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Large Pterosaur Cladogram Peters Tritosauria Huehuecuetzpalli Macrocnemus BES SC111 Large Pterosaur Cladogram Macrocnemus T4822 Macrocnemus T2472 Peters - October 2015 Dinocephalosaurus Jesairosaurus Hypuronector 225 taxa 186 characters Vallesaurus Drepanosaurus Megalancosaurus Small taxa - phylogenetic miniaturization Characiopoda Amotosaurus Langobardisaurus Tanytrachelos Embryos and juveniles Fuyuansaurus Tanystropheus MSNM BES SC1018 Fenestrasauria Tanystropheus T/2819 Pterosauria (continued) Cosesaurus Sharovipteryx Longisquama Scaphognathids Scaphognathus crassirostris MPUM 6009 Pterosauria Raeticodactylus Scaphognathus SMNS 59395 Austriadactylus SMNS 56342 Scaphognathus Maxberg sp. Austriadactylus SC332466 TM 13104 Preondactylus Gmu10157 MCSNB 2887 BM NHM 42735 Dimorphodon macronyx BSp 1986 XV 132 Peteinosaurus Ex3359 ELTE V 256-Pester specimen Carniadactylus B St 1936 I 50 n30 MCSNB 8950 Cycnorhamphus Dimorphodon? weintraubi Cycnorhamphids Moganopterus IVPP V13758 embryo Feilongus Mesadactylus holotype Yixianopterus Dendrorhynchoides JZMP embryo Haopterus Dimorphodontids GLGMV0002 SMNS 81928 Boreopterus and Anurognathids Anurognathus Zhenyuanopterus CAG IG 02-81 Hamipterus Batrachognathus Arthurdactylus Daohugoupterus Coloborhynchus spielbergi Jeholopterus Criorhynchus Ornithocheirids Ikrandraco Eudimorphodon ranzii Istiodactylus Eudimorphodon cromptonellus SMNS 1136 PAL Eudimorphodon BSp 1994 SMNK PAL 3854 Campylognathoides HLZ 50 Campylognathoides Nurhachius C zitteli SMNS 11879 Brasileodactylus C zitteli SMNS 9787 Ludodactylus Nesodactylus Guidraco C liasicus CM 11424 Cearadactylus ligabuei Rhamph interm St/Ei 8209 (n28) Cearadactylus atrox Rhamph B St 1960 I 470A (n9) Liaoningopterus Bellubrunnus Anhanguera piscator Qinglongopterus SMNS 81775 Rhamph MBH unnumbered (n20) Ornithocephalus Rhamph BMNH 37012 (n85) SOS4593 R longicaudus B St 1959 I 400 (n10) SOS4006 R longicaudus B St 1889 XI 1 (n7) Jianchangopterus R longicaudus TM 6924 (n1) Painten private pteroaur R longicaudus B St 1938 I 503a (n11) Ningchengopterus Rhamph ROM 55352 Pterodactylus AMNH 1942 R longiceps GPIT RE 7321 (n81) Pterodactylus SMF 405 - n37 Rhamph NHMW 1998z0077/0001 Pterodactylids Pterodactylus NHMW 1975/1756 R longiceps BMNH 37002 (n82) Pterodactylus scolapaciceps BSP1937 I 18 - n21 Rhamph TMP2008.41.001 Pterodactylus BSP AS V 29a R muensteri JME SOS 4785 Pterodactylus antiquus BSP AS I 739 -n4 R muensteri MTM V2008.33.1 Pterodactylus BMMS 7 R muensteri WU970001 Pterodactylus longicollum SMNS 56603 R muensteri CM 11427 (n59) Germanodactylids BSt 1967 i 276 R muensteri YPM 1778 (n33) BMNH 42736 R muensteri JME SOS 4009 (n62) BM aM 4072 R muensteri SMF R 4128 (n43) BSt ASXIX3 R muensteri TM6920/21 (n38) Germanodactylus rhamphastinus R gemmingi TM 6922/6923 (n74) Germanodactylus JME MOE 12 R gemmingi GPIH MYE 13 (n75) Rhamphorhynchus Eosipterus GMV 2117 Rhamph MJML K1597 Wenupteryx MOZ 3625 R muensteri B St 1929 I 69 (n52) Cuspicephalus Changchengopterus CYGB-0036 Germanodactylus cristatus BSt 1892 IV 1-n61 Orientognathus SMNK PAL 3830 Sordes PIN 2585-25 Phobetor Sordes PIN 2470-1 Dsungaripterus Dorygnathids Cacibuteryx Noripterus Dorygnathus Hauff sp. Nemicolopterus Dorygnathus Vienna sp. Shenzhoupterus Dorygnathus MBR 1920.16 Sinopterus Sericipterus Huaxiapterus Dorygnathus SMNS 55886 Tapejara Dorygnathus Up R 156 Dsungaripterids Tupandactylus Dorygnathus purdoni Shenzhoupterids Sinopterus liui Angustinaripterus Tapejarids Tupuxura longicristatus St/E1I Tupuxura Goshura sp. MBR 3530 n40 Thalassodromeus AMNH 5147 Elanodactylus Gnathosaurus Germanodactylus SMNK-PAL6592 Ctenochasma elegans n45 Germanodactylus NMING F15005 Gegepterus B St 1878 VI 1 Gladocephaloideus Eopteranodon Ctenochasma n65 B Aurorazhdarcho not Eosipterus D2514 Eoazhdarcho Ctenochasmatids Pterodaustro Muzquizopteryx Pterodaustro embryo Nyctosaurus FHSMVP21 Dorygnathus SMNS51827 Nyctosaurus FMNH25026 Dorygnathus SMNS 50164 Nyctosaurus UNSM93000 TM 10341 Pteranodon YPM 1179 Beipiaopterus Pteranodon FHSM 2183 Ardeadactylus longicollum Pteranodon UUPI R197 Huanhepterus Pteranodon YPM 1177 Mesadactylus BYU specimen Pteranodon KUVP 2212 skull CM 11 426 n44 Pteranodon YPM 2594 B St 1911 I 31 (n42) Pteranodon CMC VP7203 SOS2428 (n57) Pteranodon FHSM 2851 SOS 2179 Pteranodon UNSM 12167 Jidapterus Pteranodontids Pteranodon NMC41 358 Chaoyangopterus LPM-R00076 Pteranodon UALVP 24238 Microtuban Pteranodon KUVP 27821 Chaoyangopterus holotype Pteranodon FHSM VP339 Azhdarchids and kin Zhejiangopterus Quetzalcoatlus Jianchangnathus Sordes 2585 3 Pterorhynchus Changchengopterus PMOL Wukongopterus Archaeoistiodactylus Kunpengopterus Darwinopterus AMNH M8802 Darwinopterus modularis ZMNH M 8782 Darwinopterus robustodens 41H111-0309A Wukongopterids Darwinopterus linglongtaensis IVPP V 16049 Darwinopterus YH2000.
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