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0 20 40 60 80 100 Time (Ma) Papilionidae 108.3 Hesperiidae Pieridae 107.07 Riodinidae - Lycaenidae 101.63 Nymphalidae - 1 Libythea celtis 97.93 70.77 Danaus plexippus 8.3 Danaus chrysippus 82.58 Charaxes jasius Lasiommata maera 14.18 Lasiommata deidamia 11.04 Lasiommata petropolitana 79.5 Lasiommata paramegaera 21.75 Lasiommata megera Pararge xiphia 10.83 Pararge xiphioides 5.93 Pararge aegeria 19.17 Kirinia climene 64.38 7.93 Kirinia roxelana 17.27 Lopinga achine Coenonympha phryne 13.61 Coenonympha oedippus Coenonympha glycerion 17.78 Coenonympha thyrsis 1 3 7.06 Coenonympha pamphilus 10.26 Coenonympha tullia 5.7 Coenonympha amaryllis 46.1 12.28 Coenonympha rhodopensis Coenonympha dorus 11.67 Coenonympha corinna 8.85 Coenonympha hero 5.29 Coenonympha leander Coenonympha gardetta Coenonympha arcania Coenonympha orientalis Proterebia phegea 34.83 Ypthima asterope Melanargia arge 4.74 Melanargia occitanica Melanargia pherusa 14.22 Melanargia ines 44.5 13.52 Melanargia russiae 7.12 Melanargia larissa Melanargia lachesis Melanargia galathea Hipparchia statilinus Hipparchia fatua 8.79 Hipparchia tamadabae 6.33 Hipparchia wyssii Hipparchia tilosi 5.48 Hipparchia bacchus Hipparchia fidia 16.19 Hipparchia gomera 28.67 Hipparchia neomiris 4.99 Hipparchia fagi Hipparchia syriaca Hipparchia autonoe Hipparchia hermione 7.33 Hipparchia mersina Hipparchia miguelensis 35.91 Hipparchia azorina 4.71 Hipparchia maderensis Hipparchia senthes Hipparchia aristaeus Hipparchia neapolitana Hipparchia blachieri Hipparchia volgensis 19.98 Hipparchia semele Hipparchia leighebi Hipparchia pellucida Hipparchia cretica Hipparchia christenseni Hipparchia cypriensis Hipparchia sbordonii Oeneis tarpeia Oeneis bore 11.43 Oeneis ammon Oeneis melissa 6.56 Oeneis jutta Oeneis magna 3.67Oeneis norna 15.86 Oeneis polixenes Oeneis glacialis Arethusana arethusa 12.3 Minois dryas 33.15 10.12 Brintesia circe Satyrus ferula 14.46 Satyrus actaea Satyrus virbius 12.59 Chazara briseis 5.1 Chazara persephone Chazara prieuri Pseudochazara graeca 10.86 Pseudochazara amymone Pseudochazara geyeri 5.32 Pseudochazara anthelea Pseudochazara amalthea Pseudochazara cingovskii Pseudochazara orestes Pseudochazara tisiphone Pseudochazara euxina Pseudochazara williamsi Pseudochazara mercurius Hyponephele huebneri 11.88 Hyponephele lycaon 8.05 Hyponephele lupina Pyronia bathseba 27.77 13.12 Aphantopus hyperantus 15.4 Pyronia tithonus 11.94 Pyronia cecilia 17.07 Maniola cypricola Maniola halicarnassus Maniola telmessia Maniola jurtina Maniola chia Maniola nurag 29.54 Maniola megala Erebia edda 6.66 Erebia fasciata Erebia discoidalis 10.06 Erebia rossii 7.76 Erebia cyclopius Erebia embla Erebia disa 16.6 Erebia epistygne Erebia ottomana 4.99 Erebia callias Erebia rondoui Erebia hispania 13.09 Erebia nivalis Erebia calcarius Erebia tyndarus Erebia arvernensis Erebia cassioides 11.74 Erebia neleus Erebia claudina Erebia christi 7.15 Erebia pharte Erebia epiphron Erebia orientalis 10.25 Erebia jeniseiensis 9.65 Erebia dabanensis Erebia meolans 11.46 Erebia palarica 9.51 Erebia albergana 8.13 Erebia triarius Erebia polaris Erebia medusa 10.76 Erebia aethiops 9.36 Erebia pandrose Erebia sthennyo 8.39 Erebia flavofasciata Erebia gorge 9.99 Erebia pluto Erebia sudetica Erebia melampus 8.27 Erebia manto Erebia eriphyle Erebia euryale 10.44 Erebia ligea Erebia oeme 7.92 Erebia gorgone Erebia aethiopella Erebia mnestra 9.08 Erebia rhodopensis Erebia scipio 6.84 Erebia melas Erebia pronoe Erebia lefebvrei Erebia zapateri Erebia neoridas Erebia montana Erebia stiria Erebia styx 100 8 0 6 0 4 0 2 0 0 Time (Ma).
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