Unmasking Evolution
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FISHING for DUNKLEOSTEUS You’Re Definitely Gonna Need a Bigger Boat by Mark Peter
OOhhiioo GGeeoollooggyy EEXXTTRRAA July 31, 2019 FISHING FOR DUNKLEOSTEUS You’re definitely gonna need a bigger boat by Mark Peter At an estimated maximum length of 6 to 8.8 meters (20–29 sediments that eroded from the Acadian Mountains, combined feet), Dunkleosteus terrelli (Fig. 1) would have been a match for with abundant organic matter from newly evolved land plants even the Hollywood-sized great white shark from the and marine plankton, settled in the basin as dark organic movie Jaws. Surfers, scuba divers, and swimmers can relax, muds. Over millions of years, accumulation of additional however, because Dunkleosteus has been extinct for nearly 360 overlying sediments compacted the muds into black shale rock. million years. Dunkleosteus was a placoderm, a type of armored The rocks that formed from the Late Devonian seafloor fish, that lived during the Late Devonian Period from about sediments (along with fossils of Dunkleosteus) arrived at their 375–359 million years ago. Fossil remains of the large present location of 41 degrees north latitude after several species Dunkleosteus terrelli are present in the Cleveland hundred million years of slow plate tectonic movement as the Member of the Ohio Shale, which contains rocks that are North American Plate moved northward. approximately 360–359 million years old. Figure 1. A reconstruction of a fully-grown Dunkleosteus terrelli, assuming a length of 29 feet, with angler for scale. Modified from illustration by Hugo Salais of Metazoa Studio. Dunkleosteus cruised Late Devonian seas and oceans as an Figure 2. Paleogeographic reconstruction of eastern North America during apex predator, much like the great white shark of today. -
Biology in the Modern World Study Guide Southwest Minnesota State University Chapter 1
Biology in the Modern World Study Guide Southwest Minnesota State University Chapter 1 Chapter Objectives Biology and Society: Biology All Around Us 1. Describe three examples of how biology is woven into the fabric of society. The Scope of Life 2. Describe seven properties or processes we associate with life. 3. List and give an example of each level of biological organization, starting with an ecosystem and ending with atoms. 4. Describe the two main dynamic processes in an ecosystem. 5. Compare the structure of prokaryotic and eukaryotic cells. 6. Distinguish between the three domains and four eukaryotic kingdoms of life. Evolution: Biology’s Unifying Theme 7. Describe the two main points that Darwin made in his book The Origin of Species. 8. Describe the two observations that led Darwin to his inescapable conclusion. State this conclusion. 9. Compare artificial and natural selection, noting similarities and differences. The Process of Science 10. Compare discovery science and hypothesis-driven science. Provide examples of each regarding the study of trans fats in the human diet. 11. Distinguish between a hypothesis and a theory. Explain why natural selection qualifies as a scientific theory. 12. Distinguish science from other styles of inquiry. 13. Describe examples of the interdependence of technology and science. Evolution Connection: Evolution in Our Everyday Lives 14. Explain how changes in the effectiveness of antibiotics illustrate natural selection. Key Terms biology hypothesis-driven science case study life controlled experiment natural selection discovery science scientific method ecosystem theory hypothesis Chapter 2 Chapter Objectives Biology and Society: Fluoride in the Water 1. Explain how fluoride prevents tooth decay. -
Proceedings of the Indiana Academy of Science
PROCEEDINGS of the Indiana Academy of Science Founded December 29, 1885 Volume 67 1957 Richard A. Laubengayer, Editor Wabash College Crawfordsville, Indiana Spring Meeting- May 10-11 Turkey Run State Park Fall Meeting October 17-19, 1957 DePauw University Published at Indianapolis, Indiana 1958 1. The permanent address of the Academy is the Indiana State Library. 2. Instructions for Authors appear at end of this volume, p. 324. 3. Exchanges. Items sent in exchange for the Proceedings and corre- spondence concerning- exchange arrangements should be addressed: John Shepard Wright Memorial Library of the Indiana Academy of Science. c/o Indiana State Library Indianapolis 4, Indiana, 4. Proceedings may be purchased through the State Library at $3.00 per volume. 5. Reprints of technical papers can often be secured from the authors. They cannot be supplied by the State Library nor by the officers of the Academy. 6. The Constitution and By-Laws reprinted from v. (J 2 and the Member- ship List reprinted from v. 61, are available to members upon application to the Secretary. Necrologies reprinted from the various volumes can be supplied relatives anl friends of deceased members by the Secretary. 7. Officers whose names and addresses are not known to correspondents may be addressed care the State Library. The address of the editor of the present volume is Biology Department, Wabash College, Crawfordsville, Ind. Papers published in the Proceedings of the Indiana Academy of Science are abstracted or indexed in appropriate services listed here: Annotated -
'Placoderm' (Arthrodira)
Jobbins et al. Swiss J Palaeontol (2021) 140:2 https://doi.org/10.1186/s13358-020-00212-w Swiss Journal of Palaeontology RESEARCH ARTICLE Open Access A large Middle Devonian eubrachythoracid ‘placoderm’ (Arthrodira) jaw from northern Gondwana Melina Jobbins1* , Martin Rücklin2, Thodoris Argyriou3 and Christian Klug1 Abstract For the understanding of the evolution of jawed vertebrates and jaws and teeth, ‘placoderms’ are crucial as they exhibit an impressive morphological disparity associated with the early stages of this process. The Devonian of Morocco is famous for its rich occurrences of arthrodire ‘placoderms’. While Late Devonian strata are rich in arthrodire remains, they are less common in older strata. Here, we describe a large tooth-bearing jaw element of Leptodontich- thys ziregensis gen. et sp. nov., an eubrachythoracid arthrodire from the Middle Devonian of Morocco. This species is based on a large posterior superognathal with a strong dentition. The jawbone displays features considered syna- pomorphies of Late Devonian eubrachythoracid arthrodires, with one posterior and one lateral row of conical teeth oriented postero-lingually. μCT-images reveal internal structures including pulp cavities and dentinous tissues. The posterior orientation of the teeth and the traces of a putative occlusal contact on the lingual side of the bone imply that these teeth were hardly used for feeding. Similar to Compagopiscis and Plourdosteus, functional teeth were pos- sibly present during an earlier developmental stage and have been worn entirely. The morphological features of the jaw element suggest a close relationship with plourdosteids. Its size implies that the animal was rather large. Keywords: Arthrodira, Dentition, Food web, Givetian, Maïder basin, Palaeoecology Introduction important to reconstruct character evolution in early ‘Placoderms’ are considered as a paraphyletic grade vertebrates. -
Fishes Scales & Tails Scale Types 1
Phylum Chordata SUBPHYLUM VERTEBRATA Metameric chordates Linear series of cartilaginous or boney support (vertebrae) surrounding or replacing the notochord Expanded anterior portion of nervous system THE FISHES SCALES & TAILS SCALE TYPES 1. COSMOID (most primitive) First found on ostracaderm agnathans, thick & boney - composed of: Ganoine (enamel outer layer) Cosmine (thick under layer) Spongy bone Lamellar bone Perhaps selected for protection against eurypterids, but decreased flexibility 2. GANOID (primitive, still found on some living fish like gar) 3. PLACOID (old scale type found on the chondrichthyes) Dentine, tooth-like 4. CYCLOID (more recent scale type, found in modern osteichthyes) 5. CTENOID (most modern scale type, found in modern osteichthyes) TAILS HETEROCERCAL (primitive, still found on chondrichthyes) ABBREVIATED HETEROCERCAL (found on some primitive living fish like gar) DIPHYCERCAL (primitive, found on sarcopterygii) HOMOCERCAL (most modern, found on most modern osteichthyes) Agnatha (class) [connect the taxa] Cyclostomata (order) Placodermi Acanthodii (class) (class) Chondrichthyes (class) Osteichthyes (class) Actinopterygii (subclass) Sarcopterygii (subclass) Dipnoi (order) Crossopterygii (order) Ripidistia (suborder) Coelacanthiformes (suborder) Chondrostei (infra class) Holostei (infra class) Teleostei (infra class) CLASS AGNATHA ("without jaws") Most primitive - first fossils in Ordovician Bottom feeders, dorsal/ventral flattened Cosmoid scales (Ostracoderms) Pair of eyes + pineal eye - present in a few living fish and reptiles - regulates circadian rhythms Nine - seven gill pouches No paired appendages, medial nosril ORDER CYCLOSTOMATA (60 spp) Last living representatives - lampreys & hagfish Notochord not replaced by vertebrae Cartilaginous cranium, scaleless body Sea lamprey predaceous - horny teeth in buccal cavity & on tongue - secretes anti-coaggulant Lateral Line System No stomach or spleen 5 - 7 year life span - adults move into freshwater streams, spawn, & die. -
Mcgraw-HILL New York Chicago San Francisco Lisbon London Madrid Mexico City Milan New Delhi San Juan Seoul Singapore Sydney Toronto
BIOLOGY DEMYSTIFIED The Hon. Dr. Dale Pierre Layman, Ph.D. McGRAW-HILL New York Chicago San Francisco Lisbon London Madrid Mexico City Milan New Delhi San Juan Seoul Singapore Sydney Toronto [13:35 13/6/ 58 N:/4058 LAY- ebook_copyright 7.5x9.qxd 9/29/03 11:39 AM Page 1 Copyright © 2003 by The McGraw-Hill Companies, Inc. All rights reserved. Manufactured in the United States of America. Except as per- mitted under the United States Copyright Act of 1976, no part of this publication may be reproduced or distributed in any form or by any means, or stored in a database or retrieval system, without the prior written permission of the publisher. 0-07-143387-2 The material in this eBook also appears in the print version of this title: 0-07-141040-6. All trademarks are trademarks of their respective owners. Rather than put a trademark symbol after every occurrence of a trademarked name, we use names in an editorial fashion only, and to the benefit of the trademark owner, with no intention of infringement of the trademark. Where such designations appear in this book, they have been printed with initial caps. McGraw-Hill eBooks are available at special quantity discounts to use as premiums and sales promotions, or for use in corporate train- ing programs. For more information, please contact George Hoare, Special Sales, at [email protected] or (212) 904- 4069. TERMS OF USE This is a copyrighted work and The McGraw-Hill Companies, Inc. (“McGraw-Hill”) and its licensors reserve all rights in and to the work. -
Redescription of Yinostius Major (Arthrodira: Heterostiidae) from the Lower Devonian of China, and the Interrelationships of Brachythoraci
bs_bs_banner Zoological Journal of the Linnean Society, 2015. With 10 figures Redescription of Yinostius major (Arthrodira: Heterostiidae) from the Lower Devonian of China, and the interrelationships of Brachythoraci YOU-AN ZHU1,2, MIN ZHU1* and JUN-QING WANG1 1Key Laboratory of Vertebrate Evolution and Human Origins of Chinese Academy of Sciences, Institute of Vertebrate Paleontology and Paleoanthropology, Chinese Academy of Sciences, Beijing 100044, China 2University of Chinese Academy of Sciences, Beijing 100049, China Received 29 December 2014; revised 21 August 2015; accepted for publication 23 August 2015 Yinosteus major is a heterostiid arthrodire (Placodermi) from the Lower Devonian Jiucheng Formation of Yunnan Province, south-western China. A detailed redescription of this taxon reveals the morphology of neurocranium and visceral side of skull roof. Yinosteus major shows typical heterostiid characters such as anterodorsally positioned small orbits and rod-like anterior lateral plates. Its neurocranium resembles those of advanced eubrachythoracids rather than basal brachythoracids, and provides new morphological aspects in heterostiids. Phylogenetic analysis based on parsimony was conducted using a revised and expanded data matrix. The analysis yields a novel sce- nario on the brachythoracid interrelationships, which assigns Heterostiidae (including Heterostius ingens and Yinosteus major) as the sister group of Dunkleosteus amblyodoratus. The resulting phylogenetic scenario suggests that eubrachythoracids underwent a rapid diversification during the Emsian, representing the placoderm response to the Devonian Nekton Revolution. The instability of the relationships between major eubrachythoracid clades might have a connection to their longer ghost lineages than previous scenarios have implied. © 2015 The Linnean Society of London, Zoological Journal of the Linnean Society, 2015 doi: 10.1111/zoj.12356 ADDITIONAL KEYWORDS: Brachythoraci – Heterostiidae – morphology – phylogeny – Placodermi. -
Vertebrata: Placodermi) from the Famennian of Belgium
View metadata, citationGEOLOGICA and similar papers BELGICA at core.ac.uk (2005) 8/1-2: 51-67 brought to you by CORE provided by Open Marine Archive A NEW GROENLANDASPIDID ARTHRODIRE (VERTEBRATA: PLACODERMI) FROM THE FAMENNIAN OF BELGIUM Philippe JANVIER1, 2 and Gaël CLÉMENT1 1. UMR 5143 du CNRS, Département «Histoire de la Terre», Muséum National d’Histoire Naturelle, 8 rue Buff on, 75005 Paris, France. [email protected] 2. 7 e Natural History Museum, Cromwell Road, London SW7 5BD, United Kingdom (9 fi gures, 1 table and 2 plates) ABSTRACT. A new species of the arthrodire genus Groenlandaspis is described from the upper part of the Evieux Formation (Upper Famennian), based on several specimens collected from quarries at Modave and Villers-le-Temple, Liège Province, Belgium. It is the fi rst occurrence of this widespread genus in continental Europe. O is new species is characterized by an almost smooth dermal armour, except for some scattered tubercles on its skull roof, median dorsal and spinal plates. Its median dorsal plate is triangular in shape and almost perfectly equilateral in lateral aspect and bears large, spiniform denticles on its posterior edge. All these Groenlandaspis remains occur in micaceous, dolomitic claystones or siltstones probably deposited in a subtidal environment. Outcrops of the same area have yielded other vertebrate remains, such as the placoderms Phyllolepis and Bothriolepis, acanthodians, various piscine sarcopterygians (Holoptychius, dipnoans, a rhizodontid, Megalichthys, Eusthenodon and a large tristichopterid), and a tetrapod that is probably close to Ichthyostega. O e biogeographical history of the genus Groenlandaspis is briefl y outlined, and the late Frasnian-Famennian interchange of vertebrate taxa between Gondwana and Euramerica is discussed. -
The Upper Devonian Fish <I>Bothriolepis</I> (Placodermi
AUSTRALIAN MUSEUM SCIENTIFIC PUBLICATIONS Johanson, Z., 1998. The Upper Devonian fish Bothriolepis (Placodermi: Antiarchi) from near Canowindra, New South Wales, Australia. Records of the Australian Museum 50(3): 315–348. [25 November 1998]. doi:10.3853/j.0067-1975.50.1998.1289 ISSN 0067-1975 Published by the Australian Museum, Sydney naturenature cultureculture discover discover AustralianAustralian Museum Museum science science is is freely freely accessible accessible online online at at www.australianmuseum.net.au/publications/www.australianmuseum.net.au/publications/ 66 CollegeCollege Street,Street, SydneySydney NSWNSW 2010,2010, AustraliaAustralia Records of the Australian Museum (1998) Vo!. 50: 315-348. ISSN 0067-1975 The Upper Devonian Fish Bothriolepis (Placodermi: Antiarchi) from near Canowindra, New South Wales, Australia ZERINA JOHANSON Palaeontology Section, Australian Museum, 6 College Street, Sydney NSW 2000, Australia [email protected] ABSTRACT. The Upper Devonian fish fauna from near Canowindra, New South Wales, occurs on a single bedding plane, and represents the remains of one Devonian palaeocommunity. Over 3000 fish have been collected, predominantly the antiarchs Remigolepis walkeri Johanson, 1997a, and Bothriolepis yeungae n.sp. The nature of the preservation of the Canowindra fauna suggests these fish became isolated in an ephemeral pool of water that subsequently dried within a relatively short space of time. This event occurred in a non-reproductive period, which, along with predation in the temporary pool, accounts for the lower number of juvenile antiarchs preserved in the fauna. Thus, a mass mortality population profile can have fewer juveniles than might be expected. The hypothesis that a single species of Bothriolepis is present in the Canowindra fauna is based on the consistent presence of a trifid preorbital recess on the internal headshield and separation of a reduced anterior process of the submarginal plate from the posterior process by a wide, open notch. -
Chondrichthyes (Cartilaginous Fishes)
PHYLUM: CHORDATA SUBPHYLUM: VERTEBRATE (CRANIATA) SUPERCLASIS: GNATHASTOMATA (JAWED FISHES) CLASS: CHONDRICHTYES (CARTILAGINOUS FISHES) GENERAL FEATURES OF PISCES BODY PLAN SCALE TYPES CAUDAL FIN TYPES VARIATIONS OF CAUDAL FIN Pinul FISH MOUTH TYPES POSITIONS OF PELVIC FINS POSITIONS OF PECTORAL FINS LATERAL LINE CLASS I: CHONDRICHTHYES EXTERNAL FEATURES-MORPHOLOGY Trunk Head Tail Body Fusiform Body dorsoventrally compressed Median Fins Heterocercal caudal fin Placoid scales (Homologous mammalian teeth) (5-7 pairs Median fin Paired Fins It is the first time seen in Anal fin is not found cartilaginous fish Chimaeras in Squalus Teeth covered with enamel Operculum Diphycercal caudal fin MALE FEMALE Mermaid’s purse Many sharks are ovovoviparous. There is no shell in the eggs. The egg opens in the animal's body cavity (Squalus acanthias) SYSTEMATICS PHYLUM: CHORDATA SUBPHYLUM: VERTEBRATE (CRANIATA) SUPERCLASIS: GNATHASTOMATA (JAWED FISHES) CLASS I: CHONDRICHTYES (CARTILAGINOUS FISHES) SUBCLASS I: PLACODERMI (FOSSIL FORMS) Primitive jawed; Simple hyoid arch and has single (unique) gill slit. No spiracles. Body is covered with bone plate or scales. The skeleton is bone-shaped. Climatius Acanthodius Dinichthys Bothriolepis SUBCLASS II:XENACANTHIFORMES Fossil Freshwater Sharks Xenacanthus SUBCLASS III: Cladoselachiformes Fossil marine sharks SUBCLASS IV: Elasmobranchii Sharks, Skates and Rays I. ORDO: SQUALIFORMES (PLEUROTREMATA) Placoid scales or derivatives (scutes and spines) usually present 5-7 pairs of gill arches and gill slits in seperate clefts along pharynx Upper jaw not fused to cranium About 937 species Mostly marine Hexanchus griseus Heptranchias perlo Scyliorhinus sterllaris Squatina oculata Rhinobatos rhinobatos II. ORDO: RAJIFORMES (SKATES) Torpedo nobiliana Raja alba Torpedo torpedo Pteromylaeus bovinus Raja montagui (Vatoz) SUBCLASIS V: HOLOCEPHALI -CHIMAERAS It has four pairs of gills, but a pair of gill openings. -
Ptyctodontid Fishes (Vertebrata, Placodermi) from the Late Devonian Gogo Formation, Western Australia, with a Revision of the European Genus Ctenurella 0Rvig, 1960
Ptyctodontid fishes (Vertebrata, Placodermi) from the Late Devonian Gogo Formation, Western Australia, with a revision of the European genus Ctenurella 0rvig, 1960 John A. LONG Department of Earth and Planetary Sciences, The Western Australian Museum, Francis Street, Perth, Western Australia, 6000 (Australia) Long J. A. 1997. — Ptyctodontid fishes (Vertebrata, Placodermi) from the Late Devonian Gogo Formation, Western Australia, with a revision of the European genus Ctenurella 0rvlg, 1960. Geodiversitas 19 (3) : 515-555. ABSTRACT A new, almost complete specimen of the ptyctodontid placoderm Campbellodus decipiens Miles et Young, 1977 enables description of the skull toof, trunk shield, visceral skeleton, pelvic girdle, dermal scale cover, and parts of the vertebtal column. A new reconstruction of the head shield of Ctenurella gladbachensis 0rvig, 1960 from Bergisch-Gladbach permits this taxon to be genetically defined from the Gogo species pteviously referred to that genus. The Gogo form is here referred to Austroptyctodus n.g. A new spe KEYWORDS cimen of Austroptyctodus gardineri Miles et Young, 1977, together with new Ptyctodontida, observations of Chelyophorus verneuili Agassiz, 1844 and Ctenurella gladba Devonian, chensis 0rvig, 1960, shows new information fot the endocranium, the hyoid Placodermi, Gogo, arch and visceral skeleton, identifying the previously identified "metaptery- Australia, goid" elements as paired nasal bones. The large visceral skeleton bone poste Austroptyctodus n.g., rior to the jaw joint in ptyctodontids is here identified as an elongated Ctenurella, Chelyophorus. interhyal. RESUME Une nouvelle description du toit crânien, de la cuirasse thoracique, du sque lette viscéral, de la ceinture pelvienne, de l'écaillure et de quelques éléments de la colonne vertébtale est proposée à partit d'un nouveau spécimen sub complet du ptyctodonte Campbellodus decipiens Miles et Young, 1977. -
PHYLUM CHORDATA Subphylum VERTEBRATA FISHES
Natural Sciences 360 Legacy of Life Lecture 10 Dr. Stuart Sumida PHYLUM CHORDATA Subphylum VERTEBRATA FISHES So, then what’s a vertebrate…? Phylogenetic Context for Vertebrata: Vertebrates are chordates Echinodermata Hemichordata Urochordata Cephalochordata Chordata Vertebrata All vertebrates possess an embryological material known as NEURAL CREST. Neural crest gives rise to particular structures found in all vertebrates, and only in vertebrates. Phylogenetic Context for Vertebrata Echinodermata Hemichordata Urochordata Cephalochordata Vertebrata (Posess neural crest and its derivatives) EVERYONE will be able to demonstrate a cross-sectional view of a vertebrate… Remember the basic chordate features: •Dorsal Hollow Nerve Cord •Notochord •Pharyngeal Gill Slits •Post Anal Tail So what exactly is a fish…? Jawless fish Conodonts Placodermi Chondrichthyes Gnathostomata Acanthodii Actinopterygii Osteichthyes (“Bony Fish”) Sarcopterygii THE ORIGINAL CONDITION Jawless fish OF VEWRTEBRATES WAS WITHOUT JAWS Conodonts Placodermi Chondrichthyes Gnathostomata Acanthodii Actinopterygii Osteichthyes (“Bony Fish”) Sarcopterygii Jawless fish Conodonts Placodermi Chondrichthyes Gnathostomata Acanthodii Actinopterygii Osteichthyes (“Bony Fish”) Sarcopterygii CONDONTS: Originally thought not to be vertebrates, but their best known components made of same material as teeth and bones (probably from neural crest material) CONDONTS: Originally thought not to be vertebrates, but their best known components made of same material as teeth and bones (probably from