The Discovery and Naming of Papaver Orientale Sl (Papaveraceae)
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Phylogeography of a Tertiary Relict Plant, Meconopsis Cambrica (Papaveraceae), Implies the Existence of Northern Refugia for a Temperate Herb
Article (refereed) - postprint Valtueña, Francisco J.; Preston, Chris D.; Kadereit, Joachim W. 2012 Phylogeography of a Tertiary relict plant, Meconopsis cambrica (Papaveraceae), implies the existence of northern refugia for a temperate herb. Molecular Ecology, 21 (6). 1423-1437. 10.1111/j.1365- 294X.2012.05473.x Copyright © 2012 Blackwell Publishing Ltd. This version available http://nora.nerc.ac.uk/17105/ NERC has developed NORA to enable users to access research outputs wholly or partially funded by NERC. Copyright and other rights for material on this site are retained by the rights owners. Users should read the terms and conditions of use of this material at http://nora.nerc.ac.uk/policies.html#access This document is the author’s final manuscript version of the journal article, incorporating any revisions agreed during the peer review process. Some differences between this and the publisher’s version remain. You are advised to consult the publisher’s version if you wish to cite from this article. The definitive version is available at http://onlinelibrary.wiley.com Contact CEH NORA team at [email protected] The NERC and CEH trademarks and logos (‘the Trademarks’) are registered trademarks of NERC in the UK and other countries, and may not be used without the prior written consent of the Trademark owner. 1 Phylogeography of a Tertiary relict plant, Meconopsis cambrica 2 (Papaveraceae), implies the existence of northern refugia for a 3 temperate herb 4 Francisco J. Valtueña*†, Chris D. Preston‡ and Joachim W. Kadereit† 5 *Área de Botánica, Facultad deCiencias, Universidad de Extremadura, Avda. de Elvas, s.n. -
Plant Taxonomy Table
COMMON AND LATIN NAMES OF IMPORTANT PLANT TAXA LATIN NAME* COMMON NAME Abies Fir Acer Maple Acer negundo Box elder Aesculus Buckeye; Horse Chestnut Alnus Alder Ambrosia Ragweed Apiaceae [Umbelliferae] Carrot or parsley family Artemisia Sagebrush; sage; wormwood Asteraceae [Compositae] Aster or Sunflower Family Betula Birch Boraginaceae Borage family Brassicaceae [Cruciferae} Mustard family Caryophyllaceae Pinks Castanea Chestnut Compositae (Asteraceae) Aster or Sunflower family Cornus Dogwood Corylus Filbert; hazelnut Cruciferae (Brassicaceae) Mustard family Cupressaceae Junipers, cypresses, "cedars", others Cyperaceae Sedge family Ericaceae Heath family Fabaceae [Leguminosae] Pea family Fagus Beech Fraxinus Ash Gramineae (Poaceae) Grass family Juglans Walnut; butternut Labiatae (Lamiaceae) Mint family Larix Larch; tamarack Leguminosae (Fabaceae) Pea family Liliaceae Lily family Liriodendron Tulip tree or yellow poplar Nuphar Water lily Onagraceae Evening primrose family Papaveraceae Poppy family Picea Spruce Pinus Pine Plantago Plantain Poaceae [Gramineae] Grass family Polemonium Jacob's ladder Polygonaceae Buckwheat family Populus Poplar; cottonwood; aspen Potamogeton Pondweed Primulaceae Primrose family Quercus Oak Ranunculaceae Buttercup family Rosaceae Rose family Rhus sumac, incl. poison ivy, etc. Salix Willow Saxifragaceae Saxifrage family Scrophulariaceae Snapdragon family Sparganium Bur reed Thalictrum Meadow rue Tilia Linden or basswood Tsuga Hemlock Typha Cattail Ulmus Elm Umbelliferae (Apiaceae) Carrot or parsley family * Names of genera are always italicized; family names are given in Roman characters. All proper plant family name ends in -aceae; family names above that don't have this ending are old names, and the proper modern name is included in parentheses. . -
The Developmental and Genetic Bases of Apetaly in Bocconia Frutescens
Arango‑Ocampo et al. EvoDevo (2016) 7:16 DOI 10.1186/s13227-016-0054-6 EvoDevo RESEARCH Open Access The developmental and genetic bases of apetaly in Bocconia frutescens (Chelidonieae: Papaveraceae) Cristina Arango‑Ocampo1, Favio González2, Juan Fernando Alzate3 and Natalia Pabón‑Mora1* Abstract Background: Bocconia and Macleaya are the only genera of the poppy family (Papaveraceae) lacking petals; how‑ ever, the developmental and genetic processes underlying such evolutionary shift have not yet been studied. Results: We studied floral development in two species of petal-less poppies Bocconia frutescens and Macleaya cordata as well as in the closely related petal-bearing Stylophorum diphyllum. We generated a floral transcriptome of B. frutescens to identify MADS-box ABCE floral organ identity genes expressed during early floral development. We performed phylogenetic analyses of these genes across Ranunculales as well as RT-PCR and qRT-PCR to assess loci- specific expression patterns. We found that petal-to-stamen homeosis in petal-less poppies occurs through distinct developmental pathways. Transcriptomic analyses of B. frutescens floral buds showed that homologs of all MADS-box genes are expressed except for the APETALA3-3 ortholog. Species-specific duplications of other ABCE genes inB. frute- scens have resulted in functional copies with expanded expression patterns than those predicted by the model. Conclusions: Petal loss in B. frutescens is likely associated with the lack of expression of AP3-3 and an expanded expression of AGAMOUS. The genetic basis of petal identity is conserved in Ranunculaceae and Papaveraceae although they have different number of AP3 paralogs and exhibit dissimilar floral groundplans. -
A Systematic Review on Main Chemical Constituents of Papaver Bracteatum
Journal of Medicinal Plants A Systematic Review on Main Chemical Constituents of Papaver bracteatum Soleymankhani M (Ph.D. student), Khalighi-Sigaroodi F (Ph.D.)*, Hajiaghaee R (Ph.D.), Naghdi Badi H (Ph.D.), Mehrafarin A (Ph.D.), Ghorbani Nohooji M (Ph.D.) Medicinal Plants Research Center, Institute of Medicinal Plants, ACECR, Karaj, Iran * Corresponding author: Medicinal Plants Research Center, Institute of Medicinal Plants, ACECR, P.O.Box: 33651/66571, Karaj, Iran Tel: +98 - 26 - 34764010-9, Fax: +98 - 26-34764021 E-mail: [email protected] Received: 17 April 2013 Accepted: 12 Oct. 2014 Abstract Papaver bracteatum Lindly (Papaveraceae) is an endemic species of Iran which has economic importance in drug industries. The main alkaloid of the plant is thebaine which is used as a precursor of the semi-synthetic and synthetic compounds including codeine and naloxone, respectively. This systematic review focuses on main component of Papaver bracteatum and methods used to determine thebaine. All studies which assessed the potential effect of the whole plant or its extract on clinical or preclinical studies were reviewed. In addition, methods for determination of the main components, especially thebaine, which have been published from 1948 to March 2013, were included. Exclusion criteria were agricultural studies that did not assess. This study has listed alkaloids identified in P. bracteatum which reported since 1948 to 2013. Also, the biological activities of main compounds of Papaver bracteatum including thebaine, isothebaine, (-)-nuciferine have been reviewed. As thebaine has many medicinal and industrial values, determination methods of thebaine in P. bracteatum were summarized. The methods have being used for determination of thebaine include chromatographic (HPLC, GC and TLC) and non chromatographic methods. -
Functional Analyses of Genetic Pathways Controlling Petal Specification in Poppy Sinéad Drea1, Lena C
RESEARCH ARTICLE 4157 Development 134, 4157-4166 (2007) doi:10.1242/dev.013136 Functional analyses of genetic pathways controlling petal specification in poppy Sinéad Drea1, Lena C. Hileman1,*, Gemma de Martino1 and Vivian F. Irish1,2,† MADS-box genes are crucial regulators of floral development, yet how their functions have evolved to control different aspects of floral patterning is unclear. To understand the extent to which MADS-box gene functions are conserved or have diversified in different angiosperm lineages, we have exploited the capability for functional analyses in a new model system, Papaver somniferum (opium poppy). P. somniferum is a member of the order Ranunculales, and so represents a clade that is evolutionarily distant from those containing traditional model systems such as Arabidopsis, Petunia, maize or rice. We have identified and characterized the roles of several candidate MADS-box genes in petal specification in poppy. In Arabidopsis, the APETALA3 (AP3) MADS-box gene is required for both petal and stamen identity specification. By contrast, we show that the AP3 lineage has undergone gene duplication and subfunctionalization in poppy, with one gene copy required for petal development and the other responsible for stamen development. These differences in gene function are due to differences both in expression patterns and co- factor interactions. Furthermore, the genetic hierarchy controlling petal development in poppy has diverged as compared with that of Arabidopsis. As these are the first functional analyses of AP3 genes in this evolutionarily divergent clade, our results provide new information on the similarities and differences in petal developmental programs across angiosperms. Based on these observations, we discuss a model for how the petal developmental program has evolved. -
In Vitro Regeneration of Persian Poppy (Papaver Bracteatum)
Biologia 65/4: 647—652, 2010 Section Cellular and Molecular Biology DOI: 10.2478/s11756-010-0079-6 In vitro regeneration of Persian poppy (Papaver bracteatum) Sara Rostampour1,2,HalehHashemi Sohi1*&AliDehestani3 1Department of Plant Biotechnology, National Institute of Genetic Engineering and Biotechnology, P.O. Box 14155–6343 Tehran, Iran; e-mail: [email protected] 2Department of Agronomy and Plant Breeding, University of Zabol, Zabol, Iran 3Department of Plant Breeding, Faculty of Agronomic Sciences, Sari Agricultural Sciences and Natural Resources Univer- sity, Sari, Iran Abstract: Persian poppy (Papaver bracteatum Lindl.) is an important commercial source of medicinal opiates and related compounds. In this research, calli were induced from seeds, roots, cotyledons and hypocotyls of P. bracteatum at a high efficiency. The optimized callus induction media consisted of the Murashige and Skoog (MS) basic media supplemented with 1.0 mg/L 2, 4-dichlorophenoxyacetic acid (2,4-D), 0.1 mg/L kinetin and 15 mg/L ascorbic acid. The concentrations of 2,4-D and ascorbic acid were found critical to callus induction and proliferation. Subsequent subcultures resulted in excellent callus proliferation. Ascorbic acid at concentration 15 mg/L increased the callus proliferation significantly. Maximum callus growth was achieved when the explants were incubated at 25 ◦C. MS salts at full strength were found inhibitory for callus induction, while ľ MS salts were found to favor callus induction. Shoot regeneration of calli in vitro wasachievedonľ MS medium containing 0.5 mg/L benzylamine purine and 1.0 mg/L naphthalene acetic acid. Analysis of alkaloid extracts from Persian poppy tissues by high-performance liquid chromatography showed that thebaine accumulated in the tissues of plants. -
The Yearbook of Agriculture • 1961 ^
87TH CONGRESS, IST SESSION, HOUSE DOCUMENT NO. 29 THE YEARBOOK OF AGRICULTURE • 1961 ^ THE UNITED STATES DEPARTMENT OF AGRICULTURE Washington, D.G. SEEDS The Yearbook of Agriculture 1961 ¿^ f i Í THE UNITED STATES GOVERNMENT PRINTING OFFICE FOR SALE BY THE SUPERINTENDENT OF DOCUMENTS, WASHINGTON 25, D.G., PRICE $2 FOREWORD ORVILLE L. FREEMAN Secretary of Agriculture GOOD SEEDS ARE both a symbol and a foundation of the good Ufe our people have gained. A baisic factor in our realization of mankind's most sought goal, agricultural abundance, good seeds can be a means of our bringing about an Age of Plenty and an Age of Peace and Free- dom. We can use our good seeds to help end hunger and fear for the less fortunate half of the human family. So used, our seeds can be more meaningful to a hungry world than can the rocket that first carries man to the moon. This Yearbook of Agriculture seeks to provide a new and improved basis for understanding the complex order of Nature's forces so that man can better shape them in a positive and creative fashion. Seeds are ever a positive and creative force. Seeds are the germ of life, a beginning and an end, the fruit of yesterday's harvest and the promise of tomorrow's. Without an ample store of seeds there can be no national treasure, or no future for a Nation. Finding and developing better seeds is the oldest continuous service our Federal Government has rendered to our farmers—indeed, to all our people. -
Mohn (Papaver Somniferum
Mohn (Papaver somniferum L.) Synonyme (Namen dieser Pflanze in allen 76 Sprachen anzeigen) pharmazeutisch Semen Papaveris Albanisch Lulëkuqe Μήκων Altgriechisch Mekon ፓፓ Amharisch Papi , Arabisch , Khashkhash, Abu an-num, Abu al-num, Abu an-noom, Abu al-noom Մեկոն, Մեկոնի Կուտ Armenisch Mekon, Megon; Mekoni Kut, Megoni Good (Samen) Assamesisch Xaş-xaş Azeri Хаш-хаш Baskisch Lobelarr Bengali Градински мак, Опиев мак, Маково семе Bulgarisch Gradinski mak, Opiev mak; Makovo seme (Samen) Burmesisch Chinesisch 櫻粟殼 [yìng suhk hohk] (Kantonesisch) Ying suhk hohk Chinesisch 櫻粟殼 [yīng sù qiào], 罂粟 [yīng sù] (Mandarin) Ying su qiao, Ying su Dänisch Opiumvalmue (Pflanze); Birkes, Valmue-frø (Samen) Deutsch Schlafmohn, Gartenmohn, Ölmohn, Opiummohn, , Dhivehi Afihun, Kaskasaa Englisch Poppy, Opium poppy, Garden poppy Esperanto Papavo, Papavosemo Estnisch Magun, Unimagun, Moon Farsi Khash-khash, Shagheyegh Finnisch Unikko, Oopiumiunikko Französisch Pavot des jardins, Pavot somnifère, Pavot à opium Gälisch Codalion, Paipin Galizisch Mapoula, Sementes de Mapoula, Adormideira, Durmideira ყაყაჩო, ყაყაჩოს თესლი, ხოშხოში Georgisch Khoshkhoshi, Q’aq’acho, Xoshxoshi, Qaqacho; Q’aq’achos tesli, Qaqachos tesli (Samen) Παπαρούνα, Αθιόνι Griechisch Paparouna, Afioni Gujarati Hebräisch Pereg Hindi Irisch Poipín Isländisch Valmúafræ, Birki Italienisch Papavero, Papavero sonnifero 芥子, 罌粟 けし Japanisch ケシ, ポピー Keshi, Papi , Jiddisch Mondl, Mon Kannada Көкнәр Kasachisch Köknär Kashmiri Khash-khash Katalanisch Cascall, Herba dormidora 아편, 포피, 양귀비 Koreanisch Apyeon, Apyon, -
44 * Papaveraceae 1
44 * PAPAVERACEAE 1 Dennis I Morris 2 Annual or perennial herbs, rarely shrubs, with latex generally present in tubes or sacs throughout the plants. Leaves alternate, exstipulate, entire or more often deeply lobed. Flowers often showy, solitary at the ends of the main and lateral branches, bisexual, actinomorphic, receptacle hypogynous or perigynous. Sepals 2–3(4), free or joined, caducous. Petals (0–)4–6(–12), free, imbricate and often crumpled in the bud. Stamens usually numerous, whorled. Carpels 2-many, joined, usually unilocular, with parietal placentae which project towards the centre and sometimes divide the ovary into several chambers, ovules numerous. Fruit usually a capsule opening by valves or pores. Seeds small with crested or small raphe or with aril, with endosperm. A family of about 25 genera and 200 species; cosmopolitan with the majority of species found in the temperate and subtropical regions of the northern hemisphere. 6 genera and 15 species naturalized in Australia; 4 genera and 9 species in Tasmania. Papaveraceae are placed in the Ranunculales. Fumariaceae (mostly temperate N Hemisphere, S Africa) and Pteridophyllaceae (Japan) are included in Papaveraceae by some authors: here they are retained as separate families (see Walsh & Norton 2007; Stevens 2007; & references cited therein). Synonymy: Eschscholziaceae. Key reference: Kiger (2007). External resources: accepted names with synonymy & distribution in Australia (APC); author & publication abbre- viations (IPNI); mapping (AVH, NVA); nomenclature (APNI, IPNI). 1. Fruit a globular or oblong capsule opening by pores just below the stigmas 2 1: Fruit a linear capsule opening lengthwise by valves 3 2. Stigmas joined to form a disk at the top of the ovary; style absent 1 Papaver 2: Stigmas on spreading branches borne on a short style 2 Argemone 3. -
Alkaloid Profile in Relation to Different Developmental Stages of Papaver Somniferum L
ZOBODAT - www.zobodat.at Zoologisch-Botanische Datenbank/Zoological-Botanical Database Digitale Literatur/Digital Literature Zeitschrift/Journal: Phyton, Annales Rei Botanicae, Horn Jahr/Year: 2001 Band/Volume: 41_1 Autor(en)/Author(s): Shukla Sudhir, Singh S. P. Artikel/Article: Alkaloid Profile in Relation to Different Developmental Stages of Papaver somniferum L. 87-96 ©Verlag Ferdinand Berger & Söhne Ges.m.b.H., Horn, Austria, download unter www.biologiezentrum.at Phyton (Horn, Austria) Vol. 41 Fasc. 1 87-96 29. 6. 2001 Alkaloid Profile in Relation to Different Developmental Stages of Papaver somniferum L. By S. SHUKLA*)*) and S. P. SINGH*) With 2 figures Received January 17, 2000 Accepted August 28, 2000 Key words: Alkaloid, P. somniferum, morphine, codeine, thebaine, noscapine, papaverine. Summary SHUKLA S. & SINGH S. P. 2001. Alkaloid profile in relation to different develop- mental stages of Papaver somniferum L. - Phyton (Horn, Austria) 41 (1): 87-96, 2 figures. - English with German summary. The alkaloids variation and its synthesis were studied in two varieties (NBRI-1, NBRI-2) of opium poppy {Papaver somniferum L.) on fresh weight basis of different plant parts at different growth periods. In cotyledon stage (3-4 days after germina- tion) only morphine was present. In roots of two leave stage, thebaine was observed beside morphine. At bud initiation stage morphine, codeine and thebaine were pre- sent during 1994-95 but in 1995-96 thebaine was absent. During bud dropping stage (pendulous bud) the sepals, petals and anthers had morphine. When pendulous bud straightened before flowering it has morphine, codeine and thebaine in all parts in- cluding ovary. -
Opium Poppy' Cultivation and Heroin Processing in Southeast Asia
If you have issues viewing or accessing this file contact us at NCJRS.gov. u.s. Department of Justice Drug Enforcement Administration Office ofIntelligence September 1992 Opium Poppy' Cultivation and Heroin Processing in Southeast Asia . eC 7 7 s DEA- 92004 141189 U.S. Department of Justice National Institute of Justice This document has been reproduclld exactly as received from the person or organization originating It. Points of view or opinions stated In this document are those of the authors and do not necessarily represent the official position or policies of the National Institute of Justice. Permission to reproduce this : iLl i r material has been 9FUi3i~c Doroain/Drug Enforcerrent Adrm. U • S. Department of Justice to the National Criminal Justice Reference Service (NCJRS). Further reproduOllol1 outside of the NCJRS system requires permission of the ~ owner. U.S. Department of Justice Drug Enforcement Administration Office of Intelligence Washington, DC 20537 (202) 307-8100 September 1992 ADMINISTRATOR·S MESSAGE Opium Poppy Cultivation and Heroin Processing in Southeast Asia reviews in detail several complex natural and chemical procedures linked with heroin production. It further explains how the opium poppy plant of Southeast Asia is grown and harvested, how the poppy's opium is chemically converted into morphine, and fmally. how that morphine is then refmed into "China White" heroin by Southeast Asian "cooks" and "chemists." The report will interest a wide audience: those studying "source country" issues, those determining opium poppy crop estimates in mainland Southeast Asia, and those monitoring and controlling the intemational import and export of chemicals. Opium Poppy Cultivation and Heroin Processing in Southeast Asia also presents the technical relationship between opium poppy cultivation in the Golden Triangle (the moun tainous region where Burma, Laos, and Thailand share common borders) and heroin production in the laboratories of Southeast Asia. -
Early Diverging Eudicots
1/31/20 Berberidaceae - barberry family • widespread in temperate regions of Northern hemisphere - ArctoTertiary relict distribution • 3 native genera to Wisconsin + Berberis • small shrubs (Berberis) or herbs (rest of family). CA 3+3 CO 3+3 A 12-18 G 1 [monocarpic] Early Berberis thunbergii Diverging Japanese barberry Eudicots 1 2 Berberidaceae - barberry family Berberidaceae - barberry family Berberis thunbergii Japanese barberry • Puccinia graminis Fruits = berry Berberis vulgaris Berberis vulgaris Common barberry Common barberry • alternate host of the stem rust of wheat • alternate host of the stem rust of wheat • programs to irradicate plant in midwest • programs to irradicate plant in midwest 3 4 1 1/31/20 Berberidaceae - barberry family Berberidaceae - barberry family Caulophyllum thalictroides - Blue cohosh Jeffersonia diphylla - twinleaf • leaves 3X compound • Special concern species in Wisconsin. One species in • developing ovules break ovary wall so are Eastern North America and one species in Eastern Asia. naked seeds with fleshy blue seed coats • medicinally important for North American • Only member of the family that is 4-merous rather than 3- native Americans merous 5 6 Berberidaceae - barberry family Berberidaceae - barberry family Forms large colonies of deeply lobed, May-apple and other members of the peltate leaves whose umbrella like family are attacked by the same family aspect is emphasized as they emerge of rusts - Pucciniaceae. from the ground, slits in anthers Podophyllum peltatum Podophyllum peltatum May-apple