Fellows of the California Academy of Sciences in Recognition of Their Notable Contributions to One Or More of the Natural Sciences
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The Academy is governed by a distinguished group of eminent scientists who have been appointed Fellows of the California Academy of Sciences in recognition of their notable contributions to one or more of the natural sciences. Nominated by their colleagues and appointed by the Board of Trustees, the Fellows remain members of the Fellowship for life. Below is a complete list of Academy Fellows. David Ackerly Francisco J. Ayala Giacomo Bernardi Ecology and Evolutionary Biology Genetics Ichthyology Bruce M. Alberts Bruce G. Baldwin Vladimir A. Beshkov Molecular and Conservation Biology Botany Herpetology Angel C. Alcala George E. Ball Philip E. Bickler Marine Biology and Conservation, Entomology Comparative Physiology, Herpetology Neuroscience and Anesthesiology Richard T. Barber Michele L. Aldrich Marine Biology James L. Bischoff History of Science Marine Geology Anthony D. Barnosky Zeresenay Alemseged Paleontology Jeffrey M. Black Anthropology and Paleoanthropology Conservation and Ecology of Water Reginald H. Barrett Fowl Frank Almeda Applied Biology, Agricultural and Botany Environmental Science David C. Blackburn Herpetology Walter Alvarez John Barron Geology Micropaleontology Elizabeth H. Blackburn Molecular Biology Steven C. Anderson Bruce Bartholomew Systematic, Biogeographic and Botany James A. Blake Ecological Studies in Herpetology Invertebrate Zoology Gibor B. Basri Christopher Andrews Astronomy and Astrophysics Barbara A. Block Aquarium Management and Evolutionary, Cellular and Molecular Parasitology Aaron M. Bauer Physiology Herpetology Fred Andrews Carol L. Boggs Entomology Steven V. Beckwith Evolutionary Ecology Astronomy Paul H. Arnaud Stephen Bollens Entomology Shannon N. Bennett Biological Oceanography Microbiology E. N. Arnold Sarane T. Bowen Herpetology William J. Bennetta Genetics Science and Public Policy Jeanne Arnold Rauri C.K. Bowie Anthropology Robert G. Bergman Ornithology Chemistry Alissa J. Arp Justin Brashares Marine Biology Brent Berlin Ecology and Wildlife Conservation Anthropology Page 1 of 10 Academy Fellows John I. Brauman James T. Carlton Sarah Cohen Chemistry Marine Biodiversity and Conservation Marine Genetics Thomas S. Briggs Edward J. Carpenter James P. Collman Entomology Biological Oceanography and Marine Chemistry Microbiology Winslow R. Briggs Robert K. Colwell Botany James M. Carpenter Zoology Entomology Michael E. Brown Margaret W. Conkey Astronomy Kent E. Carpenter Anthropology Marine Biology and Ichthyology Robert D. Brown, Jr. Bruce Conklin Geology Mark H. Carr Bioinformatics, Genomics and Marine Ecology Molecular Pharmacology Thomas D. Bruns Mycology John Caskey Edward F. Connor Geology, Neotectonics Ecology Richard C. Brusca Marine Biology and Conservation Alan Channing T. J. Conomos Herpetology Marine Geology Robert W. Buddemeier Environmental Studies Robert L. Christiansen Claudia Corti Geology Herpetology Stephen H. Bullock Plant Ecology Christopher F. Chyba Lawrence Cory Astrobiology Zoology Margaret G. Burke Behavioral Ecology and Education Frank Cipriano Daniel P. Costa Ecology and Evolutionary Biology Vertebrate Ecology Nathalie A. Cabrol Astrobiology David A. Clague Daniel E. Crocker Volcanology Ecology and Physiology Gregor M. Cailliet Ichthyology William A. Clemens Donald Croll Paleontology Ecology of Marine Predators, Island Roy L. Caldwell Ecology and Conservation Invertebrate Zoology Lloyd S. Cluff Geology Dale P. Cruikshank Douglas G. Capone Planetary Astronomy Biological Oceanography, Marine Kenneth Coale Microbiology Oceanography Gretchen C. Daily Conservation Biology James R. Carey William P. Cochlan Entomology Biological Oceanography Howell V. Daly Entomology Sherwin Carlquist Anne C. Cohen Botany Invertebrate Systematics Marymeghan Daly Animal Systematics and Ecology, Sandra J. Carlson Daniel M. Cohen Cnidaria Paleontology Ichthyology Page 2 of 10 Academy Fellows Thomas F. Daniel Frank Drake David Epel Botany Astronomy Cell Biology and Developmental Biology Bruno David Robert C. Drewes Paleontology, Marine Biology, Herpetology Don C. Erman Evolution Fisheries Ecology Richard Dugdale Todd E. Dawson Oceanography Terry L. Erwin Physiological Plant Ecology Entomology John P. Dumbacher Donald J. De Paolo Ornithology William N. Eschmeyer Geosciences Ichthyology Thomas O. Duncan Joseph DeRisi Botany James A. Estes Molecular Biology, Genomics, Marine Ecology Parasitology, Virology, Thomas Dunne Computational Biology Geomorphology and Hydrology Sandra Faber Astronomy David Des Marais William H. Durham Geology and Astrobiology Human Biology Gregory C. Farrington Chemistry Dennis E. Desjardin David M. Eakin Mycology Physics Daphne G. Fautin Invertebrate Zoology Susan P. Desmond-Hellmann Sylvia A. Earle Oncology and Biotechnology Botany Lewis J. Feldman Botany Marian C. Diamond Anne H. Ehrlich Anatomy Conservation Biology Marcus W. Feldman Mathematical Biology Janis Dickinson Paul R. Ehrlich Behavioral Ecology, Conservation Conservation Biology Carl J. Ferraris Biology Ichthyology Emil R. Eisenhardt Rodolfo Dirzo Conservation Peggy L. Fiedler Ecology and Conservation Biology Botany Paul Ekman Daniel Doak Comparative Psychology Christopher B. Field Evolutionary Ecology Global Ecology Charles Elachi Robert V. Dowell Earth, Ocean and Atmosphere Alexei Filippenko Entomology Science from Space Astronomy John T. Doyen Peter M. Elias Paul V. Fine Entomology Medicine Ecology and Evolutionary Biology James A. Doyle Thomas S. Elias Brian Fisher Botany Botany Entomology William T. Doyle Jonathan A. Foley Botany Global Sustainability Page 3 of 10 Academy Fellows Susan L. Forsburg Alexander N. Glazer Karen Grove Molecular Biology, Cancer Biology Molecular and Conservation Biology Ecology and DNA Stephen E. Glickman Claes Gustafsson Michael S. Foster Psychology Molecular Biology and Biotechnology Marine Biology Lynda J. Goff Elizabeth Hadly Laurel R. Fox Phycology Mammalogy, Vertebrate Evolution, Ecology Climate Change David M. Golden Andrew Fraknoi Chemistry John E. Hafernik Astronomy Entomology Charles R. Goldman Laurence G. Frank Zoology David Halpern Mammalogy Physical Oceanography Michael A. Goldman Peter W. Fritsch Medical Genetics and Chromosomal Alexander Harcourt Botany Evolution Primatology Robert J. Full Edgardo D. Gomez Susan P. Harrison Zoology Marine Biology, Conservation Conservation Biology Jon S. Galehouse Deborah M. Gordon Christopher Harrold Geology Ecology Marine Biology, Conservation Newell T. Garfield George C. Gorman John Harte Physical Oceanography Evolutionary Biology Environmental Studies Jacques A. Gauthier Terrence M. Gosliner James T. Harvey Herpetology Malacology Marine Biology Jonathan B. Geller Michael H. Graham Christine A. Hastorf Marine Ecology and Marine Marine Ecology Anthropology Molecular Ecology Harry W. Greene David Haussler Wayne M. Getz Herpetology Computer Science Entomology H. Gary Greene Tyrone B. Hayes Michael T. Ghiselin Marine Geology Herpetology Invertebrate Zoology David W. Greenfield Michael S. Hedrick Gregory S. Gilbert Ichthyology Environmental Physiology and Microbial Ecology Herpetology Gary B. Griggs Rosemary G. Gillespie Geology and Oceanology Carl E. Heiles Entomology Astronomy Charles E. Griswold Gonzalo Giribet Entomology Colin O. Hermans Entomology Invertebrate Zoology Richard K. Grosberg Evolution and Invertebrate Biology Page 4 of 10 Academy Fellows Donald Heyneman Nina G. Jablonski Mimi A. Koehl Invertebrate Zoology, Tropical Anthropology Invertebrate Zoology Medicine, Epidemiology and Parasitology Rodney M. Jackson Walter D. Koenig Conservation Biology Avian Ecology Carole S. Hickman Paleontology Matthew J. James Claire Kremen Paleontology and History of Science Conservation Biology and Tessa M. Hill Entomology Earth Sciences David L. Jameson Zoology Leonard V. Kuhi Stanley Hillman Astronomy Environmental Physiology of William A. Jensen Amphibians and Herpetology Botany Eileen A. Lacey Behavioral Ecology and Mammalogy Frederick G. Hochberg, Jr. Donald C. Johanson Invertebrate Zoology Anthropology Watson M. Laetsch Botany Tori M. Hoehler Michael N. Josselyn Astrobiology and Biochemistry Ecology and Phycology Michael D. Lagios Ichthyology John P. Holdren David H. Kavanaugh Energy and Natural Resources Entomology Robert S. Lane Entomology Karen Holl James C. Kelley Restoration Ecology Oceanography Leo F. Laporte Geology Gustavo Hormiga Donald Kennedy Entomology Biological Sciences Ralph J. Larson Ichthyology Kim M. Howell William H. Kimbel Herpetology Anthropology John D. Lattin Entomology Sarah B. Hrdy Wim Kimmerer Anthropology Estuarine Ecology James V. Lawry, Jr. Anatomy James C. Ingle Patrick V. Kirch Paleontology Anthropology Burney J. Le Boeuf Biology B. L. Ingram David H. Kistner Geology Entomology Richard E. Leakey Anthropology Michael E. Irwin Richard G. Klein Entomology Paleoanthropology Walter S. Leal Chemical Ecology and Entomology Lynne A. Isbell Paul L. L. Koch Evolutionary Anthropology Paleobiology and Geobiology Gretchen LeBuhn Evolutionary Ecology Tomio Iwamoto Patrick Kociolek Ichthyology Diatom Studies Welton Lee Invertebrate Zoology Page 5 of 10 Academy Fellows Margaret S. Leinen James P. Mackey Katharine Milton Oceanography, Climate Engineering Herpetology Anthropology Janet L. Leonard David Maddison Brent D. Mishler Neurobiology and Behavior Entomology, Informatics, Computer Botany Programming and Software William A. Lester Development Richard L. Moe Physical Chemistry Phycology Rocco Mancinelli Alan E. Leviton Astrobiology John P. Monteverdi Herpetology Meteorology York T. Mandra Matthew R. Lewin