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Catalog of the Staphylinidae (Insecta: Coleoptera)

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CATALOG OF THE STAPHYLINIDAE (INSECTA: COLEOPTERA). 1758 TO THE END OF THE SECOND MILLENNIUM. V. STAPHYLININE GROUP (PART 2) STAPHYLININAE: DIOCHINI, MAOROTHIINI, OTHIINI, PLATYPROSOPINI, STAPHYLININI (AMBLYOPININA, ANISOLININA, HYPTIOMINA, PHILONTHINA) LEE H. HERMAN BULLETIN OF THE AMERICAN MUSEUM OF NATURAL HISTORY NUMBER 265 NEW YORK : 2001 Recent issues of the Bulletin may be purchased from the Museum. Lists of back issues of the Bulletin, Novitates, and Anthropological Papers published during the last five years are available at World Wide Web site http://nimidi.amnh.org. Or address mail orders to: American Museum of Natural History Library, Central Park West at 79th St., New York, N.Y. 10024. TEL: (212) 769- 5545. FAX: (212) 769-5009. E-MAIL: [email protected] CATALOG OF THE STAPHYLINIDAE (INSECTA: COLEOPTERA). 1758 TO THE END OF THE SECOND MILLENNIUM. V. STAPHYLININE GROUP (PART 2) STAPHYLININAE: DIOCHINI, MAOROTHIINI, OTHIINI, PLATYPROSOPINI, STAPHYLININI (AMBLYOPININA, ANISOLININA, HYPTIOMINA, PHILONTHINA) LEE H. HERMAN Curator, Division of Invertebrate Zoology American Museum of Natural History BULLETIN OF THE AMERICAN MUSEUM OF NATURAL HISTORY Number 265, pages 2441–3020 Issued July 18, 2001 (Parts I–VII) Copyright © American Museum of Natural History 2001 ISSN 0003-0090 CATALOG OF THE STAPHYLINIDAE, PARTS I–VII PART I, pages 1–650 Introduction Brief History of Taxonomic Studies of the Staphylinidae Research History • Biographical Sketches Overview of Classification Omaliine Group Dasycerinae • Empelinae • Glypholomatinae • Micropeplinae • Microsilphinae • Neophoninae • Omaliinae • Proteininae • Protopselaphinae PART II, pages 651–1066 Tachyporine Group Habrocerinae • Olisthaerinae • Phloeocharinae • Tachyporinae • Trichophyinae PART III, pages 1067–1806 Oxyteline Group Apateticinae • Osoriinae • Oxytelinae • Piestinae • Trigonurinae PART IV, pages 1807–2440 Staphylinine Group Euaesthetinae • Leptotyphlinae • Megalopsidiinae • Oxyporinae • Pseudopsinae • Solieriinae • Steninae PART V, pages 2441–3020 Staphylinine Group, continued Staphylininae Diochini • Maorothiini • Othiini • Platyprosopini • Staphylinini (Amblyopinina • Hyptiomina • Anisolinina • Philonthina) PART VI, pages 3021–3840 Staphylinine Group, continued Staphylininae, continued Staphylinini (Quediina • Staphylinina • Tanygnathinina • Xanthopygina • Incertae Sedis) • Xantholinini Staphylinidae: Subfamily Incertae Sedis Protactinae† PART VII, pages 3841–4218 Bibliography Index iii CATALOG OF THE STAPHYLINIDAE (INSECTA: COLEOPTERA). 1758 TO THE END OF THE SECOND MILLENNIUM ABSTRACT This catalog (published in seven parts, all released on the same day) is based on only the published lit- erature for the Staphylinidae. Of the 32 subfamilies, the following 28 are included herein: Apateticinae, Dasycerinae, Empelinae, Euaesthetinae, Glypholomatinae, Habrocerinae, Leptotyphlinae, Megalopsidiinae, Micropeplinae, Microsilphinae, Neophoninae, Olisthaerinae, Omaliinae, Osoriinae, Oxyporinae, Oxytelinae, Phloeocharinae, Piestinae, Protactinae†, Proteininae, Protopselaphinae, Pseudopsinae, Solieriinae, Staphylin- inae, Steninae, Tachyporinae, Trichophyinae, and Trigonurinae. The Aleocharinae, Paederinae, Pselphinae, and Scaphidiinae are excluded from this edition of the catalog. References to the original citation or descrip- tion are given for available family-group, genus-group, and species-group names of both extant and extinct forms. The type genus is cited for each family-group name, the type species for each genus-group name, and the type locality for each species-group name. Where appropriate, all subgenera, subspecies, or syn- onyms are listed for each valid name. Annotated subsequent references are presented for all names. Dis- tributional summaries are given for each valid taxon. Full bibliographic citations are in Part VII. A short historical review, coauthored with Alesˇ Smetana, follows the Introduction (Part I), with the main focus on biographical sketches that include many photographs. The goal of this catalog is to summarize the current state of knowledge of the family and to stimulate worldwide monographic studies. Address correspondence to: Lee H. Herman, Curator, Division of Invertebrates, American Museum of Natural History, Central Park West at 79th Street, New York, NY 10024-5192, USA ([email protected]). Please send comments, corrections, or updates to either the postal or e-mail address. Reprints of new publications would be helpful and appreciated. iv CONTENTS PART V STAPHYLININE GROUP (PART 2) Staphylininae: Diochini, Maorothiini, Othiini, Platyprosopini, Staphylinini (Amblyopinina, Anisolinina, Hyptiomina, Philonthina) Staphylinine Group, continued . 2441 Staphylininae . 2441 Diochini . 2442 Maorothiini . 2450 Othiini . 2451 Platyprosopini . 2488 Staphylinini . 2495 Amblyopinina . 2496 Anisolinina . 2505 Hyptiomina . 2510 Philonthina . 2514 PARTS I–IV, VI, VII Introduction, History, and Biographical Sketches . Part I Omaliine Group . Part I Tachyporine Group . Part II Oxyteline Group . Part III Staphylinine Group . Parts IV, VI Staphylinidae: Subfamily Incertae Sedis . Part VI Protactinae† . Part VI Bibliography . Part VII Index . Part VII v STAPHYLININAE [294 genera; 6665 species; All Regions except the Antarctic] Staphylininae Latreille, 1802. — Eichelbaum, 1909: 159 (list of genera with references; generic synonyms; number of genera; number of species in each genus; general distribution of genera). — Reitter, 1909: 105 (key to tribus). — Cameron, 1921a (characters; key to Singapore species and genera). —Sˇulc, 1922 (osmeterium; anatomy; histology; physiology). — Porta, 1926: 85 (characters; species of Italy). — Böving and Craighead, 1930: 30 (larval characters). — Cameron, 1932: 1 (characters; key to tribes). — Joy, 1932: 105 (key to British tribes). — Scheerpeltz, 1933: 1297 (world catalog supplement). — Voris, 1940 (phylogenetic relationships based on larvae). — Voris, 1939a: 289 (egg, larval and pupal characters). — Bernet Kempers, 1941: 35 (larval characters). — Scheerpeltz, 1940: 38 (key to Palaearctic genera). — Blackwelder, 1943: 393 (notes; key to tribes of West Indies). — Blackwelder, 1944: 131 (checklist of species from Mexico, Central and South America, and the West Indies). — V. Hansen, 1952: 1 (characters; Danish species). — Crowson, 1955: 34 (adult and larval characters in key). — Hatch, 1957: 48, 172 (characters; key to tribes and genera of Pacific Northwest). — Arnett, 1963: 246, 274 (key to North American genera; characters; list of North American genera). — Székessy, 1963: 46 (characters; key to genera of Hungary). — Palm, 1963b: 38 (characters; key to tribe, genera, and species of Sweden). — Steel, 1963: 54 (list of species of Malham Tarn; collecting notes; Britain). — Lohse, 1964: 169 (characters; key to central European species). — Moore, 1965: 97 (key to Nearctic genera). — Seevers, 1965: 323 (myrmecophilus species; notes on classification). — Kasule, 1966: 266, 274 (larval characters included Xantholinini; notes). — Pototskaia, 1967: 12 (larval characters; key to tribe). — Coiffait and Saiz, 1968: 346 (key to tribes and genera of Chile). — Steel and Woodroffe, 1969: 136 (list of species collected from Isle of Rhum; collecting notes). — Britton, 1970: 545 (characters of adults; Australia; notes). — Kasule, 1970: 56 (larval characters). — Moore and Legner, 1973c: 212 (key to genera of United States and Canada). — Moore and Legner, 1974g: 560 (key to genera of America north of Mexico). — Moore and Legner, 1974h: 470 (catalog of the taxonomy, biology, and ecology of develop- mental stages of species of America north of Mexico). — Coiffait, 1974: 1 (characters; key to tribes, genera and species of western Palaearctic region). — Moore and Legner, 1975: 6 (catalog; North America north of Mexico). — Topp, 1978: 307, 311 (larval characters; key to genera). — Moore and Legner, 1979: 14 (characters; tabular key to genera of North America north of Mexico). — Muona, 1979: 14 (list of Scandinavian species). — Burakowski, Mroczkowski, and Stefan´ska, 1980: 29 (catalog; Poland). — Spahr, 1981: 93 (references to specimens from amber and copal). — Watt and McColl, 1982: 19, 20 (characters; notes on New Zealand taxa). — Pajni and Kohli, 1982: 10 (comparative morphology of aedeagus). — E. Matthews, 1982: 7, 24 (notes; key to genera of Australia). — Tóth, 1983: 183 (list of species of the Hortobágy National Park, Hungary). — Tóth, 1984 (characters; key to genera and species of Hungary). 2441 2442 STAPHYLININAE Bulletin AMNH — Outerelo and Gamarra, 1985: 47 (key to genera of Iberian Peninsula). — Tóth, 1985a: 88 (list of species of Bakony Mountains, Hungary). — Dettner, 1987: 30 (defensive glands). — Drugmand 1987b (comments on distribution of species in Belgium). — Gusarov, 1989 (list of species from the Crimea; ecological and habitat notes). — Frank, 1991: 346 (larval characters). — Frania, 1992 (studies on ecdysis and metamorphosis). — Newton and Thayer, 1992: 13 (summary of current classification; discussion). — Ashe and Newton, 1993: 281 (related to Paederinae and Pseudopsinae). — Dettner, 1993a (summary and discussion of defensive secretions and exocrine glands and their bearing on phylogeny). — Hayashi, 1993b: 282 (terms for taxonomic characters). — Welch, 1993 (ovariole structure and development). — Lawrence and Newton, 1995: 830 (notes). — Sparacio, 1995: 160 (characters; species of Sicily). — Klimaszewski, Newton, and Thayer, 1996: 150 (review of subfamily for New Zealand; notes; list of New Zealand genera; number of New
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