The Academy is governed by a distinguished group of eminent scientists who have been appointed Fellows of the California Academy of Sciences in recognition of their notable contributions to one or more of the natural sciences. Nominated by their colleagues and appointed by the Board of Trustees, the Fellows remain members of the Fellowship for life.
Below is a complete list of Academy Fellows.
David Ackerly Alissa J. Arp Robert G. Bergman Plant Ecology & Evolution Marine Biology Chemistry
Bruce Alberts Frank Asaro Brent Berlin Molecular Biology Nuclear Chemistry Anthropology
Angel C. Alcala Francisco J. Ayala Giacomo Bernardi Herpetology Genetics Ichthyology
Michele L. Aldrich Bruce G. Baldwin Vladimir A. Beshkov History of Science Botany Herpetology
Zeresenay Alemseged George E. Ball James L. Bischoff Anthropology & Entomology Marine Geology Paleoanthropology Richard Barber Jeffrey M. Black Frank Almeda Marine Biology Conservation and Ecology of Botany Water Fowl Anthony D. Barnosky Paul Arnaud Jr. Paleontology David C. Blackburn Entomology Herpetology Reginald H. Barrett E. N. Arnold Environmental Science Elizabeth H. Blackburn Herpetology Molecular Biology John Barron Jeanne E. Arnold Micropaleontology James A. Blake Anthropology Invertebrate Zoology Bruce Bartholomew Walter Alvarez Botany Barbara A. Block Geology Evolutionary, Cellular, and Gibor B. Basri Molecular Physiology Steven C. Anderson Astronomy & Astrophysics Systematic, Biogeographic and Carol L. Boggs Ecological Studies in Aaron M. Bauer Evolutionary Ecology Herpetology Herpetology Stephen Bollens Christopher Andrews Steven V. W. Beckwith Biological Oceanography Aquarium Management & Astronomy Parasitology Michel Boudart Shannon N. Bennett Chemistry Fred Andrews Microbiology Entomology Sarane T. Bowen William J. Bennetta Genetics Science & Public Policy
Academy Fellows
Rauri C.K. Bowie Sherwin Carlquist Anne C. Cohen Ornithology Botany Invertebrate Systematics
Justin Brashares Sandra J. Carlson Daniel M. Cohen Ecology & Wildlife Conservation Paleontology Ichthyology
John I. Brauman James T. Carlton Sarah Cohen Chemistry Marine Biodiversity and Marine Genetics Conservation Max C. Brewer James P. Collman Geology Edward J. Carpenter Chemistry Biological Oceanography and Thomas Briggs Marine Microbiology Robert K. Colwell Entomology Zoology James M. Carpenter Winslow R. Briggs Entomology Margaret W. Conkey Botany Anthropology Kent Carpenter Michael E. Brown Marine Biology & Ichthyology Bruce R. Conklin Astronomy Bioinformatics, Genomics & Mark H. Carr Molecular Pharmacology Robert D. Brown, Jr. Marine Ecology Geology Edward Connor John Caskey Ecology Thomas D. Bruns Geology, Neotectonics Mycology T. John Conomos Alan Channing Marine Geology Richard C. Brusca Herpetology Marine Biology and Claudia Corti Conservation Robert L. Christiansen Herpetology Geology Robert W. Buddemeier Lawrence Cory Environmental Studies Christopher Chyba Zoology Astrobiology Stephen H. Bullock Daniel P. Costa Plant Ecology Frank Cipriano Vertebrate Ecology Ecology and Evolutionary Margaret G. Burke Biology Donald Croll Behavioral Ecology & Education Ecology of Marine Predators, David A. Clague Island Ecology & Conservation Gregor M. Cailliet Volcanology Ichthyology Dale P. Cruikshank William A. Clemens Planetary Astronomy Roy L. Caldwell Paleontology Invertebrate Zoology Gretchen C. Daily Lloyd Cluff Conservation Biology Douglas G. Capone Geology Biological Oceanography, Howell Daly Marine Microbiology Kenneth Coale Entomology Oceanography James R. Carey Thomas F. Daniel Entomology William P. Cochlan Botany Biological Oceanography
Academy Fellows
Todd E. Dawson John Dumbacher Sandra Faber Physiological Plant Ecology Ornithology Astronomy
Donald J. De Paolo Thomas O. Duncan Gregory C. Farrington Geosciences Botany Chemistry
Joseph DeRisi Thomas Dunne Daphne G. Fautin Genomics, Molecular Biology, Geomorphology and Hydrology Invertebrate Zoology Parasitology, Virology, Computational Biology William Durham Lewis J. Feldman Human Biology Botany Dennis E. Desjardin Mycology David M. Eakin Marcus W. Feldman Physics Mathematical Biology David Des Marais Geology and Astrobiology Sylvia A. Earle Carl J. Ferraris, Jr. Botany Ichthyology Susan Desmond-Hellmann Oncology, Biotechnology Anne H. Ehrlich Francis P. Filice Conservation Biology Zoology Marian C. Diamond Anatomy Paul R. Ehrlich Peggy L. Fiedler Conservation Biology Botany Janis Dickinson Behavioral Ecology, Emil R. Eisenhardt Alexi Filippenko Conservation Biology Conservation Astronomy
Rodolfo Dirzo Paul Ekman Brian Fisher Ecology, Conservation Biology Comparative Psychology Entomology
Daniel Doak Peter M. Elias Jonathan A. Foley Evolutionary Ecology Medicine Global Sustainability
Robert V. Dowell Thomas S. Elias Susan L. Forsburg Entomology Botany Molecular Biology: Cancer Biology, DNA John T. Doyen David Epel Entomology Cell and Developmental Biology Michael S. Foster Marine Biology James A. Doyle Don C. Erman Botany Fisheries Ecology Laurel R. Fox Ecology William T. Doyle Terry L. Erwin Botany Entomology Andrew Fraknoi Astronomy Frank Drake William N. Eschmeyer Astronomy Ichthyology Laurence G. Frank Mammalogy Robert Drewes James A. Estes Herpetology Marine Ecology Peter W. Fritsch Botany Richard Dugdale William R. Evitt Oceanography Paleontology
Academy Fellows
Robert J. Full Deborah M. Gordon Christopher Harrold Zoology Ecology Marine Biology, Conservation
Jon S. Galehouse George C. Gorman John Harte Geology Evolutionary Biology Environmental Studies
Newell T. Garfield Terrence M. Gosliner James T. Harvey Physical Oceanography Malacology Marine Biology
Jacques A. Gauthier Michael H. Graham David Haussler Herpetology Marine Ecology Computer Science Harry W. Greene Jonathan B. Geller Herpetology Tyrone B. Hayes Marine Ecology and Marine Herpetology Molecular Ecology H. Gary Greene Marine Geology Michael S. Hedrick Wayne M. Getz Environmental Physiology, Entomology David W. Greenfield Herpetology Ichthyology Michael T. Ghiselin Carl E. Heiles Invertebrate Zoology M.R.C. Greenwood Astronomy Medicine Gregory S. Gilbert Colin O. Hermans Microbial Ecology Gary B. Griggs Invertebrate Zoology Geology, Oceanography Rosemary G. Gillespie Donald Heyneman Entomology Charles W. Griswold Tropical Medicine, Entomology Epidemiology, Parasitology, Gonzalo Giribet Invertebrate Zoology Entomology Richard Grosberg Evolution and Invertebrate Carole S. Hickman Alexander N. Glazer Biology Paleontology Molecular and Conservation Biology Karen Grove Tessa M. Hill Geology Earth Sciences Stephen Glickman Claes Gustafsson Psychology Molecular Biology and Stanley Hillman Lynda J. Goff Biotechnology Environmental Physiology of Phycology Amphibians; Herpetology Elizabeth Hadly David M. Golden Mammalogy, Vertebrate Frederick G. Hochberg, Jr. Chemistry Evolution, Climate Change Invertebrate Zoology
Charles Goldman John E. Hafernik, Jr. Tori M. Hoehler Zoology Entomology Astrobiology; Biogeochemistry David Halpern Michael A. Goldman Physical Oceanography John P. Holdren Medical Genetics & Energy & Natural Resources Chromosomal Evolution Alexander Harcourt Primatology Gustavo Hormiga Edgardo D. Gomez Entomology Marine Conservation Susan P. Harrison Conservation Biology
Academy Fellows
Kim M. Howell Jean H. Langenheim James V. Lawry, Jr. Herpetology Botany Anatomy
Sarah B. Hrdy Patrick V. Kirch Richard E. Leakey Anthropology Anthropology Anthropology
James C. Ingle David H. Kistner Burney J. LeBoeuf Paleontology Entomology Biology
B. Lynn Ingram Richard G. Klein Gretchen LeBuhn Geology Paleoanthropology Evolutionary Ecology
Michael E. Irwin Paul L. Koch Welton Lee Entomology Geobiology/Paleobiology Invertebrate Zoology
Tomio Iwamoto Patrick Kociolek Janet L. Leonard Ichthyology Diatom Studies Neurobiology and Behavior
Nina G. Jablonski Mimi A. Koehl William A. Lester, Jr. Anthropology Invertebrate Zoology Physical Chemistry
Rodney M. Jackson Walter D. Koenig Alan E. Leviton Conservation Biology Avian Ecology Herpetology
Matthew J. James Claire Kremen Matthew R. Lewin Paleontology and History of Conservation Biology & Emergency Medicine Science Entomology Lowell N. Lewis David L. Jameson Leonard V. Kuhi Botany Zoology Astronomy Paul Licht William A. Jensen Eileen A. Lacey Zoology Botany Behavioral Ecology, Donald C. Johanson Mammalogy William Z. Lidicker, Jr. Anthropology Mammalogy and Ecology Watson M. Laetsch Michael Josselyn Botany Kent G. Lightfoot Ecology and Phycology Archaeology & Anthropology Michael D. Lagios David H. Kavanaugh Ichthyology David R. Lindberg Entomology Malacology Robert S. Lane James C. Kelley Entomology Jere H. Lipps Oceanography Paleontology Leo F. Laporte Donald Kennedy Geology Werner J. Loher Biological Sciences Entomology Ralph J. Larson William H. Kimbel Ichthyology Sharon Long Anthropology Plant/Microbial Symbiosis
John D. Lattin Wim Kimmerer Entomology Jerold Lowenstein Estuarine Ecology Medicine
Academy Fellows
Margaret D. Lowman Gopinathan Menon Michael W. Nachman Ecology Biochemistry Mammalogy & Evolutionary Genetics Bruce E. Lyon Fiorenza Micheli Behavioral Ecology Marine Ecology and Donald Nevins Conservation Botany James M. Lyons Plant Physiology Kathy Ann Miller Andrew T. Newberry Marine Algal Systematics Invertebrate Biology James P. Mackey Herpetology Katharine Milton Frederic H. Nichols Anthropology Invertebrate Zoology David Maddison Entomology, Informatics, Brent D. Mishler Thomas M. Niesen Computer Programming & Botany Invertebrate Biology Software Development Richard L. Moe Daniel H. Norris Rocco Mancinelli Phycology Botany Astrobiology John P. Monteverdi Peter Norvig York T. Mandra Meteorology Artificial Intelligence & Paleontology Computer Science Richard J. Mooi Marc S. Mangel Invertebrate Zoology Amos M. Nur Behavior and Ecology Geology Harold A. Mooney Geoffrey Marcy Environmental Science Richard E. Packard Astrophysics Physics Eldridge Moores Peter Marler Geology Kevin Padian Ethology Paleontology Michael J. Moratto Charles Marshall Anthropology Stephen R. Palumbi Paleontology Molecular Evolution & Craig Moritz Conservation Biology John E. McCosker Evolutionary & Conservation Aquatic Biology Genetics Theodore Papenfuss Herpetology Alistair W. McCrone David D. Morrison Geology Planetary Astronomy Lynne R. Parenti Ichthyology Jimmy A. McGuire Carol W. Mosher Herpetology, Vertebrate Chemistry Ingrid Parker Biology, Evolutionary Biology Ecology Peter B. Moyle Henry M. McHenry Ichthyology Sue Taylor Parker Biological Anthropology Anthropology Richard A. Muller Monica Medina Astrophysics Thomas Parker Evolutionary Genomics & Botany Systems Biology Michael A. Murphy Paleontology & Geology Dennis R. Parnell Botany
Academy Fellows
Robert Patterson Donald C. Potts Peter J. Richerson Botany Invertebrate Zoology Cultural Evolution
James L. Patton Jerry A. Powell Rollin C. Richmond Mammalogy Entomology Evolutionary Genetics
Saul Perlmutter Thomas M. Powell John W. Rick Astrophysics Oceanography, Limnology, Archaeology & Paleoecology Aquatic Biology John S. Pearse Neil Risch Invertebrate Zoology Mary E. Power Human Evolutionary Genetics & Ecology of Riverine Systems Epidemiology Vicki B. Pearse Invertebrate Zoology Joel R. Primack Bruce Robison Planetary Sciences Deep-Sea Biology Yvonne Pendleton Astrophysics Wojciech J. Pulawski Luiz A. Rocha Entomology Ichthyology David Perlman Journalism Alexander H. Purcell Peter U. Rodda Entomology Geology Raymond Pestrong Geology Peter R. Raimondi George K. Roderick Marine Ecology Population Genetics and Lewis Petrinovich Evolutionary Biology Ornithology Katherine Ralls Conservation Biology, Peter Rodman Theodore W. Pietsch Mammalogy Anthropology Ichthyology Janet A. Randall Barbara Rogoff Carl B. Pilcher Mammalogy Developmental Psychology Astrobiology, Astronomy, Planetary Science Galen B. Rathbun Peter Roopnarine Mammalogy Invertebrate Paleontology Edwin P. Pister Conservation Biology Peter H. Raven Terry L. Root Botany Climate Change Sensitivity of Karl S. Pister Vertebrates Civil Engineering and Education David C.F. Rentz Entomology Edward S. Ross Norman I. Platnick Entomology Entomology Vincent H. Resh Aquatic Biology Barry Roth Grant Pogson Malacology & Paleontology Molecular Population Genetics, Don Reynolds Ecological Genetics Botany Lynn J. Rothschild Astrobiology & Evolutionary George O. Poinar, Jr. Wynetka A. Reynolds Protistology Entomology Zoology Joan E. Roughgarden Katherine Pollard Mark Richards Herpetology Evolution & Computational Geophysics Biology
Academy Fellows
Eric Routman Norman J. Silberling Calvin H. Stevens Herpetology Geology Geology
Rodolfo Ruibal Paul C. Silva Jonathon H. Stillman Herpetology Botany Invertebrate Physiology & Molecular Biology Bruce Runnegar Ellen L. Simms Geology & Paleontology Botany Scott W. Stine Geography Robert Sapolsky Barry R. Sinervo Neuroendocrinology Ecology David Stoddart Geography Jose Sarukhan Leonard S. Sklar Botany Geology Sharon Y. Strauss Evolutionary Ecology Evert I. Schlinger Doris H. Sloan Entomology Geology John L. Strother Botany Russell L. Schweickart Alan R. Smith Astrobiology Botany Raymond Sullivan Geology Judith Scotchmoor Edmund H. Smith Science Education & Outreach Invertebrate Zoology James R. Sweeney Botany Eugenie C. Scott Edward L. Smith Anthropology Entomology Frank Talamantes Endocrinology Erwin Seibel Thomas B. Smith Oceanography Ecology Frank H. Talbot Marine Biology H. Bradley Shaffer George Somero Herpetology Evolutionary Biochemistry Benito C. Tan Bryology Arthur Shapiro Michael E. Soule Zoology Evolutionary and Conservation Carol Tang Biology Paleontology Howard S. Shellhammer Mammalogy Wayne P. Sousa Kimberly Tanner Marine Ecology Science Education James R. Shevock Botany Howard Spero Jill Cornell Tarter Paleoceanography Astronomy Seth Shostak Astronomy Gregory Spicer Catherine A. Tauber Invertebrate Zoology Insect Ecology, Behavior and Neil H. Shubin Physiology Evolutionary Biology Hyron Spinrad Astronomy Maurice J. Tauber Petra Sierwald Insect Ecology, Behavior and Entomology Maureen L. Stanton Physiology Population Biology Mary Silver John W. Taylor Biological Oceanography Mycology
Academy Fellows
Leighton Taylor Marvalee H. Wake R. Sanders Williams Ichthyology Zoology Medicine, Cell Biology, Genetics
Richard R. Tenaza Robert C. Walter Stanley C. Williams Mammalogy Geology Entomology
John Thompson Peter D. Ward Susan L. Williams Evolutionary Ecology Geology Marine Ecology
Robert F. Thorne Philip S. Ward Terrie Williams Botany Ant Systematics Comparative Physiology
Robin W. Thorp Marius S. Wasbauer Arthur P. Wolf Entomology Entomology Anthropology
Ignacio Tinoco Dorothy Washburn James C. Zachos Chemistry Anthropology Paleoceanography, Paleoclimatology, Marine Owen B. Toon Kenneth E. Watt Stratigraphy Atmospheric Science Zoology Frank G. Zalom Charles H. Townes Ward B. Watt Entomology Physics Evolutionary Biology Richard N. Zare Jonathan D. Trent Robert Wayne Chemistry Astrobiology, Marine Biology, Molecular Systematics and Microbiology Ermi Zhao Harold F. Weaver Herpetology Brian E. Tucker Astronomy Seismology, Seismic Safety, Adrienne L. Zihlman Geology Christian M. Wemmer Anthropology Conservation Biology James W. Valentine Irving Zucker Paleontology Cherie L.R. Wetzel Biology Botany Blaire Van Valkenburgh Vertebrate Paleontology Lisa E. White Geology & Paleontology Geerat J. Vermeij Evolutionary Biology Timothy D. White Anthropology Peter M. Vitousek Ecology Frances Wilkerson Biological Oceanography Vance T. Vredenburg Herpetology Mark Wilkinson Herpetology Peter C. Wainwright Evolution, Ecology Kippling W. Will Entomology David B. Wake Herpetology Gary C. Williams Invertebrate Zoology