Vol. 4, No. 1-4, March-December 1990 73

Dlstnbutlonal, Biological and Iaxonomlc Notes on North American (Coleoptera)

Richard L. Westcott Plallt Divisioll, 0regoll Deparlilielit of AgricullUle Salem, OR 9/310-0110, USA

Abstract care. Appreciation goes to L. H. Herman and A. Distributional, biological and taxonomic F. Newton for the loan of type material, and to data are provided for 64 species of Buprestidae G. H. Nelson for reVlewmg the manuscript. I belonging to the genera Acmaeodem Ac- am especially indebted to my son, Derek, 'llith maeoderoides, Actenodes, Agaeocera, A/srilus, out whose help I could not have generated this Buprestis, Chrysophana, Descarpentriesina, paper on our computer His patience with "the Dicerca, Hippomelas. Melanophila, Pachysche- old man" is commendable. Ius, Polycesta, Ptosima, Spectralia, Taphrocerus, Traehykele and Tyndaris. Forty six of these Acmaeodem connexa LeConte. Apparently s ecies are recorded for the first time from a the .only s~ecifica.lly recorded Washington vi, ew rv os re or s ,, are provided for 15 others. Taphrocerus sulcif- an unlikely habitat. Since it is based on a rons Fisher is recorded from the U.S. for the specimen (OSUC) originating from the O. B. first time. Evidence is given to eliminate the Johnson Collection it is further suspect. I am recorded occurrence of Agrilus arbuti Fisher in not aware of any other specimens of this spe- British Columbia and Ptosima walshii LeConte des, the known hosts of which are oaks, flOm in Califor nia. western Washington In western Oregon it is occasionally collected as far north as Polk and Introduction Yamhill Cos. The following have been collected Data prouided herein largely resulted from ieton, 31~-:J ; ~c tat 0., It7a~­ my determinations and collecting during recent 0., on, 12-"JIIII-13, aBBC, and Batns Pass, 6-VI/30- years, and are presented to voucher their inclu­ VII-69, RLWE. This is common in Cali- sio~ ~ I?ep~rt~nt ~cul­ in II of fornia and SW Oregon. tur Smltiresoman nstItutIOn atalogorieop- tela (G. II. Nelson, in pIep.). GeneIa and Acmaeodera conoidea Fall. NEW MEXICO, Eday Co., 16 KIn SSW Whites City, Il4um, 12- V-89, RLWE. Chamberlin's (1926) record from TALS are presumed new. Host records include adults taken on a plant or reared/cut from their hOsts, as indicated, and are presumed new unless marked with an asterisk (*) or otherwise clarified. Collection codens are as listed in Arnett and Samuelson (1986) unless otherwise indicated and IefeI to all data pIeceding them. V/FBC Acmaeodera constricticollis Knul!. This ~ ~rr Collecti~n ~iS~ F I to!bank indiyidu­ species has been known onIy from too hOlotyPe is an curators 0 tile mSltutIOns mentIOned collected at Phoenix, Arizona, but may be for the opportunity to examine material in their 74 Insecta Mundi confused m collectIOns under Its sIblmg, A. Acmaeodera latl/laua latl/laua Fall. NEVA· fisheri Cazier. Knull (1937) related it to A. DA, Clark Co., Grand Gulch Road, 22 air mi S. cribricollis Horn, to which it bears no resem­ Mesquite, 11/21-V-83, EMUS; Spring Mts., Lee blance CALIFORNIA, Imperial Co ,Winterha- Can, I mi W Hwy 95, 16-VI-82, Ti'allugia ven, 5-VII-64, 3-VII-66; 1.5 mi W Winterhaven, paradoxa (D.Don)EndL blossom, GHNC. 17-VI-80. Ri\JeI side Co., Coachella, 16-"'11-63, 12-VI-78, 30-VI-88; 6 mi SMecca, 30-VI-88; Acmaeodera ligulata Cazier UTAH, Wash- Thermal, 16-VI-63, all from flowers ofPluchea ington Co., Gunlock St. Beach, 17-V-84, EMUS. sericea (Nutt.) Co,\,., CLBC, DSVC, GIINC, RLWE. Specimens were reared from Atriplex Acmaeodera neoneglecta Fisher. Rearing canescens (Pursh) Nuttall, Ailzona, "Yuma Co., records from Texas are ex Leucaena pulueru­ Colorado River, Yuma, JNFBC. lenta (Schlecht)Bentham, Texas, Cameron Co., Palm Grove Sanctuarynear Brownsville, em. V- Acmaeodera diffusa Barr. NEW MEXICO, 79, FTHC; ex Sophora secundi/lora (Gomez Raton, 17 VI 35, MCZC. Ortega) Lag. ex DC., Real Co., 11 mi NW Leak- ey, em III-90; ex Ulmus crassifolia Nutt., Hidal­ Aemaeodera fisheri Fall Reared from pupa go Co, Anzalduas Park, wood coIl. IX-88, em. exFouquieria splendens Engelmann, Texas, Val IV-89, D.J. Heffern ColI. Verde Co., Seminole Can. St. Park, 23-VII-90, D.Vl. Sundbelg ColI. No lal val host fOl tiris Aemueodelu pUladis.;uncta Knull. NEVi beetle had been recorded, nor am I aware ofany MEXICO, Eddy Co., Sitting Bull Falls, 22-VIII­ otherAcmaeodera species 'which utilizes ocotillo. 85, BYUe.

Aemaeodera ftsheri Cazier. Reared from Acmaeodera pletura Barr. UTAH, Garfield Eriogonum inflatum Torr. &; Frem. collected in Co., Calf Cr., 24 V 81, EMUS; 13 mi NE Esca- California, Imperial Co., Mt. Springs and Borre- lante, 17-VI-84, CUIC. Zion Nat. Park, Watch- go, V-79, em. 'V & VI-79, BKDC. man TIail, 31-V-82, RLVIE. These specimens differ from paratypes in my collection byhaving Acmaeodera immaculata Horn. the punctures of the head coarser and almost MONTANA, Madison Co., 52 mi S Ennis, VII- leticulate, those of the plonotum slightly coars- 66; Rosebud Co , 9 mi N Lame Deer, 8-VII-65, er, and by exhibiting an indistinct subapical M'I'EC. WYOMING, Carbon Co., Hanna, Sellli­ plate on the last Visible abdommal stermte. nole Coal Mine #1, 22-"'111-76, Park Co., 16 mi SE Powell, 22-VI-82, ESUW. According to R. Acmaeodera pulchella (Herbst). NORTH LaVigne (in litt.), at the latter locality toore DAKOTA,McKenzie Co., '1'000. Roosevelt Mem. were hundreds of flying to painted Park, 24-VI-64; Pembina Co., 12-VII-62, RLWE. (01 ange) stakes, yellow corncob grit bait, flowers WYUMING, Crook Co., Devils Tower Nat. of Opuntia polyacantha Haw.* and Sarcobatus Mon., 4000', 23·VI-83, RLWE, ESUW. Ciiamb- uermiculatus (Hook.)Torr. They were observed erlin (1926) recorded it from ''Montana'' based flom noon to 3 p.m., the ail tempelatwe was on a specimen in the Liebeck Collection (MCZC) about 920 F, and mating took place almost which I have examined and found comparable Immediately as the beetles congregated at the to specimens from Carter Co., 5 mi W Alzada, fOI egoing sites. oak fOlest, 20/VI-31/VIII-89, Felgus Co., SpIing Cr, 9 mi NW Lewistown, 2-VII-88, MTEC Acmaeodera labyrmthlca Fall. Cut trom These specimens well match forms that occur Umbellularia californica (H.&A.)Nutt., Califor- o'\'er much of the eastem U.S. The Fergus Co. nia, Napa Co., Mt. St. Helena, 27-1II-79, CJSC. record is the westernmost known to me for this species and is about 460 lmi distant from any other. Chamberlin also listedA pulchella from Vol. 4, No. 1-4, March-December 1990 75

Wyommg, Medicme Bow [Carbon Co.Ras "var. Actenodes sImI FIsher. Reared from Ulmus mixta Horn", which 'Nas accorded species crassifolia Nutt. branch cut from live tree 20 status by Barr, 1975); however, it is more likely XII-88, adults em. V-89, Texas, Jackson Co., that he was referring to A. Immaculata (q.v.J. below Lake Texana Dam, D J Heffern con, These species belong to a complex, members of RLWE. I am unaware ofany previously record- which OCCUI pIactically tluoughout the U.S. and ed host fOJ tlns beetle. adjacent Canada and Mexico They are greatly in need of study. Agaeocera gentilis gentilis (Horn). CALI· FORNIA, "CIOnese", 24-\11-48, CASCo I could Acmaeodera quadrilJittata Horn Reared not. locat.e t.his on my maps except for Cronise from Yucca schldlgera Roezl ex OrtgIes, CalIfor­ valley, a SIte shown on an old AAA map, and nia, San Bernardino Co., Cedar Canyon Sum which may have been obliterated by the I 15 mit, 5400', 10-XI-74, CSLB. freeway. This is approximately 16 mi SWof Baker, San Bernardino Co. Nearby are the Acmaeodera spkaeralceae Barr. Specimens Gronese dry lakes. closely matching some from the type locality have heen seen from CALIFORNIA, San Agrilus amelanchieri KUllll BRITISH Bernardino Co., Mid Hills, 4600-5000', Sec. 14 COLUMBIA, Creston, 19-VIII-33,Amelanchier & 15, T12N, R14E, 8-V-78; Willow Wash, 2850', alnifolia, CNCI. MANITOBA, Whiteshell Provo Sec. 33, T13N, RUE, 27-Yv'"-80; GIaIlite Mts., Park, Falcon Lake, 7-VI-80, RLVV'E. TIns is a 3300', Sec. 25, T9N, R12E, 30-IV-76, EMUS; new record for Canada. The specimen from Pinyon Hills, Hwy. 138, 28 III 86, GHNC. British GoIUHlbia had been misidentified as A. NEVADA, Clark Co., Spring Mts., Lee Can., 10 arbuti Fisher (q.v.) by Bright and was recorded mi W Hwy. 95, 22-V-83, on flower Sphaeralcea by him (1987) as that species. It also bears sp.*; NEPNberry Mts., Grapevine Can., N ofHwy. determination labels of Frost, J}/ho felt it was 71, 16-VI-81, on flower Encelia farinosa Gray, either arbuti or new (at the time, amelanchieri GHNC. was undescribed). Bright (1987) included A. crataegi Frost from Alberta, probably based on Acmaeodera tuta Horn. Overwintering adUlt Fisher (1928). I find these species difficUlt to taken flOm pupal cell in Dalea schottii Tun., distinguish and questionably distinct, and in California, Riverside Co, Pinyon Crest light ofthe widely recorded western distribution (Hwy.74), 3800', 29-XII-82, FTHC. ofA. amelanchieri, the specimen(s) from Alber­ ta shuuid be Ieexamined. Acmaeoderoides humeralis (Cazier). ARI· ZONA, [La Paz Co.], 18 riii SE Parker, 22 V 70; Agrilus anxius Horn. NEVADA, Washoe Pinal Co.• 1 mi N Florence, 7-IV-72, JJMD, Co., Reno, 27-IV-88, em. from ornamental birch, RLWE. Besides a single specimen recorded Betula sp.*, NVDA, RLWE. According to R. C. from SW Texas, this species has been known Bechtel, Nevada Department of AgricUlture only from southern California. (pers. comm.), this destructive pest (known as the bronze biIchboreI) has notbeen obseI ved in Actenodes calcarata (Chevrolat) LOUISI- Nevada prior to this collection, and it appears to ANA, Cameron Parrish, 12-VI-75, LSUC. be localized in Reno.

Actenodes david; Nelson KANSAS, Wood- Agrilus arbuti Fisher This species was son Co., vic. Piqua, em. V-89 ex limbs Gleditsia recorded from British Columbia by BrIght tl iacanthos L. colI. *1-88, RLV/E. KENTUCKY, (1987) based on a specimen 1 have examined Fayette Co., Lexington, 28-VII-89, tanglefoot on and determined to be A. amelanchieri Knull G. triacanthus, SGWC. (q.v.). I have not seen specimens of A. arbuti 76 Insecta MundI collected north of southwestern Oregon. It locust fIrewood from Logan, VIII-84, RLWE. remains unknown from Canada. According to Vlilford J. Hanson (in litt.), "honey locust" probably refers to one of the thornless AgrilllS allrichalcells Redtenbacher MON- varieties of Gleditsia triacanthos L *, wbich are TANA, Wheatland Co., Deadman's Basin St. popular shade trees in Utah and elsewhere. Ree. AIea, 15-"JI-82, RLWE. DmingVIII-86, I obseI ved consideIable damage from A dilficilis to these trees in Logan. Like Agrilus cercidii Knull. NEW MEXICO, the foregoing species, it probably was imported Hidalgo Co., Peloncillo Mts., Granite Gap, 26 V on nursery stock to Utah. The nearest record 88, on Prosopis sp., WFBC. ed locality is in far eastern Colorado (Cheyenne Wells). Agrilus eliftoni Knull. NEW YORK, Living ston Co., Letchworth St. Park, U8-21)-VI-82, Agrilus gibbicollis Fall. KANSAS, Meade CUIC. Knull (1941) described this species, from Co., 30·"VI-86, D. J. Heffern ColI. MONTANA, a unique male, as without a metatibial mucro; Rosebud Co., Forsyth, 20 VI 56, MTEC. however, I discovered that specimens deter­ mined as A. cltftoni in my collection hore rneta- Agrilusgranulatus liragus Bartel & Blown. tibial mucros, a fact substantiated by G. H. The only record of this species from Oregon, Nelson (in litt.) for those in his collection. AI Austin [Grant Co.], was provided by Barter & New ton (FMNH) examined the holotype and BLOwn (1949). AIlOthel specimen plovides a found thatit too bears these mucros. Therefore, notable range extension: Klamath Co., Upper A. cli{toni vlill not keY in Fisher (1928) to A. Klamath Wildlife Refuge, 26 VI 74, 0800. In arcuatus (Say), as stated by Knull, but to A. the West, this species, like the closely relatedA. geminatus (Say). Although the male genitalia anxius, seems to be rarely collected in the field. should be used for reliable separation, A. cli,f- toni seems to be a larger species utilizing wal- Agrilus illectus FalL A notable northwest- nut, while the hosts fOI A. geminatus ale oaks. wal d I ange extension in Calif01 nia is plOvided The female of A cli,ftoni was described by by a specimen from Humboldt Co, 15 mi E Nelson (1980). Orick on Bald Hills Rd., 3-VII-75, UCDC. The elytIal vittae and sides of pronotum are indis- Agains cyanescens (Ratzeburg) UTAH, tinctly pubescent Salt Lake CIty, 20-V-78, EMOS. Adults were taken on $ymph01 iCaJ POS 01 biculatus Moench, Agrilus latin 011;S Vlaterhouse. NEW MEXI- Missouri, Jefferson Co., 14-V, EGRC, GHNC. CO, Otero Co., High Rolls, 13-VII-76, SDMC. TIlls is an introduced species and was first This species, wIllcli is widespread ill MeXico, reported about 1920, though it may have been has been known in the U.S. only from southern widesplead at that time. Honeysuclde has been AIizona. reported as a host in the IT S; other plants in Europe. Since Symphoricarpos is in the same Agrilus nevadensis Horn. A specimen family, that genus plObably sel yes as a lalVal labeled "S.Eo 01., VIII-12-17, COLL'D BY F.W. host The beetle's occurrence in Utah undoubt- NUNENMACHER" (WFBC) represents the first edly represents a secondary mtroductlOn from record from OREGON. This rarely collected east of the Rocky Mountains, likely in infested species was known definitely only by the type nursery stock from western Nevada until Nelson (1980) re- corded It from Cahforma. No host mformabon 21grilus di{{icilis Gory. UTAH, Cache Co., has been published. Specimens from Meadow Logan, 1976;9-IX-81; 24/27-VIII-83; Salt Lake Valley, Plumas Co., California were beaten only Co , Salt Lake City, I-lX-8S, all on honey locust, from Salzx lasiandra Bentham, I5-VII-78, EMUS. Specimens were reared from honey BKDC; 8-VII-83, RLWE. Several species of Vol. 4, No. 1-4, March-December 1990 77

WIllow occur at thIS localIty and all were beaten water Co., Crazy Mts., N of BIg 'l'Imber, VIII­ extensively. 76, MTEC.

Agrilus oblongus Fisher TEXAS, Bandera Descarpentriesina cyanipes (Say) MON- Co., Bandera by Medina R., 29-III-86, RLWE. TANA, Custer Co., 15 mi E Miles City, 30-VI- 56, MTEC. Agolus pensus Horn MINNESOTA, Lake Co., 10 mi N Isabella, 6-VI-80, RLWE. Descarpentriesina (errea (Melsheimer). SOUTH DAKOTA, Pennington Co., Wasta Agrilusquadriguttatusquadriguttatus Gory Park, cut ex Populus sp *, 24-VIII-66, FMBC in MONTANA, Rosebud Co., ForsYth, 20-VI-56, oIMC. Bright (198'1) mcluded TEXAS m the MTEC. range for this species; however, since I have been unaware of any specific record of it from Agrilus rubroniger Hespenheide, 1979:479 thatstate, substantiation is warranted: Tarrant ( Agriluscupricollis: Fisher, 1928:42 [misidenti Co., Fort Worth/Trinity River, 2 VII 79, RLWE. ficationD. These literature citations are provid­ ed to clarify the diseussion given by Hespen- Dicerca callosa callosa Casey. WYOMING, heide (1979), as he designated rubroniger a Crook Co., Devil's Tower Nat. Mon. ca. 4000', "new name" and his citation ''A. cupricollis 18-VI-83, ESUW. On this specimen the Fisher . ." may be confusing. A misidentifica- widLh/lengLh ratio of tire 7tix anLermal segment tion, despite the fact an actual new species was is .95/1; of the pronotum, 1.70/1 (see Nelson, involved, has no standing in nomenclature; 1975). Nelson (1975) recorded D. callosa {rosa therefore, A. rubroniger should have been Nelson from NW Wyoming, but his closest designated a new species, which would then record for D. c. callosa was from northcentral have been consistent with treatment ofthe type North Dakota. HO\Jlever, Kirk & Balsbaugh materiaL (1975) listed the latter subspecies (det. by Nelson) from Lawrence Co., SouLh Dakota, Buprestis maculativentris Say WYOMING, which is adjacent to Crook Co., Wyoming Crook Co., DeviI's Tower Nat. Mon., ca. 4000', 26-VII-83, ESUW. Dicel en pectOl osn LeConte. MONTANA, Teton Co , Choteau, 28-VI-55, MTEC Buprestls macuhpenms Gory. WISCON­ SIN, Monroe Co., 2.4 mi SW Mather, 6-VII-85, IIippomelas caelatus (LeConte). The follow- RLWE. Although it has been listed from Tex­ ingrepresents the westernmostrecord known to as, I could not find a specific record. I have me: Arizona, Yuma Co., Organ Pipe Nat. Mon., collected it from pine logs near Houston. More 2 mi w campground, 28-X-78, W. B. Warner noteworthy is its occurrence on pinyon in west ColI. Texas, Jeff DaVis Co., W base Sawtooth Mt., (26-29)-VI-79, FTHC. Hippomelas carolinensis Horn. Reared from Dalea emoryi Gray, Arizona, Yuma Co., FooL- Buprestis prospera Casey NEVADA, Was- hills, wood coIl 26-V-80, em (14-30)-VI-80, hoe Co., Reno, 7-X-53, NSDA, RLWE. BKDC.

811prestis slllcicollis (LeConte) WISCON- Rippomelas paci{iclls Chamberlin R A::l rAn SIN, Vilas Co., Sec.B, T41N, R7E, Trout Lake, from roots of Suaeda frutlcosa (L.JF'orsk., CalI­ 6-"/I-64, RUNE. fornia [Kings Co.], Hanford, VIII-61, CISC.

Chrysophana placida (LeConte). MUN Melanophila abies Champlain & KnUlL In TANA, Gallatin Co., Squaw Creek, 31-V; Sweet- the U.S. this species has been recorded only 78 Insecta Mundi from New Hampshire. NEW YORK, Essex Co., 1) Ail other records are for localItIes east of the top Mt. Vlhiteface, 27 "VI 41, CUIC. 100th Meridian; 2) Coquillette was a prominent entomologist who lived in Illinois then moved to Melanophila drllmmondi nicolayi Ohellber- southern CalifOrnia (Essig, 1955), tlilJs speci- ger. Apparently recorded only from Michigan mens collected in Illinois could have been con- and New IIampshiIe, I have seen specimens fused 01 mislabeled, 3) The tlnee otOOI U.S. from NEW YORK, Essex Co , Heart Lai

Ptosima walshii LeConte. AcCOI ding to Literature Cited Nelson (J978) this species was recorded from Arnett, R.II., and G.A. Samuelson. 1986. Cahforma based on a speCimen in the Coquill­ The and spider collections of the em Collection (USNM) flom Los Angeles Co. world. Eo J. Brill, Gainesville, 220pp. He wrote, ". . this state record is questionable and shoUld be verified." I am conVinced that it does not occur in California for three reasons: Vol. 4, No. 1-4, March-December 1990 79

Barr, W.F. 1975. 'I'axonomIc notes and new KnuII, J.N. 1937. New southwestern Buprest­ synonymies of some North American Ac- idae and Cerambycidae with notes. Ohio Jl. maeodera (Coleoptera: Buprestidae). Jour. Sci. 37(5):301-309. Kansas Em Soc 48(3)'411-421 Knull, J.N. 1941. Nine new Coleoptera (Plas- Barter, G.'I.V., and W.J. Brown. 1949. On toeeridae, Buplestidae, and Cerambycidae). the identity of Agolus auxjus Gory and Ohio Jl Sci 41 (5)'381-388 some allied species (Coleoptera: Buprest- idae). Canadian Ent. 81:245-249. Nelson, G.II. 1975. A rerision of the genus Dicerca in North America (Coleoptera' BrIght, D.E. 1987. The msects and arachnids BuprestIdae). Ent. Arb. Mus. Frey 26:8'/­ of Canada. Part 15. The metallic v.'Ood 180. boring beetles ofCanada and Alaska: Coleo- ptera: Buprestidae. Res. Br. Agnc. Canada Nelson, G.H. 1978. A reVIew of the genus Publ. 1810:1 335. Ptosima in North America (Coleoptera: Buprestidae). Coleopts. Bull. 32(4): 327-336. Chamberlin, W.J. 1926 Catalogue of the Buprestidae of North America north of Nelson, G.H. 1980. Nomenclatural changes in Mexico. Corvallis, Oregon, 289 pp. + 1 p. the family Bupres- tidae with descriptions of index. previously unknown sexeS (Coleop tel at Pan-Pacific Ent. 56(2):81-97. Essig, E.o. 1955. Official entomology in California - some comments, historical and Nelson, G.H. 1985. Clarification of the taxo- personal. Quart. Bull. California Dept. nomic status in various genera ofthe family Agric. 44(1):3 16. Buprestidae (Coleoptera). Coleopts. Bull. 39(2):133-146. Fisber, W.S. 1928. A revision of the North American species of buprestid beetles be- Skinner, H. 1903. Notes on Buprestidae longing to the genusAgrilus. U.S. Nat. Mus. (Coleoptera) with descriptions of new spe­ Bull. 145.1-347. cies. Ent. News 14.236-239.

Hespenheide, H.A. 1979. A new name and Walters, G.C., Jr. and C.L. Bellamy. 1990. distribution reeOI ds for &1 ilas (ColeopteI a, Notes on the distribution and biology of Buprestidae). Coleopts. Bull. 33(4): 479-480. certain Buprestidae (Coleoptera): Part IV. Goleopts. Bull. 44(1):113 115. Kirk, V.M., and E.U. Balsbaugh. 1975. A list of the beetles of South Dakota. South DakOta St. Oniv. l-\gric. Expt. Sta. Tech. BulL 42: 1-139. 80 Insecta MundI