Botany (the Study of Plants) General Plant Review All plants are: Eukaryotic Autotrophic Multicellular Cell Walls with cellulose Chloroplasts w/ chlorophyll a, b, and carotenoids May have waxy cuticle to prevent water loss. Stomata allow gas exchange. Plants probably evolved from green algae (charophytes) Chloroplast similarity Biochemical similarities Cell Wall similarities
Plant Reproduction Alternation of Generations – plants grow a separate, haploid organism to produce gametes; the same plant exists in two different forms during its life, although sometimes it’s hard to differentiate between the forms
Multicellular Sporophyte – (2N) Mitosis Meiosis Zygote (2N) Unicellular fertilization Spores (N)
Unicellular Gametes (egg & sperm) Multicellular Gametophyte (N) Evolutionary Trend – early plants displayed the gametophyte (n) as the dominant structure, modern plants show the sporophyte as dominant (2n) (Ex: Moss is HAPLOID, but Oak Trees are DIPLOID)
zygote
GREEN ALGA BRYOPHYTE FERN GYMNOSPERM ANGIOSPERM Classifying Plants
Plants can be divided into 2 major categories based on their characteristics: Nonvascular Plants Do NOT have specialized tissues to transport water and nutrients Instead, these plants transport water from cell-to-cell by osmosis Vascular Plants Have specialized tissues to transport water and nutrients in plants Xylem – carries water upward from roots Phloem – carries nutrients and carbohydrates produced by photosynthesis Nonvascular Plants (Bryophytes)
Short with no specialized tissues to transport water Major types: Mosses Liverworts Hornworts Nonvascular Plants/ Bryophytes
Mosses Have rhizoids that anchor them to the ground (instead of roots) Depend on water for fertilization The sperm must swim to the egg Therefore, nonvascular plants must live in MOIST environments Gametophyte is the dominant phase of the life cycle
Moss/ Bryophyte Life Cycle
Zygote grows, mature develops into a sporophyte sporophyte while still attached to gametophyte. zygote Diploid Stage Fertilization Meiosis Haploid Stage
Spores germinate. sperm- producing structure male gametophyte egg- producing female structure gametophyte Vascular Plants/Tracheophytes
Have specialized tissues to transport water and nutrients in plants called Xylem & phloem Vascular plants (tracheophytes) can be divided into 2 categories: Seedless vascular plants (spores) Seed (vascular) plants
Seedless Vascular Plants
Ferns – A Close Up Diploid sporophyte is the dominant stage Have rhizomes, which are underground stems Fronds: large “leaves” where spores develop Develop spores in sporangia on underside of fronds Reproduce using spores A sorus (plural: sori) is a cluster of sporangia Fern Life Cycle
Sporophyte still attached to gametophyte
sorus zygote rhizome Diploid Stage fertilization meiosis Haploid Stage Spores Spores develop are egg released
sperm mature gametophyte Spore germinates Seed (Vascular) Plants
Have true roots, leaves, and stems Have the ability to form seeds, which are used for reproduction Seed plants are the most dominant group of photosynthetic organisms on land There are 2 types of seed (vascular) plants: 1. gymnosperms (cones) 2. angiosperms (flowers) Seed (Vascular) Plants Gymnosperms = “cone bearers” “naked seeds” – not enclosed in ovaries Bear seeds directly on the surfaces of cones Cones = sporophyte structures that produce gametophytes (seeds) Coniferous trees are the major example Pines, junipers, spruces, etc.
section through one surface view of one cone scale ovule (houses two ovules) Pine Life Cycle ovule
surface view of one cone scale (houses a pollen-producing sac) mature sporophyte seed section through a coat pollen-producing sac zygote seeding embryo Diploid seed fertilization meiosis pollen tube Haploid microspores eggs form sperm- megaspores producing cell pollination form female gametophyte Seed (Vascular) Plants
Angiosperms = flowering plants Seeds are enclosed by an ovary Flowers are reproductive organs Evolutionary advantage attract pollinators Flowering plants contain ovaries Ovaries surround and protect seeds Ovary develops into a fruit after pollination & helps with seed dispersal when eaten Examples: Maple trees, tulips, grass sporophyte
Flowering Diploid Plant Life Double fertilization Meiosis Meiosis Haploid mitosis Cycle without microspores pollination cytoplasmic division
two sperm enter ovule female gametophyte Evolutionary Tree for Plants
Nested monophyletic groups
flowering green zygophytes, charophytes bryophytes lycophytes horsetails ferns cycads ginkgos conifers gnetophytes plants algae related groups
seed plants euphyllophytes embryophytes (land plants) vascular plants (closely related groups) Transport in Vascular Plants Transport Within Plants
Water and minerals absorbed by roots are drawn upward in the xylem to the shoots Sugar produced by photosynthesis is exported from leaves to other organs via the phloem Transport can be passive or active Short- and Long-Distance Transport in Plants
Water and sugars move differently in plants, depending on whether they’re going a short or a long distance Ie. – walking down the street or taking an airplane across the world Short-Distance Transport 1. Simple diffusion/osmosis substances in one cell move out of one cell, across the cell wall, and into another cell 2. Plasmodesmata Plasmodesmata are connections between the cytoplasm of adjacent plant cells Substances move between cells through these openings
Long-Distance Transport
Over long distances, these 3 processes (simple diffusion, apoplast,& symplast) take too long Water and solutes move through xylem and phloem by bulk flow, the movement of a fluid driven by pressure Transpiration
Transpiration is the evaporation of water from leaves and other parts of the plant; it causes a pull that brings more water up through the xylem An average maple tree loses more than 200L of water per hour during the summer! Unless this water is replaced by water absorbed by the roots, leaves will wilt and die (Cohesion-Tension Theory)
Transpiration works through a combination of evaporation, water potential, adhesion, and cohesion to pull water up the xylem
The Photosynthesis- Transpiration Compromise
Exposing leaves to the sun and opening stomata (cellular “holes” in the underside of the leaf) to allow for gas exchange helps photosynthesis, but causes transpiration to occur at a faster rate. Plants balance the loss by controlling when stomata are open. Mechanism for Stomatal Opening & Closing
Guard cells open and close stomata by changing shape using ion and hormone signals based on plant stress. Plant Structure, Growth, & Development The Diversity of Angiosperms
Angiosperms (flowering plants) can be divided into 2 major categories: Monocots – have one seed leaf (cotyledon) Dicots – have 2 seed leaves (cotyledons)
Monocots
Monocots have only 1 cotyledon (seed leaf) Examples of monocots: Corn, wheat, lilies, orchids, palms
Dicots
Dicots have 2 cotyledons (seed leaves) Examples of dicots: Roses, clover, tomatoes, oaks, daisies Woody vs. Herbaceous Plants
Angiosperms can also be subdivided into woody (produce wood) and herbaceous (do not produce wood) plants Woody plants are made of cells with thick cell walls that support the cell body Examples: trees, shrubs, vines Herbaceous plants do not produce wood as they grow, and instead have smooth stems Examples: dandelions, sunflowers Plant Life Spans The lifespan of plants, however, is genetically determined Annuals – complete their life cycle in 1 year Examples: marigolds, cucumbers (lots of garden plants) Biennials - complete their life cycle in 2 years Year 1: germinate & grow roots Year 2: grow stems & leaves, produce flowers & seeds Examples: evening primrose, celery Perennials – live for more than 2 years Examples: Maple trees, grasses, palm trees Plant Structure
Plants are made up of a root system and a shoot system
The Root System
What do roots do? Anchor the plant in the soil Absorb minerals and water Store food Types of root systems Fibrous root system Found mostly in monocots Taproot system Found mostly in dicots How do roots grow?
Roots grow down from the tip in a region called the apical meristem, where the cells are dividing quickly
The Shoot System
The shoot system consists of: vegetative shoots (which bear leaves) floral shoots (which bear flowers) Stems have 3 important functions: Producing leaves, flowers (reproduction), branches Holding leaves up to the sunlight for photosynthesis Transporting substances between roots and leaves How do stems grow?
Primary growth Increase in length Occurs by cell divisions in apical meristem (at top of shoot) Secondary growth Increase in width Occurs by cell divisions in the lateral meristems (outward growth)
The Shoot System: Leaves
Leaves are the primary photosynthetic organs of most vascular plants Most leaves have a flattened blade and a petiole, which is the stalk that attaches the leaf to the stem
Tissues in Plants
All 3 plant organs (root/stem/leaf) have dermal, vascular, and ground tissue systems Dermal Tissue System Outer protective covering, similar to our skin Protects the plant from water loss and disease The cuticle is a waxy coating that helps to prevent water loss Tissue Systems in Plants
Vascular Tissue System Carries out long-distance transport of materials within the plant Xylem and phloem are examples of vascular tissues Ground Tissue System Pith (inside vascular tissue) and cortex (outside vascular tissue) are examples of ground tissue Inner cells specialized for storage, photosynthesis, and support
Flower Structure
Flowers are the reproductive structure of angiosperms Sepals: Enclose the bud before it opens Protect flower while it’s developing Petals: Usually brightly colored to attract pollinators Flower Structure
Stamens: The male portion of a flower Made up of an anther and a filament The anther produces haploid pollen grains by meiosis Most flowers have multiple stamens Flower Structure
Carpels/Pistils: The female portion of a flower Stigma: Sticky – to trap pollen Style: Hollow tube which connects stigma and ovary Ovary: Produces female gametes (ovules) Fruit grows from an expanded ovary
Seed Dormancy
Seed dormancy means that a seed will not germinate, even if sown in a favorable place, until a specific environmental cue causes them to break dormancy Seed dormancy increases the chances that germination will occur at a time and place most advantageous to the seedling How did we break dormancy in our lab?? Stages of Seed Germination
(1) The seed absorbs water, causing it to expand and rupture its seed coat (2) The embryo resumes growth, digesting the storage materials of the endosperm (3) The radicle (embryonic root) emerges from the germinating seed (4) The shoot tip breaks through the soil surface
Stages of Seed Germination Plant Asexual Reproduction
When plant species clone themselves by asexual reproduction, it’s known as vegetative reproduction
Asexual Reproduction
Fragmentation is the separation of a parent plant into parts that re- form whole plants This type of asexual reproduction is used to produce clones from cuttings (common with houseplants)
Plant Responses to Internal & External Signals Plant Hormones
REVIEW: Hormones are chemical signals that coordinate the various parts of an organism A hormone is a compound produced in one part of the body which is then transported to other parts of the body, where it triggers responses in target cells and tissues Examples of human hormones: Adrenaline, testosterone, estrogen, epinephrine… Plant Hormones
There are 5 major classes of plant hormones: Auxin - Cytokinins Gibberellins Abscisic acid Ethylene Auxin
Stimulates stem elongation Stimulates fruit development Involved in phototropism (bending to light) and gravitropism (growing UP)
Cytokinins
Stimulate cell division and growth Stimulate cytokinesis Stimulate germination and flowering
Gibberellins
Trigger seed and bud germination Promote stem elongation and leaf growth Important in fruit growth Ethylene
Promotes fruit ripening Senescence (aging) is a progression of irreversible change that eventually leads to death Caused, at least in part, by ethylene “One bad apple spoils the whole bunch” Abscisic Acid
Induces seed dormancy Anti-gibberellin Inhibits cell growth Anti-cytokinin Inhibits fruit ripening Anti-ethylene Closes stomata during water stress, allowing many plants to survive droughts
Tropisms
Tropisms are growth responses that result in curvatures of whole plant organs toward or away from a stimuli There are three major stimuli that induce tropisms Light (Phototropism) Gravity (Gravitropism/Geotropism) Touch (Thigmotropism)
Phototropism
Phototropism is the growth of a shoot towards light This is primarily due to the action of auxin Auxin elongates the cells on the non-light side
Plant Defenses
Plants defend themselves against herbivores in several ways Physical defenses, such as thorns Chemical defenses, such as producing distasteful/toxic compounds