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New Oviraptorid Dinosaur (Dinosauria: Oviraptorosauria) from the Nemegt Formation of Southwestern Mongolia
Bull. Natn. Sci. Mus., Tokyo, Ser. C, 30, pp. 95–130, December 22, 2004 New Oviraptorid Dinosaur (Dinosauria: Oviraptorosauria) from the Nemegt Formation of Southwestern Mongolia Junchang Lü1, Yukimitsu Tomida2, Yoichi Azuma3, Zhiming Dong4 and Yuong-Nam Lee5 1 Institute of Geology, Chinese Academy of Geological Sciences, Beijing 100037, China 2 National Science Museum, 3–23–1 Hyakunincho, Shinjukuku, Tokyo 169–0073, Japan 3 Fukui Prefectural Dinosaur Museum, 51–11 Terao, Muroko, Katsuyama 911–8601, Japan 4 Institute of Paleontology and Paleoanthropology, Chinese Academy of Sciences, Beijing 100044, China 5 Korea Institute of Geoscience and Mineral Resources, Geology & Geoinformation Division, 30 Gajeong-dong, Yuseong-gu, Daejeon 305–350, South Korea Abstract Nemegtia barsboldi gen. et sp. nov. here described is a new oviraptorid dinosaur from the Late Cretaceous (mid-Maastrichtian) Nemegt Formation of southwestern Mongolia. It differs from other oviraptorids in the skull having a well-developed crest, the anterior margin of which is nearly vertical, and the dorsal margin of the skull and the anterior margin of the crest form nearly 90°; the nasal process of the premaxilla being less exposed on the dorsal surface of the skull than those in other known oviraptorids; the length of the frontal being approximately one fourth that of the parietal along the midline of the skull. Phylogenetic analysis shows that Nemegtia barsboldi is more closely related to Citipati osmolskae than to any other oviraptorosaurs. Key words : Nemegt Basin, Mongolia, Nemegt Formation, Late Cretaceous, Oviraptorosauria, Nemegtia. dae, and Caudipterygidae (Barsbold, 1976; Stern- Introduction berg, 1940; Currie, 2000; Clark et al., 2001; Ji et Oviraptorosaurs are generally regarded as non- al., 1998; Zhou and Wang, 2000; Zhou et al., avian theropod dinosaurs (Osborn, 1924; Bars- 2000). -
Implications for Predatory Dinosaur Macroecology and Ontogeny in Later Late Cretaceous Asiamerica
Canadian Journal of Earth Sciences Theropod Guild Structure and the Tyrannosaurid Niche Assimilation Hypothesis: Implications for Predatory Dinosaur Macroecology and Ontogeny in later Late Cretaceous Asiamerica Journal: Canadian Journal of Earth Sciences Manuscript ID cjes-2020-0174.R1 Manuscript Type: Article Date Submitted by the 04-Jan-2021 Author: Complete List of Authors: Holtz, Thomas; University of Maryland at College Park, Department of Geology; NationalDraft Museum of Natural History, Department of Geology Keyword: Dinosaur, Ontogeny, Theropod, Paleocology, Mesozoic, Tyrannosauridae Is the invited manuscript for consideration in a Special Tribute to Dale Russell Issue? : © The Author(s) or their Institution(s) Page 1 of 91 Canadian Journal of Earth Sciences 1 Theropod Guild Structure and the Tyrannosaurid Niche Assimilation Hypothesis: 2 Implications for Predatory Dinosaur Macroecology and Ontogeny in later Late Cretaceous 3 Asiamerica 4 5 6 Thomas R. Holtz, Jr. 7 8 Department of Geology, University of Maryland, College Park, MD 20742 USA 9 Department of Paleobiology, National Museum of Natural History, Washington, DC 20013 USA 10 Email address: [email protected] 11 ORCID: 0000-0002-2906-4900 Draft 12 13 Thomas R. Holtz, Jr. 14 Department of Geology 15 8000 Regents Drive 16 University of Maryland 17 College Park, MD 20742 18 USA 19 Phone: 1-301-405-4084 20 Fax: 1-301-314-9661 21 Email address: [email protected] 22 23 1 © The Author(s) or their Institution(s) Canadian Journal of Earth Sciences Page 2 of 91 24 ABSTRACT 25 Well-sampled dinosaur communities from the Jurassic through the early Late Cretaceous show 26 greater taxonomic diversity among larger (>50kg) theropod taxa than communities of the 27 Campano-Maastrichtian, particularly to those of eastern/central Asia and Laramidia. -
AMERICAN MUSEUM NOVITATES Published by of NATURAL HISTORY Dec
AMERICAN MUSEUM NOVITATES Published by OF NATURAL HISTORY Dec. 4, 1933 Number 679 THE AMERICAN NewMUSEUMYork City 56.81, 9 (117:51.7) TWO NEW DINOSAURIAN REPTILES FROM MONGOLIA WITH NOTES ON SOME FRAGMENTARY SPECIMENS' BY CHARLES W. GILMORE2 CONTENTS PAGE INTRODUCTION......................................................... 1 DESCRIPTION OF GENERA AND SPECIES................................... 3 Order Orthopoda.................................................... 3 Family Nodosauridae............................................. 3 Pinacosaurus grangeri, new genus, new species. 3 Order Saurischia.................................................... 9 . Mongolosaurus haplodon, new genus, new species .. .. 11 NOTES ON OTHER DINOSAURIAN OCCURRENCES IN MONGOLIA................ 16 Family Deinodontidae.............................................. 16 Family Ornithomimidae............................................. 17 Family Hadrosauridae.............................................. 18 Family Ceratopsidae............................................... 19 BIBLIOGRAPHY............. 20 INTRODUCTION The present paper gives the results of a study of the residue of the dinosaur collections made in Mongolia by various Asiatic expeditions of The American Museum of Natural History. All of the material is frag- mentary, but diagnostic parts of two of the specimens are sufficiently preserved to be worthy of detailed description. One records the presence of a new genus and species of the Nodosauridae, the other a new genus and species of the Sauropoda. The -
Dinosaur Incubation Periods Directly Determined from Growth-Line Counts in Embryonic Teeth Show Reptilian-Grade Development
Dinosaur incubation periods directly determined from growth-line counts in embryonic teeth show reptilian-grade development Gregory M. Ericksona,1, Darla K. Zelenitskyb, David Ian Kaya, and Mark A. Norellc aDepartment of Biological Science, Florida State University, Tallahassee, FL 32306-4295; bDepartment of Geoscience, University of Calgary, Calgary, AB, Canada T2N 1N4; and cDivision of Paleontology, American Museum of Natural History, New York, NY 10024 Edited by Neil H. Shubin, University of Chicago, Chicago, IL, and approved December 1, 2016 (received for review August 17, 2016) Birds stand out from other egg-laying amniotes by producing anatomical, behavioral and eggshell attributes of birds related to relatively small numbers of large eggs with very short incubation reproduction [e.g., medullary bone (32), brooding (33–36), egg- periods (average 11–85 d). This aspect promotes high survivorship shell with multiple structural layers (37, 38), pigmented eggs (39), by limiting exposure to predation and environmental perturba- asymmetric eggs (19, 40, 41), and monoautochronic egg pro- tion, allows for larger more fit young, and facilitates rapid attain- duction (19, 40)] trace back to their dinosaurian ancestry (42). For ment of adult size. Birds are living dinosaurs; their rapid development such reasons, rapid avian incubation has generally been assumed has been considered to reflect the primitive dinosaurian condition. throughout Dinosauria (43–45). Here, nonavian dinosaurian incubation periods in both small and Incubation period estimates using regressions of typical avian large ornithischian taxa are empirically determined through growth- values relative to egg mass range from 45 to 80 d across the line counts in embryonic teeth. -
Vertebrate Paleontology of the Cretaceous/Tertiary Transition of Big Bend National Park, Texas (Lancian, Puercan, Mammalia, Dinosauria, Paleomagnetism)
Louisiana State University LSU Digital Commons LSU Historical Dissertations and Theses Graduate School 1986 Vertebrate Paleontology of the Cretaceous/Tertiary Transition of Big Bend National Park, Texas (Lancian, Puercan, Mammalia, Dinosauria, Paleomagnetism). Barbara R. Standhardt Louisiana State University and Agricultural & Mechanical College Follow this and additional works at: https://digitalcommons.lsu.edu/gradschool_disstheses Recommended Citation Standhardt, Barbara R., "Vertebrate Paleontology of the Cretaceous/Tertiary Transition of Big Bend National Park, Texas (Lancian, Puercan, Mammalia, Dinosauria, Paleomagnetism)." (1986). LSU Historical Dissertations and Theses. 4209. https://digitalcommons.lsu.edu/gradschool_disstheses/4209 This Dissertation is brought to you for free and open access by the Graduate School at LSU Digital Commons. It has been accepted for inclusion in LSU Historical Dissertations and Theses by an authorized administrator of LSU Digital Commons. For more information, please contact [email protected]. INFORMATION TO USERS This reproduction was made from a copy of a manuscript sent to us for publication and microfilming. While the most advanced technology has been used to pho tograph and reproduce this manuscript, the quality of the reproduction is heavily dependent upon the quality of the material submitted. Pages in any manuscript may have indistinct print. In all cases the best available copy has been filmed. The following explanation of techniques is provided to help clarify notations which may appear on this reproduction. 1. Manuscripts may not always be complete. When it is not possible to obtain missing pages, a note appears to indicate this. 2. When copyrighted materials are removed from the manuscript, a note ap pears to indicate this. 3. -
AMERICAN MUSEUM NOVITATES Published by Number 201 Tnnaszwcas Ezusuo¶ Atmal History Nov
AMERICAN MUSEUM NOVITATES Published by Number 201 TNnAszwcAs ezusuo¶ ATmAL HisToRy Nov. 24, 1925 56.9(116 :51.7) A MESOZOIC MAMMAL SKULL FROM MONGOLIA' BY GEORGE GAY.LORD SIMPSON Among the many remarkable discoveries of the Third Asiatic Expedition of The American Museum of Natural History is a large part of the skull with associated lower jaws of a Mesozoic mammal. This unique specimen was found by A. F. Johnson in the Djadochta forma- tion at Shabarakh Usu in Mongolia. Mesozoic mammals are among the rarest of the fossils, and, while it might well have been inferred that there were multituberculates in Mongolia in Cretaceous times, this splendid confirmation is no less startling than it is welcome. Only one other partial Mesozoic mammal skull has been found, that of Tritylodon Owen (1884) from the Lower Jurassic of South Africa, and the lower jaw of the latter form is quite unknown. No Mesozoic mammal has hitherto been found in Asia, and, although mammals occur at earlier and later horizons, none has yet been found in a formation of age equiva- lent to the Djadochta. Consequently, this specimen is noteworthy, not only in being the second known skull, and the first with associated lower jaws, and as adding new morphological knowledge, but it also adds a new horizon to the stratigraphic range and a new continent to the geographic distribution. The writer is deeply indebted to Professor Osborn and Doctor Matthew for the great privilege of naming and describing this important form. The preparation, a task calling for the greatest delicacy and address, owing to the softness of the bone and the nature of the sandstone matrix, has been undertaken with great skill by Mr. -
Unexpected New Lizard from the Late Cretaceous of Southern South America Sheds Light on Gondwanan Squamate Diversity
Rev. Mus. Argentino Cienc. Nat., n.s. 23(1): 57-80, 2021 ISSN 1514-5158 (impresa) ISSN 1853-0400 (en línea) Unexpected new lizard from the Late Cretaceous of southern South America sheds light on Gondwanan squamate diversity Agustín G. MARTINELLI1, Federico L. AGNOLÍN2,3 & Martín D. EZCURRA1 1Sección Paleontología de Vertebrados, CONICET–Museo Argentino de Ciencias Naturales ‘Bernardino Rivadavia’, Avenida Ángel Gallardo 470, C1405DJR, CABA, Argentina. [email protected]. ar / [email protected]. 2Laboratorio de Anatomía Comparada y Evolución de los Vertebrados, CONICET–Museo Argentino de Ciencias Naturales “Bernardino Rivadavia”, Avenida Ángel Gallardo 470, C1405DJR, CABA, Argentina. [email protected]. 3Fundación de Historia Natural “Félix de Azara”, Departamento de Ciencias Naturales y Antropología. CEBBAD - Universidad Maimónides. Hidalgo 775, C1405BDB, CABA, Argentina. Abstract: The record of Cretaceous terrestrial lizards (Squamata) in South America is patchy, with seven species described from north-eastern and south-eastern Brazil, and few isolated records of iguanians and scincomorphans from the Argentinian Patagonia. Herein we describe a new genus and species of Cretaceous lizard, Paleochelco occultato gen. et sp. nov., based on a partial skull (MACN-Pv-N 120) discovered about three decades ago that was unnoticed in the Colección Paleovertebrados of the Museo Argentino de Ciencias Naturales “Bernardino Rivadavia”. It comes from rocks of the Upper Cretaceous Bajo de la Carpa Formation (Neuquén Group) exposed at the Campus of the Universidad Nacional del Comahue, north of Neuquén City (Neuquén Province). The new taxon was included into a broad phylogenetic dataset of squamates and it was recovered around the base of Polyglyphanodontia in a constrained analysis using a total-evidence backbone. -
Bayanzag 1-Page
THE FLAMING CLIFFS MONGOLIA’S FAMOUS FOSSIL BEDS The Flaming Cliffs in the heart of the Gobi Desert are known worldwide for containing exceptional dinosaur fossils. The cliffs, the fossils, and the plants and animals who live nearby are protected by local government as part of Bayanzag Park. As you enjoy the park, please leave everything as you find it – for science, for the environment, and for Mongolia. WHO LIVED HERE A DELICATE LIVING ECOSYSTEM The Gobi Desert is home to animals and plants that live 80 MILLION YEARS AGO? nowhere else. Follow these rules to keep Bayanzag’s living The time was the Late Cretaceous: the last era when ecosystem from going extinct like the dinosaurs: dinosaurs ruled the earth. The Flaming Cliffs were sand dunes in a desert oasis, and many animals came here to hunt, Never damage plants or collect wood. forage, and raise their young. Meet three of them: Never feed or harm wildlife. Take your trash with you when you leave. Weighing two tons and covered with FOUND A FOSSIL? spiky armor, Pinacosaurus was a formidable plant-eater first discovered Leave it alone. Fossils lose scientific data at the Flaming Cliffs in 1923. and can easily be destroyed if moved. Plus, removing fossils without a permit is illegal. Protoceratops was the first dinosaur If it’s more than a small discovered in fragment, notify the local Mongolia, at officials of Umnugovi the Flaming Aimag. Cliffs in 1922. It ate plants and Also notify the ISMD at was about the size of MongolianDinosaurs.org. Take a sheep. -
Gobekko Cretacicus Gen. Et Sp. N., a New Gekkonid Lizard from the Cretaceous of the Gobi Desert
Acta Palaeontologica Polonica Vol. 35, N~o. 1-2 pp. 67-76; pl 17, 18 W~XSZ~W,1990 MAGDALENA BORSUK-BIAEYNICKA GOBEKKO CRETACICUS GEN. ET SP. N., A NEW GEKKONID LIZARD FROM THE CRETACEOUS OF THE GOBI DESERT BORSUK-BIALYNICKA, M.: Gobekko cretackus gen. et sp. n., a new gekkonid' lizard from the Cretaceous of the Gobi Desert. Acta Palaeont. Polonica, 35, 1-2, 67-76, 1990. Gobekko cretactcus gen. et sp. n., a new representative of the family Gek- konidae sensu Kluge (1987), has been described from the ?upper Santonian andlor ?lower Campanian Djadochta Formation of the Gobi Desert. The presence of this family in the Cretaceous of Central Asia, suggested by earlier finding of fragmentary remains of Hoburogekko suchanovi Alifanov 1989 in the Aptian- -Albian of the Gobi Desert, is thus confirmed. K e y w o r d s: Reptilia, Sauria, Gekkota, Cretaceous, Mongolia. Magdalena Borsuk-Bialynlcka, Zaklad Paleobtologtt, Polska Akademta Nauk, 02- 089 Warszawa, Al. ewlrkt i Wtgury 03. Poland. Received: September 1989. INTRODUCTION The present paper concerns the remains of a gekkotan lizard collected by the Polish-Mongolian Paleontological Expeditions to the Gobi Desert (1963-1971), from the upper Cretaceous deposits of Mongolia. The remains of the Gekkota are rather rare in the Cretaceous. Of twenty five lizard genera described hitherto from the Cretaceous of the Gobi Desert (Gilmore 1943, Sulimski 1972, 1975, 1978, 1984, Borsuk- -Bialynicka 1984, 1985, 1986, 1988, Alifanov 1988, 1989 see also Estes 1983a) only two belong to the Gekkota. For the first time, the Gekkota have been recorded in the Cretaceous by Soviet paleontologists from the Mongolian locality Khobur, of the Albian-Aptian age (Alifanov 1989). -
American Museum Novitates
AMERICAN MUSEUM NOVITATES Number 3899, 44 pp. April 26, 2018 A Second Specimen of Citipati osmolskae Associated with a Nest of Eggs from Ukhaa Tolgod, Omnogov Aimag, Mongolia MARK A. NORELL,1, 2 AMY M. BALANOFF,1, 3 DANIEL E. BARTA,1, 2 AND GREGORY M. ERICKSON1, 4 ABSTRACT Adult dinosaurs preserved attending their nests in brooding positions are among the rarest vertebrate fossils. By far the most common occurrences are members of the dinosaur group Oviraptorosauria. The first finds of these were specimens recovered from the Djadokhta Forma- tion at the Mongolian locality of Ukhaa Tolgod and the Chinese locality of Bayan Mandahu. Since the initial discovery of these specimens, a few more occurrences of nesting oviraptors have been found at other Asian localities. Here we report on a second nesting oviraptorid specimen (IGM 100/1004) sitting in a brooding position atop a nest of eggs from Ukhaa Tolgod, Omnogov, Mongolia. This is a large specimen of the ubiquitous Ukhaa Tolgod taxon Citipati osmolskae. It is approximately 11% larger based on humeral length than the original Ukhaa Tolgod nesting Citipati osmolskae specimen (IGM 100/979), yet eggshell structure and egg arrangement are identical. No evidence for colonial breeding of these animals has been recovered. Reexamination of another “nesting” oviraptorosaur, the holotype of Oviraptor philoceratops (AMNH FARB 6517) indicates that in addition to the numerous partial eggs associated with the original skeleton that originally led to its referral as a protoceratopsian predator, there are the remains of a tiny theropod. This hind limb can be provisionally assigned to Oviraptoridae. It is thus at least possible that some of the eggs associated with the holotype had hatched and the perinates had not left the nest. -
From the Late Inner Mongolia
bulletin de l'institut royal des sciences naturelles de belgique sciences de la terre, 71-supp.: 5-28, 2001 bulletin van het koninklijk belgisch instituut voor natuurwetenschappen aardwetenschappen, 71-supp.: 5-28, 2001 A new Species of Protoceratops (Dinosauria, Neoceratopsia) from the Late Cretaceous of Inner Mongolia (P. R. China)1 Olivier LAMBERT, Pascal GODEFROIT, Hong LI, Chang-Young SHANG & Zhi-Ming DONG Lambert, O., Godefroit, P., Li, H., Shang, C.-Y, & Dong, Mots-clefs: Neoceratopsia, Protoceratops, Crétacé supérieur, Z.-M., 2001. - A new Species of Protoceratops (Dinosauria, Mongolie intérieure, phylogénie. Neoceratopsia) from the Late Cretaceous of Inner Mongolia (P. R. China). Bulletin de l'Institut royal des Sciences naturelles de Belgique, Introduction Sciences de la Terre, Supplement 71: 05-28, 4 pis., 13 fïgs., 2 tables; Bruxelles - Brussel, December 15, 2001. - ISSN 0374-6291. Protoceratops is probably the most emblematic dinosaur from the Gobi Desert. The first specimen was discovered Abstract in the Djadokhta Formation (Campanian) at the legendary Flaming Cliffs (Bayn Dzak, P. R. Mongolia) by the Third The basai neoceratopsian dinosaur Protoceratops hellenikorhinus nov. Central Asiatic Expédition of the AMNH on September sp. is described on the basis of material discovered by the Sino-Belgian 2, 1922 (Andrews, 1932). This skull (AMNH 6251) of a Expéditions in Inner Mongolia, in Campanian (Upper Cretaceous) deposits at Bayan Mandahu (Inner Mongolia, P.R. China). This new juvénile individual, lacking the front of the face and the species is characterised by its large size and by a series of cranial frill, was described as Protoceratops andrewsi Granger autapomorphies linked to the important development of the mandibu- & Gregory, 1923. -
Digitalcommons@University of Nebraska - Lincoln
University of Nebraska - Lincoln DigitalCommons@University of Nebraska - Lincoln Earth and Atmospheric Sciences, Department Papers in the Earth and Atmospheric Sciences of 2005 New Stratigraphic Subdivision, Depositional Environment, and Age Estimate for the Upper Cretaceous Djadokhta Formation, Southern Ulan Nur Basin, Mongolia Demberelyin Dashzeveg Geological Institute of the Mongolian Academy of Sciences, [email protected] Lowell Dingus American Museum of Natural History, [email protected] David B. Loope University of Nebraska, Lincoln, [email protected] Carl C. Swisher III Rutgers University Togtokh Dulam Mongolian Geological Survey See next page for additional authors Follow this and additional works at: https://digitalcommons.unl.edu/geosciencefacpub Part of the Earth Sciences Commons Dashzeveg, Demberelyin; Dingus, Lowell; Loope, David B.; Swisher, Carl C. III; Dulam, Togtokh; and Sweeney, Mark R., "New Stratigraphic Subdivision, Depositional Environment, and Age Estimate for the Upper Cretaceous Djadokhta Formation, Southern Ulan Nur Basin, Mongolia" (2005). Papers in the Earth and Atmospheric Sciences. 209. https://digitalcommons.unl.edu/geosciencefacpub/209 This Article is brought to you for free and open access by the Earth and Atmospheric Sciences, Department of at DigitalCommons@University of Nebraska - Lincoln. It has been accepted for inclusion in Papers in the Earth and Atmospheric Sciences by an authorized administrator of DigitalCommons@University of Nebraska - Lincoln. Authors Demberelyin Dashzeveg, Lowell Dingus,