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Die Heteropterenfauna Kretasralf Heckmann, Gerhard ZOBODAT - www.zobodat.at Zoologisch-Botanische Datenbank/Zoological-Botanical Database Digitale Literatur/Digital Literature Zeitschrift/Journal: Carolinea - Beiträge zur naturkundlichen Forschung in Südwestdeutschland Jahr/Year: 2015 Band/Volume: 73 Autor(en)/Author(s): Heckmann Ralf, Strauss Gerhard, Rietschel Siegfried Artikel/Article: Die Heteropterenfauna Kretas 83-130 ©Staatl. Mus. f. Naturkde Karlsruhe & Naturwiss. Ver. Karlsruhe e.V.; download unter www.zobodat.at Carolinea 73 (2015): 83-130, 26 Abb.; Karlsruhe, 15.12.2015 83 Die Heteropterenfauna Kretas* RALF HECKMANN, GERHARD STRAUSS & SIEGFRIED RIETSCHEL Abstract πρώτη φορά για την Ελλάδα: Dicyphus eckerleini, Pilo- A list of 230 species of Heteroptera from the Greek is- phorus simulans, Campylomma simillimum και Hete- land Crete is given. Specimens were collected in the rocapillus perpusillus. Αυτά τα έφτά και 27 ακόμα είδη years 1994, 1995, 2010, 2011 and 2012 during family καταγράφηκαν για πρώτη φορά στο νησί Κρήτη αυξάνοντας holydays of the authors. των αριθμό των Ετερόπτερων σε περίπου 491 – 525. Three species are first records for the fauna of Eu- Μια συνολική λίστα των Ετερόπτερων που βρέθηκε στην rope: Brachycarenus languidus, Eurydema blanda and Κρήτη έχει παρουσιαστεί και η Βιογεωγραφία συζητάει για Singhalesia turcica. Additionally, four species are first τα νέα καταγεγραμμένα είδη. Φωτογραφίες από 23 είδη records for Greece: Dicyphus eckerleini, Pilophorus έχουν παρουσιαστεί, ενώ τα χαρακτηριστικά γνωρίσματα simulans, Campylomma simillimum and Heterocapillus των αρσενικών Heterocapillus nitidus και perpusillus, perpusillus. These seven and 27 species more were re- καθώς και των Lepidargyrus ancorifer και syriacus έχουν corded for the first time for the island of Crete, enhanc- χαρτογραφηθεί δημιουργώντας μια βάση για μια καλύτερη ing the number of species from 491 to 525. A compiled διαφορική διάγνωση. list of all species of Heteroptera found in Crete is given and the biogeography of the new recorded species is Keywords discussed. Photographs of 23 species are presented Heteroptera, Crete, Europe, Greece, differential dia- and the male genitalia of Heterocapillus nitidus, H. per- gnosis, first record, species list, Brachycarenus lan- pusillus, Lepidargyrus ancorifer and L. syriacus are illus- guidus, Campylomma simillimum, Dicyphus eckerleini, trated to provide a basis of better differential diagnosis. Eurydema blanda, Graphosoma creticum, Heterocapil- lus perpusillus, Lepidargyrus syriacus, Pilophorus Kurzfassung simulans, Singhalesia turcica. Eine Liste von 230 Wanzenarten von der griechischen Insel Kreta wird vorgestellt. Die Tiere wurden in den Autoren Jahren 1994, 1995, 2010, 2011 und 2012 während RALF HECKMANN, Schillerstr. 13, D-78467 Konstanz; Ferienaufenthalten der Autoren gesammelt. E-Mail: [email protected]. Drei Arten sind Erstnachweise für die Fauna Euro- GERHARD STRAUSS, Mozartstr. 4, D-88400 Biberach; pas: Brachycarenus languidus, Eurydema blanda E-Mail: [email protected]. und Singhalesia turcica. Zusätzliche vier Arten sind Prof. Dr. SIEGFRIED RIETSCHEL, Waldrebenweg 6, D-76149 Erstnachweise für Griechenland: Dicyphus eckerleini, Karlsruhe; E-Mail: [email protected]. Pilophorus simulans, Campylomma simillimum und Heterocapillus perpusillus. Zusätzlich zu den schon Einleitung genannten sieben Arten wurden für Kreta weitere 27 Den ersten großen Beitrag zur Kenntnis der Arten neu nachgewiesen, womit sich die Artenzahl der Insel von bisher 491 auf 525 erhöht. Es wird eine Ge- Heter opterenfauna der griechischen Insel Kreta samtartenliste der Wanzen Kretas vorgestellt und die gibt REUTER (1891) mit Funddaten zu 101 Ar- Biogeographie der neu nachgewiesenen Arten disku- ten. In der „Bibliographie der Heteropteren der tiert. Photos von 23 Arten werden abgebildet. Zur bes- Balkanhalbinsel“ (JOSIFOV & GÖLLNER-SCHEIDING seren Unterscheidbarkeit von Heterocapillus nitidus 1984) werden 23 Arbeiten zitiert, in denen sich und perpusillus sowie von Lepidargyrus ancorifer und Angaben zu Heteropteren von Kreta finden. Bis syriacus werden deren männliche Genitalien illustriert. auf die Arbeit von JOSIFOV (1968), in der 55 Ar- ten für Kreta zitiert werden, beziehen sich diese Περίληψη Publikationen auf Beschreibungen neuer Arten, 230 Σας Παρουσιαζεται μια λιστα ειδη Ετεροπτερον Gattungsrevisionen und unspezifische Zusam- απο το ελληνικο νησι Κρητη. Τα ζώα συλλέχθηκαν κατά τα έτη 1994, 1995, 2010, 2011 και 2012 κατά τη menstellungen von Wanzenfunden aus dem Mit- διάρκεια διακοπών των συγγραφεις. Τρία από αυτά τα είδη telmeergebiet. Neun dieser Arbeiten stammen καταγράφηκαν πρώτη φορά για την πανίδα της Ευρώπης: aus der Zeit vor 1923, 13 wurden nach 1959 Brachycarenus languidus, Eurydema blanda και Sin- publiziert. Im „Verzeichnis der von der Balkan- ghalesia turcica. Επίσης τέσσερα είδη καταγράφηκαν halbinsel bekannten Heteropterenarten“ (JOSIFOV 1986) werden zusätzliche 9 Publikationen für * Unserem Freund Dr. CHRISTIAN RIEGER zu seinem 70. Kreta zitiert und insgesamt 353 Arten angege- Geburtstag am 13.12.2014 gewidmet. ben. In JOSIFOV & GÖLLNER-SCHEIDING (1984) sowie ©Staatl. Mus. f. Naturkde Karlsruhe & Naturwiss. Ver. Karlsruhe e.V.; download unter www.zobodat.at 84 Carolinea 73 (2015) in JOSIFOV (1986) wird die Publikation von REUTER Die Tiere befinden sich in den jeweiligen Samm- (1891) nicht zitiert. In dieser finden sich Angaben lungen der Autoren. Die von Dr. HARALD ZIEGLER zu weiteren 17 Arten, so dass bis 1986 nach Ab- (verstorben am 13.12. 2008) gesammelten Tiere zug zweier Synonymisierungen aus Kreta insge- befinden sich in der Sammlung G. STRAUSS und samt 368 Arten bekannt geworden sind. wurden auch von diesem bestimmt. Weitere Erkenntnisse zur kretischen Wanzenfau- na geben die Arbeiten von ERNST HEISS und sei- Abkürzungen nen Mitautoren von 1983 bis 1993 (HEISS 1983, HE = RALF HECKMANN (22.5. bis 4.6.2010) 1984 und 1985, HEISS & GÜNTHER 1986, HEISS & RI = SIEGFRIED RIETSCHEL (1.6. bis 15.6.2012) HOPP 1987, HEISS 1988, JOSIFOV & HEISS 1989 und ST = GERHARD STRAUSS (3.5. bis 17.5.2010 und HEISS, GÜNTHER, RIEGER & MALICKY 1993). In JOSI- 4.5. bis 8.5.2011) FOV (1986) werden die ersten vier dieser Arbeiten ZI = HARALD ZIEGLER (14.5. bis 30.5.1994 und zitiert und die Angaben bis auf 4 Arten übernom- 25.5. bis 4.6.1995) men. Aus dem gleichen Zeitraum sind noch die Ar- Fundstellen beiten von CARAPEZZA (1990), RIEGER (1989 & 01 Rodopos 1995a) und MATOCQ (1998) zu nennen, die fünf 35.33.58N 23.44.53E / 28.05.1995 ZI neue Arten für Kreta beschreiben. Durch die Re- 02 Omalos vision der Gattung Odontotarsus kommen zwei 35.20.00N 23.53.54E / 31.05.1995 ZI weitere Arten hinzu (GÖLLNER-SCHEIDING 1990). 03 Souda Diktamos Zuletzt geben die Arbeiten von KMENT & JINDRA 35.29.28N 24.04.27E / 03.06.1995 ZI (2006), KMENT, BRIYJA & JINDRA (2005), MATOCQ 04 Vrysses Madaro, südl. Weiden & Felsfluren (2002 & 2004) und MATOCQ & PLUOT-SIGWALT 35.23.24N 24.04.19E / 02.06.2010 HE (2006a, 2006b & 2012) Informationen über das 05 Moni Arkadi Klosterbezirk Gebäude u. Garten Vorkommen einzelner Arten auf Kreta. Im Zeit- 35.18.38N 24.03.27E / 14.06.2012 RI raum von 1983 bis 2012 sind damit ebenfalls 06 Marmara-Bucht Ardana-Schlucht 368 Arten für Kreta publiziert worden. Mit den 35.11.48N 24.03.27E / 09.06.2012 RI genannten älteren Publikationen und den Anga- 07 Chora Sfakion Imbros, Dorf Ruderal ben zu zusätzlichen 29 Arten in den Bänden der 35.15.10N 24.10.17E / „Faune de France“ (DERJANSCHI & PÉRICART 2005, 24.05.2010-02.06.2010 HE HEISS & PÉRICART 2007, MOULET 1995, PÉRICART 08 Chora Sfakion Imbros-Schlucht 1990 & 1998a, b & c, PUTSHKOV & MOULET 2009, 35.14.58N 24.10.01E / 26.05.1995 ZI RIBES & PAGOLA-CARTE 2013), in JANSSON (1986) 09 Chora Sfakion Imbros-Schlucht, Macchie und PÉRICART (1972) kommen insgesamt 491 Ar- 35.13.00N 24.10.00E / 24.05.2012 HE ten für Kreta zusammen. 20 dieser Arten (4,1 %) 10 Chora Sfakion Strand gelten bisher als endemisch für die Insel. 35.11.41N 24.09.06E / 24.05.2012 HE Eine tabellarische Gesamtartenliste der kre- 11 Georgioupoli tischen Wanzen, nach den verschiedenen Er- 35.22.26N 24.15.44E / 04.06.1995 ZI fassungszeiträumen differenziert, wird in Tab. 1 12 Georgioupoli Kalivaki-Beach Dünen, Flussufer vorgestellt. 35.22.04N 24.15.39E / 27.05.2010 HE 13 Georgioupoli Lake Kournas Ufer Material und Methoden 35.20.12N 24.16.32E / 27.05.2010 HE Um unsere Ergebnisse übersichtlich und platz- 14 Plakias Argoules, 3 km östl. Ruderal sparend darzustellen, sind die Fundorte in der 35.11.34N 24.17.12E / 24.05.2010 HE topographischen Kretakarte von West nach Ost 15 Plakias Rodalkino, 1,5 km westl. Ruderal durchnummeriert. In der Fundortliste werden 35.11.39N 24.18.11E / 01.06.2010 HE dazu die geographischen Koordinaten und das 16 Sellia Umgebung, Kulturland, Wegränder Funddatum vermerkt. In der Artenliste ist dann 35.12.21N 24.22.51E / 04.06.-12.06.2012 RI nur noch Zahl der )) und der (( mit dem jewei- 17 Plakias Plakias, Mouson-Road, Ruderal & ligen Sammler und speziellen Fundumständen Annuellenflur aufgeführt. 35.11.20N 24.24.04E / 22.05.2010 HE Die Methoden umfassten Klopfen, Keschern, 18 Plakias Plakias, Strand-SO Dünen Nachsuchen am Boden und die Untersuchung 35.10.50N 24.24.05E / spezieller Wirtspflanzen. 27.05.-29.05.2010 HE ©Staatl. Mus. f. Naturkde Karlsruhe & Naturwiss. Ver. Karlsruhe e.V.; download unter www.zobodat.at HECKMANN et al.: Die Heteropterenfauna Kretas 85 19 Plakias Ortsstrand, Dünen und Macchie 35.10.48N 24.24.00E / 23.05.2010 HE 20 Spili Kendrohori, Ruderal 35.11.34N 24.24.39E / 30.05.2010 HE 21
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