Journal of Paleontology, 95(3), 2021, p. 586–600 Copyright © The Author(s), 2020. Published by Cambridge University Press on behalf of The Paleontological Society. This is an Open Access article, distributed under the terms of the Creative Commons Attribution-NonCommercial-ShareAlike licence (http:// creativecommons.org/licenses/by-nc-sa/4.0/), which permits non-commercial re-use, distribution, and reproduction in any medium, provided the same Creative Commons licence is included and the original work is properly cited. The written permission of Cambridge University Press must be obtained for commercial re-use. 0022-3360/21/1937-2337 doi: 10.1017/jpa.2020.100 Permian millipedes from the Fort Sill fissures of southwestern Oklahoma, with comments on allied taxa and millipedes preserved in karstic environments Joseph T. Hannibal1 and William J. May2 1Cleveland Museum of Natural History, 1 Wade Oval Drive, Cleveland, Ohio 44106, USA <
[email protected]> 2Sam Noble Oklahoma Museum of Natural History, University of Oklahoma, Norman, Oklahoma 73072, USA <
[email protected]> Abstract.—Permian millipedes are rare, especially so considering the relative abundance of millipedes in Carboniferous rocks. We report an early Permian millipede fauna containing three new genera and species of millipedes (Oklahoma- soma richardsspurense new genus new species, Karstiulus fortsillensis new genus new species, and Dolesea subtila new genus new species) found in fossil-producing pockets of the Fort Sill fissures exposed in the Dolese Quarry near Richards Spur, southwest Oklahoma, USA. These are the first new genera of invertebrates to be described from this site, one of the most prolific fossil-vertebrate sites in the world.