DOCSLIB.ORG

Djvu Document

Total Page：16

File Type：pdf, Size：1020Kb

Download full-text PDF Read full-text
Djvu Document BULL. BOT. SURV. INDIA Vol. 4, Nos. 14: pp. 27-38, 1W CYTOTAXONOMY OF AMARYLLIDACEAE Cytogenctics Labmatmy, Dcwtmcnt of Botany, Cakutta Uni~wsiy,Calcutta-19 ABSTRACT Chromosome numbers are now known for about 46 genera in the family Amaryllidaceae. The lowest basic number has been found to be 5 and the highest 30. Besides these two, 6, 7, 8, 9, 10, 11, 12, 14, 15, 16, 19, 23 and 29 have also been encountered of which 11 is found in majority oc the genera. In a number of genera the derivation of one basic number from the other bas been established. In the genus Allium largest nu*er of species have been studied cytologically. The nert,largest is Narcissus followed by <ephpnthes, Crhum, Hif&astrum and Lyaris in that order. B or supernumerary chromosomes have been observed in Agnpanthq Allim, Crinum, -aa Haemanthus, Hippcmtrum, Ljcoris and ~arcisms. In this family repatterning of chromosomes through inversions (both para ahd pericentric), translocation, polypbidy (eu-, aneuo-, hetero-, auto-, and allopolyploidy) and hybridization hwr: pla ed prominent role in chromosome number evolution, karyotype alteration and speciation. In ad&tion to this, gene mutation has also pla ed significant puf in speciation. Ready vegetative propagation has helped in maintaining sterile KYbrids and thole forms with numerical md smi?Wa\ changes of chromosomes. Apomixis has also been responsible in chromosome number evolution m some genera. A consideration of taxonomic work done in Amaryllidaceae points out that Hutchinson's system of classification, based on the umbellate inflorescence rather than on the pajition of. the wary, ha been found to be more ph logenetic but the amendments proposed by several workers for ahangm in tribal and generic level could be taken into consideration and in some cases may be justified, INTRODUCTION of small chromosomes, yet the former was placed in Liliaceae and the latter in Amaryllidaceae by The family Amaryllidaceae is very interesting earlier taxonomists, On the basis df their cytologk from cytotaxonomic point of view. Cytologically, 'most of the species belonging to this family hare cal findings the7 urged for keeping. them togeiher, small number of large chromosomes and somi: In Hutchinsons classification these two genera species of the genus Allium are being extensively have been put together in Agavaceae. The genus Fureraen, which has cytolo$cal similarities used for experimental studies involvin physical (Whitaker, 1934) with Yucca an Agave, has also and chemical agents. Taxonomical~y, t1 e family has attracted added im ortance since the delimita- been grouped with these two genera by Hutchinson An'derson (rgqo, ede Lawrence, 19~1)on the tion of the famil by Rutchinson (1g34, 1959) to of only those mem gers of petaloid monocotyledons, basis of his studies on the floral anatomy whlch have umbellate inflorescence. Liliales approved of Hutchinson's inclusion of Agapantheae, and Allieae in Amaryllidaceac. Cheadle (1942)also accepted the inclusion of Aga- CYTOTAXONOMIG CONSIDERATION pantheae, Allieae and Glllieseae in Amaryllidaceae on the basis of his anatomical work on mono- Hutchinson radically changed the concept of cotyledons. Maia (1941, vide Lawrence, ~ggl) the differentiation of Liliaceae and. ~maryllida- through his studies on the pollen grab of .some Feae formerly based on the position of ova'ky and monocotyledons also came to the same conclueion, recognized the importance of the umbellate type of Lawrence (1951) commented that the classification inflorescence as the unifyin ptinciple in Amarylli- based on the position of the ovary was not constant, daceae. For this reason, {e included the tr~bes since in the genera Ophiopogon, Bomareae and Agapantheae, Gilliesieae and Allieae in Amarylli- Hemerocallis of Liliaceae both su rior and inferior daceae, which were formerly placed in Liliaceae. ovaried plants are found apd F=or this reason he On the other hand, he excluded the tribes expressed the opinion that the classification of Hy~xideae,Alstromerieae, Agaveae, Vellozieae and Hutchinson, which was based on the type of in- Gnostyleae &om Amaryllidaceac. H~tchiison's florescence, was or more fundamental imporranp classification has been more or less su ported by and stable in nature. Flory and Yarnell (193 ) gave taxonomists and in many instances eviJ? ences have an account of chromosome numbers in the fEmero- been brought forward by anatomists and cytologists callideae, Alstromerialeo, and Amaryllidales follow* to su port him. As for example, Mckelvey and ing Hutchiison's classification but they supported Sax pn333). Whitaker (19%) and Granick ('MI pointed out that Yucca and Agme are quite similar cytologically in having 5 pairs of large and 25 .pairs 28 BULLETIN OF THE BOT'AMCAL SURVEY OF INDIA [VO~.4 Stout's (1932) view that so far as the basic Traub's (1957) system, basic numbers of chromo- number is concerned the genuc Hemerocallis does somes were also taken into account in the subfamily, not fit in Liliaceae, since it has a n number of r I, tribal and generic level. Later on, Bose (1958~) while an allied genus Hbsta has a basic number of commented on the taxonomic positions of the 30. On this ground they thought that further cyto- genera Sprekeliu and I,ycoris in the light of his cy~ taxonomic study of some genera in Amaryllidaceae logical findings. might: call for an inclusion of Hemerocallis in The above discussion shows that although Hut- Amaryllidaceae. Earlier, Whitaker (1934) and chiison's (1934) new system of classification based Granick (1944) removed the genus Hosta from on t$e umbellate inflorescence rather than on the Hemerocalleae and placed ,~tin the family position of the ovary has been found to be a more Agavaceae. natural system, the amendments proposed by several Sato (1942) observed that his results of karyotype workers for shifting tribes and genera could be taken analyses in different genera of Liliaceae and related into consideration and in some cases may be families agreed with Hutchinson's (1934) system of justified. classification. He pointed out that the karyotypes of some genera in Liliaceae and Amaryllidaceae BASIC NUMBERS IN THE FAMILY AhfARYLLIDACEAE were similar in morphology, which according t& In the genus Allium largest number of species in him, showed that Hutchinson's classification based the .family Amaryllidaceae have been studled cyto- on the umbellate inflorescence was more reason- logically. The next largest is Narcissus, followed by able than that based on the position of ovary.. Zephyranthes, CTinum, Hippeastrum and Lycoris Through his studies on the karyotypes of in order. As regards the chro'mosome numbers, Phormizrm, Hosta, Hemerocallis, Leucocrinurn, Table I shows that very varied chromosome num- Hespe~ocallis,Dracaena and Cordyline he supported bers have been reported from Alliurn, Zephyranthes, Hutchinson's opinion that the Amaryllidaceae Hymenocallis and some other genera. might have originated from the tribe Hemero- A glance at Table I will also show that in this callideae of Liliaceae. Sato (1942) agreeing with family the lowest basic number i's 5, and the highest Hutchinson, also suggested the transference of is 30. Besides these two, there are n=6, 7, 8, g, 10, Alstroemerieae into Liliaceae but su ported Nakai's I I, 12, 14, IS, 16, 19,23 and 29, of which I I is found , (vide Sato 1942) modification o!! Hutchinson's in majority of genera. In a number of genera the Later, Flory (I~M)gave another list of derivation of one basic number from the other has cTtern* romosome numbers in Hemerocallideae, Alstroe- been established, as for example, Fernandes (1946) meriales, and Amaryllidales basing on Hutchinson's has shown that the basic number ,I% in Narcissus, classification but took into account the changes is derived from n=7, while in a genus like Lycorisj suggested by some workers. Sharma . and Ghosh 6 or 1 I could be taken as the orignal basic number, not only supported the removal of the tribe although Inariyama (1g51b) has favoured I I as thk oxideae from Amaryllidaceae by Hutchinson original basic number on .the basis of fusion of rod but also accepted his creatlon of the family chromosomes. On the. other hand, if the frag- Hgpxidaceae. ' Mookerjea (1955) found cytological mentation of V chrornos~mesis taken into consi- corroboration of Hutchrnson's systtm in Amarylli- deration then '6 would be the original basic number daceae? and later, working with some members in Lycoris. This aspect will be taken up in detail of Liliaceae, Mookerjea (1956) did not agree to in the following discussion. his inclusion of Funkia in Hemerocalleae. Traub' 19~7)~while supporting the delimitation of the MECHANISM OF CHROMOSOME NUMBER EVOLU- by Hutchinson, proposed some new changes TION, KARYOTYPE ALTERATION AND SPECIATfON and presented a classification of the family IN AMARYLLIDACEAE Am Ilidaceae,.which was'a third revision of their In Amaryllidaceae repatterning of chromosomes (~rax,1938 and Traub and Moldenke, 1949, vide through inversions (both ~araand pericentric) and Traub, 1957) earlier classifications. On the .basis translocation, polyploid? (eu-, aneuq hetero-, of morphological and cytological similarities with auto-, and aHopolyploidy) and hybridization have; members of the tribe Allieae, A pantheae and played prominent role in chromosome number Gllliesiae, he removed the tribe Eemerocallideae evolution, karyotype alteration and speciation. Sato from Liliaceae and placed it in Amaryllidaceae. He (1938) and others have discussed the importance of also divided
Load more

Recommended publications
  • Allium Albanicum (Amaryllidaceae), a New Species from Balkans and Its
    A peer-reviewed open-access journal PhytoKeys 119: 117–136Allium (2019) albanicum (Amaryllidaceae), a new species from Balkans... 117 doi: 10.3897/phytokeys.119.30790 RESEARCH ARTICLE http://phytokeys.pensoft.net Launched to accelerate biodiversity research Allium albanicum (Amaryllidaceae), a new species from Balkans and its relationships with A. meteoricum Heldr. & Hausskn. ex Halácsy Salvatore Brullo1, Cristian Brullo2, Salvatore Cambria1, Giampietro Giusso del Galdo1, Cristina Salmeri2 1 Department of Biological, Geological and Environmental Sciences, Catania University, Via A. Longo 19, 95125 Catania, Italy 2 Department of Biological, Chemical and Pharmaceutical Sciences and Technologies (STEBICEF), Palermo University, Via Archirafi 38, 90123 Palermo, Italy Corresponding author: Cristina Salmeri ([email protected]) Academic editor: L. Peruzzi | Received 26 October 2018 | Accepted 9 January 2019 | Published 11 April 2019 Citation: Brullo S, Brullo C, Cambria S, Giusso del Galdo G, Salmeri C (2019) Allium albanicum (Amaryllidaceae), a new species from Balkans and its relationships with A. meteoricum Heldr. & Hausskn. ex Halácsy. PhytoKeys 119: 117–136. https://doi.org/10.3897/phytokeys.119.30790 Abstract A new species, Allium albanicum, is described and illustrated from Albania (Balkan Peninsula). It grows on serpentines or limestone in open rocky stands with a scattered distribution, mainly in mountain loca- tions. Previously, the populations of this geophyte were attributed to A. meteoricum Heldr. & Hausskn. ex Halácsy, described from a few localities of North and Central Greece. These two species indeed show close relationships, chiefly regarding some features of the spathe valves, inflorescence and floral parts. They also share the same diploid chromosome number 2n =16 and similar karyotype, while seed testa micro- sculptures and leaf anatomy reveal remarkable differences.
    [Show full text]
  • Complete Chloroplast Genomes Shed Light on Phylogenetic
    www.nature.com/scientificreports OPEN Complete chloroplast genomes shed light on phylogenetic relationships, divergence time, and biogeography of Allioideae (Amaryllidaceae) Ju Namgung1,4, Hoang Dang Khoa Do1,2,4, Changkyun Kim1, Hyeok Jae Choi3 & Joo‑Hwan Kim1* Allioideae includes economically important bulb crops such as garlic, onion, leeks, and some ornamental plants in Amaryllidaceae. Here, we reported the complete chloroplast genome (cpDNA) sequences of 17 species of Allioideae, fve of Amaryllidoideae, and one of Agapanthoideae. These cpDNA sequences represent 80 protein‑coding, 30 tRNA, and four rRNA genes, and range from 151,808 to 159,998 bp in length. Loss and pseudogenization of multiple genes (i.e., rps2, infA, and rpl22) appear to have occurred multiple times during the evolution of Alloideae. Additionally, eight mutation hotspots, including rps15-ycf1, rps16-trnQ-UUG, petG-trnW-CCA , psbA upstream, rpl32- trnL-UAG , ycf1, rpl22, matK, and ndhF, were identifed in the studied Allium species. Additionally, we present the frst phylogenomic analysis among the four tribes of Allioideae based on 74 cpDNA coding regions of 21 species of Allioideae, fve species of Amaryllidoideae, one species of Agapanthoideae, and fve species representing selected members of Asparagales. Our molecular phylogenomic results strongly support the monophyly of Allioideae, which is sister to Amaryllioideae. Within Allioideae, Tulbaghieae was sister to Gilliesieae‑Leucocoryneae whereas Allieae was sister to the clade of Tulbaghieae‑ Gilliesieae‑Leucocoryneae. Molecular dating analyses revealed the crown age of Allioideae in the Eocene (40.1 mya) followed by diferentiation of Allieae in the early Miocene (21.3 mya). The split of Gilliesieae from Leucocoryneae was estimated at 16.5 mya.
    [Show full text]
  • Narcissus Pests
    Bulletin 51 HMSO 13s Od [65p] net Narcissus Pests Ministry of Agriculture, Fisheries and Food MINISTRY OF AGRICULTURE, FISHERIES AND FOOD Narcissus Pests Bulletin 51 LONDON HER MAJESTY'S STATIONERY OFFICE 197o First published June 1932 Sixth edition 197o The Ministry does not accept responsibility for any of the private or trade advertisements included in this publication. SBN 11 240351 4 Foreword THE growers of Narcissus have been very fortunate in that the pests of this valuable crop have received specialist attention for nearly forty years. Names like W. E. H. Hodson and L. N. Staniland, both past authors of this Bulletin, rank high in the list of pioneer researchers on bulb pests in this country. They have been followed with no less enthusiasm by the con- tributors to this sixth edition which brings up-to-date our knowledge of the important pests of the crop and tested and practical methods of control. Although the present Bulletin mainly follows the pattern laid down by Mr. Hodson in 1932 many sections have been extensively rewritten. Mr. H. C. Woodville has dealt with narcissus flies as he did in 1958, and Mr. H. G. Morgan with detection of pests in the field, stem and other eelworms and their control. Mr. A. L. Winfield covered bulb scale mite and the general problem of hot-water treatment of bulbs, and chemical dips to control stem eelworm, and also contributed the notes on miscellaneous pests. Mr. P. Aitkenhead dealt with bulb mites. Mr. J. F. Southey provided the section dealing with eelworms as vectors of virus diseases of the crop.
    [Show full text]
  • (Tribe Haemantheae) Inferred from Plastid and Nuclear Non-Coding DNA Sequences
    Plant Syst. Evol. 244: 141–155 (2004) DOI 10.1007/s00606-003-0085-z Generic relationships among the baccate-fruited Amaryllidaceae (tribe Haemantheae) inferred from plastid and nuclear non-coding DNA sequences A. W. Meerow1, 2 and J. R. Clayton1 1 USDA-ARS-SHRS, National Germplasm Repository, Miami, Florida, USA 2 Fairchild Tropical Garden, Miami, Florida, USA Received October 22, 2002; accepted September 3, 2003 Published online: February 12, 2004 Ó Springer-Verlag 2004 Abstract. Using sequences from the plastid trnL-F Key words: Amaryllidaceae, Haemantheae, geo- region and nrDNA ITS, we investigated the phy- phytes, South Africa, monocotyledons, DNA, logeny of the fleshy-fruited African tribe Haeman- phylogenetics, systematics. theae of the Amaryllidaceae across 19 species representing all genera of the tribe. ITS and a Baccate fruits have evolved only once in the combined matrix produce the most resolute and Amaryllidaceae (Meerow et al. 1999), and well-supported tree with parsimony analysis. Two solely in Africa, but the genera possessing main clades are resolved, one comprising the them have not always been recognized as a monophyletic rhizomatous genera Clivia and Cryp- monophyletic group. Haemanthus L. and tostephanus, and a larger clade that unites Haemanthus and Scadoxus as sister genera to an Gethyllis L. were the first two genera of the Apodolirion/Gethyllis subclade. One of four group to be described (Linneaus 1753). Her- included Gethyllis species, G. lanuginosa, resolves bert (1837) placed Haemanthus (including as sister to Apodolirion with ITS. Relationships Scadoxus Raf.) and Clivia Lindl. in the tribe among the Clivia species are not in agreement with Amaryllidiformes, while Gethyllis was classi- a previous published phylogeny.
    [Show full text]
  • Genetic Diversity and Taxonomic Studies of Allium Akaka and A
    Journal of Horticultural Research 2017, vol. 25(1): 99–115 DOI: 10.1515/johr-2017-0011 _______________________________________________________________________________________________________ GENETIC DIVERSITY AND TAXONOMIC STUDIES OF ALLIUM AKAKA AND A. ELBURZENSE NATIVE TO IRAN USING MORPHOLOGICAL CHARACTERS Sajad JAFARI1, Mohammad Reza HASSANDOKHT*1, Mahdi TAHERI2, Abdolkarim KASHI1 1 Department of Horticultural Sciences, College of Agriculture and Natural Resources, University of Tehran, Karaj, Iran 2 Soil and Water Research Department, Zanjan Agriculture and Natural Resources Research and Educa- tion Center, Agricultural Research, Education and Extension Organization (AREEO), Zanjan, Iran Received: April 2017; Accepted: June 2017 ABSTRACT Two Allium species (A. akaka S.G. Gmelin and A. elburzense W.) native to Iran are used locally as the fresh vegetables and in medical therapy. They are not cultivated, but are collected from the wild, thus, will soon be threatened with extinction. In this study, the diversity of 15 wild accessions (4 accessions of A. elburzense endemic of Iran and 11 accessions of A. akaka) collected from the north-western part of Iran were evaluated with the use of 16 qualitative and 16 quantitative characteristics. The morphological char- acters with high heritability included leaf length, flower number in umbel, inflorescence diameter, leaf dry weight, bulb fresh weight, weight of 100 seeds, seed length and seed length/width. Results of the principal component analysis indicated that 92.52% of the observed variability was explained by the first six com- ponents. The first two components explained about 64.74% of the total observed variability. The first and third hierarchical cluster analysis included all accessions of A. akaka. The accessions of A.
    [Show full text]
  • Newsletter No. 4 Autumn 2005
    The Southern African Bulb Group Newsletter No. 4 Autumn 2005 The Southern African Bulb Group was initiated by a group of enthusiasts on April 4th 2004. The objective of the group is to further the understanding of the cultivation of Southern African bulbs, where `bulbs' is used in the broad sense to encompass bulb-, corm- and tuber- possessing Southern African plants, including `dicots' such as Oxalis. In the first instance the group will be run on an informal basis, with an initial subscription of £5 invited from participants. Committee: Robin Attrill (Membership secretary and Newsletter editor), Margaret Corina (Treasurer), Stefan Rau and Terry Smale Editorial This issue of the newsletter contains a report of the meeting of the group held at Rupert Bowlby's Nursery on April 9th 2005, an article on Crinum moorei by David Corina, information on recent literature of interest to growers of Southern African bulbs, and an updated list of suppliers of seed and bulbs/corms of Southern African geophytes. In addition a balance sheet covering the first year of operation of the group is attached. With respect to the supplier list, please let me know if you are aware of other sources which should be included. As I have previously stated the group welcomes articles, and suggestions, for inclusion in future newsletters. Contributions (hand/typewritten and electronic are acceptable!) are urgently required and should be sent to the newsletter editor at 17 Waterhouse Moor, Harlow, Essex, CM18 6BA (Email [email protected] ) Publication of the next issue is scheduled for December 2005. Report on visit to Rupert Bowlby - Saturday 9 th April 2005 by David Corina About 20 members attended the event, and the Group would like to thank Rupert for his hospitality at the event and for opening his collection to the public gaze.
    [Show full text]
  • Smoke-Induced Flowering in the Fire-Lily Cyrtanthus Ventricosus
    638 S.Afr.J.Bot., 1993,59(6) Short Communications / Kort Mededelings qualitative switch from dormancy to flowering, they have shown that these gases can speed up the timing of flowering and final percentage of flowering (e.g., Imanishi 1983; Imanishi & Fortanier 1982/83). Smoke-induced flowering in the fire-lily Experiments were set up to test the hypothesis that smoke Cyrtanthus ventricosus and ethylene are the cues that induce Cyrtanthus ventricosus to switch from dormancy to flowering. Jon E. Keeley In late spring (November) 1990, approximately 10 bulbs of this species were collected from flowering plants on a Department of Biology, Occidental College, 1600 Campus very recently burned site north-west of Caledon in the Cape Road, Los Angeles, California 90041, USA Province of South Africa. They were planted in a sandy­ Received 7 May 1993; revised 5 August 1993 loam planting bed in full sunlight in Los Angeles (USA) and were maintained for a little over 2 years, during which time Flowering of certain fynbos geophytes has long been noted they remained in a vegetative state, producing a couple of to have an obligate dependence upon fire. One of these spe­ leaves each spring (February - May). In mid-spring (March cies, Cyrtanthu5 ventric05U5 (Jacq.) Willd ., is shown here to 1993) a single bulb was planted in each of eight I-gallon be stimulated to flower by smoke. Ethylene, a gaseous com­ pots of sandy-loam soil. These were maintained in full ponent of smoke, did not stimulate flowering in this species. sunlight for at least a month. The experiment consisted of three treatments, each applied to two pots in separate Dit is lank reeds bekend dat blomvorming by sekere fynbos­ chambers.
    [Show full text]
  • PLANT YOUR YARD with WILDFLOWERSI Sources
    BOU /tJ, San Francisco, "The the beautiful, old Roth Golden Gate City," pro­ Estate with its lovely for­ vides a perfect setting for mal English gardens in the 41st Annual Meeting Woodside. Visit several of the American Horticul­ gardens by Tommy tural Society as we focus Church, one of the great­ on the influence of ori­ est garden-makers of the ental gardens, plant con­ century. Observe how the servation, and edible originator of the Califor­ landscaping. nia living garden incor­ Often referred to as porated both beauty and "the gateway to the Ori­ a place for everyday ac­ ent," San Francisco is tivities into one garden the "most Asian of occi­ area. dental cities." You will Come to San Fran­ delight in the beauty of cisco! Join Society mem­ its oriental gardens as bers and other meeting we study the nature and participants as we ex­ significance of oriental plore the "Beautiful and gardening and its influ­ Bountiful: Horticulture's ence on American horti­ Legacy to the Future." culture. A visit to the Japanese Tea Garden in the Golden Gate Park, a Please send me special advance registration information for the botanical treasure, will Society's 1986 Annual Meeting in offer one of the most au­ San Francisco, California. thentic examples of Japa­ NAME ________ nese landscape artistry outside of Japan. Tour the Demonstra­ Western Plants for Amer­ ~D~SS _______ tion Gardens of Sunset Explore with us the ican Gardens" as well as CITY ________ joys and practical aspects magazine, magnificent what plant conservation of edible landscaping, private gardens open only efforts are being made STATE ZIP ____ which allows one to en­ to Meeting participants, from both a world per­ joy both the beauty and and the 70-acre Strybing spective and a national MAIL TO: Annual Meeting, American Horticultural Society, the bounty of Arboretum.
    [Show full text]
  • Allium Toksanbaicum (Amaryllidaceae), a New Species from Southeast Kazakhstan
    Phytotaxa 494 (3): 251–267 ISSN 1179-3155 (print edition) https://www.mapress.com/j/pt/ PHYTOTAXA Copyright © 2021 Magnolia Press Article ISSN 1179-3163 (online edition) https://doi.org/10.11646/phytotaxa.494.3.1 Allium toksanbaicum (Amaryllidaceae), a new species from Southeast Kazakhstan NIKOLAI FRIESEN1,2,*, POLINA VESSELOVA3, BEKTEMIR OSMONALY3, GULNARA SITPAYEVA3, ALEXANDER LUFEROV2 & ALEXANDER SHMAKOV4 1Botanical Garden, University of Osnabrück, Albrechtstrasse 29, 49076, Osnabrück, Germany; [email protected]; http://orcid.org/0000-0003-3547-3257 2I.M. Sechenov First Moscow State Medical University Ministry of Health of the Russian Federation, Department of Pharmaceutical and Natural Sciences, Izmailovsky Boulevard 8, Moscow, 105043, Russia; [email protected] 3Institute of Botany and Phytointroduction of the Committee of Forestry and Wildlife belong to the Ministry of Ecology, Geology and Natural Resources of the Republic of Kazakhstan, Almaty, 480070 Kazakhstan; [email protected]; [email protected]; [email protected] 4Altai State University, Lenina str., 61; 656049, Barnaul, Russia; [email protected] *Corresponding author Abstract Allium toksanbaicum from South East Kazakhstan is described as a new species. Molecular markers reveal a close rela- tionship to A. obliquum and some other central Asian species of the section Oreiprason. We investigated the phylogenetic relationship of the new species based on sequences of two chloroplast spacers (rpl32-trnL and trnQ-rps16) and the nuclear ribosomal DNA internal transcribed spacer (ITS) region. The new species is diploid with a chromosome number of 2n = 2x = 16. A detailed morphological description, illustrations and karyotype features of the new species are given. With its falcate leaves, the new species is very similar to A.
    [Show full text]
  • MAPEAMENTO DOS SÍTIOS DE Dnar 5S E 45S E ORGANIZAÇÃO DA CROMATINA EM REPRESENTANTES DA FAMÍLIA AMARYLLIDACEAE JAUME ST.-HIL
    EMMANUELLY CALINA XAVIER RODRIGUES DOS SANTOS MAPEAMENTO DOS SÍTIOS DE DNAr 5S E 45S E ORGANIZAÇÃO DA CROMATINA EM REPRESENTANTES DA FAMÍLIA AMARYLLIDACEAE JAUME ST.-HIL. RECIFE-PE 2015 i EMMANUELLY CALINA XAVIER RODRIGUES DOS SANTOS MAPEAMENTO DOS SÍTIOS DE DNAr 5S E 45S E ORGANIZAÇÃO DA CROMATINA EM REPRESENTANTES DA FAMÍLIA AMARYLLIDACEAE JAUME ST.-HIL. Tese apresentada ao Programa de Pós-Graduação em Botânica da Universidade Federal Rural de Pernambuco como parte dos requisitos para obtenção do título de Doutora em Botânica. Orientador: Prof. Dr. Reginaldo de Carvalho Dept° de Genética/Biologia, Área de Genética/UFRPE Co-orientador: Prof. Dr. Leonardo Pessoa Felix Dept° de Fitotecnia, UFPB RECIFE-PE 2015 ii MAPEAMENTO DOS SÍTIOS DE DNAr 5S E 45S E ORGANIZAÇÃO DA CROMATINA EM REPRESENTANTES DA FAMÍLIA AMARYLLIDACEAE JAUME ST.-HIL. Emmanuelly Calina Xavier Rodrigues dos Santos Tese defendida e _________________ pela banca examinadora em ___/___/___ Presidente da Banca/Orientador: ______________________________________________ Dr. Reginaldo de Carvalho (Universidade Federal Rural de Pernambuco – UFRPE) Comissão Examinadora: Membros titulares: ______________________________________________ Dra. Ana Emília de Barros e Silva (Universidade Federal da Paraíba – UFPB) ______________________________________________ Dra. Andrea Pedrosa Harand (Universidade Federal de Pernambuco – UFPE) ______________________________________________ Dr. Felipe Nollet Medeiros de Assis (Universidade Federal da Paraíba – UFPB) ______________________________________________ Dr. Marcelo Guerra (Universidade Federal de Pernambuco – UFPE) Suplentes: ______________________________________________ Dra. Lânia Isis Ferreira Alves (Universidade Federal da Paraíba – UFPB) ______________________________________________ Dra. Sônia Maria Pereira Barreto (Universidade Federal de Pernambuco – UFRPE) iii A minha família, em especial ao meu pai José Geraldo Rodrigues dos Santos que sempre foi o meu maior incentivador e a quem responsabilizo o meu amor pela docência.
    [Show full text]
  • (Largeflower Triteleia): a Technical Conservation Assessment
    Triteleia grandiflora Lindley (largeflower triteleia): A Technical Conservation Assessment © 2003 Ben Legler Prepared for the USDA Forest Service, Rocky Mountain Region, Species Conservation Project January 29, 2007 Juanita A. R. Ladyman, Ph.D. JnJ Associates LLC 6760 S. Kit Carson Cir E. Centennial, CO 80122 Peer Review Administered by Society for Conservation Biology Ladyman, J.A.R. (2007, January 29). Triteleia grandiflora Lindley (largeflower triteleia): a technical conservation assessment. [Online]. USDA Forest Service, Rocky Mountain Region. Available: http://www.fs.fed.us/r2/ projects/scp/assessments/triteleiagrandiflora.pdf [date of access]. ACKNOWLEDGMENTS The time spent and the help given by all the people and institutions mentioned in the References section are gratefully acknowledged. I would also like to thank the Colorado Natural Heritage Program for their generosity in making their files and records available. I also appreciate access to the files and assistance given to me by Andrew Kratz, USDA Forest Service Region 2. The data provided by the Wyoming Natural Diversity Database and by James Cosgrove and Lesley Kennes with the Natural History Collections Section, Royal BC Museum were invaluable in the preparation of the assessment. Documents and information provided by Michael Piep with the Intermountain Herbarium, Leslie Stewart and Cara Gildar of the San Juan National Forest, Jim Ozenberger of the Bridger-Teton National Forest and Peggy Lyon with the Colorado Natural Heritage Program are also gratefully acknowledged. The information provided by Dr. Ronald Hartman and B. Ernie Nelson with the Rocky Mountain Herbarium, Teresa Prendusi with the Region 4 USDA Forest Service, Klara Varga with the Grand Teton National Park, Jennifer Whipple with Yellowstone National Park, Dave Dyer with the University of Montana Herbarium, Caleb Morse of the R.L.
    [Show full text]
  • Technical Report for the Mpumalanga Biodiversity Sector Plan – MBSP 2015
    Technical Report for the Mpumalanga Biodiversity Sector Plan – MBSP 2015 June 2015 Authored by: Mervyn C. Lötter Mpumalanga Tourism &Parks Agency Private bag X1088 Lydenburg, 1120 1 Citation: This document should be cited as: Lötter, M.C. 2015. Technical Report for the Mpumalanga Biodiversity Sector Plan – MBSP. Mpumalanga Tourism & Parks Agency, Mbombela (Nelspruit). ACKNOWLEDGMENTS There are many individuals and organisations that contributed towards the success of the MBSP. In particular we gratefully acknowledge the ArcGIS software grant from the ESRI Conservation Program. In addition the WWF-SA and SANBIs Grasslands Programme played an important role in supporting the development and financing parts of the MBSP. The development of the MBSP spatial priorities took a few years to complete with inputs from many different people and organisations. Some of these include: MTPA scientists, Amanda Driver, Byron Grant, Jeff Manuel, Mathieu Rouget, Jeanne Nel, Stephen Holness, Phil Desmet, Boyd Escott, Charles Hopkins, Tony de Castro, Domitilla Raimondo, Lize Von Staden, Warren McCleland, Duncan McKenzie, Natural Scientific Services (NSS), South African National Biodiversity Institute (SANBI), Strategic Environmental Focus (SESFA),Birdlife SA, Endangered Wildlife Trust, Graham Henning, Michael Samways, John Simaika, Gerhard Diedericks, Warwick Tarboton, Jeremy Dobson, Ian Engelbrecht, Geoff Lockwood, John Burrows, Barbara Turpin, Sharron Berruti, Craig Whittington-Jones, Willem Froneman, Peta Hardy, Ursula Franke, Louise Fourie, Avian Demography
    [Show full text]