The Academy is governed by a distinguished group of eminent scientists who have been appointed Fellows of the California Academy of Sciences in recognition of their notable contributions to one or more of the natural sciences. Nominated by their colleagues and appointed by the Board of Trustees, the Fellows remain members of the Fellowship for life.
Below is a complete list of Academy Fellows.
David Ackerly Francisco J. Ayala Giacomo Bernardi Ecology and Evolutionary Biology Genetics Ichthyology
Bruce M. Alberts Bruce G. Baldwin Vladimir A. Beshkov Molecular and Conservation Biology Botany Herpetology
Angel C. Alcala George E. Ball Philip E. Bickler Marine Biology and Conservation, Entomology Comparative Physiology, Herpetology Neuroscience and Anesthesiology Richard T. Barber Michele L. Aldrich Marine Biology James L. Bischoff History of Science Marine Geology Anthony D. Barnosky Zeresenay Alemseged Paleontology Jeffrey M. Black Anthropology and Paleoanthropology Conservation and Ecology of Water Reginald H. Barrett Fowl Frank Almeda Applied Biology, Agricultural and Botany Environmental Science David C. Blackburn Herpetology Walter Alvarez John Barron Geology Micropaleontology Elizabeth H. Blackburn Molecular Biology Steven C. Anderson Bruce Bartholomew Systematic, Biogeographic and Botany James A. Blake Ecological Studies in Herpetology Invertebrate Zoology Gibor B. Basri Christopher Andrews Astronomy and Astrophysics Barbara A. Block Aquarium Management and Evolutionary, Cellular and Molecular Parasitology Aaron M. Bauer Physiology Herpetology Fred Andrews Carol L. Boggs Entomology Steven V. Beckwith Evolutionary Ecology Astronomy Paul H. Arnaud Stephen Bollens Entomology Shannon N. Bennett Biological Oceanography Microbiology E. N. Arnold Sarane T. Bowen Herpetology William J. Bennetta Genetics Science and Public Policy Jeanne Arnold Rauri C.K. Bowie Anthropology Robert G. Bergman Ornithology Chemistry Alissa J. Arp Justin Brashares Marine Biology Brent Berlin Ecology and Wildlife Conservation Anthropology
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Academy Fellows
John I. Brauman James T. Carlton Sarah Cohen Chemistry Marine Biodiversity and Conservation Marine Genetics
Thomas S. Briggs Edward J. Carpenter James P. Collman Entomology Biological Oceanography and Marine Chemistry Microbiology Winslow R. Briggs Robert K. Colwell Botany James M. Carpenter Zoology Entomology Michael E. Brown Margaret W. Conkey Astronomy Kent E. Carpenter Anthropology Marine Biology and Ichthyology Robert D. Brown, Jr. Bruce Conklin Geology Mark H. Carr Bioinformatics, Genomics and Marine Ecology Molecular Pharmacology Thomas D. Bruns Mycology John Caskey Edward F. Connor Geology, Neotectonics Ecology Richard C. Brusca Marine Biology and Conservation Alan Channing T. J. Conomos Herpetology Marine Geology Robert W. Buddemeier Environmental Studies Robert L. Christiansen Claudia Corti Geology Herpetology Stephen H. Bullock Plant Ecology Christopher F. Chyba Lawrence Cory Astrobiology Zoology Margaret G. Burke Behavioral Ecology and Education Frank Cipriano Daniel P. Costa Ecology and Evolutionary Biology Vertebrate Ecology Nathalie A. Cabrol Astrobiology David A. Clague Daniel E. Crocker Volcanology Ecology and Physiology Gregor M. Cailliet Ichthyology William A. Clemens Donald Croll Paleontology Ecology of Marine Predators, Island Roy L. Caldwell Ecology and Conservation Invertebrate Zoology Lloyd S. Cluff Geology Dale P. Cruikshank Douglas G. Capone Planetary Astronomy Biological Oceanography, Marine Kenneth Coale Microbiology Oceanography Gretchen C. Daily Conservation Biology James R. Carey William P. Cochlan Entomology Biological Oceanography Howell V. Daly Entomology Sherwin Carlquist Anne C. Cohen Botany Invertebrate Systematics Marymeghan Daly Animal Systematics and Ecology, Sandra J. Carlson Daniel M. Cohen Cnidaria Paleontology Ichthyology
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Academy Fellows
Thomas F. Daniel Frank Drake David Epel Botany Astronomy Cell Biology and Developmental Biology Bruno David Robert C. Drewes Paleontology, Marine Biology, Herpetology Don C. Erman Evolution Fisheries Ecology Richard Dugdale Todd E. Dawson Oceanography Terry L. Erwin Physiological Plant Ecology Entomology John P. Dumbacher Donald J. De Paolo Ornithology William N. Eschmeyer Geosciences Ichthyology Thomas O. Duncan Joseph DeRisi Botany James A. Estes Molecular Biology, Genomics, Marine Ecology Parasitology, Virology, Thomas Dunne Computational Biology Geomorphology and Hydrology Sandra Faber Astronomy David Des Marais William H. Durham Geology and Astrobiology Human Biology Gregory C. Farrington Chemistry Dennis E. Desjardin David M. Eakin Mycology Physics Daphne G. Fautin Invertebrate Zoology Susan P. Desmond-Hellmann Sylvia A. Earle Oncology and Biotechnology Botany Lewis J. Feldman Botany Marian C. Diamond Anne H. Ehrlich Anatomy Conservation Biology Marcus W. Feldman Mathematical Biology Janis Dickinson Paul R. Ehrlich Behavioral Ecology, Conservation Conservation Biology Carl J. Ferraris Biology Ichthyology Emil R. Eisenhardt Rodolfo Dirzo Conservation Peggy L. Fiedler Ecology and Conservation Biology Botany Paul Ekman Daniel Doak Comparative Psychology Christopher B. Field Evolutionary Ecology Global Ecology Charles Elachi Robert V. Dowell Earth, Ocean and Atmosphere Alexei Filippenko Entomology Science from Space Astronomy
John T. Doyen Peter M. Elias Paul V. Fine Entomology Medicine Ecology and Evolutionary Biology
James A. Doyle Thomas S. Elias Brian Fisher Botany Botany Entomology
William T. Doyle Jonathan A. Foley Botany Global Sustainability
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Academy Fellows
Susan L. Forsburg Alexander N. Glazer Karen Grove Molecular Biology, Cancer Biology Molecular and Conservation Biology Ecology and DNA Stephen E. Glickman Claes Gustafsson Michael S. Foster Psychology Molecular Biology and Biotechnology Marine Biology Lynda J. Goff Elizabeth Hadly Laurel R. Fox Phycology Mammalogy, Vertebrate Evolution, Ecology Climate Change David M. Golden Andrew Fraknoi Chemistry John E. Hafernik Astronomy Entomology Charles R. Goldman Laurence G. Frank Zoology David Halpern Mammalogy Physical Oceanography Michael A. Goldman Peter W. Fritsch Medical Genetics and Chromosomal Alexander Harcourt Botany Evolution Primatology
Robert J. Full Edgardo D. Gomez Susan P. Harrison Zoology Marine Biology, Conservation Conservation Biology
Jon S. Galehouse Deborah M. Gordon Christopher Harrold Geology Ecology Marine Biology, Conservation
Newell T. Garfield George C. Gorman John Harte Physical Oceanography Evolutionary Biology Environmental Studies
Jacques A. Gauthier Terrence M. Gosliner James T. Harvey Herpetology Malacology Marine Biology
Jonathan B. Geller Michael H. Graham Christine A. Hastorf Marine Ecology and Marine Marine Ecology Anthropology Molecular Ecology Harry W. Greene David Haussler Wayne M. Getz Herpetology Computer Science Entomology H. Gary Greene Tyrone B. Hayes Michael T. Ghiselin Marine Geology Herpetology Invertebrate Zoology David W. Greenfield Michael S. Hedrick Gregory S. Gilbert Ichthyology Environmental Physiology and Microbial Ecology Herpetology Gary B. Griggs Rosemary G. Gillespie Geology and Oceanology Carl E. Heiles Entomology Astronomy Charles E. Griswold Gonzalo Giribet Entomology Colin O. Hermans Entomology Invertebrate Zoology Richard K. Grosberg Evolution and Invertebrate Biology
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Academy Fellows
Donald Heyneman Nina G. Jablonski Mimi A. Koehl Invertebrate Zoology, Tropical Anthropology Invertebrate Zoology Medicine, Epidemiology and Parasitology Rodney M. Jackson Walter D. Koenig Conservation Biology Avian Ecology Carole S. Hickman Paleontology Matthew J. James Claire Kremen Paleontology and History of Science Conservation Biology and Tessa M. Hill Entomology Earth Sciences David L. Jameson Zoology Leonard V. Kuhi Stanley Hillman Astronomy Environmental Physiology of William A. Jensen Amphibians and Herpetology Botany Eileen A. Lacey Behavioral Ecology and Mammalogy Frederick G. Hochberg, Jr. Donald C. Johanson Invertebrate Zoology Anthropology Watson M. Laetsch Botany Tori M. Hoehler Michael N. Josselyn Astrobiology and Biochemistry Ecology and Phycology Michael D. Lagios Ichthyology John P. Holdren David H. Kavanaugh Energy and Natural Resources Entomology Robert S. Lane Entomology Karen Holl James C. Kelley Restoration Ecology Oceanography Leo F. Laporte Geology Gustavo Hormiga Donald Kennedy Entomology Biological Sciences Ralph J. Larson Ichthyology Kim M. Howell William H. Kimbel Herpetology Anthropology John D. Lattin Entomology Sarah B. Hrdy Wim Kimmerer Anthropology Estuarine Ecology James V. Lawry, Jr. Anatomy James C. Ingle Patrick V. Kirch Paleontology Anthropology Burney J. Le Boeuf Biology B. L. Ingram David H. Kistner Geology Entomology Richard E. Leakey Anthropology Michael E. Irwin Richard G. Klein Entomology Paleoanthropology Walter S. Leal Chemical Ecology and Entomology Lynne A. Isbell Paul L. L. Koch Evolutionary Anthropology Paleobiology and Geobiology Gretchen LeBuhn Evolutionary Ecology Tomio Iwamoto Patrick Kociolek Ichthyology Diatom Studies Welton Lee Invertebrate Zoology
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Academy Fellows
Margaret S. Leinen James P. Mackey Katharine Milton Oceanography, Climate Engineering Herpetology Anthropology
Janet L. Leonard David Maddison Brent D. Mishler Neurobiology and Behavior Entomology, Informatics, Computer Botany Programming and Software William A. Lester Development Richard L. Moe Physical Chemistry Phycology Rocco Mancinelli Alan E. Leviton Astrobiology John P. Monteverdi Herpetology Meteorology York T. Mandra Matthew R. Lewin Paleontology Richard J. Mooi Emergency Medicine Invertebrate Zoology Marc S. Mangel Lowell N. Lewis Behavior and Ecology Harold A. Mooney Botany Environmental Science Geoffrey Marcy Paul Licht Astrophysics Eldridge Moores Herpetology Geology Charles R. Marshall William Z. Lidicker, Jr. Paleontology Michael J. Moratto Mammalogy Anthropology Pamela A. Matson Kent G. Lightfoot Environmental Studies Craig C. Moritz Archaeology and Anthropology Evolutionary and Conservation John E. McCosker Genetics David R. Lindberg Aquatic Biology Malacology David D. Morrison Alistair W. McCrone Planetary Astronomy Jere H. Lipps Geology Paleontology Carol W. Mosher Jimmy A. McGuire Chemistry Werner J. Loher Herpetology, Vertebrate Biology and Entomology Evolutionary Biology Peter B. Moyle Ichthyology Sharon R. Long Henry M. McHenry Plant and Microbial Symbiosis Biological Anthropology Richard A. Muller Astrophysics Jerold Lowenstein Monica Medina Medicine Evolutionary Genomics and Systems Michael A. Murphy Biology Paleontology and Geology Margaret D. Lowman Ecology Gopinathan Menon Michael W. Nachman Biochemistry Mammalogy and Evolutionary Bruce E. Lyon Genetics Behavioral Ecology Fiorenza Micheli Marine Ecology and Conservation Donald J. Nevins James M. Lyons Botany Plant Physiology Kathy Ann Miller Marine Algal Systematics
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Academy Fellows
Andrew T. Newberry Dennis R. Parnell George O. Poinar, Jr. Invertebrate Biology Botany Entomology
Frederic H. Nichols Robert Patterson Katherine Pollard Invertebrate Zoology Botany Evolution and Computational Biology
Thomas M. Niesen James L. Patton Donald C. Potts Invertebrate Biology Mammalogy Invertebrate Zoology
Daniel H. Norris John S. Pearse Jerry A. Powell Botany Invertebrate Zoology Entomology
Peter Norvig Vicki B. Pearse Thomas M. Powell Artificial Intelligence and Computer Invertebrate Zoology Oceanography, Limnology and Science Aquatic Biology Yvonne Pendleton Amos M. Nur Astrophysics Mary E. Power Geology Ecology of Riverine Systems David Perlman Patrick M. O’Grady Journalism Joel R. Primack Evolutionary Biology Planetary Sciences Saul Perlmutter Richard E. Packard Astrophysics Wojciech J. Pulawski Physics Entomology Raymond Pestrong Kevin Padian Geology Alexander H. Purcell Paleontology Entomology Lewis Petrinovich Robert E. Page, Jr. Ornithology Peter R. Raimondi Entomology Marine Ecology Theodore W. Pietsch Stephen R. Palumbi Ichthyology Katherine Ralls Molecular Evolution and Conservation Biology and Conservation Biology Carl B. Pilcher Mammalogy Astrobiology, Astronomy, Planetary Theodore Papenfuss Science Janet A. Randall Herpetology Mammalogy Edwin P. Pister Lynne R. Parenti Conservation Biology Galen B. Rathbun Ichthyology Mammalogy Karl S. Pister Ingrid Parker Civil Engineering, Education Peter H. Raven Ecology Botany Norman I. Platnick Sue Taylor Parker Entomology David C. F. Rentz Anthropology Entomology Grant Pogson Thomas Parker Molecular Population Genetics and Vincent H. Resh Botany Ecological Genetics Aquatic Biology
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Academy Fellows
Don R. Reynolds Lynn J. Rothschild Seth Shostak Botany Astrobiology and Evolutionary Astronomy Protistology Wynetka A. Reynolds Neil H. Shubin Biochemistry Joan E. Roughgarden Evolutionary Biology Herpetology Mark Richards Petra Sierwald Geophysics Eric J. Routman Entomology Herpetology Peter J. Richerson Mary W. Silver Cultural Evolution Rodolfo Ruibal Biological Oceanography Herpetology Rollin C. Richmond W. Brian Simison Evolutionary Genetics Bruce Runnegar Genomics Geology and Paleontology John W. Rick Ellen L. Simms Archaeology and Paleoecology Robert Sapolsky Botany Neuroendocrinology Neil J. Risch Barry R. Sinervo Human Evolutionary Genetics and Jose Sarukhan Ecology Epidemiology Botany Leonard S. Sklar Bruce H. Robison Russell L. Schweickart Geology Deep-Sea Biology Astrobiology Doris H. Sloan Luiz A. Rocha Judith G. Scotchmoor Geology Ichthyology Science Education and Outreach Alan R. Smith Peter U. Rodda Eugenie C. Scott Botany Geology Anthropology Edmund H. Smith George K. Roderick Ravinder N. M. Sehgal Invertebrate Zoology Population Genetics and Evolutionary Avian Parasitology, Ecology of Biology Infectious Disease Edward L. Smith Entomology Peter Rodman Erwin Seibel Anthropology Oceanography Thomas B. Smith Ecology Barbara Rogoff H. Bradley Shaffer Developmental Psychology Herpetology George Somero Evolutionary Biochemistry Peter Roopnarine Arthur M. Shapiro Invertebrate Paleontology Zoology Victoria L. Sork Evolutionary Biology Terry L. Root Howard S. Shellhammer Climate Change Sensitivity of Mammalogy Michael E. Soule Vertebrates Evolutionary and Conservation James R. Shevock Biology Barry Roth Botany Malacology and Paleontology Wayne P. Sousa Marine Ecology
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Academy Fellows
Chelsea Specht Jill C. Tarter Vance T. Vredenburg Plant Genetics and Genomics Astronomy Herpetology
Howard J. Spero Catherine A. Tauber Peter C. Wainwright Paleoceanography Insect Ecology, Behavior and Evolution and Ecology Physiology Gregory S. Spicer David B. Wake Invertebrate Zoology Leighton R. Taylor Herpetology Ichthyology Hyron Spinrad Marvalee H. Wake Astronomy John W. Taylor Zoology Mycology Maureen L. Stanton Robert C. Walter Population Biology Richard R. Tenaza Geology Mammalogy Calvin H. Stevens Peter D. Ward Geology Robbin W. Thorp Geology Entomology Jonathon H. Stillman Philip S. Ward Invertebrate Physiology and Ignacio Tinoco Ant Systematics Molecular Biology Chemistry Marius S. Wasbauer Scott W. Stine Regina Tomlinson Entomology Geography Invertebrate Zoology Dorothy Washburn Sharon Y. Strauss O. Brian Toon Anthropology Evolutionary Ecology Atmospheric Science Kenneth E. Watt John L. Strother Jonathan D. Trent Zoology Botany Astrobiology, Marine Biology and Microbiology Ward B. Watt Raymond Sullivan Evolutionary Biology Geology Brian E. Tucker Seismology, Seismic Safety, Geology Robert Wayne James R. Sweeney Molecular Systematics Botany Ronald D. Vale Cell Biology, Public Science Education Harold F. Weaver Frank Talamantes Astronomy Endocrinology James W. Valentine Paleontology Christian M. Wemmer Frank H. Talbot Conservation Biology Marine Biology Blaire Van Valkenburgh Vertebrate Paleontology Cherie L. Wetzel Benito C. Tan Botany Bryology Geerat J. Vermeij Evolutionary Biology Lisa E. White Carol Tang Geology and Paleontology Paleontology Peter M. Vitousek Ecology Timothy D. White Kimberly Tanner Anthropology Science Education
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Academy Fellows
Frances Wilkerson Biological Oceanography
Mark Wilkinson Herpetology
Kipling W. Will Entomology
Gary C. Williams Invertebrate Zoology
R. Sanders Williams Medicine, Cell Biology and Genetics
Stanley C. Williams Entomology
Susan L. Williams Marine Ecology
Terrie Williams Comparative Physiology
Arthur P. Wolf Anthropology
James C. Zachos Paleoclimatology, Marine Stratigraphy and Paloeoceanography
Frank G. Zalom Entomology
Richard N. Zare Chemistry
Erika S. Zavaleta Community and Ecosystem Ecology, Conservation Biology
Ermi Zhao Herpetology
Adrienne L. Zihlman Anthropology
Irving Zucker Biology
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