A303 Amesbury to Berwick Down

A303 6.3 EnvironmentalStatementAppendices Appendix 8.11Invertebratesurveyreport Amesbury to Berwick Down to Berwick Amesbury Infrastructure Planning(A FormsandProcedur APFP Regulation5(2)(a) pplications: Prescribed e) Regulations2009 Planning Act2008 October2018 TR010025

Volume 16

Appendix 8.11 Invertebrate survey report

Arup Atkins Joint Venture HE551506-AA-EBD-SWI-SU-YE-000011 P01

November 2017

Notice This document and its contents have been prepared and are intended solely for Highways England’s information and use in relation to the A303 Amesbury to Berwick Down project. The Arup Atkins Joint Venture assumes no responsibility to any other party in respect of, arising out of or in connection with this document and/or its contents.

A303 Stonehenge - Amesbury to Berwick Down | HE551506 Arup Atkins Joint Venture

Document Control Document Title A303 Invertebrate Survey Report 2017 Document Reference HE551506-AA-EBD-SWI-SU-YE-000011 Author Various Document Status Final Issue to HE for Acceptance

Prepared for: Prepared by: Highways England Arup Atkins Joint Venture Temple Quay House The Hub, Aztec West 2 The Square, Temple Quay 500 Park Avenue Bristol Bristol BS1 6HA BS32 4RZ Revision History Revision Date Status Description Author

0 23/11/2017 S0 Draft Various P01 23/11/2017 S0 Draft Various

Arup Atkins Joint Venture Approvals Revision Role Name Signature Date Author Dave Boyce - 23/11/2017

Technical Checker Chloe Delgery 27/11/2017

Technical Reviewer Paul Clack 30/11/2017 P01

Checker Liz Brown 15/01/17

Approver Andy Keen 15/01/17

Author Dave Boyce - 23/11/2017

0 Technical Checker Chloe Delgery 27/11/2017

Technical Reviewer Paul Clack 30/11/2017

Highways England Reviewers Revision Title Name Signature Date A303 Stonehenge - Amesbury to Berwick Down | HE551506 Arup Atkins Joint Venture

Highways England Approval Revision Title Name Signature Date A303 Stonehenge - Amesbury to Berwick Down | HE551506 Arup Atkins Joint Venture

Table of contents

Chapter Pages Foreword 1 Executive Summary 2 1 Introduction 3 1.1 Project Background 3 1.2 Scope of the Document 3 Annexes i Annex 1 - 2017 Invertebrate Survey report ii

A303 Stonehenge - Amesbury to Berwick Down | HE551506 Arup Atkins Joint Venture

Foreword

The A303/A358 corridor is a vital connection between the South West and London and the South East. While the majority of the road has been dualled, there are still over 35 miles of single carriageway. These sections act as bottlenecks for users of the route resulting in congestion, particularly in the summer months and at weekends, delays to traffic travelling between the M3 and the South West and an increased risk of accidents. The A303 passes through the Stonehenge, Avebury and Associated Sites World Heritage Site, separating the stones from other scheduled monuments and severely limiting the enjoyment of the wider site.

The A303 Stonehenge (Amesbury to Berwick Down) scheme is part of the wider package of proposals for the A303/A358 corridor designed to transform the connectivity to and from the South West by creating an expressway. This would comprise of consistently good dual carriageway roads with grade-separated junctions, giving most users a motorway-quality journey. The A303/A358 package was identified in the 2014 National Infrastructure Plan as one of the country’s Top 40 priority infrastructure projects.

The proposals by Highways England to upgrade the A303 past Stonehenge consist of an eight mile (13 kilometre) stretch from Amesbury in the east, through the Stonehenge World Heritage Site (WHS) and the village of Winterbourne Stoke, to Berwick Down in the west. Proposals include a 1.8 mile (2.9 kilometre) tunnel with approach roads inside the WHS, a new bypass for Winterbourne Stoke (passing either north or south of the village) and improvements to existing junctions with the A345 and A360.

Highways England (HE) commissioned the Arup-Atkins Joint Venture (AAJV) to undertake the Options Phase for the scheme starting in January 2016. The AAJV was also commissioned by HE to undertake a terrestrial invertebrate survey relevant to these proposed route options, in order to de-risk the next stages of the project, due to the fast- tracked nature of the scheme. This report presents the findings of the terrestrial invertebrate surveys, which were undertaken by invertebrate specialist Dave Boyce on behalf of the AAJV. The AAJV and Dave Boyce would like to thank the landowners for their help and consideration during the course of the surveys.

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A303 Stonehenge - Amesbury to Berwick Down | HE551506 Arup Atkins Joint Venture

Executive Summary

The AAJV were commissioned by Highways England to undertake terrestrial invertebrate surveys as part of a programme of ecological surveys to inform the design of the proposed A303 Stonehenge (Amesbury to Berwick Down) Scheme.

This report presents the baseline survey results recorded during the 2017 terrestrial invertebrate surveys relevant to each of the three route options proposed at the time (1Na, 1Sa and 1Nd). It is intended that the information in this report will be used to identify and assess the potential implications of the Scheme and inform mitigation and compensation for terrestrial invertebrates.

A framework of European and national legislation, and planning policy guidance exists to protect and conserve invertebrates.

All eight sites included in the 2017 survey have been shown to have a significant invertebrate fauna. Unsurprisingly, the three sites with high-quality habitats notified as SSSIs and SACs all have important invertebrate assemblages: the two River Till sites having diverse wetland faunas and Parsonage Down NNR with an assemblage that is associated with short-sward chalk grassland. Countess Cutting CWS and Arable 1 sample sites also have an exceptionally rich invertebrate fauna found in association with early-to- mid-successional chalk grassland habitats and wide arable margins respectively. That found on the wide arable margins at the Arable 1 site was particularly noteworthy and demonstrates that the adoption of conservation headlands here is certainly having benefits for invertebrates.

Diamond Wood and the Arable 2 and Arable 3 samples sites are somewhat less diverse, but still have important invertebrate assemblages.

Forty two key invertebrate species were collected during the 2017 sampling programme. Three of these: the Leiodid beetle Ptomaphagus varicornis, the longhorn beetle Paracorymbia fulva and the silver-sided nomad bee Nomada argentata are Red Data Book species, and the Gnaphosid spider Phaeocedus braccatus is a Nationally Rare species that has also been assigned IUCN Vulnerable threat status.

These surveys fulfilled their objectives in recording the invertebrate assemblages across the route options. A complete assessment of potential impacts to invertebrates and their habitats will be undertaken within the Environmental Impact Assessment (EIA) for the preferred route option, along with details of mitigation and compensation measures as appropriate.

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A303 Stonehenge - Amesbury to Berwick Down | HE551506 Arup Atkins Joint Venture

1 Introduction 1.1 Project Background The A303 Stonehenge (Amesbury to Berwick Down) Scheme forms part of the A303/A30 trunk route, which provides vital east-west connectivity between London and the South West and is also part of the Trans-European Network- Transport (TEN-T). The A303, which runs for approximately 150 kilometres from Junction 8 of the M3 near Basingstoke towards Taunton and Exeter, serves not only long distance traffic but also intermediate regional destinations via connecting major north-south route options as well as local small and medium sized settlements along the route.

Recognising the importance of the A303/A358 Corridor and the problems along it, the Government has committed in its Road Investment Strategy (RIS) to create an 'Expressway' to the South West via the A303/A358 route by 2029. The A303 Stonehenge scheme, involving dualling the A303 between Amesbury and Berwick Down, including the construction of a tunnel at least 1.8 miles (2.9 kilometres) long as the road passes Stonehenge, has been prioritised within the first RIS period (2015/16 to 2019/20).

Following public consultation in January 2017, three routes were recommended for detailed assessment during 2017, Route Options 1Na, 1Sa and 1Nd. 1.2 Scope of the Document This report presents the baseline survey results recorded during the 2017 terrestrial invertebrate surveys. It is intended that the information in this report will be used with the results of other ecological surveys to identify and assess the potential implications of the scheme and inform mitigation and compensation for impacts to terrestrial invertebrates.

This baseline report can be used to accompany any future Development Consent Order (DCO) and associated Environmental Impact Assessment (EIA) for the Scheme.

Dave Boyce, invertebrate specialist, was commissioned to undertake these surveys on behalf of the AAJV. The full survey report which details the methodology used and describes the results of the 2017 invertebrate surveys can be found in Annex 1 of this report.

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A303 Stonehenge - Amesbury to Berwick Down | HE551506 Arup Atkins Joint Venture

Annexes

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A303 Stonehenge - Amesbury to Berwick Down | HE551506 Arup Atkins Joint Venture

Annex 1 - 2017 Invertebrate Survey report

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A303 Stonehenge Amesbury to Berwick Down Invertebrate Survey, November 2017

DC Boyce Ecologist November 2017

CONTENTS 1. INTRODUCTION ...... 1 2. METHODS ...... 1 2.1. Fieldwork ...... 1 2.2. Identification of material, data collation and report writing ...... 5 3. IMPORTANT INVERTEBRATES RECORDED ALONG THE A303 AMESBURY TO BERWICK DOWN SCHEME IN 2017 ...... 6 3.1. Key invertebrates ...... 6 3.1.1. A Gnaphosid Spider Phaeocedus braccatus (L. Koch, 1866). VU./NR...... 7 3.1.2. A Diving Beetle Hydroporus ferrugineus Stephens, 1829. NS...... 7 3.1.3. A Diving Beetle Hydroporus marginatus (Duftschmid, 1805). NS...... 7 3.1.4. A Ground Beetle Zabrus tenebrioides (Goeze, 1777). NS...... 7 3.1.5. A Ground Beetle Amara curta Dejean, 1828. NS...... 8 3.1.6. A Ground Beetle Ophonus schaubergerianus (Puel, 1937). NS...... 8 3.1.7. A Ground Beetle Licinus depressus (Paykull, 1790). NS...... 8 3.1.8. A Ground Beetle Licinus punctatulus (Fabricius, 1792). Na...... 8 3.1.9. A Leiodid Beetle Ptomaphagus varicornis (Rosenhauer, 1847). RDBK...... 9 3.1.10. A Sexton Beetle Nicrophorus interruptus Stephens, 1830. Nb...... 9 3.1.11. A Rove Beetle Sepedophilus testaceus (Fabricius, 1792). N...... 9 3.1.12. A Rove Beetle Lamprinodes saginatus (Gravenhorst, 1806). Na...... 9 3.1.13. A Rove Beetle Oxypoda exoleta Erichson, 1839. N...... 10 3.1.14. A Rove Beetle Gnypeta ripicola (Kiesenwetter, 1844). N...... 10 3.1.15. A Rove Beetle Stenus butrintensis Smetana, 1959. N...... 10 3.1.16. A Rove Beetle Sunius melanocephalus (Fabricius, 1792). N...... 10 3.1.17. A Rove Beetle Rugilus similis (Erichson, 1839). N...... 11 3.1.18. A Rove Beetle Philonthus atratus (Gravenhorst, 1802). Na...... 11 3.1.19. A Rove Beetle Philonthus parvicornis (Gravenhorst, 1802). pNS...... 11 3.1.20. A Rove Beetle Quedius nigrocaeruleus Fauvel, 1874. Nb...... 12 3.1.21. A Woodworm Beetle Anobium inexspectatum (Lohse, 1954). Nb...... 12 3.1.22. A Pollen Beetle Meligethes solidus (Kugelann, 1794). N...... 12 3.1.23. A Phalacrid Beetle Phalacrus championi Guillebeau, 1892. Na...... 12 3.1.24. A Salpingid Beetle Lissodema denticolle (Kugelann, 1794). NS...... 13 3.1.25. A Longhorn Beetle Paracorymbia fulva (De Geer, 1775). RDB3...... 13 3.1.26. A Flea Beetle Psylliodes chalcomera (Illiger, 1807). NS...... 13 3.1.27. A Weevil Hemitrichapion reflexum (Gyllenhal, 1833). Na...... 13

3.1.28. A Weevil Oxystoma cerdo (Gerstaecker, 1854). Nb...... 14 3.1.29. A Weevil Catapion pubescens (Kirby, 1811). Nb...... 14 3.1.30. A Weevil Protapion filirostre (Kirby, 1808). Nb...... 14 3.1.31. A Weevil Gymnetron melanarium (Germar, 1821). Nb...... 14 3.1.32. A Weevil Gymnetron villosulum Gyllenhal, 1838. Nb...... 15 3.1.33. A Weevil Mecinus circulatus (Marsham, 1802). Nb...... 15 3.1.34. A Weevil Tychius pusillus Germar, 1842. Nb...... 15 3.1.35. A Weevil Drupenatus nasturtii (Germar, 1824). Nb...... 15 3.1.36. A Weevil Sirocalodes quercicola (Paykull, 1792). Na...... 16 3.1.37. A Weevil Thamiocolus viduatus (Gyllenhal, 1813). Nb...... 16 3.1.38. A Weevil Zacladus exiguus (Olivier, 1807). Nb...... 16 3.1.39. A Weevil Larinus planus (Fabricius, 1792). Nb...... 16 3.1.40. Small Scabious Mining Bee Andrena marginata Fabricius, 1776. Na. ... 17 3.1.41. Plain Mini-miner Bee Andrena minutuloides Perkins, R. C. L., 1914. Na...... 17 3.1.42. Silver-sided Nomad Bee Nomada argentata Herrich-Schäffer, 1839. RDB3...... 17 4. SITE REPORTS ...... 18 4.1. Upper Till ...... 18 4.1.1. Key Invertebrate habitat features at the Upper Till ...... 19 4.2. Lower Till ...... 21 4.2.1. Key Invertebrate habitat features at the Lower Till ...... 22 4.3. Parsonage Down NNR ...... 23 4.3.1. Key Invertebrate habitat features at Parsonage Down NNR ...... 24 4.3.2. Marsh fritillary survey ...... 24 4.4. Diamond Wood ...... 26 4.4.1. Key Invertebrate habitat features at Diamond Wood ...... 26 4.5. Arable 1 ...... 28 4.5.1. Key Invertebrate habitat features at Arable 1 ...... 29 4.6. Arable 2 ...... 30 4.6.1. Key Invertebrate habitat features at Arable 2 ...... 31 4.7. Arable 3 ...... 32 4.7.1. Key Invertebrate habitat features at Arable 3 ...... 33 4.8. Countess Cutting CWS ...... 34 4.8.1. Key Invertebrate habitat features at Countess Cutting CWS ...... 34 5. CONCLUSIONS ...... 35

6. ACKNOWLEDGEMENTS ...... 35 7. REFERENCES ...... 36

1. INTRODUCTION

This report details the findings of an invertebrate survey commissioned by the Arup Atkins Joint Venture (AAJV) on sites on or adjacent to the three proposed route Options being considered during 2017 for the A303 Amesbury to Berwick Down Improvement Scheme: Option 1Na, Option 1Sa and Option 1Nd (hereafter referred to as the A303 Amesbury to Berwick Down Scheme). Eight sample sites were selected by the AAJV where invertebrate surveys needed to be undertaken. These are listed in Table 2.1. with a central grid reference, dates surveyed and any other relevant details. More information on each of these areas can be found in the Site Reports at section 3.

Some of the sample sites lie partially or completely within areas that have formal nature conservation status. Both Sites of Special Scientific Interest (SSSI) and Special Areas of Conservation (SAC) are represented. The former status denotes the site to be part of a nationally important series for wildlife, while the latter indicates that the habitats are of international importance within a Europe-wide context. Where a sample site includes such designated areas, the name of the SSSI/SAC is given in Table 2.1 below and is also mentioned in the introductory paragraph to the relevant Site Report in section 3.

2. METHODS

2.1. Fieldwork Each of the sample sites was visited by David Boyce and a field assistant at least twice over the summer of 2017. In all, eleven days’ fieldwork were undertaken; the first visits being made over a five-day period from the 26th to the 30th of June and the second over six days from the 24th to 29th of July. Sites were defined by the AAJV on the basis of the central grid references given in Table 2.1, with invertebrate surveys being carried out in the area around these – usually within no more than a 500m radius from the sample point. The days on which each site was visited are also shown in Table 2.1. During the first visit to the site, a quick walk-over reconnaissance survey was undertaken to identify the distribution of those habitats thought likely to support the most interesting invertebrate assemblages.

These features are shown in the fourth column of Table 2.1 and have constituted the main habitats sampled at each of the sample sites.

At each sample site, a range of invertebrate sampling techniques was used. In general, the sampling strategy used at each site has followed the habitat-specific invertebrate sampling protocols advocated by Natural England for assessing the impact of development on terrestrial and freshwater invertebrate assemblages (Drake et. al., 2007).

Primarily, this involved the following hand-searching techniques:

• Beating - tapping living and dead branches of trees and shrubs over a beating tray. • Ground searching – shaking tussocks, moss and litter into a white tray; looking under stones and fallen wood; and treading substrates down on wetland sites then collecting invertebrates emerging from cover. • Pond netting and sieving – on the two River Till sample sites, where there were significant aquatic habitats, a pond net was used in larger areas of open water and a kitchen sieve in small waterbodies such as sprinheads and seepages, where it was impossible to sample effectively with a net. • Sweeping – using a heavy-duty entomological sweep net to sample longer vegetation. • Spot searching - targeted collecting of flying insects that are too alert to be sampled effectively by sweeping using a lightweight net. • Tapping – tapping the foliage and flowers of invdividul plant species over a white tray.

Table 2.1. Details of invertebrate sample sites, Amesbury to Berwick Down, 2017 Sample Site: Central Grid Ref: Dates Surveyed Important Habitats: (2017): Conservation Designations: Upper Till (north of SU080417 26/06; 24/07 River; pond; trees; grassland. A303) Part River Till SSSI & River Avon SAC. Lower Till (south of SU077399 26/06; 27/06; 25/07 River; ditches; fen-meadow; A303) wet woodland. Part River Till SSSI & River Avon SAC. Parsonage Down SU062411 28/06; 28/07; 29/07 Chalk grassland. National Nature All Parsonage Down SSSI & Reserve (NNR) Salisbury Plain SAC. Diamond Wood SU105408 29/06; 27/07 Chalk grassland; arable margin; wood-edge. Arable 1 (south of SU112413 29/06; 27/07 Arable margin. Normanton Gorse) Arable 2 (north of SU113417 27/06; 29/06; 26/07 Arable restoration grassland Normanton Gorse) Arable 3 (Bowtie Field) SU138421 30/06; 26/07 Arable margin. Countess Cutting SU145421 30/06; 26/07 Early-successional chalk County Wildlife Site grassland in cutting. (CWS)

Arable habitats do not lend themselves to effective invertebrate hand sampling of the types outlined above. For this reason, it was agreed with the AAJV that at the three sites where these habitats were the primary focus of invertebrate sampling effort (Arable 1, Arable 2 and Arable 3), pitfall trapping should be employed. Initially, it had been planned to establish three pitfall trapping stations at each of the three arable sample sites. However, the reconnaissance of the Arable 2 site carried out on 27 June 2017 showed this area now to largely comprise non-arable habitats,

2 principally a large stand of moderately species-rich calcareous grassland dominated by sainfoin Onobrychis viciifolia, which is being reverted from arable production. It was felt that this site was possible to sample at least as effectively for invertebrates using active searching techniques, and so only two pitfall stations were established here.

A single pitfall station was set up at Countess Cutting. This was a late addition to the sampling programme, which it was felt would benefit from pitfalls, as the sparsely vegetated chalk grassland here was also difficult to sample for ground-active invertebrates using standard hand-searching techniques.

Each pitfall trap comprises two stacked plastic picnic cups that were dug into the ground to a depth of approximately 10-12cm, such that the rim of the cup was flush with the surface of the ground. A pitfall sample site was made up of a line of five pitfall traps spaced at 2 metre intervals. Trap lines were sited along field margins and Rights Of Way that were less likely to be disturbed by farming operations. They were not set up in areas where farm stock had access (though cattle were moved onto the first pitfall station at the Arable 2 site (AR2/PF1) between the time when the traps were set and collected. Every trap was covered with chicken wire, to minimise egress by small mammals. The traps were set up during the week of June 26th and removed in the week of July 17th.

Each pitfall trap was filled to approximately one third depth with fluid. This comprised antifreeze, the chemical constituent of which is ethylene glycol, which was diluted by about two thirds. A few drops of washing-up liquid were also added to break the surface tension of the water. At the time that the traps were removed, the catch was emptied through a tea strainer and funnel, with waster fluid being collected into a large plastic bottle. The trap contents were then carefully tapped from the tea strainer into a wide-mouthed jar. For the purposes of this survey, the five sub-samples were amalgamated into a single collecting jar. Once all the five sub-samples had been collected, undiluted IPA alcohol was added to the jar till it covered the sample and the container lid was tightly affixed after adding a waterproof data label. Table 2.2 provides details of the pitfall trapping stations, including a description of their location and eight-figure grid references that were taken from the central trap. The pitfall trapping methodology described above exactly follows the protocols set out by Luff (1996), this being the standard methodology recommended by Natural England for invertebrate impact assessments (Drake et al., 2007).

At Parsonage Down, an additional day was spent looking for marsh fritillary larval webs at the end of the July visit. The methodology employed the standardised protocols used by Butterfly Conservation (BC) for monitoring larvae of this butterfly. A full description of the methodology can be found on the UK Butterfly Monitoring Scheme website (http://www.ukbms.org/).

Table 2.2. Details of pitfall trapping stations, A303 Amesbury to Berwick Down, 2017 Sample site Sample site NGR Description Arable 1 AR1/PF1 SU11424131 Line runs E.-W. from SE edge of Normanton Gorse along junction between open ruderal strip and more established grassland AR1/PF2 SU11184143 Line runs N. -S. from SW. edge of Normanton Gorse in strip of rank grassland between arable and wood AR1/PF3 SU11334161 Line runs N.-S. from NW. edge of Normanton Gorse along junction between open ruderal strip and more established grassland Arable 2 AR2/PF1 SU11234167 Line runs S. from isolated hawthorn in boundary fence parallel to this in ‘sainfoin reversion grassland’ AR2/PF2 SU11754168 Line runs N.-S. between red-marked posts marking boundary, in strip of rank grassland between byway and fence on E. side of track, c. 150m S. of A303 Arable 3 AR3/PF1 SU13604230 Line runs E.-W. between red-marked posts marking boundary, in narrow ruderal strip between arable crop and fence AR3/PF2 SU13594201 Line runs E.-W. between red-marked posts marking boundary, in narrow ruderal strip between arable crop and A303 fence AR3/PF3 SU13734201 Line runs E.-W. between red-marked posts marking boundary, in narrow ruderal strip between arable crop and A303 fence Countess Cutting CC/PF1 SU14514213 Line runs E.-W. at western end of open cutting through chalk with short- sward ruderal chalk grassland

2.2. Identification of material, data collation and report writing Invertebrate specimens collected have been identified to species level. Taxa collected have been those thought likely to be of greatest importance in providing a thorough assessment of the importance for invertebrates of the habitats listed in Table 2.1. Chief amongst these groups has been the beetles (Coleoptera), which include large numbers of specialists associated with the wetland, chalk grassland and arable habitats that have been the primary focus of sampling during this contract. Any records of invertebrates that it was possible to identify in the field (eg, dragonflies, butteflies, macro-moths and Orthoptera) have also been kept and reported on. A full list of all invertebrates collected from each of the eight sample sites in 2017 can be found at Appendix 1. The data has also been supplied separately to the AAJV as an Excel spreadsheet.

Sections 3 and 4 provide an assessment of the invertebrate fauna of the eight sites surveyed in 2017. Based on the survey, a list of key invertebrate species is given in Section 3. For each of these, a short summary of their conservation status, UK distribution, ecology and occurrence in the 2017 sample sites is provided. Key species are those that meet one or more of the following criteria:

1) receiving statutory protection under Schedule 5 of the Wildlife and Countryside Act, 1981,as subsequently amended); 1) included in EC Annex II of the Species Directive (EC II); 2) listed in Section 41 of the NERC Act, 2006 as Species of Principal Importance for the conservation of biodiversity in England; 3) having an IUCN threat status (CR, EN, VU, NT); or 4) listed in the most recent national status review as Red Data Book (RDB), Nationally Rare (NR) or Nationally Scarce (NS, Na, Nb). An explanation of these various status categories is given in the notes that precede the key species accounts in Section 3.

In Section 4, Site Reports have been prepared for each of the eight study areas. These give a brief description of the site and identify the key species and key invertebrate habitat features present at each; the latter are defined as those required by one or more of the key species. afull list of the invertebrates recorded from each site can be found at Appendix 1.

In general, the 2017 invertebrate survey is believed to provide an excellent baseline against which to assess the terrestrial and freshwater invertebrate assemblages of each. For the most part, the weather was sufficiently warm and dry to be favourable for invertebrate sampling. The only exception to this was at Parsonage Down NNR, where it rained on both visits. For this reason, flying insects are under-represented in the sample, though some key species were still found during a short window of dry weather on 29 July. Ground searches at this site were less affected by weather conditions and still produced records of many important species. A few pitfall trap catches were lost as a result of trampling or removal by predators, but those in the remainder were very large and have provided an excellent sample at all four sites where they were employed.

3. IMPORTANT INVERTEBRATES RECORDED ALONG THE A303 AMESBURY TO BERWICK DOWN SCHEME IN 2017

3.1. Key invertebrates

Key species are those that have a formal national or international conservation status and/or are legally protected. A full list of the various designations that determine inclusion here as a key species has been given in sub-section 2.2. The species accounts in this section of the report include brief notes on their occurrence at the eight sample sites, plus comments on ecology and national distribution.

The rove beetle Philonthus parvicornis was not accorded a conservation status in the last published review of this group (Hyman & Parsons, 1994), but will become a Nationally Scarce species in the forthcoming update (Boyce, in prep.), so it has been included here as a key species. The same upcoming review has downgraded another rove beetle, Paederus fuscipes, which was found at the Upper Till in 2017. It is now known to be too widely distributed to justify Nationally Scarce status and has therefore been omitted from the key species list.

The emboldened national conservation status categories given after the scientific name that heads each Species Account are defined as follows:

VU. - IUCN Red List, Vulnerable. A taxon is Vulnerable when the best available evidence indicates that it meets any of the IUCN criteria A to E for Vulnerable. Note that the IUCN categories denote a threat based on a severe decline, as opposed to the UK NR, NS or RDB categories, which are based solely on scarcity expressed as the number of 10km squares of the National Grid (hectads) in which a species has been recently recorded (Harvey et. al., 2017). NR. - Nationally Rare. In recent National Reviews the plethora of Red Data Book categories used in earlier publications (RDB1, RDB2, RDB3, RDBI and RDBK) have been subsumed into a single category covering species occurring in 15 or fewer hectads (Harvey et. al., 2017). RDB3. – Red Data Book Category 3 - Rare. Taxa with small populations, occurring in 15 or fewer hectads of the National Grid that do not at present meet the criteria for Endangered or Vulnerable, but are nonetheless at risk (Falk, 1991a; Hyman & Parsons, 1992). N. – Nationally Scarce. Taxa which are estimated to occur within the range of 16 to 100 10km squares, but where division into Na or Nb status has not been attempted due to limited availability of information on British distribution. Second status review not yet completed and status therefore taken from first review (Hyman & Parsons, 1994). Na. – Nationally Scarce Category A. Taxa thought to occur in 30 to 16 10 km squares of the National Grid (Falk, 1991; Hyman & Parsons, 1992 & 1994). Nb. – Nationally Scarce Category B. Taxa thought to occur in between 30 and 100 10 km squares of the National Grid (Hyman & Parsons, 1992 & 1994). NS. – Nationally Scarce. In recent reviews the Na and Nb sub-divisions have been subsumed into a single category covering species occurring in 16 to 100 10km squares of the National Grid. Unlike the previous ‘N’ category, which covers the same distribution range, the amalgamation does not necessarily result from inadequate information on the British distribution (Alexander et. al., 2014; Boyce, in prep.; Foster, 2010; Telfer, 2016)

3.1.1. A Gnaphosid Spider Phaeocedus braccatus (L. Koch, 1866). VU./NR. P. braccatus is a very distinctive, mid-sized spider in which the dark abdomen has three pairs of white spots. It is one of the most important species recorded during the 2017 survey, being classed as both Vulnerable and Nationally Rare on account respectively of its significant decline and restricted British distribution. It is an inhabitant of chalk grassland and occasionally, other dry, open habitats such as lowland heaths. Sites in which it is found usually have early-successional vegetation with much bare ground. A single female was collected in June 2017 from the sparsely vegetated south-facing chalk bank at Countess Cutting CWS.

3.1.2. A Diving Beetle Hydroporus ferrugineus Stephens, 1829. NS. Within the diving beetle family Dytiscidae, Hydroporus is the largest genus, with 28 species on the British list currently. All are small and rather nondescript, with examination of the male genitalia being a very useful aid to ensure accurate identification. H. ferrugineus is one of the more distinctive species, being relatively large and reddish-yellow in colour with a rather parallel-sided body form. It has a very specialised ecology, being found in springheads where the water is at least moderately base-rich. It is believed to be partly subterranean, having been found occasionally in water brought up from underground boreholes. Adults were found in June and July at the Lower Till site, amongst the flinty substrate at the springhead that feeds one of the two main ditches crossing the southern field.

3.1.3. A Diving Beetle Hydroporus marginatus (Duftschmid, 1805). NS. Like H. ferrugineus, this is one of the larger and more easily recognised Hydroporus species. The elytra are of a pitchy yellow colour with very fine, dense punctures and the dark pronotum has broad, reddish side margins. H. marginatus is now restricted to a few sites scattered across southern England, though its historic range seems to have been wider, with older records further to the north and west in England, plus a handful of Welsh sites. It is a specialist inhabitant of lime-rich flowing water, being especially characteristic of winterbournes, such as the Till, that are subject to periodic dessication over the summer months. A male and female were netted from the main river at the Lower Till site in June 2017.

3.1.4. A Ground Beetle Zabrus tenebrioides (Goeze, 1777). NS. There is only one British species of Zabrus; a very distinctive, large black Carabid, with the pronotum having a wide border and its dorsal surface having wrinkles and coarse punctures. It is a very scarce southern species, which is known from scattered sites in southern England and south Wales. It is usually found in cultivated fields where there is abundant bare ground. A single specimen was collected in a pitfall trap at the Arable 1 site in July 2017.

3.1.5. A Ground Beetle Amara curta Dejean, 1828. NS. The genus Amara comprises a large group of mid-sized, ground beetles, most of which have a bronze-metallic ground colour. A. curta is a relatively small species, with the antennae dark except for two orange basal segments. It is a very scarce species in Britain, which occurs at scattered sites as far north as southern Scotland. Most recent localities are in southern England, where it is found in dry, open conditions in a range of habitats such as calcareous grasslands, heaths, sand dunes and brownfield sites. Often there is a significant component of bare ground in the habitats it is found in. A single adult was collected on a south-facing slope at Parsonage Down NNR in June 2017.

3.1.6. A Ground Beetle Ophonus schaubergerianus (Puel, 1937). NS. This is a black-brown, mid-sized ground beetle, with sinuate side margins to its strongly transverse pronotum. It is one of a group of closely similar Ophonus species, which may require dissection of male genitalia to ensure correct identification. It is found in a range of open habitats where short vegetation with patches of bare ground create the xerothermic conditions it requires. Larvae live in the soil, where both they, like the adults, feed on seeds. Almost all British sites are to the south of the Severn-Wash line, with a handful of additional records from the north-east of England. A single male was collected in July 2017 at Countess Cutting CWS.

3.1.7. A Ground Beetle Licinus depressus (Paykull, 1790). NS. There are only two British Licinus species, with L. depressus being distinguished from L. punctatulus by its smaller size, smooth elytra and the presence of two to three irregular puncture rows on each of the elytral striae. It is very localised in southern and eastern England, north as far as Durham and there are also a few sites in south Wales. It is a specialised snail predator, which is restricted to calcareous substrates where its prey is most numerous. Most colonies are in dry, open sites, such as calcareous grassland and chalk pits, but it has also been found occasionally in woodlands on chalk and limestone. Two adults were collected from grass tussocks at Parsonage Down NNR in July 2017 and a single individual was found in a pitfall trap at Countess Cutting CWS in the same month

3.1.8. A Ground Beetle Licinus punctatulus (Fabricius, 1792). Na. L. punctatulus is a very distinctive ground beetle of a matt-black colour, with a broad body form and an elytral sculpture consisting of large punctures set in transverse furrows. It is one of only two British Licinus; the other, L. depressus, being smaller and narrower with even elytra and much finer elytral punctures. It has a very restricted distribution in southern Britain, north as far as East Anglia and north-east Wales. It is a specialist snail predator that is restricted to dry, open calcareous grassland habitats, usually in places where vegetation cover is patchy and there is a significant component of bare ground. A single adult was found in a pitfall trap at Countess Cutting CWS in July 2017.

3.1.9. A Leiodid Beetle Ptomaphagus varicornis (Rosenhauer, 1847). RDBK. The genus Ptomaphagus includes three British species, all of which are quite small, black beetles with characteristic transverse elytral sculpturation and neat, shining pubescence. They can only be told apart by careful examination of the relative width of the fore tarsi and the length of the hind tarsi. The male aedeagus also provides useful characters. P. varicornis has a longer first segment of the hind tarsi and on average is larger than the other two species in the genus. It is a very scarce insect, which is only known from a few places in England. Most records are from southern counties, though there are a few old records north as far as Derbyshire. Its ecology is poorly understood, but the two other members of the genus are often found in association with carrion. Sites for P. varicornis are usually in dry, open places on chalk or limestone. A single male was collected along with large numbers of the commoner P. medius in a pitfall trap at the Arable 3 sample site in July 2017.

3.1.10. A Sexton Beetle Nicrophorus interruptus Stephens, 1830. Nb. The sexton beetles include five British species of large, black carrion beetles in which the elytra are marked with two bright red bands. N. interruptus can be distinguished from its congeners by its straight hind tibiae, the lack of conspicuous yellow pubescence along the front margin of the thorax and the break in the anterior red elytral band to each side of the suture. It is found in a range of open habitats, where it feeds on carrion. Specimens were found in pitfall traps at both the Arable 2 and Arable 3 sample sites in July 2017.

3.1.11. A Rove Beetle Sepedophilus testaceus (Fabricius, 1792). N. S. testaceus is a reddish-brown beetle with a strongly tapering hind body. It is one of a small group of very similar species in the genus from which it can be distinguished by its somewhat transverse apical antennal segments and less strongly dilated protarsi. Examination of the male genitalia provides a valuable confirmatory character. It is generally found in damp, well rotted heartwood of broadleaved trees, with beech and willow being especially favoured. Nationally, it is very locally distributed in England to the south and east of a line between the Humber and Severn Estuaries. A single adult was found in damp poplar heart-rot at the Upper Till site in 2017.

3.1.12. A Rove Beetle Lamprinodes saginatus (Gravenhorst, 1806). Na. Its fusiform body shape, shiny surface, orange thorax and elytra, contrasting with a black head, scutellum and hind body give L. saginatus a similar appearance to the very common species pair in the Tachyporus chrysomelinus aggregate (the T. dispar segregate was recorded in a number of sites during this study). From these L. saginatus can be recognised by its larger size and more transverse antennae. It is found at scattered sites across much of England and Wales, but seems to be absent from Scotland. It is associated with a range of ant species; usually those in the genera Lasius and Myrmica. It is usually found in open, dry sites, such as chalk grassland, coastal cliffs and lowland heaths. A single adult was shaken from a grass tussock at Parsonage Down NNR in July 2017 and a second was found in the same month in moss at Countess Cutting CWS.

3.1.13. A Rove Beetle Oxypoda exoleta Erichson, 1839. N. O. exoleta is a small, Aleocharine rove beetle with an obscure black and orange pattern that is most reliably distinguished by reference to the male and female genitalia. It is found very locally across England and Wales. The ecology of the beetle is very poorly known, though many records relate to sandy substrates in riparian habitats. It was found in pitfall traps set at the Arable 1 sample site in July 2017. Because its ecological requirements are not understood currently, this species has not been assigned to a habitat feature in the Arable 1 Site Report.

3.1.14. A Rove Beetle Gnypeta ripicola (Kiesenwetter, 1844). N. The rove beetle genus Gnypeta includes four British representatives, all of which are small black or pitchy brown insects. G. ripicola is distinguished from its congeners by its broader head, more convex eyes and longer legs and antennae. The form of the male aedeagus and female spermatheca are also extremely distinctive. Like all the species in the genus, G. ripicola is found running over bare mud at the edge of freshwater. It appears to be a relatively recently colonist, which was first described from the UK in 1980. It has spread across England as far north as Yorkshire, but still has a very localised distribution, with most sites being in the south-east. The single male recorded with the commoner G. rubrior on the muddy margins of the pond at the Upper Till site in July 2017 represents the first Wiltshire record.

3.1.15. A Rove Beetle Stenus butrintensis Smetana, 1959. N. The sub-genus Metatesnus includes a number of mid-sized black rove beetles, with grey-white hairs covering much of the body and short, strongly lobed tarsi. Like the very similar, and much commoner S. pallitarsis, S. butrintensis has yellow tarsi, and it is most readily distinguished by the form of the male genitalia. It has a very scattered distribution across southern England, with most sites being in the south- east. It is usually found in tall fen habitat, particularly in stands of greater reedmace Typha latifolia, where it is often to be found inside the leaf sheaths. In July 2017, a single male was collected from branched bur-reed Sparganium erectum growing at the edge of the river at the Lower Till sample site.

3.1.16. A Rove Beetle Sunius melanocephalus (Fabricius, 1792). N. S. melanocephalus is a rather small orange-brown rove beetle with a blackish head. It can be distinguished from the other two British Sunius species by the lack of strong reticulate microsculpture on the head and the distinctive form of the male aedeagus. It is a southern species, found as far north as south Wales and East Anglia. Most records are from dry, open sites, where it is usually found in grass tussocks and litter. A single individual was collected in July 2017 in a pitfall trap at the Arable 3 site.

3.1.17. A Rove Beetle Rugilus similis (Erichson, 1839). N. R. similis is one of a small genus of large-eyed rove beetles of very distinctive appearance in which the broad head is set onto the relatively narrow thorax with a pedicel. R. similis may be recognised from its congeners by its relatively large size, all-yellow legs and the very distinctive form of the male genitalia. It appears to be quite catholic in its habitat requirements, being recorded from litter in both wet and dry grassland sites. In 2017, a male and female were recorded at the Upper Till 1 sample site in July, these being collected in cut grass litter in an area of somewhat damp semi-improved grassland. Three further adults were found in grass tussocks at Parsonage Down NNR and two males were collected in pitfall traps set in a conservation headland at the Arable 1 site. All records date from July 2017.

3.1.18. A Rove Beetle Philonthus atratus (Gravenhorst, 1802). Na. Though it forms part of one of the largest Staphylinid genera, P. atratus is quite easy to distinguish from its congeners. It is a medium-large black species, with a slight bronzy-metallic sheen to the elytra. It has a row of four punctures on each side of the thoracic dorsum and a parenthesis-shaped line running across the basal abdominal segments. It has a wide but very scattered distribution across England and Wales, with the latter appearing to be a particular stronghold. This is certainly a wetland species, though it has also been found occasionally in other open habitats. Many of its wetland sites have expanses of bare peat and mud, often where cattle or horses are poaching the ground. A single male was found at the Upper Till 1 sample site in July 2017, running over poached, wet mud at the edge of the pond.

3.1.19. A Rove Beetle Philonthus parvicornis (Gravenhorst, 1802). pNS. P. parvicornis is a relatively small all-black member of this very large Staphylinid genus. It has a rounded head and a row of five punctures on either side of the pronotal mid-line. It can only be reliably distinguished from other similar species by examination of the male genitalia. It is usually found in cow or horse dung in a range of open situations, though it has also been recorded less frequently in other patch habitats such as compost heaps and carrion. Recent British records of P. parvicornis are confined to scattered sites in England, north as far as Yorkshire and a handful of Welsh localities, though there is an old record from Cumbria. It was not given a conservation status in the earlier national assessment of Hyman & Parsons (1994), but will be upgraded to Nationally Scarce in the forthcoming Species Status Review (Boyce, in prep.). Four males were collected from cow dung in the rough, semi- improved pasture at the Upper Till 1 sample site in July 2017.

3.1.20. A Rove Beetle Quedius nigrocaeruleus Fauvel, 1874. Nb. Quedius has 45 British species of relatively large, predaceous rove beetles. Q. nigrocaeruleus is quite easily distinguished from other members of the genus by its distinctive all-black ground colour with a slight steely-blue metallic reflection. It is an inhabitant of mammal and bird nests, where it is thought to prey on other invertebrates, such as fleas. It is most often encountered underground in mole nests, though there are some records from above ground in bird nests in tree hollows. It has been also found occasionally in nests of social insects and in fungi. It is quite widely, but very thinly distributed across England north as far as Cumbria. There is a single known Welsh site, at Newborough Warren, Anglesey but it is not currently recorded from Scotland. A single female was found in rotten heartwood of an old black poplar at the Upper Till 1 sample site in July 2017.

3.1.21. A Woodworm Beetle Anobium inexspectatum (Lohse, 1954). Nb. In general appearance, A. inexspectatum closely resembles the common woodworm A. punctatum. Reliable identification relies on microscopic examination of the male aedeagus or female ovipositor styles, the former being particularly distinctive. It has a wide, but localised distribution across England and Wales as far north as Cumbria, though the majority of recnt sites are in southern and eastern counties of England. Larvae of A. inexspectatum bore into old dead stems of ivy Hedera helix and the single male tapped from low vegetation at the Arable 3 site in June 2017 presumably originated from ivy growing in the hedge here.

3.1.22. A Pollen Beetle Meligethes solidus (Kugelann, 1794). N. The genus Meligethes includes a large number of small, black beetles that are difficult to distinguish from each other. However, M. solidus is one of the more readily recognised species, being relatively large, with prominent teeth on the tarsal claws. Both larvae and adults are found on the flowerheads of common rock-rose Helianthemum nummuralium, where they feed on the pollen. In Britain, M. solidus is restricted to chalk and limestone counties in the southern half of England as far north as Derbyshire. It is also known from a few sites in Wales. In 2017, a single specimen was tapped off flowers of common rock-rose at Parsonage Down NNR in July and another individual was collected on this plant in the remnant area of chalk grassland at Diamond Wood.

3.1.23. A Phalacrid Beetle Phalacrus championi Guillebeau, 1892. Na. The Phalacrids are a small family of shiny, oval beetles. P. championi is a predominantly black beetle with a single sutural stria on each elytron and a diagnostic comb of four or five short spines on the outer apical edge of the front tibiae. Most records of this extremely scarce beetle are from southern and eastern counties of England, but there are also a couple of recent records from eastern Scotland. Phalacrus are known to be associated with grasses and sedges, where their larvae feed on fungal spores of smuts growing on the inflorescences. There are British records of P. championi from a range of open habitats, including saltmarshes and chalk grassland. Single males were collected from a grass tussock at Parsonage Down NNR in July 2017 and from arable weeds at the Arable 1 sample site in June 2017.

3.1.24. A Salpingid Beetle Lissodema denticolle (Kugelann, 1794). NS. Its elongate body form, allied to the red head and thorax and black elytra with four orange spots makes this an easily recognised beetle. It is a dead wood specialist, which is most often found in association with dead canopy branches of a wide range of broadleaves and also occasionally Scot’s pine Pinus sylvestris. It has a wide but localised distribution across southern England as far north as the Midlands, with a few sites in the Welsh border counties and isolated sites on Anglesey and in Yorkshire. Three adults were beaten from dead hawthorn branches at the Upper Till 1 sample site in 2017.

3.1.25. A Longhorn Beetle Paracorymbia fulva (De Geer, 1775). RDB3. This is a relatively large and very distinctive longhorn beetle in which the elytra are tawny-orange with black apices and the forebody and appendages are entirely black. It was formerly said to be a dead wood species, but most of its recent localities are in open habitats where there are few if any old trees. Until the larval habits are confirmed, comments on its ecology must be regarded as provisional, but many of its modern sites are on dry, unimproved calcareous or maritime sites, where it is often found nectaring on tall umbels such as hogweed Heracleum sphondylium. Its British headquarters are on the chalk downs of southern Wiltshire and it was found widely during the 2017 study, with records from the Upper Till, Diamond Wood and Arable sample sites 1, 2 and 3. Individuals were found on blossoms of hogweed, upright hedge parsley Torilis japonica, spear thistle Cirsium vulgare and field scabious Knautia arvensis. Because its ecological requirements are not understood currently, this species has not been assigned to a habitat feature in the Site Reports (see section 4).

3.1.26. A Flea Beetle Psylliodes chalcomera (Illiger, 1807). NS. This is a metallic-blue or -green species in which the front femora are brown, and the pronotum has rounded front angles and both large and small punctures. Though it is quite widely distributed across Britain, it is exceptionally local away from southern England. Even in the latter area, it is scarce, being most frequent on very warm, open sites on the south coast. Both larvae and adults are found on a range of thistle species in the genera Carduus and Cirsium, but it seems to particularly favour musk thistle Carduus nutans. During the 2017 study it was recorded at Parsonage Down NNR and the Arable 1 sample site, in June and July respectively.

3.1.27. A Weevil Hemitrichapion reflexum (Gyllenhal, 1833). Na. H. reflexum is a small, blue-black weevil in which the elytra are sparsely pubescent. It has a scattered distribution on chalk and limestone in southern England, with a few outposts as far north as Yorkshire. It is not known to occur in Wales. The larvae feed exclusively on sainfoin Onobrychis viciifolia, and the adults are almost invariably found by beating this plant. In June 2017, two individuals were tapped from their hostplant in the restoration grassland at the Arable 2 sample site.

3.1.28. A Weevil Oxystoma cerdo (Gerstaecker, 1854). Nb. O. cerdo is a small black weevil with the rostrum markedly narrowed towards its apex. Though it is very similar to the common O. subulatum, the base of the antennae is clearer yellow in this species, and the male genitalia are also distinctive. O. cerdo is most frequent in northern England and southern Scotland andI is very locally distributed in southern England. It feeds on various species of vetch Vicia spp., with tufted vetch V. cracca being especially favoured. The Salisbury Plain area appears to be a stronghold for this beetle in southern England and it was collected from its foodplant at the Lower Till sample site in June 2017.

3.1.29. A Weevil Catapion pubescens (Kirby, 1811). Nb. C. pubescens is a small all-black weevil with a covering of sparse white pubescence. It is quite easily recognised from other superficially similar Apionids by the presence of a conspicuously hollowed-out area on the head between the eyes. Both adults and larvae are found on small yellow trefoils, such as hop trefoil T. campestre, large hop trefoil Trifolium aureum and lesser trefoil Trifolium dubium, the larvae making galls in the stems. C. pubescens occurs in a range of dry, open habitats, such as unimproved grassland, sand dunes and short ruderal vegetation, where there are good stands of its foodplants. It has a very scattered distribution across England and Wales, with most colonies lying south of a line between the Humber and Mersey estuaries. However, there are a few other records outside this area, as far north as Cumbria, but with no confirmed Scottish records. The record of a female and male at the Arable 1 sample site in June 2017 represents only the third record of C. pubescens from Wiltshire.

3.1.30. A Weevil Protapion filirostre (Kirby, 1808). Nb. P. filirostre is a small, very convex, all-black weevil. It is a scarce species restricted to southern counties of England and Wales. It is found in grassland and short, ruderal vegetation, where there are large stands of its main foodplant black medick Medicago lupulina, larvae feeding in the buds of this and other species of the genus. A single female was collected from large stands of black medick growing on a conservation headland at the Arable 1 sample site in July 2017.

3.1.31. A Weevil Gymnetron melanarium (Germar, 1821). Nb. G. melanarium is a small, all-black weevil with scattered pale setae on the elytra. It is a scarce beetle throughout its range, which is centred on southern England with scattered records north as far as Yorkshire and west into south Wales. The beetle is found in dry, open habitats, such as grassland and arable margins, where there are good populations of its foodplant. Larvae feed in galls they make in the stems of germander speedwell Veronica chamaedrys and may possibly also use other species in this genus. In July 2017, a single adult was tapped from a knapweed plant growing in the remnant area of chalk grassland at the south-western edge of Diamond Wood.

3.1.32. A Weevil Gymnetron villosulum Gyllenhal, 1838. Nb. This is a small, weevil with reddish elytra, which is most easily distinguished from its congeners by the thick-set white scales at the sides of the thorax and the dense yellowish setae covering the elytra. G. villosulum has a very scattered distribution across southern England, with almost all records being to the south-east of a line joining the Severn and Humber estuaries. Both larvae and adults feed in galled seed capsules of water-speedwells Veronica spp. During this study, it was found on blue water-speedwell Veronica anagallis-aquatica growing on sparsely vegetated, muddy river margins at both the Upper and Lower Till sample sites in both June and July 2017.

3.1.33. A Weevil Mecinus circulatus (Marsham, 1802). Nb. M. circulatus is a small, rather elongate black weevil, in which the sides of the elytra and thorax are clothed with long, white hairs. It is a warmth-loving, southern species, which is not found in Britain further north than the Midlands and north Wales. It occurs in a wide range of open habitats, especially grassland and sand dunes, with most of its British sites being coastal. It feeds on plantains, with Ribwort Plantago lanceolata and Buck’s-horn Plantain P. coronopus being known hostplants. A single specimen was found in a pitfall trap set in early-successional chalk grassland at Countess Cutting in July 2017.

3.1.34. A Weevil Tychius pusillus Germar, 1842. Nb. The genus Tychius includes 14 British species, all of which are small weevils with a covering of pale scales. Its very small size (1.4-1.8mm) and black antennal club make T. pusillus quite easy to distinguish from its congeners. It shows a markedly southern distribution in England, with sites as far north as the Midlands, but no records from Wales. It feeds on lesser trefoil Trifolium dubium and probably other small yellow- flowered trefoils, with the larvae developing inside the flowerheads. It occurs in a range of dry, open habitats, such as grassland, brownfield sites and sand dunes, where there are good stands of its hostplants. In June 2017, five adults were tapped from a mixture of lesser and hop trefoil growing on the arable margin of the Arable 1 sample site. This is the first occasion that T. pusillus has been found in Wiltshire.

3.1.35. A Weevil Drupenatus nasturtii (Germar, 1824). Nb. The sub-family Ceutorhynchinae includes many small weevils that have a distinctive squat body form. D. nasturtii is one of the larger members of the sub-family and can be easily recognised by its dark, metallic-green ground colour over which there is a sparse covering of whitish scales. It is a wetland specialist, which is found very locally throughout England and Wales, though with no known Scottish localities. Larvae devlop in the stems of water-cress Rorippa nasturtium-aquaticum growing in rivers, streams, ditches and other waterbodies. In June 2017, adults were tapped from their hostplant at the edge of the river at both the Upper and Lower Till sample sites.

3.1.36. A Weevil Sirocalodes quercicola (Paykull, 1792). Na. The genus Sirocalodes has three British representatives, all of which are small, squat weevils of a blackish ground colour. S. quercicola can be distinguished from the other two by its appendiculate claws and the large bar of whitish scales covering the base of the elytral suture. It is much the rarest of our three Sirocalodes, with recent records confined to a handful of sites, which are very thinly scattered across England, Wales and southern Scotland. Like the much commoner S. depressicollis, it feeds exclusively on fumitories Fumaria spp., with the larvae creating a gall in the stem. A single individual was found, amongst many S. depressicollis, by tapping fumitories growing on a narrow arable margin at the Arable 3 sample site. This is the first Wiltshire record of S. quercicola.

3.1.37. A Weevil Thamiocolus viduatus (Gyllenhal, 1813). Nb. T. viduatus is a widely distributed, but very local weevil that is known from scattered sites across England, Wales and southern Scotland. It is most often found on marsh woundwort Stachys palustris, though other Stachys spp are reported to also be used occasionally. Adults were tapped from marsh woundwort plants growing along the river bank at the Lower Till site in July 2017 and were also recorded in the same month on this plant where it was growing on an arable margin at the Arable 3 sample site.

3.1.38. A Weevil Zacladus exiguus (Olivier, 1807). Nb. Z. exiguus is a small black weevil with very prominent setose tubercles on the elytra and the thorax sharply turned up along its front edge. It is an extremely scarce species nationally, with a thin scatter of sites across south-eastern and south-central England. It is found in a variety of open habitats and feeds on various small-leaved cranesbills, such as herb Robert Geranium robertianum and dove’s-foot cranesbill Geranium molle, with the larvae feeding in the rootstocks of these plants. Two adults were tapped from a small-leaved cranesbill at the Arable 1 sample site in June 2017. This is the one of only two records of the beetle from Wiltshire, the first being from Hat Gate before 1938.

3.1.39. A Weevil Larinus planus (Fabricius, 1792). Nb. There is only one British Larinus and it is an unmistakeable large black weevil with conspicuous patches of yellow hairs. It is restricted to the southern part of England and Wales, where it occurs on various species of thistles of the genera Cirsium and Carduus, the larvae developing inside the flowerheads of these plants. It is reasonably widespread across southern England, and occurs very locally in Wales, but is absent from northern England and Scotland. L. planus occurs in a range of open grassland and ruderal communities. It was found at the Upper Till, Lower Till and Arable 1 sample sites in 2017, with all records coming from spear thistle Cirsium vulgare.

3.1.40. Small Scabious Mining Bee Andrena marginata Fabricius, 1776. Na. This is a variable but distinctive Andrena species in which females may be entirely dark-bodied, have orange-red gastral markings, or be predominantly orange on the abdomen. Males have a yellow-marked clypeus, but are dark bodied. The nesting biology is unrecorded in Britain, but abroad A. marginata is reported to nest both singly and in small aggregations. A. marginata shows a strong preference for collecting pollen from small scabious Scabiosa columbaria, field scabious Knautia arvensis and devil’s - bit scabious Succisa pratensis, although other pollen sources are occasionally utilised. A. marginata occurs on chalk grassland, marshy grassland, heaths, moors, sand dunes and at wood edges. A. marginata is host to the Rare (RDB 3.) cleptoparasitic bee Nomada argentata (see 3.1.41 below). In the past, A. marginata had a disjunct distribution in southern England, and in Scotland and Ireland. However, there are no modern Scottish records, and a considerable decline has occurred in Ireland. Within southern Britain A. marginata has also declined considerably, although colonies remain scattered through southern England, East Anglia and south Wales. Workers were seen in good numbers on small scabious flowers during the one brief spell of dry weather at Parsonage Down NNR on 29 July 2017.

3.1.41. Plain Mini-miner Bee Andrena minutuloides Perkins, R. C. L., 1914. Na. This small species can be difficult to separate from the closely related A. minutula, but the former is characterised by a shinier, less sculptured thoracic dorsum. Little is known of the nesting biology or pollen-collecting habits of A. minutuloides, although it is thought to require light, sparsely vegetated soils in sunny situations. It occurs in calcareous grassland, chalk heath, heathland and coastal locations. A. minutuloides is widely but thinly distributed in southern England and south Wales. A number of workers were found in July 2017 foraging on flowers of upright hedge-parsley Torilis japonica in rank calcareous grassland on the earthwork that runs up the western edge of Diamond Wood.

3.1.42. Silver-sided Nomad Bee Nomada argentata Herrich-Schäffer, 1839. RDB3. N. argentata is a red-girdled Nomada lacking yellow spots, which is strongly associated with scabious flowers. It is a cleptoparasite of the scarce mining bee Andrena marginata (see 3.1.39). The host shows a strong preference for collecting pollen from small scabious Scabiosa columbaria, field scabious Knautia arvensis and devil’s-bit scabious Succisa pratensis. Recent records are confined to southern England, East Anglia and south Wales, most frequently on chalk grassland, heaths, dunes or moorland. N. argentata has always been considerably rarer than its host, and has never been recorded from many of the host’s present and former sites, including all its Welsh and Scottish localities. Although it was recorded from widely scattered sites across southern England in the past, it has declined considerably and there are post-1970 records for mainland Britain from only 16 10km. squares (Edwards, R. & Telfer, 2001). A single male N. argentata was collected nectaring on small scabious in July 2017 at Parsonage Down NNR.

4. SITE REPORTS

The following sub-sections set out the results of the invertebrate surveys carried out at each of the eight sample sites. They include a description of the habitats and management of the site in question, along with a description of those that are of key importance for invertebrates. Key habitat features are simply defined as being those that support one or more of the key species. The key species associated with each of the key habitat features are shown in brackets after the habitat feature name. A full list of all invertebrates recorded at each of the sample sites can be found at Appendix 1.

4.1. Upper Till

This survey unit lies approximately 2 km upstream of the Lower Till site, to the north of the A303 where it passes through Winterbourne Stoke. A central grid reference for the site approximates to SU081417. The Upper Till sample site encompasses an area of cattle-grazed pasture through which the upper reaches of the River Till passes. The latter is a high-quality chalk stream and the river channel is notified as both an element of the River Till Site of Special Scientific Interest (SSSI) and the River Avon Special Area of Conservation (SAC). Invertebrate surveys were carried out here on the 26th June and 24th July 2017.

The River Till is a winterbourne and was largely dried-up by the time of the second visit in July. Vegetation within the watercourse was dominated by beds of water- cress Rorippa nasturtium-aquaticum and fool’s watercress Apium nodiflorum and there was also much water forget-me-not Myosotis scorpioides, floating sweet-grass Glyceria fluitans and blue water-speedwell Veronica anagallis-aquatica, the latter mainly being in places where cattle trampling had created areas of sparsely vegetated mud.

The pasture land around the river was agriculturally improved to semi-improved and was mostly rather species-poor, though stands around the pond were a little wetter and more diverse with scattered clumps of tufted hair-grass Deschampsia caespitosa. There was also a narrow strip of heavily grazed, relatively unimproved chalk grassland running along the eastern edge of the pond field, which had a more diverse flora including typical chalk grassland species such as stemless thistle Cirsium acaule.

The pond had much bare clayey mud, with pioneer ‘silt pond’ aquatic vegetation comprising submerged beds of stoneworts and algae with some emergent growth of amphibious bistort Persicaria amphibia. In the drawdown zone around the pond margin there was a patchy growth of the latter, with water mint Mentha aquatica and carpets of brown mosses also prominent. The sparsely vegetated appearance of the pond is maintained by trampling by cattle trampling and the massive drawdown in water levels over the summer period must also help to maintain its open character.

A feature of this site was the presence of scattered, mature specimens of a range of native trees and shrubs, including crack willow Salix cracca, ash Fraxinus excelsior, hawthorn Crataegus monogyna and what appeared to be native black poplar Populus nigra.

4.1.1. Key Invertebrate habitat features at the Upper Till The survey has identified 11 key species associated with four key habitat features. One key species, the longhorn beetle Corymbia fulva has not been assigned to a key habitat feature, as there is insufficient information on its ecological requirements.

Pond and margins (Pterostichus anthracinus; Gnypeta ripicola; Philonthus atratus) The pond and its margins were found to have a diverse wetland invertebrate assemblage that included three key species. The ground beetle P. anthracinus is usually found in well-vegetated ancient fenland sites and its presence here was somewhat surprising given the seemingly recent origin of the pond. The rove beetles G. ripicola and P. atratus are both found on freshwater margins where there are expanses of bare mud. The presence of cattle poaching the margins and the strongly fluctuating water levels are both critical in maintining suitable conditions for these two Staphylinids. A number of other local wetland beetles typical of fluctuating wetlands were also found here.

River and margins (Gymnetron villosulum; Drupenatus nasturtii) Two scarce weevil species were recorded along the River Till: D. nasturtii was found in association with water-cress beds and G. villosulum was tapped from blue water- speedwell growing on bare, poached mud in the drawdown zone. The cattle grazing the surrounding pasture have access to the river and keep conditions very open, which is certainly beneficial for G. villosulum, as blue water-speedwell requires patches of bare mud into which it can germinate. D. nasturtii requires lower levels of disturbance, such that there are less poached and grazed water-cress beds.

Semi-improved damp grassland (Pterostichus anthracinus; Rugilus similis; Larinus planus) Though it has clearly been agriculturally improved in the past, the stands of rather wet, rough grassland with tufted hair-grass near the pond were found to still have a surprisingly diverse invertebrate fauna that included three key species. The rove beetle R. similis is associated with quite tussocky unimproved or semi-improved grassland, while the ground beetle P. anthracinus is a fen species and it was surprising to find a specimen here. Two other individuals were collected on the margins of the nearby pond and this specimen is perhaps best interpreted as a stray from that site or the nearby river corridor.

Scattered old trees and shrubs (Sepedophilus testaceus; Quedius nigrocaeruleus; Lissodema denticolle). A feature of the site is the presence of a number of mature to over-mature trees, as well as some old hawthorns. Two of the key species, S. testaceus and L. denticolle, are dead wood specialists, the former being found in rotten heartwood of an old black poplar, while the latter was beaten from dead branches of an over-mature

19 hawthorn. The third key species, Q. nigrocaeruleus, is usually found in underground nests of the mole and other mammals. However, it also occurs less frequently in bird’s nests in hollow trees and this presumably explains the presence of a female in heart rot of an old black poplar.

Dung (Philonthus parvicornis) Examination of cattle dung on stands of semi-improved rough pasture showed the site to also have a diverse coprophilous invertebrate fauna that included one further key species, the rove beetle P. parvicornis.

4.2. Lower Till

The Lower Till sample site lies approximately 2 km downstream of that on the Upper Till described in the preceding sub-section and is to the south of the A303 where it passes through Winterbourne Stoke. A central grid reference for the site approximates to SU077397. The river is noticeably larger here than at the first site and still had a reasonable flow in July, when the survey site upstream had completely dried up. The River Till is a high-quality chalk stream and both the main river channel, the two main stands of wet woodland and the southern fen-meadow field to its west are notified as part of both the River Till SSSI and the River Avon SAC. This site was surveyed on the 26th June, 27th June and 25th July 2017.

The main watercourse is shallow with a flinty bottom. It is fringed with grey willow Salix cinerea and alder Alnus glutinosa carr, or in open areas with branched bur-reed Sparganium erectum, reed sweet-grass Glyceria maxima, hemlock water-dropwort Oenanthe crocata and other tall fen species. There are also two spring-fed ditches running into the river from the main fen meadow that lies to its west. These also have a predominantly open, flinty substrate, with extensive beds of water-cress Rorippa nasturtium aquaticum and fool’s water-cress Apium nodiflorum in some places, as well as a few patches of monkeyflower Mimulus moschatus.

There are stands of greater pond-sedge Carex riparia-dominated fen meadow in the southernmost of the two fields that lie to the west of the river, with this habitat being concentrated in the eastern third of the field, close to the river and its tributary ditches. The western section of this field is dominated by drier, more species-poor grassland with tall grass tussocks and abundant creeping thistle Cirsium arvense and stinging nettle Urtica dioica. The northern field has very similar rank, spoecies-poor, ruderal grassland to that described above. In its southern part, this field has some patches of more diverse neutral grassland with herbs such as red clover Trifolium pratense, common knapweed Centaurea nigra and self heal Prunella vulgaris.

The area surveyed also included two stands of wet woodland dominated by grey willow and alder; the southern area, lying around the old decoy pond and the northern block occurring along the corridor of the River Till.

4.2.1. Key Invertebrate habitat features at the Lower Till The 2017 invertebrate survey has identified eight key invertebrates, with these being placed into four key habitat features, which are listed below.

River and ditch margins (Stenus butrintensis; Drupenatus nasturtii; Gymnetron villosulum; Thamiocolus viduatus) Vegetation and exposed substrates at the edge of the River Till and the spring-fed ditches that run into it are probably the most important invertebrate habitat feature at the Lower Till site. The value of the site is enhanced by the presence of a range of vegetation structures in the riparian corridor: bare, poached mud with sparse vegetation (favoured by blue water-speedwell, the foodplant of G. villosulum); intermediate height beds of water-cress, fool’s water-cress and monkeyflower (the former being the foodplant of D. nasturtii); and tall fen with species such as branched bur-reed (used by S. butrintensis at this site).

Chalk Streams and Springs (Hyroporus ferrugineus; Hydroporus marginatus) The two small diving beetles H. ferrugineus and H. marginatus are specialist inhabitant of calcareous waterbodies. The former tends to be found only at springheads and may be largely subterranean. A number of individuals of this species were found amongst flinty gravel at the source of the large spring that feeds the more northerly of the two ditched streams crossing the southern field. H. marginatus is associated with winterbournes and two adults were collectd from the underside of a submerged branch in the main river channel in July.

Fen Meadow (Oxystoma cerdo) A single key species, the weevil O. cerdo was tapped from tufted vetch growing in the great pond-sedge-dominated fen meadow vegetation that occupies the eastern third of the southern field. The stands of fen-meadow were quite small and fragmented and lacked most of the rarer invertebrates that might be expected to occur in high-quality examples of this habitat found in the floodplain of other Wiltshire chalk streams (eg. Desmoulins’ whorl snail Vertigo moulinsiana).

Tall, ruderal vegetation (Larinus planus) The large weevil, L. planus was found in association with stands of its foodplants, thistles, in the western part of the southern field. Though it is rated as a Nationally Scarce species currently, L. planus is quite widely distributed in this part of Wiltshire.

4.3. Parsonage Down NNR

Lying approximately 1km to the west of Winterbourne Stoke and just to the north of the A303, Parsonage Down comprises a large block of nearly 200 hectares of chalk grassland. Though parts of the site were ploughed and planted with crops during the Second World War, it is very unusual in having retained species-rich chalk grassland on what is generally quite flat to shallowly sloping topography. A central grid reference for the site approximates to SU062411. The entire site lies within the Parsonage Down SSSI and Salisbury Plain SAC and it is owned and managed by Natural England (NE) as a National Nature Reserve (NNR). Only the south-eastern part of the NNR was surveyed for invertebrates in 2017, this being the area that lies closest to the A303. The site was surveyed on the 28th June, 28th July and 29th July 2017. The main emphasis of the latter day’s work was to carry out a survey for marsh fritillary larval webs. This is somewhat earlier than such surveys are normally carried out, but local NE staff indicated that marsh fritillary webs were already evident at other Wiltshire NNRs at this time and it was therefore agreed that it would be appropriate to carry out the surveys at this time.

The survey area consists of a large and structurally homogeneous stand of unimproved to semi-improved chalk grassland lying on moderate to gentle north- and south-facing slopes on either side of the central dry valley. The valley sides have species-rich grassland in which some of the main grasses are fesues Festuca spp., meadow oat-grass Avenula pratensis, crested hair-grass Koeleria macrantha and quaking grass Briza media cover. The grassland was being grazed by cattle at the time of these surveys. This maintains a short sward with a high and very diverse range of herbs; bird’s-foot trefoil Lotus corniculatus, red clover Trifolium pratense, spiny restharrow Ononis spinosa, common rock-rose Helianthemum nummularium, salad burnet Sanguisorba minor, small scabious Scabiosa columbaria and stemless thistle Cirsium acaulon are amongst the most prominent. A feature of the north- facing slopes is the abundance of devil’s-bit scabious Succisa pratensis and saw-wort Serratula tinctoria.

The flat ground in the valley bottom and on the plateau at the southern edge of the site has much less species-rich swards in which grass cover is higher, with tussocks of fescues, Yorkshire fog Holcus lanatus, cock’s-foot Dactylis glomerata dominating the sward.

4.3.1. Key Invertebrate habitat features at Parsonage Down NNR Despite very poor weather during both visits, Parsonage Down produced records of ten key species in 2017. The extremely homogeneous nature of the vegetation at Parsonage Down means that all but two of the key invertebrate species are associated with the short-sward chalk grassland that covers the great majority of the site. One key species, the weevil S. ruficornis feeds on poppies in ruderal and arable habitats, which are not present on the NNR. It is presumed to be a stray from outside the site and has therefore ot been assigned a key habitat feature in the following discussion. Two key habitat features have been identified based on the list of key species, with these being listed and discussed briefly below.

Short-sward chalk grassland (Amara curta; Licinus depressus; Lamprinodes saginatus; Rugilus similis; Meligethes solidus; Phalacrus championi; Andrena marginata; Nomada argentata) Many of the important invertebrates recorded at Parsonage Down this year have an association with chalk grassland plants. For example, the mining bee A. marginata (and therefore its rare cleptoparasite N. argentata) require an abundance of small scabious flowers and the pollen beetle is restricted to common rock-rose. The key requirement for these is to maintain a grazing regime that maintains large flowering populations of these foodplants.

Other species, such as the two ground beetles and two rove beetles that are the first four key species in the above list, are not restricted by plant species composition, but by vegetation structure, with the first three requiring a reasonably short sward. The rove beetle R. similis tends to favour grasslands of somewhat taller stature. Grazing should therefore aim to maintain a mixed sward height, though with most areas being reasonably short, as is currently the case. The Phalacrid beetle P. championi feeds on smut fungi growing on sedges and is probably also favoured by a grazing regime that maintains a short sward where small sedges are able to flourish. This is clearly a very important invertebrate habitat with a regionally important invertebrate fauna.

Tall ruderal vegetation with Carduus nutans (Psylliodes chalcomera) Where cattle and rabbits have created areas of bare substrates, there are patches of tall, ruderal vegetation with musk thistle, on which the scarce flea beetle P. chalcomera is most often found.

4.3.2. Marsh fritillary survey On the 29th July, a survey of the Parsonage Down site for marsh fritillary Euphydrya aurinia larval webs was undertaken using the standardised methodology developed by the Butterfly Monitoring Scheme. This involved walking a zig-zag transect across the area in which the butterfly’s foodplant, devil’s-bit scabious Succisa pratensis is abundant, looking for the distinctive silken tents the larvae live in communally from late-summer to early-autumn. In addition, casual searching for larval webs was carried out on the same day. It is estimated that during six man-hours spent completing the larval monitoring exercise, the surveyors came within no more than 10m of all stands of chalk grassland with abundant devil’s-bit scabious. Prior

24 discussions with NE staff had indicated that at the time of this survey, marsh fritillary larvae were evident at other Wiltshire sites, such as the Pewsey Downs NNR, indicating that if a breeding population were present here, it should have been detected by the very thorough survey undertaken.

Despite the above, no marsh fritillary larvae could be found at Parsonage Down. NE staff have confirmed that, though occasional adults are recorded on the NNR, no larval webs have ever been recorded. It therefore seems certain that, at least in the south-eastern section of the Reserve that has been the focus of the 2017 commission, there is no breeding population of the marsh fritillary butterfly.

4.4. Diamond Wood

Diamond Wood is an area of mixed plantation woodland and scrub located to the south of the A303 and approximately 1.5km to the south-west of Stonehenge. A central grid reference for the site approximates to SU104408. The site was surveyed on the 29th June and 27th July. The Arable 1 sample site, which is in the same ownership, lies approximately 1km to the north-east of this site.

The woodland is an area of plantation woodland and scrub that is made up of a mix of native and planted species, including some conifers. It is clearly of secondary origin and much of the area has dense thickets of blackthorn Prunus spinosa and hawthorn Crategus monogyna, though with some areas of more mature trees that are mostly derived from plantings. The preliminary walk-over of the woodland and scrub interior showed it was likely to be of only very moderate importance for invertebrates and subsequent surveys therefore focused on investigating the invertebrate fauna of the open areas fringing the closed-canopy woodland and scrub.

Of particular interest is the strip of undisturbed chalk grassland that is associated with the old earthbank running down much of the western edge of Diamond Wood. This is ungrazed, so much of the grassland is rank, with abundant growth of tall umbellifers such as wild parsnip Pastinaca sativa, hogweed Heracleum sphondylium and upright hedge-parsley, as well as patches of tall, ruderal thistles and neetles and some bramble Rubus fruticosus thickets. However, there are still some small patches of more species-rich grassland, with a prominent component of forbs such as common rock-rose Helianthemum nummularium, common bird’s-foot trefoil Lotus corniculatus, wild basil Clinopodium vulgare, common knapweed Centaurea nigra and pyramidal orchid Anacamptis pyramidlis. The largest area of this habitat feature was at the south-western edge of Diamond Wood, where the grassland had some very large yellow meadow ant Lasius flavus nest mounds, indicating the grassland has not been disturbed for many years.

There are also quite wide arable margins around much of the wood, with these having typical arable weeds, including rape Brassica napus and other crucifers, fumitories Fumaria spp., black medick Medicago lupulina and various clovers and trefoils Trifolium spp.

4.4.1. Key Invertebrate habitat features at Diamond Wood Th 2017 surveys have generated records of four key species at Diamond Wood. One key species, the Red Data Book longhorn beetle Paracorymbia fulva, has not been linked to a habitat feature, as its ecology is very poorly understood. The remaining three key species have been assigned to the two key habitat features listed below.

Short-sward chalk grassland (Meligethes solidus; Gymnetron melanarium) The small stands of short-sward chalk grassland still present are the single most important habitat feature still present at Diamond Wood. The lack of grazing or other forms of management has resulted in most of the relatively undisturbed chalk

26 grassland becoming rather rank and species-poor (see below). However, in a few places, notably at the south-western edge of Diamond Wood, shorter swards with low-growing forbs such as common rock-rose (the foodplant of M. solidus) and germander speedwell (required by G. melanarium) still exist.

Rank chalk grassland with abundant umbels (Andrena minutuloides) Though somewhat less botanically diverse than the short-sward grassland referred to in the preceding paragraph, the stands of rank grassland with abundant umbels, including much upright hedge-parsley did have a diverse invertebrate fauna that included one key species, A. minutuloides. Note also that though it has not been assigned to a habitat feature, a large number of individuals of the rare longhorn beetle Paracorymbia fulva were also found nectaring on umbels in these areas.

4.5. Arable 1

The Arable 1 survey site comprises the large arable field planted with rape that runs up to the block of scrub at Normanton Gorse on its southern and western margins. It is just to the south of the A303, with Stonehenge being about 1km to the north- east. A central grid reference for the site approximates to SU111414. It is adjacent to the Arable 2 block, which lies on the northern side of Normanton Gorse (see following Site Report), while the Diamond Wood survey unit is approximately 1km to the south-west (see sub-section 4.4 above). It was surveyed on the 29th June and 27th July. Three lines of pitfall traps were put out between these two dates.

The focus of invertebrate survey effort here was on the wide conservation headlands the owner has retained around the edge of the field, with these running along the entire southern and western edges of Normanton Gorse. The headlands have a two- tiered structure, with a more frequently ploughed strip adjacent to the crop and a less disturbed zone running up to Normanton Gorse. This structure creates an excellent range of vegetation structures, from sparsely vegetated soil at the crop edge, through taller and more closed growth of forbs though to rank grassland and scrub. Such gradations in structure are known to allow the development of diverse assemblages of invertebrates.

The outer strip has developed a diverse flora of typical arable weeds, interspersed with much bare chalky ground. Plants present here include rape Brassica napus and other crucifers, fumitories Fumaria spp., small-flowered cranesbills Geranium sp., black medick Medicago lupulina, lesser trefoil Trifolium dubium, hop trefoil Trifolium campestre and changing forget-me-not Myosotis discolor. There are also scattered plants of taller ruderal species, such as musk thistle Carduus nutans and weld Reseda luteola.

The less disturbed grassland strip behind has tall tussocks of grass and some elements of a more established grassland flora, such as common bird’s-foot trefoil Lotus corniculatus, red and white clovers (T. pratense and T. repens respectively) and pyramidal orchid Anacamptis pyramidalis.

4.5.1. Key Invertebrate habitat features at Arable 1 The 2017 surveys have shown this site to have an extremely rich invertebrate fauna, with 12 key species recorded. The Nationally Scarce rove beetle Oxypoda exoleta and the Red Data Book longhorn beetle Paracorymbia fulva, have not been assigned to a habitat feature, as their ecology is very poorly understood. The remaining ten key species have been assigned to the three key habitat features listed below.

Early successional arable flora with bare ground (Zabrus tenebrioides Phalacrus championi; Catapion pubescens; Protapion filirostre; Tychius pusillus; Zacladus exiguus) The richest invertebrate fauna is associated with the regularly disturbed strip along the conservation headlands, where low growth of arable weeds and patches of bare ground. Four of the five species listed above are weevils that feed on yellow- flowered trefoils (C. pubescens, T. pusillus), black medick (Protapion filirostre) and dove’s-foor cranesbill (Z. exiguus). The remaining species, P. championi, feeds on smut fungi growing on grass and sedge flowers in dry, open conditions. The rich invertebrate fauna present in this habitat feature, including one species at its only known Wiltshire locality (T. pusillus) make the conservation headlands here an invertebrate habitat of county importance.

Tall ruderal growth of thistles (Psylliodes chalcomera; Larinus planus; Urophora solstitialis) Stands of thistles were found to have a rich invertebrate fauna. Of particular importance was the presence here of good stands of musk thistle, which supported two key species (P. chalcomera and U. solstitialis). Spear thistle plants had a breeding population of the large weevil Larinus planus. Though the latter is rated as Nationally Scarce, it seems to be very widely distribued, in this part of Wiltshire at least. Also, though not assigned to a habitat feature, a single specimen of the rare longhorn beetle Paracorymbia fulva was also found nectaring on thistles here.

Tall, closed-sward grassland (Rugilus similis) The Nationally Scarce rove beetle R. similis favours quite tall grassland where there is a well-deloped tussock structure and litter layer.

4.6. Arable 2

Though it was selected as an area of arable habitat, the field on which this survey block is centred is being restored by its owners, the National Trust, to chalk grassland. It lies immediately to the south of the A303, with Stonehenge only 0.5km to the north-east. A central grid reference for the site approximates to SU115416. It is adjacent to the Arable 2 block, which is situated on the southern and western side of Normanton Gorse (see previous Site Report). It was surveyed on the 27th and 29th June and 26th July. Two lines of pitfall traps were put out between the latter two dates. The site has two large barrows, one of which has been fenced to prevent damage by cattle trampling and consequently has a rank sward dominated by tall tussocks.

The restoration grassland has a moderately herb-rich sward, which is characterised by exceptional abundance of sainfoin Onobrychis viciifolia. At the time of the June visit, the sward was quite tall, but open with other prominent forbs including red clover Trifolium pratense, bird’s-foot trefoil Lotus corniculatus, tufted vetch Vicia cracca and a few plants of pyramidal orchid Anacamptis pyramidalis. By the time of the second visit, cattle had been introduced and had grazed down the sward considerably.

The area surveyed also included the track that runs up the eastern edge of the restoration field. This has rank grassland with tall herbs such as hogweed Heracleum sphondylium, wild parsnip Pastinaca sativa and common toadflax Linaria vulgaris and scattered hawthorn bushes Crataegus monogyna.

4.6.1. Key Invertebrate habitat features at Arable 2 Th 2017 surveys showed this site to have a moderately rich invertebrate fauna that includes four key species. The Red Data Book longhorn beetle Paracorymbia fulva, has not been assigned to a habitat feature, as its ecology is poorly understood. The remaining two key species have been assigned to the two key habitat features listed below.

Restoration chalk grassland (Hemitrichapion reflexum) It is not surprising that the invertebrate fauna associated with the area of grassland that is being restored to chalk grassland from arable is still only moderately diverse, as the process of re-colonisation by invertebrates will inevitably take time. So, for example the bumblebees and butterflies recorded here lacked any of the chalk grassland specialities known from old downland sites on the Wiltshire chalk. However, it was very encouraging that one beetle, the weevil H. reflexum, which is an extremely scarce inhabitant of high-quality chalk grassland, has already colonised the restored grassland. No doubt its successful colonisation has been facilitated by the great abundance of its foodplant, sainfoin, at this site currently.

Cattle dung (Aleochara discipennis) Cattle dung had a moderately diverse invertebrate fauna that included one key species, the Nationally Scarce rove beetle Aleochara discipennis.

Mole nests (Pycnota paradoxa) The rove beetle P. paradoxa is a specialist inhabitant of mole nests.

4.7. Arable 3

This was the third area of arable habitat selected for invertebrate survey in 2017. It comprised a very large field planted with cereals and sited immediately to the north of the A303. It is located approximately 2 km to the west of Amesbury and 1.5 km east of Stonehenge. A central grid reference for the site approximates to SU138421. It was surveyed on the 30th June and 26th July. Three lines of pitfall traps were set up here between the two survey dates. This site lies immediately to the west of Countess Cutting CWS (see following sub-section).

The arable margins around the edge of the field were the focus of survey effort. For the most part, these were very narrow; no more than a metre in width, though in a few places, especially along the boundary with the A303, they were somewhat wider. The arable weed flora is rather poorly developed, but in those places where there is a wider margin, there are stands of fumitory Fumaria spp., various crucifers and a range of other plants. A feature of the boundary with the A303 is the presence of good stands of marsh woundwort Stachys palustris. This boundary also has some stands of tall, ruderal vegetation amongst which stinging nettle Urtica dioica, spear thistle Cirsium vulgare and mugwort Artemisia vulgaris are frequent.

In its eastern quarter, the A303 field edge has a wide border of scrub and chalk grassland with the latter being quite species-rich in places and having forbs such as wild basil Clinopodium vulgare, St. John’s-wort Hypericum sp. common bird’s-foot trefoil Lotus corniculatus and pyramidal orchid Anacamptis pyramidalis amongst a range of calcicolous species.

4.7.1. Key Invertebrate habitat features at Arable 3 Th 2017 surveys have generated records of seven key species. The Red Data Book longhorn beetle Paracorymbia fulva, has not been assigned to a habitat feature, as its ecology is poorly understood. The remaining three key species have been assigned to the two key habitat features listed below.

Arable margins (Sirocalodes quercicola; Thamiocolus viduatus) Despite this area having rather narrow and poorly developed arable margins compared to the Arable 1 site, where broad ‘conservation headlands’ have been retained, it was still found to have quite a rich invertebrate fauna that included two key species. Of particular note was the presence here of the weevil S. quercicola on its foodplant, fumitory, on the somewhat wider arable margins adjacent to the A303, this being its first known Wiltshire locality. A second weevil, T. viduatus, was tapped from marsh woundwort growing in the same area.

Carrion (Ptomaphagus variicornis; Nicrophorus interruptus) Unfortunately, a shrew had died in one of the pitfall traps set up here and this attracted a range of carrion-associated beetle species including the above two that are of high rarity status.

Tall, closed-sward grassland (Sunius melanocephalus) Along the arable margin that lies adjacent to the A303 a single specimen of S. melanocephalus was recorded. It is usually found in places where there is dry, tussocky grassland with a well-developed litter layer and is likely to have originated in the stands of rank grassland that border the A303 at this point.

Ivy in hedgebanks (Anobium inexspectatum) The woodworm beetle A. inexspectatum bores in old ivy stems and the singleton tapped from low-growing herbage is thought to originate in ivy growing on the

4.8. Countess Cutting CWS

Countess Cutting CWS is a south-facing cutting on the northern verge of the A303. It comprises a narrow, steeply-sloping strip of approximately 250m in length, lying about 1 km to the north-west of Amesbury. A central grid reference for the site approximates to SU146421. It was surveyed on the 30th June and 26th July. One line of pitfall traps was set up here between the above survey dates. Countess Cutting lies immediately to the east of the Arable 3 site (see preceding sub-section).

Most of the area surveyed has an early-successional vegetation cover, in which sparse clumps of grasses and forbs such as common bird’s-foot trefoil Lotus corniculatus, kidney vetch Anthyllis vulneraria, horseshoe vetch Hippocrepis comosa and salad burnet Sanguisorba minor are interspersed with patches of bare chalk and flints.

On the margins of the cutting, where slopes are less steep, a more tussocky sward has developed, with abundant tor grass Brachypodium pinnatum and taller forbs such as St. John’s-wort Hypericum sp. and wild basil Clinopodium vulgare occurring in a mosaic with patches of mixed broadleaved scrub.

4.8.1. Key Invertebrate habitat features at Countess Cutting CWS The work carried out this year has produced records of eight key species, which have been assigned to three key habitat features. One key species, the weevil S. ruficornis feeds on poppies in ruderal and arable habitats, which are not present here. It is presumed to be a stray from outside the site and has therefore ot been assigned a key habitat feature in the following discussion.

Early successional chalk grassland with bare ground (Ophonus schaubergerianus; Licinus punctatulus; Lamprinodes saginatus; Phaeocedus braccatus) This was a late addition to the sampling programme and has been subject to less collecting effort than the other seven sample sites. Despite this, the early-successional mosaic of sparse vegetation and bare chalk and flints here has an extremely interesting invertebrate fauna that includes three key species. One of these, the Gnaphosid spider Phaeocedus braccatus, is an important record, this being a Nationally Rare species of warm, sparsely vegtated calcareous grassland and heaths that is only recorded from a few places in southern England.

Closed-sward chalk grassland (Licinus depressus; Lamprinodes saginatus; Mecinus circulatus) Though it was of somewhat lower importance than the early-successional habitat described in the preceding paragraph, closed-sward grassland chalk grassland on shallower slopes also had a rich invertebrate fauna that included two key species, the snail predator L. depressus and the plantain-feeding weevil M. circulatus.

Carrion (Nicrophorus interruptus) Four specimens of the carrion-associated sexton beetle N. interruptus were collected in a pitfall trap into which a shrew had unfortunately gained access.

5. CONCLUSIONS

All eight sites included in the 2017 survey have been shown to have a significant invertebrate fauna. Unsurprisingly, the three sample sites with high-quality habitats notified as SSSIs and SACs all have very important invertebrate assemblages: the two Till sites having diverse wetland faunas that are of at least county importance, and Parsonage Down NNR having an assemblage of regional significance associated with short-sward chalk grassland. However, the Countess Cutting CWS and Arable 1 sample sites also have an exceptionally rich invertebrate fauna found in association with early- to mid-successional chalk grassland habitats and wide arable margins respectively; these are also of at least county importance for invertebrates. That found on the wide arable margins at the Arable 1 site demonstrates that the adoption of conservation headlands here is certainly having benefits for invertebrates.

Diamond Wood and the Arable 2 and Arable 3 sample sites are somewhat less diverse, but still have important invertebrate assemblages of local to county importance.

6. ACKNOWLEDGEMENTS

Many thanks to Tom Shelley, Chloe Delgery and Paul Clack of ARUP, Bristol for setting up this contract, for facilitating access to all of the sample sites so efficiently and for comments on the draft version of this report. I would like to also thank all the owners and land managers who kindly gave permission for us to carry out this survey. Particular thanks are due to the owner of the Diamond Wood and Arable 1 sites, who took the time to accompany us during survey work and explain the conservation management work being undertaking here.

7. REFERENCES

ALEXANDER, K.N.A., DODD, S. & DENTON, J.S. 2014. A review of the scarce and threatened beetles of Great Britain. The darkling beetles and their allies: Aderidae, Anthicidae, Colydiidae, Melandryidae, Meloidae, Mordellidae, Mycetophagidae, Mycteridae, Oedemeridae, Pyrochroidae, Pythidae, Ripiphoridae, Salpingidae, Scraptiidae, Tenebrionidae & Tetratomidae (Tenebrionoidea less Ciidae). Species Status No.18. Natural England Commissioned Report. NECR148. Natural England, Peterborough.

DRAKE, C.M., LOTT, D.A., ALEXANDER, K.N.A. & WEBB, J. 2007. Surveying terrestrial and freshwater invertebrates for conservation evaluation. 1st Edition. Natural England: Peterborough.

FALK, S. J. 1991a. A review of the scarce and threatened bees, ants and wasps of Great Britain. Research and survey in nature conservation. No 35. NCC, Peterborough.

FALK, S.J. 1991b. A review of the scarce and threatened flies of Great Britain (Part 1). Nature Conservancy Council. Research and survey in nature conservation. No. 39. NCC, Peterborough.

FOSTER, G.N. 2010. A review of the scarce and threatened Coleoptera of Great Britain. Part 3 Water beetles of Great Britain. Species Status 1, Joint Nature Conservation Committee, Peterborough.

FOSTER, G. N., BILTON, D. T. & NELSON, B. H. 2016. Atlas of the predaceous water beetles (Hydradephaga) of Britain and Ireland. FSC Puplications, Telford.

HARVEY, P., DAVIDSON, M., DAWSON, I., FOWLES, A., HITCHCOCK, G., LEE, P., MERRETT, P., RUSSELL-SMITH, T. & SMITH, H. 2017. A review of the scarce and threatened spiders Araneae) of Great Britain: Species Status No.22. NRW Evidence Report No. 11. Natural Resources Wales, Bangor.

HYMAN, P.S. AND PARSONS, M.S. 1992. A review of the scarce and threatened Coleoptera of Great Britain. Part 1. UK nature conservation report. No 12. JNCC, Peterborough, 1994.

HYMAN, P.S. AND PARSONS, M.S. 1994. A review of the scarce and threatened Coleoptera of Great Britain. Part 2. UK nature conservation report. No 12. JNCC, Peterborough, 1994.

LUFF, M. L. 1996. Environmental assessments using ground beetles (Carabidae) and pitfall traps. In: Eyre, M. D., ed. Environmental monitoring, surveillance and conservation using invertebrates. EMS, Publications.

LUFF, M. L. 1998. Provisional atlas of the ground beetles (Coleoptera, Carabidae) of Britain. Biological Records Centre, Huntingdon.

TELFER, M.G. 2016. A review of the beetles of Great Britain: Ground Beetles (Carabidae). Species Status No.25. Natural England Commissioned Report. NECR189. Natural England, Peterborough.

APPENDIX 1: INVERTEBRATES RECORDED ALONG THE A303 AMESBURY TO BERWICK DOWN SCHEME IN 2017

Scientific Name English Name Status Site Gridref Date Method Habitat Physa fontinalis agg. Common Bladder Snail Upper Till SU081418 26/06/2017 Pond netting Shallow pond with stoneworts

Physa fontinalis agg Common Bladder Snail Upper Till SU081418 24/07/2017 Pond netting Shallow pond with stoneworts

Stagnicola palustris Marsh Pond Snail Upper Till SU081418 26/06/2017 Pond netting Shallow pond with stoneworts

Stagnicola palustris Marsh Pond Snail Upper Till SU081418 24/07/2017 Pond netting Shallow pond with stoneworts

Anisus leucostoma Button Ram's-horn Snail Upper Till SU081418 26/06/2017 Pond netting Shallow pond with stoneworts

Succinea putris Large Amber Snail Upper Till SU081416 26/06/2017 Tapping Vegetation on river edge

Oxychilus cellarius Garlic Snail Upper Till SU081416 24/07/2017 Dead wood Rotten poplar heartwood

Monacha cantiana Kentish Snail Upper Till SU080419 26/06/2017 Tapping Thistles and teasel on bank at field edge

Cepaea hortensis White-lipped Snail Upper Till SU080419 26/06/2017 Tapping Thistles and teasel on bank at field edge

Helix aspersa Garden Snail Upper Till SU080419 26/06/2017 Tapping Thistles and teasel on bank at field edge Dead hawthorn branches, scrub woodland Polyxenus lagurus A Millipede Upper Till SU080419 26/06/2017 Beating edge Banded Demoiselle Calopteryx splendens Damselfly Upper Till SU081419 26/06/2017 Spot search Scrub woodland edge

Lestes sponsa Common Emerald Damselfly Upper Till SU081418 26/06/2017 Spot search Shallow pond

Lestes sponsa Common Emerald Damselfly Upper Till SU081418 24/07/2017 Spot search Shallow pond Common Blue-tailed Ischnura elegans Damselfly Upper Till SU081418 26/06/2017 Spot search Shallow pond

Anax imperator Emperor Dragonfly Upper Till SU081418 26/06/2017 Spot search Shallow pond Black-tailed Skimmer Orthetrum cancellatum Dragonfly Upper Till SU081418 26/06/2017 Spot search Shallow pond

Sympetrum striolatum Common Darter Dragonfly Upper Till SU081416 26/06/2017 Spot search In flight, semi-improved pasture

Sympetrum striolatum Common Darter Dragonfly Upper Till SU080419 24/07/2017 Spot search Flowery bank at field edge

Metrioptera roeselii Roesel's Bush-cricket Upper Till SU081419 26/06/2017 Spot search Singing males, scrub woodland edge

Metrioptera roeselii Roesel's Bush-cricket Upper Till SU080419 24/07/2017 Spot search Singing males, flowery bank at field edge

Tetrix subulata Slender Groundhopper Upper Till SU081418 24/07/2017 Ground search Treading, bare mud at pond edge Common Green Omocestus viridulus Grasshopper Upper Till SU081419 26/06/2017 Spot search Singing males, rough semi-improved grassland Common Green Omocestus viridulus Grasshopper Upper Till SU080419 24/07/2017 Spot search Flowery bank at field edge

Chorthippus parallelus Meadow Grasshopper Upper Till SU081418 26/06/2017 Spot search Semi-improved grassland near pond

Chorthippus parallelus Meadow Grasshopper Upper Till SU081418 24/07/2017 Spot search Semi-improved grassland near pond

Scientific Name English Name Status Site Gridref Date Method Habitat Forficula auricularia Common Earwig Upper Till SU080419 26/06/2017 Ground search Tussocks, rough semi-improved grassland

Forficula auricularia Common Earwig Upper Till SU081416 24/07/2017 Beating Dead poplar branches

Ilyocoris cimicoides Saucer Bug Upper Till SU081418 26/06/2017 Pond netting Shallow pond with stoneworts

Saldula saltatoria A Shorebug Upper Till SU081418 26/06/2017 Spot search Bare mud at pond edge

Atractotomus mali A Mirid Bug Upper Till SU082418 26/06/2017 Beating Hawthorn at field edge

Agabus didymus A Diving Beetle Upper Till SU081418 26/06/2017 Pond netting Shallow pond with stoneworts

Agabus nebulosus A Diving Beetle Upper Till SU081418 26/06/2017 Pond netting Shallow pond with stoneworts

Agabus nebulosus A Diving Beetle Upper Till SU081418 24/07/2017 Pond netting Shallow pond with stoneworts

Hydroporus discretus A Diving Beetle Upper Till SU081416 26/06/2017 Tapping Veronica on river edge

Hydroporus palustris A Diving Beetle Upper Till SU081418 26/06/2017 Pond netting Shallow pond with stoneworts

Hygrotus inaequalis A Diving Beetle Upper Till SU081418 26/06/2017 Pond netting Shallow pond with stoneworts

Elaphrus riparius A Ground Beetle Upper Till SU081418 26/06/2017 Ground search Treading, bare mud at pond edge

Elaphrus riparius A Ground Beetle Upper Till SU081418 24/07/2017 Ground search Treading, bare mud at pond edge

Clivina fossor A Ground Beetle Upper Till SU080419 26/06/2017 Ground search Tussocks, rough semi-improved grassland

Bembidion articulatum A Ground Beetle Upper Till SU081418 26/06/2017 Ground search Treading, bare mud at pond edge

Bembidion articulatum A Ground Beetle Upper Till SU081418 24/07/2017 Ground search Treading, bare mud at pond edge

Bembidion dentellum A Ground Beetle Upper Till SU081418 26/06/2017 Ground search Treading, bare mud at pond edge

Bembidion dentellum A Ground Beetle Upper Till SU081418 24/07/2017 Ground search Treading, bare mud at pond edge

Pterostichus niger A Ground Beetle Upper Till SU080419 26/06/2017 Ground search Tussocks, rough semi-improved grassland

Pterostichus niger A Ground Beetle Upper Till SU081416 24/07/2017 Dead wood Rotten poplar heartwood

Pterostichus anthracinus A Ground Beetle NS. Upper Till SU081418 26/06/2017 Ground search Treading, bare mud at pond edge Pterostichus anthracinus A Ground Beetle NS. Upper Till SU080419 26/06/2017 Ground search Tussocks, rough semi-improved grassland Pterostichus minor A Ground Beetle Upper Till SU080419 26/06/2017 Ground search Tussocks, rough semi-improved grassland

Pterostichus vernalis A Ground Beetle Upper Till SU080419 26/06/2017 Ground search Tussocks, rough semi-improved grassland

Agonum marginatum A Ground Beetle Upper Till SU081418 26/06/2017 Ground search Treading, bare mud at pond edge

Agonum viduum A Ground Beetle Upper Till SU081418 26/06/2017 Ground search Treading, bare mud at pond edge

Stenolophus mixtus A Ground Beetle Upper Till SU081418 24/07/2017 Ground search Treading, bare mud at pond edge

Helophorus brevipalpis A Water Beetle Upper Till SU081418 26/06/2017 Pond netting Shallow pond with stoneworts

Helophorus brevipalpis A Water Beetle Upper Till SU081418 24/07/2017 Pond netting Shallow pond with stoneworts

Helophorus griseus A Water Beetle Upper Till SU081418 26/06/2017 Pond netting Shallow pond with stoneworts

Helophorus minutus A Water Beetle Upper Till SU081418 26/06/2017 Pond netting Shallow pond with stoneworts

Scientific Name English Name Status Site Gridref Date Method Habitat Hydrobius fuscipes A Water Beetle Upper Till SU081418 26/06/2017 Pond netting Shallow pond with stoneworts

Laccobius minutus A Water Beetle Upper Till SU081418 24/07/2017 Pond netting Shallow pond with stoneworts

Coelostoma orbiculare A Water Beetle Upper Till SU081418 24/07/2017 Ground search Treading, bare mud at pond edge

Cercyon marinus A Water Beetle Upper Till SU081418 26/06/2017 Ground search Treading, bare mud at pond edge

Ochthebius minimus A Water Beetle Upper Till SU081418 26/06/2017 Pond netting Shallow pond with stoneworts

Ochthebius minimus A Water Beetle Upper Till SU081418 24/07/2017 Pond netting Shallow pond with stoneworts

Abraeus perpusillus A Clown Beetle Upper Till SU081416 24/07/2017 Dead wood Rotten poplar heartwood Heracleum sphondylium flowers, scrub Eusphalerum luteum A Rove Beetle Upper Till SU080419 26/06/2017 Tapping woodland edge

Ischnosoma splendidum A Rove Beetle Upper Till SU080419 26/06/2017 Ground search Tussocks, rough semi-improved grassland

Sepedophilus nigripennis A Rove Beetle Upper Till SU081418 24/07/2017 Ground search Cut grass piles, semi-improved pasture

Sepedophilus testaceus A Rove Beetle NS. Upper Till SU081416 24/07/2017 Dead wood Rotten poplar heartwood Tachinus rufipes A Rove Beetle Upper Till SU081418 24/07/2017 Ground search Cut grass piles, semi-improved pasture

Tachyporus dispar A Rove Beetle Upper Till SU081418 24/07/2017 Ground search Cut grass piles, semi-improved pasture

Tachyporus hypnorum A Rove Beetle Upper Till SU080419 26/06/2017 Ground search Tussocks, rough semi-improved grassland

Tachyporus hypnorum A Rove Beetle Upper Till SU081418 24/07/2017 Ground search Cut grass piles, semi-improved pasture

Amischa analis A Rove Beetle Upper Till SU081416 24/07/2017 Beating Dead poplar branches

Dilacra luteipes A Rove Beetle Upper Till SU081418 24/07/2017 Ground search Treading, bare mud at pond edge

Falagrioma thoracica A Rove Beetle Upper Till SU080419 26/06/2017 Ground search Tussocks, rough semi-improved grassland

Cypha seminulum A Rove Beetle Upper Till SU081416 24/07/2017 Beating Dead poplar branches

Gnypeta ripicola A Rove Beetle N. Upper Till SU081418 24/07/2017 Ground search Treading, bare mud at pond edge Gnypeta rubrior A Rove Beetle Upper Till SU081418 24/07/2017 Ground search Treading, bare mud at pond edge

Thinonoma atra A Rove Beetle Upper Till SU081418 24/07/2017 Ground search Treading, bare mud at pond edge

Anotylus inustus A Rove Beetle Upper Till SU081418 26/06/2017 Ground search Treading, bare mud at pond edge

Anotylus rugosus A Rove Beetle Upper Till SU080419 26/06/2017 Ground search Tussocks, rough semi-improved grassland

Anotylus rugosus A Rove Beetle Upper Till SU081418 24/07/2017 Ground search Cut grass piles, semi-improved pasture

Platystethus cornutus A Rove Beetle Upper Till SU081418 24/07/2017 Ground search Treading, bare mud at pond edge

Carpelimus rivularis A Rove Beetle Upper Till SU081418 26/06/2017 Ground search Treading, bare mud at pond edge

Carpelimus rivularis A Rove Beetle Upper Till SU081418 24/07/2017 Ground search Treading, bare mud at pond edge

Stenus cicindeloides A Rove Beetle Upper Till SU081418 26/06/2017 Ground search Treading, bare mud at pond edge

Stenus boops A Rove Beetle Upper Till SU081418 24/07/2017 Ground search Treading, bare mud at pond edge

Stenus comma A Rove Beetle Upper Till SU081418 24/07/2017 Ground search Treading, bare mud at pond edge

Scientific Name English Name Status Site Gridref Date Method Habitat Paederus fuscipes A Rove Beetle Upper Till SU081418 24/07/2017 Ground search Treading, bare mud at pond edge

Rugilus similis A Rove Beetle N. Upper Till SU081418 24/07/2017 Ground search Cut grass piles, semi-improved pasture Bisnius fimetarius A Rove Beetle Upper Till SU081416 24/07/2017 Dead wood Rotten poplar heartwood

Gabrius breviventer A Rove Beetle Upper Till SU081418 24/07/2017 Ground search Treading, bare mud at pond edge

Philonthus albipes A Rove Beetle Upper Till SU081418 24/07/2017 Dung Cow dung, semi-improved pasture

Philonthus atratus A Rove Beetle Na. Upper Till SU081418 24/07/2017 Ground search Treading, bare mud at pond edge Philonthus carbonarius A Rove Beetle Upper Till SU081418 24/07/2017 Spot search Resting on dock leaf, semi-improved pasture

Philonthus cruentatus A Rove Beetle Upper Till SU081418 24/07/2017 Ground search Semi-improved pasture

Philonthus intermedius A Rove Beetle Upper Till SU081418 24/07/2017 Dung Cow dung, semi-improved pasture

Philonthus micans A Rove Beetle Upper Till SU081418 26/06/2017 Ground search Treading, bare mud at pond edge

Philonthus parvicornis A Rove Beetle pNS. Upper Till SU081418 24/07/2017 Dung Cow dung, semi-improved pasture Treading, bare mud at pond edge. Including Philonthus quisquiliarius A Rove Beetle Upper Till SU081418 26/06/2017 Ground search var. inquinatus

Philonthus quisquiliarius A Rove Beetle Upper Till SU081418 24/07/2017 Ground search Treading, bare mud at pond edge

Philonthus sanguinolentus A Rove Beetle Upper Till SU081418 24/07/2017 Ground search Cut grass piles, semi-improved pasture

Philonthus varians A Rove Beetle Upper Till SU081418 24/07/2017 Dung Cow dung, semi-improved pasture

Quedius cruentus A Rove Beetle Upper Till SU081416 24/07/2017 Dead wood Rotten poplar heartwood

Quedius nigrocaeruleus A Rove Beetle Nb. Upper Till SU081416 24/07/2017 Dead wood Rotten poplar heartwood Quedius schatzmayri A Rove Beetle Upper Till SU080419 26/06/2017 Ground search Tussocks, rough semi-improved grassland

Ontholestes murinus A Rove Beetle Upper Till SU081418 24/07/2017 Dung Cow dung, semi-improved pasture

Ontholestes tessellatus A Rove Beetle Upper Till SU081418 24/07/2017 Dung Cow dung, semi-improved pasture

Gyrohypnus fracticornis A Rove Beetle Upper Till SU081418 24/07/2017 Dung Cow dung, semi-improved pasture

Xantholinus linearis A Rove Beetle Upper Till SU081418 24/07/2017 Ground search Cut grass piles, semi-improved pasture

Xantholinus longiventris A Rove Beetle Upper Till SU081418 24/07/2017 Ground search Cut grass piles, semi-improved pasture

Dorcus parallelepipedus Lesser Stag Beetle Upper Till SU077415 26/06/2017 Dead wood Rotten heartwood, old ash

Heterocerus fenestratus A Heterocerid Beetle Upper Till SU081418 26/06/2017 Ground search Treading, bare mud at pond edge

Heterocerus fenestratus A Heterocerid Beetle Upper Till SU081418 24/07/2017 Ground search Treading, bare mud at pond edge

Hemicrepidius hirtus A Click Beetle Upper Till SU082418 26/06/2017 Beating Hawthorn at field edge

Melanotus villosus agg. A Click Beetle Upper Till SU081416 24/07/2017 Dead wood Larvae, rotten willow heartwood

Rhagonycha fulva A Soldier Beetle Upper Till SU080419 26/06/2017 Spot search Rank grassland, scrub woodland edge

Rhagonycha fulva A Soldier Beetle Upper Till SU081418 24/07/2017 Spot search Flowers at pond margin

Malachius bipustulatus A Malachite Beetle Upper Till SU081416 26/06/2017 Tapping Vegetation on river edge

Scientific Name English Name Status Site Gridref Date Method Habitat Meligethes aeneus A Pollen Beetle Upper Till SU082418 26/06/2017 Tapping Ballota nigra at field edge

Meligethes ruficornis A Pollen Beetle Upper Till SU082418 26/06/2017 Tapping Ballota nigra at field edge

Olibrus aeneus A Phalacrid Beetle Upper Till SU081418 24/07/2017 Spot search Flowers at pond margin

Cryptophagus scanicus A Cryptophagid Beetle Upper Till SU081416 24/07/2017 Bird's nest Old nest in poplar

Coccidula rufa A Ladybird Upper Till SU081418 26/06/2017 Ground search Treading, bare mud at pond edge Coccinella septempunctata 7-spot Ladybird Upper Till SU081416 26/06/2017 Spot search Vegetation on river edge Coccinella septempunctata 7-spot Ladybird Upper Till SU081418 24/07/2017 Spot search Vegetation at pond margin

Harmonia axyridis Harlequin Ladybird Upper Till SU081416 26/06/2017 Beating Black poplar foliage Subcoccinella vigintiquattuorpunctata 24-spot Ladybird Upper Till SU080419 26/06/2017 Tapping Thistles and teasel on bank at field edge

Prionychus ater A Darkling Beetle Upper Till SU081416 24/07/2017 Dead wood Larvae, rotten willow heartwood

Oedemera nobilis An Oedemerid Beetle Upper Till SU080419 26/06/2017 Tapping Thistles and teasel on bank at field edge Dead hawthorn branches, scrub woodland Lissodema denticolle A Salpingid Beetle Nb. Upper Till SU080419 26/06/2017 Beating edge Salpingus planirostris A Salpingid Beetle Upper Till SU081416 26/06/2017 Beating Dead poplar branches

Salpingus planirostris A Salpingid Beetle Upper Till SU081416 24/07/2017 Beating Dead poplar branches

Paracorymbia fulva A Longhorn Beetle RDB3 Upper Till SU080419 24/07/2017 Spot search Flowery bank at field edge Phaedon armoraciae A Leaf Beetle Upper Till SU081416 26/06/2017 Tapping Rorippa nasturtium-aquaticum on river edge

Prasocuris phellandrii A Leaf Beetle Upper Till SU081416 26/06/2017 Tapping Veronica on river edge

Galerucella sagittariae A Leaf Beetle Upper Till SU081418 26/06/2017 Tapping Polygonum amphibium at pond edge Neocrepidodera transversa A Flea Beetle Upper Till SU080419 26/06/2017 Tapping Thistles and teasel on bank at field edge Neocrepidodera transversa A Flea Beetle Upper Till SU082417 24/07/2017 Spot search Bank with remnant chalk grassland Tatianaerhynchites aequatus A Weevil Upper Till SU082418 26/06/2017 Beating Hawthorn at field edge

Anthonomus pedicularius A Weevil Upper Till SU082418 26/06/2017 Beating Hawthorn at field edge

Dorytomus longimanus A Weevil Upper Till SU081416 24/07/2017 Bird's nest Old nest in poplar

Gymnetron villosulum A Weevil Nb. Upper Till SU081416 26/06/2017 Tapping Veronica on river edge Ceutorhynhus typhae A Weevil Upper Till SU081416 26/06/2017 Tapping Rorippa nasturtium-aquaticum on river edge

Drupenatus nasturtii A Weevil Nb. Upper Till SU081416 26/06/2017 Tapping Rorippa nasturtium-aquaticum on river edge Larinus planus A Weevil Nb. Upper Till SU082418 26/06/2017 Spot search Cirsium vulgare, semi-improved grassland Scolytus rugulosus A Weevil Upper Till SU082418 26/06/2017 Beating Hawthorn at field edge

Scientific Name English Name Status Site Gridref Date Method Habitat Thymelicus sylvestris Small Skipper Upper Till SU082417 24/07/2017 Spot search Bank with remnant chalk grassland

Ochlodes sylvanus Large Skipper Upper Till SU081416 26/06/2017 Spot search In flight, semi-improved pasture

Gonepteryx rhamni Brimstone Upper Till SU080419 24/07/2017 Spot search Flowery bank at field edge

Pieris brassicae Large White Upper Till SU081416 26/06/2017 Spot search In flight, semi-improved pasture

Pieris brassicae Large White Upper Till SU080419 24/07/2017 Spot search Flowery bank at field edge

Pieris rapae Small White Upper Till SU080419 26/06/2017 Spot search In flight, by bank at field edge

Pieris napi Green-veined White Upper Till SU080419 26/06/2017 Spot search In flight, by bank at field edge

Aricia agestis Brown Argus Upper Till SU080419 24/07/2017 Spot search Flowery bank at field edge

Polyommatus icarus Common Blue Upper Till SU082417 24/07/2017 Spot search Bank with remnant chalk grassland

Inachis io Peacock Upper Till SU080419 24/07/2017 Spot search Flowery bank at field edge

Vanessa atalanta Red Admiral Upper Till SU080419 24/07/2017 Spot search Flowery bank at field edge

Aglais urticae Small Tortoiseshell Upper Till SU081418 26/06/2017 Spot search Flowers at pond margin

Polygonia c-album Comma Upper Till SU081416 24/07/2017 Spot search Flowers at field edge

Melanargia galathea Marbled White Upper Till SU082418 26/06/2017 Spot search Cirsium vulgare, semi-improved grassland

Melanargia galathea Marbled White Upper Till SU080419 24/07/2017 Spot search Flowery bank at field edge

Pyronia tithonus Gatekeeper Upper Till SU080419 24/07/2017 Spot search Flowery bank at field edge

Maniola jurtina Meadow Brown Upper Till SU081418 26/06/2017 Spot search In flight, rough grassland near pond

Maniola jurtina Meadow Brown Upper Till SU081418 24/07/2017 Spot search Flowers at pond margin

Aphantopus hyperantus Ringlet Upper Till SU082418 26/06/2017 Spot search Cirsium vulgare, semi-improved grassland

Orgyia antiqua Vapourer Moth Upper Till SU082418 26/06/2017 Beating Larva, hawthorn at field edge

Tyria jacobaeae Cinnabar Moth Upper Till SU081418 26/06/2017 Spot search In flight, pond edge

Tyria jacobaeae Cinnabar Moth Upper Till SU081418 24/07/2017 Spot search Larvae on ragwort, pond margin

Eremobia ochroleuca Dusky Sallow Moth Upper Till SU080419 24/07/2017 Spot search Flowery bank at field edge

Haematopota pluvialis A Horsefly Upper Till SU080419 26/06/2017 Ground search Tussocks, rough semi-improved grassland

Empis livida A Dance Fly Upper Till SU080419 26/06/2017 Spot search Thistles and teasel on bank at field edge

Episyrphus balteatus A Hoverfly Upper Till SU081418 26/06/2017 Spot search Flowers at pond margin

Eristalinus sepulchralis A Hoverfly Upper Till SU081418 26/06/2017 Spot search In flight, pond edge

Syritta pipiens A Hoverfly Upper Till SU081418 24/07/2017 Spot search Flowers at pond margin

Sepedon sphegea A Snail-killing Fly Upper Till SU081418 26/06/2017 Spot search In flight, pond edge

Scathophaga stercoraria A Dung Fly Upper Till SU081416 26/06/2017 Dung Cow dung, semi-improved pasture

Scathophaga stercoraria A Dung Fly Upper Till SU081416 24/07/2017 Dung Cow dung, semi-improved pasture

Scientific Name English Name Status Site Gridref Date Method Habitat Mesembrina meridiana A Housefly Upper Till SU082418 26/06/2017 Dung Cow dung, semi-improved pasture

Lasius flavus Yellow Meadow Ant Upper Till SU081418 24/07/2017 Dung Cow dung, semi-improved pasture

Bombus terrestris Buff-tailed Bumblebee Upper Till SU082418 26/06/2017 Spot search Cirsium vulgare, semi-improved grassland

Bombus terrestris Buff-tailed Bumblebee Upper Till SU081418 24/07/2017 Spot search Flowers at pond margin

Bombus lapidaius Red-tailed Bumblebee Upper Till SU082418 26/06/2017 Spot search Cirsium vulgare, semi-improved grassland

Bombus pascuorum Common Carder Bumblebee Upper Till SU082418 26/06/2017 Spot search Cirsium vulgare, semi-improved grassland

Bombus pascuorum Common Carder Bumblebee Upper Till SU081416 24/07/2017 Spot search Flowers at field edge

Dicranopalpus ramosus A Harvestman Spider Upper Till SU081416 24/07/2017 Bird's nest Old nest in poplar

Araneus diadematus Garden Cross Spider Upper Till SU081416 24/07/2017 Spot search Flowers at field edge

Nuctenea umbratica An Orb-weaving Spider Upper Till SU081416 24/07/2017 Beating Dead poplar branches Dead hawthorn branches, scrub woodland Salticus cingulatus A Jumping Spider Upper Till SU080419 26/06/2017 Beating edge

Oniscus asellus Common Shiny Woodlouse Upper Till SU081416 24/07/2017 Dead wood Rotten poplar heartwood

Porcellio scaber Common Rough Woodlouse Upper Till SU081416 26/06/2017 Dead wood Under dead poplar bark

Porcellio scaber Common Rough Woodlouse Upper Till SU081416 24/07/2017 Dead wood Rotten poplar heartwood

Armadillidium vulgare Common Pill Woodlouse Upper Till SU081416 24/07/2017 Dead wood Rotten poplar heartwood

Galba truncatula Dwarf Pond Snail Lower Till SU077397 25/07/2017 Ground search Under submerged wood in river

Stagnicola palustris Marsh Pond Snail Lower Till SU077397 27/06/2017 Aquatic Under stones and branches in river

Stagnicola palustris Marsh Pond Snail Lower Till SU077397 25/07/2017 Ground search Under submerged wood in river

Radix balthica Wandering Snail Lower Till SU077397 27/06/2017 Aquatic Under stones and branches in river

Radix balthica Wandering Snail Lower Till SU077397 25/07/2017 Ground search Under submerged wood in river

Anisus leucostoma Button Ram's-horn Snail Lower Till SU077397 27/06/2017 Aquatic Under stones and branches in river

Succinea putris Large Amber Snail Lower Till SU075396 26/06/2017 Spot search Wet woodland edge

Succinea putris Large Amber Snail Lower Till SU076395 25/07/2017 Sweeping Tall marginal vegetation on riverbank

Arion ater agg. Large Black Slug Lower Till SU075396 26/06/2017 Ground search Under branch, damp neutral grassland

Arion ater agg. Large Black Slug Lower Till SU076395 25/07/2017 Sweeping Tall marginal vegetation on riverbank

Agriolimax reticulatus Reticulated Slug Lower Till SU075396 26/06/2017 Ground search Under branch, damp neutral grassland

Monacha cantiana Kentish Snail Lower Till SU077396 25/07/2017 Spot search Marshy grassland

Ashfordia granulata Silky Snail Lower Till SU076395 25/07/2017 Sweeping Tall marginal vegetation on riverbank

Trichia striolata Strawberry Snail Lower Till SU076395 25/07/2017 Sweeping Tall marginal vegetation on riverbank

Helix aspersa Garden Snail Lower Till SU077396 25/07/2017 Spot search Marshy grassland

Polycelis felina A Flatworm Lower Till SU077396 27/06/2017 Sieving Springhead

Scientific Name English Name Status Site Gridref Date Method Habitat Piscicola geometra Fish Leech Lower Till SU077397 27/06/2017 Aquatic Under stones and branches in river Banded Demoiselle Calopteryx splendens Damselfly Lower Till SU075396 26/06/2017 Spot search In flight, wet woodland edge

Aeshna grandis Brown Hawker Dragonfly Lower Till SU077396 25/07/2017 Spot search Rough, marshy grassland

Sympetrum sanguineum Bloody Darter Dragonfly Lower Till SU077396 25/07/2017 Spot search Rough, marshy grassland

Sympetrum striolatum Common Darter Drgonfly Lower Till SU077396 25/07/2017 Spot search Rough, marshy grassland

Serratella ignita Blue-winged Olive Mayfly Lower Till SU077397 27/06/2017 Aquatic Under stones and branches in river

Serratella ignita Blue-winged Olive Mayfly Lower Till SU077397 25/07/2017 Ground search Under submerged wood in river

Metrioptera roeselii Roesel's Bush-cricket Lower Till SU075396 26/06/2017 Sweeping Damp neutral grassland

Metrioptera roeselii Roesel's Bush-cricket Lower Till SU077396 25/07/2017 Spot search Rough, marshy grassland

Tetrix subulata Slender Groundhopper Lower Till SU075396 26/06/2017 Spot search Damp neutral grassland

Chorthippus parallelus Meadow Grasshopper Lower Till SU077396 25/07/2017 Spot search Marshy grassland Common Green Omocestus viridulus Grasshopper Lower Till SU075396 26/06/2017 Sweeping Damp neutral grassland Common Green Omocestus viridulus Grasshopper Lower Till SU077396 25/07/2017 Spot search Rough, marshy grassland

Forficula auricularia Common Earwig Lower Till SU075395 27/06/2017 Dead wood Beating dead willow branches, wet woodland

Forficula auricularia Common Earwig Lower Till SU077399 25/07/2017 Beating On willow

Velia caprai A Water Cricket Lower Till SU077397 27/06/2017 Aquatic Under stones and branches in river

Velia caprai A Water Cricket Lower Till SU077397 25/07/2017 Ground search Under submerged wood in river

Nepa cinerea Water Scorpion Lower Till SU076395 27/06/2017 Ground search Treading, brown mosses at ditch edge

Pentatoma rufipes Forest Shieldbug Lower Till SU075396 26/06/2017 Beating Grey willow, wet woodland

Aneurus avenius A Flatbug Lower Till SU075395 27/06/2017 Dead wood Beating dead willow branches, wet woodland

Orectochilus villosus Hairy Whirligig Beetle Lower Till SU077397 27/06/2017 Aquatic Under stones and branches in river

Orectochilus villosus Hairy Whirligig Beetle Lower Till SU077397 25/07/2017 Ground search Under submerged wood in river

Agabus didymus A Diving Beetle Lower Till SU077396 27/06/2017 Sieving Springhead

Agabus nebulosus A Diving Beetle Lower Till SU077396 25/07/2017 Sieving Chalk spring, Rough marshy grassland

Platambus maculatus A Diving Beetle Lower Till SU077397 27/06/2017 Aquatic Under stones and branches in river

Platambus maculatus A Diving Beetle Lower Till SU077397 25/07/2017 Ground search Under submerged wood in river

Ilybius fuliginosus A Diving Beetle Lower Till SU076395 27/06/2017 Ground search Treading, brown mosses at ditch edge

Hydroporus ferrugineus A Diving Beetle NS. Lower Till SU077396 27/06/2017 Sieving Springhead Hydroporus ferrugineus A Diving Beetle NS. Lower Till SU077396 25/07/2017 Sieving Chalk spring, Rough marshy grassland Hydroporus marginatus A Diving Beetle NS. Lower Till SU077397 27/06/2017 Aquatic Under stones and branches in river

Scientific Name English Name Status Site Gridref Date Method Habitat Oreodytes sanmarkii A Diving Beetle Lower Till SU077397 25/07/2017 Ground search Under submerged wood in river

Elaphrus cupreus A Ground Beetle Lower Till SU076395 27/06/2017 Ground search Treading, brown mosses at ditch edge

Elaphrus cupreus A Ground Beetle Lower Till SU077397 25/07/2017 Ground search Bare mud, river edge

Elaphrus riparius A Ground Beetle Lower Till SU077397 25/07/2017 Ground search Bare mud, river edge

Loricera pilicornis A Ground Beetle Lower Till SU077397 27/06/2017 Ground search Bare mud, river edge

Dyschirius aeneus A Ground Beetle Lower Till SU077397 27/06/2017 Ground search Bare mud, river edge

Bembidion dentellum A Ground Beetle Lower Till SU076395 27/06/2017 Ground search Treading, brown mosses at ditch edge

Bembidion dentellum A Ground Beetle Lower Till SU077397 25/07/2017 Ground search Bare mud, river edge Bembidion qudrimaculatum A Ground Beetle Lower Till SU077397 25/07/2017 Ground search Bare mud, river edge

Bembidion guttula A Ground Beetle Lower Till SU077397 27/06/2017 Ground search Bare mud, river edge

Pterostichus minor A Ground Beetle Lower Till SU077397 25/07/2017 Ground search Bare mud, river edge

Pterostichus nigrita A Ground Beetle Lower Till SU077397 25/07/2017 Ground search Bare mud, river edge

Agonum viduum A Ground Beetle Lower Till SU077397 27/06/2017 Ground search Bare mud, river edge

Stenolophus mixtus A Ground Beetle Lower Till SU077397 27/06/2017 Ground search Bare mud, river edge

Dromius meridionalis A Ground Beetle Lower Till SU077399 25/07/2017 Beating Dead willow branches

Helophorus brevipalpis A Water Beetle Lower Till SU077396 27/06/2017 Sieving Springhead

Helophorus brevipalpis A Water Beetle Lower Till SU077397 25/07/2017 Ground search Bare mud, river edge

Laccobius bipunctatus A Water Beetle Lower Till SU077397 27/06/2017 Ground search Bare mud, river edge

Coelostoma orbiculare A Water Beetle Lower Till SU077396 27/06/2017 Sieving Springhead

Cercyon marinus A Water Beetle Lower Till SU077397 25/07/2017 Ground search Bare mud, river edge

Cercyon sternalis A Water Beetle Lower Till SU077397 27/06/2017 Aquatic Under stones and branches in river

Cercyon tristis A Water Beetle Lower Till SU077397 27/06/2017 Ground search Bare mud, river edge

Cercyon tristis A Water Beetle Lower Till SU077397 25/07/2017 Ground search Bare mud, river edge

Ochthebius minimus A Water Beetle Lower Till SU077396 27/06/2017 Sieving Springhead

Ochthebius minimus A Water Beetle Lower Till SU077396 25/07/2017 Sieving Chalk spring, Rough marshy grassland

Oiceoptoma thoracicum A Carrion Beetle Lower Till SU075395 27/06/2017 Spot search On vegetation, wet woodland

Rybaxis longicornis A Rove Beetle Lower Till SU077397 27/06/2017 Ground search Bare mud, river edge

Amischa analis A Rove Beetle Lower Till SU075396 26/06/2017 Sweeping Damp neutral grassland

Philhygra volans A Rove Beetle Lower Till SU077397 27/06/2017 Ground search Bare mud, river edge

Philhygra volans A Rove Beetle Lower Till SU077397 25/07/2017 Ground search Bare mud, river edge

Deinopsis erosa A Rove Beetle Lower Till SU077397 27/06/2017 Ground search Bare mud, river edge

Scientific Name English Name Status Site Gridref Date Method Habitat Deinopsis erosa A Rove Beetle Lower Till SU077397 25/07/2017 Ground search Bare mud, river edge

Cypha seminulum A Rove Beetle Lower Till SU075395 27/06/2017 Dead wood Beating dead willow branches, wet woodland

Myllaena intermedia A Rove Beetle Lower Till SU077397 27/06/2017 Ground search Bare mud, river edge

Gnypeta rubrior A Rove Beetle Lower Till SU077397 25/07/2017 Ground search Bare mud, river edge

Anotylus rugosus A Rove Beetle Lower Till SU077396 27/06/2017 Ground search Litter around springhead

Platystethus cornutus A Rove Beetle Lower Till SU077396 27/06/2017 Ground search Litter around springhead

Platystethus cornutus A Rove Beetle Lower Till SU077397 25/07/2017 Ground search Bare mud, river edge

Carpelimus erichsoni A Rove Beetle Lower Till SU077397 27/06/2017 Ground search Bare mud, river edge

Carpelimus rivularis A Rove Beetle Lower Till SU077397 25/07/2017 Ground search Bare mud, river edge

Stenus cicindeloides A Rove Beetle Lower Till SU077397 27/06/2017 Ground search Bare mud, river edge Sedge litter, sedge-bed at ditch edge, marshy Stenus cicindeloides A Rove Beetle Lower Till SU077396 25/07/2017 Ground search grassland Sedge litter, sedge-bed at ditch edge, marshy Stenus latifrons A Rove Beetle Lower Till SU077396 25/07/2017 Ground search grassland

Stenus butrintensis A Rove Beetle N. Lower Till SU077397 25/07/2017 Ground search Sparganium erectum by river Sedge litter, sedge-bed at ditch edge, marshy Stenus picipennis A Rove Beetle Lower Till SU077396 25/07/2017 Ground search grassland Sedge litter, sedge-bed at ditch edge, marshy Stenus pubescens A Rove Beetle Lower Till SU077396 25/07/2017 Ground search grassland

Stenus boops A Rove Beetle Lower Till SU077396 27/06/2017 Ground search Litter around springhead

Stenus boops A Rove Beetle Lower Till SU077397 25/07/2017 Ground search Bare mud, river edge

Gabrius breviventer A Rove Beetle Lower Till SU077397 27/06/2017 Ground search Bare mud, river edge

Philonthus quisquiliarius A Rove Beetle Lower Till SU077397 27/06/2017 Ground search Bare mud, river edge Bare mud, river edge. Forma typica and var Philonthus quisquiliarius A Rove Beetle Lower Till SU077397 25/07/2017 Ground search inquinatus

Philonthus umbratilis A Rove Beetle Lower Till SU077397 27/06/2017 Ground search Bare mud, river edge Rorippa nasturtium-aquaticum, rough marshy Scirtes hemisphaerica A Marsh Beetle Lower Till SU077396 25/07/2017 Tapping grassland

Elmis aenea A Riffle Beetle Lower Till SU077397 27/06/2017 Aquatic Under stones and branches in river

Heterocerus fenestratus A Heterocerid Beetle Lower Till SU077397 25/07/2017 Ground search Bare mud, river edge

Hemicepidius hirtus A Click Beetle Lower Till SU076395 26/06/2017 Sweeping Carex riparia fen meadow

Rhagonycha fulva A Soldier Beetle Lower Till SU077396 27/06/2017 Spot search Carex riparia fen meadow

Rhagonycha fulva A Soldier Beetle Lower Till SU077396 25/07/2017 Spot search Rough, marshy grassland

Anobium punctatum A Woodworm Beetle Lower Till SU075395 27/06/2017 Dead wood Beating dead willow branches, wet woodland

Kateretes rufilabris A Pollen Beetle Lower Till SU077396 25/07/2017 Ground search Sedge litter, sedge-bed at ditch edge

Scientific Name English Name Status Site Gridref Date Method Habitat Stachys sylvatica, tall marginal vegetation on Meligethes brunnicornis A Pollen Beetle Lower Till SU076395 25/07/2017 Tapping riverbank

Telmatophilus caricis A Cryptophagid Beetle Lower Till SU077397 26/06/2017 Spot search Sparganium erectum by river

Dacne bipustulata An Erotylid Beetle Lower Till SU077399 25/07/2017 Fungi Daedaleopsis confragosa, on willow

Coccidula rufa A Ladybird Lower Till SU077397 27/06/2017 Ground search Bare mud, river edge Rorippa nasturtium-aquaticum, rough marshy Coccidula rufa A Ladybird Lower Till SU077396 25/07/2017 Tapping grassland Propylea quatturodecimpunctata 14-spot Ladybird Lower Till SU076395 26/06/2017 Sweeping Carex riparia fen meadow Propylea quatturodecimpunctata 14-spot Ladybird Lower Till SU076395 25/07/2017 Sweeping Tall marginal vegetation on riverbank Calvia quattuordecimguttata Cream-spot Ladybird Lower Till SU075396 26/06/2017 Beating Grey willow, wet woodland

Harmonia axyridis Harlequin Ladybird Lower Till SU075396 26/06/2017 Beating Grey willow, wet woodland

Harmonia axyridis Harlequin Ladybird Lower Till SU077399 25/07/2017 Beating On willow Subcoccinella vigintiquattuorpunctata 24-spot Ladybird Lower Till SU076395 25/07/2017 Sweeping Tall marginal vegetation on riverbank Mycetophagus multipunctatus A Fungus Beetle Lower Till SU077399 25/07/2017 Fungi Daedaleopsis confragosa, on willow

Oedemera nobilis An Oedemerid Beetle Lower Till SU076395 26/06/2017 Sweeping Carex riparia fen meadow

Donacia simplex A Reed Beetle Lower Till SU077397 26/06/2017 Spot search Sparganium erectum by river

Chrysolina herbacea A Leaf Beetle Lower Till SU075396 26/06/2017 Spot search Mentha aquatica, wet woodland edge

Chrysolina herbacea A Leaf Beetle Lower Till SU077396 25/07/2017 Tapping Mentha aquatica, Rough marshy grassland Mentha aquatica, tall marginal vegetation on Chrysolina polita A Leaf Beetle Lower Till SU076395 25/07/2017 Tapping riverbank

Phaedon tumidulus A Leaf Beetle Lower Till SU076395 26/06/2017 Sweeping Carex riparia fen meadow

Phaedon tumidulus A Leaf Beetle Lower Till SU077397 25/07/2017 Ground search Bare mud, river edge Rorippa nasturtium-aquaticum, rough marshy Prasocuris junci A Leaf Beetle Lower Till SU077396 25/07/2017 Tapping grassland

Aphthona nonstriata A Flea Beetle Lower Till SU075396 26/06/2017 Spot search Iris pseudacorus, wet woodland edge

Aphthona nonstriata A Flea Beetle Lower Till SU077396 25/07/2017 Tapping Iris pseudacorus, Rough marshy grassland

Psylliodes chrysocephala A Flea Beetle Lower Till SU076395 26/06/2017 Sweeping Carex riparia fen meadow

Apion cruentatum A Weevil Lower Till SU076395 26/06/2017 Sweeping Carex riparia fen meadow

Oxystoma cerdo A Weevil Nb. Lower Till SU076395 26/06/2017 Tapping Vicia cracca, Carex riparia fen meadow Catapion seniculus A Weevil Lower Till SU075396 26/06/2017 Sweeping Damp neutral grassland

Protapion apricans A Weevil Lower Till SU075396 26/06/2017 Sweeping Damp neutral grassland

Scientific Name English Name Status Site Gridref Date Method Habitat Rorippa nasturtium-aquaticum, rough marshy Protapion fulvipes A Weevil Lower Till SU077396 25/07/2017 Tapping grassland Propylea quatturodecimpunctata A Weevil Lower Till SU077397 26/06/2017 Sweeping Carex riparia fen meadow Veronica anagallis-aquatica, bare mud, river Gymnetron villosulum A Weevil Nb. Lower Till SU077397 27/06/2017 Ground search edge Veronica anagallis-aquatica, bare mud, river Gymnetron villosulum A Weevil Nb. Lower Till SU077397 25/07/2017 Tapping edge Tychius picirostris A Weevil Lower Till SU075396 26/06/2017 Sweeping Damp neutral grassland Ceutorhynchus pallidactylus A Weevil Lower Till SU076395 26/06/2017 Tapping Rorippa nasturtium-aquaticum, ditch edge

Ceutorhynchus typhae A Weevil Lower Till SU076395 26/06/2017 Tapping Rorippa nasturtium-aquaticum, ditch edge Mentha aquatica, tall marginal vegetation on Datonychus melanostictus A Weevil Lower Till SU076395 25/07/2017 Tapping riverbank

Drupenatus nasturtii A Weevil Nb. Lower Till SU076395 26/06/2017 Tapping Rorippa nasturtium-aquaticum, ditch edge Nedyus quadrimaculatus A Weevil Lower Till SU076395 25/07/2017 Sweeping Tall marginal vegetation on riverbank

Poophagus sisymbrii A Weevil Lower Till SU076395 26/06/2017 Tapping Rorippa nasturtium-aquaticum, ditch edge Stachys palustris, tall marginal vegetation on Thamiocolus viduatus A Weevil Nb. Lower Till SU076395 25/07/2017 Tapping riverbank Rhinoncus perpendicularis A Weevil Lower Till SU077398 25/07/2017 Tapping Persicaria hydropiper, rank, ruderal grassland

Euophryum confine A Weevil Lower Till SU077399 25/07/2017 Fungi Daedaleopsis confragosa, on willow

Larinus planus A Weevil Nb. Lower Till SU076396 26/06/2017 Spot search Cirsium vulgare, rank, ruderal grassland Anthophila fabriciana Nettle Tap Moth Lower Till SU075396 26/06/2017 Spot search In flight, damp neutral grassland

Thymelicus sylvestris Small Skipper Lower Till SU077398 25/07/2017 Spot search Rank, ruderal grassland

Ochlodes venata Large Skipper Lower Till SU075396 26/06/2017 Spot search In flight, damp neutral grassland

Ochlodes venata Large Skipper Lower Till SU077396 25/07/2017 Spot search Rough, marshy grassland

Gonepteryx rhamni Brimstone Lower Till SU077396 25/07/2017 Spot search Marshy grassland

Pieris rapae Small White Lower Till SU075396 26/06/2017 Spot search In flight, damp neutral grassland

Pieris napi Green-veined White Lower Till SU075396 26/06/2017 Spot search In flight, damp neutral grassland

Pieris napi Green-veined White Lower Till SU077396 25/07/2017 Spot search Rough, marshy grassland

Aricia agestis Brown Argus Lower Till SU077396 25/07/2017 Spot search Rough, marshy grassland

Vanessa atalanta Red Admiral Lower Till SU077396 25/07/2017 Spot search Marshy grassland

Cynthia cardui Painted Lady Lower Till SU075396 26/06/2017 Spot search In flight, damp neutral grassland

Cynthia cardui Painted Lady Lower Till SU077396 25/07/2017 Spot search Rough, marshy grassland

Aglais urticae Small Tortoiseshell Lower Till SU076396 26/06/2017 Spot search On thistles, rank ruderal grassland

Scientific Name English Name Status Site Gridref Date Method Habitat Polygonia c-album Comma Lower Till SU077396 25/07/2017 Spot search Marshy grassland

Argynnis aglaja Dark Green Fritillary Lower Till SU077396 27/06/2017 Spot search In flight, Carex riparia fen meadow

Pyronia tithonus Gatekeeper Lower Till SU077396 25/07/2017 Spot search Rough, marshy grassland

Maniola jurtina Meadow Brown Lower Till SU075396 26/06/2017 Spot search In flight, damp neutral grassland

Maniola jurtina Meadow Brown Lower Till SU077396 25/07/2017 Spot search Rough, marshy grassland

Aphantopus hyperantus Ringlet Lower Till SU075396 26/06/2017 Spot search In flight, damp neutral grassland

Callimorpha dominula Scarlet Tiger Moth Lower Till SU075396 26/06/2017 Spot search In flight, wet woodland edge

Autographa gamma Silver Y Moth Lower Till SU075396 26/06/2017 Spot search In flight, damp neutral grassland

Chrysopilus cristatus A Snipe Fly Lower Till SU075396 26/06/2017 Sweeping Wet woodland edge

Chloromyia formosa A Soldierfly Lower Till SU076395 26/06/2017 Sweeping Carex riparia fen meadow

Chrysotoxum bicinctum A Hoverfly Lower Till SU076395 26/06/2017 Spot search Carex riparia fen meadow

Episyrphus balteatus A Hoverfly Lower Till SU076395 26/06/2017 Sweeping Carex riparia fen meadow

Syritta pipiens A Hoverfly Lower Till SU076395 26/06/2017 Sweeping Carex riparia fen meadow

Volucella bombylans A Hoverfly Lower Till SU077397 27/06/2017 Spot search On vegetation by river

Volucella pellucens A Hoverfly Lower Till SU075396 26/06/2017 Spot search On flowers, wet woodland edge

Xylota segnis A Hoverfly Lower Till SU075396 26/06/2017 Spot search On foliage, wet woodland

Sicus ferrugineus A Thick-headed Fly Lower Till SU076395 25/07/2017 Spot search Tall marginal vegetation on riverbank

Mesembrina meridiana A House Fly Lower Till SU075396 26/06/2017 Spot search Wet woodland edge

Eriothrix rufomaculata A Parasite Fly Lower Till SU077398 25/07/2017 Spot search Rank, ruderal grassland

Phasia hemiptera A Parasite Fly Lower Till SU076395 25/07/2017 Spot search Tall marginal vegetation on riverbank

Bombus lucorum White-tailed Bumblebee Lower Till SU076395 25/07/2017 Spot search Tall marginal vegetation on riverbank

Bombus terrestris Buff-tailed Bumblebee Lower Till SU075396 26/06/2017 Spot search On flowers, wet woodland edge

Bombus hortorum Garden Bumblebee Lower Till SU077396 25/07/2017 Spot search On flowers, marshy grassland

Bombus lapidarius Red-tailed Bumblebee Lower Till SU077396 25/07/2017 Spot search Rough, marshy grassland

Bombus hypnorum Tree Bumblebee Lower Till SU075396 26/06/2017 Spot search Flowers, wet woodland edge

Bombus pascuorum Common Carder Bumblebee Lower Till SU077396 25/07/2017 Spot search On flowers, marshy grassland

Araneus diadematus Garden Cross Spider Lower Till SU077396 25/07/2017 Spot search Rough, marshy grassland

Nuctenea umbratica An Orb-weaving Spider Lower Till SU075395 27/06/2017 Dead wood Beating dead willow branches, wet woodland

Nuctenea umbratica An Orb-weaving Spider Lower Till SU077399 25/07/2017 Beating On willow

Gammarus pulex Freshwater Shrimp Lower Till SU077396 27/06/2017 Sieving Springhead

Porcellio scaber Common Rough Woodlouse Lower Till SU077399 25/07/2017 Beating On willow

Scientific Name English Name Status Site Gridref Date Method Habitat Pupilla muscorum Moss Chrysalis Snail Parsonage Down NNR SU056409 28/06/2017 Ground search Grass tussocks, chalk downland

Pupilla muscorum Moss Chrysalis Snail Parsonage Down NNR SU061412 28/07/2017 Ground search Grass tussocks, chalk downland

Arion ater agg. Large Black Slug Parsonage Down NNR SU056409 28/06/2017 Spot search Chalk downland

Cernuella virgata Striped Snail Parsonage Down NNR SU056409 28/06/2017 Ground search Grass tussocks, chalk downland

Trichia hispida A Snail Parsonage Down NNR SU056409 28/06/2017 Ground search Grass tussocks, chalk downland

Trichia hispida A Snail Parsonage Down NNR SU061412 28/07/2017 Ground search Grass tussocks, chalk downland

Chorthippus parallelus Meadow Grasshopper Parsonage Down NNR SU056409 28/06/2017 Spot search Singing males, chalk downland

Chorthippus parallelus Meadow Grasshopper Parsonage Down NNR SU061412 28/07/2017 Spot search Chalk downland Common Green Omocestus viridulus Grasshopper Parsonage Down NNR SU061412 28/07/2017 Spot search Chalk downland

Metrioptera roeselii Roesel's Bush-cricket Parsonage Down NNR SU056409 28/06/2017 Spot search Singing males, chalk downland

Forficula auricularia Common Earwig Parsonage Down NNR SU056409 28/06/2017 Ground search Grass tussocks, chalk downland

Podops inuncta Turtle Bug Parsonage Down NNR SU056409 28/06/2017 Ground search Grass tussocks, chalk downland

Peritrechus geniculatus A Ground Bug Parsonage Down NNR SU061412 28/07/2017 Ground search Grass tussocks, chalk downland

Pithanus maerkelii A Plant Bug Parsonage Down NNR SU061412 28/07/2017 Ground search Grass tussocks, chalk downland

Trechus quadristriatus A Ground Beetle Parsonage Down NNR SU063411 28/07/2017 Ground search Grass tussocks, chalk downland

Pterostichus madidus A Ground Beetle Parsonage Down NNR SU056409 28/06/2017 Ground search Grass tussocks, chalk downland

Pterostichus madidus A Ground Beetle Parsonage Down NNR SU061412 28/07/2017 Ground search Grass tussocks, chalk downland

Calathus fuscipes A Ground Beetle Parsonage Down NNR SU056409 28/06/2017 Ground search Grass tussocks, chalk downland

Calathus fuscipes A Ground Beetle Parsonage Down NNR SU061412 28/07/2017 Ground search Grass tussocks, chalk downland

Amara curta A Ground Beetle NS. Parsonage Down NNR SU056411 28/06/2017 Ground search Grass tussocks, chalk downland Amara familiaris A Ground Beetle Parsonage Down NNR SU061412 28/07/2017 Ground search Grass tussocks, chalk downland

Licinus depressus A Ground Beetle NS. Parsonage Down NNR SU061412 28/07/2017 Ground search Grass tussocks, chalk downland Badister bullatus A Ground Beetle Parsonage Down NNR SU061412 28/07/2017 Ground search Grass tussocks, chalk downland

Demetrias atricapillus A Ground Beetle Parsonage Down NNR SU063411 28/07/2017 Ground search Grass tussocks, chalk downland

Paradromius linearis A Ground Beetle Parsonage Down NNR SU061412 28/07/2017 Ground search Grass tussocks, chalk downland

Megasternum concinnum A Hydrophilid Beetle Parsonage Down NNR SU061412 28/07/2017 Ground search Grass tussocks, chalk downland

Silpha atrata A Carrion Beetle Parsonage Down NNR SU056409 28/06/2017 Ground search Grass tussocks, chalk downland

Silpha atrata A Carrion Beetle Parsonage Down NNR SU061412 28/07/2017 Ground search Grass tussocks, chalk downland

Metopsia clypeata A Rove Beetle Parsonage Down NNR SU063411 28/07/2017 Ground search Grass tussocks, chalk downland

Lamprinodes saginatus A Rove Beetle Na. Parsonage Down NNR SU063411 28/07/2017 Ground search Grass tussocks, chalk downland Sepedophilus nigripennis A Rove Beetle Parsonage Down NNR SU061412 28/07/2017 Ground search Grass tussocks, chalk downland

Scientific Name English Name Status Site Gridref Date Method Habitat Tachyporus atriceps A Rove Beetle Parsonage Down NNR SU056409 28/06/2017 Ground search Grass tussocks, chalk downland

Tachyporus dispar A Rove Beetle Parsonage Down NNR SU061412 28/07/2017 Ground search Grass tussocks, chalk downland

Tachyporus hypnorum A Rove Beetle Parsonage Down NNR SU056409 28/06/2017 Ground search Grass tussocks, chalk downland

Tachyporus hypnorum A Rove Beetle Parsonage Down NNR SU061412 28/07/2017 Ground search Grass tussocks, chalk downland

Tachyprous nitidulus A Rove Beetle Parsonage Down NNR SU061412 28/07/2017 Ground search Grass tussocks, chalk downland

Tachyporus tersus A Rove Beetle Parsonage Down NNR SU061412 28/07/2017 Ground search Grass tussocks, chalk downland Litter at mouth of rabbit burrows, chalk Aleochara cuniculorum A Rove Beetle Parsonage Down NNR SU059409 28/06/2017 Ground search grassland Litter at mouth of rabbit burrows, chalk Drusilla canaliculata A Rove Beetle Parsonage Down NNR SU059409 28/06/2017 Ground search grassland

Drusilla canaliculata A Rove Beetle Parsonage Down NNR SU061412 28/07/2017 Ground search Grass tussocks, chalk downland

Stenus aceris A Rove Beetle Parsonage Down NNR SU061412 28/07/2017 Ground search Grass tussocks, chalk downland

Stenus ossium A Rove Beetle Parsonage Down NNR SU061412 28/07/2017 Ground search Grass tussocks, chalk downland

Stenus flavipes A Rove Beetle Parsonage Down NNR SU061412 28/07/2017 Ground search Grass tussocks, chalk downland

Paederus littoralis A Rove Beetle Parsonage Down NNR SU061412 28/07/2017 Ground search Grass tussocks, chalk downland

Rugilus similis A Rove Beetle N. Parsonage Down NNR SU061412 28/07/2017 Ground search Grass tussocks, chalk downland Philonthus carbonarius A Rove Beetle Parsonage Down NNR SU061412 28/07/2017 Ground search Grass tussocks, chalk downland

Philonthus cruentatus A Rove Beetle Parsonage Down NNR SU056409 28/06/2017 Dung Cow dund, chalk downland

Philonthus tenuicornis A Rove Beetle Parsonage Down NNR SU056409 28/06/2017 Dung Cow dund, chalk downland

Philonthus varians A Rove Beetle Parsonage Down NNR SU056409 28/06/2017 Dung Cow dung, chalk downland

Quedius levicollis A Rove Beetle Parsonage Down NNR SU056409 28/06/2017 Ground search Grass tussocks, chalk downland

Quedius semiobscurus A Rove Beetle Parsonage Down NNR SU056409 28/06/2017 Ground search Grass tussocks, chalk downland

Quedius semiobscurus A Rove Beetle Parsonage Down NNR SU061412 28/07/2017 Ground search Grass tussocks, chalk downland

Tasgius globulifer A Rove Beetle Parsonage Down NNR SU056409 28/06/2017 Ground search Grass tussocks, chalk downland

Tasgius globulifer A Rove Beetle Parsonage Down NNR SU061412 28/07/2017 Ground search Grass tussocks, chalk downland

Tasgius morsitans A Rove Beetle Parsonage Down NNR SU063411 28/07/2017 Ground search Grass tussocks, chalk downland

Gyrohypnus fracticornis A Rove Beetle Parsonage Down NNR SU056409 28/06/2017 Dung Cow dung, chalk downland

Megalinus glabratus A Rove Beetle Parsonage Down NNR SU056409 28/06/2017 Dung Cow dung, chalk downland

Xantholinus linearis A Rove Beetle Parsonage Down NNR SU061412 28/07/2017 Ground search Grass tussocks, chalk downland

Trachys troglodytes A Jewel Beetle Parsonage Down NNR SU059409 28/06/2017 Spot search Larval mines and egg spots on Succisa leaves

Agrypnus murinus A Click Beetle Parsonage Down NNR SU059409 28/06/2017 Ground search Grass tussocks, chalk downland

Agrypnus murinus A Click Beetle Parsonage Down NNR SU063411 28/07/2017 Ground search Grass tussocks, chalk downland

Scientific Name English Name Status Site Gridref Date Method Habitat Aplotarsus incanus A Click Beetle Parsonage Down NNR SU059409 28/06/2017 Ground search Grass tussocks, chalk downland

Agriotes sputator A Click Beetle Parsonage Down NNR SU061412 28/07/2017 Ground search Grass tussocks, chalk downland

Meligethes solidus A Pollen Beetle N. Parsonage Down NNR SU063411 28/07/2017 Tapping Helianthemum nummularium, chalk downland Phalacrus championi A Phalacrid Beetle Na. Parsonage Down NNR SU061412 28/07/2017 Ground search Grass tussocks, chalk downland Oulema melanopus agg. A Leaf Beetle Parsonage Down NNR SU063411 28/07/2017 Ground search Grass tussocks, chalk downland

Chrysolina banksii A Leaf Beetle Parsonage Down NNR SU056409 28/06/2017 Ground search Grass tussocks, chalk downland

Chrysolina banksii A Leaf Beetle Parsonage Down NNR SU061412 28/07/2017 Ground search Grass tussocks, chalk downland

Sermylassa halensis A Leaf Beetle Parsonage Down NNR SU061412 28/07/2017 Ground search Grass tussocks, chalk downland

Neocrepidodera ferruginea A Flea Beetle Parsonage Down NNR SU056409 28/06/2017 Ground search Grass tussocks, chalk downland

Neocrepidodera ferruginea A Flea Beetle Parsonage Down NNR SU061412 28/07/2017 Tapping Lotus corniculatus, chalk downland

Sphaeroderma rubidum A Flea Beetle Parsonage Down NNR SU063411 28/07/2017 Ground search Grass tussocks, chalk downland

Sphaeroderma testaceum A Flea Beetle Parsonage Down NNR SU056409 28/06/2017 Ground search Cirsium vulgare, chalk downland

Psylliodes chalcomera A Flea Beetle NS. Parsonage Down NNR SU056409 28/06/2017 Tapping Carduus nutans, chalk downland Psylliodes chrysocephala A Flea Beetle Parsonage Down NNR SU061412 28/07/2017 Tapping Lotus corniculatus, chalk downland

Ceratapion carduorum A Weevil Parsonage Down NNR SU056409 28/06/2017 Ground search Cirsium vulgare, chalk downland

Holotrichapion ononis A Weevil Parsonage Down NNR SU056409 28/06/2017 Ground search Ononis spinosa, chalk downland

Ischnopterapion loti A Weevil Parsonage Down NNR SU059409 28/06/2017 Ground search Grass tussocks, chalk downland

Ischnopterapion loti A Weevil Parsonage Down NNR SU061412 28/07/2017 Tapping Lotus corniculatus, chalk downland

Ischnopterapion virens A Weevil Parsonage Down NNR SU061412 28/07/2017 Ground search Grass tussocks, chalk downland

Protapion apricans A Weevil Parsonage Down NNR SU061412 28/07/2017 Tapping Trifolium pratense, chalk downland

Mecinus pascuorum A Weevil Parsonage Down NNR SU063411 28/07/2017 Ground search Grass tussocks, chalk downland

Stenocarus ruficornis A Weevil Nb. Parsonage Down NNR SU061412 28/07/2017 Ground search Grass tussocks, chalk downland Otiorhynchus clavipes A Weevil Parsonage Down NNR SU063411 28/07/2017 Ground search Grass tussocks, chalk downland

Otiorhynchus ligneus A Weevil Parsonage Down NNR SU063411 28/07/2017 Ground search Grass tussocks, chalk downland

Sitona hispidulus A Weevil Parsonage Down NNR SU061412 28/07/2017 Ground search Grass tussocks, chalk downland

Sitona hispidulus A Weevil Parsonage Down NNR SU063411 28/07/2017 Ground search Grass tussocks, chalk downland

Sitona lineatus A Weevil Parsonage Down NNR SU059409 28/06/2017 Ground search Grass tussocks, chalk downland

Sitona lineatus A Weevil Parsonage Down NNR SU063411 28/07/2017 Ground search Grass tussocks, chalk downland

Sitona sulcifrons A Weevil Parsonage Down NNR SU056411 28/06/2017 Ground search Grass tussocks, chalk downland

Zygaena filipendulae Six-spot Burnet Parsonage Down NNR SU063411 29/07/2017 Spot search Scabiosa columbaria, chalk downland

Myelois circumvoluta Thistle Ermine Parsonage Down NNR SU056409 28/06/2017 Ground search Cirsium vulgare, chalk downland

Scientific Name English Name Status Site Gridref Date Method Habitat Thymelicus sylvestris Small Skipper Parsonage Down NNR SU056411 28/06/2017 Spot search Chalk downland

Thymelicus sylvestris Small Skipper Parsonage Down NNR SU063411 29/07/2017 Spot search Chalk downland

Aricia agestis Brown Argus Parsonage Down NNR SU063411 29/07/2017 Spot search Chalk downland

Cynthia cardui Painted Lady Parsonage Down NNR SU063411 29/07/2017 Spot search Chalk downland

Argynnis aglaja Dark Green Fritillary Parsonage Down NNR SU056411 28/06/2017 Spot search Chalk downland

Melanargia galathea Marbled White Parsonage Down NNR SU056409 28/06/2017 Spot search Chalk downland

Melanargia galathea Marbled White Parsonage Down NNR SU061412 28/07/2017 Spot search Chalk downland

Maniola jurtina Meadow Brown Parsonage Down NNR SU056409 28/06/2017 Spot search Chalk downland

Maniola jurtina Meadow Brown Parsonage Down NNR SU061412 28/07/2017 Spot search Chalk downland

Aphantopus hyperantus Ringlet Parsonage Down NNR SU056411 28/06/2017 Spot search Chalk downland

Coenonympha pamphilus Small Heath Parsonage Down NNR SU056409 28/06/2017 Spot search Chalk downland

Autographa gamma Silver Y Moth Parsonage Down NNR SU061412 28/07/2017 Spot search Chalk downland

Nephrotoma guestfalica A Cranefly Parsonage Down NNR SU056409 28/06/2017 Sweeping Chalk downland

Chloromyia formosa A Soldierfly Parsonage Down NNR SU056409 28/06/2017 Sweeping Chalk downland

Leptogaster cylindrica A Robberfly Parsonage Down NNR SU056409 28/06/2017 Sweeping Chalk downland

Episyrphus balteatus A Hoverfly Parsonage Down NNR SU063411 29/07/2017 Spot search Scabiosa columbaria, chalk downland

Eristalis pertinax A Hoverfly Parsonage Down NNR SU063411 29/07/2017 Spot search Scabiosa columbaria, chalk downland

Scathophaga stercoraria A Dung Fly Parsonage Down NNR SU056409 28/06/2017 Dung Cow dung, chalk downland

Mesembrina meridiana A House Fly Parsonage Down NNR SU056411 28/06/2017 Dung Cow dung, chalk downland

Prosena siberita A Parasite Fly Parsonage Down NNR SU063411 29/07/2017 Spot search Scabiosa columbaria, chalk downland

Eriothrix rufomaculata A Parasite Fly Parsonage Down NNR SU061412 28/07/2017 Spot search Chalk downland

Nowickia ferox A Parasite Fly Parsonage Down NNR SU063411 28/07/2017 Spot search Scabiosa columbaria, chalk downland

Andrena marginata Small Scabious Mining Bee Na. Parsonage Down NNR SU063411 29/07/2017 Spot search Scabiosa columbaria, chalk downland Bombus hortorum Garden Bumblebee Parsonage Down NNR SU061412 28/07/2017 Spot search Chalk downland

Bombus lapidarius Red-tailed Bumblebee Parsonage Down NNR SU056409 28/06/2017 Spot search On flowers, chalk downland

Bombus lapidarius Red-tailed Bumblebee Parsonage Down NNR SU061412 28/07/2017 Spot search Chalk downland

Nomada argentata Silver-sided Nomad Bee RDB3. Parsonage Down NNR SU063411 29/07/2017 Spot search Scabiosa columbaria, chalk downland Lasioglossum albipes Bloomed Furrow Bee Parsonage Down NNR SU063411 29/07/2017 Spot search Scabiosa columbaria, chalk downland

Melitta haemorrhoidalis Gold-tailed Melitta Bee Parsonage Down NNR SU063411 29/07/2017 Spot search Campanula rotundifolia, chalk downland

Lasius flavus Yellow Meadow Ant Parsonage Down NNR SU061412 28/07/2017 Ground search Grass tussocks, chalk downland

Lasius niger Black Garden Ant Parsonage Down NNR SU056409 28/06/2017 Ground search Grass tussocks, chalk downland

Scientific Name English Name Status Site Gridref Date Method Habitat Lasius niger Black Garden Ant Parsonage Down NNR SU061412 28/07/2017 Ground search Grass tussocks, chalk downland

Myrmica scabrinodis A Red Ant Parsonage Down NNR SU061412 28/07/2017 Ground search Grass tussocks, chalk downland

Nemastoma bimaculatum A Harvestman Spider Parsonage Down NNR SU063411 28/07/2017 Ground search Grass tussocks, chalk downland

Araneus diadematus Garden Cross Spider Parsonage Down NNR SU061412 28/07/2017 Ground search Grass tussocks, chalk downland

Argiope bruennichi Zebra Spider Parsonage Down NNR SU061412 28/07/2017 Spot search Chalk downland

Agelena labyrinthica An Agelenid Spider Parsonage Down NNR SU056409 28/06/2017 Spot search Webs, chalk downland

Philoscia muscorum Striped Woodlouse Parsonage Down NNR SU056409 28/06/2017 Ground search Grass tussocks, chalk downland

Philoscia muscorum Striped Woodlouse Parsonage Down NNR SU061412 28/07/2017 Ground search Grass tussocks, chalk downland Platyarthrus hoffmannseggii Ant Woodlouse Parsonage Down NNR SU061412 28/07/2017 Ground search Grass tussocks, chalk downland Litter at mouth of rabbit burrows, chalk Oniscus asellus Common Shiny Woodlouse Parsonage Down NNR SU059409 28/06/2017 Ground search grassland

Armadillidium vulgare Common Pill Woodlouse Parsonage Down NNR SU056409 28/06/2017 Ground search Grass tussocks, chalk downland

Monacha cantiana Kentish Snail Diamond Wood SU105406 29/06/2017 Spot search Remnant chalk grassland, wood edge On vegetation, remnant chalk grassland, wood Monacha cantiana Kentish Snail Diamond Wood SU105406 27/07/2017 Spot search edge Remnant chalk grassland on earthwork at Cepaea nemoralis Brown-lipped Snail Diamond Wood SU103408 27/07/2017 Sweeping wood edge Common Green Omocestus viridulus Grasshopper Diamond Wood SU105406 27/07/2017 Spot search Singing, remnant chalk grassland, wood edge Singing male, remnant chalk grassland, wood Chorthippus parallelus Meadow Grasshopper Diamond Wood SU105406 29/06/2017 Spot search edge

Chorthippus parallelus Meadow Grasshopper Diamond Wood SU105406 27/07/2017 Spot search Singing, remnant chalk grassland, wood edge

Metrioptera roeselii Roesel's Bush-cricket Diamond Wood SU105406 29/06/2017 Spot search Singing, remnant chalk grassland, wood edge

Metrioptera roeselii Roesel's Bush-cricket Diamond Wood SU105406 27/07/2017 Spot search Singing, remnant chalk grassland, wood edge

Conocephalus discolor Long-winged Conehead Diamond Wood SU103408 27/07/2017 Sweeping Remnant chalk grassland, wood edge

Forficula auricularia Common Earwig Diamond Wood SU105406 29/06/2017 Sweeping Remnant chalk grassland, wood edge

Forficula auricularia Common Earwig Diamond Wood SU106407 27/07/2017 Beating Crataegus monogyna, wood edge Centaurea nigra, remnant chalk grassland, Dolycoris baccarum Sloe Bug Diamond Wood SU105406 27/07/2017 Tapping wood edge

Pentatoma rufipes Forest Bug Diamond Wood SU105408 29/06/2017 Beating Ligustrum vulgare, woodland

Eurydema oleracea Brassica Shieldbug Diamond Wood SU103408 29/06/2017 Sweeping Crucifers, arable margin Grass tussocks, remnant chalk grassland, Rhopalus subrufus A Rhopalid Bug Diamond Wood SU105406 27/07/2017 Ground search wood edge

Aneurus avenius A Flatbug Diamond Wood SU107407 29/06/2017 Spot search Dead branches, wood edge

Bembidion lampros A Ground Beetle Diamond Wood SU105406 29/06/2017 Ground search Under stone, arable margin

Scientific Name English Name Status Site Gridref Date Method Habitat Pterostichus madidus A Ground Beetle Diamond Wood SU105406 29/06/2017 Ground search Under stone, arable margin Grass tussocks, remnant chalk grassland, Calathus melanocephalus A Ground Beetle Diamond Wood SU105406 27/07/2017 Ground search wood edge

Amara convexior A Ground Beetle Diamond Wood SU107407 29/06/2017 Ground search Under stone, arable margin

Ophonus rufibarbis A Ground Beetle Diamond Wood SU105406 29/06/2017 Ground search Under stone, arable margin

Badister bullatus A Ground Beetle Diamond Wood SU107407 29/06/2017 Ground search Under stone, arable margin Grass tussocks, remnant chalk grassland, Bolitobius cingulatus A Rove Beetle Diamond Wood SU105406 27/07/2017 Ground search wood edge Grass tussocks, remnant chalk grassland, Sepedophilus nigripennis A Rove Beetle Diamond Wood SU105406 27/07/2017 Ground search wood edge Grass tussocks, remnant chalk grassland, Tachyporus hypnorum A Rove Beetle Diamond Wood SU105406 27/07/2017 Ground search wood edge Grass tussocks, remnant chalk grassland, Tachyporus solutus A Rove Beetle Diamond Wood SU105406 27/07/2017 Ground search wood edge Achillea millefolium, remnant chalk grassland Tachyporus solutus A Rove Beetle Diamond Wood SU103408 27/07/2017 Tapping on earthwork at wood edge

Atheta crassicornis A Rove Beetle Diamond Wood SU105408 29/06/2017 Carrion Dead squirrel, woodland

Drusilla canaliculata A Rove Beetle Diamond Wood SU105406 29/06/2017 Ground search Under stone, arable margin

Stenus flavipes A Rove Beetle Diamond Wood SU107407 27/07/2017 Tapping Centaurea nigra, wood edge Grass tussocks, remnant chalk grassland, Stenus ochropus A Rove Beetle Diamond Wood SU105406 27/07/2017 Ground search wood edge

Xantholinus linearis A Rove Beetle Diamond Wood SU107407 29/06/2017 Ground search Under stone, arable margin On flowers, remnant chalk grassland, wood Rhagonycha fulva A Soldier Beetle Diamond Wood SU105406 29/06/2017 Spot search edge On Pastinaca sativa, remnant chalk grassland, Rhagonycha fulva A Soldier Beetle Diamond Wood SU105406 27/07/2017 Spot search wood edge

Malthinus balteatus A Soldier Beetle Diamond Wood SU107407 29/06/2017 Beating Ligustrum vulgare, wood edge

Malthinus seriepunctatus A Soldier Beetle Diamond Wood SU107407 29/06/2017 Beating Ligustrum vulgare, wood edge Remnant chalk grassland on earthwork at Hemicrepidius hirtus A Click Beetle Diamond Wood SU103408 27/07/2017 Sweeping wood edge

Hemicoelus fulvicorne A Woodworm Beetle Diamond Wood SU107407 29/06/2017 Beating Dead branches, wood edge Linaria vulgaris, remnant chalk grassland on Brachypterolus vestitus A Pollen Beetle Diamond Wood SU103408 27/07/2017 Tapping earthwork at wood edge Clinopodium vulgare, remnant chalk grassland, Brachypterus glaber A Pollen Beetle Diamond Wood SU105406 27/07/2017 Tapping wood edge

Meligethes bidens A Pollen Beetle Diamond Wood SU107407 29/06/2017 Tapping Clinopodium vulgare, arable margin Clinopodium vulgare, remnant chalk grassland, Meligethes bidens A Pollen Beetle Diamond Wood SU105406 27/07/2017 Tapping wood edge

Scientific Name English Name Status Site Gridref Date Method Habitat Helianthemum nummularium, remnant chalk Meligethes solidus A Pollen Beetle N. Diamond Wood SU103408 27/07/2017 Tapping grassland on earthwork at wood edge Stilbus testaceus A Phalacrid Beetle Diamond Wood SU107407 27/07/2017 Tapping Centaurea nigra, wood edge Grass tussocks, remnant chalk grassland, Rhyzobius litura A Ladybird Diamond Wood SU105406 27/07/2017 Ground search wood edge Propylea quatturodecimpunctata 14-spot Ladybird Diamond Wood SU103408 29/06/2017 Sweeping Crucifers, arable margin Propylea Centaurea nigra, remnant chalk grassland, quatturodecimpunctata 14-spot Ladybird Diamond Wood SU105406 27/07/2017 Tapping wood edge Tytthaspis sedecimpunctata 15-spot Ladybird Diamond Wood SU107407 27/07/2017 Tapping Centaurea nigra, wood edge Coccinella septempunctata 7-spot Ladybird Diamond Wood SU103408 29/06/2017 Sweeping Crucifers, arable margin Coccinella On vegetation, remnant chalk grassland, wood septempunctata 7-spot Ladybird Diamond Wood SU105406 27/07/2017 Spot search edge On flowers, remnant chalk grassland, wood Harmonia axyridis Harlequin Ladybird Diamond Wood SU105406 27/07/2017 Spot search edge Subcoccinella Centaurea nigra, remnant chalk grassland, vigintiquattuorpunctata 24-spot Ladybird Diamond Wood SU105406 27/07/2017 Tapping wood edge

Oedemera lurida An Oedemerid Beetle Diamond Wood SU105406 29/06/2017 Sweeping Remnant chalk grassland, wood edge On flowers and in flight, remnant chalk Oedemera lurida An Oedemerid Beetle Diamond Wood SU105406 27/07/2017 Spot search grassland, wood edge

Salpingus planirostris A Salpingid Beetle Diamond Wood SU103408 29/06/2017 Beating Dead elder, scrub woodland Grass tussocks, remnant chalk grassland, Omonadus formicarius An Anthicid Beetle Diamond Wood SU105406 27/07/2017 Ground search wood edge

Paracorymbia fulva A Longhorn Beetle RDB3. Diamond Wood SU107407 29/06/2017 Spot search Heracleum sphondylium, wood edge On Torilis japonica, remnant chalk grassland, Paracorymbia fulva A Longhorn Beetle RDB3. Diamond Wood SU103408 27/07/2017 Spot search wood edge Grass tussocks, remnant chalk grassland, Oulema melanopus agg. A Leaf Beetle Diamond Wood SU105406 27/07/2017 Ground search wood edge Remnant chalk grassland on earthwork at Psylliodes chrysocephala A Leaf Beetle Diamond Wood SU103408 27/07/2017 Sweeping wood edge

Sphaeroderma rubidum A Flea Beetle Diamond Wood SU107407 27/07/2017 Tapping Centaurea nigra, wood edge

Sphaeroderma testaceum A Flea Beetle Diamond Wood SU107407 27/07/2017 Tapping Centaurea nigra, wood edge Centaurea nigra, remnant chalk grassland, Cassida vibex A Tortoise Beetle Diamond Wood SU105406 27/07/2017 Tapping wood edge Lathyrus pratensis and Vicia cracca, remnant Eutrichapion viciae A Weevil Diamond Wood SU103408 27/07/2017 Tapping chalk grassland on earthwork at wood edge Centaurea nigra, remnant chalk grassland, Gymnetron melanarium A Weevil Nb. Diamond Wood SU105406 27/07/2017 Tapping wood edge Linaria vulgaris, remnant chalk grassland on Rhinusa antirrhini A Weevil Diamond Wood SU103408 27/07/2017 Tapping earthwork at wood edge

Scientific Name English Name Status Site Gridref Date Method Habitat Rhamphus oxyacanthae A Weevil Diamond Wood SU106407 27/07/2017 Beating Crataegus monogyna, wood edge

Tychius picirostris A Weevil Diamond Wood SU105406 29/06/2017 Tapping Trifolium and Medicago, arable margin Ceutorhynchus pallidactylus A Weevil Diamond Wood SU103408 29/06/2017 Sweeping Crucifers, arable margin

Sirocalodes depressicollis A Weevil Diamond Wood SU103408 29/06/2017 Sweeping Fumaria, arable margin

Nedyus quadrimaculatus A Weevil Diamond Wood SU103408 29/06/2017 Sweeping Remnant chalk grassland, wood edge Lathyrus pratensis and Vicia cracca, remnant Sitona lineatus A Weevil Diamond Wood SU103408 27/07/2017 Tapping chalk grassland on earthwork at wood edge

Sitona suturalis A Weevil Diamond Wood SU103408 29/06/2017 Sweeping Arable margin

Ischnopterapion virens A Weevil Diamond Wood SU105406 29/06/2017 Tapping Trifolium and Medicago, arable margin

Stenopterapion tenue A Weevil Diamond Wood SU105406 29/06/2017 Tapping Trifolium and Medicago, arable margin

Protapion fulvipes A Weevil Diamond Wood SU105406 29/06/2017 Tapping Trifolium and Medicago, arable margin

Hypera postica A Weevil Diamond Wood SU105406 29/06/2017 Tapping Trifolium and Medicago, arable margin

Sitona lineatus A Weevil Diamond Wood SU105406 29/06/2017 Tapping Trifolium and Medicago, arable margin On flowers, remnant chalk grassland, wood Thymelicus lineola Essex Skipper Diamond Wood SU105406 29/06/2017 Spot search edge

Thymelicus sylvestris Small Skipper Diamond Wood SU105406 29/06/2017 Spot search In flight, remnant chalk grassland, wood edge On flowers and in flight, remnant chalk Thymelicus sylvestris Small Skipper Diamond Wood SU105406 27/07/2017 Spot search grassland, wood edge On flowers and in flight, remnant chalk Gonepteryx rhamni Brimstone Diamond Wood SU105406 27/07/2017 Spot search grassland, wood edge

Pieris brassicae Large White Diamond Wood SU105406 29/06/2017 Spot search In flight, remnant chalk grassland, wood edge On flowers and in flight, remnant chalk Pieris brassicae Large White Diamond Wood SU105406 27/07/2017 Spot search grassland, wood edge

Pieris rapae Small White Diamond Wood SU105406 29/06/2017 Spot search In flight, remnant chalk grassland, wood edge On flowers and in flight, remnant chalk Pieris rapae Small White Diamond Wood SU105406 27/07/2017 Spot search grassland, wood edge On flowers and in flight, remnant chalk Aricia agestis Brown Argus Diamond Wood SU105406 27/07/2017 Spot search grassland, wood edge On flowers and in flight, remnant chalk Polyommatus icarus Common Blue Diamond Wood SU105406 27/07/2017 Spot search grassland, wood edge

Vanessa atalanta Red Admiral Diamond Wood SU105406 29/06/2017 Spot search In flight, remnant chalk grassland, wood edge On flowers and in flight, remnant chalk Vanessa atalanta Red Admiral Diamond Wood SU105406 27/07/2017 Spot search grassland, wood edge On flowers and in flight, remnant chalk Aglais urticae Small Tortoiseshell Diamond Wood SU105406 27/07/2017 Spot search grassland, wood edge

Melanargia galathea Marbled White Diamond Wood SU105406 29/06/2017 Spot search In flight, remnant chalk grassland, wood edge

Pyronia tithonus Gatekeeper Diamond Wood SU105406 27/07/2017 Spot search On flowers and in flight, chalk grassland

Scientific Name English Name Status Site Gridref Date Method Habitat Maniola jurtina Meadow Brown Diamond Wood SU105406 29/06/2017 Spot search In flight, remnant chalk grassland, wood edge On flowers and in flight, remnant chalk Maniola jurtina Meadow Brown Diamond Wood SU105406 27/07/2017 Spot search grassland, wood edge

Aphantopus hyperantus Ringlet Diamond Wood SU105406 29/06/2017 Spot search In flight, remnant chalk grassland, wood edge On flowers, remnant chalk grassland, wood Eremobia ochroleuca Dusky Sallow Moth Diamond Wood SU105406 27/07/2017 Spot search edge

Autographa gamma Silver Y Moth Diamond Wood SU105406 27/07/2017 Spot search In flight, remnant chalk grassland, wood edge On Torilis japonica, remnant chalk grassland, Nephrotoma cornicina A Cranefly Diamond Wood SU103408 27/07/2017 Spot search wood edge

Chloromyia formosa A Soldierfly Diamond Wood SU105406 29/06/2017 Spot search In flight, remnant chalk grassland, wood edge Remnant chalk grassland on earthwork at Chloromyia formosa A Soldierfly Diamond Wood SU103408 27/07/2017 Sweeping wood edge On Pastinaca sativa, remnant chalk grassland, Thereva nobilitata Common Stiletto Fly Diamond Wood SU103408 27/07/2017 Spot search wood edge On Heracleum sphondylium, remnant chalk Cheilosia illustrata A Hoverfly Diamond Wood SU105406 29/06/2017 Spot search grassland, wood edge

Chryotoxum bicinctum A Hoverfly Diamond Wood SU105406 29/06/2017 Spot search In flight, remnant chalk grassland, wood edge On flowers, remnant chalk grassland, wood Episyrphus balteatus A Hoverfly Diamond Wood SU105406 29/06/2017 Spot search edge Remnant chalk grassland on earthwork at Episyrphus balteatus A Hoverfly Diamond Wood SU103408 27/07/2017 Sweeping wood edge Remnant chalk grassland on earthwork at Eristalis tenax A Hoverfly Diamond Wood SU103408 27/07/2017 Sweeping wood edge Remnant chalk grassland on earthwork at Myathropa florea A Hoverfly Diamond Wood SU103408 27/07/2017 Sweeping wood edge Remnant chalk grassland on earthwork at Scaeva pyrastri A Hoverfly Diamond Wood SU103408 27/07/2017 Sweeping wood edge Pastinaca sativa, remnant chalk grassland on Sphaerophoria interrupta A Hoverfly Diamond Wood SU103408 27/07/2017 Sweeping earthwork at wood edge Remnant chalk grassland on earthwork at Syritta pipiens A Hoverfly Diamond Wood SU103408 27/07/2017 Sweeping wood edge Remnant chalk grassland on earthwork at Scathophaga stercoraria A Dung Fly Diamond Wood SU103408 27/07/2017 Sweeping wood edge

Mesembrina meridiana A House Fly Diamond Wood SU105406 29/06/2017 Spot search Resting on fencepost, wood edge On flowers, remnant chalk grassland, wood Eriothrix rufomaculata A Parasite Fly Diamond Wood SU105406 27/07/2017 Spot search edge On Torilis japonica, remnant chalk grassland, Andrena minutuloides Plain Mini-miner Bee Na. Diamond Wood SU103408 27/07/2017 Spot search wood edge On flowers and in flight, remnant chalk Bombus terrestris Buff-tailed Bumblebee Diamond Wood SU105406 27/07/2017 Spot search grassland, wood edge On flowers, remnant chalk grassland, wood Bombus lapidarius Red-tailed Bumblebee Diamond Wood SU105406 29/06/2017 Spot search edge

Scientific Name English Name Status Site Gridref Date Method Habitat On flowers and in flight, remnant chalk Bombus lapidarius Red-tailed Bumblebee Diamond Wood SU105406 27/07/2017 Spot search grassland, wood edge On flowers, remnant chalk grassland, wood Bombus pratorum Early Bumblebee Diamond Wood SU105406 29/06/2017 Spot search edge On flowers, remnant chalk grassland, wood Bombus pascuorum Common Carder Bumblebee Diamond Wood SU105406 29/06/2017 Spot search edge On flowers and in flight, remnant chalk Bombus pascuorum Common Carder Bumblebee Diamond Wood SU105406 27/07/2017 Spot search grassland, wood edge On Torilis japonica, remnant chalk grassland, Nomada fabriciana Fabricius' Nomad Bee Diamond Wood SU103408 27/07/2017 Spot search wood edge

Sphecodes ephippius Bare-saddled Blood Bee Diamond Wood SU103408 27/07/2017 Sweeping Remnant chalk grassland, wood edge On Torilis japonica, remnant chalk grassland, Sphecodes geoffrellus Geoffroy's Blood Bee Diamond Wood SU103408 27/07/2017 Spot search wood edge Pastinaca sativa, remnant chalk grassland on Tiphia femorata A Beetle-killing Wasp Diamond Wood SU103408 27/07/2017 Spot search earthwork at wood edge

Lasius flavus Yellow Meadow Ant Diamond Wood SU105406 29/06/2017 Spot search Anthills, remnant chalk grassland, wood edge Grass tussocks, remnant chalk grassland, Lasius niger Black Garden Ant Diamond Wood SU105406 29/06/2017 Ground search wood edge

Philoscia muscorum Striped Woodlouse Diamond Wood SU107407 29/06/2017 Ground search Under stone, arable margin Grass tussocks, remnant chalk grassland, Philoscia muscorum Striped Woodlouse Diamond Wood SU105406 27/07/2017 Ground search wood edge

Porcellio scaber Common Rough Woodlouse Diamond Wood SU107407 29/06/2017 Beating Dead branches, wood edge

Armadillidium vulgare Common Pill Woodlouse Diamond Wood SU105406 29/06/2017 Ground search Under stone, arable margin Grass tussocks, remnant chalk grassland, Armadillidium vulgare Common Pill Woodlouse Diamond Wood SU105406 27/07/2017 Ground search wood edge

Monacha cantiana Kentish Snail Arable 1 SU112414 27/07/2017 Sweeping Arable margin

Cepaea nemoralis Brown-lipped Snail Arable 1 SU113416 29/06/2017 Sweeping Rank arable margin

Cepaea nemoralis Brown-lipped Snail Arable 1 SU112414 27/07/2017 Spot search Arable margin

Helix aspersa Garden Snail Arable 1 SU112414 27/07/2017 Sweeping Arable margin

Metrioptera roeselii Roesel's Bush-cricket Arable 1 SU112414 29/06/2017 Spot search Singing, rank arable margin

Metrioptera roeselii Roesel's Bush-cricket Arable 1 SU112414 27/07/2017 Sweeping Arable margin Common Green Omocestus viridulus Grasshopper Arable 1 SU114413 27/07/2017 Pitfall trap Arable margin

Forficula auricularia Common Earwig Arable 1 SU112414 27/07/2017 Sweeping Arable margin

Sehirus luctuosus Forget-me-not Shieldbug Arable 1 SU114413 27/07/2017 Pitfall trap Arable margin

Dolycoris baccarum Sloe Bug Arable 1 SU112414 27/07/2017 Sweeping Arable margin

Rhopalus subrufus A Rhopalid Bug Arable 1 SU112414 27/07/2017 Tapping Carduus nutans, arable margin

Carabus violaceus A Ground Beetle Arable 1 SU114413 27/07/2017 Pitfall trap Arable margin

Scientific Name English Name Status Site Gridref Date Method Habitat Nebria brevicollis A Ground Beetle Arable 1 SU114413 27/07/2017 Pitfall trap Arable margin

Nebria salina A Ground Beetle Arable 1 SU114413 27/07/2017 Pitfall trap Arable margin

Notiophilus biguttatus A Ground Beetle Arable 1 SU114413 27/07/2017 Pitfall trap Arable margin

Notiophilus palustris A Ground Beetle Arable 1 SU113416 29/06/2017 Ground search Litter at mouth of rabbit burrow, arable margin

Bembidion lampros A Ground Beetle Arable 1 SU114413 27/07/2017 Pitfall trap Arable margin

Bembidion properans A Ground Beetle Arable 1 SU114413 27/07/2017 Pitfall trap Arable margin

Bembidion obtusum A Ground Beetle Arable 1 SU114413 27/07/2017 Pitfall trap Arable margin

Poecilus cupreus A Ground Beetle Arable 1 SU114413 27/07/2017 Pitfall trap Arable margin

Poecilus versicolor A Ground Beetle Arable 1 SU114413 27/07/2017 Pitfall trap Arable margin

Pterostichus madidus A Ground Beetle Arable 1 SU114413 27/07/2017 Pitfall trap Arable margin

Pterostichus melanarius A Ground Beetle Arable 1 SU114413 27/07/2017 Pitfall trap Arable margin

Calathus cinctus A Ground Beetle Arable 1 SU114413 27/07/2017 Pitfall trap Arable margin

Calathus fuscipes A Ground Beetle Arable 1 SU114413 27/07/2017 Pitfall trap Arable margin

Calathus melanocephalus A Ground Beetle Arable 1 SU114413 27/07/2017 Pitfall trap Arable margin

Anchomenus dorsalis A Ground Beetle Arable 1 SU114413 27/07/2017 Pitfall trap Arable margin

Zabrus tenebrioides A Ground Beetle NS. Arable 1 SU114413 27/07/2017 Pitfall trap Arable margin Amara convexior A Ground Beetle Arable 1 SU114413 27/07/2017 Pitfall trap Arable margin

Amara familiaris A Ground Beetle Arable 1 SU113416 29/06/2017 Ground search Under stone, arable margin

Amara ovata A Ground Beetle Arable 1 SU114413 27/07/2017 Pitfall trap Arable margin

Amara similata A Ground Beetle Arable 1 SU114413 27/07/2017 Pitfall trap Arable margin

Harpalus rufipes A Ground Beetle Arable 1 SU114413 27/07/2017 Pitfall trap Arable margin

Harpalus affinis A Ground Beetle Arable 1 SU114413 27/07/2017 Pitfall trap Arable margin

Harpalus rubripes A Ground Beetle Arable 1 SU114413 27/07/2017 Pitfall trap Arable margin

Harpalus tardus A Ground Beetle Arable 1 SU114413 27/07/2017 Pitfall trap Arable margin

Syntomus foveatus A Ground Beetle Arable 1 SU114413 27/07/2017 Pitfall trap Arable margin

Silpha atrata A Carrion Beetle Arable 1 SU114413 27/07/2017 Pitfall trap Arable margin

Silpha tristis A Carrion Beetle Arable 1 SU114413 27/07/2017 Pitfall trap Arable margin

Tachinus rufipes A Rove Beetle Arable 1 SU114413 27/07/2017 Pitfall trap Arable margin

Tachyporus dispar A Rove Beetle Arable 1 SU114413 27/07/2017 Pitfall trap Arable margin

Aloconota gregaria A Rove Beetle Arable 1 SU114413 27/07/2017 Pitfall trap Arable margin

Microdota amicula A Rove Beetle Arable 1 SU114413 27/07/2017 Pitfall trap Arable margin

Scientific Name English Name Status Site Gridref Date Method Habitat Mocyta fungi s.s. A Rove Beetle Arable 1 SU114413 27/07/2017 Pitfall trap Arable margin

Falagrioma thoracica A Rove Beetle Arable 1 SU113416 29/06/2017 Ground search Litter at mouth of rabbit burrow, arable margin

Falagrioma thoracica A Rove Beetle Arable 1 SU114413 27/07/2017 Pitfall trap Arable margin

Drusilla canaliculata A Rove Beetle Arable 1 SU114413 27/07/2017 Pitfall trap Arable margin

Oxypoda exoleta A Rove Beetle N. Arable 1 SU114413 27/07/2017 Pitfall trap Arable margin Rugilus similis A Rove Beetle N. Arable 1 SU114413 27/07/2017 Pitfall trap Arable margin Philonthus cognatus A Rove Beetle Arable 1 SU114413 27/07/2017 Pitfall trap Arable margin

Philonthus laminatus A Rove Beetle Arable 1 SU114413 27/07/2017 Pitfall trap Arable margin

Quedius schatzmayri A Rove Beetle Arable 1 SU114413 27/07/2017 Pitfall trap Arable margin Swarming around hawthorn bushes along track Amphimallon solstitialis Summer Chafer Arable 1 SU116413 26/07/2017 Spot search at dusk

Agrypnus murinus A Click Beetle Arable 1 SU111414 29/06/2017 Spot search Singing, rank arable margin

Agriotes obscurus A Click Beetle Arable 1 SU114413 27/07/2017 Pitfall trap Arable margin

Agriotes sputator A Click Beetle Arable 1 SU114413 27/07/2017 Pitfall trap Arable margin

Rhagonycha fulva A Soldier Beetle Arable 1 SU113416 29/06/2017 Sweeping Rank arable margin

Rhagonycha fulva A Soldier Beetle Arable 1 SU112414 27/07/2017 Spot search Arable margin

Meligethes aeneus A Pollen Beetle Arable 1 SU114413 27/07/2017 Pitfall trap Arable margin

Phalacrus championi A Phalacrid Beetle Na. Arable 1 SU112414 29/06/2017 Sweeping Arable margin Stilbus testaceus A Phalacrid Beetle Arable 1 SU112414 27/07/2017 Tapping Carduus nutans, arable margin

Rhyzobius litura A Ladybird Arable 1 SU112414 27/07/2017 Sweeping Arable margin Propylea quatturodecimpunctata 14-spot Ladybird Arable 1 SU112414 27/07/2017 Sweeping Arable margin Psyllobora vigintiduopunctata 22-spot Ladybird Arable 1 SU112414 27/07/2017 Sweeping Arable margin Coccinella septempunctata 7-spot Ladybird Arable 1 SU112414 27/07/2017 Spot search Arable margin

Harmonia axyridis Harlequin Ladybird Arable 1 SU112414 27/07/2017 Spot search Arable margin

Oedemera lurida An Oedemerid Beetle Arable 1 SU113416 29/06/2017 Sweeping Rank arable margin

Oedemera nobilis An Oedemerid Beetle Arable 1 SU113416 29/06/2017 Sweeping Rank arable margin

Oedemera nobilis An Oedemerid Beetle Arable 1 SU112414 27/07/2017 Spot search Arable margin

Paracorymbia fulva A Longhorn Beetle RDB3. Arable 1 SU113416 29/06/2017 Sweeping On Knautia arvensis, rank arable margin Gastrophysa polygoni A Leaf Beetle Arable 1 SU112414 27/07/2017 Sweeping Arable margin

Phyllotreta nigripes A Flea Beetle Arable 1 SU112414 27/07/2017 Tapping Reseda luteola, arable margin

Scientific Name English Name Status Site Gridref Date Method Habitat Sphaeroderma rubidum A Flea Beetle Arable 1 SU112414 27/07/2017 Tapping Carduus nutans, arable margin

Psylliodes chalcomera A Flea Beetle NS. Arable 1 SU112414 27/07/2017 Tapping Carduus nutans, arable margin Psylliodes chrysocephala A Flea Beetle Arable 1 SU112414 27/07/2017 Tapping Carduus nutans, arable margin

Cassida rubiginosa A Tortoise Beetle Arable 1 SU112414 27/07/2017 Tapping Carduus nutans, arable margin

Ceratapion carduorum A Weevil Arable 1 SU112414 27/07/2017 Tapping Carduus nutans, arable margin

Ceratapion gibbirostre A Weevil Arable 1 SU114412 29/06/2017 Tapping Carduus nutans, arable margin

Holotrichapion aethiops A Weevil Arable 1 SU114413 26/07/2017 Tapping Vicia craca, arable margin

Catapion pubescens A Weevil Nb. Arable 1 SU114412 29/06/2017 Tapping Trifolium medium etc, arable margin Ischnopterapion virens A Weevil Arable 1 SU114412 29/06/2017 Tapping Trifolium medium etc, arable margin

Ischnopterapion virens A Weevil Arable 1 SU114413 27/07/2017 Pitfall trap Arable margin

Stenopterapion tenue A Weevil Arable 1 SU114412 29/06/2017 Tapping Medicago lupulina, arable margin

Protapion apricans A Weevil Arable 1 SU114413 26/07/2017 Sweeping Arable margin

Protapion filirostre A Weevil Nb. Arable 1 SU114413 26/07/2017 Tapping Medicago lupulina, arable margin Protapion fulvipes A Weevil Arable 1 SU114412 29/06/2017 Tapping Trifolium medium etc, arable margin

Protapion fulvipes A Weevil Arable 1 SU114413 27/07/2017 Pitfall trap Arable margin

Protapion nigritarse A Weevil Arable 1 SU114412 29/06/2017 Tapping Trifolium medium etc, arable margin

Tychius junceus A Weevil Arable 1 SU114412 29/06/2017 Tapping Trifolium medium etc, arable margin

Tychius picirostris A Weevil Arable 1 SU114412 29/06/2017 Tapping Trifolium medium etc, arable margin

Tychius picirostris A Weevil Arable 1 SU114413 27/07/2017 Pitfall trap Arable margin

Tychius pusillus A Weevil Nb. Arable 1 SU114412 29/06/2017 Tapping Trifolium medium etc, arable margin Ceutorhynchus obstrictus A Weevil Arable 1 SU112414 27/07/2017 Tapping Reseda luteola, arable margin Ceutorhynchus pallidactylus A Weevil Arable 1 SU112414 27/07/2017 Tapping Reseda luteola, arable margin

Mogulones asperifoliarum A Weevil Arable 1 SU114412 29/06/2017 Tapping Myosotis etc, arable margin

Zacladus exiguus A Weevil Nb. Arable 1 SU114412 29/06/2017 Tapping Geranium, arable margin Sirocalodes depressicollis A Weevil Arable 1 SU114412 29/06/2017 Tapping Fumaria, arable margin

Phyllobius viridiaeris A Weevil Arable 1 SU114412 29/06/2017 Tapping Trifolium medium etc, arable margin

Sitona hispidulus A Weevil Arable 1 SU114413 27/07/2017 Pitfall trap Arable margin

Sitona humeralis A Weevil Arable 1 SU114413 26/07/2017 Sweeping Arable margin

Sitona humeralis A Weevil Arable 1 SU114413 27/07/2017 Pitfall trap Arable margin

Sitona lineatus A Weevil Arable 1 SU114412 29/06/2017 Tapping Trifolium medium etc, arable margin

Sitona lineatus A Weevil Arable 1 SU114413 26/07/2017 Sweeping Arable margin

Scientific Name English Name Status Site Gridref Date Method Habitat Larinus planus A Weevil Nb. Arable 1 SU114413 29/06/2017 Spot search Cirsium vulgare, arable margin Thymelicus lineola Essex Skipper Arable 1 SU112414 29/06/2017 Spot search On flowers, rank arable margin

Pieris brassicae Large White Arable 1 SU112414 27/07/2017 Spot search Arable margin

Pieris napi Green-veined White Arable 1 SU112414 29/06/2017 Spot search In flight, rank arable margin

Aricia agestis Brown Argus Arable 1 SU112414 27/07/2017 Spot search Arable margin

Polyommatus icarus Common Blue Arable 1 SU112414 27/07/2017 Spot search Arable margin

Pyronia tithonus Gatekeeper Arable 1 SU112414 27/07/2017 Spot search Arable margin

Maniola jurtina Meadow Brown Arable 1 SU112414 29/06/2017 Spot search In flight, rank arable margin

Maniola jurtina Meadow Brown Arable 1 SU112414 27/07/2017 Spot search Arable margin

Coenonympha pamphilus Small Heath Arable 1 SU112414 29/06/2017 Spot search In flight, rank arable margin

Chloromyia formosa a Soldierfly Arable 1 SU114413 27/07/2017 Pitfall trap Arable margin

Empis livida A Dance Fly Arable 1 SU113416 29/06/2017 Sweeping Rank arable margin

Episyrphus balteatus A Hoverfly Arable 1 SU112414 29/06/2017 Spot search On flowers, rank arable margin

Urophora solstitialis A Picture-winged Fly RDB3. Arable 1 SU112414 27/07/2017 Tapping Carduus nutans, arable margin Bombus lucorum White-tailed Bumblebee Arable 1 SU112414 27/07/2017 Spot search Arable margin

Bombus terrestris Buff-tailed Bumblebee Arable 1 SU112414 27/07/2017 Spot search Arable margin

Bombus lapidarius Red-tailed Bumblebee Arable 1 SU112414 29/06/2017 Spot search On flowers, rank arable margin

Bombus lapidarius Red-tailed Bumblebee Arable 1 SU112414 27/07/2017 Spot search Arable margin

Bombus pascuorum Common Carder Bumblebee Arable 1 SU112414 29/06/2017 Spot search On flowers, rank arable margin

Bombus pascuorum Common Carder Bumblebee Arable 1 SU112414 29/06/2017 Spot search In flight, rank arable margin

Myrmica sabuleti A Red Ant Arable 1 SU114413 27/07/2017 Pitfall trap Arable margin

Myrmica scabrinodis A Red Ant Arable 1 SU114413 27/07/2017 Pitfall trap Arable margin

Araneus diadematus Garden Cross Spider Arable 1 SU112414 27/07/2017 Spot search Arable margin

Candidula intersecta Wrinkled Snail Arable 2 SU112416 29/07/2017 Pitfall trap Restoration grassland

Cepaea nemoralis Brown-lipped Snail Arable 2 SU117416 27/06/2017 Spot search Scrubby grassland by track

Metrioptera roeselii Roesel's Bush-cricket Arable 2 SU115416 27/06/2017 Sweeping Restoration grassland Singing, restoration grassland with much Metrioptera roeselii Roesel's Bush-cricket Arable 2 SU115416 27/07/2017 Spot search sainfoin Common Green Omocestus viridulus Grasshopper Arable 2 SU115416 27/06/2017 Sweeping Restoration grassland

Carabus violaceus A Ground Beetle Arable 2 SU112416 29/07/2017 Pitfall trap Restoration grassland

Nebria salina A Ground Beetle Arable 2 SU112416 29/07/2017 Pitfall trap Restoration grassland

Scientific Name English Name Status Site Gridref Date Method Habitat Poecilus versicolor A Ground Beetle Arable 2 SU112416 29/07/2017 Pitfall trap Restoration grassland

Pterostichus madidus A Ground Beetle Arable 2 SU112416 29/07/2017 Pitfall trap Restoration grassland

Pterostichus melanarius A Ground Beetle Arable 2 SU112416 29/07/2017 Pitfall trap Restoration grassland

Calathus fuscipes A Ground Beetle Arable 2 SU112416 29/07/2017 Pitfall trap Restoration grassland

Calathus melanocephalus A Ground Beetle Arable 2 SU112416 29/07/2017 Pitfall trap Restoration grassland

Anchomenus dorsalis A Ground Beetle Arable 2 SU112416 29/07/2017 Pitfall trap Restoration grassland

Amara convexior A Ground Beetle Arable 2 SU112416 29/07/2017 Pitfall trap Restoration grassland

Amara lunicollis A Ground Beetle Arable 2 SU112416 29/07/2017 Pitfall trap Restoration grassland

Amara ovata A Ground Beetle Arable 2 SU112416 29/07/2017 Pitfall trap Restoration grassland

Harpalus rufipes A Ground Beetle Arable 2 SU112416 29/07/2017 Pitfall trap Restoration grassland

Harpalus affinis A Ground Beetle Arable 2 SU112416 29/07/2017 Pitfall trap Restoration grassland

Badister bullatus A Ground Beetle Arable 2 SU112416 29/07/2017 Pitfall trap Restoration grassland

Paradromius linearis A Ground Beetle Arable 2 SU115416 27/06/2017 Sweeping Restoration grassland

Megasternum concinnum A Hydrophilid Beetle Arable 2 SU112416 29/07/2017 Pitfall trap Restoration grassland Cow dung, restoration chalk grassland with Hister unicolor A Clown Beetle Arable 2 SU113416 26/07/2017 Dung abundant sainfoin

Tachinus rufipes A Rove Beetle Arable 2 SU112416 29/07/2017 Pitfall trap Restoration grassland

Tachyporus nitidulus A Rove Beetle Arable 2 SU112416 29/07/2017 Pitfall trap Restoration grassland Cow dung, restoration chalk grassland with Aleochara discipennis A Rove Beetle NS. Arable 2 SU113416 26/07/2017 Dung abundant sainfoin Cow dung, restoration chalk grassland with Aleochara lanuginosa A Rove Beetle Arable 2 SU113416 26/07/2017 Dung abundant sainfoin

Amischa nigrofusca A Rove Beetle Arable 2 SU112416 29/07/2017 Pitfall trap Restoration grassland

Dinaraea angustula A Rove Beetle Arable 2 SU112416 29/07/2017 Pitfall trap Restoration grassland

Pycnota paradoxa A Rove Beetle N. Arable 2 SU112416 29/07/2017 Pitfall trap Restoration grassland Falagrioma thoracica A Rove Beetle Arable 2 SU112416 29/07/2017 Pitfall trap Restoration grassland

Drusilla canaliculata A Rove Beetle Arable 2 SU112416 29/07/2017 Pitfall trap Restoration grassland Cow dung, restoration chalk grassland with Philonthus albipes A Rove Beetle Arable 2 SU113416 26/07/2017 Dung abundant sainfoin

Philonthus carbonarius A Rove Beetle Arable 2 SU112416 29/07/2017 Pitfall trap Restoration grassland

Philonthus cognatus A Rove Beetle Arable 2 SU112416 29/07/2017 Pitfall trap Restoration grassland Cow dung, restoration chalk grassland with Philonthus cruentatus A Rove Beetle Arable 2 SU113416 26/07/2017 Dung abundant sainfoin

Philonthus laminatus A Rove Beetle Arable 2 SU112416 29/07/2017 Pitfall trap Restoration grassland

Scientific Name English Name Status Site Gridref Date Method Habitat Cow dung, restoration chalk grassland with Philonthus sanguinolentus A Rove Beetle Arable 2 SU113416 26/07/2017 Dung abundant sainfoin Cow dung, restoration chalk grassland with Philonthus splendens A Rove Beetle Arable 2 SU113416 26/07/2017 Dung abundant sainfoin Cow dung, restoration chalk grassland with Philonthus varians A Rove Beetle Arable 2 SU113416 26/07/2017 Dung abundant sainfoin

Quedius levicollis A Rove Beetle Arable 2 SU112416 29/07/2017 Pitfall trap Restoration grassland Cow dung, restoration chalk grassland with Gyrohypnus fracticornis A Rove Beetle Arable 2 SU113416 26/07/2017 Dung abundant sainfoin

Aplotarsus incanus A Click Beetle Arable 2 SU115416 27/06/2017 Sweeping Restoration grassland

Agriotes sputator A Click Beetle Arable 2 SU112416 29/07/2017 Pitfall trap Restoration grassland

Meligethes aeneus A Pollen Beetle Arable 2 SU115416 27/06/2017 Tapping On Onobrychis viciifolia, restoration grassland Propylea quatturodecimpunctata 14-spot Ladybird Arable 2 SU115416 27/06/2017 Sweeping Restoration grassland Coccinella septempunctata 7-spot Ladybird Arable 2 SU115416 27/06/2017 Sweeping Restoration grassland Coccinella Linaria vulgaris, rank grassland along track septempunctata 7-spot Ladybird Arable 2 SU117416 26/07/2017 Tapping edge Subcoccinella vigintiquattuorpunctata 24-spot Ladybird Arable 2 SU115416 27/06/2017 Tapping On Vicia cracca, restoration grassland Subcoccinella Linaria vulgaris, rank grassland along track vigintiquattuorpunctata 24-spot Ladybird Arable 2 SU117416 26/07/2017 Tapping edge

Oedemera lurida An Oedemerid Beetle Arable 2 SU115416 27/06/2017 Sweeping Restoration grassland

Oedemera nobilis An Oedemerid Beetle Arable 2 SU115416 27/06/2017 Sweeping Restoration grassland

Paracorymbia fulva A Longhorn Beetle RDB3. Arable 2 SU115416 27/06/2017 Spot search On Knautia arvensis, restoration grassland Agapanthia villosoviridescens A Longhorn Beetle Arable 2 SU114415 27/06/2017 Spot search On Heracleum sphondylium, scrub edge

Cryptocephalus moraei A Leaf Beetle Arable 2 SU115416 27/06/2017 Sweeping Restoration grassland

Cryptocephalus moraei A Leaf Beetle Arable 2 SU112416 29/07/2017 Pitfall trap Restoration grassland

Phyllobius viridiaeris A Weevil Arable 2 SU115416 27/06/2017 Sweeping Restoration grassland

Sitona humeralis A Weevil Arable 2 SU115416 27/06/2017 Sweeping Restoration grassland

Sitona lineatus A Weevil Arable 2 SU115416 27/06/2017 Sweeping Restoration grassland

Hypera venusta A Weevil Arable 2 SU115416 27/06/2017 Sweeping Restoration grassland

Hypera venusta A Weevil Arable 2 SU112416 29/07/2017 Pitfall trap Restoration grassland

Hemitrichapion reflexum A Weevil Na. Arable 2 SU115416 27/06/2017 Tapping On Onobrychis viciifolia, restoration grassland Holotrichapion pisi A Weevil Arable 2 SU115416 27/06/2017 Sweeping Restoration grassland

Holotrichapion aethiops A Weevil Arable 2 SU115416 27/06/2017 Tapping On Vicia cracca, restoration grassland

Scientific Name English Name Status Site Gridref Date Method Habitat Ischnopterapion loti A Weevil Arable 2 SU115416 27/06/2017 Sweeping Restoration grassland

Ischnopterapion virens A Weevil Arable 2 SU115416 27/06/2017 Sweeping Restoration grassland

Protapion apricans A Weevil Arable 2 SU115416 27/06/2017 Sweeping Restoration grassland

Protapion assimile A Weevil Arable 2 SU115416 27/06/2017 Sweeping Restoration grassland

Protapion trifolii A Weevil Arable 2 SU115416 27/06/2017 Sweeping Restoration grassland Linaria vulgaris, rank grassland along track Rhinusa antirrhini A Weevil Arable 2 SU117416 26/07/2017 Tapping edge

Thymelicus sylvestris Small Skipper Arable 2 SU115416 27/06/2017 Spot search In flight and on flowers, restoration grassland

Ochlodes sylvanus Large Skipper Arable 2 SU117416 27/06/2017 Spot search In flight, scrubby grassland by track

Gonepteryx rhamni Brimstone Arable 2 SU114415 27/06/2017 Spot search In flight and on flowers, scrub edge

Pieris rapae Small White Arable 2 SU117416 27/06/2017 Spot search In flight, scrubby grassland by track

Polyommatus icarus Common Blue Arable 2 SU115416 27/06/2017 Spot search In flight and on flowers, restoration grassland

Melanargia galathea Marbled White Arable 2 SU115416 27/06/2017 Spot search In flight and on flowers, restoration grassland

Pyronia tithonus Gatekeeper Arable 2 SU114415 27/06/2017 Spot search In flight and on flowers, scrub edge Larvae, Urtica dioica, scrubby grassland by Inachis io Peacock Arable 2 SU117416 27/06/2017 Spot search track

Aglais urticae Small Tortoiseshell Arable 2 SU117416 27/06/2017 Spot search On flowers, scrubby grassland by track

Maniola jurtina Meadow Brown Arable 2 SU115416 27/06/2017 Spot search In flight and on flowers, restoration grassland On flowers and in flight, restoration grassland Maniola jurtina Meadow Brown Arable 2 SU115416 27/07/2017 Spot search with much sainfoin

Aphantopus hyperantus Ringlet Arable 2 SU115416 27/06/2017 Spot search In flight and on flowers, restoration grassland

Coenonympha pamphilus Small Heath Arable 2 SU115416 27/06/2017 Spot search In flight and on flowers, restoration grassland On flowers and in flight, restoration grassland Coenonympha pamphilus Small Heath Arable 2 SU115416 27/07/2017 Spot search with much sainfoin

Chloromyia formosa a Soldierfly Arable 2 SU117416 27/06/2017 Spot search On flowers, scrubby grassland by track

Paracrocera orbiculus An Acrocerid Fly Arable 2 SU115416 27/06/2017 Spot search In flight, restoration grassland

Machimus atricapillus A Robberfly Arable 2 SU115416 27/06/2017 Sweeping Restoration grassland

Leptarthrus brevirostris A Robberfly Arable 2 SU115416 27/06/2017 Sweeping Restoration grassland

Leptogaster cylindrica A Robberfly Arable 2 SU115416 27/06/2017 Sweeping Restoration grassland

Empis livida A Dance Fly Arable 2 SU115416 27/06/2017 Spot search In flight and on flowers, scrub edge

Episyrphus balteatus A Hoverfly Arable 2 SU115416 27/06/2017 Spot search On flowers, restoration grassland

Volucella bombylans A Hoverfly Arable 2 SU115416 27/06/2017 Spot search In flight and on flowers, scrub edge

Physocephala rufipes A Big-headed Fly Arable 2 SU115416 27/06/2017 Sweeping Restoration grassland

Scientific Name English Name Status Site Gridref Date Method Habitat Cow dung, restoration grassland with much Mesembrina meridiana A House Fly Arable 2 SU115416 27/07/2017 Dung sainfoin

Apis mellifera Honeybee Arable 2 SU117416 27/06/2017 Spot search On flowers, scrubby grassland by track

Bombus terrestris Buff-tailed Bumblebee Arable 2 SU115416 27/06/2017 Spot search In flight and on flowers, restoration grassland

Bombus lucorum White-tailed Bumblebee Arable 2 SU117416 27/06/2017 Spot search On flowers, scrubby grassland by track

Bombus lapidarius Red-tailed Bumblebee Arable 2 SU115416 27/06/2017 Spot search In flight and on flowers, restoration grassland

Bombus hypnorum Tree Bumblebee Arable 2 SU117416 27/06/2017 Spot search On flowers, scrubby grassland by track

Bombus jonellus Heath Bumblebee Arable 2 SU115416 27/06/2017 Spot search In flight and on flowers, restoration grassland

Bombus pratorum Early Bumblebee Arable 2 SU117416 27/06/2017 Spot search On flowers, scrubby grassland by track

Bombus pascuorum Common Carder Bumblebee Arable 2 SU117416 27/06/2017 Spot search On flowers, scrubby grassland by track

Myrmica ruginodis A Red Ant Arable 2 SU112416 29/07/2017 Pitfall trap Restoration grassland

Myrmica sabuleti A Red Ant Arable 2 SU112416 29/07/2017 Pitfall trap Restoration grassland

Myrmica scabrinodis A Red Ant Arable 2 SU112416 29/07/2017 Pitfall trap Restoration grassland

Porcellio scaber Common Rough Woodlouse Arable 2 SU112416 29/07/2017 Pitfall trap Restoration grassland

Aegopinella nitidula Dull Glass Snail Arable 3 SU137420 26/07/2017 Pitfall trap Arable margin

Candidula intersecta Wrinkled Snail Arable 3 SU137420 26/07/2017 Pitfall trap Arable margin

Monacha cantiana Kentish Snail Arable 3 SU137420 30/06/2017 Sweeping Arable margin

Monacha cantiana Kentish Snail Arable 3 SU137420 26/07/2017 Pitfall trap Arable margin

Trichia striolata Strawberry Snail Arable 3 SU137420 26/07/2017 Pitfall trap Arable margin

Trichia hispida Hairy Snail Arable 3 SU137420 26/07/2017 Pitfall trap Arable margin

Cepaea nemoralis Brown-lipped Snail Arable 3 SU137420 30/06/2017 Sweeping Arable margin

Cepaea nemoralis Brown-lipped Snail Arable 3 SU137420 26/07/2017 Pitfall trap Arable margin

Helix aspersa Garden Snail Arable 3 SU137420 30/06/2017 Sweeping Arable margin

Helix aspersa Garden Snail Arable 3 SU137420 26/07/2017 Pitfall trap Arable margin

Metrioptera roeselii Roesel's Bush-cricket Arable 3 SU141420 30/06/2017 Spot search Singing, scrubby chalk grassland

Chorthippus parallelus Meadow Grasshopper Arable 3 SU137420 30/06/2017 Sweeping Singing, arable margin

Chorthippus parallelus Meadow Grasshopper Arable 3 SU137420 26/07/2017 Pitfall trap Arable margin

Eurydema oleracea Brassica Bug Arable 3 SU137420 26/07/2017 Tapping Brassica napus, arable margin

Eurydema oleracea Brassica Bug Arable 3 SU137420 26/07/2017 Sweeping Arable margin

Carabus violaceus A Ground Beetle Arable 3 SU137420 26/07/2017 Pitfall trap Arable margin

Nebria brevicollis A Ground Beetle Arable 3 SU137420 26/07/2017 Pitfall trap Arable margin

Pterostichus madidus A Ground Beetle Arable 3 SU137420 26/07/2017 Pitfall trap Arable margin

Scientific Name English Name Status Site Gridref Date Method Habitat Pterostichus melanarius A Ground Beetle Arable 3 SU137420 30/06/2017 Ground search Under stone, arable margin

Pterostichus melanarius A Ground Beetle Arable 3 SU137420 26/07/2017 Pitfall trap Arable margin

Calathus fuscipes A Ground Beetle Arable 3 SU137420 30/06/2017 Sweeping Arable margin

Calathus fuscipes A Ground Beetle Arable 3 SU137420 26/07/2017 Pitfall trap Arable margin

Anchomenus dorsalis A Ground Beetle Arable 3 SU137420 30/06/2017 Ground search Litter, arable margin

Anchomenus dorsalis A Ground Beetle Arable 3 SU137420 26/07/2017 Pitfall trap Arable margin

Curtonotus aulicus A Ground Beetle Arable 3 SU137420 26/07/2017 Pitfall trap Arable margin

Harpalus rufipes A Ground Beetle Arable 3 SU137420 26/07/2017 Pitfall trap Arable margin

Harpalus affinis A Ground Beetle Arable 3 SU137420 26/07/2017 Pitfall trap Arable margin

Badister bullatus A Ground Beetle Arable 3 SU137420 26/07/2017 Pitfall trap Arable margin

Demetrias atricapillus A Ground Beetle Arable 3 SU137420 26/07/2017 Sweeping Arable margin

Megasternum concinnum A Hydrophilid Beetle Arable 3 SU137420 26/07/2017 Pitfall trap Arable margin

Sciodrepoides watsoni A Leiodid Beetle Arable 3 SU137420 26/07/2017 Pitfall trap Arable margin

Ptomaphagus medius A Leiodid Beetle Arable 3 SU137420 26/07/2017 Pitfall trap Arable margin RDBK Ptomaphagus varicornis A Leiodid Beetle . Arable 3 SU137420 26/07/2017 Pitfall trap Arable margin Silpha laevigata A Carrion Beetle Arable 3 SU137420 26/07/2017 Pitfall trap Arable margin

Nicrophorus interruptus A Sexton Beetle Nb. Arable 3 SU137420 26/07/2017 Pitfall trap Arable margin Omalium caesum A Rove Beetle Arable 3 SU137420 26/07/2017 Pitfall trap Arable margin

Tachinus rufipes A Rove Beetle Arable 3 SU137420 26/07/2017 Pitfall trap Arable margin

Tachyporus hypnorum A Rove Beetle Arable 3 SU137420 26/07/2017 Pitfall trap Arable margin

Tachyporus nitidulus A Rove Beetle Arable 3 SU141420 30/06/2017 Tapping Hypericum, scrubby chalk grassland

Tachyporus nitidulus A Rove Beetle Arable 3 SU137420 26/07/2017 Pitfall trap Arable margin

Tachyporus obtusus A Rove Beetle Arable 3 SU137420 26/07/2017 Sweeping Arable margin

Aleochara curtula A Rove Beetle Arable 3 SU137420 26/07/2017 Pitfall trap Arable margin

Aleochara bipustulata A Rove Beetle Arable 3 SU137420 26/07/2017 Pitfall trap Arable margin

Alaobia trinotata A Rove Beetle Arable 3 SU137420 30/06/2017 Ground search Litter, arable margin

Atheta xanthopus A Rove Beetle Arable 3 SU137420 26/07/2017 Pitfall trap Arable margin

Atheta crassicornis A Rove Beetle Arable 3 SU137420 26/07/2017 Pitfall trap Arable margin

Atheta oblita A Rove Beetle Arable 3 SU137420 26/07/2017 Pitfall trap Arable margin

Mocyta fungi s.s. A Rove Beetle Arable 3 SU137420 26/07/2017 Pitfall trap Arable margin

Mycetota laticollis A Rove Beetle Arable 3 SU137420 26/07/2017 Pitfall trap Arable margin

Scientific Name English Name Status Site Gridref Date Method Habitat Falagrioma thoracica A Rove Beetle Arable 3 SU137420 26/07/2017 Pitfall trap Arable margin

Drusilla canaliculata A Rove Beetle Arable 3 SU137420 26/07/2017 Ground search Under stone, arable margin

Drusilla canaliculata A Rove Beetle Arable 3 SU137420 26/07/2017 Pitfall trap Arable margin

Anotylus inustus A Rove Beetle Arable 3 SU137420 30/06/2017 Tapping Alliaria petiolata, arable margin

Anotylus sculpturatus A Rove Beetle Arable 3 SU137420 30/06/2017 Ground search Litter, arable margin

Sunius melanocephalus A Rove Beetle N. Arable 3 SU137420 26/07/2017 Pitfall trap Arable margin Rugilus rufipes A Rove Beetle Arable 3 SU137420 30/06/2017 Ground search Litter, arable margin

Rhagonycha fulva A Soldier Beetle Arable 3 SU141420 30/06/2017 Tapping Hypericum, scrubby chalk grassland

Anobium inexspectatum A Woodworm Beetle Nb. Arable 3 SU137420 30/06/2017 Tapping Alliaria petiolata, arable margin Meligethes bidens A Pollen Beetle Arable 3 SU141420 30/06/2017 Tapping Clinopodium vulgare, scrubby chalk grassland Propylea quatturodecimpunctata 14-spot Ladybird Arable 3 SU137420 26/07/2017 Sweeping Arable margin Coccinella septempunctata 7-spot Ladybird Arable 3 SU137420 26/07/2017 Sweeping Arable margin

Harmonia axyridis Harlequin Ladybird Arable 3 SU141420 30/06/2017 Spot search On foliage, scrubby chalk grassland Subcoccinella vigintiquattuorpunctata 24-spot Ladybird Arable 3 SU137420 26/07/2017 Sweeping Arable margin Subcoccinella vigintiquattuorpunctata 24-spot Ladybird Arable 3 SU137420 26/07/2017 Pitfall trap Arable margin

Oedemera lurida An Oedemerid Beetle Arable 3 SU137420 30/06/2017 Sweeping Arable margin

Oedemera nobilis An Oedemerid Beetle Arable 3 SU141420 30/06/2017 Tapping Hypericum, scrubby chalk grassland

Paracorymbia fulva A Longhorn Beetle RDB3. Arable 3 SU137420 30/06/2017 Sweeping Arable margin Phyllotreta diademata A Flea Beetle Arable 3 SU137420 26/07/2017 Pitfall trap Arable margin

Phyllotreta nigripes A Flea Beetle Arable 3 SU137420 26/07/2017 Sweeping Arable margin

Psylliodes chrysocephala A Flea Beetle Arable 3 SU137420 30/06/2017 Tapping Alliaria petiolata, arable margin

Psylliodes chrysocephala A Flea Beetle Arable 3 SU137420 26/07/2017 Pitfall trap Arable margin

Ceratapion carduorum A Weevil Arable 3 SU137420 30/06/2017 Tapping Cirsium vulgare, arable margin

Ceratapion gibbirostre A Weevil Arable 3 SU137420 30/06/2017 Tapping Cirsium vulgare, arable margin

Taeniapion urticarium A Weevil Arable 3 SU137420 30/06/2017 Sweeping Arable margin

Ceutorhynchus obstrictus A Weevil Arable 3 SU137420 30/06/2017 Tapping Alliaria petiolata, arable margin

Ceutorhynchus obstrictus A Weevil Arable 3 SU137420 26/07/2017 Sweeping Arable margin

Ceutorhynchus typhae A Weevil Arable 3 SU137420 26/07/2017 Pitfall trap Arable margin

Sirocalodes depressicollis A Weevil Arable 3 SU137420 30/06/2017 Tapping Fumaria, arable margin

Scientific Name English Name Status Site Gridref Date Method Habitat Sirocalodes depressicollis A Weevil Arable 3 SU137420 26/07/2017 Tapping Fumaria, arable margin

Sirocalodes quercicola A Weevil Na. Arable 3 SU137420 26/07/2017 Tapping Fumaria, arable margin Thamiocolus viduatus A Weevil Nb. Arable 3 SU137420 30/06/2017 Spot search Stachys palustris, arable margin Thamiocolus viduatus A Weevil Nb. Arable 3 SU137420 26/07/2017 Tapping Stachys palustris, arable margin Barypeithes pellucidus A Weevil Arable 3 SU137420 30/06/2017 Tapping Alliaria petiolata, arable margin

Barypeithes pellucidus A Weevil Arable 3 SU137420 26/07/2017 Pitfall trap Arable margin

Thymelicus lineola Essex Skipper Arable 3 SU141420 30/06/2017 Spot search In flight, scrubby chalk grassland

Thymelicus sylvestris Small Skipper Arable 3 SU137420 30/06/2017 Spot search In flight, arable margin

Vanessa atalanta Red Admiral Arable 3 SU135420 30/06/2017 Spot search In flight, arable margin

Aglais urticae Small Tortoiseshell Arable 3 SU141420 30/06/2017 Spot search On flowers, scrubby chalk grassland

Maniola jurtina Meadow Brown Arable 3 SU141420 30/06/2017 Spot search In flight, scrubby chalk grassland

Aphantopus hyperantus Ringlet Arable 3 SU135420 30/06/2017 Spot search In flight, arable margin

Empis livida A Dance Fly Arable 3 SU137420 30/06/2017 Sweeping Arable margin

Episyrphus balteatus A Hoverfly Arable 3 SU135420 30/06/2017 Spot search On flowers, arable margin

Scaeva pyrastri A Hoverfly Arable 3 SU141420 30/06/2017 Spot search On flowers, scrubby chalk grassland

Bombus lapidarius Red-tailed Bumblebee Arable 3 SU137420 26/07/2017 Spot search On flowers, arable margin

Bombus pascuorum Common Carder Bumblebee Arable 3 SU137420 26/07/2017 Spot search On flowers, arable margin

Lasius flavus Yellow Meadow Ant Arable 3 SU141420 30/06/2017 Spot search Nest mounds, scrubby chalk grassland

Lasius niger Black Garden Ant Arable 3 SU137420 30/06/2017 Ground search Under stone, arable margin

Myrmica ruginodis A Red Ant Arable 3 SU137420 26/07/2017 Pitfall trap Arable margin

Nemastoma bimaculatum A Harvestman Spider Arable 3 SU137420 30/06/2017 Ground search Litter, arable margin

Trichoniscus pusillus Common Pygmy Woodlouse Arable 3 SU137420 30/06/2017 Ground search Under stone, arable margin

Philoscia muscorum Striped Woodlouse Arable 3 SU137420 30/06/2017 Ground search Under stone, arable margin

Oniscus asellus Common Shiny Woodlouse Arable 3 SU137420 30/06/2017 Ground search Litter, arable margin

Oniscus asellus Common Shiny Woodlouse Arable 3 SU137420 26/07/2017 Pitfall trap Arable margin

Porcellio scaber Common Rough Woodlouse Arable 3 SU137420 30/06/2017 Ground search Under bark, dead branch, arable margin

Armadillidium vulgare Common Pill Woodlouse Arable 3 SU137420 30/06/2017 Ground search Litter, arable margin

Armadillidium vulgare Common Pill Woodlouse Arable 3 SU137420 26/07/2017 Pitfall trap Arable margin

Discus rotundatus Radiated Snail Countess Cutting SU145421 26/07/2017 Pitfall trap Chalk cutting with patchy vegetation

Arion ater agg. Large Black Slug Countess Cutting SU145421 30/06/2017 Ground search Under stone, chalk cutting

Aegopinella nitidula Dull Glass Snail Countess Cutting SU145421 30/06/2017 Ground search Under stone, chalk cutting

Scientific Name English Name Status Site Gridref Date Method Habitat Candidula intersecta Wrinkled Snail Countess Cutting SU145421 26/07/2017 Pitfall trap Chalk cutting with patchy vegetation

Cernuella virgata Striped Snail Countess Cutting SU145421 30/06/2017 Ground search Moss and litter, chalk cutting

Cernuella virgata Striped Snail Countess Cutting SU145421 26/07/2017 Pitfall trap Chalk cutting with patchy vegetation

Helicella itala Heath Snail Countess Cutting SU145421 30/06/2017 Ground search Moss and litter, chalk cutting

Helicella itala Heath Snail Countess Cutting SU145421 26/07/2017 Pitfall trap Chalk cutting with patchy vegetation

Monacha cantiana Kentish Snail Countess Cutting SU145421 30/06/2017 Sweeping Chalk cutting with patchy vegetation

Monacha cantiana Kentish Snail Countess Cutting SU145421 26/07/2017 Pitfall trap Chalk cutting with patchy vegetation

Trichia striolata Strawberry Snail Countess Cutting SU145421 30/06/2017 Ground search Moss and litter, chalk cutting

Trichia striolata Strawberry Snail Countess Cutting SU145421 26/07/2017 Pitfall trap Chalk cutting with patchy vegetation

Cepaea nemoralis Brown-lipped Snail Countess Cutting SU145421 30/06/2017 Sweeping Chalk cutting with patchy vegetation

Cepaea nemoralis Brown-lipped Snail Countess Cutting SU145421 26/07/2017 Pitfall trap Chalk cutting with patchy vegetation

Helix aspersa Garden Snail Countess Cutting SU145421 30/06/2017 Ground search Moss and litter, chalk cutting

Metrioptera roeselii Roesel's Bush-cricket Countess Cutting SU145421 30/06/2017 Sweeping Chalk cutting with patchy vegetation

Chorthippus parallelus Meadow Grasshopper Countess Cutting SU145421 30/06/2017 Sweeping Chalk cutting with patchy vegetation

Carabus violaceus A Ground Beetle Countess Cutting SU145421 26/07/2017 Pitfall trap Chalk cutting with patchy vegetation

Bembidion lampros A Ground Beetle Countess Cutting SU145421 26/07/2017 Pitfall trap Chalk cutting with patchy vegetation

Pterostichus madidus A Ground Beetle Countess Cutting SU145421 26/07/2017 Pitfall trap Chalk cutting with patchy vegetation

Pterostichus melanarius A Ground Beetle Countess Cutting SU145421 26/07/2017 Pitfall trap Chalk cutting with patchy vegetation

Calathus cinctus A Ground Beetle Countess Cutting SU145421 30/06/2017 Ground search Moss and litter, chalk cutting

Calathus fuscipes A Ground Beetle Countess Cutting SU145421 30/06/2017 Ground search Moss and litter, chalk cutting

Calathus fuscipes A Ground Beetle Countess Cutting SU145421 26/07/2017 Pitfall trap Chalk cutting with patchy vegetation

Anchomenus dorsalis A Ground Beetle Countess Cutting SU145421 30/06/2017 Ground search Moss and litter, chalk cutting

Amara bifrons A Ground Beetle Countess Cutting SU145421 26/07/2017 Pitfall trap Chalk cutting with patchy vegetation

Amara montivaga A Ground Beetle Countess Cutting SU145421 26/07/2017 Pitfall trap Chalk cutting with patchy vegetation

Curtonotus aulicus A Ground Beetle Countess Cutting SU145421 30/06/2017 Ground search Moss and litter, chalk cutting

Curtonotus aulicus A Ground Beetle Countess Cutting SU145421 26/07/2017 Pitfall trap Chalk cutting with patchy vegetation

Harpalus rufipes A Ground Beetle Countess Cutting SU145421 26/07/2017 Pitfall trap Chalk cutting with patchy vegetation

Harpalus affinis A Ground Beetle Countess Cutting SU145421 26/07/2017 Pitfall trap Chalk cutting with patchy vegetation

Ophonus puncticeps A Ground Beetle Countess Cutting SU145421 30/06/2017 Ground search Under stone, chalk cutting

Ophonus puncticeps A Ground Beetle Countess Cutting SU145421 26/07/2017 Pitfall trap Chalk cutting with patchy vegetation Ophonus schaubergerianus A Ground Beetle NS. Countess Cutting SU145421 30/06/2017 Ground search Moss and litter, chalk cutting

Scientific Name English Name Status Site Gridref Date Method Habitat Licinus depressus A Ground Beetle NS. Countess Cutting SU145421 26/07/2017 Pitfall trap Chalk cutting with patchy vegetation Licinus punctatulus A Ground Beetle NS. Countess Cutting SU145421 26/07/2017 Pitfall trap Chalk cutting with patchy vegetation Syntomus foveatus A Ground Beetle Countess Cutting SU145421 30/06/2017 Ground search Moss and litter, chalk cutting

Saprinus semistriatus A Clown Beetle Countess Cutting SU145421 26/07/2017 Pitfall trap Chalk cutting with patchy vegetation

Sciodrepoides watsoni A Leiodid Beetle Countess Cutting SU145421 26/07/2017 Pitfall trap Chalk cutting with patchy vegetation

Ptomaphagus medius A Leiodid Beetle Countess Cutting SU145421 26/07/2017 Pitfall trap Chalk cutting with patchy vegetation

Nicrophorus interruptus A Sexton Beetle Nb. Countess Cutting SU145421 26/07/2017 Pitfall trap Chalk cutting with patchy vegetation Lamprinodes saginatus A Rove Beetle Na. Countess Cutting SU145421 30/06/2017 Ground search Moss and litter, chalk cutting Aleochara curtula A Rove Beetle Countess Cutting SU145421 26/07/2017 Pitfall trap Chalk cutting with patchy vegetation

Aleochara lata A Rove Beetle Countess Cutting SU145421 26/07/2017 Pitfall trap Chalk cutting with patchy vegetation

Dinaraea angustula A Rove Beetle Countess Cutting SU145421 26/07/2017 Pitfall trap Chalk cutting with patchy vegetation

Philhygra palustris A Rove Beetle Countess Cutting SU145421 26/07/2017 Pitfall trap Chalk cutting with patchy vegetation

Drusilla canaliculata A Rove Beetle Countess Cutting SU145421 30/06/2017 Ground search Moss and litter, chalk cutting

Drusilla canaliculata A Rove Beetle Countess Cutting SU145421 26/07/2017 Pitfall trap Chalk cutting with patchy vegetation

Platystethus capito A Rove Beetle Countess Cutting SU145421 26/07/2017 Pitfall trap Chalk cutting with patchy vegetation

Ocypus aeneocephalus A Rove Beetle Countess Cutting SU145421 30/06/2017 Ground search Moss and litter, chalk cutting

Ocypus aeneocephalus A Rove Beetle Countess Cutting SU145421 26/07/2017 Pitfall trap Chalk cutting with patchy vegetation

Platydracus stercorarius A Rove Beetle Countess Cutting SU145421 26/07/2017 Pitfall trap Chalk cutting with patchy vegetation

Tasgius morsitans A Rove Beetle Countess Cutting SU145421 26/07/2017 Pitfall trap Chalk cutting with patchy vegetation

Amphimallon solstitialis Summer Chafer Countess Cutting SU145421 26/07/2017 Pitfall trap Chalk cutting with patchy vegetation

Chaetophora spinosa A Pill Beetle Countess Cutting SU145421 26/07/2017 Pitfall trap Chalk cutting with patchy vegetation

Rhagonycha fulva A Soldier Beetle Countess Cutting SU145421 30/06/2017 Sweeping Chalk cutting with patchy vegetation

Meligethes bidens A Pollen Beetle Countess Cutting SU145421 30/06/2017 Tapping Clinopodium vulgare, chalk cutting

Oedemera lurida An Oedemerid Beetle Countess Cutting SU145421 30/06/2017 Sweeping Chalk cutting with patchy vegetation

Oedemera nobilis An Oedemerid Beetle Countess Cutting SU145421 30/06/2017 Sweeping Chalk cutting with patchy vegetation

Chrysolina polita A Leaf Beetle Countess Cutting SU145421 26/07/2017 Pitfall trap Chalk cutting with patchy vegetation

Sermylassa halensis A Leaf Beetle Countess Cutting SU145421 30/06/2017 Ground search Moss and litter, chalk cutting

Hemitrichapion waltoni A Weevil Countess Cutting SU145421 30/06/2017 Tapping Hippocrepis comosa, chalk cutting

Diplapion confluens A Weevil Countess Cutting SU145421 26/07/2017 Pitfall trap Chalk cutting with patchy vegetation

Tychius stephensi A Weevil Countess Cutting SU145421 30/06/2017 Tapping Anthyllis vulneraria, chalk cutting

Mecinus circulatus A Weevil Nb. Countess Cutting SU145421 26/07/2017 Pitfall trap Chalk cutting with patchy vegetation

Scientific Name English Name Status Site Gridref Date Method Habitat Stenocarus ruficornis A Weevil Nb. Countess Cutting SU145421 26/07/2017 Pitfall trap Chalk cutting with patchy vegetation Otiorhynchus ligneus A Weevil Countess Cutting SU145421 26/07/2017 Pitfall trap Chalk cutting with patchy vegetation

Chloromyia formosa A Soldierfly Countess Cutting SU145421 30/06/2017 Sweeping Chalk cutting with patchy vegetation

Thymelicus lineola Essex Skipper Countess Cutting SU145421 30/06/2017 Spot search In flight, chalk cutting

Pieris brassicae Large White Countess Cutting SU145421 30/06/2017 Spot search In flight, chalk cutting

Pieris rapae Small White Countess Cutting SU145421 30/06/2017 Spot search In flight, chalk cutting

Pieris napi Green-veined White Countess Cutting SU145421 30/06/2017 Spot search In flight, chalk cutting

Vanessa atalanta Red Admiral Countess Cutting SU145421 30/06/2017 Spot search In flight, chalk cutting

Thereva nobilitata Common Stiletto Fly Countess Cutting SU145421 26/07/2017 Pitfall trap Chalk cutting with patchy vegetation

Episyrphus balteatus A Hoverfly Countess Cutting SU145421 30/06/2017 Sweeping Chalk cutting with patchy vegetation

Scaeva pyrastri A Hoverfly Countess Cutting SU145421 30/06/2017 Ground search On flowers, chalk cutting

Lasius flavus Yellow Meadow Ant Countess Cutting SU145421 30/06/2017 Ground search Under stone, chalk cutting

Lasius niger Black Garden Ant Countess Cutting SU145421 30/06/2017 Ground search Under stones, chalk cutting

Lasius niger Black Garden Ant Countess Cutting SU145421 26/07/2017 Pitfall trap Chalk cutting with patchy vegetation

Myrmica sabuleti A Red Ant Countess Cutting SU145421 30/06/2017 Ground search Under stone, chalk cutting

Myrmica sabuleti A Red Ant Countess Cutting SU145421 26/07/2017 Pitfall trap Chalk cutting with patchy vegetation

Myrmecina graminicola Woodlouse Ant Countess Cutting SU145421 30/06/2017 Ground search Moss and litter, chalk cutting

Myrmecina graminicola Woodlouse Ant Countess Cutting SU145421 26/07/2017 Pitfall trap Chalk cutting with patchy vegetation

Agelena labyrinthica An Agelenid Spider Countess Cutting SU145421 30/06/2017 Spot search Web in tussock, chalk cutting VU./N Pheocedus braccatus A Gnaphosid Spider R. Countess Cutting SU145421 30/06/2017 Ground search Moss and litter, chalk cutting Trachyzelotes pedestris A Gnaphosid Spider Countess Cutting SU145421 30/06/2017 Ground search Moss and litter, chalk cutting

Philoscia muscorum Striped Woodlouse Countess Cutting SU145421 26/07/2017 Pitfall trap Chalk cutting with patchy vegetation

Oniscus asellus Common Shiny Woodlouse Countess Cutting SU145421 26/07/2017 Pitfall trap Chalk cutting with patchy vegetation

Armadillidium vulgare Common Pill Woodlouse Countess Cutting SU145421 30/06/2017 Ground search Under stones, chalk cutting

Armadillidium vulgare Common Pill Woodlouse Countess Cutting SU145421 26/07/2017 Pitfall trap Chalk cutting with patchy vegetation

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