are abelisaurids has been attempted by SAMPSON et al. Jubbulpuria tenuis and Laevisuchus indicus, but (1996). In this work, it has been considered that pre­ thought to be indeterminate (NORMAN, 1990). Other maxillary dentition of Majungasaurus crenatissimus forms which remain unidentified and await assess­ (DEPRET, 1896) (from Upper Cretaceous of Mada- ment are Coeluroides largus, Oryptosauroides 380 STA TUS AND PALAEOBIOLOGY OF THE LATE CRETACEOUS INDIAN THEROPODS 2f s' .Lavario ~ Muwodo , ,~ Ohl011.- , . timnton. (Colcrete ) 23 f~~~~ ~ _H ..U :!w :-:......:-:-:.: . a-ritty Sand~tont ~ 0=0_' , , km 1_0~00 • • • • • • Oronite r • • • ~ Lom.to Formation (LotI' Cr.tQcI'OU~ ) ~ Nf'!tho n ••ly found I'llfp!toid 1'99\. I~I Ntnt of earlier known 1ophl'riCo' dino\our 1'99", I:::J ISOnH of Titano!tDurid" and The-ropod" Fig. 1 - Location of newly found ellipsoidal eggs (elongatoolithid) in Kheda district, Gujarat. grandis, Omithomimoides mobilis and O. (?) bara­ more recently elongatoolithid and megaloolithid egg simlensis. Further, D. grandis is considered by MOL­ nests have been recorded from the same stra­ NAR (1990) to possess no carnosaurian characters, tigraphic level in this area. At least 60 eggs (ellipsoi­ while ornithomimosaurids are not present south of dal) have been located in more than half a dozen Mongolian localities (BARSBOLD & OSMOLSKA, nests in addition to stray eggs and eggshell debris 1990). (MOHABEY, 1996b.) A single nest has yielded thir­ teen eggs. In a single nest the eggs are disposed PRESENT
Recommended publications
  • Factors Controlling Detrital Mineralogy of the Sandstone of the Lameta Formation (Cretaceous), Jabalpur Area, Madhya Pradesh, India

    Factors Controlling Detrital Mineralogy of the Sandstone of the Lameta Formation (Cretaceous), Jabalpur Area, Madhya Pradesh, India

    FactorsProc Indian Controlling Natn Sci Acad Detrital 74 No.2 Mineralogy pp. 51-56 (2008)of the Sandstone of the Lameta Formation 51 Research Paper Factors Controlling Detrital Mineralogy of the Sandstone of the Lameta Formation (Cretaceous), Jabalpur Area, Madhya Pradesh, India AHM AHMAD ANSARI*, SM SAYEED** and AF KHAN*** Department of Geology, Aligarh Muslim University, Aligarh 202 002 (UP) (Received 7 February 2008; Accepted 6 May 2008) Cretaceous (Maastrichtian) deposits of the Lameta Formation crop out along the eastern part of Jabalpur basin on isolated hills and along the banks of Narmada River near Jabalpur city. The quartzarenite composition with little amounts of feldspar, mica, rock fragments and heavy minerals, are medium to fine grained, moderately sorted to poorly sorted and subangular to subrounded. The study suggests that palaeoclimate, distance of transport and source rock composition influenced the detrital mineralogy of the sandstone. By using Suttner and Dutta diagram, the mean values of the ratio were plotted and that indicate a humid Paleoclimate in this area. The plate tectonic setting and provenance of the sandstone were interpreted using the Dickinson’s method of detrital modes and Qt-F-L, Qm-F-Lt, Qp-Lv-Ls and Qm-P-K triangular diagrams. The petrofacies analysis of the Lameta Formation suggest mainly craton interior in a rifted continental margin basin setting. The plot of various quartz types on diamond diagram after [17] reflects Plutonic terrain. The probable provenance of these sandstones is Mahakoshal and Jabalpur Groups. Key Words: Cretaceous; Lameta Formation; Jabalpur; Mineralogy; Madhya Pradesh; India 1. Introduction Table 1. Stratigraphy of Lameta Formation, Jabalpur area (Madhya Pradesh); Tandon et al.
  • A Partial Skeleton of the Tyrannosaurid Dinosaur Aublysodon the Upper Cretaceous of New Mexico Thomas M

    A Partial Skeleton of the Tyrannosaurid Dinosaur Aublysodon the Upper Cretaceous of New Mexico Thomas M

    J. Paleont., 64(6), 1990, pp. 1026-1032 Copyright © 1990, The Paleontological Society 0022-3360/90/0064-1026S03.00 A PARTIAL SKELETON OF THE TYRANNOSAURID DINOSAUR AUBLYSODON THE UPPER CRETACEOUS OF NEW MEXICO THOMAS M. LEHMAN AND KENNETH CARPENTER Department of Geosciences, Texas Tech University, Lubbock 79409, and Department of Earth Sciences, Denver Museum of Natural History, City Park, Denver, Colorado 80205 Abstract—A fragmentary tyrannosaurid skull and postcranial skeleton from the Kirtland Shale of northwestern New Mexico is the most complete specimen of a carnivorous dinosaur known from these strata. The specimen is identified as Aublysodon cf. A. mirandus on the basis of its narrow frontals, V-shaped frontal-parietal suture, and nondenticulate incisiform premaxillary tooth. The D-shaped cross section of the premaxillary tooth, rugose postorbital, well-developed footed pubis, and proximally constricted third metatarsal confirm the assignment of Aublysodon to the Tyrannosauridae. The limb bones are gracile and similar in proportions to those of Albertosaurus', however, the tibia and metatarsals are shorter relative to the femur. The distal end of the tibia exhibits a unique medial emargination not reported in other tyrannosaurids. INTRODUCTION imen (OMNH 10131) was collected in San Juan County, New Mexico, northeast of Chaco Canyon in June, 1940, by J. W. arge carnivorous dinosaurs were first reported from Up­ Stovall and D. E. Savage of the University of Oklahoma. The per Cretaceous continental deposits in the San Juan Basin remains comprise several skull fragments, an incomplete den­ Lof northwestern New Mexico by Barnum Brown (1910, p. 268)tary, parts of both femora, a tibia, pubis, metatarsals, several who found “disassociated limb bones ..
  • [PDF] Dinosaur Eggshell from the Red Sandstone Group of Tanzania

    [PDF] Dinosaur Eggshell from the Red Sandstone Group of Tanzania

    Journal of Vertebrate Paleontology 24(2):494±497, June 2004 q 2004 by the Society of Vertebrate Paleontology NOTE DINOSAUR EGGSHELL FROM THE RED SANDSTONE GROUP OF TANZANIA MICHAEL D. GOTTFRIED1, PATRICK M. O'CONNOR2, FRANKIE D. JACKSON3, ERIC M. ROBERTS4, and REMEGIUS CHAMI5, 1Mich- igan State University Museum, East Lansing, Michigan, 48824, [email protected]; 2College of Osteopathic Medicine, Ohio University, Athens, Ohio, 45701; 3Department of Earth Sciences, Montana State University, Bozeman, Montana, 59717; 4Department of Geology and Geophysics, University of Utah, Salt Lake City, Utah, 84112, 5Antiquities Unit, P.O. Box 2280, Dar es Salaam, Tanzania Investigations over the last several decades at Gondwanan Mesozoic Although the age of the Red Sandstone Group is poorly understood (see localities have signi®cantly expanded our knowledge of the diversity Damblon et al., 1998), a Cretaceous age is suggested at this site based and distribution of Southern Hemisphere dinosaurs. These records are on (1) the overall composition of the fauna, which includes titanosaurid? primarily based on skeletal remains, but included among them are in- sauropods and both avian and nonavian theropods, as well as osteo- stances of preserved eggshell, notably from Argentina (e.g., Calvo et glossomorph ®shes, and (2) the possibility that these deposits may be al., 1997; Chiappe et al., 1998) and India (e.g., Khosla and Sahni, 1995). approximately coeval with the Cretaceous dinosaur beds of Malawi (Ja- In general, however, dinosaur eggshell is relatively poorly known from cobs et al., 1990), which lie ca. 200 km southeast of the Mbeya region. Gondwana, and from Africa in particular.
  • Jurassic Algae of the Perachora-Peninsula: Biostratigraphical and Paleoecological Implications

    Jurassic Algae of the Perachora-Peninsula: Biostratigraphical and Paleoecological Implications

    ZOBODAT - www.zobodat.at Zoologisch-Botanische Datenbank/Zoological-Botanical Database Digitale Literatur/Digital Literature Zeitschrift/Journal: Beiträge zur Paläontologie Jahr/Year: 1994 Band/Volume: 19 Autor(en)/Author(s): Dragastan Ovidiu, Gielisch Hartwig, Richter Detlef K., Grewer Till, Kaziur Thomas, Kube Bärbel, Radusch Christoph Artikel/Article: Jurassic algae of the Perachora-Peninsula: Biostratigraphical and paleoecological implications 49-81 ©Verein zur Förderung der Paläontologie am Institut für Paläontologie, Geozentrum Wien Beitr. Paläont., 19:49-81, Wien 1994 Jurassic algae of the Perachora-Peninsula: Biostratigraphical and paleoecological implications Jurassische Algen der Perachora Halbinsel: Biostratigraphische und paläoökologische Folgerungen by DRAGASTAN, Ovidiu,* GIELISCH, Hartwig**, RICHTER, Detlev K.**, GREWER, TiU**, KAZIUR, Thomas**, KUBE, Bärbel **& RADUSCH, Christoph** DRAGASTAN, O., GIELISCH, H„ RICHTER, D.K., GREWER, T„ KAZIUR, T„ KUBE, B. & RADUSCH, C., 1994. Jurassic algae of the Perachora-Peninsula: Biostratigraphical and paleoecological implications. — Beitr. Palaont., 19:49-81, 9 Figures, 6 Plates, Wien. ceae), Rivularia (Rivulariaceae), Alpinelia graeca n.sp. Contents (Scytonemataceae), Girvanella, Hedstroemia (Porostro- Abstract, Zusammenfassung.................................................. 49 mata) and Microproblematicae. 1. Introduction..............................................................................50 Biostratigraphically, a succession of algae and foramini- 2. Geological setting.................................................................
  • Perinate and Eggs of a Giant Caenagnathid Dinosaur from the Late Cretaceous of Central China

    Perinate and Eggs of a Giant Caenagnathid Dinosaur from the Late Cretaceous of Central China

    ARTICLE Received 29 Jul 2016 | Accepted 15 Feb 2017 | Published 9 May 2017 DOI: 10.1038/ncomms14952 OPEN Perinate and eggs of a giant caenagnathid dinosaur from the Late Cretaceous of central China Hanyong Pu1, Darla K. Zelenitsky2, Junchang Lu¨3, Philip J. Currie4, Kenneth Carpenter5,LiXu1, Eva B. Koppelhus4, Songhai Jia1, Le Xiao1, Huali Chuang1, Tianran Li1, Martin Kundra´t6 & Caizhi Shen3 The abundance of dinosaur eggs in Upper Cretaceous strata of Henan Province, China led to the collection and export of countless such fossils. One of these specimens, recently repatriated to China, is a partial clutch of large dinosaur eggs (Macroelongatoolithus) with a closely associated small theropod skeleton. Here we identify the specimen as an embryo and eggs of a new, large caenagnathid oviraptorosaur, Beibeilong sinensis. This specimen is the first known association between skeletal remains and eggs of caenagnathids. Caenagnathids and oviraptorids share similarities in their eggs and clutches, although the eggs of Beibeilong are significantly larger than those of oviraptorids and indicate an adult body size comparable to a gigantic caenagnathid. An abundance of Macroelongatoolithus eggs reported from Asia and North America contrasts with the dearth of giant caenagnathid skeletal remains. Regardless, the large caenagnathid-Macroelongatoolithus association revealed here suggests these dinosaurs were relatively common during the early Late Cretaceous. 1 Henan Geological Museum, Zhengzhou 450016, China. 2 Department of Geoscience, University of Calgary, Calgary, Alberta, Canada T2N 1N4. 3 Institute of Geology, Chinese Academy of Geological Sciences, Beijing 100037, China. 4 Department of Biological Sciences, University of Alberta, Edmonton, Alberta, Canada T6G 2E9. 5 Prehistoric Museum, Utah State University, 155 East Main Street, Price, Utah 84501, USA.
  • A Troodontid Dinosaur from the Latest Cretaceous of India

    A Troodontid Dinosaur from the Latest Cretaceous of India

    ARTICLE Received 14 Dec 2012 | Accepted 7 Mar 2013 | Published 16 Apr 2013 DOI: 10.1038/ncomms2716 A troodontid dinosaur from the latest Cretaceous of India A. Goswami1,2, G.V.R. Prasad3, O. Verma4, J.J. Flynn5 & R.B.J. Benson6 Troodontid dinosaurs share a close ancestry with birds and were distributed widely across Laurasia during the Cretaceous. Hundreds of occurrences of troodontid bones, and their highly distinctive teeth, are known from North America, Europe and Asia. Thus far, however, they remain unknown from Gondwanan landmasses. Here we report the discovery of a troodontid tooth from the uppermost Cretaceous Kallamedu Formation in the Cauvery Basin of South India. This is the first Gondwanan record for troodontids, extending their geographic range by nearly 10,000 km, and representing the first confirmed non-avian tetanuran dinosaur from the Indian subcontinent. This small-bodied maniraptoran dinosaur is an unexpected and distinctly ‘Laurasian’ component of an otherwise typical ‘Gondwanan’ tetrapod assemblage, including notosuchian crocodiles, abelisauroid dinosaurs and gondwanathere mammals. This discovery raises the question of whether troodontids dispersed to India from Laurasia in the Late Cretaceous, or whether a broader Gondwanan distribution of troodontids remains to be discovered. 1 Department of Genetics, Evolution, and Environment, University College London, London WC1E 6BT, UK. 2 Department of Earth Sciences, University College London, London WC1E 6BT, UK. 3 Department of Geology, Centre for Advanced Studies, University of Delhi, New Delhi 110 007, India. 4 Geology Discipline Group, School of Sciences, Indira Gandhi National Open University, New Delhi 110 068, India. 5 Division of Paleontology and Richard Gilder Graduate School, American Museum of Natural History, New York, New York 10024, USA.
  • Abstract Book.Pdf

    Abstract Book.Pdf

    Welcome! Welcome to the VI Symposium on Dinosaur Eggs and Babies, the return of this periodic gathering to the Iberian Peninsula, when it hatched eighteen years ago. From the slopes of the Pyrenees, we have followed the first steps of dinosaurs through France, Argentina, the United States and China. Today, we come back and see the coast where the first theropod embryos were discovered twenty years ago. Since the end of the last century, Paleoology, much like other branches of palaeontology, has evolved thanks to the advance of new methodologies and analytical tools, becoming a progressively more interdisciplinary area of knowledge. Dinosaur babies and embryos, rare findings back when these meetings started, seem to be everywhere now that we learn to look for them under the light of the microscope. New astonishing specimens allow us to understand how Mesozoic dinosaurs mate and reproduce. Oology, our parent discipline in the modern world, has made great advances in understanding the form and function of the egg, and its applications on poultry industry are countless. More than thirty contributions evidence that our field remains small but alive and healthy. We hope that you find in this Symposium an opportunity to share knowledge and open new lines of collaboration. And do not forget to enjoy your stay in Portugal. The host committee CONTENTS How to get to the FCT 6 Acknowledgements 10 PROGRAM 11 ABSTRACTS 14 THE FIRST ORNITHOMIMID EMBRYO IN A SHELL WITH A SINGLE STRUCTURAL LAYER: A CHALLENGE TO ORTHODOXY 15 Araújo R., Lamb J., Atkinson P., Martins R. M. S., Polcyn M.J., Fernandez V.
  • Implications for Predatory Dinosaur Macroecology and Ontogeny in Later Late Cretaceous Asiamerica

    Implications for Predatory Dinosaur Macroecology and Ontogeny in Later Late Cretaceous Asiamerica

    Canadian Journal of Earth Sciences Theropod Guild Structure and the Tyrannosaurid Niche Assimilation Hypothesis: Implications for Predatory Dinosaur Macroecology and Ontogeny in later Late Cretaceous Asiamerica Journal: Canadian Journal of Earth Sciences Manuscript ID cjes-2020-0174.R1 Manuscript Type: Article Date Submitted by the 04-Jan-2021 Author: Complete List of Authors: Holtz, Thomas; University of Maryland at College Park, Department of Geology; NationalDraft Museum of Natural History, Department of Geology Keyword: Dinosaur, Ontogeny, Theropod, Paleocology, Mesozoic, Tyrannosauridae Is the invited manuscript for consideration in a Special Tribute to Dale Russell Issue? : © The Author(s) or their Institution(s) Page 1 of 91 Canadian Journal of Earth Sciences 1 Theropod Guild Structure and the Tyrannosaurid Niche Assimilation Hypothesis: 2 Implications for Predatory Dinosaur Macroecology and Ontogeny in later Late Cretaceous 3 Asiamerica 4 5 6 Thomas R. Holtz, Jr. 7 8 Department of Geology, University of Maryland, College Park, MD 20742 USA 9 Department of Paleobiology, National Museum of Natural History, Washington, DC 20013 USA 10 Email address: [email protected] 11 ORCID: 0000-0002-2906-4900 Draft 12 13 Thomas R. Holtz, Jr. 14 Department of Geology 15 8000 Regents Drive 16 University of Maryland 17 College Park, MD 20742 18 USA 19 Phone: 1-301-405-4084 20 Fax: 1-301-314-9661 21 Email address: [email protected] 22 23 1 © The Author(s) or their Institution(s) Canadian Journal of Earth Sciences Page 2 of 91 24 ABSTRACT 25 Well-sampled dinosaur communities from the Jurassic through the early Late Cretaceous show 26 greater taxonomic diversity among larger (>50kg) theropod taxa than communities of the 27 Campano-Maastrichtian, particularly to those of eastern/central Asia and Laramidia.
  • Abelisaurid Pedal Unguals from the Late Cretaceous of India

    Abelisaurid Pedal Unguals from the Late Cretaceous of India

    AsociaciónPaleontológicaArgentina.PublicaciónEspecial7 ISSN0328-347X VIIInternationalSymposiumon MesozoicTerrestrialEcosystems:145-149Buenos. Aires,30-6-2001 AbeliSaurid Pedal unguals from the Late Cretaceous of India Fernando E. NOVAS!and Saswati BANDYOPADHYAY2 Abstraet. The ungual phalanges of theropod dinosaurs discovered in the Lameta Formation (Maastrichtian),centralIndia,exhibitdistinctivecharactersunknownin othertheropods.Hence,their tax- onomic identificationremained obscure since their original description in 1933.Recent discoveriesof abelisauridtheropods in Patagoniaindicate that the phalangesfromIndia belong to the foot of the~e~i- nosaurs. New informationon the foot skeletonof this group of CretaceousGondwanan predators lS m- cluded herein. Key words. Theropoda. Abelisauridae. Unguals. Cretaceous. Gondwana. IntroducTion Xenotarsosaurus; Bonaparte and Novas, 1985; Martínez et al., 1986; Bonaparte et al., 1990), India The Late Cretaceous (Maastrichtian) Lameta ilndosuchus; Chatterjee and Rudra, 1996), and Formation of central India has yielded dissociated el- Madagascar (Majungatholus; Sampson et al., 1998). ements of a variety of predatory dinosaurs. The ma- This considerable amount of information about terials were described by Frederich von Huene (in abelisaurid anatomy has served as a valuable guide Huene and Matley, 1933), who recognized nine to resolve confusing taxonomic aspects of the Indian theropod species from a fossil quarry at Bara Simla theropods. Hill (Madhya Pradesh, India). They are: lndosuchus Currently, lndosuchus, lndosaurus, and also rapiorius Huene, Indosaurus matleyi Huene, Compso- Laeoisuchue, are interpreted as members of suchus solus Huene, Laeoisuchus indicus Huene, Abelisauridae (Bonaparte and Novas, 1985; Molnar, [ubbulpuria tenuis Huene, Coeluroides largus Huene, 1990; Chatterjee and Rudra, 1996; Sampson et al., Dryptosauroides grandis Huene, Ornithomimoides mo- 1998; Novas and Bandyopadhyay, 1999). Moreover, bilis Huene, and Ornithomimoides (?) barasimlensis the controversial taxa "Compsosuchus", "Drypto- Huene.
  • Cáscaras De Huevos De Dinosaurios De La Formación Allen (Campaniano-Maastrichtiano), En Salitral Moreno, Provincia De Río Negro, Argentina

    Cáscaras De Huevos De Dinosaurios De La Formación Allen (Campaniano-Maastrichtiano), En Salitral Moreno, Provincia De Río Negro, Argentina

    AMEGHINIANA (Rev. Asoc. Paleontol. Argent.) - 43 (3): 513-528. Buenos Aires, 30-9-2006 ISSN 0002-7014 Cáscaras de huevos de dinosaurios de la Formación Allen (Campaniano-Maastrichtiano), en Salitral Moreno, provincia de Río Negro, Argentina María Edith SIMÓN1 Abstract. DINOSAUR EGGSHELLS FROM THE ALLEN FORMATION (CAMPANIAN-MAASTRICHTIAN), AT SALITRAL MORENO, RÍO NEGRO PROVINCE, ARGENTINA. A detailed description of two morphotypes of dinosaur eggshells from Salitral Moreno, Rio Negro Province (Allen Formation, Upper Cretaceous) is presented. One of them is described as Patagoolithus salitralensis, a new oogenus et oospecies nov. belonging to the Megaloolithidae, and the second is referred as ?Faveoloolithidae, oogenus et oospecies indet. A detailed ultrastructural description of both morphotypes is provided, as well as, the chemical and mineralogical analysis of ?Faveoloolithidae oogen. et oospec. indet. In the studied sections, three eggshell-bearing levels were recognized. In the lower one (NH1) Patagoolithus salitralensis is associated to a diverse fauna of di- nosaurs, chelonians and invertebrates, and the few shell fragments are dispersed, suggesting deposition in an alluvial plain or disintegrated eggs. In NH2 and NH3, only eggshells of ?Faveoloolithidae oogenus et oospecies indet. were found. NH2 shows lateral variations in the inferred degree of transport and den- sity of the shell fragments in different sections. NH3 bears small shell fragments of the same morphotype decreasing in number towards the top. NH1 and NH3 bear rizolithes and alveolar structures, characteris- tic of calcretes, which could be indicative of palaeosoils. Resumen. En el presente trabajo se describen en detalle dos morfotipos de cáscaras de huevos de di- nosaurios hallados en Salitral Moreno (Formación Allen, Cretácico Superior), en la provincia de Río Negro.
  • Dinosaur Species List E to M

    Dinosaur Species List E to M

    Dinosaur Species List E to M E F G • Echinodon becklesii • Fabrosaurus australis • Gallimimus bullatus • Edmarka rex • Frenguellisaurus • Garudimimus brevipes • Edmontonia longiceps ischigualastensis • Gasosaurus constructus • Edmontonia rugosidens • Fulengia youngi • Gasparinisaura • Edmontosaurus annectens • Fulgurotherium australe cincosaltensis • Edmontosaurus regalis • Genusaurus sisteronis • Edmontosaurus • Genyodectes serus saskatchewanensis • Geranosaurus atavus • Einiosaurus procurvicornis • Gigantosaurus africanus • Elaphrosaurus bambergi • Giganotosaurus carolinii • Elaphrosaurus gautieri • Gigantosaurus dixeyi • Elaphrosaurus iguidiensis • Gigantosaurus megalonyx • Elmisaurus elegans • Gigantosaurus robustus • Elmisaurus rarus • Gigantoscelus • Elopteryx nopcsai molengraaffi • Elosaurus parvus • Gilmoreosaurus • Emausaurus ernsti mongoliensis • Embasaurus minax • Giraffotitan altithorax • Enigmosaurus • Gongbusaurus shiyii mongoliensis • Gongbusaurus • Eoceratops canadensis wucaiwanensis • Eoraptor lunensis • Gorgosaurus lancensis • Epachthosaurus sciuttoi • Gorgosaurus lancinator • Epanterias amplexus • Gorgosaurus libratus • Erectopus sauvagei • "Gorgosaurus" novojilovi • Erectopus superbus • Gorgosaurus sternbergi • Erlikosaurus andrewsi • Goyocephale lattimorei • Eucamerotus foxi • Gravitholus albertae • Eucercosaurus • Gresslyosaurus ingens tanyspondylus • Gresslyosaurus robustus • Eucnemesaurus fortis • Gresslyosaurus torgeri • Euhelopus zdanskyi • Gryponyx africanus • Euoplocephalus tutus • Gryponyx taylori • Euronychodon
  • Late Cretaceous) Record of Ornithischia from Africa Matthew .C Lamanna University of Pennsylvania

    Late Cretaceous) Record of Ornithischia from Africa Matthew .C Lamanna University of Pennsylvania

    University of Pennsylvania ScholarlyCommons Departmental Papers (EES) Department of Earth and Environmental Science September 2004 From dinosaurs to dyrosaurids (Crocodyliformes): Removal of the post-Cenomanian (Late Cretaceous) record of Ornithischia from Africa Matthew .C Lamanna University of Pennsylvania Joshua B. Smith University of Pennsylvania, [email protected] Yousry S. Attia Egyptian Geological Survey and Mining Authority Peter Dodson University of Pennsylvania, [email protected] Follow this and additional works at: http://repository.upenn.edu/ees_papers Recommended Citation Lamanna, M. C., Smith, J. B., Attia, Y. S., & Dodson, P. (2004). From dinosaurs to dyrosaurids (Crocodyliformes): Removal of the post-Cenomanian (Late Cretaceous) record of Ornithischia from Africa . Retrieved from http://repository.upenn.edu/ees_papers/31 Copyright The ocS iety of Vertebrate Paleontology. Use for profit not allowed. Reprinted from: Journal of Vertebrate Paleontology, Volume 24, Issue 3, 2004, pages 764-768. Publisher URL: http://www.vertpaleo.org/ This paper is posted at ScholarlyCommons. http://repository.upenn.edu/ees_papers/31 For more information, please contact [email protected]. From dinosaurs to dyrosaurids (Crocodyliformes): Removal of the post- Cenomanian (Late Cretaceous) record of Ornithischia from Africa Abstract Ornithischian dinosaurs are uncommon elements in Late Cretaceous faunal assemblages of many Gondwanan landmasses, particularly Africa. The best-documented post-Cenomanian record of purported ornithischian body fossils from Africa consists of a left umeh rus, with associated cranial and costal fragments, from the Santonian-Campanian Quseir Formation of Kharga Oasis, Egypt (Fig. 1 ) (Awad and Ghobrial, 1966). We show that this specimen pertains instead to a dyrosaurid crocodyliform, and restrict known African ornithischian body fossils to pre-Turonian sediments.