The Herpetology of Erie County, Pennsylvania: A Bibliography

Revised 2 nd Edition

Brian S. Gray and Mark Lethaby

Special Publication of the Natural History Museum at the Tom Ridge Environmental Center, Number 1

2 Special Publication of the Natural History Museum at the Tom Ridge Environmental Center

The Herpetology of Erie County, Pennsylvania: A Bibliography Revised 2 nd Edition

Compiled by

Brian S. Gray [email protected]

and

Mark Lethaby Natural History Museum at the Tom Ridge Environmental Center, 301 Peninsula Dr., Suite 3, Erie, PA 16505 [email protected]

Number 1 Erie, Pennsylvania 2017

Cover image: Smooth Greensnake, Opheodrys vernalis from Erie County, Pennsylvania.

3

Introduction

Since the first edition of The herpetology of Erie County, Pennsylvania: a bibliography (Gray and Lethaby 2012), numerous articles and books have been published that are pertinent to the literature of the region’s amphibians and reptiles. The purpose of this revision is to provide a comprehensive and updated list of publications for use by researchers interested in Erie County’s herpetofauna. We have made every effort to include all major works on the herpetology of Erie County. Included are the works of Atkinson (1901) and Surface (1906; 1908; 1913) which are among the earliest to note amphibians and or reptiles specifically from sites in Erie County, Pennsylvania. The earliest publication to utilize an Erie County specimen, however, may have been that of LeSueur (1817) in his description of Graptemys geographica (Lindeman 2009). While the bibliography is quite extensive, we did not attempt to list everything, such as articles in local newspapers, and unpublished reports, although some of the more significant of these are included. The authors welcome any literature citations that we may have missed, so that we may consider their inclusion in future editions of this bibliography.

Milk Snake, Lampropeltis triangulum from West Springfield, Erie County.

4

Bibliography of Erie County Herpetology

Anderson, E. 1983. Turtle attracts statewide attention. Erie Morning News, Sept. 15.

Anderson, E. 1989a. Nature’s way: Amphibians and turtles. Erie Morning News. April 13, 1989.

Anderson, E. 1989b. Nature’s way: The lucky frog. Erie Morning News. April 17, 1989.

Anderson, E. 2009. Nature's Way. Turtle focus of research. Erie Times-News Monday, August 31, 2009. 4B.

Anderson, E. and J. Stull. 1995. Nature Watch: 1985-1995 Erie, Pennsylvania and nearby areas. RNS Printing. Erie, PA 125 pp.

Aquatic Ecology Associates. 1977a. First interim data report for the period March 3, 1977 to May 20, 1977 for United States Steel Corporation. 600 Grant Street, Pittsburg, Pennsylvania 15230, May 31, 1977.

Aquatic Ecology Associates. 1977b. Aquatic and terrestrial ecology studies, Lake Front Plant, Conneaut, Ohio, summary report for the period May 21, 1977 to June 15, 1977. Ecology Associates, Pittsburgh, Pennsylvania.

Atkinson, D. A. 1901. The reptiles of Allegheny County, Pennsylvania. Annals of the Carnegie Museum 1:145-157.

Bartlett, J. 1999. Reptile study leaves unsettled questions. Erie Times Morning News. Monday, December 27, 1999, 4B.

Bartlett, J. 2000. Turtles, ducks might carry PCBs, mercury. Erie Times News. April 24, 2000. 1B and 8B.

Bartlett, J. 2001. Park puzzled by turtle deaths. Erie Times News vol.1 (193): 1A and 7A. April 13, 2001.

Bartlett, J. 2002. Lake Erie mudpuppies dying. Erie Times News vol. 2 (272): 1A and 7A. July 1, 2002.

Bartlett, J. 2003. Park staff a little shell-shocked. Erie Times News 3 (247): 1A and 4A. June 10, 2003.

Bates, S. P.1884. History of Erie County, Pennsylvania: Containing a history of the county; its townships, towns, villages, schools, churches, industries, etc.; portraits of early settlers and prominent men; biographies; history of Pennsylvania,

5 statistical and miscellaneous matter, etc., etc. Warner, Beers and Co., Chicago.

Black, D. 1998. ‘Hunters’ play shell game of turtle round-up. Erie Times News Sunday, August 16, 1998.

Bogart, J. and M. W. Klemens. 2008. Additional distributional records of Ambystoma laterale, A. jeffersonianum (Amphibia: Caudata) and their unisexual kleptogens in Northeastern North America. American Museum Novitates, Number 3627. 58 pp.

Brewer, M.A. and J. L. Schnars. 2007. The response of neonate snapping turtle (Chelydra serpentina serpentina ) hatchlings to a moisture gradient. Regional Science Consortium at the Tom Ridge Environmental Center at Presque Isle, 3rd Annual Research Symposium (abstract).

Campbell, J.M., J.K. Bissell, H.N. Cunningham, Jr., E.C. Masteller, and J. Stull. Biodiversity of Presque Isle. Pp. 405-422. In : Majumdar, S.K., F.J. Brenner, J.E. Lovich, J.F. Schalles, and E.W. Miller (eds.). Biological Diversity: Problems and Challenges. The Pennsylvania Academy of Sciences, Easton, Pennsylvania.

Carnegie Museum of Natural History: collection and distribution records. Provided by Stephen Rogers, collection manager.

Collins, H. H. 1932. Observations on the Life History of the Vermilion-spotted Newt, Triturus viridescens , in Western Pennsylvania Proc. Pa. Acad. Sci. 6:164-165.

Conant, R. 1975. Peterson field guide to reptiles and amphibians of eastern and central North America. Houghton Mifflin, Boston.

Ernst C. 1985. Blanding’s Turtle. In : Genoways, H. H., and F. J. Brenner (eds). Species of Special Concern in Pennsylvania. Special Publication of Carnegie Museum of Natural History, No. 11. Pittsburgh, PA.

Ernst, C. H. and E. M. Ernst. 1971. The taxonomic status and zoogeography of the Painted Turtle, Chrysemys picta , in Pennsylvania. Herpetologica 27(4):390-396.

Ernst, C. H. and J. E. Lovich. 2009. Turtles of the United States and Canada. Second edition. Johns Hopkins University Press, Baltimore. 827 pp.

Fontanella, F. M., C. R. Feldman, M. E. Siddall, and F. T. Burbrink. 2008. Phylogeography of Diadophis punctatus : extensive lineage diversity and repeated patterns of historical demography in a transcontinental snake. Molecular Phylogenetics and Evolution 46(2008):1049-1070.

Gray, B. 1999. Observations of amphibians and reptiles at Asbury Woods and along the Greenway Trail: With a preliminary list of species. The Gray Press (privately printed), Erie, PA. 18 pp. +10 plates.

6 Gray, B. 2000. Scaleless snake. Reptiles Magazine Vol.8(12):6

Gray, B. S. 2002a. Observations on the herpetofauna of the Asbury Woods Greenway in Pennsylvania. Bull. Chicago Herp. Soc. 37(2):21-24.

Gray, B. S. 2002b. Natural History Notes: Thamnophis sirtalis sirtalis (Eastern Garter Snake) Diet. Herpetol. Rev. 33(2):142-143.

Gray, B. S. 2002c. Natural History Notes: Storeria dekayi dekayi (Northern Brownsnake) and S. occipitomaculata occipitomaculata (Northern Red-bellied Snake). Morphology. Herpetol. Rev. 33(3):216.

Gray, B. S. 2002d. An addition to the herpetofauna of the Asbury Woods Greenway, Erie, Pennsylvania. Bull. Chicago Herp. Soc. 37(11):196-197.

Gray, B. S. 2004. Report on a population of spotted turtles, Clemmys guttata in western Erie County, Pennsylvania. Bull. Chicago Herp. Soc. 39(2):21-29.

Gray, B. S. 2005a. Checklist of the amphibians & reptiles of the Asbury Woods Greenway. Pamphlet. Privately printed.

Gray, B. S. 2005b. Note on the distribution of the Short-headed Gartersnake ( Thamnophis brachystoma ) in Erie County, Pennsylvania. Bull. Chicago Herp. Soc. 40(6):105- 106.

Spotted Salamander, Ambystoma maculatum from Asbury Woods, Erie County.

7

Gray, B. S. 2006. The reptiles and amphibians of the Asbury Woods Greenway, Erie County, Pennsylvania. Bull. Maryland Herp. Soc. 42 (2):115- 126.

Gray, B. S. 2007a. The herpetofauna of a national superfund site in Erie, Pennsylvania. Bull. Maryland Herp. Soc. Vol. 43(3):129-133.

Gray, B. S. 2007b. Inventory of amphibians along Wolf Run, Erie, Pennsylvania. Bull. Chicago Herp. Soc. Vol. 42(12):194.

Immature Grey Treefrog, Hyla versicolor from Asbury Woods, Erie County.

Gray, B. S. 2008a. A note on site fidelity for ecdysis in the Northern Brown Snake, Storeria dekayi dekayi . Bull. Chicago Herp. Soc. 43(3):

Gray, B. S. 2008b. Observations on the diet of the Shorthead Garter Snake, Thamnophis brachystoma. J. Kansas Herpetol. No. 25 (March 2008):24-28.

Gray, B. S. 2008c. A study of carapace and plastron patterns in the spotted turtle, Clemmys guttata , and their use as a technique for individual recognition. Bull. Chicago Herp. Soc. 43(7):109-114.

Gray, B. S. 2009a. Arboreal ecdysis in the Eastern Garter Snake ( Thamnophis sirtalis sirtalis ). J. Kansas Herpetol. 29:11.

Gray, B. S. 2009b. Recent observations of the herpetofauna of a former National Superfund Site in Erie County, Pennsylvania. J. Kansas Herpetol. 31:9-11.

Gray, B. S. 2009c. Anomalous scutellation in an Eastern Garter Snake from Erie County, Pennsylvania. Bull. Chicago Herp. Soc. 44(9):137-138.

Gray, B. S. 2009d. Estimating snout to vent length from data acquired from the shed skins of the Northern Brown Snake, Storeria dekayi dekayi . J. Kansas Herpetol. 32:17-19.

Gray, B. S. 2009e. Aberrant pattern in the Allegheny Dusky Salamander (Desmognathus ochrophaeus ) from Erie County, Pennsylvania. Bull. Chicago Herp. Soc. 44(11):166.

Gray, B. S. 2010. Distribution of native and exotic earthworms in the eastern US: implications for the ecology of vermivorous snakes. Bull. Chicago Herp. Soc. 45(5):73-86.

Gray, B. S. 2011a. Longevity and growth of a captive Shorthead Garter Snake, Thamnophis brachystoma . J. Kansas Herpetol. 38:10-11.

Gray, B. S. 2011b. Seasonal activity and natural history observations of five snake species from the Central Lowland Province of Erie County, Pennsylvania. J. Kansas Herpetol. 39:14-21.

Gray, B. S. 2011c. Random shots (photograph of a Spring Peeper) in Erie Reader. September 21 – October 4, 2011. Vol. 1(19):2.

Gray, B. S. 2011d. Random shots (photograph of a Blanding’s Turtle) in Erie Reader. October 5 – October 18, 2011. Vol. 1(20):2.

Gray, B. S. 2011e. A Guide to the Reptiles of Erie County, Pennsylvania. Privately Printed. Erie, PA. 88 pp.

Gray, B. S. 2011f. A Guide to the Amphibians of Erie County, Pennsylvania. Privately Printed. Erie, PA. 76 Pp.

Gray, B. S. 2012. A note on the diet of the Common Garter Snake, Thamnophis sirtalis , in Pennsylvania. Bull. Chicago Herp. Soc. 47(8):97-98.

Gray, B. S. 2013 a. Aggregations of Brownsnakes, Storeria dekayi (Holbrook, 1836), at a site in northwestern Pennsylvania. Bull. Chicago Herp. Soc. 48(3):33-38.

Gray, B. S. 2013 b. Natural history notes: Storeria dekayi (Dekay’s Brownsnake) Diet. Herpetol. Rev. 44(3):527.

Gray, B. S. 2013c. The diet of Dekay’s Brownsnake, Storeria dekayi at a site in Erie County, Pennsylvania. Regional Science Consortium at the Tom Ridge Environmental Center at Presque Isle, 9 th annual research symposium proceedings (abstract of poster presentation).

9

Common Five-lined Skink, Plestiodon fasciatus .

Gray, B. S. 2013 d. Observations on litters of Dekay’s Brownsnake, Storeria dekayi from an urban population in northwest Pennsylvania. Bull. Maryland Herp. Soc. 49(1-4):30-39.

Gray, B. S. 2013 e. The relative importance of slugs and earthworms in the diet of Dekay’s Brownsnake, Storeria dekayi , from northwestern Pennsylvania. Bull. Chicago Herp. Soc. 48(12):157-165.

Gray, B. S. 2014 a. Natural history of Dekay’s Brownsnake, Storeria dekayi (Holbrook 1836) at a site in northwestern Pennsylvania. J. N. Amer. Herpetol. 2014(1):28- 39.

Gray, B. S. 2014 b. Natural history notes: Storeria dekayi (Dekay’s Brownsnake) Cold-tolerance. Herpetol. Rev. 45(1):148.

Gray, B. S. 2014 c. Seasonal activity and temperature relationships of the eastern gartersnake, Thamnophis sirtalis sirtalis (Linnaeus, 1758), from an urban population in Erie County, Pennsylvania. Bull. Maryland Herp. Soc. 50(1- 2):22-33.

10 Gray, B. S. 2014 d. Kyphoscoliosis in a Dekay’s Brownsnake, Storeria dekayi , from Erie County, Pennsylvania, USA. Bull. Chicago Herp. Soc. 49(9):127-128.

Gray, B. S. 2014 e. An introduction to identifying prey remains in fecal samples of snakes of the genus Storeria . Bull. Chicago Herp. Soc. 49(10):142-146.

Gray, B. S. 2014 f. The defensive behaviors of Dekay’s Brownsnake, Storeria dekayi : An illustrated review of a diverse repertoire. Collinsorum 3(2-4):20-27.

Gray, B. S. 2015 a. Natural history notes: Storeria dekayi (Dekay’s Brownsnake) Diet. Herpetol. Rev. 46(1):108.

Gray, B. S. 2015 b. A Study of the Defensive Behaviors of Free-ranging Dekay’s Brownsnakes, Storeria dekayi (Holbrook, 1836). J. N. American Herpetol. 2015(1):43-52.

Gray, B. S. 2015 c. Natural History Observations of the Common Gartersnake, Thamnophis sirtalis from Northwest Pennsylvania. Bull. Chicago Herp. Soc. 50(5):61-72.

Gray, B.S. 2015 d. The scavenging of snake carcasses by the Spotted Snake Millipede, Blaniulus guttulatus . Collinsorum 4(2):3.

Gray, B. S. 2015 e. Natural history notes: Storeria dekayi (Dekay’s Brownsnake) Population size, density and biomass. Herpetol. Rev. 46(4):650-651.

Gray, B. S. 2016. Associations between Dekay’s Brownsnake, Storeria dekayi , and invertebrates. Bull. Chicago Herp. Soc. 51(2):25-28.

Gray, B. S. and P. Curtis. 2003. A note on sexual dichromatism in Clemmys guttata from Erie County, Pennsylvania. Bull. Chicago Herp. Soc. 38(6):120-121.

Gray, B. S. and M. Lethaby. 2004. A den site utilized by the Northern Red-bellied Snake, Storeria occipitomaculata occipitomaculata , in Pennsylvania. Bull. Maryland Herp. Soc. 40(3):94-96.

Gray, B. S. and M. Lethaby. 2008. The amphibians and reptiles of Erie County, Pennsylvania. Bull. Maryland Herp. Soc. 44(2):49-69.

Gray, B. S. and M. Lethaby. 2010. Observations of limb abnormalities in amphibians from Erie County, Pennsylvania. J. Kansas Herpetol. 35:14-16.

Gray, B. S. and M. Lethaby. 2012. An addendum to The Amphibians and Reptiles of Erie County, Pennsylvania. Bull. Maryland Herp. Soc. 48(1-4):11-19.

Gray, B. S., M. Lethaby, N. McAllister, and S. Bloomstine. 2013. Natural history notes.

11 Storeria occipitomaculata (Red-bellied Snake) Diet. Herpetol. Rev. 44(2):335.

Gray, B., H. M. Smith, and D. Chiszar. 2003. Further anomalies in the litters of a garter snake from a hazardous waste site. Bull. Chicago Herp. Soc. 38(1):4-6.

Gray, B., H. M. Smith, J. Woodling, and D. Chiszar. 2001. Some bizarre effects on snakes, supposedly from pollution, at a site in Pennsylvania. Bull. Chicago Herp. Soc. 36(7):144-148.

Green, D. M. and C. Parent. 2003. Variable and asymmetric introgression in a hybrid zone in the toads, Bufo americanus and Bufo fowleri . Copeia 2003(1):34-43.

Hahn, T. 2011. A jaw-dropper. Erie Times-News. Saturday, July 2, 2011.

Hudson, H.B. 1930. The distribution and habitat preference of the urodele amphibian Triturus viridescens , at Presque Isle, Erie, Pennsylvania. Proc. Pa. Acad. Sci. 4:55-58.

Hughes, J.R. and J.L. Schnars. 2007. Geographic Movements of the Common Snapping Turtle ( Chelydra serpentina serpentina ) on Presque Isle State Park, Erie Pennsylvania. Regional Science Consortium at the Tom Ridge Environmental Center at Presque Isle, 3rd Annual Research Symposium (abstract).

Hughes, J.R. and J.L. Schnars. 2007. A Herpetological Inventory of Presque Isle State Park, Erie, Pennsylvania Regional Science Consortium at the Tom Ridge Environmental Center at Presque Isle, 3rd Annual Research Symposium (abstract).

Hughes, J. R. and J. L. Schnars. 2008. Geographic movements of the common snapping turtle ( Chelydra s. serpentina ) on Presque Isle State Park, Erie, Pennsylvania. Proceedings of the Regional Science Consortium at the Tom Ridge Environmental Center at Presque Isle, 4 th Annual Research Symposium (Abstract).

Hulse, A. C. and K. L. Hulse. 1992. Geographic distribution. New county records for amphibians and reptiles from Pennsylvania. Herpetol. Rev. 23(2):62-64.

Hulse, A.C., C.J. McCoy and E.J. Censky.2001. Amphibians and Reptiles of Pennsylvania and the Northeast. Ithaca and London. Cornell University Press.

Jones, A. and C. Wilson. 2015. A reptile species inventory in Presque Isle State Park. Regional Science Consortium at the Tom Ridge Environmental Center at Presque Isle, 11th Annual Research Symposium (abstract).

King, R.B. 1988. Polymorphic populations of the garter snake Thamnophis sirtalis near Lake Erie. Herpetologica 44:451-458.

12

Snapping Turtle, Chelydra serpentina from Millcreek Township, Erie County.

LeSueur, C. A. 1817. An account of an American species of tortoise, not noticed in the systems. Journal of the Academy of Natural Sciences of Philadelphia 1:86-87.

Lethaby, M. 1990. Geographic distribution. Ambystoma jeffersonianum. Herpetol. Rev. 21(4):94.

Lethaby, M. 2001. Amphibians and reptiles on Presque Isle State Park: Summary of current knowledge.Unpublished report to Presque Isle State Park.

Lethaby, M. 2004. Natural History Notes: Thamnophis brachystoma . Maximum size. Herpetol. Rev. 35(1):73.

Lethaby, M. 2011a. Geographic distribution: Opheodrys vernalis . Herpetol. Rev. 42(1):115.

Lethaby, M. 2011b. Geographic distribution: Pantherophis spiloides . Herpetol. Rev. 42(1):115.

13

Lethaby, M. 2011c. Natural History Notes: Thamnophis brachystoma . Reproduction: maximum litter size. Herpetol. Rev. 42(4):620.

Lethaby, M. 2011d. Natural history notes: Thamnophis sauritus septentrionalis . Reproduction: litter size. Herpetol. Rev. 42(4):621.

Lethaby, M. and B. S. Gray. 2015. Natural history notes: Thamnophis brachystoma (Short-headed Gartersnake). Population size, density and biomass. Herpetol. Rev. 46(4):652.

Lethaby, M. and B. S. Gray. 2016. Natural history notes: Thamnophis brachystoma (Short-headed Gartersnake). Movements and site fidelity. Herpetol. Rev. 47(3):485.

Lethaby, M. and J. Tucci. 1990. Geographic distribution. Eumeces fasciatus . Herpetol. Rev. 21(4):96 .

Lindeman, P.V. 2006. Zebra and Quagga Mussels (Dreissena spp.) and Other Prey of a Lake Erie Population of Common Map Turtles (Emydidae: Graptemys geographica ). Copeia, 2006(2): 268-273.

Lindeman, P. V. 2009. On the type locality and type specimen of Testudo geographica LeSueur 1817. Chelonian Cons. Biol. 95-98.

Lindeman, P. V. 2013. The Map Turtle and Sawback Atlas: Ecology, evolution, distribution, and conservation. Animal Natural History Series Vol. 12. Univ. Oklahoma Press, Norman, OK.

McCoy, C.J. 1982. Amphibians and reptiles in Pennsylvania: check-list, bibliography, and atlas of distribution. Carnegie Museum of Natural History Special Publication No. 6, Pittsburgh, PA. 91pp.

McCoy, C. J. 1990. Status of Pennsylvania amphibians and reptiles. Final report submitted to the Pennsylvania Fish Commission, and The Wild Resources Conservation Fund. 112 pp.

McCoy, C.J. and A.V. Bianculli. 1966. The distribution and dispersal of Heterodon platyrhinos in Pennsylvania. J. Ohio Herp. Soc. 5: 153-158.

McCoy, C. J., K. C. Parkes, and J. C. Sacco. 1994. Diversity of terrestrial vertebrates in Pennsylvania. Pp. 315-325. In : Majumdar, S.K., F.J. Brenner, J.E. Lovich, J.F. Schalles, and E.W. Miller (eds.). Biological Diversity: Problems and Challenges. The Pennsylvania Academy of Sciences, Easton, Pennsylvania.

McKinstry, D. M. 1973. Blood parasites in snakes of northwestern Pennsylvania.

14 Journal of Parasitology 59(2):343.

McKinstry, D.M. 1975. Notes on the herpetology of Presque Isle State Park, Erie, Pennsylvania. Bull. Maryland Herp. Soc. 11:21-26.

McKinstry, D. M. and H. N. Cunningham. 1980. Reptiles and amphibians of select Lake Erie streams in northwestern Pennsylvania. Bull. Maryland Herp. Soc. 16(3):88- 93.

McKinstry, D. M. and H. N. Cunningham. 1989. Assessment of amphibian and reptile populations on Presque Isle State Park, Erie, Pennsylvania. Final report submitted to Wild Resources Conservation Board. Harrisburg, PA.

McKinstry, D. M., H. N. Cunningham, Jr. and M. A. Lethaby. 1991. Assessment of amphibian and reptile populations on Presque Isle State Park, Erie, Pennsylvania (abstract). J. Pa. Acad. Sci. 64(supplement and index):198.

McKinstry, D. M., H. N. Cunningham, Jr., M. A. Lethaby and P. R. McKinstry. 1989. Assessment of amphibian and reptile populations on Presque Isle State Park, Erie, Pennsylvania-A preliminary report (abstract). J. Pa. Acad. Sci. 63(1):52.

McKinstry, D. M., H. N. Cunningham, M. A. Lethaby and T. A. Shrout. 1999. An inventory of amphibians and reptiles of French Creek, Erie County, Pennsylvania through diurnal opportunistic collecting. J. Pa. Acad. Sci. 73(3):106-112.

McKinstry, D.M. and S. Felege. 1974. Snakes of northwestern Pennsylvania. Bull. Maryland Herp. Soc. 10:29-31.

McKinstry, D.M., M. Lethaby, and H. Cunningham. 1991. Amphibians and Reptiles of Presque Isle State Park, Erie County, Pennsylvania. J. Pa. Acad. Sci. 65(1):17-23

McKinstry, D.M., M. Lethaby, H.E. Donachy, and J.D. Wester. 1990. Geographic distribution. Thamnophis sirtalis sirtalis . Herpetol. Rev. 21(2):42.

McKinstry, D.M., R. McPherson, and N. McAllister. 1987. Herpetology of Presque Isle State Park, Erie, Pennsylvania. Bull. Maryland Herp. Soc. 23:58-64.

McPherson, Roger. 1982. Amphibians and reptiles of Presque Isle State Park, Erie, Pennsylvania. Report for the Presque Isle State Park Management Plan. Biology Dept. Clarion University of Pennsylvania, Clarion, PA.

Meshaka, W. E., Jr., and J. T. Collins. 2009. A Pocket Guide to Pennsylvania Snakes. The Commonwealth of Pennsylvania. 53 pp

15 Meshaka, W. E., Jr., and J. T. Collins. 2010. A Pocket Guide to Pennsylvania Frogs and Toads. The Commonwealth of Pennsylvania. 40 pp

Moseley, E. L. 1901. The python in Pennsylvania. Science. New series 14(361):852-853.

Wood Frog, Lithobates sylvaticus .

Netting, M. G. 1932. Blanding’s Turtle, Emys blandingii (Holbrook), in Pennsylvania. Copeia 1932:173-174.

Ostrander, S. J. 2000. Great natural areas in western Pennsylvania. Stackpole Books. Mechanicsburg, Pennsylvania.

Patterson, J.C. and P.V. Lindeman. 2006. Effects of Zebra and Quagga Mussel (Dreissena spp.) Invasion on the Feeding Habits of the Stinkpot ( Sternotherus odoratus ) on Presque Isle, Northwestern Pennsylvania. Regional Science Consortium at the Tom Ridge Environmental Center at Presque Isle, 3rd Annual Research Symposium (abstract).

Patterson, J.C. and P.V. Lindeman. 2009. Effects of Zebra and Quagga Mussel (Dreissena spp.) Invasion on the Feeding Habits of the Stinkpot ( Sternotherus odoratus ) on Presque Isle, Northwestern Pennsylvania. Northeastern Naturalist 16(3):365-374.

Price, A.H. 1978. New locality records and range extensions for Thamnophis brachystoma (Reptilia: Serpentes) in Pennsylvania. Bull. Maryland Herp. Soc. 14:260-263.

Richmond, N. D. 1952. First record of the Green Salamander in Pennsylvania, and other range extensions in Pennsylvania, Virginia and West Virginia. Annal. Carnegie Mus. Vol. 32: 313-318.

Ryan, K.M. and P.V. Lindeman. 2007. Reproductive allometry in the Common Map Turtle, Graptemys geographica . American Midl. Nat.158: 49-59.

Schnars, J. L. 2005. Effects of PCBs on developing embryos in the Common Snapping Turtle ( Chelydra serpentina serpentina ). Regional Science Consortium at the Tom Ridge Environmental Center at Presque Isle, 1st Annual Research Symposium (abstract).

Schnars, J. L. 2006. Using non-invasive techniques to determine mercury levels in

16 Snapping Turtle ( Chelydra serpentina serpentina ) tissues. Regional Science Consortium at the Tom Ridge Environmental Center at Presque Isle, 2nd Annual Research Symposium (abstract).

Shaffer, L. L. 1991. Pennsylvania amphibians and reptiles. Pennsylvania Fish Commission. Harrisburg, Pennsylvania.

Steele, M. A., M. C. Brittingham, T. J. Maret, and J. F. Merrit (eds.). 2010. Terrestrial Vertebrates of Pennsylvania: A Complete Guide to Species of Special Concern. The John Hopkins University Press, Baltimore. 507 pp.

Surface, H. A. 1906. The serpents of Pennsylvania. Monthly Bull. Div. Zool. PA. Dept. Agr. 4:114-202.

Surface, H. A. 1908. First report on the economic features of turtles of Pennsylvania. Zool. Bull. Div. Zool. PA Dept. Agr. 6:107-195.

Surface, H. A. 1913.First report on the economic features of the amphibians of Pennsylvania. Zool. Bull. Div. Zool. PA Dept. Agr. 3:68-153.

Blanding’s Turtle, Emydoidea blandingii from Presque Isle State Park, Erie County, Pennsylvania.

17

Swanson, P. L. 1948. Notes on the amphibians of Venango County, Pennsylvania. American Midl. Nat. 40(2):362-371.

Swanson, P. L. 1952. The reptiles of Venango County, Pennsylvania. Amer. Midl. Nat. 47(1):161-182.

The Lake Erie Region Conservancy. 2008. Pennsylvania Lake Erie Watershed Conservation Plan. Available at: http://lerc.wikispaces.com/conservation+plan

Tryon, Jr., C. A. 1974. Vertebrates of Pennsylvania and adjacent areas. Revised edition. The Pymatuning Laboratory of Ecology. Linesville, Pennsylvania.

Walker, M. 2000. Presque Isle State Park: A Scenic Tour of the Peninsula. Matthew D. Walker Publishing, Erie, Pennsylvania.

Wellington, R. J. 2005. Longevity of a captive deformed Northern Leopard Frog, Rana pipiens, in Erie County, Pennsylvania. Bull. Chicago Herp. Soc. 40(6):110-111.

Wellington, R. J. 2009. An overview of concerns and issues relating to the Mudpuppy, Necturus maculosus maculosus in Lake Erie/Presque Isle Bay, Erie County, Pennsylvania. Bull. Chicago Herp. Soc. 44(3):38-41.

Westcott, S. 1999. The summer of Presque Isle. Erie Times News. Morning News. Friday, May 28, 1999. 1A and 16A.

Williams, Samuel H. 1931. Preliminary report on the Animal Ecology of Presque Isle, Lake Erie, Pennsylvania. Proc. of the Pa. Acad. of Sci.5:88-97.

18

Northern Ribbonsnake, Thamnophis sauritus septentrionalis from Millcreek Township, Erie County.

19

20