Ministerium für Landwirtschaft, Umwelt und ländliche Räume des Landes Schleswig-Holstein

Die Käfer Schleswig-Holsteins

Rote Liste

Band 2 Herausgeber: Ministerium für Landwirtschaft, Umwelt und ländliche Räume des Landes Schleswig-Holstein (MLUR)

Erarbeitung durch: Landesamt für Landwirtschaft, Umwelt und ländliche Räume des Landes Schleswig-Holstein Hamburger Chaussee 25 24220 Flintbek Tel.: 0 43 47 / 704-0 www.llur.schleswig-holstein.de

Ansprechpartner: Arne Drews (Tel. 0 43 47 / 704-360)

Autoren: Stephan Gürlich Roland Suikat Wolfgang Ziegler

Titelfoto: Macroplea mutica (RL 1), Langklauen-Rohrblattkäfer, 7 mm, Familie Blattkäfer, Unterfamilie Schilfkäfer galt bereits als ausgestorben. Die Art konnte aber in neuerer Zeit in den Seegraswiesen der Orther Reede auf Fehmarn wieder nachgewiesen werden. Die Käferart vollzieht ihren gesamten Lebenszyklus vollständig submers und gehört gleichzeitig zu den ganz wenigen Insektenarten, die im Salzwasser leben können. Dieses einzige an der schleswig-hol- steinischen Ostseeküste bekannte Vorkommen ist in den dortigen Flachwasserzonen durch Wassersport gefährdet. (Foto: R. Suikat)

Herstellung: Pirwitz Druck & Design, Kronshagen

Dezember 2011

ISBN: 978-3-937937-54-0

Schriftenreihe: LLUR SH – Natur - RL 23 Band 2 von 3

Diese Broschüre wurde auf Recyclingpapier hergestellt.

Diese Druckschrift wird im Rahmen der Öffentlichkeitsarbeit der schleswig- holsteinischen Landesregierung heraus- gegeben. Sie darf weder von Parteien noch von Personen, die Wahlwerbung oder Wahlhilfe betreiben, im Wahl- kampf zum Zwecke der Wahlwerbung verwendet werden. Auch ohne zeit- lichen Bezug zu einer bevorstehenden Wahl darf die Druckschrift nicht in einer Weise verwendet werden, die als Partei- nahme der Landesregierung zu Gunsten einzelner Gruppen verstanden werden könnte. Den Parteien ist es gestattet, die Druckschrift zur Unterrichtung ihrer eigenen Mitglieder zu verwenden.

Die Landesregierung im Internet: www.landesregierung.schleswig-holstein.de Inhalt - Band 2

Rote Liste und Checkliste der Käfer Schleswig-Holsteins von FHL Band 2 bis 6 – Carabidae bis Byrrhidae –

Familie Deutscher Name Seite

Carabidae Laufkäfer...... 5 Hygrobiidae Schlammschwimmer...... 20 Haliplidae Wassertreter ...... 20 Noteridae Ruderschwimmer ...... 21 Dytiscidae Schwimmkäfer...... 21 Gyrinidae Taumelkäfer ...... 25 Microsporidae Kugelkäfer...... 26 Hydraenidae Langtaster-Wasserkäfer...... 26 Hydrochidae Rippen-Wasserkäfer...... 27 Spercheidae Tümpelkäfer...... 27 Georissidae Uferschlammkäfer ...... 27 Hydrophilidae Wasserfreunde ...... 27 Histeridae Stutzkäfer...... 31 Silphidae Aaskäfer...... 34 Agyrtidae Dicktaster-Aaskäfer...... 35 Leptinidae Pelzflohkäfer ...... 35 Cholevidae Nestkäfer ...... 35 Colonidae Kolonistenkäfer ...... 37 Leiodidae Trüffelkäfer, Schwammkugelkäfer ...... 37 Scydmaenidae Ameisenkäfer...... 39 Ptiliidae Federflügler...... 41 Staphylinidae Kurzflügler...... 43 Lycidae Rotdeckenkäfer...... 90 Lampyridae Leuchtkäfer, Glühwürmchen...... 90 Cantharidae Weichkäfer...... 90 Drilidae Schneckenhauskäfer...... 92 Malachiidae Malachitkäfer, Zipfelkäfer...... 92 Dasytidae Wollhaarkäfer part...... 93 Phloiophilidae Winter-Rindenkäfer ...... 94 Cleridae Buntkäfer ...... 94 Derodontidae Lärchenkäfer ...... 94 Trogositidae Flachkäfer, Jagdkäfer ...... 95 Peltidae Flachkäfer...... 95 Lophocateridae Flachkäfer...... 95 Lymexylonidae Werftkäfer...... 95

3 Elateridae Schnellkäfer ...... 95 Eucnemidae Kamm-, Dornhalskäfer ...... 98 Lissomidae Binden-Dornhalskäfer...... 98 Throscidae Hüpfkäfer ...... 99 Buprestidae Prachtkäfer...... 99 Clambidae Punktkäfer...... 100 Dascillidae Wiesenkäfer...... 101 Scirtidae Jochkäfer, Sumpffieberkäfer...... 101 Eucinetidae Wiesenkäfer, Faulkäfer ...... 101 Dryopidae Klauenkäfer ...... 101 Elmidae Hakenkäfer...... 102 Heteroceridae Sägekäfer...... 102 Psephenidae Sumpfwiesenkäfer...... 103 Limnichidae Ufer-Pillenkäfer ...... 103 Dermestidae Speckkäfer, Pelzkäfer...... 103 Nosodendridae Saftkäfer...... 104 Byrrhidae Pillenkäfer ...... 104

Fußnoten (Anmerkungen)...... 105

Legende zur Roten Liste und Checkliste der Käfer Schleswig-Holstein

4

Anmerkung Letzter Nachweis Letzter

1957 Verantwortung SH Verantwortung

**

Verantwortung D Verantwortung

in SH in Verbreitungstyp Verbreitungstyp Wertgebende Begleitart Charakteristische Arten

Charakter- art i.e.S.

Risikofaktoren Risiko

-D StD-DI Kü [1] =WLa kurzfristig )

↑ ↓ ↓↓ ( langfristig

Kriterien

Bestand Schutzstatus ., 1798 § s <<< = - I LL I ., 1784) § ss <<< BST L., 1758 § mh = = = ., 1786 § ss (<) = = W (H BST ., 1764 § sh = = = (F., 1787)(F., §§ ex !! 1870 H

Taxon (L., 1758) § ex ., 1822 § ss << ., 1784 § s << = = HMo, CaH, WLa ÜLL L., 1758 § mh < = - D HMo EJ (L., 1758) § ss < L., 1758 § sh = = = BST M D F., 1775F., § s <<< = - I L., 1758 § mh = = = L., 1758 § es <<< = = CaH sö L., 1758 § h = = = L., 1761 § es << = - DF L., 1761 § ss << = - I HMo fDÜ SzW, [2] H L., 1758 § mh < = - I L., 1761 § mh << = - I ! L., 1758 § ss << = - F HMo CaH, ErH ! [3] (= Calosoma maderae auropunctatum (Hbst., 1784)) 1787)) (= Calosoma reticulatum (F.,

Cicindela sylvatica Cicindela hybrida Cicindela maritima Cicindela campestris Calosoma inquisitor Calosoma sycophanta Calosoma auropunctatum Callisthenes reticulatus Carabus coriaceus Carabus violaceus Carabus intricatus Carabus problematicus Carabus granulatus Carabus clatratus Carabus cancellatus Carabus auratus Carabus convexus Carabus nitens Carabus arcensis Carabus nemoralis Carabus hortensis

Grund Kat. Änd. Kat. Grund ↑ ↓

Kat +/- Kat RL 2010 RL 1 3 1-RK 0 0-R 2-RK 2 1 2+R 2 3 1 2-RK * * * * * V+MK V-RK G Carabidae (Laufkäfer)

5

Anmerkung

Letzter Nachweis Letzter

Verantwortung SH Verantwortung

Verantwortung D Verantwortung

in SH in Verbreitungstyp Verbreitungstyp Wertgebende Begleitart Charakteristische Arten

Charakter- art i.e.S.

Risikofaktoren Risiko

-I = kurzfristig )

↓ ↓↓ ( langfristig

Kriterien

Bestand Schutzstatus ., 1833 s = = = KüN [4] ., 1812) mh = = = ., 1812) h = = = ., 1793) mh = = = UFT TRH ., 1863 mh < = - I CaH Mgr, ., 1826 es << ., 1829 ss = = = WLa (D W EJ UFT ., 1854 s = = - I (L., 1758) s << ., 1790 § s < = - FW WLa, haW

Taxon (F., 1779)(F., h = = = ., 1812 mh = = = AUV (L., 1758) h = = = D ELLW (F., 1776)(F., mh = = = F URT (D AB (L., 1758) mh = = = F., 1775F., ss << = = Röh, ÜGr [5] (L., 1758) mh < = - I AYK UFT C (H .L P (F., 1775)(F., sh = = = D (F., 1792)(F., sh = = = (L., 1758) mh = = = AIRM F (L., 1758) s << = = SKü, StU (= Leistus rufescens (F., 1775)) (= Leistus rufescens (F., (= Notiophilus pusillus Wtrh., 1833) (= Notiophilus hypocrita auct. nec Curt., 1829)

Carabus glabratus Cychrus caraboides Leistus rufomarginatus Leistus terminatus Leistus ferrugineus Nebria livida Nebria brevicollis Nebria salina Notiophilus aestuans Notiophilus aquaticus Notiophilus palustris Notiophilus germinyi Notiophilus substriatus Notiophilus rufipes Notiophilus biguttatus Omophron limbatum Blethisa multipunctata Elaphrus uliginosus Elaphrus cupreus Elaphrus riparius Loricera pilicornis

Grund Kat. Änd. Kat. Grund ↓ Kat +/- Kat

+M +M +M RL 2010 RL 2 3+ M 3 1-R 2 * * * * * * * * * * * * * V+MK V+MK V-RK

6 1958 1962 [7,8] 1900 ** *** -- DI DI SbW SzP SzW, StS-I Kü KüN - IFW CaH Mgr BDü = SzW KüN == = StD = = ) ) ) ) ) ) ) ) ) ↓ ↓ ↓ ↓ ↓ ↓ ↓ ↓ ↓ ↓↓ ( ( ( ( ( ( ( ( ( , 1825) es (<) ., 1854 ss < , 1846 ex , 1846 ss < = - D SKü StU ., 1915) ex ., 1781) sh = = = TURM AWS , 1790) mh = = = ., 1784) s = ., 1830) s < AGN D (S , 1843 mh = = = SzB Kü [6] ., 1827) s < = - I StD ., 1784) sh = = = UTZEYS UTZEYS HR W CHRK BST OSSI P P ., 1798) es << ., 1810) ss << = = HMo, NMo ., 1825) mh = = = YLL BST ., 1837 s << ., 1827 mh = = = (A (S ., 1825)., 1825) ss < s = = = Mgr (H ., 1790) s < = = fHo (R R EJ (G (H EJ EJ E (L., 1758) mh < YLL AYK CHAUM ., 1784) mh = = = ., 1837 sh = = = AYK (D ., 1831 ex RFk Hgld S R TEPH (P (D (D (G (P E S BST EJ (F., 1792)(F., s < (F., 1792)(F., mh < D (H (L., 1758) sh = = = (= Clivina contracta (Fourcr., 1785)) (= Clivina contracta (Fourcr., 1915) (= Dyschirius luedersi Wagn., (= Dyschirius similis Petri, 1891) secalis (Payk., 1790)) (= Trechus rivularis (Gyll., 1810)) (= Trechus (= Lasiotrechus discus (Hbst., 1784))

Clivina fossor Clivina collaris Dyschirius thoracicus Dyschirius obscurus Dyschirius neresheimeri Dyschirius nitidus Dyschirius politus Dyschirius impunctipennis Dyschirius chalceus Dyschirius salinus Dyschirius aeneus Dyschirius tristis Dyschirius intermedius Dyschirius laeviusculus Dyschirius angustatus Dyschirius globosus Broscus cephalotes Miscodera arctica Thalassophilus longicornis Epaphius secalis Epaphius rivularis rubens Trechus fulvus Trechus quadristriatus Trechus obtusus Trechus Blemus discus micros Trechoblemus K K K ↓ ↓ ↓ ↓ ↓ +M +M 1 2+MK 2 0-R 2+ M 3 3 3 1 0-R 2 2+MK 0 * * * * * * * * * * V-R V-R V-R V+ M

7 Anmerkung

[10]

Letzter Nachweis Letzter Verantwortung SH Verantwortung

**

Verantwortung D Verantwortung

in SH in Verbreitungstyp Verbreitungstyp Wertgebende Begleitart Charakteristische Arten

Charakter- art i.e.S.

Risikofaktoren Risiko

- D gFG- DI ElbO - gFG ID- IN CaH - DI gFG, kFG - DI StS Elb [9] gFG Kü ! = = = = gFG= ElbO gFG KüN [11] kurzfristig ) ) ) ) ) ) )

↑ ↑ ↑ ↓ ↓ ↓ ↓ ↓ ↓ ↓ ↓↓ ( ( ( ( ( ( ( langfristig

Kriterien

Bestand Schutzstatus (L., 1761) s < , 1821 es << = = Elb ., 1787) s = = = ., 1812 ss << ., 1822) s > ., 1828) sh = = = , 1825 s < = = ., 1802) s < ., 1845 es = = = ., 1827 es << HR RAPIEZ ., 1784) sh = = = LL ., 1797) es << ., 1831) s > A ICOL .-W., 1828).-W., es < D OL TEPH HUNB (I YLL (F., 1792)(F., es = = = K EJ TURM

Taxon (N BST ., 1795) h = = = (S (T ., 1790) ss <<< G ANZ S L L (D ISCH (F., 1792)(F., ex gFG ElbO 1961 (H (P (F (O ., 1810) ss = (O (L., 1761) ss << YLL (G (= Tachys micros (Fisch.-W., 1828)) micros (Fisch.-W., (= Tachys inaequalis auct. nec Kol., 1845) (= Tachys parvulus (Dej., 1831)) (= Tachys bisulcatus (Nicol., 1822)) (= Tachys

Paratachys micros Elaphropus diabrachys Elaphropus parvulus Porotachys bisulcatus nana Tachyta Bembidion striatum Bembidion argenteolum Bembidion velox Bembidion litorale Bembidion pygmaeum Bembidion nigricorne Bembidion lampros Bembidion properans Bembidion punctulatum Bembidion pallidipenne Bembidion bipunctatum Bembidion ruficolle Bembidion dentellum Bembidion obliquum Bembidion varium Grund Kat. Änd. Kat. Grund K

↑ ↑ ↓ ↓ ↓ ↓ * * Kat +/- Kat

+R +MK +M RL 2010 RL 1-R 2+ M 0-R 1-R 2+R 1 2 1 1-R 3+MK * * * * * * * V+ M R-M R- M

8 [12] [15] [16] 1920 1908 * ** ** ** ** - ID StS- KüN I SKü KüO -I sö == gFG === Röh, SzB SzW Feh KüO [18] ) ) ) ) ) ) ) ↓ ↓ ↓ ↓ ↓ ↓ ↓ ↓↓ ( ( ( ( ( ( ( , 1930 s = ÜLLER ., 1806) ss = = = Elb ., 1821 ss > M ERV ONOV ., 1802) es << , 1866 s < = = SKü, StU , 1942 es = = - D ! [14] ., 1835 s < , 1825 sh = = = S (L., 1761) sh = = = ., 1839 ss << = = gFG KüN ., 1812) s > , 1825 es = = = kFG [13] ., 1812) ex ., 1831 mh = = = Kü (D , 1825 ss << = = ErH, HMo ., 1831 es = = = ., 1812) ex (!) 1937 , 1823 sh = = = EJ ESM , 1825 mh = = = ., 1812) ss < ARSH UFT UFT EJ ., 1810 mh = = = ., 1821 s = = = TURM TEPH D AY ., 1827 s < = = SKü KüO ASSATI ROTCH ., 1831 ex ! 1907 ., 1849) es = UFT W ., 1837 ss < D S TURM (F., 1801)(F., ex (D S TURM S F (D (M UFT C R EJ S YLL ERV (F., 1792)(F., mh = = = SzB Kü [17] S (D ., 1914 mh = = = YLL TURM E S D (D G AWS S G ET N (D (= Bembidion nitidulum (Marsh., 1802)) 1930) (= Bembidion andreae polonicum Müller, (= Bembidion tetragrammum illigeri Net., 1914) (= Bembidion pusillum Gyll., 1827)

Bembidion semipunctatum Bembidion ephippium Bembidion prasinum Bembidion monticola Bembidion deletum Bembidion stephensii Bembidion milleri kulti Bembidion lunatum Bembidion bruxellense Bembidion maritimum Bembidion tetracolum Bembidion cruciatum polonicum Bembidion femoratum Bembidion saxatile Bembidion testaceum Bembidion fluviatile Bembidion modestum Bembidion illigeri Bembidion schueppelii Bembidion gilvipes Bembidion fumigatum Bembidion assimile Bembidion clarkii Bembidion normannum Bembidion minimum Bembidion tenellum Bembidion humerale Bembidion quadrimaculatum Bembidion quadripustulatum K K K ↓ * ↓ ↓ +M +M +MK +M 1 3+MK 2+MK 1 3-R 2 2+ M 0 0 0 1-R 2 0-R * * * * * * * * * * * V+ M V+ M V+MK R-M R+ M

9

Anmerkung

Letzter Nachweis Letzter Verantwortung SH Verantwortung

* ** **

Verantwortung D Verantwortung

in SH in Verbreitungstyp Verbreitungstyp Wertgebende Begleitart Charakteristische Arten

Charakter- art i.e.S.

Risikofaktoren Risiko

= KüN =SzW = = StU kurzfristig ) ) ) )

↓ ↓ ↓ ↓ ( ( ( ( langfristig

Kriterien

Bestand Schutzstatus , 1930 ? ? = , 1777) ss = = = ., 1827 mh = = = OEZE ÜLLER ., 1785) mh > = = SzB ., 1796) h = = = (G ., 1822) ss << = = SzW KüN AHLB ., 1810) es << ., 1802) mh = = = KüN S ., 1824 mh > = = Kü [19] ., 1821 s = J. M ANZ ., 1812) s < , 1879 s < ., 1870 mh = = = YLL OURCR ERM (P , 1819) ss << = - D SbW KüN (F., 1779)(F., mh = = = ., 1821) s << = = kFG WAQ, Elb+ [20]

Taxon ERV ., 1797) mh = = = UFT (F ERM (G ARSH (G EYD S G EDEL (L., 1761) mh = = = ERV (D H (F., 1792)(F., mh = = = (M ANZ B EACH (S (P (L (= Bembidion unicolor Chaud., 1850) (= Bembidion lateralis (Leach, 1819)) 1821) (= Bembidion harpaloides Serv., (= Bembidion quinquestriatum Gyll., 1810) (= Asaphidion flavipes part. )

Bembidion doris Bembidion articulatum Bembidion octomaculatum Bembidion obtusum Bembidion biguttatum Bembidion aeneum Bembidion mannerheimii Bembidion neresheimeri Bembidion guttula Bembidion lunulatum Bembidion iricolor Cillenus lateralis Ocys harpaloides Ocys quinquestriatus Asaphidion pallipes Asaphidion flavipes Asaphidion curtum Pogonus luridipennis Pogonus chalceus

Grund Kat. Änd. Kat. Grund ↓ Kat +/- Kat

+M +M +M RL 2010 RL 1 3 1-R 3 2 3+MK * * * * * * * * * * * V+MK D

10 1909 -I HMo -IDBDü sö - I BDü, CaH ==sö HMo==Mgr =Mgrsö =Mgrsö =Mgr =Mgr [21] = [23] ) ) ) ) ) ) ) ) ↑ ↑ ↓ ↓ ↓ ↓ ↓ ↓ ↓ ↓ ↓↓ ↓↓ ( ( ( ( ( ( ( ( ., 1923 ss ? ., 1812) es << ., 1828) es << = - DI StS, SzB KüO [22] UFT CHAUB ., 1783) ss << , 1828 s = = = ., 1812) s < S TEPH (D ., 1812) s < ITT ., 1812) s > (S UFT ., 1768) mh = = = , 1838) es = = = , 1774) sh = = = ., 1812) es < = = gFG Elb UFT .M ., 1844 es = UFT , 1818 s < ., 1812) ss < (F., 1787)(F., h = = = ., 1875 es << = - I WAQ ! (D (L., 1758) es ? ., 1831) s > = = (D EER ILL ., 1797) ex (D UFT ., 1781) sh = = = EER ., 1797) s < UFT EJ TRÖM ., 1829 ss < G (P (H (D HAUD AHLB TURM ETTERSTEDT (D ANZ EJ ANZ E (S (D C S S Z CHRK D (P (P (D (S (L., 1758) h = = = (= Patrobus septentrionis FHL nec Dejean, 1828) (= Ophonus signaticornis (Duft., 1812)) 1774)) (= Pseudoophonus rufipes (De Geer, (= Pseudoophonus griseus (Panz., 1797)) (= Platus calceatus (Duft., 1812)) 1775)) (= Harpalus aeneus (F., (= Harpalus fuliginosus (Duft., 1812)) (= Harpalus winkleri Schaub., 1923) (= Harpalus quadripunctatus Dej., 1829)

Patrobus australis Patrobus assimilis Patrobus atrorufus Perigona nigriceps Anisodactylus binotatus Anisodactylus nemorivagus Anisodactylus poeciloides Diachromus germanus nitens Trichotichnus Harpalus signaticornis Harpalus rufipes Harpalus griseus Harpalus calceatus Harpalus flavescens Harpalus froelichii Harpalus hirtipes Harpalus affinis Harpalus distinguendus Harpalus smaragdinus Harpalus solitaris Harpalus xanthopus winkleri Harpalus latus Harpalus luteicornis Harpalus laevipes K ↓ * * +M +M 3 1-R 2+ M 1-RK 1 3 2 1 3 1 0 3 2 * * * * * * * * V+MK R+ M R-M

11

Anmerkung

Letzter Nachweis Letzter

Verantwortung SH Verantwortung

Verantwortung D Verantwortung

in SH in Verbreitungstyp Verbreitungstyp Wertgebende Begleitart Charakteristische Arten

Charakter- art i.e.S.

Risikofaktoren Risiko

-I Mgr BDü -I- ID BDü KDü, Mgr, - I SKü, StU = === = SKü SKü, StU kurzfristig ) ) ) ) ) ) )

↓ ↓ ↓ ↓ ↓ ↓ ↓ ↓↓ ↓↓ ( ( ( ( ( ( ( langfristig

Kriterien

Bestand Schutzstatus , 1828 ss = ., 1781) mh = = = ., 1828) s < , 1806) ss << ., 1812) s << , 1818 mh = = = ., 1812) ss << = = BDü Mgr, CHRK , 1818 es << , 1818 s < ., 1821 ss << = = KDü, BDü ., 1829 es < ., 1812) h = = = TEPH UFT (S

Taxon , 1837 ss << UFT ., 1812) ss < EJ ., 1812) s < = = ., 1797) h = = = (S ERV (D ANNERHEIM UFT UENSEL TURM D (D S (F., 1792)(F., mh = = = TURM EER TURM S UFT UFT M S (D ANZ (Q S H (D (D (P (= Harpalus rufitarsis (Duft., 1812)) 1801)) (= Harpalus vernalis (F., 1792)) (= Harpalus rufibarbis (F., (= Harpalus punctatulus Duftschmid, 1812) 1837) (= Harpalus melleti Heer, (= Harpalus rupicola Sturm, 1818) (= Harpalus puncticeps (Steph., 1828))

Harpalus rubripes Harpalus rufipalpis Harpalus neglectus Harpalus autumnalis Harpalus picipennis Harpalus pumilus Harpalus servus Harpalus tardus Harpalus modestus Harpalus anxius Harpalus serripes Ophonus rufibarbis Ophonus laticollis Ophonus melletii Ophonus rupicola Ophonus puncticeps Stenolophus teutonus

Grund Kat. Änd. Kat. Grund ↓ ↓

Kat +/- Kat RL 2010 RL 2- M 3 2 2 3+MK 1-R 1 3 1 1-R 2 * * * * * V-MK

12 -- I-I D CaH, ErH ErH BDü = == SzW sö =Mgr KüN ) ) ) ) ) ↓ ↓ ↓ ↓ ↓ ↓↓ ↓↓ ( ( ( ( ( ., 1828 s < = - I ÜGr WAQ, , 1871 mh = = = SzB Kü [6] ., 1846 s << = = CaH, Mgr TEPH ., 1812) es = = = ., 1783) es << , 1825) ss << = - I BDü, Mgr [24] , 1824) h = = = ., 1821) mh = = = ., 1797) sh = = = S , 1825) mh = = = ., 1827) ss << ., 1828) s << ., 1812) mh = = = , 1825) mh = = = ., 1796) mh = = = , 1888 s << = - I HMo, fDÜ ., 1812) mh < ., 1827) h = = = , 1824) sh = = = ., 1784) mh = = = UFT ROTCH HAUD ERV ., 1812) es = = = sö ., 1790) h = = = ANZ YLL UFT C ., 1796) mh = = = C TURM ., 1829) es., 1829) << mh > = = (D , 1912 s = TURM YLL CHALL UFT ANZ TEPH (S BST , 1785) s < = - I CaH, Mgr (L., 1761) mh < TURM (P (G TURM UFT (D AYK EJ EJ (S (S (P (S TURM (G (D (S (S (H ANZ (S (P CHILSKY (D (D (D (S ACZO (P ESKE S (L., 1758) mh = = = L (L (= Dicheirotrichus pubescens (Paykull, 1790)) (= Bradycellus collaris (Payk., 1798)) 1787)) (= Acupalpus dorsalis (F.,

Stenolophus skrimshiranus Stenolophus mixtus Dicheirotrichus gustavii placidus Trichocellus cognatus Trichocellus Bradycellus ruficollis Bradycellus verbasci Bradycellus harpalinus Bradycellus csikii Bradycellus caucasicus Acupalpus elegans Acupalpus flavicollis Acupalpus brunnipes Acupalpus meridianus Acupalpus parvulus Acupalpus dubius Acupalpus luteatus Acupalpus exiguus Anthracus consputus Stomis pumicatus Poecilus punctulatus Poecilus lepidus Poecilus cupreus Poecilus versicolor Pterostichus longicollis Pterostichus strenuus Pterostichus diligens Pterostichus vernalis Pterostichus nigrita K K K ↓ ↓ ↓ ↓ ↓ +M +M +M 1-R 2-R 1 3-MK 3-RK 3+MK 1 2 1-RK * * * * * * * * * * * * * * * V-R V-R V-R R+ M R-MK

13

Anmerkung

Letzter Nachweis Letzter Verantwortung SH Verantwortung

**

Verantwortung D Verantwortung

in SH in Verbreitungstyp Verbreitungstyp Wertgebende Begleitart Charakteristische Arten

Charakter- art i.e.S.

Risikofaktoren Risiko

-I = BDü = ==Mgr, = HMo, NMo, Röh kurzfristig ) ) ) ) )

↓ ↓ ↓ ↓ ↓ ↓↓ ( ( ( ( ( langfristig

Kriterien

Bestand Schutzstatus ., 1852 s < (F., 1787)(F., h = = = ., 1783) mh = = = ETZN L ITT ., 1784) ss << ., 1798) s < = = ., 1798) sh = = = , 1837 mh ? = = ., 1802) s (<) ., 1812) s < .M LL (L., 1758) sh = = = ., 1783) sh = = = LL , 1802) s < = - D KDü Kü BST ., 1850) mh = (I , 1777) h = = = ., 1828) ss < = = ., 1827) h = = = (I ., 1790) s < EER ILL UFT ., 1827) h = = = (H H EJ ARSH

Taxon (P (F., 1775)(F., es = = = [25] (D YLL AYK CHALL (D OEZE ., 1798) mh = = = (M OTSCH (P (G AHLB (S ARSHAM (G LL (I (M (S (M (= Pterostichus nigrita part. ) (= Pterostichus angustatus (Duft., 1812)) (= Calathus ochropterus (Duft., 1812))

Pterostichus rhaeticus Pterostichus anthracinus Pterostichus gracilis Pterostichus minor Pterostichus macer Pterostichus oblongopunctatus Pterostichus quadrifoveolatus Pterostichus niger Pterostichus melanarius Pterostichus aterrimus Pterostichus madidus Abax parallelepipedus Synuchus vivalis Calathus fuscipes Calathus erratus Calathus ambiguus Calathus micropterus Calathus melanocephalus Calathus mollis Calathus cinctus

Grund Kat. Änd. Kat. Grund K

↓ ↓ Kat +/- Kat

+M RL 2010 RL 3 3 1-R 3 3 3+MK * * * * * * * * * * V+ M V-R R- M G

14 1956 - I HMo, ErH ! = ÜGr = gFG, ÜGr =HMo ) ) ↑ ↓ ↓ ↓↓ ( ( , 1777) ss ? , 1994 s (<) = - I WAQ ., 1827) h = = = ., 1783) s << = = ., 1790) ss << = = BDü, CaH, Mgr ., 1763) h = = = (L., 1758) ex syn 1959 [26] ., 1809) sh = = = , 1824) s < , 1935) es << OEZE YLL ., 1828 mh = = = ., 1783) ex BST AYK ., 1812) es <<< = = ONT (L., 1758) mh = = = , 1828 h = = = EJ ANZ (G (G ., 1822) s << = = gFG kFG ., 1784) sh = = = CHMIDT (H ., 1827) ss < = - I gFG, ÜGr Elb (P ., 1797) mh = = = (P (L., 1758) mh = = = D ., 1828 ss < = = gFG Elb+ ., 1812) ss << = = Röh S ., 1827) s << = = HMo (P UFT TURM ., 1809) ss << ELLEN BST CHALL EJ (S ICOL (D ANZ UFT (H YLL AHLB D EJEAN (S (H ANZ (P (L., 1758) s < = = Röh (N D (S (D (G (P (= Calathus piceus (Marsh., 1802)) (= Pristonychus terricola (Hbst., 1783)) (= Olisthopus rotundicollis (Marsham, 1812)) (= Agonum moestum part. ) (= Agonum afrum (Duft., 1812)) (= Agonum pelidnum (Payk., 1798)) (= Platynus dorsalis (Pont., 1763))

Calathus rotundicollis Dolichus halensis Sphodrus leucophthalmus Laemostenus terricola Olisthopus rotundatus Agonum sexpunctatum Agonum ericeti Agonum viridicupreum Agonum gracilipes Agonum marginatum Agonum muelleri Agonum dolens Agonum versutum Agonum viduum Agonum duftschmidi Agonum emarginatum Agonum lugens Agonum micans Agonum scitulum Agonum piceum Agonum gracile Agonum munsteri Agonum fuliginosum Agonum thoreyi Anchomenus dorsalis ↓ ↓ +M +K 1 2+MK 3+ M 3-MK 0-R 1 0-R 2 2 3+ M 1 3+ M 3 * * * * * * * * * * V-MK G-RK

15

Anmerkung Letzter Nachweis Letzter

1909

Verantwortung SH Verantwortung

Verantwortung D Verantwortung

in SH in Verbreitungstyp Verbreitungstyp Wertgebende Begleitart Charakteristische Arten

Charakter- art i.e.S.

Risikofaktoren Risiko

-I = gFG ElbO =Mgr kurzfristig )

↓ ↓↓ ↓↓ ( langfristig

Kriterien

Bestand Schutzstatus ., 1825) es << ., 1784) h = = = NNH ., 1790) sh = = = , 1777) ss <<< = - I BST (M (H , 1825 ex AYK ., 1797) h = = = ., 1837 h = = = ., 1828 mh = = = ., 1831 es = = - I OEZE

Taxon (P EJ ., 1810) s << = = WAQ ., 1810) sh = = = (G ., 1832 es = = = gFG ElbO ! [27] ANZ ., 1810) sh = = = TURM , 1825 es = = = D , 1774) sh = = = TEPH (F., 1796)(F., mh = = = (P S YLL YLL CHDTE S YLL IMM ., 1828 s < S ., 1947 es = = = EER (G (G Z (G TURM EJ (G (F., 1792)(F., mh = S ASS D F (= Agonum livens (Gyll., 1810)) (= Platynus longiventris (Mnnh., 1825)) (= Platynus assimilis (Payk., 1790)) (= Platynus ruficornis (Gze., 1777)) (= Platynus obscurus (Hbst., 1784))

Platynus livens Limodromus longiventris Limodromus assimilis Paranchus albipes Oxypselaphus obscurus Zabrus tenebrioides Amara plebeja Amara tricuspidata Amara strenua Amara kulti Amara similata Amara ovata Amara montivaga Amara nitida Amara convexior Amara communis Amara curta Amara lunicollis Amara aenea

Grund Kat. Änd. Kat. Grund ↓

Kat +/- Kat RL 2010 RL 0 1 3 1-R 1 3 * * * * * * * * * V+MK R+ M R- M R-MK

16 1890 <1900 <1900 -I- I CaH Mgr - ID- D SKü, StU StD SzW, Kü = == CaH ! =BDü = = = =Mgr = = SKü, StU ) ) ) ) ) ↓ ↓ ↓ ↓ ↓ ↓↓ ↓↓ ↓↓ ↓↓ ↓↓ ↓↓ ↓↓ ↓↓ ↓↓ ( ( ( ( ( ., 1790) ex ., 1832 ss < ., 1802) mh = ., 1859 ex AYK ., 1827) ss < ., 1781) es <<< IMM ., 1798) ex (P ARSH Z ., 1850 s (<) = = ., 1812) ss = ., 1783) ss << = = Röh ., 1790) s << OTSCH ., 1812) s < = = AHLB AYK ., 1812) s << = = BDü, Mgr ., 1812) sh = = = (M ., 1797) s < ., 1832 ss << , 1831 ss ? = - I CHRK M (F., 1787)(F., mh << UFT , 1833 s << = = ., 1810) mh = = = ., 1832 es << ., 1790) mh < = = ., 1812) ss << (S AYK (P UFT ., 1810) mh = = = (S HAUD (F., 1792)(F., mh = = = UFT ., 1812) es << ., 1798) mh = = = (D (P UFT ., 1812) s = ., 1797) mh = = = CHALL ANZ ., 1776) mh = = = YLL IMM UFT AYK C (D ILLA ., 1831 mh = = = IMM (D YLL Z ., 1828 ss < (D (P (S V EJEAN Z UFT (P (G AYK UFT (D EJ ÜLL ANZ EJ (G D D (P (D (D (P D (M (= Amara helleri Gredler, 1868) (= Amara helleri Gredler,

Amara eurynota Amara spreta Amara famelica Amara familiaris Amara anthobia Amara lucida Amara tibialis Amara quenseli silvicola Amara ingenua Amara fusca Amara cursitans Amara municipalis Amara bifrons Amara infima Amara praetermissa Amara brunnea Amara fulva Amara consularis Amara majuscula Amara apricaria Amara aulica Amara gebleri Amara convexiuscula Amara equestris Chlaenius tristis Chlaenius nigricornis Chlaenius nitidulus Chlaenius vestitus Chlaenius sulcicollis Chlaenius quadrisulcatus Chlaenius costulatus Oodes helopioides ↓ ↓ ↓ ↓ ↓ +M 3 3+ M 2+MK 3 2-R 0 3 1-R 2-R 1-R 2 1 0 2+ M 2 0 1 1-R * * * * * * * * * V+MK V+MK V-MK G G

17

Anmerkung

Letzter Nachweis Letzter

Verantwortung SH Verantwortung

Verantwortung D Verantwortung

in SH in Verbreitungstyp Verbreitungstyp Wertgebende Begleitart Charakteristische Arten

Charakter- art i.e.S.

Risikofaktoren Risiko

-I- ID Mgr - I Mgr KDü, CaH BDü =Röh ==WAQ ==Röh KDü WAQ gFG, ÜGr = =Mgr [28] kurzfristig ) ) ) ) ) ) ) )

↓ ↓ ↓ ↓ ↓ ↓ ↓ ↓

↓↓ ( ( ( ( ( ( ( ( ↓↓↓ langfristig

Kriterien

Bestand Schutzstatus , 1844 s < = = Röh ., 1819 mh = = = ., 1785) es << ., 1803) s < ., 1813) s < ., 1824) mh > = = ., 1813 ss < , 1925 es = = = sö , 1815 mh = = = AM YLL ON (F., 1775)(F., s < S ., 1837 mh = = = ., 1810 es = = - I Mgr sö ., 1798) h = = = ., 1790) ss << ERM B UEL ., 1812) ss < ., 1797) s < OFFM

Taxon (G OURCR (L., 1758) mh = = = (L., 1758) mh < = = ., 1812) mh = = = P (L., 1767) mh = = = TURM (G YLL AYK (H (F UFT S , 1833 es ? G ANZ HAUD CHRK (P UFT OTSCHULSKY C (D (P (S (L., 1758) ss ? (D M ILLA V (= Badister bipustulatus (F., 1792 nec 1775)) (= Badister bipustulatus (F., (= Badister anomalus (Perris, 1866))

Oodes gracilis Licinus depressus Badister unipustulatus Badister bullatus Badister lacertosus Badister meridionalis Badister sodalis Badister dorsiger Badister dilatatus Badister peltatus Badister collaris Panagaeus cruxmajor Panagaeus bipustulatus Odacantha melanura Masoreus wetterhallii Lebia chlorocephala Lebia cruxminor Demetrias atricapillus Demetrias monostigma Demetrias imperialis Cymindis humeralis Cymindis angularis

Grund Kat. Änd. Kat. Grund ↓ Kat +/- Kat

+M +M RL 2010 RL 2 3 1 1 2 1-R 1 3+MK 3+MK 1 3+MK * * * * * * * * * V+ M R- M

18 1887 ** ** ==Röh BDü, CaH = ) ) ) ↓ ↓ ↓ ( ( ( , 1991 ss < = - D KDü, Mgr Kü ., 1825 mh = = = CIAKY ., 1825 es ? = = Feh EJ , 1777) s > = = Mgr S D EJ ., 1785) mh = = = , 1827) ss < = - I SKü, StU ., 1826) s = , 1871 es = = = ., 1812) es ? D (L., 1758) h = = = , 1834 mh = = = ! EJ ., 1825 es = = = Feh OEZE .-W., 1824.-W., es < = - I BDü, CaH, Mgr ., 1827 es = = = ., 1798) mh = = = ., 1795) h = = = UFT , 1790) mh = = = (L., 1758) es << EJ (D L (L., 1761) h = = = (G LL TURM OURCR (D D ROTCH (I ISCH (F., 1794)(F., es = = = TEPH (O (S RULLE (L., 1758) ex OSSI (F C F S B (R (F., 1787)(F., mh = = = (= Dromius marginellus (Fabricius, 1794)) (= Dromius longiceps Dej., 1826) (= Dromius linearis (Ol., 1795)) 1801)) (= Dromius quadrinotatus (Panzer, (= Dromius quadrisignatus Dej., 1825) (= Dromius sigma (Rossi, 1790)) (= Dromius notatus Steph., 1827) (= Dromius melanocephalus Dej., 1825) 1991) (= Dromius crucifer confusus Sciaky,

Cymindis macularis Cymindis vaporariorum Dromius agilis Dromius angustus Dromius meridionalis Dromius schneideri Dromius fenestratus Dromius quadrimaculatus Paradromius longiceps Paradromius linearis Calodromius spilotus Philorhizus quadrisignatus Philorhizus sigma Philorhizus notatus Philorhizus melanocephalus Philorhizus crucifer confusus Syntomus foveatus Syntomus truncatellus Lionychus quadrillum Microlestes minutulus Microlestes maurus Brachinus crepitans K * +M +M 1 1 2-RK 0 1 2 * * * * * * * * * * V+MK R+ M R- M R-M R- M R+ M

19

Anmerkung Letzter Nachweis Letzter

1908 Verantwortung SH Verantwortung

*

Verantwortung D Verantwortung

in SH in Verbreitungstyp Verbreitungstyp Wertgebende Begleitart Charakteristische Arten

Charakter- art i.e.S.

Risikofaktoren Risiko

-IDokG = =kFG === SzG HMo, dyG= StS obG Kü [29] [30] kurzfristig ) ) ) )

↑ ↓ ↓ ↓ ↓ ↓↓ ↓↓ ( ( ( ( langfristig

Kriterien

Bestand Schutzstatus ., 1802) s = = - I ., 1877 h = = = ., 1783) s < = = , 1875 mh = = = , 1834 es << = - I okG [31] ., 1844 s < = = , 1834 s = = = ., 1805) mh = = = ., 1877 mh = = = ERH ARSH ., 1794) ss < ., 1868 s < , 1774) h = = =

Taxon , 1836 mh < = - I ., 1828 s = = - I ., 1887 ss < G UFT ANNH CHALL TURM (M ., 1837 ss < ., 1822 ex ERH ., 1779)) ANZ R EER (F., 1775)(F., ss <<< TURM S EHNCKE UBÉ (D G (S EIDL M (P TEPH E S A BST HOMS (F., 1787)(F., ss < (G S W S ICOL T (H (F., 1801)(F., ss < N

Hygrobia tarda

Hygrobia hermanni (= Brychius elevatus Peltodytes caesus Haliplus varius Haliplus obliquus Haliplus confinis Haliplus lineatocollis Haliplus ruficollis Haliplus heydeni Haliplus fluviatilis Haliplus wehnckei Haliplus lineolatus Haliplus immaculatus Haliplus apicalis Haliplus fulvicollis Haliplus furcatus Haliplus laminatus Haliplus flavicollis Haliplus fulvus Haliplus variegatus

Grund Kat. Änd. Kat. Grund ↓ Kat +/- Kat

+M RL 2010 RL 0 3+ K 2 2+MK 1 2 2 3 1-R * * * * * * V+ M V+MK V-RK V+ M V+MK Hygrobiidae (Schlammschwimmer) Haliplidae (Wassertreter)

20 [34] 1926 1964 ** ** - I StS- I obG -- I I WAQ HMo, dyG NMo NMo, dyG dyG [33] [35] = =okG ) ) ↑ ↓ ↓ ↓↓ ↓↓ ↓↓ ( ( ., 1783) h = = = ., 1812) s < = - I SzG StS Kü [32] CHALL ., 1828) ex ., 1907 ex , 1808) ss = HR (S , 1836 ss = ., 1828) s = ., 1813) es << = - I okG (A , 1835 h = = = , 1818) es ? ., 1841 mh = = = TEPH ORBR ., 1808) h = = = YLL UBE ., 1783) mh < = - I TEVEN ., 1781) ss < (S V ., 1776) h = = = , 1835 es << ., 1869 mh = = = TEPH A ., 1810) h = = = (G UNZE TURM (S YLL , 1774) mh < = - I HP (S ., 1827) mh = = = CHDTE S ÜLL (K ., 1798) h = = = S (G CHRK S YLL (F., 1781)(F., mh = = = (F., 1787)(F., s = = = CHALL EER TURM (L., 1761) h = = = YLL (M (S S (G (F., 1777)(F., h = = = AYK (S (G (G (L., 1761) mh = = = (P (= Guignotus pusillus (F., 1781)) (= Guignotus pusillus (F., (= Bidessus hamulatus (Gyll., 1813)) (= Coelambus lautus (Schaum, 1843)) (= Hydroporus piceus auct. nec Steph., 1828)

Noterus clavicornis Noterus crassicornis Hyphydrus ovatus Hydroglyphus pusillus Hydroglyphus hamulatus Bidessus unistriatus Bidessus grossepunctatus Hydrovatus cuspidatus Coelambus impressopunctatus Coelambus parallelogrammus Coelambus novemlineatus Coelambus nigrolineatus Coelambus confluens Hygrotus versicolor Hygrotus inaequalis Hygrotus decoratus Hydroporus scalesianus Hydroporus angustatus Hydroporus umbrosus Hydroporus tristis Hydroporus glabriusculus Hydroporus notatus Hydroporus gyllenhalii Hydroporus palustris Hydroporus incognitus Hydroporus striola K * +M +M 2 1 2 1 3+MK 3 0 2+MK 0 * * * * * * * * * * * * * * V+MK V-RK R-M Noteridae (Ruderschwimmer) Dytiscidae (Schwimmkäfer)

21

Anmerkung Letzter Nachweis Letzter

1973

Verantwortung SH Verantwortung

Verantwortung D Verantwortung

in SH in Verbreitungstyp Verbreitungstyp Wertgebende Begleitart Charakteristische Arten

Charakter- art i.e.S.

Risikofaktoren Risiko

-- I I NMo, HMo dyG-I ErH, ÜGr -I QuB QuB [35] == HMo HMo, dyG dyG=QuB [36] [37] kurzfristig ) ) ) ) ) )

↓ ↓ ↓ ↓ ↓ ↓ ↓↓ ( ( ( ( ( ( langfristig

Kriterien

Bestand Schutzstatus , 1868 ex , 1835 ss (<) , 1845 s > = = ., 1822 mh = = = , 1835 s = = - I HMo, dyG , 1835) es << = - I ÜGr , 1776) es << ., 1808) mh = = = (L., 1758) h = = = , 1835 s < , 1836 es = = = QuB ., 1859 s < ., 1871 es = ICOL ., 1835) s < = - I dyG, HMo, NMo YLL TURM CHAUM TURM HP TURM N S S UBE CHAUM ÜLLER ., 1829) ss << = - ID kFG (G S S , 1836 ss << TURM AIRM (L., 1767) ss << = = dyG (S A S

Taxon S F TEPH (F., 1787)(F., mh < = - I (M UBÉ (S TEPH (F., 1787)(F., mh = = = (F., 1781)(F., h = = = (F., 1775)(F., mh << = = (F., 1792)(F., s = A (S (= Hydroporus melanocephalus (Marsh., 1802)) 1787)) (= Hydroporus dorsalis (F.,

Hydroporus erythrocephalus Hydroporus morio Hydroporus obscurus Hydroporus elongatulus Hydroporus rufifrons Hydroporus planus Hydroporus pubescens Hydroporus fuscipennis Hydroporus discretus Hydroporus nigrita Hydroporus memnonius Hydroporus obsoletus Hydroporus melanarius Hydroporus longicornis Hydroporus neglectus Suphrodytes dorsalis Graptodytes granularis Graptodytes bilineatus Graptodytes pictus Laccornis oblongus Porhydrus lineatus Deronectes latus

Grund Kat. Änd. Kat. Grund ↓ ↓ Kat +/- Kat

+M +M RL 2010 RL 1-RK 1 1 1 1-R 3 3 2 3 3+MK 1 0-R * * * * * * V-MK V-RK V+MK R+ M

22 - I ErH, fDÜ obG - I StS SzG Kü [42] = = WLa WAQ== HMo, dyG HMo, dyG [41] sö [40] [29] ) ) ) ) ) ↑ ↓ ↓ ↓ ↓ ↓ ( ( ( ( ( ., 1835) mh = = = (F., 1792)(F., s (<) = - I kFG [38] ., 1824) es < ., 1868 es << ACORD ., 1826) ss (<) = = kFG [39] ., 1802) s ? (F., 1787)(F., s < = = , 1774) mh << = = ., 1794) mh << = = (L AR ., 1796) mh = = = ., 1771) s < = = ., 1798) es < ., 1828) es << , 1836 ss << = = QuB AHLB (F., 1775)(F., mh = .H EER AHLB ANZ ARSH ., 1808) s < = - I kFG ANZ (S (L., 1758) mh = = = AYK ., 1798) s < = - I QuB (P (S UBÉ (L., 1758) h = = = (G ., 1837 s < = = ORST (L., 1767) h = = = TEPH ., 1808) mh = = = (P (F., 1787)(F., mh = = = (M (P YLL R ., 1798) mh < = = A EMM (F (F., 1801)(F., mh = = = AYK (S (L., 1761) mh = = = E ., 1837 ss < = - I dyG sö [40] YLL (G G R (P AYK E (G (P (= Potamonectes canaliculatus (Lacord., 1835)) 1775)) (= Potamonectes depressus (F., (= Potamonectes elegans (Panz., 1794)) (= Oreodytes rivalis (Gyll., 1827)) 1837 (nec !)) (= Agabus nigroaeneus Er., (= Agabus melanocornis Zimm., 1915)

Stictotarsus duodecimpustulatus Nebrioporus canaliculatus Nebrioporus depressus Nebrioporus elegans Oreodytes sanmarkii Scarodytes halensis Laccophilus minutus Laccophilus hyalinus Copelatus haemorrhoidalis Platambus maculatus Agabus subtilis Agabus erichsoni Agabus chalconatus Agabus montanus Agabus neglectus Agabus guttatus Agabus melanarius Agabus bipustulatus Agabus striolatus Agabus sturmii Agabus wasastjernae Agabus fuscipennis Agabus uliginosus Agabus paludosus Agabus nebulosus Agabus conspersus Agabus affinis ↑ ↓ +R +M +M 1 1-R 2 3+MK 1 1 3 3 2+MK 2 * * * * * * * * * * * V-MK V-MK V+ M V+ M D G

23

Anmerkung Letzter Nachweis Letzter

1933 1939 Verantwortung SH Verantwortung

**

Verantwortung D Verantwortung

in SH in Verbreitungstyp Verbreitungstyp Wertgebende Begleitart Charakteristische Arten

Charakter- art i.e.S.

Risikofaktoren Risiko

- I ErH ÜGr, =kFG = okG, euG kurzfristig ) ) )

↓ ↓ ↓ ( ( ( langfristig

Kriterien

Bestand Schutzstatus ., 1835) mh = = = ., 1778) es << ., 1867 mh = = = , 1825) h = = = ., 1776) h = = = ., 1828) s << = = dyG ., 1794) s << = = dyG ., 1870 s << = = HMo, dyG ACORD ., 1867 ex , 1790) s << = = ErH, fDÜ, dyG

Taxon EAY ., 1808) es = = = (L HOMS ARR ., 1837 s < ERGSTR T ., 1795) s < CHRK ., 1808) mh = = = ., 1854 ex HUNB R L HOMS (H YLL (F., 1781)(F., mh = = = (B (S E HOMS RAHM T ., 1808) mh = = = (F., 1792)(F., h = = = (T (M’L YLL (O T (F., 1781)(F., mh = = = , 1774) mh = = = (G (B YLL (G HOMS T EER (G (G (= Ilybius obscurus (Marsh., 1802)) (= Nartus grapei (Gyll., 1808)) (= Rhantus pulverosus (Steph., 1828)) (= Rhantus frontalis (Marsham, 1802))

Agabus unguicularis Agabus congener Agabus clypealis Agabus didymus Agabus undulatus Agabus labiatus Ilybius fenestratus Ilybius ater Ilybius fuliginosus Ilybius subaeneus Ilybius quadriguttatus Ilybius similis Ilybius angustior Ilybius guttiger Ilybius aenescens Nartus grapii Rhantus suturalis Rhantus notatus Rhantus suturellus Rhantus bistriatus

Grund Kat. Änd. Kat. Grund K

↓ *

Kat +/- Kat RL 2010 RL 0 0 3 3 3 3 3+ M 1-R 3+MK * * * * * * * * * * R-M

24 [29] ** -I - I okG [48] = HMo, dyG = HMo, dyG ==== HMo dyG, euG dyG, HMo dyG, euG, okG = HMo [36] [46] dyG [47] [46] ) ) ) ) ) ) ) ↑ ↓ ↓ ↓ ↓ ↓ ↓ ↓ ↓↓ ( ( ( ( ( ( ( , 1774) ss << , 1834) ss (<) = = dyG, ÜGr [44] , 1774) §§, II,IV es <<< ., 1763) ss << = = obG, okG , 1795) s < = = dyG, HMo [43] EER ., 1776 es << ., 1778 s = = = ., 1811 s << ., 1822) mh = = = , 1774) mh = = = TURM (G EER ONT ., 1828 h = = = ., 1771) mh = = = HR ÜLL ., 1808 s << = = HMo dyG [47] , 1865 es = ., 1882 s << = = ÜGr (S OPPE A (G (P , 1836 ss < EER ., 1837 s < = = dyG, HMo, NMo M ICOL (L., 1758) s < = = F., 1801F., s < HP YLL R (H ., 1835 ss < = = euG, kFG [48] ERGSTR ., 1808 mh = = = TEPH S ORST (G E (L., 1758) es = = = LARK G (N B UBÉ ., 1882 es < = - I ÜGr sö L., 1758 mh = = = (L., 1758) h = = = S L., 1758 §§, II,IV es <<< = = dyG, HMo [45] (F C YLL A HP TEPH F., 1798F., es << G S S (L., 1758) mh = = = (L., 1758) ss < (= Hydaticus stagnalis (F., 1787)) (= Hydaticus stagnalis (F., (= Hydaticus laevipennis Thoms., 1867)

Rhantus exsoletus Rhantus latitans Colymbetes fuscus Colymbetes paykulli Colymbetes striatus Hydaticus transversalis Hydaticus modestus Hydaticus aruspex Hydaticus seminiger Graphoderus bilineatus Graphoderus zonatus Graphoderus cinereus Graphoderus austriacus Acilius sulcatus Acilius canaliculatus Dytiscus latissimus Dytiscus semisulcatus Dytiscus dimidiatus Dytiscus marginalis Dytiscus circumcinctus Dytiscus circumflexus Dytiscus lapponicus Cybister lateralimarginalis Gyrinus minutus Gyrinus aeratus Gyrinus marinus Gyrinus distinctus Gyrinus substriatus Gyrinus natator ↓ ↓ ↓ ↓ 3+ M 1 2-R 1-R 3+ K 2+MK 2-R 1 3 3 2 1-R 3+ M 1+ K 1 2+ M * * * * * * * * * V+ M V+ M V+ M R- M Gyrinidae (Taumelkäfer) Gyrinidae (Taumelkäfer)

25 Anmerkung

[42] [51]

Letzter Nachweis Letzter Verantwortung SH Verantwortung

* *

Verantwortung D Verantwortung

in SH in Verbreitungstyp Verbreitungstyp Wertgebende Begleitart Charakteristische Arten

Charakter- art i.e.S.

Risikofaktoren Risiko

== okG, euG = =kFG =kFG =kFG kurzfristig ) ) ) )

↓ ↓ ↓ ↓ ↓↓ ↓↓ ( ( ( ( langfristig

Kriterien

Bestand Schutzstatus , 1838 es = ., 1776) mh < = - I kFG , 1885 s = = - I SzG KüN ., 1829 mh = = = SzG Kü ., 1833 es < ALTL EY ., 1824 es <<< = = kFG ÜLL ., 1830 ss < = = QuB ., 1832 s << = = StS R W , 1855 ss << = = okG [49] ., 1824 ss < = = kFG

Taxon TEPH ., 1824 ss < TRH ., 1837 s < = = , 1927 s = (M ERM S , 1807 s = URT ., 1794 s = = = R W G C ERM E OY ., 1888 es = = = 1967 [50] ENETR ERM CRIBA UG CHS J G G S D.T., 1877D.T., ss < K M O UW K (= Sphaerius acaroides Waltl, 1848) (= Sphaerius acaroides Waltl, (= Hydraena sternalis auct. nec Rey) (= Hydraena bohemica Hrbácek, 1951)

Gyrinus suffriani Gyrinus caspius Gyrinus paykulli Orectochilus villosus Microsporus acaroides Hydraena palustris Hydraena britteni Hydraena riparia Hydraena reyi Hydraena melas Hydraena nigrita Hydraena pygmaea Hydraena pulchella Hydraena testacea Hydraena gracilis Ochthebius dilatatus Ochthebius auriculatus

Grund Kat. Änd. Kat. Grund K

↓ Kat +/- Kat

+M +M RL 2010 RL 1 3 0- M 1 1 2 2 2 3+ M 3+ M * * V+MK V+MK V+ M V-R V+MK Microsporidae (Kugelkäfer) Hydraenidae (Langtaster-Wasserkäfer) Hydraenidae (Langtaster-Wasserkäfer)

26 * ** -I sö - IF StU SGKü =SzG = ) ) ) ↑ ↓ ↓ ↓ ( ( ( ., 1783) s < = - I , 1988 ss (<) = - I NMo, euG [53] ., 1794) mh = = - I ., 1808 es <<< ., 1783) s < = - I ., 1860 es (<) = = Röh , 1794) s << , 1797) mh = = = YLL ., 1844 s << = = ., 1824 mh = = = CHALL ERGE ., 1858 ss < ., 1835 es (<) = = ÜGr ., 1798) mh = = = Kü [52] ., 1802) ss < = - I obG ., 1824 mh = = - I ., 1791) es = = = KüN G HUNB B ., 1805) ss (<) (S CHALL OSSI ULS (T ., 1793) s = = = AYK ERM OTSCH ., 1798 mh = = = ERBST (L., 1758) s = = = , 1881 mh = = = , 1885 mh = = = (S OSC TEPH (R ERM (F., 1792)(F., h = =UFT = M ARSH G (P LL M BST S (H I EY G 1777F., mh = = = EYRHFF (B (D P R (M (H EDEL B (= Limnebius truncatulus Thoms., 1853)

Ochthebius bicolon Ochthebius minimus Ochthebius pusillus Ochthebius marinus Ochthebius viridis Limnebius truncatellus Limnebius papposus Limnebius parvulus Limnebius crinifer Limnebius nitidus Limnebius aluta Limnebius atomus Hydrochus elongatus Hydrochus ignicollis Hydrochus carinatus Hydrochus brevis Hydrochus megaphallus Spercheus emarginatus Georissus crenulatus Helophorus rufipes Helophorus nubilus Helophorus tuberculatus Helophorus grandis Helophorus aquaticus ↓ ↓ +MK +MK +M +M 2+MK 2 3+ M 2+ M 2+MK 1-R 2+ M 2-R 3 3 * * * * * * * * * * * R+ M G D Georissidae (Uferschlammkäfer) Hydrochidae (Rippen-Wasserkäfer) Hydrochidae (Rippen-Wasserkäfer) Hydrophilidae (Wasserfreunde) Spercheidae (Tümpelkäfer)

27

Anmerkung Letzter Nachweis Letzter

1957 Verantwortung SH Verantwortung

**

Verantwortung D Verantwortung

in SH in Verbreitungstyp Verbreitungstyp Wertgebende Begleitart Charakteristische Arten

Charakter- art i.e.S.

Risikofaktoren Risiko

- D kFG [54] == obG = kurzfristig ) ) )

↓ ↓ ↓ ↓↓ ( ( ( langfristig

Kriterien

Bestand Schutzstatus ., 1860 ss < = - ID SzG Kü , 1986 s ? = - I Elb ., 1885 es = = = ., 1868 s < = = , 1881 h = = = ., 1846 ss << (L., 1758) mh < UW ., 1802) ss < ., 1868 h = = = ., 1844 sh = = = ., 1853 ex F., 1781F., mh = = = NGUS K , 1885 es <<< = = OTSCH F., 1787F., s = = = A ULS , 1836 s << = = obG EDEL ., 1793 s < = - I ULS EY HOMS M B ARSH M 1775)(F., h = = = T

Taxon ., 1837 ss < R (L., 1761) h = = = ., 1808) mh = = = Kü HOMS M F., 1792F., mh = = = HOMS BST R F., 1775F., mh = = = T T F., 1792F., (M mh = = = TURM E H YLL S (G (= Helophorus guttulus brevipalpis Bedel, 1881)

Helophorus aequalis Helophorus arvernicus Helophorus brevipalpis Helophorus nanus Helophorus pumilio Helophorus redtenbacheri Helophorus laticollis Helophorus strigifrons Helophorus flavipes Helophorus obscurus Helophorus asperatus Helophorus dorsalis Helophorus fulgidicollis Helophorus granularis Helophorus minutus Helophorus paraminutus Helophorus griseus Coelostoma orbiculare Sphaeridium bipustulatum Sphaeridium marginatum Sphaeridium scarabaeoides Sphaeridium lunatum Cercyon littoralis

Grund Kat. Änd. Kat. Grund K K

↓ ↓ ↓ * ↓

Kat +/- Kat RL 2010 RL 3 1-R 2-R 0-R 1+ K 2+MK 3 2+MK * * * * * * * * * * * V-R V-MK R-M G

28 ** == StD= = Kü = = StS, SzB SzG KüO ) ) ) ) ↑ ↓ ↓ ↓ ↓ ( ( ( ( ↓↓↓ ., 1802) sh = = = ., 1794) es << ., 1829 h = = = , 2009 s > ARSH ANZ ., 1802) s = = = , 1788) es = = - I StS ., 1802) s = = = ., 1851 mh = = = ., 1790) mh = = = , 1807) mh = = = TEPH ., 1884 mh = = = (L., 1758) mh = = = ORST (P ., 1824) es < ., 1829 es << (M (F., 1775)(F., s < (F., 1775)(F., mh = = = S ., 1808) ss < V ., 1802) h = = = ., 1798) mh = = = HP ÜST ., 1853 mh = = = ., 1801) mh = = = ARSH S ARSH K ., 1873 mh = = = (L., 1758) mh = = = TURM ERM YLL TEPH AYK ., 1918 mh = = = LL ., 1837 ss < = = REYSSL HP SCHACH (M ., 1798) mh = = = R (L., 1761) mh = = = (M (I S (S ARSH (L., 1758) sh = = = (G (P (G HP S HOMS (P E (Z (F., 1792)(F., h = = = ., 1801) mh = = = S T (M AYK LL (P (I (= Cercyon lugubris (Ol., 1790)) 1969) (= Cercyon alni Vogt, (= Megasternum boletophagum auct.)

Cercyon depressus Cercyon ustulatus Cercyon obsoletus Cercyon castaneipennis Cercyon impressus Cercyon haemorrhoidalis Cercyon melanocephalus Cercyon marinus Cercyon bifenestratus Cercyon lateralis Cercyon laminatus Cercyon unipunctatus Cercyon quisquilius Cercyon atricapillus Cercyon terminatus Cercyon pygmaeus Cercyon granarius Cercyon tristis Cercyon convexiusculus Cercyon sternalis Cercyon analis Megasternum obscurum Cryptopleurum minutum Cryptopleurum crenatum Cryptopleurum subtile Paracymus aeneus Hydrobius fuscipes Limnoxenus niger Anacaena globulus Anacaena limbata K ↓ ↓ * +M 1 1-R 1 1 1-MK 3-R 3+ M * * * * * * * * * * * * * * * * * * * * * * *

29

Anmerkung Letzter Nachweis Letzter

1950 1950

Verantwortung SH Verantwortung

Verantwortung D Verantwortung

in SH in Verbreitungstyp Verbreitungstyp Wertgebende Begleitart Charakteristische Arten

Charakter- art i.e.S.

Risikofaktoren Risiko

== = obG obG = StS, SzG KüO kurzfristig ) ) ) )

↓ ↓ ↓ ↓ ( ( ( ( langfristig

Kriterien

Bestand Schutzstatus ., 1797) mh = = = , 1884 ss ? ? = ., 1792) s < = = obG ., 1802) s << = = HMo, dyG L BST ., 1863) mh = = = ., 1776) mh = = = ., 1874) ex ., 1849 ss < , 1878) ss < (O ., 1829) h = = = (H , 1829) ss < ., 1855 ex ., 1869 s = = = ARSH ., 1771) es ? = = ÜLL OTT HP (F., 1792)(F., mh = = = ., 1794) mh = = = HOMSON REDL (F., 1775)(F., s = EDEL TEPH (M

Taxon S (R T OTSCH (M (G (S (B ORST (F., 1801)(F., mh = = = (F., 1801)(F., s << = = kFG OTSCH M (L., 1758) h = = = (F TEPHENS HUNB M (F., 1792)(F., s < = = StS, SzG, SzB (S (T (= Laccobius alutaceus Thoms., 1868) (= Laccobius biguttatus Gerh.) (= Enochrus quadripunctatus fuscipennis Thoms., 1884)

Anacaena lutescens Laccobius striatulus Laccobius sinuatus Laccobius obscuratus Laccobius bipunctatus Laccobius minutus Laccobius colon Laccobius gracilis Helochares lividus Helochares obscurus Helochares punctatus Enochrus melanocephalus Enochrus ochropterus Enochrus quadripunctatus Enochrus fuscipennis Enochrus halophilus Enochrus bicolor Enochrus testaceus Enochrus affinis Enochrus coarctatus Cymbiodyta marginella

Grund Kat. Änd. Kat. Grund K K

↓ ↓ * ↓ Kat +/- Kat

+M +M RL 2010 RL 2-R 2 3-MK 0-R 0 2+MK 3 * * * * * * * * * V+ M V+ M V-R R-M D

30 1926 1960 ** -- I I dyG obG, okG obG - D [30] nWZ2 SSKüO KüO =StS =StS = ) ) ) ) ) ↑ ↓ ↓ ↓ ↓ ↓ ( ( ( ( ( ., 1857 es = , 1802) s < = - D ., 1797) mh = = = ., 1799) ex ., 1822) § s << = - I euG ., 1797) mh = = = ., 1825) ss < OLL BST , 1979) s = = = ., 1803) s = = = W ANZ SCHZ ., 1792) es << = - DF nWZ1 (H REUTZ ., 1839 s = = - D , 1842) ss < = - D UWR2, nWZ1 (L., 1758) mh < = = R ., 1808) es << HARP ARSHAM , 1972 ex UWR2, nWZ1 1901 (P , 1888 es << (E ., 1834 es = = = nWZ2 sö (C E BST ., 1792) ex ., 1839 s << R ANAAR (C UBE OFFM TEV R (M E , 1879 ss ? ? = sö (L., 1758) § s << = - I euG (H (K E (A BST AZUR (H (S UWERT (H M (L., 1761) ss << K EWIS L (= Hydrophilus caraboides (L., 1758)) (= Hydrous piceus (L., 1758)) (= Hydrous aterrimus (Eschz., 1822)) 1792)) picipes (F., (= Teretrius (= Chartabraeus globulus (Creutz., 1799)) (= Abraeus globosus (Hoffm., 1803)) (= Acritus atomarius (Aubé, 1842)) (= Myrmetes piceus (Payk., 1809))

Chaetarthria seminulum Hydrochara caraboides Hydrophilus piceus Hydrophilus aterrimus Berosus signaticollis Berosus luridus Berosus spinosus Berosus fulvus fabricii Teretrius Plegaderus saucius Plegaderus vulneratus Plegaderus caesus Plegaderus dissectus Chetabraeus globulus Abraeus granulum Abraeus perpusillus Acritus minutus Acritus nigricornis Acritus komai Acritus homoeopathicus Aeletes atomarius Myrmetes paykulli K ↓ * +M 1-R 1 3 1-RK 2+MK 2 2+MK 2 0 1 2+MK 0 0 * * * * * V+MK R-MK R-M D Histeridae (Stutzkäfer)

31

Anmerkung Letzter Nachweis Letzter

1909 Verantwortung SH Verantwortung

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Verantwortung D Verantwortung

in SH in Verbreitungstyp Verbreitungstyp Wertgebende Begleitart Charakteristische Arten

Charakter- art i.e.S.

Risikofaktoren Risiko

== = SKü= = BDü KDü, =StDKDüKü KüO =nWZ2 = kurzfristig ) ) ) ) ) ) ) )

↓ ↓ ↓ ↓ ↓ ↓ ↓ ↓ ( ( ( ( ( ( ( ( langfristig

Kriterien

Bestand Schutzstatus ., 1830) es < = = StD KDü KüN TEPH , 1792) ex (S , 1862) ss < = = ., 1862) s = = = OSSI ., 1849 s < ., 1792) s < , 1790) mh < ., 1792) s = = = Kü (R ., 1792) s = = = ARS , 1917 mh = = = ., 1798) mh = = = Kü ., 1805) ss < ., 1792) mh < = - D BST ARSEUL UG BST ., 1827) es << (M (L., 1758) s = = = (H ., 1798) es < OTSCH UFT AYK CRIBA (K BST (M UZAT (H YLL

Taxon (P A M (S (D (H ., 1834) s < ., 1809) ss < AYK (G R (P (E ., 1839 es = = = AYK R (F., 1775)(F., ss << = = (P E (= Gnathoncus nanus (Scriba, 1790)) (= Gnathoncus schmidti Rtt., 1894) (= Saprinus cuspidatus Ihss., 1949) (= Baeckmanniolus maritimus (Steph., 1830))

Gnathoncus rotundatus Gnathoncus nannetensis Gnathoncus communis Gnathoncus buyssoni Saprinus rugifer Saprinus planiusculus Saprinus semistriatus Saprinus lautus Saprinus virescens Saprinus immundus Saprinus aeneus Chalcionellus decemstriatus Hypocaccus rugifrons Hypocaccus metallicus Hypocaccus rugiceps Hypocaccus dimidiatus maritimus Dendrophilus punctatus Dendrophilus pygmaeus Carcinops pumilio Paromalus flavicornis

Grund Kat. Änd. Kat. Grund K K K K

↓ * ↓ ↓ ↓ Kat +/- Kat

+M +M RL 2010 RL 3-R 3-MK 3-R 1-R 2-MK 2 3 1 2 0 2 * * * * * * V-R V+MK R-M

32 - D-I [55] sö = = = =nWZ1sö = ) ) ) ) ) ↑ ↓ ↓ ↓ ↓ ↓ ↓↓ ( ( ( ( ( ., 1792) h = = = ., 1781) ss << ., 1792) s < ., 1803)., 1854) mh = ss = < = = = ., 1803) s < = = , 1776) s << ., 1794) es < = = nWZ2 sö BST ., 1813) mh = = = BST ., 1783) ss << = = nWZ1 ARS ., 1854) mh = = = CHRK ., 1824) s << OFFM ., 1792) mh = = = (H (H BST (M ., 1819) h = = = ., 1771) ss << OFFM (S ERM ÜLLER (H HUNB UG ARS (H ERM (H (G (T (K (M , 1776) s > (M ORST (G AHLB ., 1834) es = = = nWZ2 sö (F (S R , 1792) es << (E ULZER (S OSSI (R (= Onthophilus punctatus sensu FHL Bd. 12) 1792)) (= Onthophilus sulcatus (F., 1834)) (= Cylister lineare (Er., (= Cylister angustatum (Hoffm., 1803)) (= Platysoma frontale (Payk., 1798)) (= Paralister stercorarius (Hoffm., 1803)) (= Paralister bipustulatus (Schrk., 1781)) (= Paralister purpurascens (Hbst., 1792)) (= Paralister neglectus (Germ., 1813)) (= Paralister ventralis (Mars., 1854)) (= Paralister carbonarius (Hoffm., 1803)) (= Paralister stigmosus (Mars., 1854)) (= Hister striola Sahlb., 1819) (= Hister terricola Germ., 1824) (= Hister merdarius Hoffm., 1803)

Paromalus parallelepipedus Hololepta plana Onthophilus striatus Onthophilus punctatus Platysoma compressum Platysoma lineare Platysoma ferrugineum Eblisia minor Margarinotus obscurus Margarinotus bipustulatus Margarinotus purpurascens Margarinotus neglectus Margarinotus ventralis Margarinotus carbonarius Margarinotus punctiventer Margarinotus striola Margarinotus terricola Margarinotus merdarius K K ↑ ↓ ↓ ↓ ↓ * ↓ +R 1-R 2-R 2 2+ M 1-R 1-R 3-MK 2-R 3+MK * * * * * * * V-MK R-M

33

Anmerkung Letzter Nachweis Letzter

1950 1860

Verantwortung SH Verantwortung

Verantwortung D Verantwortung

in SH in Verbreitungstyp Verbreitungstyp Wertgebende Begleitart Charakteristische Arten

Charakter- art i.e.S.

Risikofaktoren Risiko

= = == CaH, Mgr = kurzfristig ) ) ) )

↓ ↓ ↓ ↓

( ( ( ( ↓↓↓ langfristig

Kriterien

Bestand Schutzstatus , 1807 s < ., 1781) mh = = = ., 1783 sh = = = ., 1824 mh = = = ., 1834) mh = = = CHRK ., 1825 ss < = = sö ., 1767) mh = = = R BST , 1790 es <<< ETT (S (L., 1758) ex H Z ERSCHEL (E ., 1789) ss << LED L H (F., 1775)(F., mh = = = HARP L., 1758 ex ., 1817) es <<< ., 1837 es <<<

Taxon (G C (O CRIBA R (L., 1758) sh = = = F., 1801F., es << = = (L., 1758) s << = = S E , 1852 ss < = = WAQ ERM (L., 1758) mh = = = (G L., 1758 mh = = = RUQUI T (= Hister cadaverinus Hoffm., 1803) 1834)) (= Grammostethus marginatus (Er.,

Margarinotus brunneus Margarinotus marginatus Hister quadrimaculatus Hister unicolor Hister quadrinotatus Hister helluo Hister bissexstriatus Atholus bimaculatus Atholus duodecimstriatus Atholus corvinus Hetaerius ferrugineus Necrophorus germanicus Necrophorus humator Necrophorus sepultor Necrophorus investigator Necrophorus fossor Necrophorus vespilloides Necrophorus vespillo Necrophorus vestigator Necrodes littoralis

Grund Kat. Änd. Kat. Grund K

↓ ↓ ↓ Kat +/- Kat

+M RL 2010 RL 1 3-R 3+ M 1-R 0 0-R 3+ M 1 2 1 3 * * * * * * * * * Silphidae (Aaskäfer)

34 1945 -ID -ID nWZ2 =sö ) ) ↓ ↓ ↓↓ ( ( , 1777) mh = = = ., 1772) s << = = WLa ., 1845) h = = = , 1815) mh = = = ., 1847) ss = OEZE COP OSH ., 1793) es <<< = = (S HAUD (G PENCE , 1839) ss < = = WLa WAQ, ., 1817 mh = = = ., 1851) s = (R ., 1776) es <<< = = (C R (F., 1775)(F., mh = = = BST (L., 1758) mh = = = (S , 1815) mh = = = (L., 1758) h = = = ÜLL (K ÜLL (H , 1815) h = = = ., 1840 es = = = WAQ TURM M ., 1783 s < = = (F., 1792)(F., ex (M (S (L., 1758) h = = = (L., 1758) s << ., 1798) mh = = = AST PENCE BST PENCE C LL ., 1798 h = = = L., 1758 s << = = (S H (I (S LL I (= Xylodrepa quadripunctata (L., 1761 nec 1758)) 1889) (= Ptomaphagus medius Rey, (= Nargus wilkini (Spence, 1815))

Thanatophilus rugosus Thanatophilus sinuatus Thanatophilus dispar Oiceoptoma thoracica Blitophaga opaca Blitophaga undata Xylodrepa quadrimaculata Silpha carinata Silpha obscura Silpha tristis Phosphuga atrata Agyrtes bicolor Agyrtes castaneus Leptinus testaceus Ptomaphagus varicornis Ptomaphagus subvillosus Ptomaphagus sericatus Nemadus colonoides Nargus velox Nargus wilkinii Nargus anisotomoides Choleva spadicea Choleva agilis K ↓ ↓ +M 2-R 0-R 1 3+ M 1 3 3 3 * * * * * * * * * * * * V-MK V+MK R+ M Leptinidae (Pelzflohkäfer) Cholevidae (Nestkäfer) Agyrtidae (Dicktaster-Aaskäfer) Agyrtidae (Dicktaster-Aaskäfer)

35

Anmerkung

Letzter Nachweis Letzter Verantwortung SH Verantwortung

***

Verantwortung D Verantwortung

in SH in Verbreitungstyp Verbreitungstyp Wertgebende Begleitart Charakteristische Arten

Charakter- art i.e.S.

Risikofaktoren Risiko

= = = = kurzfristig ) ) )

↓ ↓ ↓ ↓↓ ( ( ( langfristig

Kriterien Bestand

es = = = E Schutzstatus , 1915) mh = = = , 1815) h = = = , 1937 ., 1798) s < ANZ PENCE PENCE HSSEN , 1815) h = = = (P ., 1846 ss <<< (S ., 1846 mh = = = (S ., 1900 s = = = ., 1798) mh = = = ., 1922 es = = =

Taxon & I ..1846 es = = = ., 1837 s = = = ., 1922 mh = = = ., 1837 mh = = = ., 1807 mh = = = R , 1915 es = = = ., 1852 s < ., 1918 mh = = = R , 1815) mh = = = ., 1794) s = ELLN AYK ERH R E ELLN PENCE (F., 1781)(F., s = = = E ., 1793) h = = = K ATR RITT ELLN K EANN G (P K L RITT ENICK (S ETRI B J ANZ K P B ANZ (P PENCE (F., 1792)(F., h = = = (P (S ssp. holsatica B

Choleva septentrionis Choleva oblonga Choleva reitteri Choleva glauca Choleva angustata Choleva fagniezi Choleva jeanneli Choleva elongata Sciodrepoides watsoni Sciodrepoides fumatus Catops subfuscus Catops longulus Catops coracinus Catops grandicollis Catops kirbii Catops tristis Catops chrysomeloides Catops neglectus Catops morio Catops nigriclavis Catops fuscus Catops fuliginosus Catops nigricans

Grund Kat. Änd. Kat. Grund K *

↓ ↓ Kat +/- Kat

+M RL 2010 RL 1-R 3 3 * * * * * * * * * * * * * * * V-R R-M R+ M R- M R+ M

36 1936 1920 1922 ** = = = =WLa = =WLa ) ) ↑ ↑ ↓ ↓ ↓↓ ↓↓ ( ( ., 1813) ss < ., 1822) es << , 1807) es = = = YLL ., 1841) ss < = - I Mgr ., 1840) ex ., 1840 mh = = = AHLB ., 1822) ex (G TURM ., 1931 s = = = , 1807) ex (S ., 1807) h = = = ., 1822) ss << = = WLa URT ., 1797 es << (S ÄRK ., 1837 h = = = ., 1841) ss < ., 1829 ss <<< ., 1822) mh = = = CHM ., 1841) s < = - I BDü KDü, StD Kü R AHLB ., 1837 ss << = = ., 1845) s = = = ROG ., 1852 es = = = (C M ., 1858 ss < = = WLa (S R E ATR R BST R K (S R TURM ., 1837 mh = = = E CHM AHLB K (L H TEPH ., 1845) es = = = KDü KüN CHM (E AHLB K R (S R S (S (S E (F., 1792)(F., h = = = (S ., 1850 s = (S (E R K (= Catops nigritus Er., 1837) (= Catops nigritus Er., (= Catops westi Krog., 1931) (= Hydnobius strigosus Schm., 1841) (= Leiodes anglica (Rye, 1873))

Catops picipes Apocatops nigritus Fissocatops westi Colon latum Colon fuscicorne Colon angulare Colon dentipes Colon barnevillei Colon brunneum Colon appendiculatum Colon calcaratum Colon serripes Colon viennense Colon bidentatum maerkelii Triarthron Hydnobius latifrons Hydnobius punctatus Hydnobius multistriatus Leiodes ciliaris Leiodes furva Leiodes pallens Leiodes rubiginosa Leiodes triepkii Leiodes rugosa Leiodes oblonga K K ↑ ↓ ↓ * * +M +R +M 2 2 0 3 0 2 2- M 0 3+MK 3+ K 1-R 2 1-R * * * * * * * * * R+ K R-M R-M Leiodidae (Trüffelkäfer, Schwammkugelkäfer) Leiodidae (Trüffelkäfer, Colonidae (Kolonistenkäfer)

37

Anmerkung Letzter Nachweis Letzter

1909 1912 Verantwortung SH Verantwortung

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Verantwortung D Verantwortung

in SH in Verbreitungstyp Verbreitungstyp Wertgebende Begleitart Charakteristische Arten

Charakter- art i.e.S.

Risikofaktoren Risiko

-I = = kurzfristig ) )

↓ ↓ ↓↓ ( ( langfristig

Kriterien

Bestand Schutzstatus ., 1841) es = = = , 1807) es << = = ., 1798) h = = = ., 1841) es = = = ., 1898 ss =

Taxon CHM ., 1802) h = = = ., 1855) es = = = ., 1841) ss = , 1807) es = = = ., 1841) ex , 1872 es = = = SKü ., 1841) mh = = = AYK ., 1852) es = = = (S ., 1797) ex CHM R TURM (F., 1787)(F., s = = = YE (P AHLB ., 1884 ss < CHM R (S (K AIRM S (S ARSH CHM CHM TT TURM ANZ (S (F R (S (S (M (P (S (= Leiodes curta (Fairm.)) (= Leiodes calcarata (Er.)) (= Leiodes distinguenda part. ) (= Leiodes clavicornis (Rye)) (= Leiodes nigrita sensu FHL Bd3 ) (= Leiodes ovalis (Schm.)) (= Leiodes scita sensu FHL nec Erichson )

Leiodes silesiaca Leiodes lucens Leiodes longipes Leiodes flavescens Leiodes polita Leiodes lunicollis Leiodes picea Leiodes obesa Leiodes rufipennis Leiodes bicolor Leiodes brunnea Leiodes ruficollis Leiodes ferruginea Leiodes gallica Leiodes nigrita Leiodes badia

Grund Kat. Änd. Kat. Grund ↓

Kat +/- Kat RL 2010 RL 2+ M 0-R 3 2+MK 0 * * * * R+ M R- M R+ M R+ M R- M R+ M G

38 1927 - ID CaH, Mgr = = ) ↑ ↓ ↓↓ ( , 1855) es = = - D nWZ1 ., 1841 es << , 1807 s << ., 1833) ss < = = WLa sö ., 1827) mh = = = ., 1845 es = = - I BDü, Mgr Elb CHM R , 1807 ss < = - D WLa; nWZ1 ., 1792) mh = = = S E ., 1863 s = = = YLL ., 1866 ss < = = ., 1813) ex ., 1792) s < = = TURM ., 1845 mh = = = ., 1813) s ? AHLB , 1817) h = = = , 1807) h = = = R AIRMAIRE S HP BST (G RIS ., 1829 ss < = = (L., 1758) mh = = = ., 1810 es <<< = = nWZ1 sö (S E ., 1798) mh = = = YLL BST (F., 1792)(F., mh = = = (F S YLL ., 1812) mh = = = ., 1840) mh = = = TURM B ., 1845 mh = = = ., 1794) ss << = = nWZ2 sö (H (F., 1792)(F., mh = = = ECK R S (G (H YLL (G HR AYK TURM E UG TEPH URT (B (F., 1792)(F., mh = = = G (P (A S (S (K (C (= Leiodes parvula (Sahlberg)) 1812)) (= Cyrtusa minuta (Ahr., (= Cyrtusa pauxilla (Schm., 1841)) (= Agathidium sphaerulum Rtt., 1898)

Leiodes gyllenhali Colenis immunda Agaricophagus cephalotes Cyrtusa subtestacea Liocyrtusa minuta Liocyrtusa vittata Anisotoma humeralis Anisotoma axillaris Anisotoma castanea Anisotoma glabra Anisotoma orbicularis Liodopria serricornis Amphicyllis globus Amphicyllis globiformis Agathidium marginatum Agathidium haemorrhoum Agathidium varians Agathidium convexum Agathidium rotundatum Agathidium confusum Agathidium nigrinum Agathidium nigripenne Agathidium atrum Agathidium seminulum Agathidium laevigatum Agathidium badium Euthiconus conicicollis ↓ * 1 1+ K 3 1- M 2- M 2-R 3 2+MK 0 3 1 * * * * * * * * * * * * * * * V-MK Scydmaenidae (Ameisenkäfer)

39 Anmerkung

[56]

Letzter Nachweis Letzter Verantwortung SH Verantwortung

**

Verantwortung D Verantwortung

in SH in Verbreitungstyp Verbreitungstyp Wertgebende Begleitart Charakteristische Arten

Charakter- art i.e.S.

Risikofaktoren Risiko

-D -D =sö kurzfristig )

↑ ↓ ↓↓ ( langfristig

Kriterien

Bestand Schutzstatus , 1822 es = = = WLa , 1822) h = = = , 1822) h = = = , 1822) s = = = , 1866) ss ? , 1844) s = = = , 1822) es = = = UNZE ., 1845) s < = - D UNZE , 1822) h = = = ., 1845) es = = - D .K , 1844) s < = - D WLa UNZE UNZE ., 1899 es = = = .K ., 1830 mh = = = UNZE .K .K ÜLL , 1884) es = = = CaH ., 1861) s = = - D UNZE HAUD ., 1925 s = HAUD CHAUM ÜLL , 1920 es = = = .K , 1825) s < = - D nWZ2 M ÜLL .K (C TEPH ÜLL (S (C ., 1806) s < = - D CHAUFUSS ., 1879 es < CHAUM ANGLB ULS (M ACH S ÜLL

Taxon TT (M G (S OKAY (S ÜLL (M ENNY OWLER (M ATR M R L (M ., 1861 s = = - D (F (L (M ., 1813) es = = = WLa sö (D ULS YLL M (G (= Stenichnus harwoodianus Will., 1927) (= Euconnus unicus Franz, 1957)

Euthia plicata Euthia linearis Euthia scydmaenoides Cephennium thoracicum Cephennium gallicum Neuraphes elongatulus Neuraphes angulatus Neuraphes carinatus Neuraphes ruthenus Neuraphes talparum Neuraphes plicicollis Scydmoraphes helvolus Scydmoraphes minutus Stenichnus scutellaris Stenichnus godarti Stenichnus collaris Stenichnus bicolor Stenichnus poweri Microscydmus nanus Microscydmus minimus Euconnus pubicollis Euconnus campestris Grund Kat. Änd. Kat. Grund K K

↓ * * * Kat +/- Kat

+M RL 2010 RL 3-RK 1-R 3-RK 1-RK 3 3 * * * * * * * V+MK V+MK V+MK R-M R- M R- M R-M R- M R- M

40 1945 1916 1938 WAQ, WLa WAQ, -D StD Kü = ↑ ↓↓ , 1822) es << = = ErH, HMo, , 1822) ss << = = , 1822 mh = = = , 1822) ss < = = WLa , 1822 s = UNZE ., 1844) s < = = UNZE ., 1869 mh = = = UNZE ., 1802) es = = = ., 1845) s = = = .K .K ., 1851 mh = = = ., 1844) ss < = = nWZ2 .K ., 1855) es << ., 1827) s < = - D Kü ., 1923) es = = - D ., 1855 es = = = UNZE LLIB UNZE R ANK K ., 1808) sh = = = ÜLL ., 1845) ss = = = ., 1868) ss < = - D UWR2, nWZ1 sö ARSH ., 1845 mh = = = ÜLL .K .K YLL (A , 1841) sh = = = ÜLL ., 1845 mh = = = W R R ALID ACH ., 1844) mh = = = HAUD ., 1845 s < = - D nWZ2 R ., 1844) ex ., 1798) mh = = = ., 1861) ex ., 1971 ss = = = ANNH E (G M (M (M YLL R (M E (E HOMS ÜLL ÜLL (H (C EER LL (M ES ATTH E T (M (I LLIB (G LLIB M B (H ATTH (M (M (A (A ., 1845 ex (M R E (= Ptenidium myrmecophilum (Motsch., 1845)) (= Ptilium minutissimum auct. nec (Ljungh, 1804)) (= Ptilium exaratum FHL )

Euconnus claviger Euconnus pragensis Euconnus maeklinii Euconnus denticornis Euconnus rutilipennis Euconnus hirticollis Euconnus fimetarius Scydmaenus tarsatus Scydmaenus rufus Nossidium pilosellum Ptenidium gressneri Ptenidium laevigatum Ptenidium turgidum Ptenidium intermedium Ptenidium fuscicorne Ptenidium punctatum Ptenidium formicetorum Ptenidium pusillum Ptenidium nitidum Actidium coarctatum Oligella foveolata Oligella intermedia Oligella insignis Micridium halidaii Ptilium exaratum Ptilium affine Ptilium myrmecophilum Ptilium canaliculatum K K ↑ ↓ * * * +R +K 2 2-RK 1 3+ M 0 0-R 3+ M 0 2- M 1 3 3 * * * * * * * * * * * * * V+ M R-M R-M Ptiliidae (Federflügler)

41

Anmerkung

Letzter Nachweis Letzter

Verantwortung SH Verantwortung

Verantwortung D Verantwortung

in SH in Verbreitungstyp Verbreitungstyp Wertgebende Begleitart Charakteristische Arten

Charakter- art i.e.S.

Risikofaktoren Risiko

= = kurzfristig )

↑ ↓ ( langfristig

Kriterien

Bestand Schutzstatus , 1867) mh = = = ., 1845) s = = = EY ., 1855) mh = = = ., 1934 s = = = ., 1884) ss = = = .R AB ILLM , 1966) es = = = , 1960 es = = = , 1887) mh = = = ., 1857) es = = = ., 1975 ss ? ., 1858 ss < ., 1860) es = = = .L (G ., 1844) mh = = = ULS

Taxon ., 1834) mh = = = ATTH OSSK , 1966) es = = = , 1841) h = = = , 1841) s < = - D YBAS R ., 1845) mh = = = LACH ., 1839) mh < = - D ATTH AIRM (M ATTH LLIB OHNS , 1841) mh = = = (M R AIRM TRAND (F J (F (D EER ., 1845) s < = - D M EWM EER S YBAS (A (F (E (M R UER EER (H (N (H (D (E (G (H (= Nanoptilium kunzei (Heer, 1841)) (= Nanoptilium kunzei (Heer, (= Nanoptilium brevicolle (Matth., 1860)) (= Baeocrara litoralis (Thoms., 1855))

Euryptilium saxonicum Ptiliola kunzei Ptiliola brevicolle Ptiliolum schwarzi Ptiliolum wuesthoffi Ptiliolum spencei Ptiliolum fuscum Ptinella britannica Ptinella limbata Ptinella denticollis Ptinella aptera Ptinella tenella Ptinella errabunda Pteryx suturalis Pteryx splendens Nephanes titan Smicrus filicornis Baeocrara variolosa Baeocrara japonica Bambara fusca Bambara contorta Grund Kat. Änd. Kat. Grund K K K

* * * * *

Kat +/- Kat RL 2010 RL 3+MK 3-RK 2 * * * * * * * * * * * * V-RK R-M R-M R-M R-MK R-MK

42 ======) ) ) ↑ ↑ ↓ ↓ ↓ ↓↓ ( ( ( , 1815 s < = - D nWZ2 , 1954 s < = - D PENCE ., 1790 mh = = = ., 1830 mh = = = L ., 1844) mh = = = O ., 1845) sh = ., 1844) mh = = = & S , 1971 ss < ., 1845) sh = = = ., 1792) mh = = = ANNH ., 1844) es = = = , 1838) ss < = = ., 1852) h = = = ., 1877) mh = = = ., 1845) mh = = = LLIB ., 1935 ss < , 1774) sh = = = AMANINI M ANNH ., 1914 s = = = ., 1935 ss = , 1841) mh = = = R ., 1872) ss = ., 1935 es = = = QuB BST , 1825) mh = = = IRBY UNDT ILLM ., 1865) mh = = = T OTSCH ., 1935 es = = = (L., 1758) mh = = = (A LLIB ., 1793) mh = = = ., 1845 s < ALTL K S (E (M R (H EER AEKL ATTH (G EER (A OSSK (M ., 1845) mh = = = AFFR E ATTH OSSK OSSK (W ANZ ENNY ATTH R (G R (H R OSSK (M (M R R (P (D (E R (M (M (= Acrotrichis fratercula (Matth., 1878)) (= Acrotrichis fraterna Johns.)

Acrotrichis grandicollis Acrotrichis montandonii Acrotrichis chevrolatii Acrotrichis thoracica Acrotrichis sericans Acrotrichis dispar Acrotrichis brevipennis Acrotrichis pumila Acrotrichis silvatica Acrotrichis cognata Acrotrichis insularis Acrotrichis arnoldi Acrotrichis intermedia Acrotrichis atomaria Acrotrichis lucidula Acrotrichis sitkaensis Acrotrichis henrici Acrotrichis rosskotheni Acrotrichis fascicularis Acrotrichis rugulosa Siagonium quadricorne Scaphidium quadrimaculatum Scaphisoma agaricinum Scaphisoma boleti Scaphisoma assimile Scaphisoma balcanicum Phloeocharis subtilissima Bibloporus bicolor Bibloporus minutus K ↓ ↓ ↓ * * +M 3 3-MK 2-R 3+MK 3+MK 2-R 3-R * * * * * * * * * * * * * * * * * * * R+ M R-MK R+ M Staphylinidae (Kurzflügler)

43

Anmerkung Letzter Nachweis Letzter

1919 Verantwortung SH Verantwortung

*

Verantwortung D Verantwortung

in SH in Verbreitungstyp Verbreitungstyp Wertgebende Begleitart Charakteristische Arten

Charakter- art i.e.S.

Risikofaktoren Risiko

=sö =WAQ = = = = =nWZ1 kurzfristig ) ) ) ) )

↑ ↓ ↓ ↓ ↓ ↓ ↓↓ ( ( ( ( ( langfristig

Kriterien

Bestand Schutzstatus ., 1881 es = = = TT R , 1833) s = ., 1881 es = = = ., 1816) mh = = = TT UBE ., 1814) ex ., 1880) ss < , 1825 s < ., 1876) es = = - D nWZ1 R ., 1857) es = = - D nWZ1 , 1844) es = = = sö ., 1841) mh = = = (A TT TT ., 1816) s < ., 1861 mh = = = ., 1816) mh = = = , 1895 ss < ., 1835 mh = = = EICHB ., 1888 es = ., 1910 mh = ., 1881 s < = - D nWZ2 (R UBÉ (R AFFR ., 1816) mh = = = ENNY AIRM , 1833 es = = = (R ULS TT RIMM

Taxon D (A (R (F R EICHB EICHB M AFFR (G UILLB AUVEL UBÉ OTSCH (R EICHB R F (R A G M (R (= Euplectus bohemicus Mach., 1935) (= Euplectus fauveli Guillb., 1888) (= Grammoplectus spinolae (Aubé, 1844))

Bibloporus mayeti Bibloplectus tenebrosus Bibloplectus spinosus Bibloplectus ambiguus Bibloplectus minutissimus Euplectus nanus Euplectus piceus Euplectus infirmus Euplectus bescidicus Euplectus sanguineus Euplectus signatus Euplectus bonvouloiri narentinus Euplectus bonvouloiri rosae Euplectus punctatus Euplectus karsteni Euplectus mutator Euplectus brunneus Euplectus duponti Leptoplectus spinolae Plectophloeus nubigena Plectophloeus nitidus

Grund Kat. Änd. Kat. Grund K K K K

↑ ↓ ↓ * * ↓ ↓ Kat +/- Kat

+R +M RL 2010 RL 2-R 2-R 0 3 3-R 1 3-R 1 1-RK * * * * * * * V+MK R-M R+ M R- M R-M

44 1954 * ** - D StW Feh ↓↓ ., 1836) s < = = -G., 1836) s = = = SzB Kü [6] OTSCH , 1817) s < = = , 1817) s < = = (M ., 1792) ex , 1904 es = = = KüN ., 1816) h = = = , 1790 es = , 1817) ss < = = WLa CHMIDT , 1833) es = = = nWZ2 ., 1816) s = = - D , 1833 s < = = WLa WAQ, BST ., 1816) s < = - D nWZ2 , 1844) es = = = EACH EACH ., 1816) s = = = ., 1988 es = = - D (S , 1825) mh = = = , 1833) mh = = = (L (L (H ., 1805) s < = = UBE ES UBÉ EICHB ., 1806) ss < = = WAQ sö AFFRAY EACH UBE , 1998 mh = = = ., 1803) s = = = ., 1880 es < = = WAQ EICHB , 1825 s < = = WAQ ., 1816) mh = = = B A (A R UBÉ EICHB (L (R ., 1792 mh < = = EICHB ENNY TT (A ANZ (R ANZ REYSSLER ANZ R (A (R (R (P (D BST P ENNY (P (P ., 1800) mh = = = ERCHE EICHB H D Z (R AYK (P (= Bryaxis glabricollis (Reichenbach, 1816)) (= Rybaxis laminata (Motsch., 1836)) 1904) (= Brachygluta haematica simplicior Raffray, (= Brachygluta haematica sinuata (Aubé, 1833)) impressus (Panz., 1805)) (= Trissemus (= Metopsia clypeata sensu FHL Bd 4)

Plectophloeus fischeri brevicorne Trimium sulcicollis Trichonyx Amauronyx maerkelii Batrisodes venustus Batrisodes unisexualis Bythinus macropalpus Bythinus burrellii Bryaxis puncticollis Bryaxis curtisii Bryaxis bulbifer Bryaxis clavicornis niger Tychus monilicornis Tychus Rybaxis longicornis s.str. Rybaxis longicornis f. laminata Brachygluta fossulata Brachygluta simplicior Brachygluta sinuata Brachygluta helferi Reichenbachia juncorum Fagniezia impressa Pselaphaulax dresdensis Pselaphus heisei mucronatus Tyrus Claviger testaceus Metopsia similis K ↓ * +M +M +M 3 3+ M 2+ M 1 0-R 1-RK 3 * * * * * * * * * * V+ M V+MK V+ M V+ M V-MK V+ M V+ M R+ M R+ M R-M

45 Anmerkung

[57] Letzter Nachweis Letzter

1947 Verantwortung SH Verantwortung

***

Verantwortung D Verantwortung

in SH in Verbreitungstyp Verbreitungstyp Wertgebende Begleitart Charakteristische Arten

Charakter- art i.e.S.

Risikofaktoren Risiko

-I kFG = =WLa = =(!) kurzfristig ) )

↑ ↓ ↓

↓↓ ( ( ↓↓↓ langfristig

Kriterien

Bestand Schutzstatus , 1857 es = = = AL V ., 1802) mh = = = , 1857) ex , 1789) mh = = = , 1941) ss << ., 1904 mh = = = , 1964 es = , 1828 s < , 1817) mh = = = ARSH ., 1859 es = = = ., 1872 mh = = = ÄRKEL R ANGAS AYKULL K ECK (M ., 1810) s = = = ., 1839 es < URTIS (F., 1792)(F., h = = = ., 1857 es = = = (M (F., 1792)(F., mh = = = (P (K R ., 1840 s = = = (B CHATZM C R

Taxon OCHH AQUELINE DU (F., 1789)(F., mh < = = ., 1840 mh = = = R ERSTENS ., 1834 ss = E YLL S K J R H E K E (G TEPH S (= Megarthrus depressus sensu FHL Bd 4 ) (= Megarthrus sinuatocollis sensu FHL Bd 4 ) (= Proteinus macropterus sensu FHL Bd 4 )

Megarthrus prosseni Megarthrus depressus Megarthrus denticollis Megarthrus nitidulus Proteinus ovalis Proteinus brachypterus Proteinus atomarius Proteinus laevigatus Micropeplus tesserula Micropeplus fulvus Micropeplus marietti Micropeplus longipennis Micropeplus porcatus Micropeplus ripicola Micropeplus caelatus Eusphalerum minutum Eusphalerum sorbicola Eusphalerum signatum Eusphalerum sorbi Eusphalerum torquatum

Grund Kat. Änd. Kat. Grund ↓ ↓ ↓ * Kat +/- Kat

+M RL 2010 RL 1 2-R 1 0-R 1-R * * * * * * * * * * * * R+ M R-MK R- M

46 1944 * ** *** -ID nWZ2 -ID nWZ1 -ID nWZ1 - D StD Kü =WAQ = = =WLa =nWZ1 = = ) ) ↑ ↓ ↓ ↓↓ ↓↓ ↓↓ ↓↓ ↓↓ ↓↓ ↓↓ ↓↓ ( ( ., 1856) ss < = - ID nWZ2 ., 1932) ss < AB . & L ERNHAUER ., 1827 es < = = RFk Hgld (F., 1787)(F., ss << ., 1903 ss < ., 1806 mh = = = AIRM ., 1800) ss < (B ., 1912) ss < ., 1834) mh = = = , 1940 ss < = = WLa YLL ., 1857 s < (F RAV ., 1789) mh = = = ., 1834) s = = = ., 1810) ss = ., 1810 mh = = = G AYK ERNH ., 1806) ss < ., 1806 mh = = = G ., 1789) sh = = = TEPH ERNH B (P ., 1795) s < AYK ., 1840) ss ? YLL YLL ., 1857 es = = = L ., 1857 s < = - D StD Kü R TEPH RAV (S RAV ., 1840) ex ANSEN HOMS (B R AYK (P R ., 1813) es << G (G R T G H K (O (E (S (G K (P (E YLL (G (= Eusphalerum florale (Panz., 1793)) 1802)) (= Elonium striata (Grav., (= Elonium pliginskii (Bernh., 1913)) (= Elonium sulcula (Steph., 1834)) (= Phyllodrepa pygmaea (Payk., 1800)) 1840)) (= Phyllodrepa rufula (Er., (= Phyllodrepa ioptera (Steph., 1834)) 1840)) (= Phyllodrepa vilis (Er., (= Phyllodrepa gracilicornis (Fairm. & Lab., 1856))

Eusphalerum tenenbaumi Acrulia inflata Acrolocha minuta Acrolocha pliginskii Acrolocha sulcula Phyllodrepa melanocephala Phyllodrepa salicis Phyllodrepa nigra Phyllodrepa puberula Phyllodrepa floralis Phyllodrepa melis Hapalaraea pygmaea Hypopycna rufula Dropephylla ioptera Dropephylla vilis Dropephylla gracilicornis Omalium laeviusculum Omalium riparium Omalium validum Omalium rivulare Omalium oxyacanthae Omalium exiguum Omalium littorale Omalium caesum K K K ↑ ↓ ↓ * ↓ ↓ +R 1 3-R 3+MK 2-R 1 2-R 1 0 2 2+ M 3 3 2 2-R * * * * * * * * V+MK R-M

47 Anmerkung

[58] Letzter Nachweis Letzter

1916 Verantwortung SH Verantwortung

**

Verantwortung D Verantwortung

in SH in Verbreitungstyp Verbreitungstyp Wertgebende Begleitart Charakteristische Arten

Charakter- art i.e.S.

Risikofaktoren Risiko

-ID nWZ2 = = = = WLa nWZ2 kurzfristig ) )

↑ ↑ ↓ ↓ ↓↓ ( ( langfristig

Kriterien

Bestand Schutzstatus , 1873) es = = = EAY L ., 1867 mh = = = AC , 1880 mh = = = ., 1827) sh = = = (M ., 1802) mh = = = , 1783) es = = = SbW KüN EY HOMS ., 1802) s < YLL T ., 1806) mh = = = ., 1834) ss << = = ., 1810) ss < ., 1834 mh = = = ., 1838) s = = = ARSH ., 1840) ss < (G RAV ., 1840) ex R RAV R TRÖM YLL ., 1902 es = = = . & R ., 1877 s < = = ETT ., 1902 ss ? (G (M TEPH ., 1792) mh = (E TEPH (E (S

Taxon (G (Z (G S (S ULS ERH AYK ERNH ERNH G M B (P B (= Phloeonomus monilicornis (Gyll., 1810)) (= Phloeonomus bosnicus Bernh., 1902) (= Phloeonomus plana (Payk., 1792)) (= Phloeonomus lapponica (Zett., 1838)) (= Xylodromus brunnipennis (Steph., 1834)) (= Lathrimaeum atrocephalum (Gyll., 1827))

Omalium rugatum Omalium italicum Omalium excavatum Phloeonomus pusillus Phloeonomus punctipennis Xylostiba monilicornis Xylostiba bosnica Phloeostiba plana Phloeostiba lapponica Paraphloeostiba gayndahensis Xylodromus depressus Xylodromus affinis Xylodromus concinnus Xylodromus testaceus Philorinum sordidum Micralymma marinum Orochares angustatus Anthobium atrocephalum Grund Kat. Änd. Kat. Grund K

K

↑ ↓ * ↓ * Kat +/- Kat

+R +M RL 2010 RL 2 2 3-R 0 2-R * * * * * * * * * V+ M R-M R- M R+ K

48 *** -D kFG -ID nWZ2 =QuB, kFG ==QuB, =kFGQuB gFG, kFG =WLa ) ) ) ) ↓ ↓ ↓ ↓ ↓↓ ↓↓ ( ( ( ( , 1856 es = = = EY ., 1802) s << = - I HMo, NMo , 1957) es = = - ID kFG RAV ., 1821 es = = = kFG ., 1834 ss < . & R AGEL , 1777) sh = = = (G ., 1834 mh = = = ., 1802) h = = = (F ., 1830 es < ., 1821) mh = = = ÜLL ULS ., 1839) s < = = gFG ., 1827) ss < = = gFG sö [59] ., 1830) s < = = TEPH (L., 1758) s < ., 1806) s < R ., 1871 mh = = = ., 1802) s << = = kFG M ., 1800) ss << = = WAQ OEZE ., 1967) ss ? ? = M ., 1806) mh = = = S ., 1810) mh = = = ÜLL TEPH YLL ARSH (E , 1953 ss < (F., 1793)(F., mh = = = ANNH RAV S (G RAV AUV AYK RAV (M YLL ANNH (G MET F ., 1839 ss < (M M (G (P (G R (G (S (G OHSE (M E L (F., 1793)(F., s = = = (= Lathrimaeum unicolor (Marsh., 1802)) (= Coryphium letzneri Schwarz, 1873) (= Acrognathus mandibularis (Gyll., 1827)) 1856)) (= Ancyrophorus flexuosus (Muls. & Rey, (= Ochthephilus rivularis (Smetana, 1967)) bilineatus (Steph., 1834)) (= Trogophloeus similis (Smet., 1967)) (= Trogophloeus

Anthobium unicolor Olophrum piceum Olophrum fuscum Olophrum assimile Arpedium quadrum Eucnecosum brachypterum Acidota crenata Acidota cruentata Lesteva punctata Lesteva sicula heeri Lesteva longoelytrata Lesteva hanseni Lesteva pubescens Anthophagus praeustus Anthophagus caraboides Coryphium angusticolle Syntomium aeneum Deleaster dichrous Coprophilus striatulus Manda mandibularis Planeustomus palpalis Ochthephilus flexuosus Ochthephilus andalusiacus Carpelimus bilineatus Carpelimus similis K K K ↓ ↓ ↓ * ↓ ↓ 2- M 2 3+ M 1-R 3+ M 1 2-RK 2-R 3-R 3-R 2-R * * * * * * * * * V+ M V-MK R+ M R-M D

49 Anmerkung

[60]

Letzter Nachweis Letzter Verantwortung SH Verantwortung

**

Verantwortung D Verantwortung

in SH in Verbreitungstyp Verbreitungstyp Wertgebende Begleitart Charakteristische Arten

Charakter- art i.e.S.

Risikofaktoren Risiko

- D SzP SzW, Kü [61] = SzB SzW, kurzfristig ) )

↓ ↓ ( ( langfristig

Kriterien

Bestand Schutzstatus ., 1946) ss << = = OUB (R , 1835) s = = = , 1934) s = = = ., 1901) es <<< ENICK ., 1823) s = = = Kü ., 1844) ss = ., 1802) mh = = = ., 1860) mh = = = , 1871) ss ? ? = ., 1806) h = = = ERNH , 1942) s = = = ., 1844) ss < = = RAV AHLB ACORDAIRE (L. B IESW RAV (B HARP (G (L (S OTSCH

Taxon (K ALM (G IESW (S (P (M (K (= Trogophloeus augustae Bernh., 1901) (= Trogophloeus rivularis Motsch., 1860) (= Trogophloeus 1844) obesus Kiesw., (= Trogophloeus 1802) fuliginosus Grav., (= Trogophloeus lindrothi Palm, 1942) (= Trogophloeus impressus Lacordaire, 1835) (= Trogophloeus heidenreichi L. Benick, 1934) (= Trogophloeus 1806)) corticinus (Grav., (= Trogophloeus (= Carpelimus subtilicornis (Roub., 1946)) ganglbaueri Bernh., 1901) (= Trogophloeus foveolatus (Sahlb., 1823)) (= Trogophloeus 1844) halophilus Kiesw., (= Trogophloeus

Carpelimus erichsoni Carpelimus rivularis Carpelimus obesus Carpelimus fuliginosus Carpelimus lindrothi Carpelimus impressus Carpelimus heidenreichi Carpelimus corticinus Carpelimus manchuricus subtilicornis Carpelimus ganglbaueri Carpelimus foveolatus Carpelimus halophilus

Grund Kat. Änd. Kat. Grund Kat +/- Kat

+M +M +M +M RL 2010 RL 3 2 1 3+MK * * * * * * * D

50 [61] 1909 ** ** - D SbW SzP, SzW, KüN = =WAQ = =WAQ ) ↓ ↓↓ ↓↓ ↓↓ ( ↓↓↓ , 1870) s < = - D StU ELVE S ., 1895) ss = ., 1834) s = = = ., 1900) es = = = ., 1839) mh = = = ., 1802) h = = = ., 1830) mh = = = ., 1861 es << = = StD Kü R HP ., 1849) ss < ., 1802) mh = = = AUDI DI ANGLB TEPH ., 1802) ss < ., 1806) ss < (E (S ., 1839) es = = = ., 1806 mh = = = (B (S ., 1904 mh = = = (G ., 1839) s < = = ARSH R RAV ANNH RAV RAV ., 1839) es = R HOMS OCHH R ., 1839 s < = = WAQ (E RAV T (G (M AUV (G (G (E (M R (H G F (E (F., 1775)(F., sh = = = E (L., 1767) ex (= Trogophloeus schneideri (Ganglb., 1895)) (= Trogophloeus despectus Baudi di Selve, 1870) (= Trogophloeus 1839) exiguus Er., (= Trogophloeus zealandicus (Shp., 1900)) (= Trogophloeus lindbergi (Scheerpeltz, 1937)) (= Trogophloeus gracilis (Mannh., 1830)) (= Trogophloeus 1839) subtilis Er., (= Trogophloeus 1839) elongatulus Er., (= Trogophloeus arcuatus (Steph., 1834)) (= Trogophloeus 1806) (= Oxytelus insecatus Grav., (= Oxytelus rugifrons Hochh., 1849) 1775)) (= Oxytelus rugosus (F., 1862) (= Oxytelus perrisi Fauv.,

Carpelimus schneideri Carpelimus despectus Carpelimus exiguus Carpelimus zealandicus Carpelimus pusillus Carpelimus gracilis Carpelimus subtilis Carpelimus elongatulus Thinodromus arcuatus Aploderus caelatus Aploderus caesus Oxytelus sculptus Oxytelus migrator Oxytelus fulvipes Oxytelus piceus Oxytelus laqueatus Anotylus insecatus Anotylus rugifrons Anotylus rugosus Anotylus maritimus K K K ↓ ↓ ↓ ↓ * * ↓ +M +M 2-R 3 2-R 2-R 2+ M 0-R 1 2-R * * * * * * * * V+ M V+ M R-M R-M

51

Anmerkung

Letzter Nachweis Letzter

Verantwortung SH Verantwortung

Verantwortung D Verantwortung

in SH in Verbreitungstyp Verbreitungstyp Wertgebende Begleitart Charakteristische Arten

Charakter- art i.e.S.

Risikofaktoren Risiko

= = = = kurzfristig ) ) ) )

↓ ↓ ↓ ↓ ( ( ( ( langfristig

Kriterien

Bestand Schutzstatus , 1785) mh = = = , 1799) sh = = = ., 1830 es = = = ., 1861 s = = = ., 1867) ss <<< ., 1802) mh = = = ., 1806) h = = = ., 1832) ss << = = SzP SzW, ., 1839) ss << = = LOCK ANNH EOFFROY R ., 1867) es << RAV AND RAV HOMS , 1963) s = = = ., 1802) es << = = (B T M (E (G (P

Taxon ., 1806) s = (G (G ., 1867) es << AHLB AND RAV OHSE (S RAV (P AND (G (L (G (P (= Oxytelus inustus Grav., 1806) (= Oxytelus inustus Grav., 1806) (= Oxytelus sculpturatus Grav., (= Oxytelus mutator Lohse, 1963) 1802) (= Oxytelus nitidulus Grav., 1839) (= Oxytelus complanatus Er., (= Oxytelus clypeonitens Pand., 1867) (= Oxytelus saulcyi Pand., 1867) (= Oxytelus fairmairei Pand., 1867) (= Oxytelus tetracarinatus (Block, 1799))

Anotylus inustus Anotylus sculpturatus Anotylus mutator Anotylus nitidulus Anotylus complanatus Anotylus clypeonitens Anotylus saulcyi Anotylus fairmairei Anotylus tetracarinatus Platystethus arenarius Platystethus cornutus Platystethus alutaceus Platystethus nodifrons Platystethus nitens

Grund Kat. Änd. Kat. Grund K

↓ ↓ ↓ Kat +/- Kat

+M +M RL 2010 RL 2 2- M 2-MK 1-R 1-R 1 * * * * * * V-R R+ M

52 [62] * ** ** - ID SzW StS KüO = SKü= SKü KüO =nWZ2 =nWZ1 KüO ) ) ↓ ↓ ↓↓ ↓↓ ↓↓ ( ( , 1986 s < = - D SzP SzW, KüN [62] , 1978 mh = = = KüN ERMAN H ., 1929 ss < = = SzW KüN OHSE , 1984 es = = = ., 1907 es < = - D L ILL ., 1825) s < = - D SKü KüO ., 1866 s < = = kFG ., 1898 s = = - D StD gFG Kü ., 1839 s = = = R ., 1827) s << = = CaH, Mgr W , 1912 s < = - D StD gFG Kü , 1799) h = = = ., 1872 es < = = SKü, StU OHSE , 1866 ss < = - D SzP SzW, Kü ONDR ., 1784) mh = = = ., 1940 ss < E F., 1793F., es <<< ., 1806) s = = = ERM ., 1806) mh = = = L B OY ., 1840 ss < YLL ., 1839 mh = = = , 1839 es = = - D StU ., 1839 mh = = = CHDTE J R (G AUV ., 1872 ss < = - I Mgr BST CHNEID R R RAV RAV S LOCK F (G ., 1911 mh = = = E OTTH S E ., 1840 ss < E (H EER (G T R (L., 1758) ss < (G HP (B AUV CHIØDTE H E F S S (= Bledius bicornis atlanticus Lohse, 1978) 1935) (= Bledius germanicus Wagn., (= Bledius larseni Hansen, 1940) 1878 nec FHL) (= Bledius subsinuatus Muls. & Rey, (= Bledius fracticornis (Payk., 1790)) (= Bledius arenarius (Payk., 1800))

Bledius bicornis jutlandensis Bledius diota Bledius spectabilis frisius Bledius limicola Bledius tricornis Bledius defensus Bledius pallipes Bledius terebrans Bledius filipes Bledius longulus Bledius opacus Bledius gallicus Bledius dissimilis Bledius occidentalis Bledius femoralis Bledius atricapillus Bledius praetermissus Bledius nanus Bledius baudii Bledius erraticus Bledius subniger Bledius fergussoni Bledius subterraneus Bledius tibialis Thinobius flagellatus Oxyporus rufus Oxyporus maxillosus K K K ↓ * ↓ ↓ +M 2+ M 3+MK 3 3 2 2+MK 2-R 3-RK 1 2+MK 1 2-RK 1-R 3+ M 2-R * * * * * * * * * V+ M V-RK R-M

53

Anmerkung Letzter Nachweis Letzter

1948 Verantwortung SH Verantwortung

*

Verantwortung D Verantwortung

in SH in Verbreitungstyp Verbreitungstyp Wertgebende Begleitart Charakteristische Arten

Charakter- art i.e.S.

Risikofaktoren Risiko

-I Mgr = HMo, NMo = HMo, ErH kurzfristig ) )

↓ ↓

( ( ↓↓↓ langfristig

Kriterien

Bestand Schutzstatus , 1861 es << EY .R ., 1810 mh = = = ., 1851 es << = - ID fDÜ ., 1833 s < = = , 1864 s = = = Elb ULS , 1863 mh = = = ., 1763) sh = = =

Taxon M ., 1839 mh = = = YLL , 1810 sh = = = ., 1838 es = = = ., 1840 s < = = SKü, StU, kFG R COP TEPH ., 1821 s < = = kFG G ., 1839 ss < = = ., 1873 ss << = = Röh R ., 1839 ex CRIBA ONTE HOMS E ., 1839 mh = = = S R ., 1830 es << ., 1839 mh = = = R (S E ., 1839 ss << ETT T (L., 1758) mh = = = S R C R E ., 1789) sh = = = R ÜLL E Z E AUV E E E L JUNGH F M L AYK ANNH (P M (= Stenus bipunctatus Er., 1839) (= Stenus bipunctatus Er., (= Stenus excubitor sensu FHL Bd. 4 ) 1857) (= Stenus rogeri Kr.,

Stenus biguttatus Stenus comma Stenus guttula Stenus fossulatus Stenus aterrimus Stenus juno Stenus calcaratus Stenus ater Stenus longitarsis Stenus gallicus Stenus lustrator Stenus proditor Stenus sylvester Stenus clavicornis Stenus providus Stenus bimaculatus Stenus palposus Stenus boops Stenus incrassatus Stenus melanarius Stenus subdepressus

Grund Kat. Änd. Kat. Grund ↓ ↓ Kat +/- Kat

+M RL 2010 RL 1-R 1 3+ M 0-R 1 2 1 * * * * * * * * * * V+ M V-MK V+ M R+ K

54 1921 * -I =HMo =WAQ = == SzW Kü [63] ) ↑ ↓ ↓↓ ↓↓ ↓↓ ( ↓↓↓ ., 1843 s < = - I HMo, NMo ., 1802) s < = = gFG SzW, Kü ., 1827 h = = = , 1994 ss < = - I , 1966 s << = = NMo WAQ, ., 1827 mh = = = ANNH ., 1833 mh = = = ., 1833 mh = = = , 1884 s < = - ID StS ., 1870 es << = - ID NMo HMo YLL , 1884 es = = = , 1810 s = ARSH M ., 1827 ss < EY ., 1802 es <<< ., 1933 s < = - ID YLL ., 1833 mh = = = ., 1802 ss << = = Röh G EY UTHZ ., 1806 ss << = = HMo, NMo R ., 1839 ss < ., 1833 mh = = = ., 1833 s < = - I G TEPH SSING TEPH ., 1839 s < (M R YLL R P ., 1839 mh = = = ., 1806 es << RAV ., 1806 ss << = = WAQ S RAV HOMS R A S ., 1839 h = = = R E ., 1839 ex G TEPH RAV TEPH T G E R G JUNGH R E S TEPH S TEPH RAV G RAV E L E S S G G (= Stenus rossicus Bernh., 1903) (= Stenus cautus sensu FHL Bd. 4 ) 1839) (= Stenus vafellus Er., 1840) (= Stenus exiguus Er., (= Stenus simillimus Benick, 1949) (= Stenus problematicus Kevan & Allen, 1962)

Stenus morio Stenus excubitor Stenus atratulus Stenus melanopus Stenus canaliculatus Stenus nitens Stenus fuscipes Stenus argus Stenus europaeus Stenus cautus Stenus pusillus Stenus nanus Stenus assequens Stenus contumax Stenus circularis Stenus humilis Stenus carbonarius Stenus glabellus Stenus opticus Stenus formicetorum Stenus intermedius Stenus crassus Stenus brunnipes Stenus nigritulus Stenus latifrons Stenus fulvicornis Stenus tarsalis ↑ ↓ +R 2 1 1 3 0 3+MK 2-RK 2- M 3 2-MK 3-MK 3 2 3 2- M 1-R * * * * * * * * * V+ M R- M

55

Anmerkung

Letzter Nachweis Letzter Verantwortung SH Verantwortung

**

Verantwortung D Verantwortung

in SH in Verbreitungstyp Verbreitungstyp Wertgebende Begleitart Charakteristische Arten

Charakter- art i.e.S.

Risikofaktoren Risiko

- D kFG [65] = =NMo === =QuB kFG, euG [64] kurzfristig ) ) ) )

↓ ↓ ↓ ↓

↓↓ ( ( ( ( ↓↓↓ ↓↓↓ langfristig

Kriterien

Bestand Schutzstatus ., 1833 s < = = QuB TEPH ., 1851 ss << = = HMo, ErH S HOMS ., 1833 mh = = = ., 1783) h = = = ., 1810) ss < T ., 1856 s << = = HMo ., 1947 mh = = = ., 1827 mh = = = YLL ., 1833 s < , 1804 mh < = = TEPH ., 1806 s << = - ID Mgr CaH, BDü, KDü, ., 1824 h = = = ., 1833 ss < , 1884 ss < OSH CHALL ., 1914 es = = = ., 1833 s << = = Röh S (G

Taxon YLL R ., 1833 h = = = ACH TT RAV (S ., 1840 ss < TEPH ., 1802 s << = = G ERM R ., 1784) s << R G ASEY TEPH M S ., 1865 es << ., 1839 h = = = TEPH JUNGH G E S ., 1840 s << = = Röh, NMo C R L S RAV TEPH R BST E AUV S G E F (H (= Edaphus bluehweissi Scheerp., 1936)

Stenus bohemicus Stenus similis Stenus solutus Stenus cicindeloides Stenus kiesenwetteri Stenus fornicatus Stenus pallitarsis Stenus niveus Stenus binotatus Stenus pubescens Stenus umbratilis Stenus flavipes Stenus nitidiusculus Stenus picipennis Stenus bifoveolatus Stenus picipes Stenus picipes brevipennis Stenus impressus Stenus palustris Stenus geniculatus Stenus pallipes Dianous coerulescens Edaphus beszedesi

Grund Kat. Änd. Kat. Grund ↓ ↓ ↓ ↓

Kat +/- Kat RL 2010 RL 2 2 2-R 3 3 1-R 1 2 2-R 3 2-R 3+ M 2 * * * * * * * * V+ M R+ M

56 1898 1951 1957 1957 == Mgr öH = ) ) ↑ ↓ ↓ ( ( , 1804) mh = = = ., 1844 s = = = ., 1835 mh = = = , 1785) s > = = ANNH JUNGH M ACORD (L , 1839) s < ., 1789) mh = = = ., 1830) es = = = L , 1908) es = = = ., 1802) es << = = ., 1826 ex ., 1806) ss < = - I Mgr ., 1802 ss < ., 1867) mh = = = ., 1839 ex EOFFROY OY EER ., 1836) h = = = R AYK ., 1789) ss < = - I Mgr ., 1839) ss << = = WLa RAV (F., 1781)(F., ex URT RAV (J E RAV R (G ANNH (H (P AUV C (G ., 1854) ss < = - ID sö G (G AYK ., 1839) es = = = ., 1840 ex ., 1858) s = (L., 1758) h = = = (E ERM (F R R R R (M (P E (G (K (E (K (= Paederus ruficollis (F., 1781)) (= Paederus ruficollis (F., 1806)) (= Astenus filiformis (Latr., (= Astenus longelytratus Palm, 1936) 1785)) (= Stilicus angustatus (Geoffroy, 1840)) (= Stilicus subtilis (Er., (= Stilicus rufipes (Germ., 1836)) 1839)) (= Stilicus geniculatus (Er., (= Stilicus orbiculatus (Payk., 1789)) 1867) (= Stilicus erichsoni Fauv.,

Euaesthetus bipunctatus Euaesthetus ruficapillus Euaesthetus laeviusculus Paederus fuscipes Paederus riparius Paederus littoralis Paederidus ruficollis Astenus procerus Astenus pulchellus Astenus lyonessius Astenus gracilis Rugilus angustatus Rugilus subtilis Rugilus rufipes Rugilus geniculatus Rugilus orbiculatus Rugilus erichsoni Medon castaneus Medon dilutus Medon piceus Medon brunneus Medon fusculus Medon ripicola K K ↑ ↓ ↓ ↓ ↓ * ↓ +M +R 2-RK 3-R 2-RK 0 0-R 2 2-R 0-R 2- M 2 0-R * * * * * * * * * R+ M R-M R+ M

57

Anmerkung Letzter Nachweis Letzter

1910 Verantwortung SH Verantwortung

*

Verantwortung D Verantwortung

in SH in Verbreitungstyp Verbreitungstyp Wertgebende Begleitart Charakteristische Arten

Charakter- art i.e.S.

Risikofaktoren Risiko

=sö = = Feh kurzfristig

↑ ↓↓ ↓↓ langfristig

Kriterien

Bestand Schutzstatus ., 1837) es = = - D [66] ., 1802) s = = = ., 1857) ss = ., 1840) s = = = R , 1839) es = RAV ORDM OLL (E ., 1833) mh = = = (G ., 1802) mh = = = ., 1827) es << , 1872 es = = = sö (N (W (F., 1792)(F., es = = = Elb ., 1859) mh = = = ., 1843 ex Mgr sö 1961 ., 1840) ex RAV YLL R R

Taxon TEPH ., 1840 s < = - I Mgr RICHSON (G (K (G (E R (S IESW (E E ., 1857) s = = = K ., 1795) s = = = R L OLLASTON (K W (O (= Hypomedon bicolor (Ol., 1795)) 1792)) (= Hypomedon melanocephalus (F., 1857)) (= Chloecharis debilicornis (Woll., 1855) (= Scopaeus cognatus Muls. & Rey, (= Scopaeus didymus sensu FHL Bd. 4 ) (= Scopaeus minimus sensu FHL Bd. 4 ) (= Scopaeus furcatus Binaghi, 1935) 1802) (= Lathrobium multipunctum Grav.,

Medon apicalis Sunius bicolor Sunius melanocephalus Hypomedon debilicornis Pseudomedon obscurellus Pseudomedon obsoletus Lithocharis ochracea Lithocharis nigriceps Scopaeus laevigatus Scopaeus sulcicollis Scopaeus minutus Scopaeus pusillus Scopaeus ryei Scopaeus minimus Domene scabricollis Lobrathium multipunctum

Grund Kat. Änd. Kat. Grund K

↑ ↓ ↓ * Kat +/- Kat

+M +R +M +M RL 2010 RL 1-R 1-RK 3+MK 0-R 1 0 * * * * * * * * R-M R+ K

58 1939 =HMo = = gFG Kü ) ) ↓ ↓ ↓↓ ( ( , 1977) es << ., 1936 mh = = = ., 1841 mh = = = ., 1851 s < = = ., 1800) mh = = = ., 1846 es = = = Elb UONA ., 1802) mh = = = , 1841 mh = = = OL ., 1806) h = = = , 1839 ex ., 1837 s = = = ÄRK AYK ., 1789) s < = = (M ., 1909) ss < = = K ., 1813 s << = = HMo ., 1802 h = = = ., 1833 mh = = = OCHH RAV (P M TT ., 1884) s < = = SzW ONISTH EER RAV EER AYK ., 1857 mh = = = H ., 1822) ss = D TT H (G YLL (R R RAV H (L., 1767) s < (G (P ORDM ., 1839 es = = = (F., 1792)(F., h = = = K TEPH G G (R R N S ICOL E (N (= Lathrobium gracile Hampe, 1866) 1802) (= Lathrobium terminatum Grav., (= Lathrobium fennicum Renk., 1938) (= Lathrobium quadratum (Payk., 1789)) (= Lathrobium volgense Hochh., 1851) (= Lathrobium ripicola Czwal., 1888) (= Lathrobium muelleri Bernh., 1899) 1806) (= Lathrobium filiforme Grav., (= Lathrobium microps Benick, 1942) (= Cryptobium fracticorne (Payk., 1800)) (= Cryptobium collare Rtt., 1884)

Tetartopeus sphagnetorum Tetartopeus terminatus Tetartopeus rufonitidus Tetartopeus quadratus Tetartopeus Lathrobium rufipenne Lathrobium elongatum Lathrobium geminum Lathrobium pallidipenne Lathrobium laevipenne Lathrobium castaneipenne Lathrobium fulvipenne Lathrobium brunnipes Lathrobium fovulum Lathrobium impressum Lathrobium longulum Lathrobium pallidum Lathrobium dilutum Achenium humile Ochthephilum fracticorne Ochthephilum collare Leptacinus formicetorum Leptacinus intermedius K ↓ ↓ +M 3-R 1 3 3 0-R 3+MK * * * * * * * * * * * V-MK V+ M V-MK R- M R+ M

59

Anmerkung

Letzter Nachweis Letzter

Verantwortung SH Verantwortung

Verantwortung D Verantwortung

in SH in Verbreitungstyp Verbreitungstyp Wertgebende Begleitart Charakteristische Arten

Charakter- art i.e.S.

Risikofaktoren Risiko

= kurzfristig )

↓ ( langfristig

Kriterien

Bestand Schutzstatus ., 1827) mh = = = YLL (G ., 1776) mh = = = ., 1833 h = = = ., 1789) mh = = = , 1839 h = = = ., 1833) s < = = ., 1827) s < ÜLL AYK ., 1849 s < = - I EER ., 1833) mh = = = TEPH , 1839) ss < = = ., 1795) es = = = ., 1956 s << = = (P H S (M YLL AC TEPH ., 1795) sh = = = LIV J

Taxon EER L ., 1806) mh = = = (G OIFF (S TEPH (F., 1787)(F., ss << = = (O C (H (O (F., 1787)(F., s = = - I (S RAV (G (= Leptacinus othioides sensu FHL Bd. 12 ) 1802)) (= Leptacinus linearis (Grav., (= Gyrohypnus liebei sensu FHL Bd. 12 ) 1913)) (= Gyrohypnus scoticus (Joy, (= Xantholinus semirufus sensu FHL Bd. 4 ) (= Xantholinus clairei Coiffait, 1956) (= Xantholinus rhenanus Coiff., 1962)

Leptacinus sulcifrons Leptacinus batychrus Leptacinus pusillus Phacophallus parumpunctatus Gauropterus fulgidus Nudobius lentus Gyrohypnus punctulatus Gyrohypnus fracticornis Gyrohypnus atratus Gyrohypnus angustatus Xantholinus tricolor Xantholinus elegans Xantholinus laevigatus Xantholinus linearis Xantholinus gallicus Xantholinus longiventris

Grund Kat. Änd. Kat. Grund K K

* ↓ Kat +/- Kat

+M RL 2010 RL 2- M 3-MK 3-R 3 3-MK * * * * * * * * V-MK V-RK R-M

60 * *** =sö =sö = = ) ) ) ↓ ↓ ↓ ↓↓ ( ( ( , 1848) s < = = , 1853) ss << = = EY ELVE S . & R ., 1802) mh = = = ., 1799) mh = = = ., 1806) s < = = ULS ., 1835) ss < ., 1833) s < = = AUDI DI ., 1802) ss < RAV ., 1802) s < = - ID ., 1802) mh = = = ., 1802) es <<< , 1777) mh = = = ., 1802) ss < = = SzW [67] ., 1806) mh = = = ., 1833 h = = = AC (M RAV (B ., 1956 s < = = SzW Kü ., 1832) es <<< = = (G ., 1806) mh < = - D Kü RAV ., 1802) mh = = = ., 1802) mh = = = RAV RAV (L REUTZ TEPH ., 1875 ss < ., 1802) s = = = , 1837 es <<< = - ID StD KüO RAV ., 1806) s << = = HMo NMo RAV (G RAV ., 1833 s < = - I (G OEZE OIFF (G TEPH (S (G RAV (C RAV (G RAV (G TEPH (G RAV PPH S RAV C ., 1789) ss << = = nWZ2 (G (G (G E (S (G OLME TEPH (G (G S AYK H (P (= Xantholinus dissimilis Coiff., 1956) (= Baptolinus affinis (Payk., 1789)) (= Othius melanocephalus (Gravenhorst, 1806)) 1843) (= Othius myrmecophilus Kiesenwetter, (= Neobisnius cerrutii Grid., 1943) (= Philonthus immundus (Gyll., 1810))

Xantholinus dvoraki Megalinus glabratus Atrecus affinis Othius punctulatus Othius angustus Othius subuliformis Neobisnius villosulus Neobisnius procerulus Neobisnius lathrobioides Erichsonius signaticornis Erichsonius cinerascens Cafius xantholoma Remus sericeus Philonthus nigrita Philonthus fumarius Philonthus umbratilis Philonthus rufipes Philonthus debilis Philonthus atratus Philonthus albipes Philonthus alpinus Philonthus concinnus Philonthus corruscus Philonthus ebeninus Philonthus intermedius Philonthus laminatus K K ↓ ↓ ↓ ↓ 1- M 2 3-RK 3+ M 2- M 1 3-MK 2-R 2-R 3 2-R 1-R * * * * * * * * * V-RK V-MK V+ M V-MK V-MK

61 Anmerkung

[63] Letzter Nachweis Letzter

1949 Verantwortung SH Verantwortung

**

Verantwortung D Verantwortung

in SH in Verbreitungstyp Verbreitungstyp Wertgebende Begleitart Charakteristische Arten

Charakter- art i.e.S.

Risikofaktoren Risiko

= = kurzfristig ) )

↓ ↓ ( ( langfristig

Kriterien

Bestand Schutzstatus , 1853 mh = = = EY , 1941 ex ., 1832) mh = = = ., 1869 ss < = = WAQ ., 1835) es < = - ID ., 1802) sh = = = . & R TRAND ., 1860 h = = = ., 1832 sh = = = ÉNÉTR ., 1806) es = = = Röh SzW, KüO ., 1790) mh < = = S AUV RAV ., 1802) mh = = = ., 1802) ss << = = sö ., 1867 s << = = F ULS ., 1937 s << = = M (M ., 1789) mh = = = ., 1874 ss > ACORD (G RAV HP M

Taxon (G TEPH HOMS (F., 1792)(F., mh < = = RAV RAV S (L HP (G S T AYK OTTH S (G (G T (P (L., 1758) 1787)(F., mh << = = s << (= Philonthus carbonarius (Gyll., 1810)) (= Philonthus fuscipennis (Mannh., 1830)) (= Philonthus chalceus sensu FHL Bd. 4 ) (= Philonthus varius (Gyll., 1810)) 1802)) (= Philonthus bimaculatus (Grav.,

Philonthus tenuicornis Philonthus cognatus Philonthus mannerheimi Philonthus politus Philonthus succicola Philonthus addendus Philonthus decorus Philonthus nitidus Philonthus spinipes Philonthus rotundicollis Philonthus carbonarius Philonthus nitidicollis Philonthus cruentatus Philonthus pseudovarians Philonthus jurgans Philonthus varians Philonthus splendens Philonthus punctus Philonthus binotatus

Grund Kat. Änd. Kat. Grund ↓ ↓ Kat +/- Kat

+M RL 2010 RL 2-R 0-R 3-MK 3 1-RK 2+ M 3 * * * * * * * * * V-MK V+MK R- M

62 - ID Mgr CaH =HMoNMo = = SzW Kü = = = = ) ) ) ) ) ) ) ↓ ↓ ↓ ↓ ↓ ↓ ↓ ↓↓ ( ( ( ( ( ( ( , 1956 mh = = = ., 1802) s < , 1764) h = = = ., 1810) h = = = OHSE RAV ., 1832 mh = = = ., 1832 s < = = ., 1949 es <<< = = sö ., 1802) ss < = = .L ., 1802) s = = = (G YLL ., 1874 s < = = ., 1867) es << ., 1802) s < = - ID ÜLLER EN ., 1802) ss < = = HP RAV ., 1802) s < RAV (G ARR TEPH ., 1844 s < ., 1802) mh = = = ., 1802) es = = - I Mgr CaH B TEPH ., 1937 ss < = = S J RAV (M S ., 1839 ss < S (G (G RAV ., 1802) mh < = = RAV R HOMS RAV RAV (G ., 1846) ss < ENK E (G IESW (T (G RAV ., 1874) s < (G (G R K UC HP (G (L (F., 1792)(F., es << (S (= Philonthus agilis (Grav., 1806)) (= Philonthus agilis (Grav., 1793)) (= Philonthus fulvipes (F., 1792)) (= Philonthus tenuis (F., 1802)) (= Philonthus subuliformis (Grav., (= Philonthus sparsus Luc., 1846) 1802)) (= Philonthus cephalotes (Grav., (= Philonthus nigriventris Thoms., 1867) 1802)) (= Philonthus sordidus (Grav., (= Philonthus parcus Shp., 1874)

Philonthus ventralis Philonthus quisquiliarius Philonthus corvinus Philonthus sanguinolentus Philonthus salinus Philonthus rectangulus Philonthus discoideus Philonthus lepidus Philonthus coprophilus Philonthus longicornis Philonthus parvicornis Philonthus micans Philonthus micantoides Philonthus furcifer Philonthus rubripennis Philonthus marginatus Rabigus tenuis Bisnius subuliformis Bisnius sparsus Bisnius cephalotes Bisnius nigriventris Bisnius sordidus Bisnius parcus K K ↓ ↓ ↓ ↓ ↓ 3-MK 3-MK 3+MK 2-R 3-R 1-R 3 1 3-R 3+MK 1 3-MK 1 2-R * * * * * * * V-MK V-MK

63

Anmerkung Letzter Nachweis Letzter

1917 Verantwortung SH Verantwortung

**

Verantwortung D Verantwortung

in SH in Verbreitungstyp Verbreitungstyp Wertgebende Begleitart Charakteristische Arten

Charakter- art i.e.S.

Risikofaktoren Risiko

-IDkFG =WLa = gFG= Elb kurzfristig )

↓ ↓↓ ↓↓ ↓↓ ( langfristig

Kriterien

Bestand Schutzstatus , 1848) es < , 1976) ss < = = , 1850) s < = - D ., 1897) ss < = = ELVE , 1876 es = = = , 1946) ss = ., 1947 es < = = ., 1802) mh = = = UBÉ S ., 1837) mh = = = EY RUNNE ., 1837) ex (A ., 1802) h = = = RAV ., 1802) s = ., 1846) mh = = =

Taxon (B ANGLB ., 1910 ss < = - ID fDÜ SzW, KüN ., 1802) s << = - ID Mgr CaH, BDü ., 1837) ss < (G TRAND RAV OL CHEERP ORDM RAV . & R HP (G (S RAV S AUDI DI ORDM S (G (K (N (G ULS (G (B (N ORDM M (N (= Philonthus pseudoparcus Brunne, 1976) (= Philonthus spermophili Ganglb., 1897) 1802)) (= Philonthus fimetarius (Grav., (= Philonthus puella Nordm., 1837) 1802)) (= Philonthus nitidulus (Grav., 1806)) (= Gabrius vernalis (Grav., (= Gabrius velox sensu FHL Bd. 4 )

Bisnius pseudoparcus Bisnius spermophili Bisnius fimetarius Bisnius puella Bisnius nitidulus Gabronthus thermarum Gabrius osseticus Gabrius exiguus Gabrius astutoides Gabrius lividipes Gabrius splendidulus Gabrius trossulus Gabrius piliger Gabrius keysianus Gabrius nigritulus Gabrius austriacus

Grund Kat. Änd. Kat. Grund K K

↓ ↓ ↓

Kat +/- Kat RL 2010 RL 2+MK 1-R 3-MK 3 2-R 2-RK 3+MK 0 2+MK 2- M * * * * V-R R+ M

64 1932 1991 - I-ID CaH, ErH = = = = = ) ) ) ) ↑ ↓ ↓ ↓ ↓ ↓↓ ↓↓ ( ( ( ( , 1774) ss << , 1785) mh = = = ., 1851 ss << = = , 1774) s < = - ID EER (F., 1793)(F., ss << = - ID Mgr EER , 1910 h = = = G EMM ., 1806) s < (F., 1801)(F., ex G E ., 1795) mh = = = ., 1763) mh < G L ., 1798 ex EOFFROY , 1835) h = = = L., 1758 mh << = = CaH, ErH, NMo E ., 1763) s = = = (D HARP RAV , 1790) mh > (O ED COP (G (D S , 1781) s < = = ., 1906) ss < = - ID SKü C (G ., 1802) ss < COP (L., 1758) mh < (S PERK (L., 1758) mh = = = OSSI (S ., 1764) mh = = = (S RAV , 1913 es <<< = = (F., 1781)(F., mh = = = (R ERNH ., 1802) mh = = = OY (G ÜLL CHRANK (B J (S RAV (M (L., 1758) es << (G (= Gabrius pennatus Shp., 1910) (= Gabrius subnigritulus Rtt., 1909 n. nudum) pubescens sensu FHL Bd. 4 ) (= Trichoderma (= Ocypus hypsibatus (Bernh., 1899)) (= Ocypus similis semialatus sensu FHL Bd. 4 ) 1802)) (= Ocypus ater (Grav., (= Ocypus compressus (Marsh., 1802)) (= Ocypus winkleri (Bernh., 1906))

Gabrius breviventer Gabrius appendiculatus Gabrius toxotes Creophilus maxillosus Ontholestes tessellatus Ontholestes murinus Emus hirtus Platydracus fulvipes Platydracus latebricola Platydracus chalcocephalus Platydracus stercorarius Dinothenarus pubescens Staphylinus erythropterus Staphylinus caesareus Staphylinus dimidiaticornis Ocypus olens Ocypus ophthalmicus Ocypus nitens Ocypus brunnipes Ocypus fuscatus Ocypus picipennis Ocypus aeneocephalus ater Tasgius morsitans Tasgius winkleri Tasgius K K ↑ ↓ ↓ ↓ ↓ +M +M +R 0 3 2- M 0 1- M 2 1-R 1-RK 3+MK 2 1-R * * * * * * * * * * V-R V+ M V-MK V-R

65

Anmerkung

Letzter Nachweis Letzter Verantwortung SH Verantwortung

**

Verantwortung D Verantwortung

in SH in Verbreitungstyp Verbreitungstyp Wertgebende Begleitart Charakteristische Arten

Charakter- art i.e.S.

Risikofaktoren Risiko

-D -ID nWZ1 SWKü =SzW = =sö kurzfristig ) ) ) )

↓ ↓ ↓ ↓ ↓↓ ( ( ( ( langfristig

Kriterien

Bestand Schutzstatus ., 1806) s = = = ., 1979 ss < = = RAV ., 1856 ss < = - ID nWZ1 (G AB ., 1832) ss < = = ., 1876 s = = = ., 1802) ss < ., 1802) mh = = = ., 1867) mh = = = ., 1785) ss < = = ., 1860 s = = = SRAELS I AUV RAV RAV , 1839) mh = = = . & L ., 1839 ss < F OLL ÉNÉTR R ., 1800) ss < (G ., 1857 mh = = = (G HOMS

Taxon ., 1802) mh = = = ., 1840 es << ., 1847 ss < = - ID nWZ2 E W EER R ., 1924 s < = = R ., 1868 s << (M (T AIRM K AYK R EOFFROY E (H RAV RAV F K RID (P ., 1840 mh = = = (G G (G (F., 1787)(F., s < = - ID nWZ2 R G E (= Ocypus melanarius (Heer, 1839)) (= Ocypus melanarius (Heer, 1785)) (= Ocypus globulifer (Geoffroy., (= Quedius ventralis (Arag., 1830))

Tasgius melanarius Tasgius globulifer Tasgius Heterothops binotatus Heterothops praevius Heterothops niger Heterothops stiglundbergi Heterothops quadripunctulus Heterothops dissimilis Heterothops minutus Euryporus picipes dilatatus Velleius Quedius brevis Quedius truncicola Quedius infuscatus Quedius microps Quedius lateralis Quedius longicornis Quedius ochripennis Quedius nigrocaeruleus Quedius puncticollis Quedius invreae

Grund Kat. Änd. Kat. Grund K

↓ ↓ ↓ Kat +/- Kat

+M RL 2010 RL 2 2-R 2-RK 2 3 3 3 2 2-R 1-R 3+ M * * * * * * * * * V+ M

66 [68] * = SzW= KüO = ) ) ) ↓ ↓ ↓ ( ( ( ., 1802) h = = = ARSH , 1848 es << (M ., 1802) ss < = - ID ELVE ., 1833) s = = = S ., 1861 s < = - ID SzW Kü ., 1860 s < = - D nWZ2 ., 1908 s < = - ID ARSH ., 1806) mh = = = ., 1806) mh = = = ., 1802) h = = = ., 1850 es << = - ID QuB , 1832) s < = - ID Mgr ., 1806) s < ., 1830) s = = = ., 1833) mh = = = TEPH (M ., 1830) mh < = - D ., 1845 ss < = = kFG QuB, RAV RAV AIRM , 1960 ss < RAV ., 1839 mh < = - D ., 1843 es = = = F ERNH (S HOMS ., 1795) h = = = ., 1790) mh = = = RAV ., 1839 mh = = = R IESW (G (G ., 1857 s < = - I BDü, Mgr AUDI DI ., 1839 mh = = = L ., 1806) s < = - D nWZ2 B T R (G RULLÉ E K R TEPH R ANNH B IESW (G AHLB E K AYK ORGE (O E ELLN K (B (S RAV (F., 1793)(F., ss < = - D K (S K (P (M (G (= Quedius assimilis (Nordm., 1837)) 1802) n. nudum) (= Quedius tristis (Grav., (= Quedius hispanicus Bernh., 1898) (= Quedius pseudoumbrinus Lohse, 1958) (= Quedius humeralis sensu FHL Bd. 4 )

Quedius fulgidus Quedius cruentus Quedius brevicornis Quedius mesomelinus mesom. Quedius maurus Quedius xanthopus Quedius scitus Quedius cinctus Quedius fuliginosus Quedius curtipennis Quedius levicollis Quedius balticus Quedius simplicifrons Quedius molochinus Quedius picipes Quedius umbrinus Quedius nigriceps Quedius suturalis Quedius maurorufus Quedius nemoralis Quedius fumatus Quedius auricomus Quedius scintillans Quedius lucidulus Quedius riparius Quedius semiaeneus Quedius semiobscurus K ↓ ↓ +M 3 2 3-R 3 3 2-RK 2+MK 3 3+MK 3 3+MK 1 1-R * * * * * * * * * * * V-RK V+MK R-MK

67

Anmerkung Letzter Nachweis Letzter

1957

Verantwortung SH Verantwortung

Verantwortung D Verantwortung

in SH in Verbreitungstyp Verbreitungstyp Wertgebende Begleitart Charakteristische Arten

Charakter- art i.e.S.

Risikofaktoren Risiko

-ID kFG = =HMo kurzfristig )

↓ ↓↓ ↓↓ ( langfristig

Kriterien

Bestand Schutzstatus , 1875 s < = - ID ., 1850 ss << = = HMo EY , 1965 s < = - ID , 1969 ss < = = , 1876 s < = - ID Mgr ., 1806) h = = = IESW ., 1784) ex ., 1895 ss < = = BDü, Mgr [69] ., 1835 s = = = ., 1830 mh = = = EY K AGEL . & R RAV ., 1839) es < F AC BST ., 1806) h = = = R L ., 1923 ss < TRAND (G ., 1810) mh = = = ., 1833) mh = = = (H ULS ANNH S ., 1833) ss = (E ANGLB . & R RAV M G M YLL

Taxon UNST (G ULS TEPH ., 1802) mh = = = TEPH (G M (S M (S RAV (G (= Quedius aridulus Jansson, 1939) (= Mycetoporus baudueri sensu FHL Bd. 4 ) (= Mycetoporus hellieseni Strand, 1950) (= Mycetoporus brunneus (Marsh., 1802)) 1857) (= Mycetoporus ruficornis Kr., (= Mycetoporus bimaculatus sensu FHL Bd. 4 )

Quedius nitipennis Quedius fulvicollis Quedius boopoides Quedius persimilis Quedius boops Acylophorus wagenschieberi Acylophorus glaberrimus Atanygnathus terminalis Habrocerus capillaricornis pilicornis Trichophya Mycetoporus mulsanti Mycetoporus erichsonanus Mycetoporus baudueri Mycetoporus lepidus Mycetoporus longulus Mycetoporus bimaculatus Mycetoporus despectus

Grund Kat. Änd. Kat. Grund ↓ ↓ ↓ Kat +/- Kat

+M RL 2010 RL 2-R 3+MK 2-R 2 3 3 3 3 1 0-R * * * * * * *

68 1965 ** -IDNMo = ) ↑ ↓ ( , 1968 ss < = = WAQ AGEL ., 1856 s = , 1847) s = = = F , 1847) s << = = NMo ., 1832) mh = = = ., 1832) s = = = ., 1806) mh = = = AB , 1925) ex ., 1835 es < = = ., 1875 es << = - ID ., 1830 mh = = = RAV ., 1806 s < = - ID TEPH , 1835) es = = = ÄKLIN ÄKLIN TEPH AUV TEPH (G . & L (S ., 1806) ss << ELLÉN F (S RAV (M (M S ANNH ., 1839) mh = = = PERK G (H ., 1839) mh = = = R RAV ., 1794) es < = - D nWZ1 R M (F., 1777)(F., mh = = = L AIRM (S (E (L., 1760) mh = = = (G F (E (O (= Mycetoporus splendens (Marsh., 1802)) (= Mycetoporus aequalis Thoms., 1868) (= Mycetoporus brucki sensu FHL Bd. 4 ) (= Mycetoporus bergrothi Hell., 1925) (= Mycetoporus longicorne Maekl., 1847) 1806)) (= Mycetoporus splendidum (Grav., (= Bryoporus crassicornis (Mäkl., 1847)) 1777)) (= Bolitobius thoracicus (F., 1839)) (= Bolitobius exoletus (Er., 1839)) (= Bolitobius trinotatus (Er., (= Bolitobius lunulatus (L., 1760)) (= Bolitobius striatus (Ol., 1794)) (= Bryocharis cingulata Mannh., 1830)

Mycetoporus nigricollis Mycetoporus glaber Mycetoporus forticornis Mycetoporus clavicornis Mycetoporus niger Mycetoporus rufescens Mycetoporus eppelsheimianus Mycetoporus punctus Ischnosoma bergrothi Ischnosoma longicorne Ischnosoma splendidum Bryoporus cernuus Bryophacis crassicornis Lordithon thoracicus Lordithon exoletus Lordithon trinotatus Lordithon lunulatus Carphacis striatus Bolitobius cingulatus K ↑ ↓ ↓ * +R +M 3+ M 0-R 2+ M 3 3 1-R 1 1 * * * * * * * * * * R-M

69

Anmerkung

Letzter Nachweis Letzter

Verantwortung SH Verantwortung

Verantwortung D Verantwortung

in SH in Verbreitungstyp Verbreitungstyp Wertgebende Begleitart Charakteristische Arten

Charakter- art i.e.S.

Risikofaktoren

Risiko

kurzfristig langfristig

Kriterien

Bestand Schutzstatus , 1832) ss < = - ID ., 1832) mh = = = ., 1802) mh = = = ., 1802) s < = - D nWZ2 ., 1832) mh = = = TEPH ., 1806) s = = = RAV RAV , 1902) ss < = - ID sö TEPHENS (S ., 1832) mh = = = TEPH (G (G RAV (S (S UZE (F., 1793)(F., mh = = = (G TEPH (L., 1758) h = = = (L (F., 1775)(F., sh = = =

Taxon ., 1839 mh = = = (S R (F., 1781)(F., h = = = (L., 1767) mh = = = E (= Bryocharis analis F., 1789 nec 1787) (= Bryocharis analis F., 1806)) (= Bolitobius inclinans (Grav., (= Conosoma littoreus (L., 1758)) 1792)) (= Conosoma testaceus (F., (= Conosoma marshami (Steph., 1832)) (= Conosoma immaculatus (Steph., 1832)) 1802)) (= Conosoma pedicularius (Grav., (= Conosoma obtusus (Luze, 1902)) (= Sepedophilus lividus (Erichson, 1839)) 1802)) (= Conosoma bipunctatus (Grav.,

Bolitobius castaneus Parabolitobius inclinans Sepedophilus littoreus Sepedophilus testaceus Sepedophilus marshami Sepedophilus immaculatus Sepedophilus pedicularius Sepedophilus obtusus Sepedophilus nigripennis Sepedophilus bipunctatus nitidulus Tachyporus obtusus Tachyporus solutus Tachyporus hypnorum Tachyporus

Grund Kat. Änd. Kat. Grund

Kat +/- Kat RL 2010 RL 2 2+MK 3 * * * * * * * * * * *

70 1966 1909 - ID CaH, Mgr sö ==NMo SzW Kü = =WAQ = =HMoNMo ) ) ) ) ↑ ↓ ↓ ↓ ↓ ↓↓ ↓↓ ( ( ( ( ., 1869 ss < = - ID CaH, Mgr , 2006 es = AND ., 1876 es << = = sö ., 1806 mh < = = ., 1843 ss < P ., 1806) s << = - ID BDü, CaH, Mgr (L., 1758) h = = = RAV RAV ., 1832 mh = = = AHLB ., 1800) s << G CHÜLKE ANNH ., 1806 mh = = = ., 1802 h = = = ., 1810 ex S ., 1802 s < ., 1810 es << ., 1802 h = = = (G ., 1789) h = = = S (L., 1758) mh = = = ., 1832) mh = = = ., 1839 ss = M AYK ., 1802 mh = = = ., 1806 mh = = = (F., 1793)(F., ex TEPH ., 1855 s = = = RAV R RAV (F., 1781)(F., mh = = = YLL YLL ., 1839 ss < RAV RAV (P R AYK S R E G G G G G G RAV K RAV E (L., 1758) mh = = = TEPH (P G G (L., 1758) sh = = = (S (L., 1767) mh = = = (= Tachyporus signifer sensu FHL Bd. 4 ) (= Tachyporus macropterus Steph., 1832) (= Tachyporus macropterus sensu FHL Bd. 4 ) (= Tachyporus 1802) signatus Grav., (= Tachinus (= Leucoparyphus silphoides (L., 1767))

Tachyporus chrysomelinus Tachyporus dispar Tachyporus tersus Tachyporus atriceps Tachyporus quadriscopulatus Tachyporus transversalis Tachyporus pulchellus Tachyporus pusillus Tachyporus scitulus Tachyporus corpulentus Tachyporus Lamprinodes saginatus lignorum Tachinus humeralis Tachinus proximus Tachinus subterraneus Tachinus bipustulatus Tachinus pallipes Tachinus fimetarius Tachinus rufipes Tachinus laticollis Tachinus marginellus Tachinus corticinus Tachinus rufipennis Tachinus elongatus Tachinus Coproporus immigrans Cilea silphoides Deinopsis erosa Gymnusa brevicollis K ↓ ↓ ↓ ↓ * 2+MK 2-RK 2 0-R 2-R 2 2+ M 2-R 1-R 3 0 * * * * * * * * * * * * * * * * R-M

71 Anmerkung

[70] [70] Letzter Nachweis Letzter

1914 1945 Verantwortung SH Verantwortung

* **

Verantwortung D Verantwortung

in SH in Verbreitungstyp Verbreitungstyp Wertgebende Begleitart Charakteristische Arten

Charakter- art i.e.S.

Risikofaktoren Risiko

- ID-D SbW gFG KüN =HMo = kurzfristig ) ) )

↑ ↓ ↓ ↓ ( ( ( langfristig

Kriterien

Bestand Schutzstatus , 1870) ss < EY ., 1838) s < = - I . & R ., 1838) ss < = - I kFG ., 1845 ex ., 1838) s < = - I ATTH ., 1835) mh = = = ., 1837 mh = = = ULS ., 1837 ex

Taxon ., 1806) s = = = ., 1806) mh = R ., 1853 s < = - I R ATTH AC ., 1830 mh = = = IESW ., 1858 mh = = = (M ., 1806) s = = = ., 1837) ss < = - D nWZ2 E R (M E ., 1871 es << ATTH K R ., 1837) ss < = - ID SbW gFG Kü (L RAV RAV K (M ., 1837 s < HP RAV (E R AL (G (G (M IESW S ANNH ., 1862 mh = = = E (G K (H R M K (= Diglotta submarina sensu FHL Bd. 5 ) (= Diglotta mersa sensu FHL Bd. 5 ) (= Oligota flavicornis (Lac., 1835) 1837)) (= Oligota apicatus (Er.,

Gymnusa variegata Myllaena dubia Myllaena intermedia Myllaena kraatzi Myllaena elongata Myllaena brevicornis Myllaena gracilis Myllaena minuta Myllaena infuscata Diglotta mersa Diglotta sinuaticollis Pronomaea rostrata Oligota granaria Oligota parva Oligota inflata Oligota pusillima Oligota pumilio Holobus flavicornis Holobus apicatus

Grund Kat. Änd. Kat. Grund ↓ Kat +/- Kat

+M RL 2010 RL 3 3 1 0-R 2 0 2 3-RK 2-RK 3-RK 2 * * * * * * * *

72 1961 -D -D =nWZ2 = = = = =WLa ) ) ) ) ) ) ↑ ↑ ↓ ↓ ↓ ↓ ↓ ↓ ( ( ( ( ( ( , 1939 es = = = ., 1832 s = = - I , 1935 s = ., 1867 s = = = ., 1937 h = = = ., 1802) s < , 1866 ex ., 1806) mh = = = TRAND , 1839 ss << = = WLa , 1924 ss < = - D ., 1830 mh = ., 1830) mh = = = S TEPH , 1966 es = = = WLa ., 1857) es = = = ÜSTH RAV S ., 1837 es = HOMS EER ARSH ., 1837 ss = TRAND ., 1837 mh = ., 1800) h = = = ., 1800) s << = = ., 1839 mh = = = R T R R H W (G S ., 1839) ss < R ROTCH ANNH E ANNH (M E R E E C AYK ., 1839) mh = = = AYK VORAK , 1902) es = = = ., 1863) es < = = M OTSCH ENDELER R ., 1839) s < (E (M (P D (P R (E RIS W (M UZE (E (B (L (= Hypocyphtus longicornis (Payk., 1800)) (= Hypocyphtus laeviusculus Mannh., 1830) (= Hypocyphtus tarsalis Luze, 1902) 1839) (= Hypocyphtus discoideus Er., (= Cypha nitida (Palm, 1936)) (= Cypha hanseni (Palm, 1949)) 1839) (= Hypocyphtus pulicarius Er., (= Hypocyphtus punctum Motsch., 1857)

Cypha longicornis Cypha laeviuscula Cypha tarsalis Cypha discoidea Cypha apicalis Cypha seminulum Cypha pulicaria Cypha punctum Hygronoma dimidiata Encephalus complicans Gyrophaena pulchella Gyrophaena affinis Gyrophaena rousi Gyrophaena nana Gyrophaena gentilis Gyrophaena poweri Gyrophaena minima Gyrophaena congrua Gyrophaena williamsi Gyrophaena fasciata Gyrophaena bihamata Gyrophaena lucidula Gyrophaena transversalis Gyrophaena joyi Gyrophaena joyioides K K K K ↓ ↓ ↓ ↓ ↓ 3-R 2-RK 2- M 2- M 2+MK 3-R 1+ K 0-R 3-MK 3-R * * * * * * * * * V-RK V-R R- M R- M R- M R+ M

73 Anmerkung

[71] Letzter Nachweis Letzter

1961 Verantwortung SH Verantwortung

** ** ***

Verantwortung D Verantwortung

in SH in Verbreitungstyp Verbreitungstyp Wertgebende Begleitart Charakteristische Arten

Charakter- art i.e.S.

Risikofaktoren Risiko

- D nWZ2 -D = = kurzfristig ) ) )

↑ ↓ ↓ ↓ ( ( ( langfristig

Kriterien

Bestand Schutzstatus , 1859) ex ., 1839) mh = = = , 1838) mh = = = R ., 1832) es = = = , 1922) s = = = (E , 1830) s = = = CRIBA ., 1941 ss = ALTL ., 1830) mh = = = , 1845 s = = = TEPH ., 1934 ss = (S ., 1802) es = = - D ., 1839 s = = = (W (S

Taxon R ., 1838 ss < = - D StD Kü ., 1839 s = = = ODERO ., 1802) mh = = = R ., 1810) mh = = = JÖB E RAV ANNH ., 1859 ss < = - D StD Kü TRAND ÄKLIN E S ., 1837 ss < R URT ., 1850) es = = = KüN (D S (L., 1758) mh ? R ., 1837) ss < = - D WAQ (G YLL RAV K M (M C R E ANNERHEIM (G (G (E IESW ., 1839 es = = = (M R (K E (= Gyrophaena angustata (Steph., 1832)) (= Pragensiella marchii (Dodero, 1922))

Gyrophaena manca Gyrophaena strictula Gyrophaena polita Gyrophaena boleti Agaricochara latissima Cyphea curtula Placusa depressa Placusa tachyporoides Placusa incompleta Placusa atrata Placusa pumilio Homalota plana Anomognathus cuspidatus Silusa rubiginosa Silusa rubra Phytosus spinifer Phytosus balticus Arena tabida Rhopalocerina clavigera Thecturota marchii Leptusa pulchella Leptusa norvegica Grund Kat. Änd. Kat. Grund K K

K K

↓ ↓ * * ↓ ↓ * Kat +/- Kat

+M +M +M RL 2010 RL 2 1-RK 3-R 2-R 2 0-R 2 * * * * * * * * * * * R-M R+ M R-M G-R

74 [8] 1938 1945 *** -- D D- D nWZ1 nWZ1 sö nWZ2 -ID- Mgr D RFk, StD Hgld = = = ) ) ) ) ) ) ) ↓ ↓ ↓ ↓ ↓ ↓ ↓ ↓↓ ( ( ( ( ( ( ( , 1920) es = = = ., 1839) s = = = R (E ., 1850) es = ., 1832) s < AMERON ., 1947 mh = = = ., 1806) es = = = (C ., 1856 s ? = - D ., 1802) es < IESW ., 1875 ss < R TEPH ., 1806) s < ., 1837 s < = - D kFG ., 1864 ex ., 1844 ex ., 1802) mh = = = , 1839 ss < (K HP K ., 1837 s < = - D kFG ., 1837 mh = = = RAV R RAV (S R R S ., 1802) h = = = ., 1839) mh = = = HP ., 1795) mh = = = E CHEERP RAV (G E L E R RAV ., 1839) mh = = = (G EER ÄRK S ., 1837 ss < S R RAV R (G H (E (G ., 1837 ss < (O M E (E R (G E (= Bolitochara lunulata (Payk., 1789)) 1944) (= Autalia brevicornis Blair, (= Falagria sulcata (Payk., 1789)) (= Falagria concinnus ) (= Falagria thoracica Steph., 1832) 1802)) (= Falagria nigra (Grav., (= Bohemiellina paradoxa Mach., 1941) (= Xenusa lohmanderi Bernh., 1927) 1837)) (= Ischnopoda constricta (Er., 1837)) (= Ischnopoda coarctata (Er.,

Leptusa fumida Leptusa ruficollis Euryusa castanoptera Euryusa optabilis Euryusa sinuata Bolitochara obliqua Bolitochara bella Bolitochara mulsanti Bolitochara pulchra Autalia impressa Autalia longicornis Autalia rivularis Autalia puncticollis Cordalia obscura Falagria sulcatula Falagria caesa Myrmecocephalus concinnus Falagrioma thoracica Anaulacaspis nigra Bohemiellina flavipennis Myrmecopora sulcata constricta Tachyusa coarctata Tachyusa K K K ↓ ↓ ↓ ↓ ↓ +M 2+MK 2 3 1-R 1 3 3 2-R 3-R 0-R 0-R 2 * * * * * * * * R- M R- M G-RK

75

Anmerkung

Letzter Nachweis Letzter Verantwortung SH Verantwortung

* *

***

Verantwortung D Verantwortung

in SH in Verbreitungstyp Verbreitungstyp Wertgebende Begleitart Charakteristische Arten

Charakter- art i.e.S.

Risikofaktoren Risiko

-D kFG =kFG = == = =NMo NMo, Röh, SzW =NMo =NMo =NMo kurzfristig ) ) ) ) ) ) ) ) ) )

↓ ↓ ↓ ↓ ↓ ↓ ↓ ↓ ↓ ↓ ( ( ( ( ( ( ( ( ( ( langfristig

Kriterien

Bestand Schutzstatus , 1981 es = = = NMo ., 1876) es = ENICK ., 1869) s < B ., 1802) es = = = kFG sö AHLB , 1981 es = HP ., 1830) mh < (S (S ., 1837) es = = = kFG ., 1837) ss < ., 1837) s = = = ., 1860) es = R ., 1802 s = = = ARSH ALM R , 1866) ss < R ., 1839) s < = = ., 1837) s < P ., 1844) s ? = - D kFG (E ANNH

Taxon RIS (E R R (E YE (M ., 1939 s < = = RAV ., 1806) mh = = = (B (E (E G (R (M ., 1837 ss < IESW RAV R OTTH E (K ., 1856) s < (G T R (K (= Ischnopoda scitula (Er., 1837)) (= Ischnopoda scitula (Er., 1806)) atra (Grav., (= Tachyusa (= Atheta fleischeri (Epph., 1892)) (= Schistoglossa benicki Lohse, 1981)

Tachyusa scitula Tachyusa Thinonoma atra Ischnopoda leucopus Ischnopoda umbratica Dasygnypeta velata Gnypeta ripicola Gnypeta carbonaria Gnypeta rubrior Brachyusa concolor Dacrila fallax Callicerus obscurus Schistoglossa viduata Schistoglossa drusilloides Schistoglossa gemina Schistoglossa pseudogemina Schistoglossa curtipennis Schistoglossa bergvalli Schistoglossa aubei Boreophilia eremita

Grund Kat. Änd. Kat. Grund K K K

↓ ↓ ↓ ↓ ↓

Kat +/- Kat RL 2010 RL 3-R 2 2 2-R 3 1 1-R 1 3-R * * * V-MK V-R V-MK R+ M R- M R- M G

76 1945 -IDkFG -D kFG -ID WLa = =WLa ) ) ) ) ) ↓ ↓ ↓ ↓ ↓ ( ( ( ( ( , 1853) mh = = = , 1852) es = = = EY ., 1830) s = = = , 1959) es = = = ElbO EY , 1988 ex , 1873 es = = = , 1954) es = = = ., 1854) ss < = = EY OHSE ANNH . & R R ., 1830) mh = = = , 1861) es = = = OHSE , 1839) es << ., 1864) ss < = = ., 1832) ss < . & R ., 1832) mh = = = (K EY ., 1844) ss < L (M ., 1856) s < = - ID kFG ULS ., 1806) mh = = = & L ., 1839) s = = = ., 1913) es = , 1952) ss < RUNDIN .R EER TRH . & R ULS R ANNH ., 1837) mh = = = ., 1869) s = = = ÄRK TEPH ., 1837) mh = = = (M ., 1839) sh = = = R TEPH RAV (B (H (E R HP AST R ULS (W (M (S , 1841) mh = = = ., 1802) sh = = = (M ULS (E (S (M HOMS ERNH (G (E (L ENICK (S (E M (T (M (B RAV EER (B (G (H (= Hydrosmecta thinobioides (Kr., 1854)) (= Hydrosmecta thinobioides (Kr., 1839)) (= Atheta hepatica (Er., (= Amischa cavifrons (Shp., 1869)) 1856)) (= Amischa soror (Kr., (= Nehemitropia sordida (Marsh., 1802)) 1806)) (= Nothotecta flavipes (Grav., (= Nothotecta confusa (Märk., 1844))

Hydrosmecta longula Hydrosmecta paralongula Hydrosmecta subtilissima Aloconota planifrons Aloconota mihoki Aloconota sulcifrons Aloconota insecta Aloconota subgrandis Aloconota gregaria Aloconota languida Aloconota ultima Aloconota longicollis Aloconota coulsoni Enalodroma hepatica Pycnota paradoxa Amischa analis Amischa bifoveolata Amischa nigrofusca Amischa decipiens Amischa forcipata Amidobia talpa Nehemitropia lividipennis Notothecta flavipes Notothecta confusa anceps Lyprocorrhe Neohilara subterranea K K ↓ ↓ ↓ * ↓ +M +M +M 3+ M 2-R 3-RK 1-R 1-R 0-R 2 2 3 * * * * * * * * * * * * R+ M R- M R- M R-M R- M

77

Anmerkung Letzter Nachweis Letzter

1959 Verantwortung SH Verantwortung

*

Verantwortung D Verantwortung

in SH in Verbreitungstyp Verbreitungstyp Wertgebende Begleitart Charakteristische Arten

Charakter- art i.e.S.

Risikofaktoren Risiko

SWKü =SzW = == SzW = kurzfristig ) ) ) ) )

↓ ↓ ↓ ↓ ↓ ( ( ( ( ( langfristig

Kriterien

Bestand Schutzstatus , 1853) es < EY , 1977 es = , 1853) s < = - D SzP SzW, Kü ., 1886) es = = = . & R EY ., 1867) mh = = = OHSE PPH ., 1837) es < L ., 1802) es = = = R ULS ., 1837) s < = = ., 1810) mh = = = ., 1831) ss < = - D WLa sö (E ., 1806) sh = = = ., 1850) s = = = R ., 1856) es = = = (E . & R R ., 1839) es = = = sö (M

Taxon RAV ., 1802) s = = = (E YLL HOMS R RAV ., 1837) s < ., 1837) mh = = = ., 1839) s < = - ID BDü, Mgr (K IESW AHLB (G R ULS R R (T (E (G ., 1839) ex ., 1839) ss = (G RAV (F., 1798)(F., mh = = = R (K (S R (E (E (E (M (G (E (E (= Atheta marina (Muls. Rey, 1853)) (= Atheta marina (Muls. Rey, (= Atheta rudiventris Epph., 1886) (= Liogluta oblongiuscula (Sharp, 1869))

Alaobia scapularis Brundinia meridionalis Brundinia marina Dochmonota clancula Dochmonota rudiventris luteicornis Tomoglossa heydemanni Tomoglossa Ousipalia caesula Geostiba circellaris Dinaraea angustula Dinaraea aequata Dinaraea linearis Dinaraea arcana Paranopleta inhabilis Dadobia immersa Plataraea nigrifrons Plataraea brunnea Liogluta pagana Liogluta granigera Liogluta longiuscula Liogluta microptera

Grund Kat. Änd. Kat. Grund K

↓ ↓ ↓ ↓ Kat +/- Kat

+M RL 2010 RL 3-R 0-R 3 1 2 1-R 3 1-R * * * * * * * V-MK R- M R- M R- M R- M G

78 - ID gFG sö =kFG = =NMo = = = =WLa ) ) ) ) ) ) ↓ ↓ ↓ ↓ ↓ ↓ ↓↓ ↓↓ ( ( ( ( ( ( ., 1856) s < = = HOMS ., 1830) s < = = ., 1856) s < , 1873) s = = = ., 1940 es = (T ., 1852) s = = = ., 1856) s < = = WAQ EY ., 1867) ss = , 1954 ss < = = WAQ sö ., 1832) mh = = = ., 1802) mh = = = , 1913 ss < = = WAQ , 1932 es = = = ., 1856) ss < , 1839) h = = = ., 1806) ss < ., 1859) es = = = gFG Elb ., 1844) mh = = = ANNH R ERNH HOMS ., 1856) ss < = - ID HMo NMo, fDÜ , 1859) s < OY ., 1869) es < = = ., 1852) es = = - I gFG Elb ., 1856) ss = RAV RIS J B (K ., 1856) s = = = RAV EER (T HOMS HOMS TEPH R (M . & R HOMS HP , 1913 s < ., 1837) s < = = R ., 1909) ss = = = IESW (G ., 1837) s < = = (B R (T (G (T (H (S ., 1837) s = = = (K RUNDIN (T R (S LL OY (K ANSSEN CRIBA R HOMS (K ULS HOMS B (E J (E (E H ., 1837) s < = = (T (S (E (T R (M (E (= Liogluta nitidula (Kr., 1856)) (= Liogluta nitidula (Kr., (= Atheta similata Benick, 1975) 1913) (= Atheta nannion Joy,

Liogluta alpestris Atheta arctica Atheta elongatula Atheta hygrobia Atheta hygrotopora Atheta luridipennis Atheta terminalis Atheta grisea Atheta botildae Atheta tmolosensis Atheta gyllenhalii Atheta melanocera Atheta malleus Atheta volans Atheta obtusangula Atheta sequanica Atheta palustris Atheta debilis Atheta scotica Atheta parca Atheta deformis Atheta ripicola Atheta fallaciosa Atheta luteipes Atheta vilis Atheta occulta Atheta fungivora Atheta monticola Atheta excellens Atheta euryptera K K K K K ↓ ↓ ↓ ↓ ↓ ↓ ↓ +M +M 3+ M 3-R 2 3-R 2-RK 1-RK 3-R 3-R 3+ M 2-R 1-R 2-R 3 * * * * * * * * * V+ M V+ M V-MK V-MK V-MK V-MK R- M R- M

79

Anmerkung Letzter Nachweis Letzter

1944 1932

Verantwortung SH Verantwortung

Verantwortung D Verantwortung

in SH in Verbreitungstyp Verbreitungstyp Wertgebende Begleitart Charakteristische Arten

Charakter- art i.e.S.

Risikofaktoren Risiko

= = kurzfristig )

↓ ↓↓ ( langfristig

Kriterien

Bestand Schutzstatus , 1948 ex , 1948 ss = = = ., 1931 ss = , 1948 ss = = = ., 1867 es = = = ., 1950 mh = = = ., 1859) es = = =

Taxon ., 1856) s = = = ., 1802) s = = = , 1866) ss = = = ., 1930 mh = = = ., 1832) mh = = = , 1948 es = = = ., 1860) s = = = ., 1802) es < RUNDIN ., 1832) ex RIS ERNH ., 1869) es = = = RUNDIN ., 1856) es = = = ., 1844) s = = = ILL B RAV ., 1869) ss = = = RIS R ., 1837) es = = = HOMS B ., 1856) ss = = = B (B RAV , 1839) h = = = HP TEPH R RUNDIN T R W HP HOMS CRIBA (G (B TEPH (K CHEERP ÄRK (S B (G (S (E RUNDIN (K S (S (T (S EER (S B (M (H (= Atheta nigricornis (Thoms., 1852))

Atheta divisa Atheta vaga Atheta harwoodi Atheta liturata Atheta nigritula Atheta corvina Atheta sodermani Atheta nitella Atheta palleola Atheta benickiella Atheta amicula Atheta pittionii Atheta inquinula Atheta atricolor Atheta ganglbaueri Atheta subtilis Atheta atomaria Atheta minuscula Atheta glabricula Atheta indubia Atheta liliputana Atheta boreella Atheta cribrata

Grund Kat. Änd. Kat. Grund ↓ ↓ ↓ Kat +/- Kat

+M +M RL 2010 RL 3+MK 1-R 0-R 0-R * * * * * * * * * * * * * R+ K R- M R- M R-MK R-MK R-MK

80 = = ) ) ↓ ↓ ( ( , 1873) s = = = , 1848) mh = = = , 1873) mh = = = EY EY ., 1830) s = = = ., 1944 ss = = = ELVE ., 1856) mh = = = S ., 1856) s = = = ., 1830) s << = = ., 1806) h = = = ., 1850) s = = = . & R ., 1867 mh = = = ANNH . & R ., 1856) es << ., 1860) mh = = = ., 1869) s = = = ERNH ., 1856) ss < = = nWZ2 ., 1839) ss < ., 1856) h = = = ., 1869) s = = = RAV HOMS , 1938 es = = = ULS B (M RIS R HP R ULS ANNH HOMS IESW ., 1837) mh = = = HP (T (G ., 1839) s = = = ., 1856) mh = = = ., 1869) ss = = = ., 1837) s = = = R (E (S (B (K HOMS AUDI DI (T HOMS (M ., 1806) sh = = = R R (K R ., 1837) h = = = (M (S (M ., 1837) mh = = = T HP (E ., 1837)., 1837) h h ======HOMS R (T (B ENICK R (E (K (E ., 1856) mh = = = R R (S RAV (T (E R B (E (E (E (G (K (= Atheta griepi Scheerpeltz, 1954) (= Atheta delecta Benick, 1975) (= Atheta crebrepunctata Benick, 1940)

Atheta voeslauensis Atheta sodalis Atheta gagatina Atheta pallidicornis Atheta hybrida Atheta trinotata Atheta cadaverina Atheta subglabra Atheta serrata Atheta picipes Atheta melanaria Atheta clientula Atheta orphana Atheta orbata Atheta fungi Atheta negligens Atheta amplicollis Atheta subsinuata Atheta zosterae Atheta nigra Atheta dadopora Atheta canescens Atheta sordidula Atheta celata Atheta celata Atheta hypnorum Atheta castanoptera Atheta triangulum Atheta xanthopus Atheta graminicola ↓ ↓ * +M +M +M +M +M 2-R 3-MK 1-R 3 * * * * * * * * * * * * * * * * * * * * * * * * * R- M

81

Anmerkung

Letzter Nachweis Letzter

Verantwortung SH Verantwortung

Verantwortung D Verantwortung

in SH in Verbreitungstyp Verbreitungstyp Wertgebende Begleitart Charakteristische Arten

Charakter- art i.e.S.

Risikofaktoren Risiko

=QuB, kFG =QuB, = = kurzfristig ) )

↓ ↓ ↓↓ ( ( langfristig

Kriterien

Bestand Schutzstatus ., 1926 mh = = = , 1852) ss = EY CHEERP , 1873) es = = = EY , 1954 ex HMo 1960 ., 1856) ss < = - ID nWZ1 ., 1852) ss < ., 1856) s < = = , 1938 es = = = ., 1852) s = = = . & R . & S ., 1852) s < = - ID nWZ2 ., 1856) mh = = = ., 1952) s < = = ., 1867) s <

Taxon R ., 1869) mh = = = ., 1832) mh = = = . & R ., 1869) mh = = = ULS HOMS ENICK HOMS (K HOMS HP ., 1914) ss = = = (F., 1792)(F., sh = = = RUNDIN ERNH B HP (T HOMS ULS (T HOMS (T (S ., 1856) mh = = = HOMS B (M HOMS TEPH B OH R (S (T (T ., 1839) h = = = ., 1837) ss < = = ., 1839) mh = = = (T (T (M (S R (J R R (K (E (E (E (= Atheta pertyi sensu FHL Bd. 5 ) (= Atheta repanda sensu FHL Bd. 5 )

Atheta ebenina Atheta aquatilis Atheta incognita Atheta aquatica Atheta aeneicollis Atheta laticollis Atheta coriaria Atheta intermedia Atheta ravilla Atheta basicornis Atheta myrmecobia Atheta nidicola Atheta oblita Atheta boletophila Atheta strandiella Atheta pilicornis Atheta fungicola Atheta britanniae Atheta crassicornis Atheta macrocera Atheta puncticollis Atheta cauta

Grund Kat. Änd. Kat. Grund K

↓ ↓ ↓ Kat +/- Kat

+M RL 2010 RL 2 3 3 3+ M 3-R 0-R 2-R * * * * * * * * * * * V+ M V+ M R- M R- M

82 * ** =WLa = ) ) ↓ ↓ ( ( ., 1926 es < = - D KDü KüN CHEERP , 1852) s = = = ., 1858) ss < = = WLa ., 1933) es = = = ., 1856) s < , 1852) s = = = . & S EY EY AM ., 1860) ss < ., 1810) mh = = = ., 1802) s = = = ., 1870 s = = = ., 1830) mh = = = ., 1802) h = = = OTSCH (C ., 1802) mh = = = ., 1802) h = = = ERNH , 1940) ss = = = HOMS RIS . & R ., 1856) h = = = YLL RAV B ., 1856) mh = = = RAV (T (M . & R (B RAV RAV R (G ANNH (G ., 1837) s < = - ID BDü, Mgr HOMS ., 1869) s = = = ULS ., 1806) mh < = - D StD Kü (G LLEN ., 1837) s = = = R T (G (K (G ULS HP R HOMS (M (A (M (E RAV (S (E (T (M (G (= Atheta exigua (Er., 1837)) (= Atheta exigua (Er., 1856)) (= Atheta sylvicola (Kr., 1802)) (= Atheta pygmaea (Grav., 1802)) (= Atheta obfuscata (Grav., (= Acrotona consanguinea (Epph., 1875)) (= Acrotona rassii Muona, 1993) 1852)) (= Atheta parens (Muls.Rey, (= Atheta muscorum (Bris., 1860)) 1802)) (= Atheta aterrima (Grav., (= Acrotona pusilla (Brundin, 1952)) (= Atheta parvula (Mannh., 1831))

Atheta ischnocera Atheta setigera Atheta laevana Atheta nigripes Atheta atramentaria Atheta marcida Atheta cinnamoptera Atheta longicornis Atheta varendorffiana Atheta vestita Acrotona exigua Acrotona sylvicola Acrotona pygmaea Acrotona obfuscata Acrotona troglodytes Acrotona pseudotenera Acrotona parens Acrotona muscorum Acrotona aterrima Acrotona benicki Acrotona parvula K K * ↓ +M +M 1 3-R 3 3+ M 2 * * * * * * * * * * * * * * V-RK R-M

83 Anmerkung

[72]

Letzter Nachweis Letzter Verantwortung SH Verantwortung

* *

Verantwortung D Verantwortung

in SH in Verbreitungstyp Verbreitungstyp Wertgebende Begleitart Charakteristische Arten

Charakter- art i.e.S.

Risikofaktoren Risiko

- D StD Kü =NMoHMo kurzfristig ) )

↓ ↓ ( ( langfristig

Kriterien

Bestand Schutzstatus ., 1802) mh = = = RAV ., 1856) ss < = = ., 1844) s = = = ., 1861 s << = - D StD SbW Kü , 1984 mh = = = R (G ., 1839) es = = = WLa (K , 1851 s < R ÄRK

Taxon ., 1832) ss = = - I OHSE ., 1802) mh = = = (E HOMS (F., 1787)(F., h = = = (M ., 1842) s < = = WLa L ., 1837) mh = = = T ARDY ., 1806) s << = - D WLa R ., 1800) s << ., 1789) mh = = = RAV ., 1837) mh = = = TEPH H ÄRK (E R (G RAV (S AYK AYK (E (M (G (P (P (= Aleuonota rufotestacea (Kr., 1856)) (= Aleuonota rufotestacea (Kr., 1839)) (= Aleuonota gracilenta (Er., (= Halobrecta puncticeps (Thoms., 1852)) (= Zyras limbatus (Payk., 1789)) 1806)) (= Zyras funestus (Grav., 1802)) (= Zyras humeralis (Grav., (= Zyras cognatus (Märk., 1842))

Alevonota rufotestacea Alevonota gracilenta Halobrecta algae Halobrecta flavipes Alianta incana Pachnida nigella immigrata Trichiusa Thamiaraea cinnamomea Thamiaraea hospita Drusilla canaliculata Zyras collaris Zyras haworthi Pella limbata Pella funesta Pella humeralis Pella cognata

Grund Kat. Änd. Kat. Grund Kat +/- Kat

+M +M +M RL 2010 RL 2 3+ M 3 2-RK 2-RK * * * * * * * * V-MK R+ M G

84 =BDü == BDü, CaH ) ) ) ↓ ↓ ↓ ( ( ( ., 1789) s = = = CaH ., 1830) mh = = = ., 1900 es = = = sö (F., 1792)(F., es << ., 1806) es = = = BDü, CaH , 1850) es = = = ., 1802) h = = = ., 1914 mh = = = ., 1860 es < AYK AUV ., 1800) mh = = = ., 1884 mh = = = RAV ., 1856) es << (P F ANNH RIS UBÉ RAV ., 1843 ss < = = CaH ., 1830 mh = = = R ., 1802) es = = = sö ., 1856 s < = - I B (G ., 1806) es = = = CaH (A AYK R ., 1837 ss << = = (G (K (M PPH R ONISTH K RAV ., 1845) s < = - D WLa E (P IESW E RAV D ., 1837 s < = = ANNH ., 1802) s < = - D WLa K (G R ÄRK (G M E RAV (M (G (= Zyras lugens (Grav., 1802)) (= Zyras lugens (Grav., (= Zyras laticollis (Märk., 1842)) 1792)) (= Lomechusa strumosa (F., (= Atemeles emarginata (Payk., 1789)) 1806)) (= Atemeles paradoxa (Grav., (= Atemeles pubicollis Bris., 1860) (= Phloeopora corticalis sensu FHL Bd. 5 ) 1856)) (= Phloeodroma concolor (Kr., (= Phloeopora angustiformis sensu FHL Bd. 5 ) (= Phloeopora bernhaueri Lohse, 1984) (= Ilyobates subopacus Palm, 1935) (= Ilyobates hoelzeli Scheerp., 1947)

Pella lugens Pella laticollis Lomechusoides strumosus Lomechusa emarginata Lomechusa paradoxa Lomechusa pubicollis Dinarda dentata Dinarda maerkelii Phloeopora teres Phloeopora concolor Phloeopora testacea Phloeopora nitidiventris Phloeopora corticalis Phloeopora scribae Ilyobates bennetti Ilyobates nigricollis Ilyobates propinquus Calodera nigrita Calodera riparia Calodera uliginosa Calodera rufescens K K ↓ * * * 3 3+MK 2 3 3 1 1-R 1 * * * * * * * V+ M R+ M R- M R-M R- M R-M

85

Anmerkung

Letzter Nachweis Letzter Verantwortung SH Verantwortung

***

Verantwortung D Verantwortung

in SH in Verbreitungstyp Verbreitungstyp Wertgebende Begleitart Charakteristische Arten

Charakter- art i.e.S.

Risikofaktoren Risiko

=NMo kurzfristig )

↓ ( langfristig

Kriterien

Bestand Schutzstatus ., 1839) s < = = SKü, StU, kFG SzW, ., 1837) s < = = R R ., 1865) es = = = QuB ., 1835) es = = = , 1996 es = = = Elb (E ., 1830 ss < = - I gFG Elb (E ., 1905 es = = = ., 1837) es = = = AC AUV R ., 1802) mh = = = , 1954 es = = = SzW KüN (L

Taxon (F (E ., 1849) mh = = = SSING ., 1870 es = = = kFG ANNH ERNH RAV ., 1915 s < ., 1832) h = = = ., 1851 mh < = = A ., 1906) mh = = = ., 1837 ss < = = WAQ B HP M ENICK (G EDT R HP ILL S B E S RAV (R TEPH ., 1837 mh = = = M R (G (S E (= Ityocara rubens (Er., 1837)) (= Ityocara rubens (Er., 1837)) (= Parocyusa rubicunda (Er., 1839)) (= Parocyusa longitarsis (Er., (= Meotica hanseni Scheerp., 1954) (= Meotica lubecensis Benick, 1954)

Calodera aethiops Calodera protensa Calodera cochlearis Calodera rubens rubicunda Tetralaucopora longitarsis Tetralaucopora Amarochara umbrosa Amarochara bonnairei Amarochara forticornis Ocalea badia Ocalea picata Ocalea rivularis Ocalea latipennis Pentanota meuseli Meotica pallens Meotica winkleri Meotica exilis Meotica exillima

Grund Kat. Änd. Kat. Grund K K

* *

Kat +/- Kat RL 2010 RL 3+ M 3 2 * * * * * * V+ M V+ M R- M R-M R-M R- M R+ K R- M R+ M

86 -I - I BDü, CaH, Mgr = =QuB = = ) ) ) ) ) ↓ ↓ ↓ ↓ ↓ ↓↓ ( ( ( ( ( , 1852) s = = = , 1861 mh = = = , 1832) h = = = , 1832) mh = = = EY ., 1830) ss < = - I CaH, Mgr EY ., 1856) es = = = , 1861 es < EY ., 1960) mh = = = ., 1830 mh = = = AB ., 1844 s = = = ., 1867) es < ., 1839) es = = = , 1850 mh = = = ANNH . & R ., 1802) mh = = = R . & R ., 1837) es = = = ElbO TEPHENS ÄRK R TEPHENS (E (M UBÉ ., 1842 mh = = = . & R ANNH RAV . & L ULS ., 1802) h = = = (S ., 1856 s < OTSCH ULS (E M HOMS ., 1839 ss < = - I CaH, Mgr (S A , 1850) s < = = ., 1839 s = = = R R M ., 1837 ss < = = WAQ (G ., 1837 ss < R (T (M M ULS ÄRK RAV K ., 1856 es << ., 1837) s < = = R E (M R E AIRM R R E UBÉ ., 1856 ss < E M M (G K R (F (E (A K (= Meotica capitalis (Muls. & Rey, 1873)) (= Meotica capitalis (Muls. & Rey, (= Deubelia picina (Aubé, 1850)) (= Ocyusa nitidiventris sensu FHL Bd. 5 (?) ) (= Chilomorpha longitarsis (Thoms., 1867)) (= Oxypoda lividipennis sensu FHL Bd. 5 ) (= Oxypoda umbrata (Gyll., 1810)) (= Oxypoda huetheri Scheerp., 1956)

Meotica filiformis Ocyusa picina Ocyusa maura Cousya nigrata Cousya longitarsis Zoosetha inconspicua Mniusa incrassata Hygropora cunctans Oxypoda elongatula Oxypoda procerula Oxypoda opaca Oxypoda longipes Oxypoda vittata Oxypoda acuminata Oxypoda spectabilis Oxypoda funebris Oxypoda lentula Oxypoda induta Oxypoda brevicornis Oxypoda vicina Oxypoda abdominalis Oxypoda rufa Oxypoda togata Oxypoda exoleta Oxypoda alternans Oxypoda praecox K K ↓ ↓ ↓ ↓ ↓ * * +M 2 1-R 1-R 1-R 3 2-R 2-R 2 3 * * * * * * * * * * * * V-MK V-MK R+ K R-M R-M

87

Anmerkung

Letzter Nachweis Letzter

Verantwortung SH Verantwortung

Verantwortung D Verantwortung

in SH in Verbreitungstyp Verbreitungstyp Wertgebende Begleitart Charakteristische Arten

Charakter- art i.e.S.

Risikofaktoren Risiko

-D = = kurzfristig ) )

↓ ↓ ↓↓ ( ( langfristig

Kriterien

Bestand Schutzstatus , 1875 es = = = ElbO EY ., 1830) mh = = = ., 1806) s > = = nWZ2 ., 1827) s = = = ., 1841 s = = = ., 1832) h = = = ., 1832) mh = = = ., 1852) s = = = ., 1830) s < ., 1830) mh = = = ., 1844) es << ., 1842) es = . & R ., 1802) mh = = = RAV ., 1839) ss = = - D WLa YLL ÄRK ANNH R TEPH ., 1837) mh = = = (G (G ÄRK ÄRK TEPH ULS M RAV R AIRM ., 1856 s = = = (E

Taxon (M ., 1837) mh = = = ., 1856 s < = = (S ANNH ANNH R (S ., 1837 es = = = M (F R R (G (E ., 1802) mh = = = (M (M K R K (M (M (E E ., 1871 s ? ? = RAV HP (G S (= Oxypoda difficilis Roub., 1931) (= Oxypoda amoena Fairm. & Lab., 1856) (= Haploglossa pulla (Gyll., 1827))

Oxypoda testacea Oxypoda brachyptera Oxypoda tarda Oxypoda annularis Oxypoda flavicornis Oxypoda nigrocincta Oxypoda formiceticola Oxypoda recondita Oxypoda haemorrhoa Stichoglossa semirufa Ischnoglossa prolixa Dexiogya corticina Thiasophila angulata Thiasophila inquilina Crataraea suturalis Haploglossa gentilis Haploglossa villosula Haploglossa nidicola Haploglossa marginalis Haploglossa picipennis morion Tinotus

Grund Kat. Änd. Kat. Grund K K

* ↓ Kat +/- Kat

+M +M +M RL 2010 RL 1 1 3-R * * * * * * * * * * * * * V+ M R+ K R-M D G

88 1960 1961 =sö ======) ) ) ) ) ↑ ↑ ↓ ↓ ↓ ↓ ↓ ( ( ( ( ( , 1858 ex , 1864 s = = = , 1850 s < = = ., 1806 mh = = = RAATZ ., 1802 mh = = = ., 1858 s < = = ., 1830 s < ., 1856 ex , 1984 ss < = = , 1965 es = = = K ., 1802 mh = R ., 1827 ex R , 1833 es = , 1777) s << = = ., 1810 ss < UBÉ RAV ., 1830 s = ., 1810 mh = = = RAV K K ., 1837) mh = = = A ., 1802 es << G , 1839 sh = = = (L., 1761) h = = = RAV R G YLL ., 1856 s = = = OLLASTON YLL (L., 1758) ss < = = ., 1806 es < = = LUG ANNH YLL R G OEZE G ., 1862 es < = = (E G , 1836 ss < RAV K ANNH G EER W K M R IKOVSKY IKOVSKY RAV G AY (G H K L L M S G (= Aleochara ripicola Muls. & Rey, 1874) (= Aleochara ripicola Muls. & Rey, (= Aleochara diversa (Sahlb., 1876)) (= Aleochara albovillosa Bernh., 1901) 1837)) (= Rheochara spadicea (Er., (= Aleochara verna sensu FHL Bd. 5) 1874)) (= Aleochara pauxilla (Muls. & Rey,

Aleochara curtula Aleochara haematoptera Aleochara brevipennis Aleochara puberula Aleochara intricata Aleochara tristis Aleochara inconspicua Aleochara sparsa Aleochara stichai Aleochara kamila Aleochara funebris Aleochara lanuginosa Aleochara lygaea Aleochara villosa Aleochara sanguinea Aleochara fumata Aleochara moerens Aleochara laevigata Aleochara haemoptera Aleochara cuniculorum Aleochara spadicea Aleochara ruficornis Aleochara bilineata Aleochara binotata Aleochara bipustulata Aleochara verna K ↑ ↓ ↓ ↓ ↓ ↓ * +K +R 1-R 3-M 2 1 3+MK 2- M 3 0-R 0-R 3-MK 2 0-R 2-R * * * * * * * * * * V+ M V+ M R-MK

89

Anmerkung

Letzter Nachweis Letzter Verantwortung SH Verantwortung

* * **

Verantwortung D Verantwortung

in SH in Verbreitungstyp Verbreitungstyp Wertgebende Begleitart Charakteristische Arten

Charakter- art i.e.S.

Risikofaktoren Risiko

- D-D StD Kü sö =WLa kurzfristig ) ) )

↓ ↓ ↓ ( ( ( langfristig

Kriterien

Bestand Schutzstatus , 1777) s < = - I NMo ., 1858 mh << OEZE , 1774) s = = - D (G (L., 1758) mh = = = ., 1806 mh < = - D StD Kü ., 1784) ss < = - D nWZ1 sö EER OTSCH (L., 1767) es = = = sö ., 1829) s < = - D nWZ2 RAV

Taxon ., 1798) s < BST (G M ., 1807 s < = = G (H ., 1856 mh < = - D Kü AYK F., 1792F., h = = = HEVR (L., 1758) s << = = R ALL (P (C F K (F., 1787)(F., s < L., 1758 h = = = (= Polystomota grisea (Kr., 1856)) (= Polystomota grisea (Kr., 1806)) (= Emplenota obscurella (Grav., (= Polystomota punctatella (Motsch., 1858))

Aleochara grisea Aleochara obscurella Aleochara punctatella Dictyopterus aurora Pyropterus nigroruber Platycis minutus Platycis cosnardi sanguineus Lygistopterus Lampyris noctiluca Lamprohiza splendidula Phosphaenus hemipterus Podabrus alpinus Cantharis fusca Cantharis rustica Cantharis pellucida

Grund Kat. Änd. Kat. Grund K K

↓ ↓

Kat +/- Kat RL 2010 RL 3-MK 3-R 2+MK 3 3+MK 3-R 3 * * * V-RK V-RK V-RK V-MK R+ M Lampyridae (Leuchtkäfer, Glühwürmchen) Lampyridae (Leuchtkäfer, Cantharidae (Weichkäfer) Cantharidae (Weichkäfer) Lycidae (Rotdeckenkäfer) Lycidae

90 1952 = NMo= kFG, WAQ [73] ) ) ↓ ↓ ( ( ss << = = SzW Kü ., 1835) s < ., 1851 es = = = ALD ., 1785) mh = = = HARP IESW ., 1864 h = = = (F ., 1845) ss < = = kFG ., 1764) h = = = ., 1802) s < = = S K ., 1851 ex , 1871 mh = = = ., 1776) h = = = ., 1843 mh < = = ., 1864 ss < = = nWZ2 ., 1764) mh = = = ., 1807 s < ., 1763) sh = = = ÜLL , 1947 mh = = = ., 1790) s < = = OURCR ETZN , 1777 mh = = = , 1923 mh = = = ÜLL (L., 1758)HOMS mh = = = L ARSH AUDI UFFR ÜLL (L., 1758) s < = = IC (F (L T ALL (M COP S ANNH B SHE F., 1792F., mh < = = P (O F (M HOMS (M L., 1758 mh = = = OEZE (M A (S L., 1758 h = = = M T (L., 1767) es = = = G L., 1758 h = = = L., 1758 h = = = (F., 1775)(F., mh < = = (= Cantharis bicolor Hbst., 1784) (= Cantharis violacea Payk., 1798) (= Podistra rufotestacea (Letzn., 1845)) (= Malthinus flaveolus (Hbst., 1786))

Cantharis fulvicollis Cantharis thoracica Cantharis paludosa Cantharis obscura Cantharis lateralis Cantharis nigricans Cantharis decipiens Cantharis livida Cantharis rufa Cantharis rufa darwiniana Cantharis cryptica Cantharis pallida Cantharis figurata Ancistronycha cyanipennis Absidia rufotestacea Rhagonycha lutea Rhagonycha fulva Rhagonycha testacea Rhagonycha limbata Rhagonycha lignosa Rhagonycha atra Rhagonycha gallica Silis ruficollis Malthinus punctatus Malthinus seriepunctatus Malthinus balteatus Malthinus facialis Malthinus biguttatus Malthinus frontalis K * ↓ ↓ 3 3 3-R 3 0-R 2 * * * * * * * * * * * * * * * * * * V-MK V-MK V+ M R-M R- M

91

Anmerkung Letzter Nachweis Letzter

1865 Verantwortung SH Verantwortung

*

Verantwortung D Verantwortung

in SH in Verbreitungstyp Verbreitungstyp Wertgebende Begleitart Charakteristische Arten

Charakter- art i.e.S.

Risikofaktoren Risiko

======nWZ2 =WLa kurzfristig ) ) )

↓ ↓ ↓ ↓↓ ↓↓ ↓↓ ↓↓ ( ( ( langfristig

Kriterien

Bestand Schutzstatus , 1846) es << = - D sö ., 1852 ss << = - D ., 1806) mh = = = ., 1789) ss < ., 1852 ss < ÄKLIN IESW ., 1852 mh = = = ., 1852 ss < ., 1852 ss < ., 1835) s < = = , 1838) s = = = ATR K ., 1852 es = = = Elb ., 1852 s < ., 1840) es = = = ., 1824) s < ., 1790) es <<< = = ., 1852 es = = = (M AYK (L L

Taxon IESW IESW REB IESW (P AST IESW ALTL K K (L., 1758) mh = = = (O IESW IESW K (F., 1787)(F., es << = - D nWZ1 sö K ERM (B IESW (C ., 1812 ss < K K (W K (G (L., 1767) ex HR A

Malthodes debilis Malthodes flavoguttatus Malthodes dispar Malthodes fibulatus Malthodes misellus Malthodes maurus Malthodes fuscus Malthodes minimus Malthodes guttifer Malthodes marginatus Malthodes mysticus Malthodes pumilus Malthodes spathifer Malthodes lobatus Malthodes crassicornis Malthodes brevicollis Drilus concolor albicans Troglops cephalotes Troglops Hypebaeus flavipes

Grund Kat. Änd. Kat. Grund K

↓ ↓ ↓

Kat +/- Kat RL 2010 RL 3 2-R 2-R 3 1+ K 2 0 1+ K 1 2-R 2+MK 1+ K * * * * V-MK R- M R+ M R- M Drilidae (Schneckenhauskäfer) Zipfelkäfer) Malachiidae (Malachitkäfer,

92 1896 ** -ID - D nWZ1 sö = = =nWZ2 = ↓↓ ↓↓ ↓↓ ↓↓ ↓↓ ↓↓↓ , 1784) ss << ., 1802) ss < ., 1783) mh = = = ., 1832) mh > = = ., 1861) es << ., 1840) mh = = = ARRER ., 1790) ss < = - ID KDü KüO ARSH ., 1785) ex L CHALL ENETR (H , 1830 h = = = AST IESW ., 1776) h = = = ., 1790) s < = = ID WLa , 1774) mh = = = (M (S (F., 1792)(F., ss < = = sö (O L (L., 1758) h = = = ., 1798) mh = = = (M (K (C (L., 1758) s << = = ., 1840) es << = - ID KDü KüO ., 1790) es = = = (F., 1775)(F., mh = = = ÜLL L R (F., 1787)(F., mh = = = EER (O ., 1801) ss = OURCR (L., 1758) es << AYK (E (O (M (F., 1801)(F., es = = - ID sö (G LL (F (L., 1758) es << (P TEPHENS (I S (L., 1761) ss < = = (= Malachius marginellus Ol., 1790) 1787) (= Malachius viridis F., (= Haplocnemus impressus (Marsh., 1802)) 1792)) (= Haplocnemus nigricornis (F., (= Dasytes cyaneus ) 1867) (= Dasytes aerosus Kiesw.,

Charopus flavipes Charopus concolor Malachius aeneus Malachius bipustulatus Clanoptilus marginellus Cordylepherus viridis Anthocomus coccineus Anthocomus fasciatus Anthocomus bipunctatus Cerapheles terminatus Paratinus femoralis Sphinginus lobatus Ebaeus thoracicus Ebaeus pedicularius Axinotarsus ruficollis Axinotarsus pulicarius Axinotarsus marginalis Aplocnemus impressus Aplocnemus nigricornis memnonia Trichoceble Dasytes niger Dasytes caeruleus Dasytes plumbeus Dasytes aeratus Dasytes fusculus K ↓ ↓ ↓ ↓ * +M 3-MK 1 2 1 3-MK 1-R 2 3-MK 1-R 1-R 1 2-R 0 * * * * * * * * * * V+MK R-M Dasytidae (Wollhaarkäfer part.) Dasytidae (Wollhaarkäfer

93

Anmerkung

Letzter Nachweis Letzter

Verantwortung SH Verantwortung

Verantwortung D Verantwortung

in SH in Verbreitungstyp Verbreitungstyp Wertgebende Begleitart Charakteristische Arten

Charakter- art i.e.S.

Risikofaktoren Risiko

-D -D sö = = kurzfristig )

↓

↓↓ ↓↓ ( ↓↓↓ langfristig

Kriterien

Bestand Schutzstatus ., 1785) es << = - D KDü Hgld [74] ., 1830 s < = - D ., 1838) ss = = = , 1794) mh < = = OURCR , 1775) s < ., 1846 ss = , 1837) ss <<< (F ETT (L., 1758) mh = = = TEPH EER OSSI S

Taxon (Z , 1775) ss << = = OSH (G (R R TURM EER (L., 1758) es << (F., 1775)(F., es << = - D (S (L., 1758) s < = = (G (L., 1758) s < = - D nWZ2 (L., 1758) s << = - D nWZ2 (= Psilothrix cyaneus (Ol., 1790)) 1843) (= Opilo germanus Chevr., (= Thanasimus rufipes (Brahm, 1797))

Psilothrix viridicoeruleus Dolichosoma lineare Phloiophilus edwardsii elongatus Tillus Opilo mollis Opilo domesticus Thanasimus formicarius Thanasimus femoralis apiarius Trichodes Korynetes caeruleus Necrobia ruficollis Necrobia violacea Necrobia rufipes Laricobius erichsonii

Grund Kat. Änd. Kat. Grund K K

↓ ↓ Kat +/- Kat

+M RL 2010 RL 2 3-R 2-RK 1 1-RK 1+ K 1 2-R 3 3+MK * * * V-MK Derodontidae (Lärchenkäfer) Phloiophilidae (Winter-Rindenkäfer) Phloiophilidae (Winter-Rindenkäfer) Cleridae (Buntkäfer)

94 -D - D UWR1, nWZ1 = ) ) ↓ ↓ ↓↓ ( ( ., 1776) s << = = ., 1821) ss < = - D nWZ2 ., 1829) ex nWZ1 1948 , 1777) s < = - D CHRK ., 1865) es < , 1925) s < ., 1830) ss < = - D nWZ1 öH ÜLL ., 1830) ss < = = HMo (L., 1761) mh = = = (L., 1758) ss << (S ., 1835) ss < = - D nWZ2 ., 1784) mh = = = SCHZ (M OEZE ., 1784) s < = - D ORN (L., 1761) mh = = = (E BST TEPH (F., 1787)(F., es = = = nWZ2 sö TEPH (L., 1758) mh = = = CHDTE (G (D BST (L., 1758) ex nWZ1 1887 (F., 1792)(F., ex nWZ1 1924 (S (H (S (S ACORD , 1905) s << = - D nWZ1 (L., 1758) mh = = = (F., 1787)(F., es < = - DW nWZ1 (H (L., 1758) ss < = - D nWZ2 L., 1758 ex UWR2, nWZ1 1951 (L (L., 1758) mh = = = YE (R

Nemosoma elongatum mauritanicus Tenebroides Thymalus limbatus Grynocharis oblonga Hylecoetus dermestoides navale Lymexylon Ampedus erythrogonus Ampedus rufipennis Ampedus balteatus Ampedus praeustus Ampedus cardinalis Ampedus sanguineus Ampedus cinnabarinus Ampedus pomonae Ampedus sanguinolentus Ampedus pomorum Ampedus hjorti Ampedus triangulum Ampedus nigroflavus Ampedus elongatulus Ampedus nigrinus Procraerus tibialis Elater ferrugineus Sericus brunneus ↓ ↓ ↓ +M 2 3 3 3 2 2 3 1-R 0 0-R 0 0-R 3 1 2 1-RK 2 * * * * * * R- M Lymexylonidae (Werftkäfer) (Werftkäfer) Lymexylonidae Lophocateridae (Flachkäfer) Trogositidae (Flachkäfer, Jagdkäfer) (Flachkäfer, Trogositidae Peltidae (Flachkäfer) Elateridae (Schnellkäfer)

95

Anmerkung Letzter Nachweis Letzter

1954

Verantwortung SH Verantwortung

Verantwortung D Verantwortung

in SH in Verbreitungstyp Verbreitungstyp Wertgebende Begleitart Charakteristische Arten

Charakter- art i.e.S.

Risikofaktoren Risiko

-ID Mgr -I =fHo kurzfristig ) ) )

↓ ↓ ↓ ( ( ( langfristig

Kriterien

Bestand Schutzstatus , 1829) ss << ., 1764) s < = = ELERIN ÜLL ., 1800) mh = = = ., 1830) mh = = = ., 1817) ss < = = WLa (P ., 1783) ex ., 1827) s < ., 1785) es = = = sö (M AYK ., 1784) mh = = = (L., 1758) h = = = (L., 1758) s < TEPH YLL

Taxon (P (L., 1758) mh = = = (S (L., 1758) h = = = BST (L., 1758) s < = - I CHÖNH (G CHALL OURCR (F., 1781)(F., ss < = = Elb (S (L., 1758) sh = = = (H (S (L., 1761) mh = = = (L., 1758) h = = = (L., 1758) mh = = = (F (L., 1767) h = = = (F., 1792)(F., mh = = = (= Agriotes aterrimus (L., 1761)) 1792)) (= Melanotus niger (F., (= Adelocera murina (L., 1758))

Dalopius marginatus Agriotes acuminatus Agriotes ustulatus Agriotes pilosellus Agriotes lineatus Agriotes obscurus Agriotes sputator Ectinus aterrimus Synaptus filiformis Adrastus pallens Adrastus rachifer Melanotus rufipes Melanotus castanipes Melanotus punctolineatus Agrypnus murina Ctenicera pectinicornis Actenicerus sjaelandicus Prosternon tessellatum Anostirus castaneus Haplotarsus incanus Selatosomus aeneus

Grund Kat. Änd. Kat. Grund K K

↓ ↓ ↓ ↓

Kat +/- Kat RL 2010 RL 3+ M 3-R 3 3-R 0-R 3 1-R * * * * * * * * * * * * V-MK R- M

96 1926 ** - D nWZ2 -D- D sö -- ID UWR2, nWZ1 ID StD, gFG gFG =WLa = = CaH =kFG ) ) ) ) ) ) ) ) ↓ ↓ ↓ ↓ ↓ ↓ ↓ ↓ ↓↓ ( ( ( ( ( ( ( ( ., 1847) ss << (F., 1792)(F., s = = - ID ., 1799) ss < = = sö ., 1835) mh < = - D OSH ., 1790) s = = - D ., 1783 es = = - D nWZ1 , 1774) es < , 1774) es << ., 1784) s = = = ., 1853) ss < L ANZ ., 1790) mh = = = ., 1852) ss < = - ID BDü KDü, KüN L (R ITT ., 1799) ex (F., 1792)(F., ss << ., 1767) h = = = (P (F., 1801)(F., h = = = (O OH EER EER , 1785) ss < = = BST (L., 1767) ss < (L., 1758) s < OH ACORD (O .M G (L., 1761) s < (B (G ÜLL ANZ (H E (L (B (L., 1758) h = = = ILL ESKE (P (F., 1792)(F., s < (D (M (L., 1758) mh = = = P (L (L., 1758) h = = = (F., 1792)(F., mh = = = (= Selatosomus nigricornis (Panz., 1799)) 1792)) (= Selatosomus impressus (F., (= Hypoganus cinctus (Payk., 1800)) (= Cidnopus minutus (L., 1758)) (= Cidnopus parvulus (Panz., 1799)) 1785)) (= Stenagostus villosus (Fourcr., (= Pseudathous niger (L., 1758)) (= Pseudathous hirtus (Hbst., 1784))

Selatosomus cruciatus Mosotalesus nigricornis Mosotalesus impressus Calambus bipustulatus Hypoganus inunctus Denticollis rubens Denticollis linearis Cidnopus pilosus Cidnopus aeruginosus Kibunea minuta Nothodes parvulus Limonius aeneoniger Stenagostus rufus Stenagostus rhombeus Hemicrepidius niger Hemicrepidius hirtus Crepidophorus mutilatus Athous haemorrhoidalis Athous vittatus Athous subfuscus Hypnoidus riparius Oedostethus quadripustulatus Negastrius sabulicola Negastrius pulchellus Negastrius arenicola K ↓ ↓ ↓ 3-R 2 3 1-R 2 1-RK 0 1-R 1 3+ M 3+ M 1 3 2 * * * * * * * * V+MK V+MK V+MK

97

Anmerkung Letzter Nachweis Letzter

1919

Verantwortung SH Verantwortung

Verantwortung D Verantwortung

in SH in Verbreitungstyp Verbreitungstyp Wertgebende Begleitart Charakteristische Arten

Charakter- art i.e.S. Risikofaktoren

DNW Risiko

--ID D- nWZ2 D nWZ2 nWZ2 [75] =sö kurzfristig )

↑ ↑ ↓ ↓↓ ( langfristig

Kriterien

Bestand Schutzstatus , 1976 s < = - ID ., 1763) ex OHSE ., 1840 es = ., 1784) mh = = = COP L ., 1784) ss < = - ID sö R E (S ., 1800) es = = - BST ., 1840 ss < = - ID BDü, CaH, Mgr ., 1812 s << BST ., 1874) s > = = R (L., 1758) s < = = ., 1796) ss < = - D

Taxon (H (L., 1761) ss <<< AYK (H HR E (F., 1792)(F., s = = = (P A ANZ ., 1902 ss = HOMS TT (P , 1955 s > = = R (T ALM P (= Hypocoelus olexai Palm, 1955) (= Hypocoelus cariniceps Rtt., 1902) (= Hypocoelus foveicollis (Thoms., 1874)) (= Xylobius corticalis (Payk., 1800)) (= Drapetes biguttatus (Pill. Mitt., 1783))

Cardiophorus nigerrimus Cardiophorus gramineus Cardiophorus ruficollis Cardiophorus asellus Dicronychus cinereus Dicronychus equisetioides Dicronychus equiseti Melasis buprestoides Eucnemis capucina Dirhagus pygmaeus Hylis olexai Hylis cariniceps Hylis foveicollis Xylophilus corticalis Drapetes cinctus

Grund Kat. Änd. Kat. Grund ↑ Kat +/- Kat

+M +M +M RL 2010 RL 2+ K 3 2+MK 3+R 2-RK 2-RK 0 1 2+MK 1 * * * * V-MK Eucnemidae (Kamm-, Dornhalskäfer) Lissomidae (Binden-Dornhalskäfer)

98 1948 1936 -D-D--D D sö sö nWZ2 sö sö = = = ) ↑ ↑ ↓ ↓↓ ↓↓ ↓↓ ↓↓ ( , 2005 mh = = = , 1774) § ex , 2002 h = = = , 1839 § es = = = EER ., 1849 § ss = ., 1837) § ss < = = ., 1835 § mh = = = ., 1859 ss = = = (L., 1758) s = = = ., 1833) § es = = = sö (G (L., 1767) h = = = ORY UONA ., 1857 § ss = EDT ., 1839) § es = = = L., 1758 § es = = = ., 1803) § mh = = = ATZ EBL ESEIGNEUR ONV (F., 1794)(F., ex R ., 1837) § s = .G M LL (L., 1758) mh = = = L ., 1803) § mh = (R B ATZ ., 1921 ss > = = (G (I ACORD ., 1859 ss < IESW (F., 1775)(F., ss > LL (F., 1777)(F., s = = = ATZ AST L TT (R (I K C R (F., 1777)(F., § ss > (R ONV B (= Throscus dermestoides (L., 1767)) 1859) (= Throscus carinifrons Bonv., 2005) (= Throscus meybohmi Leseigneur, (= Throscus leseigneuri Muona, 2002) (= Throscus duvali sensu FHL Bd.6) (= Throscus caucasicus Rtt., 1921)

Trixagus dermestoides Trixagus carinifrons Trixagus meybohmi Trixagus leseigneuri Trixagus exul Trixagus atticus Trixagus Buprestis octoguttata Melanophila acuminata Phaenops cyanea Anthaxia salicis Anthaxia quadripunctata Anthaxia godeti Chrysobothris affinis Agrilus biguttatus Agrilus laticornis Agrilus angustulus Agrilus sulcicollis Agrilus olivicolor Agrilus convexicollis Agrilus cyanescens Agrilus subauratus Agrilus betuleti Agrilus pratensis Agrilus viridis K * ↑ ↑ ↓ ↓ * * * +M +M +R +R 3+MK 2+MK 3 2-R 3+MK 0 2+MK 0-R * * * * * * * * * * * * R+ M R-M R-M R- M Throscidae (Hüpfkäfer) Buprestidae (Prachtkäfer)

99

Anmerkung

Letzter Nachweis Letzter

Verantwortung SH Verantwortung

Verantwortung D Verantwortung

in SH in Verbreitungstyp Verbreitungstyp Wertgebende Begleitart Charakteristische Arten

Charakter- art i.e.S.

Risikofaktoren Risiko

- I-I -I BDü [76] = = kurzfristig ) ) )

↑ ↓ ↓ ↓ ↓↓ ( ( ( langfristig

Kriterien Bestand

mh > = = Schutzstatus , 1832 § ss > ., 1802) mh = = = ., 1857 § ss < , 1817) mh = = = ., 1849 mh = = = , 2003 ss ? ., 1790) § es << . ss = = = nWZ2 ., 1817 § ss << ARSH , 1774) mh = = = L IESW ECK

Taxon EDT ., 1911 s > = = ., 1850) es = = = K (M ÉNÉTRIÉS , 1891 § es ? = - F YLL ., 1914 es = = = (O EER R TT (B EC G EY TT ., 1790) § ss (<) = - ID M ATZEB LACKBURN R (G L R (L R R EYBOHM B (L., 1758) § s = = = (O M (= Agrilus aurichalceus Redt., 1849) 1891) troglodytes subglabra Rey, (= Trachys pumilus FHL ) (= Trachys 1960) (= Clambus radula E.-Y.,

Agrilus viridis fagi Agrilus cuprescens Agrilus sinuatus Aphanisticus pusillus minutus Trachys troglodytes Trachys subglabra Trachys scrobiculatus Trachys Calyptomerus dubius Clambus lohsei Clambus pubescens Clambus simsoni Clambus punctulum Clambus gibbulus Clambus pallidulus Clambus armadillo Clambus nigrellus Grund Kat. Änd. Kat. Grund K

K *

* * * Kat +/- Kat

+MK RL 2010 RL 1-RK 3- 2+MK 1 1+ K * * * * * * * * * V+MK R-M R- M Clambidae (Punktkäfer)

100 [77] 1910 * -IDMgr =QuB =QuB = = = ) ) ) ) ) ↓ ↓ ↓ ↓ ↓ ↓↓ ( ( ( ( ( ., 1818) ss < ., 1821) s < = - D nWZ2 ., 1924 es = ERM ., 1971 mh = = = ., 1859) es = = - ID fDÜ ErH , 1868 mh = = = ÜLL (G UNST RIS (M ., 1830 s < = = M , 1985 ss < .B ., 1787) mh = = = (L., 1767) mh = = = ., 1855 s < LAUSN ., 1799 h = = = ., 1793) s = ., 1868 es = = = QuB K OURNIER TEPH AYK ANZ T (F., 1792)(F., s = = = AIRM (L., 1767) mh = = = P HUNB S HOMS ., 1944 s < = = HMo (L., 1758) ex (P OURN YHOLM (F (F., 1798)(F., s < (T T T N YH (L., 1767) mh = = = N (L., 1758) h = = = (= Helodes minuta (L., 1767)) (= Helodes pseudominuta Klausn., 1971) 1868) (= Helodes elongata sensu Klausnitzer nec Tourn., 1798)) (= Helodes marginata (F., (= Cyphon phragmiteticola Nyholm, 1955)

Dascillus cervinus Elodes minuta Elodes pseudominuta Elodes tricuspis Elodes marginata Microcara testacea Cyphon coarctatus Cyphon palustris Cyphon ruficeps Cyphon kongsbergensis Cyphon ochraceus Cyphon variabilis Cyphon laevipennis Cyphon pubescens Cyphon padi Cyphon hilaris Prionocyphon serricornis Scirtes hemisphaericus Scirtes orbicularis Eucinetus haemorrhoidalis Dryops striatellus K ↓ ↓ 1-R 3-R 3 2 3+MK 2 0 1-RK * * * * * * * * * V+MK V-MK V-MK R+ M Eucinetidae (Wiesenkäfer, Faulkäfer) Eucinetidae (Wiesenkäfer, Dascillidae (Wiesenkäfer) Scirtidae (Jochkäfer, Sumpffieberkäfer) Scirtidae (Jochkäfer, Dryopidae (Klauenkäfer)

101 Anmerkung

[78] Letzter Nachweis Letzter

1954 Verantwortung SH Verantwortung

** **

Verantwortung D Verantwortung

in SH in Verbreitungstyp Verbreitungstyp Wertgebende Begleitart Charakteristische Arten

Charakter- art i.e.S.

Risikofaktoren Risiko

==KüN SzB [6] kurzfristig

↑ ↑ langfristig

Kriterien

Bestand Schutzstatus ., 1784) h = = = ., 1843 mh = = = ., 1806) es = = = ., 1806) mh = = = kFG ., 1843 mh = = = ., 1828 mh = = = SzB Kü [6] ., 1838 s = ., 1828 s = ., 1827) ss << = - I okG öH ., 1843 s < = = ., 1785) s < = - I NMo IESW HUNB ÜLL ÜLL K UER IESW (T URT YLL (F., 1787)(F., s < = = TEPH , 1936 es << = - ID SzG Kü (M ., 1793) mh = = = (M G K C IESW S ., 1909 ss < = - I NMo öH (G

Taxon , 1841) ex K OURCR ANZ DW ., 1798 ss < = - ID kFG ., 1886 mh = = = (F EER E ., 1847) es < = - ID SzG Kü (P ., 1806) mh = = = ., 1847) mh = = = OLLOW R R OZ (H ATR B (E ÜLL L G (E (M

Dryops ernesti Dryops luridus Dryops auriculatus Dryops similaris Dryops griseus Dryops anglicanus Dryops nitidulus Elmis maugetii Elmis aenea Esolus parallelepipedus Oulimnius tuberculatus Oulimnius troglodytes Limnius volckmari Heterocerus flexuosus Heterocerus obsoletus Heterocerus marginatus Heterocerus fenestratus Heterocerus fusculus Heterocerus hispidulus Heterocerus intermedius Heterocerus maritimus

Grund Kat. Änd. Kat. Grund K

↑ ↑ ↓ * Kat +/- Kat

+M +M +M +R +R RL 2010 RL 2-RK 0-R 1 1 3 2 1 * * * * * * * * * * * V-MK V-MK R-M Elmidae (Hakenkäfer) Heteroceridae (Sägekäfer)

102 1895 1964 ** -D - ID StD KDü-D KüO - D- D nWZ2 WLa; nWZ2 = =Mgr = = = ) ) ) ) ) ) ↑ ↓ ↓ ↓ ↓ ↓ ↓ ↓↓ ↓↓ ↓↓ ( ( ( ( ( ( ., 1852 es = ., 1799) es = = = ÜST , 1807) es << = = ., 1790 es << ., 1808) ex K , 1973 es = = = ., 1840 es = = = , 1774 es = ., 1849) s = ., 1863) ss < , 1950 ss < , 1791) es = ., 1840 ex sö 1944 ., 1797) es << = - ID sö YLL OL , 1825) es << = = SKü, StU REUTZ AST EER TURM RAHM AST BST (G ALIK C (S ICHH F., 1775F., s < = = ., 1802 ss << (C ., 1792 s = G B UFT C K (S RAHM (H HANTIEV ., 1818 es << = - IN QuB sö LL , 1774 es = = = (E L., 1758 mh = = = L., 1758 ss << = = I UG (F., 1792)(F., ex Z (D (B F., 1781F., es = = = (L., 1758) s < = - D WLa K EER ERM (L., 1758) mh = = = G G (F., 1792)(F., ss = (=Globicornis marginata (PAYK., 1798)) (=Globicornis marginata (PAYK.,

Eubria palustris Limnichus pygmaeus Limnichus sericeus Dermestes maculatus Dermestes frischii Dermestes murinus Dermestes laniarius Dermestes undulatus Dermestes szekessyi Dermestes gyllenhalii Dermestes ater Dermestes peruvianus Dermestes haemorrhoidalis Dermestes bicolor Dermestes lardarius Attagenus smirnovi Attagenus unicolor Attagenus pellio angustum Trogoderma versicolor Trogoderma glabrum Trogoderma Globicornis nigripes Globicornis emarginata Globicornis corticalis Megatoma undata Ctesias serra Anthrenus pimpinellae K K K K K ↓ * * ↓ ↓ ↓ 1-R 3 1 0 3+MK 0 2 1 1-R 2-MK 1 2+MK 2 0 1 2 1-R * * * V-R V-MK R-M R-MK R+ M R-M R-MK Psephenidae (Sumpfwiesenkäfer) Limnichidae (Ufer-Pillenkäfer) Limnichidae (Ufer-Pillenkäfer) Dermestidae (Speckkäfer, Pelzkäfer) Dermestidae (Speckkäfer,

103

Anmerkung

Letzter Nachweis Letzter

Verantwortung SH Verantwortung

Verantwortung D Verantwortung

in SH in Verbreitungstyp Verbreitungstyp Wertgebende Begleitart Charakteristische Arten

Charakter- art i.e.S.

Risikofaktoren Risiko

- D nWZ2 -I HMo =sö =WLa = CaH = kurzfristig ) ) ) )

↓ ↓ ↓ ↓ ↓↓ ↓↓ ( ( ( ( langfristig

Kriterien

Bestand Schutzstatus ., 1783) ss << = - ID Mgr sö , 1843) es = = = ., 1839 es = = - D ., 1790) ss < = - D nWZ1 L , 1794) es = = = CHALL OPE ., 1771) s < = - I Mgr (O (L., 1758) ss < , 1771) mh = = = (S (H , 1934) ss < OTSCH ., 1771) mh < = = OSSI (F., 1794)(F., mh = = = ., 1798) ss < (L., 1761) mh = = =

Taxon ., 1842 es << M LL (R ., 1939) es = = = ORST ., 1789 s < = = (I ALM L (L., 1767) mh = = = (F., 1794)(F., s < ORST (F ILL TEFF (P (F., 1775)(F., s < = = O ORSTER (F S (M (F (F., 1781)(F., ss < (L., 1758) mh = = = (= Cytilus auricomus (Duft., 1825)) (= Syncalypta spinosa (Rossi, 1794))

Anthrenus scrophulariae Anthrenus verbasci Anthrenus museorum Anthrenus fuscus hirtus Trinodes Thylodrias contractus Anthrenocerus australis Reesa vespulae Nosodendron fasciculare Simplocaria semistriata Morychus aeneus Lamprobyrrhulus nitidus Cytilus sericeus Byrrhus fasciatus Byrrhus pilula Byrrhus arietinus Byrrhus pustulatus Porcinolus murinus Chaetophora spinosa Curimopsis nigrita Curimopsis setigera

Grund Kat. Änd. Kat. Grund K K K

↓ ↓ ↓ * * ↓

Kat +/- Kat RL 2010 RL 2-R 1-RK 2-R 1-R 3-RK 2-R 2 3 1 2 * * * * * * V-MK V-MK R-M R-M R-MK Byrrhidae (Pillenkäfer) Nosodendridae (Saftkäfer)

104 Fußnoten

[lfd. Nr.] Anmerkung [1] durch ökologischen Landbau begünstigt [2] SzW auf Torf, Salzröhricht [3] Die noch vorhandenen Vorkommen erscheinen derzeit stabil bzw. gesichert. [4] Binnenlandfunde sind extrem selten, z.B. auf Spülfeldern oder in Gruben. [5] beweidetes Feuchtgrünland (Störstellen) [6] an der einzigen Binnenland-Salzstelle (Brennermoor) seit rund 40 Jahren keine aktuellen Funde [7] Trotz intensiver Nachsuche nicht mehr gefunden. [8] nur Helgoland; einziges Vorkommen in D [9] Außerhalb der Elbtalaue extrem selten (z.B. auf Spülfeldern). [10] Wegen des starken Nutzungsdrucks durch Strandtourismus höher eingestuft, insbesondere an der Ostsee. [11] an der Ostseeküste extrem selten (nur alte Funde) [12] Bindung an dynamische Küstenstrukturen; schluffiges Substrat [13] in Kombination mit alten Waldstandorten [14] Erst seit 1989 aus dem Gebiet bekannt (Segrahn). Vorkommen in Segrahn erloschen, die Grube bei Mehlbek wird gerade geflutet. Aktuell keine stabilen Vorkommen bekannt. [15] gebunden an den Tideeinfluss [16] historisch auch von Steilufern der Elbe und der Nordseeküste be- kannt [17] die Art ist ausgesprochen flugaktiv, daher auch vereinzelte Binnen- landfunde; an der einzigen Binnenland-Salzstelle (Brennermoor) seit rund 40 Jahren keine aktuellen Funde [18] bevorzugt in lichten Salzröhrichten [19] An der Ostseeküste wertgebend für Salzwiesen. [20] beschränkt auf Weidenauwald [21] benötigt offene Torfflächen [22] früher auch im Brennermoor (zuletzt 1942) [23] Durch die „Inwertsetzung“ von Brachflächen ist ein deutlicher Rückgang absehbar. [24] Ufer temporärer Kleingewässer auf Sand-Rohboden [25] Isolierte Vorkommen innerhalb der norddeutschen Tiefebene. [26] in feuchten Vorratskellern [27] Funde außerhalb der Elbtalaue sind extrem selten (Einzelfälle) [28] auf kalkhaltigen Sanden; Abhängigkeit von Gehäuseschnecken

105 [lfd. Nr.] Anmerkung [29] bevorzugt LRT 7140 [30] bevorzugt LRT 3130 [31] LRT 7140 und 3140 [32] Neben den Küstenvorkommen nur aus den Qualmwasserbereichen der Elbe bei Besenhorst bekannt. [33] bevorzugt LRT 7140 und *91D0 [34] bevorzugt LRT *91E0 und *91D0 [35] bevorzugt LRT 7140 und 7230 [36] bevorzugt LRT *91D0 und 7110 [37] bevorzugt LRT 7110 [38] Gefährdung durch Degradation von Brandungsufern. [39] bisher fast ausschließlich im nördlichen Landesteil [40] bevorzugt LRT *91D0 [41] bevorzugt Waldtümpel in LRT 9130 [42] bevorzugt LRT 1150 [43] bevorzugt LRT 7110 und 7140 [44] Bindung an stark gefährdete Lebensräume: sonnendurchflutete Qualmwasserbiotope und Moorgewässer. [45] letzter Nachweis 1975 im Hechtmoor, Kreis Schleswig-Flensburg [46] bevorzugt dyG in LRT 7140 [47] bevorzugt dyG in LRT *7110 [48] bevorzugt lichte Röhrichte an Brandungsufern und Seeabflüssen [49] bevorzugt im Röhrichtsaum von Verlandungsmooren an Seen [50] vermutlich durch Gewässerräumung ausgestorben [51] Die Art ist ausgesprochen flugaktiv, daher auch vereinzelte Binnen- landfunde. [52] nur vereinzelt auch Binnenlandfunde [53] Wegen Bindung an hochwertige Gewässer (Krebsscherenbestände, Niedermoore) stark gefährdet; bevorzugt LRT 7230. [54] Bindung an Sandbänke und dynamische Uferstellen [55] Durch systematische Beseitigung alter Pappeln gefährdet. [56] bisher nur von Amrum bekannt [57] Bei den Vorkommen in Schleswig-Holstein handelt es sich um die einzigen in Mitteleuropa! [58] in Muschelschill; früher am Kreidefelsen Helgoländer Düne [59] Schwerpunkt in der Elbtalaue [60] unter oligohalinen Bedingungen, bisher nur Schellbruch/Lübeck, frü- her im Brennermoor [61] dynamische Küstenstrukturen [62] nur auf sandigem Substrat [63] Salzröhricht

106 [lfd. Nr.] Anmerkung [64] Stenus niveus Fauv., 1865 - trotz intensiver Nachsuche augen- scheinlich verschwunden [65] Bindung an die Sprühwasserzone [66] Die Grube bei Lägerdorf mit dem einzigen bei uns bekannten Vor- kommen soll geflutet werden. [67] Früher mehrfach auch im Siedlungsbereich, heute bevorzugt im Spülsaum der Küste. [68] an der Ostsee sehr selten [69] bevorzugt im Verbuschungs- und Vorwaldstadium [70] gebunden an den Porenlufthorizont [71] an der Ostseeküste ‘es’ [72] Von der Ostseeküste keine aktuellen Funde (2010) [73] auf Sumpfdotterblumen-Wiesen [74] nur Helgoland [75] Bindung an stehende Starkholzstrukturen und grenzwertig zu ‘es’ [76] an Sandsegge [77] bisher nur auf Sylt [78] in oligotrophen, bevorzugt kalkreichen Seen des östlichen Hügellan- des [79] Küstendünen Sonderfall Sylt; sonst Waldbewohner (Eschenwald) [80] aktuell nur auf Sylt [81] Trotz intensiver Nachsuche nicht wieder gefunden. [82] Freilandfund, eine Verschleppung ist aber nicht ganz auszuschlie- ßen. [83] Bindung an starkes Eichentotholz (Stubben) [84] Die einzige Population durch intensiven Einschlag von alten Bäu- men gefährdet (aktuell 2008/09). [85] Wegen Bindung an (ur)alte Eichen als gefährdet eingestuft. [86] Durch Einstellung der Beweidung am Dummersdorfer Ufer ist das einzige bekannte Vorkommen nachweislich erloschen. [87] Durch Einstellung der Beweidung am Dummersdorfer Ufer auch das letzte verbliebene Vorkommen nachweislich erloschen. [88] Einziges aktuelles Vorkommen bei uns: Elbinsel Pagensand. [89] Das einzige bekannte Vorkommen befindet sich auf Sylt. Nicht akut bedroht, solange (extensive) Pferdehaltung besteht. [90] Das einzige Vorkommen in SH sind schon immer die Geesthachter Dünen; dort starker Rückgang durch Bebauung und Freizeitnutzung. [91] Abhängigkeit von Kompostwirtschaft. Verschlechterung der Situati- on durch verstärkten Einsatz in Biogasanlagen.

107 [lfd. Nr.] Anmerkung [92] Die Art stand bisher auf ‘0’, es gibt jedoch einen Einzelnachweis als Totfund. Dieser Fund liegt außerhalb aller historisch bekannten Vor- kommen, so dass vorerst eine Verschleppung nicht auszuschließen ist. [93] Alte Funde wurden lange als verschleppte Tiere interpretiert, was aus heutiger Sicht nicht schlüssig erscheint (keine Art der Holzlä- ger). Aktuelle unbelegte Beobachtung Büchen Hellberg, Frau Den- ker ca. 1995 am Licht. Eindeutige Schlupflöcher Segrahn (Suikat). [94] Unstete Vorkommen in einem begrenzten Raum (Hamburger Moor- gürtel). [95] außerhalb der Elbtalaue äußerst selten [96] Bindung an Linum. In den 1990er Jahren durch verstärkten Anbau von Linum usitatissimum zeitweilig häufig gewesen, seit etwa 2005 sehr stark rückläufig bis verschwunden. [97] An der Ostsee deutlich seltener und auch als gefährdet anzusehen. [98] Bindung an sandige Salzwiesen und kalkreiche Trittfluren; daher langfristiger Rückgang als sicher anzunehmen. [99] Bindung an Linum. In den 1990er Jahren durch verstärkten Anbau von Linum usitatissimum zeitweilig häufig gewesen, seit etwa 2005 sehr stark rückläufig bis verschwunden. [100] bisher nur von den Geestinseln bekannt [101] Strandwall mit Gehölzbewuchs (Weißdorn) [102] bisher nur an der Nordsee [103] Langfristiger Trend nicht sicher, da früher als ssp. von suturalis be- trachtet und nicht immer davon getrennt. [104] xylobiont; an natürlichen und verbauten Hölzern [105] an Störstellen mit Butomus [106] Pionierfluren mit Krötenbinse [107] Erstnachweis 1970, danach mehrfach im südlichen SH, allerdings seit Mitte der achtziger Jahre wieder im Rückgang! [108] Erstmeldung 1959, danach mehrfach gefunden. In den letzten 10 Jahren drastische Abnahme. Letzte Meldung 1998. [109] xylobiont an Buche und Heidekraut; alte Waldrandstrukturen [110] Endemit der Nordseeküste [111] Lebensraum von Mahd abhängig [112] nur noch im Bereich Borghorster / Besenhorster Elbsandwiesen [113] sowohl an Calluna als auch an Erica [114] aktuell nur auf Helgoland [115] Die einzige bekannte Fundstelle wurde abgegraben. [116] Trotz intensiver Nachsuche am einzigen bekannten Standort nicht mehr gefunden.

108 [lfd. Nr.] Anmerkung [117] Die Art ist nur durch einen jüngst aufgetauchten Altfund für unser Gebiet belegt - ‘Sleswig’ (Schleswig) Juli 1883 Coll. Andersen (Jan Pedersen vid.)

109 110 Legende zur Roten Liste und Checkliste der Käfer Schleswig-Holsteins

Rote Liste: 0 = ausgestorben oder verschollen, 1 = vom Aussterben bedroht, 2 = stark gefährdet, 3 = gefährdet, R = extrem seltene Arten, G = Gefährdung unbekannten Ausmaßes, V = Vorwarnliste, * = ungefährdet, D = Datenlage unzureichend.

Kategorieänderung: + = niedrigere Einstufung gegenüber der alten Roten Liste („Herabstufung“), – = höhere Einstufung („Heraufstufung“), also Verschärfung der Gefährdungssituation, ohne Eintrag = Einstufung unverändert.

Grund der veränderten Einstufung: R = reale Veränderung des Gefährdungsgrades bzw. der Bestandssituation ggf. mit Angabe der Richtung (↓ Abnahme / ↑ Zunahme des Bestandes), K = Kenntniszuwachs, M = Methodik der Bewertung, insbesondere Änderungen im Kriteriensystem, * = Taxon erst nach 1994 im Gebiet aufgetreten bzw. als Art berücksichtigt, Na = Veränderung des Gefährdungsgrades bzw. der Bestandssituation durch Naturschutzmaßnahmen.

Schutzstatus: §§ = streng geschützt, § = besonders geschützt nach BArtSchV; II bzw. IV = streng geschützte Arten nach Anhang II bzw. IV der FFH-Richtlinie.

Bestand: ex = ausgestorben, es = extrem selten, ss = sehr selten, s = selten, mh = mäßig häufig, h = häufig, sh = sehr häufig.

Langfristiger Bestandstrend: <<< = sehr starker Rückgang, << = starker Rückgang, < = mäßiger Rückgang, (<) = Rückgang, Ausmaß unbekannt, > = deutliche Zunahme, ? = historische Datenlage unklar.

Kurzfristiger Bestandstrend: ↓↓↓ = sehr starker Rückgang, ↓↓ = starker Rückgang, (↓) = mäßiger Rückgang, = = gleich bleibend,

↑ = nachweisliche Zunahme, ? = aktuelle Datenlage unklar.

Risiko: – = negativ wirksame Faktoren vorhanden, = = keine zusätzlich negativ wirksamen Faktoren feststellbar (bzw. als nicht zusätz- lich relevant eingestuft).

Risikofaktoren: A = Bindung an stärker abnehmende Arten, D = Verstärkte direkte, konkret absehbare menschliche Einwirkung, F = Fragmentierung / Isolation, I = Verstärkte indirekte, konkret absehbare menschliche Einwirkung, N = Abhängigkeit von nicht langfristig gesicherten Naturschutzmaßnahmen, W = Wiederbesiedlung unwahrscheinlich (z.B. durch geringes Ausbreitungsvermögen).

Charakteristische Arten: Für die Bewertung des Erhaltungszustandes insbesondere der FFH-Lebensraumtypen relevante Arten (siehe Erläuterung in Band 2, Kapitel 6 Seite 20). Küstenbiotope und Salzstellen Gewässerbiotope einschließlich deren Ufer SbW Sandbänke und Watten (= 1110, 1140) Q Quellbereiche (enthält *7220) RFk Riffe / Felsenküste semiaquatisch (= 1170) obG oligotrophe basenarme Gewässer SKü Steilküste terrestrisch (= 1230) (= 3110, 3130) StD dynamischer Strandkomplex mit Spülsaum und okG oligotrophe kalkhaltige Gewässer (= 3140) Primärdüne (= 1210, 2110) euG eutrophe Stillgewässer (= 3150) StS Strandseen (= *1150, 1160 ) dyG dystrophe Stillgewässer (= 3160) StW Strandwall mehrjährig (= 1220) kFG kleine Fließgewässer - Bäche, Flüsse (enthält KDü Küstendünen (= 2120, *2130, *2140, *2150, 3260) 2460, 2170) gFG große Fließgewässer - Elbe, Flussunterläufe fDÜ feuchte Dünentäler (= 2190) (enthält 1130, 3270) SzP Salzpionierflur, Quellerwatt (= 1310) StU Steilufer Binnenland (Prallhänge dynamischer SzW Salzwiese der Küsten (= 1330) Gewässer, Abbruchkanten an Seen) SzB Salzwiese des Binnenlandes (= *1340) Wälder SzG Salzbeeinflusste Kleingewässer (= 1330 part.) WLa Laubwald (enthält 9110, 9120, 9130, 9160, Trockenbiotope Binnenland *9180, 9190) BDü Binnendünen (= 2310, 2320, 2330) WAQ Au- und Quellwälder (= *91E0) Mgr (Sand-)Magerrasen (enthält *6120, 6210/*6210, Sonstige Biotope / Qualitäten *6230) ÜGr Überflutungsgrünland (enthält 6440, 6510 part.) CaH Calluna-Heiden (= Trockenheide 4030, 5130) haW historisch alte Waldstandorte Moore u.a. Feuchtbiotope Indikatoren für Strukturqualität und Habitattradition HMo Hochmoor- und Übergangsmoor (= *7110, in Gehölzbiotopen 7140, 7150, 7120 inkl. *91D0) UWR Urwaldrelikt-Arten auf Bundesebene NMo Niedermoor/Sümpfe (enthält 7230) nWZ Zeiger für Naturnähe auf Landesebene ErH Erica-Heide (= Feuchtheide 4010) fHo feuchte Hochstaudenfluren (= 6430) Röh Röhrichte (enthält *7210) Verbreitungstyp in SH: Kennzeichnung von Arten mit offenkundig eingeschränkter Verbreitung innerhalb des Landes. Kü = Küstenart (Nord- und Ostsee), KüN = Küstenart, nur Nordsee, KüO = Küstenart, nur Ostsee, öH = östliches Hügelland (Jungmoräne), Feh = Fehmarn, Elb = Elbtal (auf das Urstromtal begrenzt), ElbO = Elbtal zwischen Lauenburg und Hamburg, ElbW = Elbtal zwischen Mündung und Hamburg, Elb+ = Elbe und Nebengewässer (Unterläufe), Hgld = Helgoland, sö = nur im Südosten (kontinentaler Einfluss), W = nur im Westen.

Verantwortung D: Deutschlands Verantwortung im globalen Maßstab. !! = in besonders hohem Maße verantwortlich, ! = in hohem Maße verantwortlich, (!) = in besonderem Maße für hochgradig isolierte Vorposten verantwortlich, E = Endemit (globale Alleinverantwortung).

Verantwortung SH: Schleswig-Holsteins Verantwortung innerhalb Deutschlands. *** = ausschließliche Verantwortlichkeit (bundes- weit nur aus Schleswig-Holstein bekannt), ** = sehr hohe Verantwortlichkeit (bundesweit aus bis zu zwei weiteren faunistischen Regionen bzw. Bundesländern bekannt und Schleswig-Holstein aus naturräumlichen Gründen von tragender Bedeutung), * = hohe Verantwortlichkeit (bundesweit aus bis zu drei weiteren faunistischen Regionen bzw. Bundesländern bekannt, Schleswig-Holstein in der Mitverantwortung).

Letzter Nachweis: Bei Arten der Kategorie 0 (ausgestorben oder verschollen) wird das Jahr des letzten bekannten Nachweises ange- geben.

Anmerkungen: Ergänzende Informationen in fortlaufender Nummerierung (siehe Ende des Bandes).