Biosystematica ISSN: 0973-7871(online)

New Genera and Species of Idiocerine (: Cicadellidae) from India, Sri Lanka and Myanmar

C.A. VIRAKTAMATH Department of Entomology, University of Agricultural Sciences, GKVK, Bangalore 560 065, India

Corresponding author: [email protected]

ABSTRACT - Five genera of Idiocerinae, Burmascopus gen. nov. (type species: Idiocerus fasciolatus Distant) from Myanmar, Ceylonoscopus gen. nov. (type species: Ceylonoscopus quadripunctatus sp. nov.), Lankacerus gen. nov. (type species: Lankacerus rotundus sp. nov.), Neoscopus sp. nov. (type species: Neoscopus ceylonensis sp. nov) all three from Sri Lanka and Nilgiriscopus gen. nov. (type species: Niligiriscopus transversus sp. nov.) from south India are described and the type species are described and illustrated. A key to the known idiocerine genera of the Indian subcontinent is also provided. KEY WORDS - Hemiptera, Cicadellidae, Idiocerinae, new genus, key, India, Sri Lanka, Myanmar

Introduction and Viraktamath (1998) revised the genus from the subcontinent. Maldonado-Capriles (1961, 1970) revised The Idiocerinae is a small subfamily of the the genus Balocha (type species Balocha tricolor family Cicadellidae distributed in all the zoogeographic Distant) and Kameswara Rao & Ramakrishnan (1979) regions. The leafhoppers of the subfamily are recognized described two species from India. Maldonado-Capriles by the following set of characters. Head wider than (1964) in addition to transferring a number of species pronotum, ocelli on face, as far apart from each other earlier placed in Idiocerus to Idioscopus, revised the as they are from adjacent eye; facial sutures extending genus Busonia and described the genera Busoniomimus beyond antennal pits; distance between antennal bases (Maldonado-Capriles, 1977) and Paraidioscopus greater than that between ocelli; pronotum with short, (Maldonado-Capriles, 1964). Viraktamath & Sohi (1994) rounded lateral margins; forewing with large appendix; showed the occurrence of the true species of the genus hind wing with four apical closed cells. Idiocerus from the Sub-Himalayan region. Two genera Distant (1908, 1916) treated species of Idiocerinae with rugose pronotum, Tasnimocerus Ghauri (1975) from the Indian subcontinent in the genera Idiocerus (type species: Tasnimocerus clypeatus Ghauri) and Lewis, Balocha Distant and Busonia Distant. Baker Jogocerus Viraktamath (1979a) (type species: (1915) established the genus Idioscopus (type species: Jogocerus freytagi Viraktamath) were described. Idiocerus clypealis Lethierry) and Viraktamath (1973, Viraktamath and Parvathi (2002) described the genus 1976, 1979a, 1979b) added species to this genus from Periacerus (type species; Idioscopus lalithae the subcontinent. Anufriev (1970) described the genus Viraktamath). At present 10 genera and 43 species are Amritodus (type species: Idiocerus atkinsoni Lethierry) known to occur in the subcontinent.

© Prof. T.C. Narenderan Trust for Taxonomy Biosystematica, 2007, 1(1): 21-30 http://www.tcntrust.org/journal.php 21 22 C.A. VIRAKTAMATH In the Indian subcontinent Idiocerinae breed on trees - Clypellus narrower at base compared to apex; and shrubs. Most species of the genera Idioscopus and aedeagus with well developed dorsal apodeme; Amritodus use species of Anacardiaceae as breeding pygophore with a ventral process ...... hosts (Viraktamath, 1989), whereas those of Balocha ...... Jogocerus Viraktamath use species of Sizygium of the family Myrtaceae 6. Face with frontal suture beyond antennal pit .... 7 (Viraktamath unpublished)) and those of Idiocerus use Cedrus (Coniferae) (Viraktamath & Sohi, 1994). At least - Face without frontal sutures beyond antennal pits; three species of Idioscopus and two species of Amritodus inner apical cell very large...... Busonia Distant are pestiferous on mango (Mangifera indica L.) 7. Forewing with third apical cell stalked (Viraktamath, 1989). Busoniomimus manjunathi (pedunculate); breeding on plants of the genus Viraktamath & Viraktamath causes damage to Garcinia Sizygium ...... Balocha Distant combogia (Mathew et al. 2002). - Forewing without stalked apical cells ...... 8 Examination of leafhoppers in the Natural History Museum, London (BMNH) and U.S. National Museum 8. Hind femoral spinulation 2+1 ...... 9 of Natural History, Smithsonian Institution, Washington, - Hind femoral spinulation 2+0 ..... Idiocrus Lewis D.C. USA (USNM) from south India, Sri Lanka and Myanmar revealed the presence of new genera and 9. Male pygophore with triangular projection on species which are described and illustrated here. ventral margin; aedeagus pustulate...... Busoniamimus Maldonado-Capriles - Male pygophore and aedeagus not as above ... 10 Key to Idiocerine genera of the Indian subcontinent 10. Male style with abbreviated apical apophysis (Fig. 8); aedeagus longer than style ...... 1. Forewing without cross veins before apical cross ...... Burmascopus gen. nov. veins (Figs 36, 40) ...... 2 - Male style with elongate apical apophysis (Fig.14); - Forewing with at least one cross vein before aedeagus shorter than style ...... 11 apical cross veins (Figs 5, 22) ...... 3 11. Male subgenital plate with stouter short setae in 2. Clavus with one vein (Fig. 36); male pygophore and addition to hair-like setae (Fig. 11, 15)...... anal collar without processes (Figs 27) ...... Ceylonoscopus gen. nov...... Lankacerus gen. nov. - Male subgenital plate with only long hair-like setae - Clavus with two veins; male pygophore and anal ...... 12 collar with processes (Fig. 39)...... Neoscopus gen. nov. 12. Male style apical apophysis with expanded processes laterally (Fig. 26) ...... 3. Pronotum with transverse rugae ...... 4 ...... Periacerus Viraktamath & Parvathi - Pronotum without transverse rugae ...... 6 - Male style with apical apophysis without such 4. Clypellus parallel sided (Fig. 64); both anal collar expansions ...... 13 and pygophore without processes (Fig. 55)...... 13. Aedeagal shaft slender, sinuate (Fig. 54) ...... Nilgiriscopus gen. nov...... Paraidioscopus Maldonado-Capriles - Clypellus either narrowed or widened at apex, not - Aedeagus not as above (Figs 24, 51) ...... 14 parallel sided; either anal collar or pygophore or both with processes ...... 5 14. Aedeagus with elongated preatrium, shaft comparatively short (Fig. 24, 25) ...... 5. Clypellus broad at base; aedeagus with poorly ...... Amritodus Anufriev developed dorsal apodeme, pygophore without ventral process ...... Tasnimocerus Ghauri - Aedeagus without preatrium, shaft elongate (Fig. 51); if preatrium long, and shaft short, then apex of

Biosystematica, 2007, 1(1) New Genera and Species of Idiocerine Leafhoppers 23 shaft is drawn out into a slender (Fig. 53), recurved MYANMAR [BMNH, examined]. process and style with a lateral tooth near apex (Fig. Sexually dimorphic. 52) ...... Idioscopus Baker Female. Ochraceous. Vertex with a stripe on either side of median line, brownish; a spot on anterior area on Burmascopus gen. nov. either side, visible in dorsal and ventral aspect, black. Face with brown stripe across ocelli, areas surrounding Type species: Idiocerus fasciolatus Distant 1908:186 each ocellus on outside margin whitish, a spot on either Ochraceous with brown markings, sexually dimorphic. side of inner lower margin, black. Lateral margins of Head broader than pronotum, shagreen. Face frontal suture below antennal pits, clypeus and inner shagreen. Ocelli closer to adjacent eyes than to each margin of genae along side, dark brown. Clypellus and other. Facial sulci reaching ocelli. Antennal ledges adjacent area on lower part of lora dark brown. prominent. Clypellus as broad at base as at apex, but narrowed at midlength. Lora slightly raised above level of genae. Labium extending beyond mesocoxae. A spine on each side of gena below eye. Pronotum and scutellum shagreen. Forewing with two subapical cells, inner subapical cell open behind, four apical cells, outer apical cell longest; clavus with two veins. Hind femoral spinulation 2+1. hind tibial spinulation R1 17-18, R2 6,

R3 7-8. Hind basitarsus with three platellae. Male eighth sternite medially produced, lateral angles lobe-like. Male pygophore with basal anterior fracture, dorsal apodeme on anterior margin well developed with a ventral process. Anal collar with a process. Subgenital plate narrow, long, with marginal long hair-like setae. Style short, with subapical lobe absent, apical apophysis short directed caudally with a row of marginal setae on lateral margin of body and another row near apex. Connective with anterior margin trilobed, lateral margin concave. Aedeagus elongate, slender, preatrium longer than shaft, dorsal apodeme poorly developed, shaft without appendages but with lateral flange near apex, gonopore subapical. Remarks. This genus is peculiar in having the style Figs. 1-10. Burmascopus fasciolatus (Distant). 1. Head and thorax, with the abbreviated apophysis. Burmascopus resembles female type; 2. Face, female; 3. Head and thorax, male type; 4. Face, male; 5. Forewing; 6. Male genital capsule, lateral view; 7. Amritodus in having an anal collar, ventral pygophore Male eighth sternite; 8. Male style and connective; 9. Aedeagus, processes and elongate preatrium to the aedeagus. cephalic view; 10. Aedeagus, lateral view. However, the new genus differs from Amritodus in having the vertex shagreen, pygophore with a basal Pronotum with a curved stripe on either side of median fracture and the style with apical apophysis abbreviated. line, light brown, a large lateral spot and an anterior mesal rather triangular spot dark brown. Scutellum with basal triangles black, area between them dark brown; a round Burmascopus fasciolatus (Distant) comb. nov. discal spot on either side of median line but anterior to (Figs 1-10) median sulcus, dark brown. Forewing with costal margin Idiocerus fasciolatus Distant 1908: 186; Maldonado- in basal half, a broad area on clavus along scutellum, Capriles 1964: 93, figs 10,11. Types 1 Male, 1 Female, claval commissure, apices of outer claval vein, short

Biosystematica, 2007, 1(1) 24 C.A. VIRAKTAMATH segment of vein M stramineous; a large chocolate brown apical cell largest; clavus with two veins. Hind femoral spot on costa and veins dark brown. spinulation 2+1;hind tibial spinulation R117±1, R2 9, R3 8; hind basitarsus with 3 platellae. Basal apodeme at Male. Similar to female but facial markings more base of abdomen well developed. Male eighth sternite bold and dark; black spots on anterior part of head absent. with anterior apodeme, caudal margin convexly produced Male genitalia. Pygophore with ventral process in middle. dorso-ventrally curved not reaching caudal margin. Anal Male pygophore without anterior basal fracture, with collar process, hook-like. Aedeagus widest at midlength, both dorsal and ventral apodemes on anterior margin, shaft abruptly curved dorsally with crenulate lateral tergum short. Anal segment 10 large roof-like, ventrally flanges in cephalic view. produced into spatulate process, 11th segment well Female genitalia. Hind margin of seventh sternite sclerotized, pigmented. Pygophore lobes with a ventral with a median shallow concavity in the middle. process. Subgenital plate with two rows of stouter setae Measurements. Male 4.3 mm long, 1.3 mm wide in basal half, apical half more membranous with hair- across eyes. Female 4.4 mm long, 1.6 mm wide across like setae. Style elongate, arched with a strong ventral eyes. angle. Connective T-shaped, stem longer than combined width of arms. Aedeagus with elongate socle and atrial Material examined. MYANMAR: Type 1 Female, rim, dorsal apodeme poorly developed, shaft cylindrical, ‘Type’ ‘Idiocerus fasciolatus Dist. type’ ‘Myitta’ with lamellate appendages near apex, gonopore subapical ‘(Doherty)’ ‘Distant Coll. 1911-383’. 1 Male, ‘Tenass. on ventral surface. Vall.’ ‘Myitta’ ‘(Doherty)’ ‘Distant Coll. 1911-393’ (BMNH). Female with first pair of gonapophyses with striate sculpture. Second pair of gonapophysis with teeth Remarks. Burmascopus fasciolatus may be confined to distal 0.33. confused with species of Amritodus as it has pronotal black spots in both the male and female but can be easily Remarks. Ceylonoscopus is characteristic in having distinguished from species of Amritodus by the stout setae arranged medially in two rows on the abbreviated style and by the smooth upper part of face subgenital plate in addition to the long hair-like setae. and vertex (striate in species of Amritodus). Ceylonoscopus quadripunctatus sp. nov. Ceylonoscopus gen. nov. (Figs 11-23) Type species: Ceylonoscopus quadripunctatus sp. nov. Uniformly ochraceous brown to light brown with a lateral submarginal spot on pronotum surrounded by pale halo, Ochraceous to light brown with black spots on pronotum black; caudal half of scutellum some times paler. and forewings. Forewing with a spot on cross vein, black. A spot at Robust. Face convex, smooth, vertex rounded to midlength on costal margin dark brown, apical region face, smooth slightly narrower in middle than next to fuscous. eyes. Ocelli closer to lateral eyes than to midline, Male genitalia. Anterior marginal tergal apodemes clypellus with concave lateral margin, slightly wider on pygophore wide apart, sternal apodemes close apically compared to width at base, extending beyond together. Tenth segment collar-like, produced into blade- genal curve. Lora small, slightly raised. Labium like process with caudal prolongation; pygophore lobes extending beyond mesotharacic coxae but not reaching rounded with a short spine-like ventral process not metathoracic coxae. Pronotum declivous anteriorly reaching dorsal angle. Aedeagal shaft in lateral aspect smooth, 2.2 times as wide as long. Scutellum longer curved, with lamellate process directed dorsad, before than pronotum, anterior half declivous caudally, median apex, gonopore subapical. sulcus angular, posterior half slightly raised. Forewing with 2 subapical cells, inner subapical cell closed behind, Female genitalia. Hind margin of seventh sternite outer subapical cell open behind, apical cells 4, outer with a median broad projection caudally, lateral angles

Biosystematica, 2007, 1(1) New Genera and Species of Idiocerine Leafhoppers 25 prominent. Second pair of gonapophyses with seven teeth in distal 0.33 length. Lankacerus gen. nov. Measurements. Male 4.9-5.2 mm long, 1.9-2.0 mm Type species: Lankacerus rotundus sp. nov. wide across eyes. Female 5.7 mm long, 2.2 mm wide across eyes. Vertex of uniform length, upper part of face and vertex shagreen. Ocelli closer to median line than to adjacent Material examined. SRI LANKA: Holotype Male, eye. Clypellus as wide at base as at apex, lateral margins Rat. Dist. Gilimale Lumber Mill, 115 ft (29 m), black narrowed to 0.75, then abruptly widened and rounded light, 20-25.x.1976, G.F. Hevel, R.E. Dietz IV, S. apically. Lora narrow, small. Labium extending slightly Karunaratne, D.W. Balasooriya Collectors (USNM). beyond mesocoxae. A seta on gena below eye. Pronotum Paratypes: 4 Male, 1 Female, data as for holotype twice as wide as median length, slightly declivous (USNM). anteriorly. Scutellum longer than pronotum. Forewing Remarks. Black spots on pronotum and forewing clavus with one vein, subapical cells two, both open distinguish this species from other species of Idiocerinae behind, apical cells four, outer apical cell largest. Hind from the subcontinent. femoral spinulation 2+1; hind tibial spinulation R1 14, R2

5, R3 6; hind basitarsus with three platellae. Male seventh

Figs. 11-19. Ceylonoscopus quadripunctatus sp. nov. 11. Male Figs. 20-26. Species of Idiocerinae: Ceylonoscopus genital capsule, lateral view. 12. Aedeagus and connective, lateral quadripunctatus sp. nov. 20. Head and thorax, male; 21. Face, male. view; 13. Aedeagus, caudal view; 14. Connective and style; 15. 22. Forewing; 23. Abdominal apodemes, male. 24. Aedeagus of Subgenital plate; 16. Male eighth sternite; 17. Female seventh Amritodus atkinsoni, lateralview; 25. Aedeagus caudal view; 26. sternite; 18, 19. Female second pair of gonapophysis. Male style of Periacerus, different views.

Biosystematica, 2007, 1(1) 26 C.A. VIRAKTAMATH sternite without anterior apodeme, caudal margin the clavus has two cells in rotundus compared to only convexly rounded. one in ceylonensis. However, the structure of male genitalia is entirely different in these species. Male pygophore, without anterior basal fracture and anterior apodemes, with rounded caudal lobe. Tenth segment broad, with lateral margin lobe-like but lacking Neoscopus gen. nov. process. Eleventh segment and anal segment well developed. Subgenital plate, of uniform width, longer Type species: Neoscopus ceylonensis sp. nov. than pygophore, with sparse, marginal long hair-like Vertex of uniform length, upper part of face and vertex setae. Style curved, with a basal triangular projection at shagreen. Ocelli closer to adjacent eye than to median base of elongate apical apophysis, ventral margin of line. Clypellus slightly wider basally than at apex, lateral latter smooth. Aedeagus simple, slightly compressed margins narrowed to 0.75 then abruptly widened. Lora laterally, with long preatrium, dorsal apodeme short, of normal size. Labium exceeding mesocaxae. A seta gonopore subapical on ventral margin. on gena below eye. Pronotum twice as wide as median Remarks. Lankacerus resembles Neoscopus gen. length. Scutellum longer than pronotum. Forewing nov. externally but differs in having a single claval vein clavus with two claval veins, subapical cells two, both (two in Neoscopus), lacking both an anal collar process open behind, apical cells four, outer apical cell largest. and pygophore prcess (well developed in Neoscopus), Hind femoral spinulation 2+1, hind tibial spinulation and having the ventral margin of the style apical R1 16-19, R2 8-9, R3 8-9; hind basitarsus with three apophysis smooth (serrated in Neoscopus). It has a platellae. Male seventh sternite with well developed superficial resemblance to Philipposcopus Maldonado anterior apodeme, caudal margin medially convexly Capriles from which it differs in the shape of the rounded. clypellus (parallel sided in Philipposcopus but not so in Male pygophore with well developed anterior tergal Lankacerus). The male pygophore has apodemes on its apodeme, without anterior basal fracture, lobe with a anterior margin and also a caudo-dorsal spine in recurved ventral marginal process. Anal collar well Philipposcopus; both structures that are absent in developed with ventrally directed process. Subgenital Lankacerus. plate of uniform width, longer than pygophore, with marginal long hair-like setae. Style well developed with angulate projection near base of elongate apical Lankacerus rotundus sp. nov. apophysis, ventral margin serrated. Connective T- (Figs 27-36) shaped. Aedeagus narrowed in basal 0.33 then abruptly Ochraceous, a halo surrounding each ocellus, a spot on broadened, with lateral angular projection, gonopore either side of median line on vertex, black; a stripe along subapical on ventral margin. hind margin of pronotum and entire scutellum fuscous. Female eighth sternite produced medially, ninth Basal half of costal margin, claval commissure, apical sternum well developed. First pair of gonapophyses with regions of third apical cell of forewing, fuscous. striate sculpturing. Second pair of gonapophyses with Male genitalia: Pygophore without processes, prominent teeth occupying distal 0.75 length. caudo-dorsal angle infolded. Aedeagus with apex Remarks. Neoscopus shares some of the rounded, ventral margins sinuate, devoid of processes. characters of both Philipposcopus Maldonado-Capriles Measurements. Male 4.8 mm long, 1.5 mm wide and Angusticella Maldonado-Capriles, which it across eyes. resembles closely. The three genera have shagreen vertex and lack a basal fracture of the male pygophore. Material examined. SRI LANKA: Holotype Male, Neoscopus shares the caudal spine of male pygophore Peradeniya, Ceylon, xii.1910 (BMNH). and structure of style with Philipposcopus but differs Remarks. Externally L. rotundatus resembles from it in having a serrated ventral margin of the style Neoscopus ceylonensis sp. nov. in coloration, but has apical apophysis (smooth in Philipposcopus); anal collar black spots on the vertex absent in ceylonensis) and process well developed (absent in Philipposcopus).

Biosystematica, 2007, 1(1) New Genera and Species of Idiocerine Leafhoppers 27 process, recurved anteriorly and spine-like. Anal collar process directed ventrally, anchor-shaped with caudal projection stouter than cephalic and a subapical tooth on cephalic margin of stem. Aedeagus with caudal U- shaped excavation in dorsal aspect, lateral projection with both outer and inner angles dentate. Female genitalia. Hind margin of seventh sternite produced medially with concave margin, eighth sternite broader than seventh sternite. Second pair of gonapophyses with seven prominent teeth, caudo- ventral margin smooth. Measurements. Male 4.5 mm long, 1.6 mm wide across eyes. Female 5.0 mm long, 1.7 mm wide across eyes. Material examined. Holotype Male, SRI LANKA: Mate Dist., Sigiriya, 243m, 13-14.xi.1976, Black light,

Figs. 27-36. Lankacerus rotundus sp. nov. 27. Male genital capsule; 28. Aedeagus and connective, lateral view; 29. Aedeagus, caudal view; 30. Connective; 31. Tenth segment, lateral view; 32. Style, lateral view; 33. Male eighth sternite; 34. Head and thorax, male; 35. Face, male; 36. Forewing.

Neoscopus differs from Angusticella in having the second apical cell of the forewing not pedunculate (pedunculate in Angusticella), pygophore with a caudal spine (absent in Angusticella) and serrated ventral margin of the style apical apophysis (smooth in Angusticella).

Neoscopus ceylonensis sp. nov. (Figs 37-50) Ochraceous. A halo around each ocellus, basal triangles on scutellum, a spot on prosternum behind procoxae, black. Hind margin of pronotum and a large spot on scutellum between basal triangles faintly fuscous, apices Figs. 37-45. Neoscopus ceylonensis sp. nov. 37. Head and thorax, male; 38. face, male; 39. Male genital capsule; 40. Forewing; 41. of forewing margined by fuscous. Claws dark brown. Style; 42, Connectve and aedeagus, cephalic view; 43. Connective Male genitalia. Pygophore with strong ventral and aedeagus, lateral view; 44. Female seventh and eighth sternites; 45. Female ovipositor.

Biosystematica, 2007, 1(1) 28 C.A. VIRAKTAMATH dorsad of ocelli and vertex transversely rugose. Ocelli high on face, closer to adjacent eyes than to median line. Clypellus parallel sided, rectangular, outer margin of gena below eyes slightly inwardly curved, face including eyes wider than long, lora slightly raised above genal surface. Pronotum transversely rugose, 2.6 times as wide as long. Scutellum as long as pronotum. Forewing with two claval veins, two closed anteapical cells. Hind femoral spinulation 2+1; hind tibial spinulation

R1 13, R2 7, R3 8; hind basitarsus with 3 platellae. Male pygophore longer than its depth, without anterior basal fracture, caudo-ventral lobe inwardly curved, devoid of process, dorsal anterior apodeme well developed. Anal collar without process. Subgenital plates longer than pygophore, parallel sided with a few hair- like setae. Style arcuate with ventral distinctly bilobed margin. Aedeagus with well developed preatrium, gonopore terminal, shaft simple with short denticles. Female with striate first pair of gonapophysis. Second pair of gonapophysis with dentate area occupying distal 0.75 length. Remarks: This is the third genus of the subcontinent with transverse rugae on the pronotum, the other two being Tasnimocerus Ghauri (from Simla) and Jogocerus Viraktamath (from Jog Falls). Niligiriscopus differs Figs. 46-54. Species of Idiocerinae. Neoscopus ceylonensis sp. from both Jogocerus and Tasnimocerus in having a nov. 46. Female first pair of gonapophysis; 47-48. Enlarged view at arrow a and b of fig. 46; 49. Female second pair of gonapophysis; parallel sided clypellus, lacking both anal collar process 50. Male eighth sternite. 51. Idioscopus clypelais (Lethierry), and pygophore process and in having only two subapical aedeagus, lateral view. 52. Idioscopus spectabilis Viraktamath, apo- cells in the forewing. Jogocerus differs from physis of style. 53. Idioscopus jayashriae Viraktamath and Nilgiriscopus in having a basal fracture to the Viraktamath, aedeagus, lateral view. 54. Paraidiocerus harrisi Viraktamath, aedeagus, lateral view. pygophore, which is absent in Nilgiriscopus as well as in Tasnimocerus. coll. G.F. Hevel, R.E. Dietz IV, S. Karunaratne, W. Balasooriya, (USNM). Paratype 1 Female, SRI LANKA: Peradeniya, Ceylon, 18.vii.1910 (BMNH). Nilgiriscopus transversus sp. nov. (Figs 55-65) Remarks. This species closely resembles Lankacerus rotundus in external appearance except in Lemon yellow. Pronotum with a small dark brown spot lacking the black spots on the vertex. Its male genitalia on anterior margin on either side of median line (less are however entirely different. prominent compared to male or absent in female). Scutellum with basal triangles, black. Forewing with corial veins and costal margin fuscous. Nilgiriscopus gen. nov. Forewing with radial vein not very prominent, two Type species: Nilgiriscopus transversus sp. nov. subapical cells and 3 apical cells, inner subapical cell open behind. Apodeme at base of abdomen well Lemon yellow with black spots on pronotum and developed. Male eighth sternite with a rather rounded scutellum. Vertex, pronotum and scutellum (not scutum) caudal margin. transversely striated. Head wider than pronotum, region

Biosystematica, 2007, 1(1) New Genera and Species of Idiocerine Leafhoppers 29 Measurements. Male 4.4 mm long, 1.5 mm wide across eyes. Female 5.1 mm long, 1.7 mm wide across eyes. Material examined. Holotype, Male, INDIA: Tamil Nadu, Madras, Coonoor, 5.15, T.V. Campbell (BMNH). Paratypes: 2 Females, data as for holotype (BMNH). Remarks. In one female the lateral margins of vertex are piceous adjacent to the eyes, whereas the other female lacks both these spots as well as the spots on the pronotum. Among the genera having transverse striae on the pronotum this species can be recognized by its clypellus, which is as broad at base as at apex.

Acknowledgments I thank Mr. M.D. Webb (BMNH) and Dr Froeshner (USNM) for their hospitality during my visits and for making available the material for my study on loan. Mr. Webb also gave his critical comments on an earlier draft of this paper.

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