Annotated Checklist of the (Coleoptera) from Louisiana Author(s): Christopher E. Carlton, Ted C. MacRae, Alexey K. Tishechkin, Victoria L.Bayless and Wood Johnson Source: The Coleopterists Bulletin, 72(2):351-367. Published By: The Coleopterists Society https://doi.org/10.1649/0010-065X-72.2.351 URL: http://www.bioone.org/doi/full/10.1649/0010-065X-72.2.351

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ANNOTATED CHECKLIST OF THE BUPRESTIDAE (COLEOPTERA) FROM LOUISIANA

CHRISTOPHER E. CARLTON Louisiana State Museum Louisiana State University Agricultural Center Baton Rouge, LA 70808, USA [email protected]

TED C. MACRAE c/o Monsanto Company 700 Chesterfield Parkway West Chesterfield, MO 63017-1732, USA

ALEXEY K. TISHECHKIN Plant Pest Diagnostics Center, CDFA 3294 Meadowview Road Sacramento, CA 95832, USA

VICTORIA L. BAYLESS Louisiana State Arthropod Museum Louisiana State University Agricultural Center Baton Rouge, LA 70808, USA

AND

WOOD JOHNSON USDA Forest Service, Forest Health Protection 2500 Shreveport Hwy, Pineville, LA 71360, USA

ABSTRACT We report 110 species of Buprestidae from the state of Louisiana, 103 species based on 5,047 specimens examined and seven from previous literature records. Fifteen new state records are reported, including the first published records of Agrilus planipennis Fairmaire, 1888 (emerald ash borer), which is now established in the northern parishes of the state. Two new state records are based on specimens taken during biomonitoring surveys using the buprestid specialist wasp Cerceris fumipennis (Say, 1837). Potential larval and host plants of buprestid species in Louisiana and collection date ranges of specimens examined are provided.

Key Words: metallic wood borers, emerald ash borer, taxonomic checklist, forest pests, Cerceris fumipennis

DOI.org/10.1649/0010-065X-72.2.351

The establishment and spread of Agrilus plani- particularly on native buprestids and buprestid pennis Fairmaire, 1888 (emerald ash borer, EAB) predators and parasitoids. To that end, we updated has generated a surge of interest in collecting identifications and created a database of buprestid and surveying buprestid across the poten- holdings in the Louisiana State Arthropod tial North American range of EAB. This inva- Museum (Louisiana State University, Baton Rouge, sive pest of ash, Fraxinus spp. (Oleaceae), is Louisiana, USA) (LSAM). We also obtained Lou- spreading throughout eastern North America and isiana records from entomology collections in other is now established as far south and west as southeastern US states and the US National Mu- northern Louisiana and northeast Texas (Harrison seum. We included records of buprestids collected County, texasforestservice.tamu.edu/content/article. from burrow aggregations of Cerceris fumipennis aspx?id524246, accessed 4 October 2017). (Say, 1837) (Hymenoptera: Crabronidae) during Establishing effective survey and monitoring surveys at several localities in central and south- protocols and determining the pre-EAB diversity central Louisiana (results reported by Johnson et al. and distribution of buprestids are critical in 2015). Cerceris fumipennis is a specialized predator assessing impacts of EAB on native flora and fauna, of buprestid beetles, which has been widely used

351 352 THE COLEOPTERISTS BULLETIN 72(2), 2018 as a biomonitoring tool for buprestids in eastern cyberfloralouisiana.com, accessed 12 July 2016) North America (Marshall et al. 2005). Historical and/or the US Department of Agriculture Plants museum records and recent C. fumipennis buprestid Database (plants.usda.gov/java/, accessed 13 July records are compiled here into a checklist of 2016). buprestid beetle occurrence, abundance, and dis- tributions in Louisiana. We included notes on po- RESULTS tential larval and adult host plants in Louisiana, as well as comments on collection date ranges and This checklist includes records based on 5,047 collecting events. adult buprestid specimens. Records from FSCA, MEM, and TAMU represented 63, 102, and 306 specimens, respectively. Twenty-three species are MATERIAL AND METHODS also represented by Louisiana specimens in the This work is based on authoritatively identified USNM, none of which are unique records. Seven specimens housed in the LSAM, Mississippi En- additional Louisiana “state only” records are cited tomological Museum (MEM, Mississippi State, from Nelson et al. (2008) and included literature MS), US National Museum of Natural History, citations. Seventeen species are represented by Smithsonian Institution (USNM, Washington, DC), single specimens. Texas A&M Collection (TAMU, College With the inclusion of previously published re- Station, TX), Florida State Collection of cords, 110 species are recorded for Louisiana, in- (FSCA, Gainesville, FL), and the personal collec- cluding 15 reported here as new state records, two tion of TCM. Most unidentified or questionably based on specimens captured from C. fumipennis identified Louisiana specimens from the LSAM and wasps and one represented by EAB. Species MEM were sent to one of the authors (TCM) for represented in surveys using C. fumipennis identification or confirmation. Specimens within the are highlighted in the checklist comments by genera Eschscholtz and “C. fumipennis prey”. Dejean were identified by R. L. Westcott (Oregon Department of Agriculture, Salem). TAMU speci- mens were identified by the former curator, Edward CHECKLIST OF THE BUPRESTIDAE Riley. Records based on USNM specimens were OF LOUISIANA compiled by AKT. Additional identifications were provided by Crystal Maier, Field Museum of Nat- Family Buprestidae ural History, Chicago, IL and confirmed by TCM. Subfamily LSAM specimen data and those obtained during Tribe Polycestini C. fumipennis monitoring were entered into the 1. Polycesta (Tularensia) elata LeConte, 1858 LSAM’s Specify 6 database. When georeference Parish records (n 5 1): Concordia Parish (LSAM). coordinates were not provided by collectors (the Potential larval host in Louisiana: Platanus vast majority of pre-2000 records), they were ap- occidentalis L. proximated retroactively using the Geolocate fea- Collection date range: 24 May–7 June (trap ture in Specify for place names or through searches service interval). using Google Earth or Google Maps. Georeference Comments: Collected using canopy Malaise trap. sources are indicated in Specify to allow users to Tribe Ptosimini interpret levels of precision. A downloaded file of 2. Ptosima gibbicollis (Say, 1823) all specimen data is available on request. Parish records (n 5 4): Union and West Collecting methods and other relevant data Feliciana Parishes (LSAM). recorded on labels of specimens examined are Potential larval host in Louisiana: Cercis noted in comments. Collection date ranges are canadensis L. Adults reported also on based on earliest and latest dates on labels, with Prunus serotina Ehrh. and Sassafras albi- high concentrations of monthly occurrences noted. dum (Nutt.) Nees. Additional potential host plants in Louisiana, both Collection date range: 18 March–8 April. larval and adult, were deduced by comparing hosts Comments: Collection methods unspecified. reportedbyNelsonet al. (2008) (and other pre- viously reported hosts omitted from that work, as 3. Ptosima idolynae Frost, 1923 compiled by TCM and R. L. Westcott) and two Parish records (n 5 2): East Baton Rouge later works (Hansen et al. 2012; MacRae and Parish (LSAM). Basham 2013) with known distributions of plant Potential larval host in Louisiana: Crataegus species in Louisiana. Occurrences of questionable sp. Adults reported also on Gleditsia tri- host species were checked by consulting records acanthos L. in the Cyberflora of Louisiana database (www. Collection date range: 6 May. THE COLEOPTERISTS BULLETIN 72(2), 2018 353

Comments: New state record: East Baton Tangipahoa, Vernon, Webster, and Winn Rouge Par., Highland Rd. Park, 6 May Parishes (LSAM, MEM, TAMU, USNM). 2011, collected as C. fumipennis prey, Potential larval hosts in Louisiana: Acer sp., 30.3491°N, 91.0741°W, W. Virgets (n 5 2). Crataegus sp., Gleditsia triacanthos, and Taxodium distichum (L.) Rich. Adults re- Tribe Haplostethini ported also on Acer saccharinum L., Dio- 4. Mastogenius crenulatus Knull, 1934 spyros virginiana L. and Quercus alba. Parish records (n 5 46): Ascension, Concordia, Collection date range: 12 March–24 August, East Baton Rouge, Morehouse, Plaque- mostly during June (n 5 17). mines, Natchitoches, St. Tammany, West Comments: Collected by hand ‘on flowers,’ Baton Rouge, and West Feliciana Parishes using Malaise traps, and as C. fumipennis prey. (FSCA, LSAM, TAMU). 8. Acmaeodera (s. str.) texana LeConte, 1860 Potential larval hosts in Louisiana: Carya Parish records (n 5 1): Claiborne Parish (TAMU). illinoinensis (Wangenh.) K. Koch, Cercis Potential larval hosts in Louisiana: Unknown. canadensis, Juglans nigra L., Quercus alba Collection date range: 6 July. L., and Quercus phellos L. Adults reported Comments: Collection method unspecified. also on Acer saccharum Marsh., Salix sp., and Quercus falcata Michx. 9. Acmaeodera (s. str.) tubulus (Fabricius, 1801) Collection date range: 12 March–3 May, six Parish records (n 5 118): Acadia, Avoyelles, reared specimens emerged February–May. Bossier, Catahoula, DeSoto, East Baton Comments: Collected at light, by flight in- Rouge, Grant, Iberville, Jackson, Lafayette, tercept traps, reared from ‘miscellaneous Natchitoches, St. Landry, St. Tammany, dead limbs’, from unspecified coarse, Vernon, Washington, Webster, and Winn woody debris, and by sweeping. Parishes (LSAM, MEM, TAMU, USNM). Potential larval hosts in Louisiana: Betula 5. Mastogenius subcyaneus (LeConte, 1860) 5 nigra L., Carya glabra (Mill.) Sweet, Carya Parish records (n 2): West Feliciana Parish illinoinensis, Carya ovata (Mill.) K. Koch, (LSAM). Carya tomentosa (Lam.) Nutt., Celtis occi- Potential larval hosts in Louisiana: Carya dentalis L., Cercis canadensis, Crataegus illinoinensis, Cercis canadensis, Ostrya viridis L., Gleditsia triacanthos, Juglans virginiana (Mill.) K. Koch, and Quercus sp. nigra, Ostrya virginiana, Quercus alba, Salix (prob. Q. palustris Muenchh.). Adults re- sp., Ulmus crassifolia Nutt., and Ulmus rubra ported also on Cornus sp. – Muhl. Adults reported also on Juniperus Collection date range: 6 April 8 May. virginiana L., Quercus velutina Lam., Sapin- Comments: Collected using pitfall trap and dus saponaria L., and Taxodium distichum. method unspecified. Collection date range: 8 March–9 September, mostly during April (n 5 84) and May Tribe Acmaeoderini (n 5 25). 6. Acmaeodera (s. str.) ornata (Fabricius, 1775) Comments: Collected by hand, beating, using Parish records (n 5 2): Bossier and Caddo flight intercept, Lindgren funnel, Malaise, Parishes (LSAM, MEM). and sticky traps, sweeping ‘Aster sp.,’ and Potential larval hosts in Louisiana: Unknown. as C. fumipennis prey. Adults reported on Persea borbonia (L.) Spreng. and Quercus alba. Subfamily Collection date range: 17–30 April. Tribe Poecilonotini Comments: New state record: Bossier Parish, 10. cyanipes (Say, 1823) Barksdale Air Force Base, 30 Apr. 1996, on Parish records (n 5 4): Bossier and East Baton Coreopsis lanceolata in calcareous prairie, Rouge Parish (LSAM, MEM). 32.4913°N, 93.5152°W, D. M. Pollock (n 5 Potential larval hosts in Louisiana: Populus 1); Caddo Parish, 5 mi N Vivian on LA-1, deltoides Marsh., Robinia pseudoacacia L., 17 Apr.1969, on flower, 32.9616°N, and Salix nigra Marsh. 93.9935°W, M. S. Strother (n 5 1). Collection date range: 13–22 May. Comments: Collected by beating and as C. 7. Acmaeodera (s. str.) pulchella (Herbst, 1801) fumipennis prey. Parish records (n 5 50): Acadia, Bossier, Caddo, Cameron, Catahoula, DeSoto, East 11. Poecilonota thureura (Say, 1832) Baton Rouge, Jefferson, Jefferson Davis, Parish records (n 5 43): East Baton Rouge Natchitoches, Red River, St. Tammany, Parish (LSAM). 354 THE COLEOPTERISTS BULLETIN 72(2), 2018

Potential larval host in Louisiana: Salix nigra. Potential larval hosts in Louisiana: Carya sp. Adults reported also on Cercis canadensis. and Quercus alba. Adults reported also on Collection date range: 6 May–2 July, mostly Pinus sp. and Taxodium distichum. during May (n 5 39). Collection date range: 16 April–12 November. Comments: Collected only as C. fumipennis prey. Comments: Collecting methods unspecified. Tribe Chrysochroini 17. caudata LeConte, 1860 12. georgiana (LeConte, 1857) Literature record only. Parish records (n 5 6): St. Bernard, St. Tammany, Potential larval hosts in Louisiana: Alnus sp. and Washington Parishes (LSAM, USNM). and Betula nigra. Adults reported also on Potential larval hosts in Louisiana: Pinus Crataegus sp. and Prunus persica (L.) echinata Mill., Pinus palustris Mill., and Batsch. Pinus taeda L. Comments: This is mainly a northern, trans- Collection date range: 15 February–5 August. continental North American species with Comments: Collected at UVand mercury vapor Louisiana as the southernmost record lights. (Nelson 1975). 13. Chalcophora virginiensis (Drury, 1770) 18. Dicerca juncea Knull, 1958 Parish records (n 5 71): Assumption, Bienville, Parish records (n 5 11): Orleans and St. Caddo, Calcasieu, DeSoto, East Baton Tammany Parishes (LSAM, USNM). Rouge, East Feliciana, Franklin, Grant, Potential larval hosts in Louisiana: Unknown. Iberville, La Salle, Livingston, Natch- Adults reported on Acer sp., Cyrilla race- itoches, Orleans, Rapides, St. Tammany, miflora L., Prunus persica, and Quercus sp. Tangipahoa, Vernon, and Washington Collection date range: 18 April–8 September. Parishes (LSAM, MEM, TAMU). Comments: Collected using Malaise traps. Potential larval hosts in Louisiana: Pinus echinata, P. palustris, and P. taeda. 19. Dicerca lepida LeConte, 1857 Collection date range: 9 March–4 November, Parish records (n 5 2): Franklin and Natch- with no apparent pattern between dates. itoches Parishes (FSCA, LSAM). Comments: Collected at light, by hands on Potential larval hosts in Louisiana: Crataegus ‘loblolly pine,’ flight intercept, Lindgren fun- coccinea L. and Ostrya virginiana. Adults nel and pitfall traps, and rearing from ‘cypress.’ reported also on Quercus sp. and Ulmus rubra. Collection date range: 16 April-17 May. 14. campestris (Say, 1823) Comments: Collected using canopy Malaise trap. Parish records (n 5 9): Acadia, Franklin, Grant, Natchitoches, and Winn Parishes and 20. Dicerca lurida (Fabricius, 1775) ‘Louisiana’ (LSAM, USNM). Parish records (n 5 304): Avoyelles, Caddo, Potential larval hosts in Louisiana: Acer sac- Concordia, DeSoto, East Baton Rouge, charum, Fagus grandifolia Ehrh., Platanus Evangeline, Franklin, Grant, Livingston, occidentalis, Salix nigra, and Tilia ameri- Madison, Natchitoches, Orleans, Plaque- cana L. Adults reported also on Lirioden- mines, St. Landry, St. Martin, St. Tammany, dron sp. and Quercus sp. Tensas, Webster, and Winn Parishes Collection date range: 17 January–18 May. (LSAM, TAMU, USNM). Comments: Collected by hand, using canopy Potential larval hosts in Louisiana: Carpinus Malaise traps, and as C. fumipennis caroliniana Walt., Carya cordiformis prey. (Wangenh.) K. Koch, Carya glabra, Carya 15. Texania langeri (Chevrolat, 1853) laciniosa (Michx. f.) G. Don, Prunus car- Literature record only. oliniana (Mill.) Ait., Salix exigua Nutt., and Potential larval hosts in Louisiana: Unknown. Tilia americana. Adults reported also on Comments: The type locality of this species is Carya ovata, Quercus alba, Quercus fal- “Greenville, pres de la Nouvelle Orleans.” cata, and Quercus velutina. Collection date range: 20 January–11 Novem- Greenville was a city formerly in Jefferson 5 5 Parish. The city was annexed by New ber, mostly in May (n 96) and June (n Orleans and became part of Orleans Parish. 99), five reared specimens emerged in June and September. Tribe Comments: Collected by hand, using canopy 16. Dicerca asperata (Laporte and Gory, 1837) Malaise and Malaise traps, reared from Parish records (n 5 4): Assumption, Grant, and ‘dead hickory limbs’ and ‘miscellaneous Iberville Parishes (LSAM). dead limbs’, and as C. fumipennis prey. THE COLEOPTERISTS BULLETIN 72(2), 2018 355

21. (Fabricius, 1781) Collection date range: 18 April–17 July, mostly Parish records (n 5 103): Bossier, Concordia, in May (n 5 91) and June (n 5 31), two East Baton Rouge, Franklin, Grant, Iber- reared specimens emerged in June and July. ville, Orleans, and West Feliciana Parishes Comments: Collected at light, by hand, in (FSCA, LSAM, MEM, TAMU, USNM). canopy and ground-based Malaise and Potential larval host in Louisiana: Diospyros flight intercept traps, and reared from virginiana. Adults reported also on Carya ‘hickory limbs’,alsoasC. fumipennis prey. sp. and Quercus sp. Collection date range: 30 April–9 October, 26. Actenodes auronotata Gory and Laporte, mostly in May (n 5 16) and June (n 5 53). 1837 5 Comments: Collected by hand and using canopy Parish records (n 1): Jefferson Parish (LSAM). Malaise traps, also as C. fumipennis prey. Potential larval hosts in Louisiana: Avicennia germinans (L.) L., Baccharis halimifolia L., 22. Dicerca punctulata (Schoenherr, 1817) and Taxodium distichum. Parish records (n 5 7): East Baton Rouge, Collection date: 16 July. Webster,andWinnParishes(LSAM, Comments: New state record: Jefferson Par- TAMU). ish, Grand Isle, 16 July 2004, blacklight, Potential larval hosts in Louisiana: Pinus echi- 29.2277°N, 90.0122°W, A. K. Tishechkin nata, Pinus elliottii Engelm., and Pinus taeda. (n 5 1). Collection date range: 15 May–17 June. Comments: Collection methods unspecified, 27. Actenodes calcaratus (Chevrolat, 1835) and as C. fumipennis prey. Parish records (n 5 1): Cameron Parish (LSAM). Potential larval hosts in Louisiana: Vachellia 23. Dicerca spreta (Gory, 1841) farnesiana (L.) Wight & Arn. Adults also Parish records (n 5 9): Allen, Orleans, and West reported also on Parkinsonia sp. and Pro- Feliciana Parishes (LSAM). sopis glandulosa Torr. Potential larval host in Louisiana: Nyssa syl- Collection date: 12 June. vatica Marsh. Comments: Collection method unspecified. Collection date range: 1 March–19 June, seven reared specimens emerged in June– 28. Actenodes davidi Nelson, 1979 September. Parish records (n 5 152): East Baton Rouge and Comments: Reared from unidentified wood. Winn Parishes (LSAM). Other collection methods unspecified. Potential larval host in Louisiana: Gleditsia triacanthos. 24. (Spinthoptera) drummondi (Laporte Collection date range: 6 May–12 July, mostly in and Gory, 1837) May (n 5 99). Literature record only. Comments: Collected only as C. fumipennis prey. Potential larval hosts in Louisiana: Unknown. Adults reported on Carya illinoinensis, Tribe Euphorbia sp., Gleditsia triacanthos, 29. (s. str.) consularis Gory, 1840 Gossypium sp., Salix sp., and Tamarix sp. Parish records (n 5 59): East Baton Rouge, Comments: Nelson (1986) recorded specimens Franklin, Grant, Livingston, and Winn collected in Rapides Parish. Parishes (LSAM). Potential larval host in Louisiana: Pinus taeda. Tribe Collection date range: 16 April–14 October, Tribe Actenodini mostly in May (n 5 41). 25. Actenodes acornis (Say, 1833) Comments: Collected at light and by beating, 5 Parish records (n 130): Bossier, Concordia, also as C. fumipennis prey. East Baton Rouge, St. Tammany, Union, West Feliciana, and Winn Parishes (FSCA, 30. Buprestis (s. str.) lineata Fabricius, 1775 LSAM, TAMU). Parish records (n 5 142): Bienville, Caddo, Potential larval hosts in Louisiana: Acer rubrum East Baton Rouge, East Feliciana, Grant, L., Acer saccharum, Betula nigra, Carya Lincoln, Livingston, Natchitoches, Oua- glabra, Carya illinoinensis, , chita, Rapides, St. Tammany, West Felic- Carya ovata, Cornus florida L., Fagus iana, and Winn Parishes (LSAM, MEM). grandifolia, Fagus sylvatica L., Quercus Potential larval hosts in Louisiana: Pinus laurifolia Michx., and Q. velutina. Adults echinata, Pinus palustris, and P. taeda. reported also on Cercis canadensis, Diospyros Collection date range: 9 April–28 October, virginiana,andQuercus prinoides Willd. mostly in May (n 5 73) and June (n 5 30). 356 THE COLEOPTERISTS BULLETIN 72(2), 2018

Comments: Collected at light and by hand; also Potential larval hosts in Louisiana: Pinus pal- using Lindgren funnel and pitfall traps, and ustris and Pinus echinata. as C. fumipennis prey. Collection date: 15 June. Comments: Collecting method unspecified. 31. Buprestis (s. str.) maculipennis Gory, 1840 Parish records (n 5 133): East Baton Rouge, 36. lecontei (Gory, 1841) Grant, Livingston, Rapides, and Winn Parish records (n 5 6): “Magnolia, Louisiana,” Parishes (LSAM). presumed Assumption Parish (LSAM, USNM). Potential larval host in Louisiana: Pinus taeda Potential larval host in Louisiana: Taxodium and Taxodium distichum. distichum. Collection date range: 5 May–4 October, Collection date: 25 March. mostly in May (n 5 94). Comments: Reared from ‘cypress.’ Theentirese- Comments: Collected using pheromone and ries is from a single rearing event during 1924. Lindgren funnel traps, and as C. fumipennis prey. Tribe Anthaxiini 37. Agrilaxia flavimana (Gory, 1841) 32. Buprestis (Cypriacis) striata Fabricius, 1775 Parish records (n 5 11): Caddo, Concordia, Parish records (n 5 2): East Baton Rouge and East Baton Rouge, St. Tammany, and Winn Parishes (LSAM). Webster Parishes (LSAM, MEM). Potential larval host in Louisiana: Pinus Potential larval hosts in Louisiana: Cercis palustris. canadensis, Pinus echinata, Prunus Collection date range: 27 February–18 May. domestica L., Quercus alba, Quercus Comments: Collection method unspecified and macrocarpa Michx., Quercus muehlenber- as C. fumipennis prey. gii Engelm., Quercus stellata Wangenh., Quercus virginiana, and Ulmus crassifolia. 33. Buprestis (Knulliobuprestis) rufipes Olivier, Collection date range: 7–25 May. 1790 5 Comments: Collected at light, using canopy Parish records (n 350): Acadia, Ascension, Malaise traps, and by sweeping. Bossier, Catahoula, East Baton Rouge, Grant, Iberville, Lincoln, Livingston, Rap- 38. Anthaxia (Haplanthaxia) cyanella Gory, 1841 ides, St. James, St. Tammany, West Felic- Parish records (n 5 12): East Baton Rouge, iana, and Winn Parishes (FSCA, LSAM, Tensas, and West Feliciana Parishes (LSAM). MEM, TAMU, USNM). Potential larval hosts in Louisiana: Amelanchier Potential larval hosts in Louisiana: Acer sac- arborea (Michx. f.) Fern., Betula nigra, charinum, Acer saccharum, Carya sp., Carya sp., Cercis canadensis, Crataegus Fagus grandifolia, Liriodendron tulipifera coccinea, Gleditsia triacanthos, Quercus L., Nyssa sylvatica, Quercus alba, Quercus nigra, Quercus velutina,andVitis sp. Adults palustris, Quercus virginiana, Ulmus reported also on Ostrya virginiana, Prunus americana, and Ulmus rubra. Adults re- americanus Marsh., Rhus aromatica Ait., ported also on Platanus occidentalis. and Ulmus rubra. Collection date range: 6 March–4 October, Collection date range: 17 May–22 July. Nine mostly in May (n 5 203) and June (n 5 81), reared specimens emerged June–July. 19 reared specimens emerged in April–August. Comments: Collected using boll weevil pher- Comments: Collected at light and by hand and omone and sticky traps. Reared from un- using Lindgren funnel and pitfall traps. specified coarse, woody debris. Reared from unspecified coarse, woody ı debris; also as C. fumipennis prey. 39. Anthaxia (Haplanthaxia) dichroa B´l´y, 1991 Parish records (n 5 3): East Baton Rouge and 34. Buprestis (Stereosa) apricans Herbst, 1801 Grant Parishes (FSCA, LSAM). Parish records (n 5 10): East Feliciana, Grant, Potential larval hosts in Louisiana: Vitis spp. and Winn Parishes (LSAM). Collection date range: 9–17 May (trap service Potential larval hosts in Louisiana: Pinus interval) and June (date unspecified). echinata, Pinus palustris, and Pinus taeda. Comments: Collected using boll weevil pher- Collection date range: 9 April–7 July. omone trap and reared from dead limbs Comments: Collected using Lindgren funnel (emerged June). traps, also as C. fumipennis prey. 40. Anthaxia (Haplanthaxia) fisheri Obenberger, 35. Buprestis (Stereosa) decora Fabricius, 1775 1928 Parish records (n 5 1): East Baton Rouge Parish Parish records (n 5 4): DeSoto, St. Landry, and (LSAM). Tensas Parishes (FSCA, LSAM). THE COLEOPTERISTS BULLETIN 72(2), 2018 357

Potential larval hosts in Louisiana: Prunus L., Gleditsia triacanthos, Juglans nigra, americana. Adults reported also on Acer sp., Malus sp., Quercus alba, Quercus rubra, Gleditsia triacanthos,andPrunus serotina. Rubus sp., and Ulmus alata Michx. Collection date range: 27 April–18 May. Collection date range: 8 April–12 May. Comments: Collected using boll weevil pher- Comments: Collected at light and boll weevil omone trap. pheromone traps. 41. Anthaxia (Haplanthaxia) quercata (Fabricius, Tribe Xenorhipidini 1801) 44. Xenorhipis brendeli LeConte, 1866 Parish records (n 5 81): Bossier, Caddo, Cat- Parish records (n 5 3): East Baton Rouge and ahoula, East Baton Rouge, East Feliciana, Franklin Parishes (LSAM). Grant, Jefferson Davis, Livingston, Natch- Potential larval hosts in Louisiana: Betula nigra, itoches, Rapides, St. Helena, St. Landry, St. Carya glabra, Carya illinoinensis, Carya Tammany, and Webster Parishes (FSCA, laciniosa, Carya ovata, Cercis canadensis, LSAM, MEM, TAMU). and Quercus alba. Adults reported also on Potential larval hosts in Louisiana: Pinus Fagus sp., Gleditsia triacanthos, Quercus echinata and Pinus sp. possibly taeda; virginiana, and Quercus velutina. records of larvae (but not adults) utilizing Collection date range: 15 May–17 July. angiosperms probably not this buprestid Comments: Collected using canopy Malaise species (MacRae 2006). Adults reported traps, also as C. fumipennis prey. on Carya tomentosa, Corylus americana Walt., Prunus americana, Quercus cocci- Tribe Melanophilini nea, Quercus laurifolia, Quercus palustris, 45. Melanophila atropurpurea (Say, 1823) Quercus rubra, Quercus stellata, Quercus Parish records (n 5 1): St. Tammany Parish velutina,andSassafras albidum. (LSAM). Collection date range: 19 April–31 May, mostly 5 Potential larval host in Louisiana: Unknown, in April (n 49). possibly Pinus spp. Comments: Collected at light, by beating and Collection date: 13 August. using boll weevil pheromone traps. Comments: New State Record: St. Tammany 42. Anthaxia (Haplanthaxia) viridicornis (Say, 1823) Parish, 4.2 mi N Abita Springs, 30.5162°N, Parish records (n 5 2): East Baton Rouge Parish 89.9547°W, collected at mercury vapor 5 (LSAM). light, 13 Aug. 2007, V. Brou (n 1). Potential larval hosts in Louisiana: SaIix car- 46. Melanophila notata (Laporte and Gory, 1837) oliniana, Salix exigua, and Salix nigra; Parish records (n 5 1): East Baton Rouge Parish records of larvae utilizing Carya spp., (LSAM). Quercus spp., and Ulmus spp. probably not Potential larval hosts in Louisiana: Pinus spp. this buprestid species (MacRae 2006). Adults reported also on Pinus echinata. Adults reported also on Populus sp. and Collection date: 10 August. Salix babylonica L. Comments: Collecting method unspecified for Collection date range: 26 February–23 April. this specimen collected in 1926. Comments: Collected by beating and sweeping. 47. Phaenops aeneola (Melsheimer, 1845) 43. Anthaxia (Haplanthaxia) viridifrons Gory, Parish records (n 5 6): Caddo, Calcasieu, 1841 5 Grant, St. Helena, Tangipahoa, and Vernon Parish records (n 15): Bossier, Catahoula, East Parishes. Baton Rouge, Grant, St. Landry, St. Martin, Potential larval hosts in Louisiana: Unknown, and Tensas Parishes (FSCA, LSAM, MEM). probably Pinus spp. Adults reported on Potential larval hosts in Louisiana: Amelanchier Pinus echinata. arborea, Carya cordiformis, Carya glabra, Collection date range: 13 May–5 June. Carya illinoinensis, Carya ovata, Carya Comments: All specimens collected by beating tomentosa, Morus sp., Ulmus americana, Pinus spp. Ulmus crassifolia, and Ulmus rubra; re- cords of larvae utilizing Salix spp. and Vitis 48. Phaenops obtusa (Horn, 1882) spp. probably not this buprestid species Literature record only. (MacRae 2006). Adults reported also on Potential larval hosts in Louisiana: Pinus spp. Acer saccharum, Asimina triloba (L.) Comments: Nelson and Westcott (1976) Dunal, Carya cordiformis, Cercis cana- recorded a specimen beaten from Pinus sp. in densis, Crataegus sp., Fraxinus americana Livingston Parish on 16 May. 358 THE COLEOPTERISTS BULLETIN 72(2), 2018

Tribe Phrixiini Iberville, Livingston, Rapides, Richland, 49. Spectralia gracilipes (Melsheimer, 1845) Sabine, St. Landry, St. Martin, St. Tam- Parish records (n 5 19): Concordia, East Baton many, West Feliciana, and Winn Parishes Rouge, West Feliciana, and Winn Parishes (LSAM, TAMU, USNM). (LSAM). Potential larval hosts in Louisiana: Baccharis Potential larval hosts in Louisiana: Fraxinus sp., halimifolia, Betula nigra, Carya illinoi- Ostrya virginiana, Quercus alba, Quercus nensis, Celtis laevigata, Cercis canadensis, macrocarpa,andQuercus stellata. Adults , Ficus sp., Juglans reported also on Celtis sp. and Crataegus sp. nigra, Quercus falcata (see Comments), Collection date range: 4 May–3 August, mostly Taxodium distichum, Ulmus crassifolia, and in May (n 5 8). Seven reared specimens Ximenia americana L. Adults reported also emerged in May. on Morella cerifera (L.) Small, Prunus Comments: Collected by hand, using canopy persica, Salix sp., and Vitis sp. Malaise traps, and by rearing from un- Collection date range: 7 February–28 November, specified coarse, woody debris; also as C. especially April (n 5 12) and May (n 5 11); 69 fumipennis prey. reared specimens emerged February–July. Comments: Collected by hand on ‘oak’ and ‘live Tribe oak’, using canopy Malaise, flight intercept, 50. adelpha Harold, 1869 5 Lindgren funnel, Malaise, sticky traps, Parish records (n 17): Concordia Parish ‘window pane trap on cypress’, and reared (LSAM). from Quercus falcata, ‘dead cypress limbs’, Potential larval hosts in Louisiana: Amelanchier ‘dead pecan limbs’, ‘dead hickory limbs’, arborea, Carya glabra, Carya illinoinensis, ‘miscellaneous dead limbs’, and unspecified Carya laciniosa, Carya ovata, Carya coarse, woody debris, also as C. fumipennis tomentosa, Prosopis glandulosa.Adultsre- prey. ported also on Fraxinus pennsylvanica Marsh. and Quercus sp. 54. Chrysobothris cribraria Mannerheim, 1837 Collection date range: 30 April–15 July, mostly Parish records (n 5 26): East Baton Rouge, in June (n 5 8). Natchitoches, Rapides, St. Tammany, Comments: All specimens collected by canopy Washington, and Winn Parishes (FSCA, Malaise traps. LSAM, MEM, TAMU). 51. Chrysobothris caddo Wellso and Manley, Potential larval host in Louisiana: Pinus echi- 2007 nata. Adults reported also on Pinus pal- Parish records (n 5 54): East Baton Rouge and ustris and Pinus taeda. Collection date range: 1 April–17 June, also 25 Winn Parishes (LSAM). 5 5 Potential larval hosts in Louisiana: Celtis lae- October (n 3), mostly in May (n 8). Comments: Collected by hand, using Leggett vigata Willd. and Cercis canadensis. ‘ ’ Collection date range: 27 April–9 July, mostly trap, and by beating pine. Also as C. in May (n 5 47). fumipennis prey. Comments: Collected only as C. fumipennis prey. 55. Chrysobothris dentipes (Germar, 1824) 52. Chrysobothris chlorocephala Gory, 1841 Parish records (n 5 22): Natchitoches, St. Parish records (n 5 4): East Baton Rouge and Tammany, and Winn Parishes (LSAM, Evangeline Parishes (FSCA, TAMU). MEM, USNM). Potential larval hosts in Louisiana: Acer sp., Potential larval hosts in Louisiana: Unknown, Amelanchier arborea, Betula nigra, Carya probably Pinus spp. Adults reported on glabra, Carya illinoinensis, Cercis cana- Pinus echinata, Pinus elliottii, Pinus pal- densis, Quercus nigra, Quercus stellata, ustris, and Pinus taeda. and Vitis sp. Adults reported also on Acer Collection date range: 18 May–16 June, mostly rubrum and Quercus alba. in May (n 5 14). Collection date range: Collected May; emerged Comments: Collection methods unspecified May and June. and as C. fumipennis prey. Comments: Collected by rearing from ‘dead ’ ‘ ’ ‘ 56. (Olivier, 1790) pecan limbs , dead hickory limbs ,and misc. 5 dead limbs.’ Parish records (n 13): Acadia, East Baton Rouge, East Feliciana, and St. Tammany 53. Chrysobothris chrysoela Illiger, 1800 Parishes (LSAM, USNM). Parish records (n 5 112): Avoyelles, Con- Potential larval hosts in Louisiana: Acer negundo cordia, East Baton Rouge, Lincoln, L., Acer rubrum, Acer saccharinum, Carpinus THE COLEOPTERISTS BULLETIN 72(2), 2018 359

caroliniana, Carya illinoinensis, Cercis can- marilandica Muenchh., Quercus palustris, adensis, Cornus florida, Crataegus crus-galli Quercus shumardii Buckl., Quercus stellata, L., Malus spp., Platanus occidentalis, Populus Quercus velutina, and Ulmus americana. deltoides, Prunus armeniaca L., Prunus Collection date range: 30 March–12 September, domestica, Prunus persica, Prunus serotina, mostly in May (n 5 16), 12 reared speci- Tilia americana, Ulmus Americana,and mens emerged March–May. Ulmus parvifolia Jacq. Many larval host plants Comments: Collected at light, by hand, using reported for this species prior to the recent Lindgren funnel and pitfall traps, and reared revision of the C. femorata species-group from unspecified coarse, woody debris; also (Wellso and Manley 2007) likely refer to as C. fumipennis prey. other species, especially oaks, which this 60. Chrysobothris rugosiceps Melsheimer, 1845 species does not normally utilize. Adults re- 5 ported also on Betula nigra, Fraxinus penn- Parish records (n 12): Bossier, Caddo, sylvanica, Gleditsia triacanthos, Salix exigua, Concordia, East Feliciana, Grant, Lincoln, Sorbus aucuparia L., Tilia cordata Mill., and and Winn Parishes (LSAM, MEM). Ulmus rubra. Potential larval hosts in Louisiana: Castanea Collection date range: 9 April–16 July, mostly dentata Borkh., Quercus alba,andQuercus in June (n 5 5). macrocarpa. Adults reported also on Carya Comments: Collected using Malaise traps and ovata, Quercus marilandica, Quercus pal- by sweeping, also as C. fumipennis prey. ustris, Quercus stellata,andQuercus velutina. Collection date range: 16 April–7 August. 57. Chrysobothris purpureovittata purpureovittata Comments: Collected using canopy Malaise Horn, 1886 and Lindgren funnel traps. Parish records (n 5 1): Bossier Parish (USNM). Potential larval hosts in Louisiana: Carya illi- 61. Chrysobothris scitula Gory, 1841 5 noinensis, Celtis laevigata, Celtis tenuifolia Parish records (n 13): East Baton Rouge, Nutt., Cercis canadensis, Crataegus viridis, Madison, and West Feliciana Parishes Gleditsia triacanthos, Koelreuteria pan- (LSAM, MEM). iculata Laxmann, Parkinsonia aculeata L., Potential larval hosts in Louisiana: Unknown. Prunus sp., and Ulmus crassifolia. Adults Adults reported on Quercus sp. – reported also on Vachellia farnesiana, Collection date range: 30 April 12 June, two Crataegus phaenopyrum (L. f.) Medik., reared specimens emerged in May and June. Diospyros virginiana, Prosopis glandulosa, Comments: Collected at light, by hand, by Prunus americana, and Quercus sp. sweeping, and by rearing from unspecified Collection date: 10 May. coarse, woody debris; also as C. fumipennis Comments: Collection methods unspecified. prey. 58. Chrysobothris pusilla Gory and Laporte, 62. Say, 1839 5 1837 Parish records (n 14): Concordia, East Baton Parish records (n 5 1): Ouachita Parish (LSAM). Rouge, East Feliciana, Orleans, Rapides, West Potential larval hosts in Louisiana: Unknown, Feliciana, and Winn Parishes (LSAM, MEM). probably Pinus spp. Adults reported also on Potential larval hosts in Louisiana: Acacia sp., Pinus echinata. Acer rubrum, Acer saccharum, Ame- Collection date: 6 June. lanchier arborea, Betula nigra, Carya Comments: Collected by ‘beating pine.’ glabra, Carya illinoinensis, Carya ovata, Castanea dentata, Celtis laevigata, Celtis 59. Chrysobothris quadriimpressa Gory and tenuifolia, Cercis canadensis, Crataegus Laporte, 1837 viridis, Diospyros virginiana, Fagus grandi- Parish records (n 5 41): Acadia, Concordia, folia, Fraxinus americana, Fraxinus pennsyl- East Baton Rouge, Grant, Iberville, Rap- vanica, Gleditsia triacanthos, Juglans nigra, ides, Webster, West Feliciana, and Winn Prosopis sp., Quercus alba, Quercus macro- Parishes (LSAM, MEM, USNM). carpa, Quercus muehlenbergii, Quercus stel- Potential larval hosts in Louisiana: Juglans lata, Taxodium distichum, Ulmus alata, Ulmus nigra, Liquidambar styraciflua L., Quercus rubra,andVitis sp. Adults reported also on alba, Quercus coccinea, Quercus rubra, Acer saccharinum and Quercus palustris. and Sapindus saponaria. Adults reported Collection date range: 23 April–7 July. Two also on Celtis laevigata, Diospyros virgin- reared specimens emerged May–June. iana, Fraxinus pennsylvanica, Juglans Comments: Collected at light, using canopy nigra, Quercus imbricaria Michx., Quercus Malaise, flight intercept, and Lindgren funnel 360 THE COLEOPTERISTS BULLETIN 72(2), 2018

traps, and reared from unspecified coarse, Tribe woody debris; also as C. fumipennis prey. 66. Agrilus (s. str.) abductus Horn, 1891 Literature record only. 63. Chrysobothris shawnee Wellso and Manley, Potential larval hosts in Louisiana: Unknown. 2007 Adults reported on Cornus florida, Gle- 5 Parish records (n 168): Bossier, Concordia, ditsia triacanthos, Quercus stellata, and East Baton Rouge, East Feliciana, Evan- Zanthoxylum clava-herculis L. geline, Franklin, Grant, St. Tammany, Comments: Nelson and MacRae (1990) recorded Tensas, and Winn Parishes (LSAM, MEM). specimens collected in Tangipahoa Parish on Potential larval hosts in Louisiana: Castanea 25 April. dentata, Quercus palustris, Quercus phellos, Quercus rubra,andQuercus stellata. Adults 67. Agrilus (s. str.) acutipennis Mannerheim, 1837 reported also on Celtis laevigata, Fraxinus Parish records (n 5 1): Bossier Parish (MEM). pennsylvanica, Quercus alba, Quercus cocci- Potential larval hosts in Louisiana: Quercus nea, Quercus marilandica, Quercus michauxii alba and Quercus lyrata Walt. Adults re- Nutt., Quercus palustris,andQuercus velutina. ported also on Amelanchier arborea, Betula Collection date range: 29 March–5 November, sp., Carya cordiformis, Corylus americana, mostly in May (n 5 76) and June (n 5 33). Diospyros virginiana, Juglans nigra, Populus Comments: Collected at light, using canopy sp., Ostrya virginiana, Quercus imbricaria, Malaise, and Malaise traps; also as C. Quercus laurifolia, Quercus macrocarpa, fumipennis prey. Quercus marilandica, Quercus stellata,and Quercus velutina. 64. Chrysobothris viridiceps Melsheimer, 1845 Collection date: 30 April. Parish records (n 5 32): Concordia, Franklin, Comments: Collected using Leggett trap. St. Tammany, West Feliciana, and Winn 68. Agrilus (s. str.) arcuatus (Say, 1825) Parishes (LSAM). 5 Potential larval hosts in Louisiana: Acer Parish records (n 20): East Baton Rouge and rubrum, Carya illinoinensis, Pinus sp., Grant Parishes (LSAM). Prosopis glandulosa, Prunus sp., Quercus Potential larval hosts in Louisiana: Castanea alba, Quercus macrocarpa, Quercus stel- dentata, Fagus grandifolia, Quercus alba, lata, and Ulmus crassifolia. Adults reported Quercus ilicifolia Wangenh., and Quercus also on Acer saccharinum, Carya ovata, velutina. Adults reported also on Carya sp., Quercus macrocarpa, Quercus mar- Juglans nigra, Quercus shumardii, and ilandica, Quercus velutina,andUlmus Quercus stellata. Collection date range: 5 May–27 June, most in americana. 5 Collection date range: 30 April–17 July, with May (n 18). specimens evenly distributed between the Comments: Collected at light, using boll weevil extreme dates; 13 reared specimens emerged pheromone traps, and by hand; also as April–June. C. fumipennis prey. Comments: Collected by sweeping, using 69. Agrilus (s. str.) benjamini Fisher, 1928 canopy Malaise traps, and reared from un- Parish records (n 5 2): East Baton Rouge specified coarse, woody debris; also as C. andSt.TammanyParishes(LSAM, fumipennis prey. TCMC). Potential larval hosts in Louisiana: Unknown. Subfamily Adults reported on Quercus alba and Quercus Tribe Aphanisticini macrocarpa. 65. Aphanisticus cochinchinae seminulum Oben- Collection date range: 12 April–28 May berger, 1929 Comments: New state record: East Baton Rouge Parish records (n 5 3) West Feliciana Parish Parish,SiegenLane,300ydSofPerkinsRd, (LSAM). 12 April 1999, B. J. Viator (n 5 1); St. Potential larval hosts in Louisiana: Saccharum Tammany Parish, 4.2 mi NE Abita Springs, officinarum L., Tripsacum dactyloides (L.) 30.5162°N, 89.9547°W, 28 May 2000, V. L. Adults reported also on Paspalum liv- Brou (n 5 1). idum Trin. Collection date: 4 June. 70. Agrilus (s. str.) bilineatus (Weber, 1801) Comments: New state record: W. Feliciana Parish records (n 5 42): Assumption, Parish, Feliciana Preserve/Natural Area, Avoyelles, Concordia, East Baton Rouge, 30°47.68N 91°15.24W, sweeping, 4 Jun. Franklin, St. Tammany, West Feliciana, and 2006, M. Gimmel (n 5 3). Winn Parishes (LSAM). THE COLEOPTERISTS BULLETIN 72(2), 2018 361

Potential larval hosts in Louisiana: Castanea Collection date range: 11–13 May. dentata, Fagus sp., Quercus alba, Quercus Comments: New state record: East Baton coccinea, Quercus lyrata, Quercus macro- Rouge Par., Highland Rd. Park, 11,13 May carpa, Quercus muehlenbergii, Q. michauxii, 2011, collected as C. fumipennis prey, Quercus rubra, Quercus stellata, Quercus 30.3491°N, 91.0741°W, W. Virgets (n 5 2). texana Buckley, and Quercus velutina. Adults 75. Agrilus (s. str.) defectus LeConte, 1860 reported also on Q. nigra and Q. palustris. Parish records (n 5 2) East Baton Rouge and Collection date range: 26 April–9 September, Grant Parishes (LSAM). mostly (n 5 29) in May and early June. Potential larval hosts in Louisiana: Quercus Comments: Collected at light, using canopy alba, Quercus muehlenbergii,andQuercus Malaise, Lindgren funnel, and sticky traps; stellata. Adults reported also on Carya also as C. fumipennis prey. ovata, Carya tomentosa, Cercis canadensis, 71. Agrilus (s. str.) carpini Knull, 1923 Crataegus sp., Quercus macrocarpa, Quer- Parish records (n 5 1): Assumption Parish (LSAM). cus marilandica,andQuercus velutina. Potential larval hosts in Louisiana: Carpinus Collection date range: 28 March–4 April. caroliniana, Fagus grandifolia, and Ostrya Comments: New state record: East Baton virginiana. Rouge Parish, Baton Rouge, 30.5018N, Collection date: 2 June. 90.1063W, 4 April 2005, N. H. Nguyen, Hg Comments: Collecting method unspecified. vapor light trap (n 5 1); Grant Parish, Kisatchie National, Forest, Stuart Lake Rd. 72. Agrilus (s. str.) celti Knull, 1920 W of Pollock, 31.5124N, 92.4406W, 28 Parish records (n 5 6): Avoyelles, Bossier, East March 2011, ETOH baited Lindren funnel Baton Rouge, St. Landry, and West Baton trap (n 5 1). Rouge Parishes (LSAM, TAMU). Potential larval hosts in Louisiana: Celtis lae- 76. Agrilus (s. str.) difficilis Gory, 1841 vigatus and Celtis occidentalis. Adults re- Parish records (n 5 385): Assumption, ported also on Celtis tenuifolia, Cercis Avoyelles, East Baton Rouge, La Salle, St. canadensis, Corylus americana,andGle- Martin, and Tensas Parishes (LSAM). ditsia triacanthos. Potential larval hosts in Louisiana: Gleditsia Collection date range: 2 April–25 May. triacanthos and Zanthoxylum clava-herculis. Comments: New state record: Avoyelles Parish, Collection date range: 4 April–23 August, 25 May 1979 (n 5 1); Bossier Parish, near mostly (n 5 347) during May and June. Flag Lake, Barksdale AFB, 21 May 1996, E. Comments: Collected at light, by hand on a G. Riley (n 5 1);EastBatonRougeParish, sugar cane plant, ‘fogging cypress’, and Bayou Fountain Apts., 30.3622N, 91.1247W, using boll weevil pheromone and sticky hand collected, 9 May 2006, A. K. Tishechkin traps; also as C. fumipennis prey. (n 5 2); St. Landry Parish, Thistlethwaite 77. Agrilus (s. str.) egenus Gory, 1841 Wildlife Management Area, 30.6623N, Parish records (n 5 9): Bossier, Catahoula, East 91.9983W, 7 May 1989, D. Rider (n 5 1); Baton Rouge, and Natchitoches Parishes West Baton Rouge Parish, 30.4581N, (LSAM, MEM, USNM). 91.3239W, 2 May 1973, boll weevil phero- Potential larval hosts in Louisiana: Robinia mone trap (n 5 1). pseudoacacia. 73. Agrilus (s. str.) cephalicus LeConte, 1860 Collection date range: 6 April–22 May. Parish records (n 5 10): Bossier, Pointe Cou- Comments: Collected at light and by beating pee, and St. Tammany Parishes (LSAM, vegetation. Other methods unspecified; also MEM, TAMU, USNM). as C. fumipennis prey. Potential larval hosts in Louisiana: Cornus 78. Agrilus (s. str.) fallax Say, 1833 florida. Adults reported also on Corylus Parish records (n 5 52): Bossier, East Baton americana and Juglans nigra. Rouge, Jefferson, Pointe Coupee, St. Collection date range: 18 April–20 July. Landry, St. Tammany, and Tensas Parishes Comments: Collected at lights. Other methods (LSAM, MEM, USNM). unspecified. Potential larval hosts in Louisiana: Celtis 74. Agrilus (s. str.) concinnus Horn, 1891 laevigata, Celtis tenuifolia, Cornus flor- Parish records (n 5 2): East Baton Rouge Parish ida,andGleditsia triacanthos. Adults (LSAM). reported also on Crataegus sp. probably Potential larval hosts in Louisiana: Hibiscus viridis, Diospyros virginiana, Juglans laevis All. and Hibiscus lasiocarpos Cav. nigra,andTilia sp. 362 THE COLEOPTERISTS BULLETIN 72(2), 2018

Collection date range: 6 April–8August. Iberville, Jefferson, Natchitoches, Rapides, St. Specimens evenly distributed between dates. Landry, and Tensas Parishes (LSAM, TAMU). Comments: Collected at light, by hand, and using Potential larval hosts in Louisiana: Baccharis flight intercept and sticky (‘on cypress’)traps; sp., Celtis laevigata, and Celtis laevigata also as C. fumipennis prey. var. reticulata (Torr.) L. Benson. Adults reported also on Sapindus saponaria var. 79. Agrilus (s. str.) fuscipennis Gory, 1841 5 drummondii (H. & A.) L. Benson. Parish records (n 2): Bossier and East Baton Collection date range: 14 March–10 July, Rouge (LSAM, MEM). mostly (n 5 89) during April, 34 reared Potential larval hosts in Louisiana: Diospyros specimens emerged April–December. virginiana. Adults reported also on Oxy- Comments: Collected at light, by hand, using dendrum arboreum (L.) DC. – boll weevil pheromone, canopy Malaise, and Collection date range: 24 30 April. sticky (‘on cypress’)traps,‘in dead sugarberry’, Comments: New state record: Bossier Parish, reared from ‘hackberry’ and ‘miscellaneous Barksdale Air Force Base, 30 Apr. 1996, dead limbs.’ Also as C. fumipennis prey. Leggett trap in calcareous prairie, 32.4913°N, 93.5152° W, D. M. Pollock (n 5 1); East 84. Agrilus (s. str.) macer LeConte, 1858 Baton Rouge Parish, Baton Rouge, 25.iv.1979, Parish records (n 5 1,083) Concordia, East 30.4505°N, 91.1544°E, D. Demorvelle Baton Rouge, La Salle, and Orleans Par- (n 5 1). ishes (LSAM, MEM, TAMU). 80. Agrilus (s. str.) geminatus (Say, 1823) Potential larval hosts in Louisiana: Celtis lae- Parish records (n 5 1): West Feliciana Parish vigata. Adults reported also on Celtis lae- (LSAM). vigata var. reticulata. Collection date range: 6 May–23 August, Potential larval hosts in Louisiana: Quercus 5 5 phellos, Quercus marilandica, and Quercus mostly during May (n 534) and June (n velutina. Adults reported also on Carya 311); two reared specimens emerged during tomentosa, Juglans nigra, Quercus alba, February; emergence dates of 54 reared and Quercus coccinea. specimens were not recorded. Comments: Collected using canopy Malaise and Collection date: 8 May. ‘ ’ Comments: Collecting method unspecified. sticky traps; reared from Celtis sp. and unspecified tree limbs; also as C. fumipennis 81. Agrilus (s. str.) imbellis Crotch, 1873 prey as the most abundant (n 5 1,023, all East Literature record only. Baton Rouge Parish) species in our surveys. Potential larval hosts in Louisiana: Unknown. 85. Agrilus (s. str.) oblongus Fisher, 1928 Adults reported on flowers of Bigelowia 5 nudata (Michx.) DC., Lechea sp., Heli- Parish records (n 7). Catahoula, West Baton anthemum rosmarinifolium Pursh, and Rouge, and Tensas Parishes (LSAM). Potential larval hosts in Louisiana: Vitis sp. Rudbeckia sp. – Comments: Nelson et al. (2008) listed Louisiana Collection date range: 3 May 5 June in the distribution of this species, although no Comments: All specimens collected in boll supporting literature reference was found. weevil pheromone traps. 86. Agrilus (s. str.) obsoletoguttatus Gory, 1841 82. Agrilus (s. str.) lacustris LeConte, 1860 5 Parish records (n 5 15): Bossier, Cameron, Parish records (n 222): East Baton Rouge, DeSoto, and Evangeline Parishes (FSCA, Iberville, Jefferson, Pointe Coupee, Rap- LSAM, MEM, TAMU). ides, St. Landry, St. Martin, St. Tammany, Potential larval hosts in Louisiana: Croton Tensas, and West Feliciana Parishes capitatus Michx. and Solanum elaeagnifo- (LSAM, MEM, TAMU). lium Cav. Adults reported also on Croton Potential larval hosts in Louisiana: Aesculus punctatus Jacq. pavia L., Carpinus caroliniana, Carya sp., Collection date range: 19 March–21 June. Fagus grandifolia, Gleditsia triacanthos, Comments: Five specimens collected by hand Ostrya virginiana, Quercus falcata (see on Croton capitatus; also using Malaise Comments), Quercus prinoides, Quercus trap. Other methods unspecified. rubra, and Salix sp. Adults reported also on Acer saccharum, Diospyros virginiana, 83. Agrilus (s. str.) lecontei lecontei Saunders, Juglans nigra, Quercus alba, Quercus 1871 imbricaria, Quercus marilandica, Quercus Parish records (n 5 160). Acadia, Catahoula, palustris, Quercus velutina, Ulmus ameri- Concordia, East Baton Rouge, Grant, cana, and Ulmus rubra. THE COLEOPTERISTS BULLETIN 72(2), 2018 363

Collection date range: 26 February–22 June, bark; same locality, 1 May 2015, adults re- mostly (n 5 19) in April. Many of these covered from z-3-hexanol baited funnel traps records are from rearing chambers, which in Fraxinus, C. W. Johnson and J. R. Meeker. likely are not representative of adult activity 90. Agrilus (s. str.) politus (Say, 1825) under natural conditions. 5 Comments: Collected at light, by beating’,branch Parish records (n 11): East Baton Rouge bagging’, using boll weevil pheromone, flight Parish (LSAM). intercept, and sticky traps, and by rearing from Potential larval hosts in Louisiana: Salix bab- ‘ ’ ‘ ylonica and Salix nigra. Quercus falcata, hickory limbs , mis- – cellaneous dead limbs’, and unspecified Collection date range: 27 April 13 May. coarse, woody debris; also as C. fumipennis Comments: All but one specimen collected as prey. This was the most abundant of 35 beetle C. fumipennis prey. species reared from Q. falcata twigs by Ferro 91. Agrilus (s. str.) pseudofallax Frost, 1923 et al. (2009). Parish records (n 5 9): Caddo, East Baton 87. Agrilus (s. str.) olentangyi Champlain and Rouge, Madison, St. Landry, and St. Tam- Knull, 1925 many Parishes (FSCA, LSAM, USNM). Parish records (n 5 1): Caddo Parish (USNM). Potential larval hosts in Louisiana: Gleditsia tri- acanthos. Adults reported also on Quercus sp. Potential larval hosts in Louisiana: Rhus glabra – L. Adults reported also on Celtis laevigata, Collection date range: 12 April 18 July, all but Celtis tenuifolia, Maclura pomifera (Raf.) one specimen collected during April. C.K. Schneid., Ptelea trifoliata L., and Comments: Collecting methods unspecified. Quercus sp. 92. Agrilus (s. str.) quadriguttatus quadriguttatus Collection date: 7 April Gory, 1841 Comments: Fisher (1928) also recorded specimens Parish records (n 5 116): Concordia and East collectedinMadisonParishon7–9 April. Baton Rouge Parishes (LSAM). Potential larval hosts in Louisiana: Alnus sp. 88. Agrilus (s. str.) otiosus (Say, 1833) and Salix nigra. Adults reported also on Parish records (n 5 2): St. Landry and Tensas Populus nigra L. and Salix interior Rowlee. Parishes (LSAM). Collection date range: 6 May–10 July, mostly Potential larval hosts in Louisiana: Carya (n 5 106) during May. cordiformis, Carya laciniosa, Carya illi- Comments: One specimen collected using noinensis, Carya ovata, Carya tomentosa, canopy Malaise traps, remainder as C. and Sassafras albidum. fumipennis prey. Collection date range: 27 April–21 May. Comments: Collected by hand ‘on pecan tree’; 93. Agrilus (s. str.) ruficollis Fabricius, 1787 other method unspecified. Parish records (n 5 92): East Baton Rouge, East Feliciana, Jefferson, Lafayette, Livingston, 89. Agrilus (s. str.) planipennis Fairmaire, 1888 5 Natchitoches, Orleans, St. Landry, St. Parish records (n 5, plus survey reports): Tammany, Tensas, Webster, and West Bossier, Bienville, Claiborne, Jackson, Feliciana Parishes (LSAM, TAMU). Lincoln, Morehouse, Ouachita, Union, and Potential larval hosts in Louisiana: Rubus spp. Webster Parishes (LSAM, Webster Parish Collection date range: 19 February–21 May, only; other Parish records reported by mostly (n 5 51) during April, 27 reared Richard Miller, Louisiana Department Ag- specimens also emerged during April. riculture and Forestry (www.laforestry.com/ Comments: Collected at light, by beating, News/tabid/112/ArticleID/138/EAB-spreads- sweeping, using Malaise, flight intercept, to-nine-parishes.aspx#prettyPhoto, accessed and sticky traps; also reared from 4 October 2017). ‘blackberry.’ Potential larval hosts in Louisiana: Chionan- thus virginicus L. and Fraxinus spp. 94. Agrilus (s. str.) subcinctus Gory, 1841 Collection date range: Webster Parish larvae Parish records (n 5 2): Caddo and St. Landry collected during February; adults dur- Parishes (LSAM, USNM). ing May. Potential larval hosts in Louisiana: Fraxinus Comments: New state record: Webster Parish, americana, Fraxinus pennsylvanica. Adults 8.3 km N of Shongaloo, 1 May 2015, Hand reported also on Carya sp., Ligustrum sp., Collected, ex. Green Ash Tree 30.0167°N, and Toxicodendron radicans (L.) Kuntze. 93.3152°W, 5 Feb. 2015, C. W. Johnson and Collection date range: 7–27 April. J. R. Meeker, larvae collected in Fraxinus Comments: Collecting method unspecified. 364 THE COLEOPTERISTS BULLETIN 72(2), 2018

95. Agrilus (s. str.) vittaticollis (Randall, 1838) 99. Leiopleura carbonata (LeConte, 1860) Parish records (n 5 2): East Baton Rouge and Literature record only. St. Tammany Parishes (LSAM). Potential larval hosts in Louisiana: Unknown. Potential larval hosts in Louisiana: Amelanchier Comments: The unique holotype of this species arborea, Aronia arbutifolia (L.) Pers., Cra- is labeled as having been collected in Lou- taegus sp., Malus sp., and Pyrus communis isiana. However, it is likely a mislabeled L. Adults reported also on Malus sylvestris Central or South American specimen, and its (L.) Mill., Oxydendrum arboreum, Prunus inclusion in the fauna of the USA should be virginiana L., and Quercus velutina. considered doubtful (Hespenheide 2003). Collection date range: 4–29 April. Comments: New state record: East Baton Rouge 100. Pachyschelus laevigatus (Say, 1839) 5 Par., Baton Rouge, at light, 30.4505°N, Parish records (n 49): Cameron, Catahoula, 91.1544°W; St. Tammany Parish, E. Fk. Lit. Claiborne, East Baton Rouge, Iberville, Jef- Bogue Falaya R., LA 1082 NE of Covington, ferson, Natchitoches, Rapides, Webster, and 29 April 1980 sweeping foliage, 30.4901°N, West Feliciana Parishes (LSAM, TAMU). 90.0686°W, D. A. Rossman (n 5 1). Potential larval hosts in Louisiana: Desmodium ciliare (Muhl. ex Willd.) DC., Desmodium 96. Paragrilus tenuis (LeConte, 1866) obtusum (Muhl. ex Willd.) DC., Desmodium Parish records (n 5 1): Caddo Parish (LSAM). viridiflorum (L.) DC., Lespedeza capitata Potential larval hosts in Louisiana: Hibiscus Michx., and Lespedeza virginica (L.) Britton. laevis, Hibiscus lasiocarpos, and Hibiscus Adults reported also on Desmodium can- moscheutos L. escens (L.) DC., Desmodium cuspidatum Collection date: 30 June. (Muhl. ex Willd.) DC. ex Loud., Desmodium Comments: New state record: Caddo Parish, glutinosum (Muhl. ex Willd.) Wood, Des- Shreveport, 30.vi.1974, J. R. Barr Sr. modium paniculatum (L.) DC., Desmodium (n 5 1). perplexum Schub., Lespedeza bicolor Turcz., and Lespedeza hirta (L.) Hornem. Tribe Tracheini Collection date range: April–11 October, most 97. Brachys fasciferus Schwartz, 1878 5 5 collected during May (n 14) and July Parish records (n 1): St. Landry Parish (n 5 15). (LSAM). Comments: Collected at light, by hand, ‘on Potential larval hosts in Louisiana: Unknown. Desmodium’, and by sweeping. Adults reported on Quercus virginiana and Rubus trivialis Michx. 101. Pachyschelus nicolayi Obenberger, 1925 Collection date: 27 April. Parish records (n 5 7): Orleans, St. Tammany, and Comments: New state record: St. Landry Parish, Webster Parishes (FSCA, TAMU, USNM). 3 mi. E. of Washington, 27 April 1986, Potential larval hosts in Louisiana: Apios 30.6300°N, 92.0062°W, D. A. Rider and E. G. americana Medik. and Wisteria frutescens Riley (n 5 1). (L.) Poir. Collection date range: 11 April–19 May. 98. Brachys ovatus (Weber, 1801) Comments: Collecting methods unspecified. Parish records (n 5 84): Assumption, DeSoto, East Baton Rouge, Jefferson, Morehouse, 102. Pachyschelus purpureus (Say, 1833) Natchitoches, Plaquemines, Rapides, St. Parish records (n 5 1): East Baton Rouge Landry, St. Tammany, Tangipahoa, Vernon, (FSCA). Webster, and West Feliciana Parishes Potential larval hosts in Louisiana: Geranium (FSCA, LSAM, MEM, TAMU, USNM). maculatum L. Adults reported also on Potential larval hosts in Louisiana: Quercus incana Carya sp., Fraxinus americana, Juglans W. Bartram and Quercus rubra. Adults re- nigra, and Toxicodendron radicans. ported also on Quercus alba, Quercus falcata, Collection date: 23 May. Quercus imbricaria, Quercus laevis Walt., Comment. Nelson et al. (2008) listed Louisiana Quercus marilandica, Quercus shumardii, within the distribution of this species, possibly Quercus stellata,andQuercus velutina. based on this specimen collected during 1988. Collection date range: 20 March–14 October, mostly during April (n 5 42) and May (n 5 17). 103. Taphrocerus agriloides Crotch, 1873 Comments: Collected by hand, sweeping, us- Parish records (n 5 2): Iberville Parish (LSAM). ing boll weevil pheromone, flight in- Potential larval hosts in Louisiana: Unknown. tercept, Leggett, Malaise traps, and as C. Adults reported on Eleocharis sp. and fumipennis prey. Nemastylis geminiflora Nutt. THE COLEOPTERISTS BULLETIN 72(2), 2018 365

Collection date range: 5 April–12 June. 108. Taphrocerus laevicollis LeConte, 1878 Comments: New state record: Iberville Parish, 1 Parish records (n 5 1): Jefferson Parish (FSCA). mi S Hwy 30, 12.vi.1994, L. M. Rodriguez, Potential larval hosts in Louisiana: Unknown. under log (n 5 1); St. Gabriel, 5.iv.2005, L. Collection date range: 8–9 September Eisenberg, sweeping vegetation (n 5 1). Comments: This is the specimen on which Nelson’s (1987) record was based. 104. Taphrocerus albodistinctus Knull, 1954 Parish records (n 5 26): Ascension, Assumption, 109. Taphrocerus nicolayi Obenberger, 1924 East Baton Rouge, Grant, Jefferson, Natch- Parish records (n 5 43): Beauregard, Cameron, itoches, Vernon, St. Martin, Washington, and Catahoula, East Baton Rouge, Iberville, West Feliciana Parishes (LSAM). Livingston, Natchitoches, St. Tammany, Potential larval hosts in Louisiana: Unknown. Tangipahoa, Webster, and West Baton Rouge Collection date range: 12 March–21 November, Parishes (FSCA, LSAM, MEM, TAMU). mostly (n 5 14) March and April. Potential larval hosts in Louisiana: Unknown. Comments: Collecting methods mostly un- Adults reported on Carex hyalinolepis and specified; some collected by beating grasses, Carex vulpinoidea Michx. using flight intercept traps, and sweeping. Collection date range: 20 February–6 October. Comments: Collecting methods unspecified. 105. Taphrocerus cylindricollis Kerremans, 1896 Parish records (n 5 21): Natchitoches, St. Mary, 110. Taphrocerus schaefferi Nicolay and Weiss, 1920 St. Tammany, and West Feliciana Parishes 5 (LSAM). Parish records (n 9): East Baton Rouge, Evan- Potential larval hosts in Louisiana: Unknown. geline, St. Tammany, Washington, Webster, Adults reported on undetermined Cyperaceae. and West Feliciana Parishes (LSAM, MEM). Collection date range: 29 March–8 September. Potential larval host in Louisiana: Cyperus Comments: Collecting methods unspecified on esculentus L. Adults reported also on Carex most specimens, some collected by flight vulpinoidea and Desmodium sp. (see intercept trap and sweeping. Comments). Collection date range:12 April–31 October. 106. Taphrocerus gracilis (Say, 1825) Comments: Collected by beating, by hand ‘on Parish records (n 5 11): Acadia, Ascension, Desmodium’, and by sweeping. Other col- East Baton Rouge, Livingston, St. Tam- lecting methods unspecified. many, and West Baton Rouge Parishes (FSCA, LSAM). Potential larval hosts in Louisiana: Rhynchospora DISCUSSION corniculata (Lam.) Gray. Adults reported also feeding on Carex hyalinolepis Steud. and col- Johnson et al. (2015) reported 35 species of lected on Baccharis halimifolia, Cephalanthus buprestids as prey of C. fumipennis, based on occidentalis L., and Rumex verticillatus L. multiyear surveys, mostly at two localities, one in Collection date: 4 March–4 August. Catahoula Parish, central Louisiana, another in East Comments: Collection method mostly un- Baton Rouge Parish, south-central Louisiana. Total specified, some collected by sweeping. buprestid specimens represented in that study was 1551. Those records are included in the number 107. Taphrocerus howardi Obenberger, 1934 reported in this checklist. Thus, recent surveys Parish records (n 5 52): Acadia, Ascension, based on C. fumipennis surveys represent approx- Cameron, DeSoto, East Baton Rouge, East imately one-third of the total number of buprestid Feliciana, Iberville, Jefferson, Lafayette, specimens known from the state. Despite the con- Livingston, Orleans, Pointe Coupee, St. venience of stealing buprestids from heavily laden, Tammany, and West Feliciana Parishes slow flying wasps, the results present a skewed (FSCA, LSAM). survey method due to the biological preferences of Potential larval hosts in Louisiana: Unknown. the wasps (Hellman and Fierke 2014) and the un- Adults reported also on Fimbristylis even distribution of C. fumipennis aggregations, puberula (Michx.) Vahl and Scirpus sp. which tend to be concentrated in human-created, Collection date range: 11 March–4 November. open spaces such as parks and sports fields. For Comments: Collected by hand ‘on aquatic example, if the results from Johnson et al. (2015) vegetation’ and ‘ex. grass’, flight intercept and our museum records were accepted as typical, trap, and sweeping, including ‘mixed the city of Baton Rouge would appear to be the grasses and forbs’, ‘riverside vegetation’, epicenter of diversity and abundance of buprestids and Trifolium incarnatum L. for the state. Although “more data are needed” has 366 THE COLEOPTERISTS BULLETIN 72(2), 2018 become a tired clich´e in studies of this sort, it is ACKNOWLEDGMENTS certainly a true statement in this case. Clearly, surveys utilizing C. fumipennis are a valuable part of We thank Richard Miller (Louisiana Department EAB and general buprestid surveys, and the doc- of Agriculture and Forestry) for information about umentation of new state records based on C. EAB Louisiana survey results. Participants in fumipennis captures in this and other projects (e.g., C. fumipennis surveys in addition to the authors Westcott and Thomas 2015) is a testament to their included Michael Ferro, Forest Huval, Brittany value in adding to the overall knowledge of Owens, Warren Virgets, Jong-Seok Park, and buprestid beetle diversity and distributions. Surveys Mikhail Tishechkin. We thank Crystal Maier, at several localities using C. fumipennis wasps to do Edward Riley, and Richard Westcott for identifi- the collecting produced valuable additional data that cations. We thank Terrence Schiefer (MEM), are qualitatively and quantitatively different from Edward Riley (TAMU), Paul Skelley, and Kyle previous museum data. Schnepp (both FSCA) for making specimens and The negative impact of EAB on ash trees data in their care available for study. This publi- (Fraxinus spp.) in forests and urban tree plantings cation is approved by the Director, Louisiana Ag- is well documented (Poland and McCullough ricultural Experiment Station as manuscript number 2006), but the secondary impact on existing 2016-234-27539. This work is/was supported by the buprestid diversity is less well understood. Such USDA National Institute of Food and Agriculture impacts may include declines in native buprestid Hatch Project LAB94244. abundance and diversity due to EAB-triggered increases in abundances of buprestid predators REFERENCES CITED and parasitoids. At least one buprestid, A. sub- cinctus, may experience direct negative effects, as Ferro, M. L., M. L. Gimmel, K. E. Harms, and C. E. dead ash twigs are its only known larval food Carlton. 2009. The beetle community of small oak (although initially its numbers may climb as the twigs in Louisiana, with a literature review of Coleoptera from fine woody debris. 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