Elephant Evolution
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Open Thesis Final V2.Pdf
The Pennsylvania State University The Graduate School Department of the Geosciences TAXONOMIC AND ECOLOGIC IMPLICATIONS OF MAMMOTH MOLAR MORPHOLOGY AS MEASURED VIA COMPUTED TOMOGRAPHY A Thesis in Geosciences by Gregory J Smith 2015 Gregory J Smith Submitted in Partial Fulfillment of the Requirements for the Degree of Master of Science August 2015 ii The thesis of Gregory J Smith was reviewed and approved* by the following: Russell W. Graham EMS Museum Director and Professor of the Geosciences Thesis Advisor Mark Patzkowsky Professor of the Geosciences Eric Post Director of the Polar Center and Professor of Biology Timothy Ryan Associate Professor of Anthropology and Information Sciences and Technology Michael Arthur Professor of the Geosciences Interim Associate Head for Graduate Programs and Research *Signatures are on file in the Graduate School iii ABSTRACT Two Late Pleistocene species of Mammuthus, M. columbi and M. primigenius, prove difficult to identify on the basis of their third molar (M3) morphology alone due to the effects of dental wear. A newly-erupted, relatively unworn M3 exhibits drastically different characters than that tooth would after a lifetime of wear. On a highly-worn molar, the lophs that comprise the occlusal surface are more broadly spaced and the enamel ridges thicken in comparison to these respective characters on an unworn molar. Since Mammuthus taxonomy depends on the lamellar frequency (# of lophs/decimeter of occlusal surface) and enamel thickness of the third molar, given the effects of wear it becomes apparent that these taxonomic characters are variable throughout the tooth’s life. Therefore, employing static taxonomic identifications that are based on dynamic attributes is a fundamentally flawed practice. -
1.1 První Chobotnatci 5 1.2 Plesielephantiformes 5 1.3 Elephantiformes 6 1.3.1 Mammutida 6 1.3.2 Elephantida 7 1.3.3 Elephantoidea 7 2
MASARYKOVA UNIVERZITA PŘÍRODOVĚDECKÁ FAKULTA ÚSTAV GEOLOGICKÝCH VĚD Jakub Březina Rešerše k bakalářské práci Využití mikrostruktur klů neogenních chobotnatců na příkladu rodu Zygolophodon Vedoucí práce: doc. Mgr. Martin Ivanov, Dr. Brno 2012 OBSAH 1. Současný pohled na evoluci chobotnatců 3 1.1 První chobotnatci 5 1.2 Plesielephantiformes 5 1.3 Elephantiformes 6 1.3.1 Mammutida 6 1.3.2 Elephantida 7 1.3.3 Elephantoidea 7 2. Kly chobotnatců a jejich mikrostruktura 9 2.1 Přírůstky v klech chobotnatců 11 2.1.1 Využití přírůstků v klech chobotnatců 11 2.2 Schregerův vzor 12 2.2.1 Stavba Schregerova vzoru 12 2.2.2 Využití Schregerova vzoru 12 2.3 Dentinové kanálky 15 3 Sedimenty s nálezy savců v okolí Mikulova 16 3.1 Baden 17 3.2 Pannon a Pont 18 1. Současný pohled na evoluci chobotnatců Současná systematika chobotnatců není kompletně odvozena od jejich fylogeneze, rekonstruované pomocí kladistických metod. Diskutované skupiny tak mnohdy nepředstavují monofyletické skupiny. Přestože jsou taxonomické kategorie matoucí (např. Laurin 2005), jsem do jisté míry nucen je používat. Některým skupinám úrovně stále přiřazeny nebyly a zde této skutečnosti není přisuzován žádný význam. V této rešerši jsem se zaměřil hlavně na poznatky, které následovaly po vydání knihy; The Proboscidea: Evolution and Paleoecology of Elephants and Their Relatives, od Shoshaniho a Tassyho (1996). Chobotnatci jsou součástí skupiny Tethytheria společně s anthracobunidy, sirénami a desmostylidy (Shoshani 1998; Shoshani & Tassy 1996; 2005; Gheerbrant & Tassy 2009). Základní klasifikace sestává ze dvou skupin. Ze skupiny Plesielephantiformes, do které patří čeledě Numidotheriidae, Barytheriidae a Deinotheridae a ze skupiny Elephantiformes, do které patří čeledě Palaeomastodontidae, Phiomiidae, Mammutida, Gomphotheriidae, tetralofodontní gomfotéria, Stegodontidae a Elephantidae (Shoshani & Marchant 2001; Shoshani & Tassy 2005; Gheerbrant & Tassy 2009). -
Mammoths and Mastodons
AMERI AN MU E.UM OF ATU R I HI 1 R Mammoths and Mastodons By W. D. MATTHEW . THE. AMERICAN MASTODON Model by Charles R. Knight, based upon The Warren Mastodon skeleton in the American Museum of Natural History No. 43 Of THE GUIDE LEAFLET SERIES.-NOVE.MBER, 1915. Aft, O.~born THE WARREN MASTODON SKELETON IN THE AMERICAN MUSEUM . Mammoths and Mastodons A guide to th collections of fossil proboscideans in the Ameri an Museum of Natural History By W. D. MATTHEW 0 TE.i. T Pag 1. L ~TROD T RY. Di tribulion. Early Di coYerie . .............. ~. THE ExTL -cT ELEPHA_~T . The tru mammoth-~ la kan mamm th - k 1 t n from Indiana- ize of the mammoth-th Columbian 1 - phant- th Imperial lephant-extin t 011 World elephant - Plio n and Plei tocene elephant ' of Inclia-e,·olution f lephant from n1a to don ......................... .. ............... .. ..... ... 3. THE ... :\IERICAX :MA TODOX. Teeth f the ma tod n-habit an 1 en Yironment-the w·arren ma t don-male and femal kull -di tribu- tion of the ... merican ma todon . 1 z ..J.. THE LATER TERTB.RY 11A TODOX . The two-tu ked mat don Dibelodon-the long-jawed ma. t don Tetralophodon-the b aked ma todon Rhyncotherium-the primitiYe four tu k d ma tod n . Trilophod n-the Dinotherium.. 1.5 THE EARLY TERTL\.RY AxcE TOR OF THE 11A TODOX . Palreoma tod n - M reritherium-character. and affinitie . I 6. THE E'.'OL TIOX OF THE PRoBo CIDEA. D ubtful po ition of :i\I rither ium-Palreoma todon a primiti, prob cidean-Dinoth rium an aberrant ide-branch-Tril phodon de.:,cended from Palreoma todon branching into everal phYla in ~Ii cene and Plioc ne- Dib lodon phylum in ~ ~ orth and outh America-~Ia tod n phylum-elephant phylum-origin and di per al of th probo cidea and th ir proo·re ,i,·e exti11ctio11 . -
Filling a Gap in the Proboscidean Fossil Record: a New Genus from The
Filling a gap in the proboscidean fossil record: a new genus from the Lutetian of Senegal Rodolphe Tabuce, Raphaël Sarr, Sylvain Adnet, Renaud Lebrun, Fabrice Lihoreau, Jeremy Martin, Bernard Sambou, Mustapha Thiam, Lionel Hautier To cite this version: Rodolphe Tabuce, Raphaël Sarr, Sylvain Adnet, Renaud Lebrun, Fabrice Lihoreau, et al.. Filling a gap in the proboscidean fossil record: a new genus from the Lutetian of Senegal. Journal of Paleontology, Paleontological Society, 2019, pp.1-9. 10.1017/jpa.2019.98. hal-02408861 HAL Id: hal-02408861 https://hal.archives-ouvertes.fr/hal-02408861 Submitted on 8 Dec 2020 HAL is a multi-disciplinary open access L’archive ouverte pluridisciplinaire HAL, est archive for the deposit and dissemination of sci- destinée au dépôt et à la diffusion de documents entific research documents, whether they are pub- scientifiques de niveau recherche, publiés ou non, lished or not. The documents may come from émanant des établissements d’enseignement et de teaching and research institutions in France or recherche français ou étrangers, des laboratoires abroad, or from public or private research centers. publics ou privés. 1 Filling a gap in the proboscidean fossil record: a new genus from 2 the Lutetian of Senegal 3 4 Rodolphe Tabuce1, Raphaël Sarr2, Sylvain Adnet1, Renaud Lebrun1, Fabrice Lihoreau1, Jeremy 5 E. Martin2, Bernard Sambou3, Mustapha Thiam3, and Lionel Hautier1 6 7 1Institut des Sciences de l’Evolution, UMR5554, CNRS, IRD, EPHE, Université de 8 Montpellier, Montpellier, France <[email protected]> 9 <[email protected]> <[email protected]> 10 <[email protected]> <[email protected] > 11 2Univ. -
Paleobiogeography of Trilophodont Gomphotheres (Mammalia: Proboscidea)
Revista Mexicana deTrilophodont Ciencias Geológicas, gomphotheres. v. 28, Anúm. reconstruction 2, 2011, p. applying235-244 DIVA (Dispersion-Vicariance Analysis) 235 Paleobiogeography of trilophodont gomphotheres (Mammalia: Proboscidea). A reconstruction applying DIVA (Dispersion-Vicariance Analysis) María Teresa Alberdi1,*, José Luis Prado2, Edgardo Ortiz-Jaureguizar3, Paula Posadas3, and Mariano Donato1 1 Departamento de Paleobiología, Museo Nacional de Ciencias Naturales, CSIC, José Gutiérrez Abascal 2, 28006, Madrid, España. 2 INCUAPA, Departamento de Arqueología, Universidad Nacional del Centro, Del Valle 5737, B7400JWI Olavarría, Argentina. 3 LASBE, Facultad de Ciencias Naturales y Museo, Universidad Nacional de La Plata, Paseo del Bosque S/Nº, B1900FWA La Plata, Argentina. * [email protected] ABSTRACT The objective of our paper was to analyze the distributional patterns of trilophodont gomphotheres, applying an event-based biogeographic method. We have attempted to interpret the biogeographical history of trilophodont gomphotheres in the context of the geological evolution of the continents they inhabited during the Cenozoic. To reconstruct this biogeographic history we used DIVA 1.1. This application resulted in an exact solution requiring three vicariant events, and 15 dispersal events, most of them (i.e., 14) occurring at terminal taxa. The single dispersal event at an internal node affected the common ancestor to Sinomastodon plus the clade Cuvieronius – Stegomastodon. A vicariant event took place which resulted in two isolated groups: (1) Amebelodontinae (Africa – Europe – Asia) and (2) Gomphotheriinae (North America). The Amebelodontinae clade was split by a second vicariant event into Archaeobelodon (Africa and Europe), and the ancestors of the remaining genera of the clade (Asia). In contrast, the Gomphotheriinae clade evolved mainly in North America. -
Foraging Ecology and Conservation Biology of African Elephants: Ecological and Evolutionary Perspectives on Elephant-Woody Plant Interactions in African Landscapes
Foraging ecology and conservation biology of African elephants: Ecological and evolutionary perspectives on elephant-woody plant interactions in African landscapes Item Type Thesis Authors Dudley, Joseph Paine Download date 27/09/2021 15:01:40 Link to Item http://hdl.handle.net/11122/9523 INFORMATION TO USERS This manuscript has been reproduced from the microfilm master. UMI films the text directly from the original or copy submitted. Thus, some thesis and dissertation copies are in typewriter free, while others may be from any type of computer printer. The quality of this reproduction is dependent upon the quality of the copy submitted. Broken or indistinct print, colored or poor quality illustrations and photographs, print bleedthrough, substandard margins, and improper alignment can adversely affect reproduction. In the unlikely event that the author did not send UMI a complete manuscript and there are missing pages, these will be noted. Also, if unauthorized copyright material had to be removed, a note will indicate the deletion. Oversize materials (e.g., maps, drawings, charts) are reproduced by sectioning the original, beginning at the upper left-hand comer and continuing from left to right in equal sections with small overlaps. Each original is also photographed in one exposure and is included in reduced form at the back o f the book. Photographs included in the original manuscript have been reproduced xerographically in this copy. Higher quality 6” x 9” black and white photographic prints are available for any photographs or illustrations appearing in this copy for an additional charge. Contact UMI directly to order. UMI A Bell & Howell Information Company 300 North Zed) Road, Ann Arbor MI 48106-1346 USA 313/761-4700 800/521-0600 Reproduced with permission of the copyright owner. -
Manus Descriptions of an Undescribed Mastodon from the Latest Miocene-Earliest Pliocene Gray Fossil Site, with Comparisons to Other North American Proboscidean Taxa
East Tennessee State University Digital Commons @ East Tennessee State University Electronic Theses and Dissertations Student Works 12-2019 Manus Descriptions of an Undescribed Mastodon from the Latest Miocene-Earliest Pliocene Gray Fossil Site, with Comparisons to other North American Proboscidean Taxa Brenna Hart-Farrar East Tennessee State University Follow this and additional works at: https://dc.etsu.edu/etd Part of the Geology Commons, and the Paleobiology Commons Recommended Citation Hart-Farrar, Brenna, "Manus Descriptions of an Undescribed Mastodon from the Latest Miocene-Earliest Pliocene Gray Fossil Site, with Comparisons to other North American Proboscidean Taxa" (2019). Electronic Theses and Dissertations. Paper 3680. https://dc.etsu.edu/etd/3680 This Thesis - unrestricted is brought to you for free and open access by the Student Works at Digital Commons @ East Tennessee State University. It has been accepted for inclusion in Electronic Theses and Dissertations by an authorized administrator of Digital Commons @ East Tennessee State University. For more information, please contact [email protected]. Manus Descriptions of an Undescribed Mastodon from the Latest Miocene-Earliest Pliocene Gray Fossil Site, with Comparisons to other North American Proboscidean Taxa _________________________________ A thesis presented to the faculty of the Department of Geosciences East Tennessee State University In partial fulfillment of the requirements for the degree Master of Science in Geosciences _________________________________ by Brenna J. Hart-Farrar December 2019 _________________________________ Steven C. Wallace, Chair Chris. Widga Blaine W. Schubert Keywords: Gray Fossil Site, Mammut, Mastodon, Morphology, Manus ABSTRACT Manus descriptions of an Undescribed Mastodon from the Latest Miocene-Earliest Pliocene Gray Fossil Site, with Comparisons to other North American Proboscidean Taxa by Brenna J. -
Resolving the Relationships of Paleocene Placental Mammals
Biol. Rev. (2015), pp. 000–000. 1 doi: 10.1111/brv.12242 Resolving the relationships of Paleocene placental mammals Thomas J. D. Halliday1,2,∗, Paul Upchurch1 and Anjali Goswami1,2 1Department of Earth Sciences, University College London, Gower Street, London WC1E 6BT, U.K. 2Department of Genetics, Evolution and Environment, University College London, Gower Street, London WC1E 6BT, U.K. ABSTRACT The ‘Age of Mammals’ began in the Paleocene epoch, the 10 million year interval immediately following the Cretaceous–Palaeogene mass extinction. The apparently rapid shift in mammalian ecomorphs from small, largely insectivorous forms to many small-to-large-bodied, diverse taxa has driven a hypothesis that the end-Cretaceous heralded an adaptive radiation in placental mammal evolution. However, the affinities of most Paleocene mammals have remained unresolved, despite significant advances in understanding the relationships of the extant orders, hindering efforts to reconstruct robustly the origin and early evolution of placental mammals. Here we present the largest cladistic analysis of Paleocene placentals to date, from a data matrix including 177 taxa (130 of which are Palaeogene) and 680 morphological characters. We improve the resolution of the relationships of several enigmatic Paleocene clades, including families of ‘condylarths’. Protungulatum is resolved as a stem eutherian, meaning that no crown-placental mammal unambiguously pre-dates the Cretaceous–Palaeogene boundary. Our results support an Atlantogenata–Boreoeutheria split at the root of crown Placentalia, the presence of phenacodontids as closest relatives of Perissodactyla, the validity of Euungulata, and the placement of Arctocyonidae close to Carnivora. Periptychidae and Pantodonta are resolved as sister taxa, Leptictida and Cimolestidae are found to be stem eutherians, and Hyopsodontidae is highly polyphyletic. -
Karyotype Determination of Rock Hyrax Procavia Capensis in Saudi Arabia
© 2015 The Japan Mendel Society Cytologia 80(3): 287–293 Karyotype Determination of Rock Hyrax Procavia capensis in Saudi Arabia Saud A. Al-Dakan and Abdulaziz A. Al-Saleh* Zoology Department, College of Science, King Saud University, P.O. Box 2455, Riyadh 11451, Saudi Arabia Received November 1, 2014; accepted May 31, 2015 Summary Procavia capensis is considered as a small mammalian animal which belongs to order Hyracoidea, and it is the only species of the order that has been found in Saudi Arabia. Therefore, karyotype analysis of this species has been carried out and the finding summarized as follows. The diploid chromosome number is 54. In the karyotype analysis, the somatic chromosomes were catego- rized into three groups: 21 pairs of acrocentric, 2 pairs of submetacentric and 3 pairs of metacentric chromosomes. The sex chromosomes are one submetacentric X chromosome and one acrocentric Y chromosome. The lengths of chromosomes varied between 1.6–7.6 µm, and the Y chromosome is the shortest. The FN is 65 in the male and 66 in the female, while the FNa is 62. The karyotype formula of Procavia capensis could be deduced as: a sm a sm a sm m (2=54);Ln 14+ L 1 + M 14 + M 2 ++ S 15 S 2 + S 6 Key words Karyotype, Rock hyrex, Chromosome, Procavia capensis. The rock hyrax (Procavia capensis) is one of small mammalian herbovorous animals that lives in small family groups ranging from 10 to 80 members headed by a dominant adult male which defends and watches over the group (Turner and Watson 1965, Grzimek 1975, Skinner and Smithers 1990, Estes 1991, Kingdon 1991, Manharth and Harris-Gerber 2002). -
City's Limits: Can Verreaux's Eagle Survive Urbanization?
See discussions, stats, and author profiles for this publication at: https://www.researchgate.net/publication/289670312 City's limits: Can Verreaux's Eagle survive urbanization? Article · July 2007 CITATIONS READS 0 12 4 authors, including: Robert Simmons University of Cape Town 117 PUBLICATIONS 1,378 CITATIONS SEE PROFILE Some of the authors of this publication are also working on these related projects: Climate change View project Namibian Red data birds View project All content following this page was uploaded by Robert Simmons on 14 January 2016. The user has requested enhancement of the downloaded file. and bush encroachment, the raptors second pair live on the west side of the which need pristine woodlands rather Cape Peninsula mountain chain, in the than those that have been burnt and Silvermine area of the Table Mountain CITY’S logged, and some, such as the harriers National Park, just north of Chapman’s and the African Fish-Eagle, which live Peak. There they have bred successfully Can Verreaux’s Eagles survive urbanisation? around the fringes of degraded and for three years, monitored weekly by drained wetlands. their finder and eagle-advocate Lucia What of our montane eagles? Can Rodrigues. These pairs share similarities species, such as Verreaux’s (Black) Eagles and differences that allow us to assess Aquila verreauxii, which live along- if Verreaux’s Eagles can survive human side man, continue to thrive in their encroachment. mountain retreats? Two closely studied When the Roodekrans pair began Below An adult Verreaux’s Eagle arrives LIMITS examples, one from Johannesburg and to bring chickens to their nest, it was at its nest on the Roodekrans in the sub- the other from Cape Town, allow an apparent that the dassie (rock hyrax) urbs of Johannesburg. -
The African Elephant Under Threat
AFROTHERIAN CONSERVATION Newsletter of the IUCN/SSC Afrotheria Specialist Group Number 11 Edited by PJ Stephenson October 2015 Afrotherian Conservation is published annually by the To help PJ focus on our conservation work, Chris IUCN Species Survival Commission Afrotheria Specialist and Mathilde Stuart have kindly agreed to take on the Group to promote the exchange of news and information role of editing the next edition of Afrotherian Conservation. on the conservation of, and applied research into, golden Their contacts are in the guidelines for submissions (page moles, sengis, hyraxes, tenrecs and the aardvark. 17). We hope you’ll send them plenty of material for the next edition. Published by IUCN, Gland, Switzerland. © 2015 International Union for Conservation of Nature Galen Rathbun, Cambria, California, USA and Natural Resources & ISSN: 1664-6754 PJ Stephenson, Gland, Switzerland 1 October 2015 Find out more about the Group on our website at http://afrotheria.net/ASG.html and follow us on Twitter @Tweeting_Tenrec Message from the Chairs Galen Rathbun & PJ Stephenson Co-Chairs, IUCN/SSC Afrotheria Specialist Group It’s been a busy twelve months for the group. Sadly, 2015 started with the terrible news that Peter Vogel had passed away. Peter was a global expert on shrews but he was a long-term member of the group due to his specialist knowledge of otter-shrews; he was one of the few biologists to capture and study these illusive afrotheres. We include an obituary to Peter on page 8 and send our Chequered sengi (Rhynchocyon cirnei) by Jonathan Kingdon condolences to his family, friends and colleagues. -
Tour Report 7 – 25 November 2015
The Best of Ethiopia Naturetrek Tour Report 7 – 25 November 2015 African Fish Eagle Walia Ibex Ethiopian Wolf White-cheeked Turaco Report & images compiled by Eric Barnes Naturetrek Mingledown Barn Wolf's Lane Chawton Alton Hampshire GU34 3HJ UK T: +44 (0)1962 733051 E: [email protected] W: www.naturetrek.co.uk The Best of Ethiopia Tour Report Tour Participants: Abiy Dagne and Eric Barnes (leaders) with ten Naturetrek clients Introduction For most people, Ethiopia won’t be top of their list of places to visit for an enjoyable holiday, nor would they think of it as a wildlife hotspot. Those that take the trip, in my experience, have been universally pleased that they took a leap of faith and travelled to a wildlife paradise full of character and surprises. “Eric, this Lammergeier is far too close for us to photograph” and “Don’t stop, it’s only another Arabian Bustard” were comments from this year’s tour that, perhaps, give some idea of the great views and wealth of wildlife experiences. Unfortunately, it is unclear how long this situation will last. In the past two years there have been changes relating to uncontrolled grazing in their National Parks. It is clear that the wildlife will suffer severely over the next decade, so if you are thinking of going, go sooner rather than later! Day 1 Saturday 7th November In flight to Ethiopia. Day 2 Sunday 8th November Our Ethiopian Airline flight touched down as smooth as silk, slightly ahead of schedule. We got our visas, picked up our adjoining clients and headed off for a quick breakfast and wander around the Ghion Hotel.