OF the NETHERLANDS INTRODUCTION Until Now Very
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Pilot Project for the Definition of Environment-Friendly Measures To
European Commission - DG Environment Walloon Regional Government Province of Limburg - The Netherlands Province of Limburg - Belgium Waterboard Roer & Overmaas 3LORWSURMHFW IRUWKHGHILQLWLRQRIHQYLURQPHQWIULHQGO\PHDVXUHV WRUHGXFHWKHULVNIRUIODVKIORRGV LQWKH*HXO5LYHUFDWFKPHQW %HOJLXPDQGWKH1HWKHUODQGV B4-3040/97/730/JNB/C4 by S. Dautrebande, J.G.B. Leenaars, J.S. Smitz & E. Vanthournout (eds.) May 2000 European Commission - DG Environment Walloon Regional Government Province of Limburg - The Netherlands Province of Limburg - Belgium Waterboard Roer & Overmaas 7$%/(2)&217(176 5SCENARIO DEVELOPMENT .......................................................................................................... 59 5.1 INTRODUCTION........................................................................................................................... 59 5.2 APPROACH ................................................................................................................................. 59 5.3 IDENTIFICATION OF ALL POTENTIAL MEASURES ......................................................................... 60 5.4 GROUPING OF MEASURES ........................................................................................................... 60 5.5 RANKING OF MEASURES PER ‘MODEL-BASED’ GROUP ................................................................ 61 5.6 SELECTION OF SCENARIOS TO BE USED IN THE STUDY ................................................................ 62 +<'52/2*,&$/678'< 6.1 INTRODUCTION.......................................................................................................................... -
Comparative Functional Morphology of Attachment Devices in Arachnida
Comparative functional morphology of attachment devices in Arachnida Vergleichende Funktionsmorphologie der Haftstrukturen bei Spinnentieren (Arthropoda: Arachnida) DISSERTATION zur Erlangung des akademischen Grades doctor rerum naturalium (Dr. rer. nat.) an der Mathematisch-Naturwissenschaftlichen Fakultät der Christian-Albrechts-Universität zu Kiel vorgelegt von Jonas Otto Wolff geboren am 20. September 1986 in Bergen auf Rügen Kiel, den 2. Juni 2015 Erster Gutachter: Prof. Stanislav N. Gorb _ Zweiter Gutachter: Dr. Dirk Brandis _ Tag der mündlichen Prüfung: 17. Juli 2015 _ Zum Druck genehmigt: 17. Juli 2015 _ gez. Prof. Dr. Wolfgang J. Duschl, Dekan Acknowledgements I owe Prof. Stanislav Gorb a great debt of gratitude. He taught me all skills to get a researcher and gave me all freedom to follow my ideas. I am very thankful for the opportunity to work in an active, fruitful and friendly research environment, with an interdisciplinary team and excellent laboratory equipment. I like to express my gratitude to Esther Appel, Joachim Oesert and Dr. Jan Michels for their kind and enthusiastic support on microscopy techniques. I thank Dr. Thomas Kleinteich and Dr. Jana Willkommen for their guidance on the µCt. For the fruitful discussions and numerous information on physical questions I like to thank Dr. Lars Heepe. I thank Dr. Clemens Schaber for his collaboration and great ideas on how to measure the adhesive forces of the tiny glue droplets of harvestmen. I thank Angela Veenendaal and Bettina Sattler for their kind help on administration issues. Especially I thank my students Ingo Grawe, Fabienne Frost, Marina Wirth and André Karstedt for their commitment and input of ideas. -
Nuevos Datos Sobre Megabunus Diadema (Fabricius, 1779) (Opiliones: Phalangiidae)
Revista Ibérica de Aracnología, nº 22 (30/06/2013): 102–106. NOTA CIENTÍFICA Grupo Ibérico de Aracnología (S.E.A.). ISSN: 1576 - 9518. http://www.sea-entomologia.org/ Nuevos datos sobre Megabunus diadema (Fabricius, 1779) (Opiliones: Phalangiidae) Izaskun Merino-Sáinz1, Fernando A. Fernández-Álvarez1 & Carlos E. Prieto2 1 Departamento de Biología de Organismos y Sistemas, Universidad de Oviedo. C/Catedrático Uría s/n 33071 Oviedo (España) – [email protected] – [email protected] 2 Departamento de Zoología y Biología Celular Animal, Facultad de Ciencia y Tecnología, Universidad del País Vasco (UPV/EHU). Apdo. 644, 48080 Bilbao (España) – [email protected] Resumen: Megabunus diadema (Fabricius, 1779) es un falángido inconfundible con distribución europea occidental. Es conocido de Islandia, Noruega, Islas Feroe, Islas Británicas, Normandía y Península Ibérica, donde se extiende a lo largo de los Pirineos y la Cordille- ra Cantábrica. Los nuevos registros aportados (primeras citas para Andorra, Álava, Bizkaia, Burgos, La Rioja y Zamora) dan continuidad a la distribución ya conocida, extienden su área conocida al extremo norte del Sistema Ibérico y aportan nueva información sobre su fenología. Palabras clave:Opiliones, Phalangiidae, distribución, fenología, Península Ibérica. New data on Megabunus diadema (Fabricius, 1779) (Opiliones: Phalangiidae) Abstract: Megabunus diadema (Fabricius, 1779) is an unmistakable phalangiid with a West-European distribution range. It has been re- corded from Iceland, Norway, the Faroe Islands, the British Isles, Normandy and the Iberian Peninsula, where it occurs along the Pyre- nees and Cantabrian Mountains. The new data (first records from Andorra, Alava, Bizkaia, Burgos, La Rioja and Zamora) give continuity to the distribution already known, extend its known range to the northern end of the Sistema Ibérico mountains and provide new informa- tion on the phenology of the species. -
Opiliones, Palpatores, Caddoidea)
Shear, W. A. 1975 . The opilionid family Caddidae in North America, with notes on species from othe r regions (Opiliones, Palpatores, Caddoidea) . J. Arachnol . 2:65-88 . THE OPILIONID FAMILY CADDIDAE IN NORTH AMERICA, WITH NOTES ON SPECIES FROM OTHER REGION S (OPILIONES, PALPATORES, CADDOIDEA ) William A . Shear Biology Departmen t Hampden-Sydney, College Hampden-Sydney, Virginia 23943 ABSTRACT Species belonging to the opilionid genera Caddo, Acropsopilio, Austropsopilio and Cadella are herein considered to constitute the family Caddidae . The subfamily Caddinae contains the genu s Caddo ; the other genera are placed in the subfamily Acropsopilioninae. It is suggested that the palpatorid Opiliones be grouped in three superfamilies : Caddoidea (including the family Caddidae) , Phalangioidea (including the families Phalangiidae, Liobunidae, Neopilionidae and Sclerosomatidae ) and Troguloidea (including the families Trogulidae, Nemostomatidae, Ischyropsalidae an d Sabaconidae). North American members of the Caddidae are discussed in detail, and a new species , Caddo pepperella, is described . The North American caddids appear to be mostly parthenogenetic, an d C. pepperella is very likely a neotenic isolate of C. agilis. Illustrations and taxonomic notes ar e provided for the majority of the exotic species of the family . INTRODUCTION Considerable confusion has surrounded the taxonomy of the order Opiliones in North America, since the early work of the prolific Nathan Banks, who described many of ou r species in the last decade of the 1800's and the first few years of this century. For many species, no additional descriptive material has been published following the original de- scriptions, most of which were brief and concentrated on such characters as color and body proportions . -
Harvest-Spiders 515
PROVISIONAL ATLAS OF THE REF HARVEST-SPIDERS 515. 41.3 (ARACHNIDA:OPILIONES) OF THE BRITISH ISLES J H P SANKEY art å • r yz( I is -..a .e_I • UI II I AL _ A L _ • cta • • .. az . • 4fe a stir- • BIOLOGICAL RECORDS CENTRE Natural Environment Research Council Printed in Great Britain by Henry Ling Ltd at the Dorset Press, Dorchester, Dorset ONERC Copyright 1988 Published in 1988 by Institute of Terrestrial Ecålogy Merlewood Research Station GRANGE-OVER-SANDS Cumbria LA1/ 6JU ISBN 1 870393 10 4 The institute of Terrestrial Ecology (ITE) was established in 1973, from the former Nature Conservancy's research stations and staff, joined later by the Institute of Tree Biology and the Culture Centre of Algae and Protozoa. ITO contribbtes to, and draws upon, the collective knowledge of the 14 sister institutes which make up the Natural Environment Research Council, spanning all the environmental sciences. The Institute studies the factors determining the structure, composition and processes of land and freshwater systems, and of individual plant and animal species. It is developing a sounder scientific basis for predicting and modelling environmental trends arising from natural or man-made change. The results of this research are available to those responsible for the protection, management and wise use of our natural resources. One quarter of ITE's work is research commissioned by customers, such as the Department of Environment, the European Economic Community, the Nature Conservancy Council and the Overseas Development Administration. The remainder is fundamental research supported by NERC. ITE's expertise is widely used by international organizations In overseas projects and programmes of research. -
De Hooiwagens 1St Revision14
Table of Contents INTRODUCTION ............................................................................................................................................................ 2 CHARACTERISTICS OF HARVESTMEN ............................................................................................................................ 2 GROUPS SIMILAR TO HARVESTMEN ............................................................................................................................. 3 PREVIOUS PUBLICATIONS ............................................................................................................................................. 3 BIOLOGY ......................................................................................................................................................................... 3 LIFE CYCLE ..................................................................................................................................................................... 3 MATING AND EGG-LAYING ........................................................................................................................................... 4 FOOD ............................................................................................................................................................................. 4 DEFENCE ........................................................................................................................................................................ 4 PHORESY, -
Arachnida, Solifugae) with Special Focus on Functional Analyses and Phylogenetic Interpretations
HISTOLOGY AND ULTRASTRUCTURE OF SOLIFUGES Comparative studies of organ systems of solifuges (Arachnida, Solifugae) with special focus on functional analyses and phylogenetic interpretations HISTOLOGIE UND ULTRASTRUKTUR DER SOLIFUGEN Vergleichende Studien an Organsystemen der Solifugen (Arachnida, Solifugae) mit Schwerpunkt auf funktionellen Analysen und phylogenetischen Interpretationen I N A U G U R A L D I S S E R T A T I O N zur Erlangung des akademischen Grades doctor rerum naturalium (Dr. rer. nat.) an der Mathematisch-Naturwissenschaftlichen Fakultät der Ernst-Moritz-Arndt-Universität Greifswald vorgelegt von Anja Elisabeth Klann geboren am 28.November 1976 in Bremen Greifswald, den 04.06.2009 Dekan ........................................................................................................Prof. Dr. Klaus Fesser Prof. Dr. Dr. h.c. Gerd Alberti Erster Gutachter .......................................................................................... Zweiter Gutachter ........................................................................................Prof. Dr. Romano Dallai Tag der Promotion ........................................................................................15.09.2009 Content Summary ..........................................................................................1 Zusammenfassung ..........................................................................5 Acknowledgments ..........................................................................9 1. Introduction ............................................................................ -
Durham E-Theses
Durham E-Theses A study of the activity of six species of harvest spiders (arachnida, opiliones) Lenton, Graham M How to cite: Lenton, Graham M (1974) A study of the activity of six species of harvest spiders (arachnida, opiliones), Durham theses, Durham University. Available at Durham E-Theses Online: http://etheses.dur.ac.uk/8916/ Use policy The full-text may be used and/or reproduced, and given to third parties in any format or medium, without prior permission or charge, for personal research or study, educational, or not-for-prot purposes provided that: • a full bibliographic reference is made to the original source • a link is made to the metadata record in Durham E-Theses • the full-text is not changed in any way The full-text must not be sold in any format or medium without the formal permission of the copyright holders. Please consult the full Durham E-Theses policy for further details. Academic Support Oce, Durham University, University Oce, Old Elvet, Durham DH1 3HP e-mail: [email protected] Tel: +44 0191 334 6107 http://etheses.dur.ac.uk A STUDY OF THE ACTIVITY OF SIX SPECIES OF HARVEST - SPIDERS (ARACHNIDA, OPILIONES). by GRAHAM M. LENTON B.Sc (READING), Dip.Ed. (VALES) Being A thesis submitted as part of the requirements for the degree of Master of Science (Advcinced Course in Ecology) in the University of Durham, September 1970. a7 NOV 1974 . SECTION , LfBRARl, CONTENTS PAGE I INTRODUCTION 1 II MATERIALS AND METHODS 3 a) The Species b) The study site, Meteorological Data, Time periods. -
Behavioral Roles of the Sexually Dimorphic Structures in the Male Harvestman, Phalangium Opilio (Opiliones, Phalangiidae)
1763 Behavioral roles of the sexually dimorphic structures in the male harvestman, Phalangium opilio (Opiliones, Phalangiidae) Rodrigo H. Willemart, Jean-Pierre Farine, Alfredo V. Peretti, and Pedro Gnaspini Abstract: In various animal species, male sexual dimorphic characters may be used during intrasexual contests as orna- ments to attract females, or to hold them before, during, or after copulation. In the well-known harvestman, Phalangium opilio L., 1758, the behavioral functions of these male sexually dimorphic structures have never been studied in detail. Therefore, in addition to a morphometric study, 21 male contests and 43 sexual interactions were analyzed. Our observa- tions revealed that during contests, the male cheliceral horns form a surface by which the contestants use to push each other face-to-face while rapidly tapping their long pedipalps against the pedipalps of the opponent, occasionally twisting the opponent’s pedipalp. Scanning electron micrographs revealed contact mechanoreceptors on the pedipalp that would de- tect the intensity–frequency of contact with the contender’s pedipalp. Larger males won almost all contests, whereas the loser rapidly fled. During sexual interactions, the longer pedipalps of the male held legs IV of the female, whereas males with shorter pedipalps held the female by legs III. No contact with the male pedipalps and chelicerae by the females was visible before, during, or after copulation. Soon after copulating, males typically bent over the female, positioning their cheliceral horns against the females’s dorsum. Consequently, our data show that the cheliceral horns and the longer pedi- palps of the male seem to play an important role, during both intersexual and intrasexual encountering. -
The Phylogeny of Fossil Whip Spiders Russell J
Garwood et al. BMC Evolutionary Biology (2017) 17:105 DOI 10.1186/s12862-017-0931-1 RESEARCH ARTICLE Open Access The phylogeny of fossil whip spiders Russell J. Garwood1,2*, Jason A. Dunlop3, Brian J. Knecht4 and Thomas A. Hegna4 Abstract Background: Arachnids are a highly successful group of land-dwelling arthropods. They are major contributors to modern terrestrial ecosystems, and have a deep evolutionary history. Whip spiders (Arachnida, Amblypygi), are one of the smaller arachnid orders with ca. 190 living species. Here we restudy one of the oldest fossil representatives of the group, Graeophonus anglicus Pocock, 1911 from the Late Carboniferous (Duckmantian, ca. 315 Ma) British Middle Coal Measures of the West Midlands, UK. Using X-ray microtomography, our principal aim was to resolve details of the limbs and mouthparts which would allow us to test whether this fossil belongs in the extant, relict family Paracharontidae; represented today by a single, blind species Paracharon caecus Hansen, 1921. Results: Tomography reveals several novel and significant character states for G. anglicus; most notably in the chelicerae, pedipalps and walking legs. These allowed it to be scored into a phylogenetic analysis together with the recently described Paracharonopsis cambayensis Engel & Grimaldi, 2014 from the Eocene (ca. 52 Ma) Cambay amber, and Kronocharon prendinii Engel & Grimaldi, 2014 from Cretaceous (ca. 99 Ma) Burmese amber. We recovered relationships of the form ((Graeophonus (Paracharonopsis + Paracharon)) + (Charinus (Stygophrynus (Kronocharon (Charon (Musicodamon + Paraphrynus)))))). This tree largely reflects Peter Weygoldt’s 1996 classification with its basic split into Paleoamblypygi and Euamblypygi lineages; we were able to score several of his characters for the first time in fossils. -
A Stable Phylogenomic Classification of Travunioidea (Arachnida, Opiliones, Laniatores) Based on Sequence Capture of Ultraconserved Elements
A stable phylogenomic classification of Travunioidea (Arachnida, Opiliones, Laniatores) based on sequence capture of ultraconserved elements The Harvard community has made this article openly available. Please share how this access benefits you. Your story matters Citation Derkarabetian, Shahan, James Starrett, Nobuo Tsurusaki, Darrell Ubick, Stephanie Castillo, and Marshal Hedin. 2018. “A stable phylogenomic classification of Travunioidea (Arachnida, Opiliones, Laniatores) based on sequence capture of ultraconserved elements.” ZooKeys (760): 1-36. doi:10.3897/zookeys.760.24937. http://dx.doi.org/10.3897/zookeys.760.24937. Published Version doi:10.3897/zookeys.760.24937 Citable link http://nrs.harvard.edu/urn-3:HUL.InstRepos:37298544 Terms of Use This article was downloaded from Harvard University’s DASH repository, and is made available under the terms and conditions applicable to Other Posted Material, as set forth at http:// nrs.harvard.edu/urn-3:HUL.InstRepos:dash.current.terms-of- use#LAA A peer-reviewed open-access journal ZooKeys 760: 1–36 (2018) A stable phylogenomic classification of Travunioidea... 1 doi: 10.3897/zookeys.760.24937 RESEARCH ARTICLE http://zookeys.pensoft.net Launched to accelerate biodiversity research A stable phylogenomic classification of Travunioidea (Arachnida, Opiliones, Laniatores) based on sequence capture of ultraconserved elements Shahan Derkarabetian1,2,7 , James Starrett3, Nobuo Tsurusaki4, Darrell Ubick5, Stephanie Castillo6, Marshal Hedin1 1 Department of Biology, San Diego State University, San -
IIIIIIIIIIH Raadsnummer: BP/389
GEMEENTE Gulpen-Wittem GEMEENTEBESTUUR VAN GULPEN-WITTEM Kenmerk : G. 19.00844 IIIIIIIIIIH Raadsnummer: BP/389 Dossiercode: Ondenverp: Bedrijfsuitbreiding - enverplaatsing landbouwbedrijf Voncken Trintelen De raad van de gemeente Gulpen-Wittem Gezien het voorstel van burgemeester en wethouders d.d. 9 april 2019 inzake bovengenoemd onderwerp. Gelet op de Gemeentewet, de algemene wet Bestuursrecht alsmede het verzoek zoals geformuleerd in de brief van de Provincie Limburg omtrent de bedrijfsontwikkeling casus Voncken Trintelen Besluit 1) In beginsel medewerking te verlenen aan een pilot waarin de haalbaarheid van een mogelijke bedrijfsvestiging voor Voncken binnen Trintelen of directe omgeving breed onderzocht wordt, onder voorwaarden dat: • de pilot (ontwerpend onderzoek) een landschappelijk acceptabele invulling oplevert en daarnaastde ontwikkeling getuigt van duurzame bedrijfsvoeringen een bijdrage levert aan maatschappelijke doelstellingen (maatschappelijk verantwoord ondernemerschap); • de bestaande bedrijfslocatie aan de Eyserweg wordt gesaneerd, herbestemd en overtollige bedrijfsbebouwing wordt gesloopt. • het college de agrarische ondernemer Voncken een bedrijfsvisie en businessplan laat opstellen, dat geëntis op de agrariër van de toekomst ineen kwetsbaar- landschap. Met de volgende aandachtspunten o hoogwaardige architectuur o duurzame bedrijfsvormingzoveel mogelijk gericht op een circulaire economie GEMEENTE Gulpen-Wittem o boerderij die ook een bijdrage levert aan doelstellingen op het gebied van klimaat, energie en milieu o