"- - 70.096

60.096

50.096

40.096

30.096

20.096

10.096

0.0%

Fig. 1 Most abundant Apiformes calculated as a proportion of the total abundance of Apiformes in the collection period. Pinon Canyon Maneuver Site, 2008. 04% 1 j 0.391>

0.2%

0.1%

0.0%

Fig. 2 Least abundant Apiformes species calculated as a proportion of the total abundance of Apiformes in the collection period. Pinon Canyon Maneuver Site, 2008.7

Fig. 3 Most abundant Carabidae species calculated as a proportion of the total abundance of Carabidae in the collection period. Pinon Canyon Maneuver Site, 2008.

Fig. 4 Least abundant Carabidae species calculated as a proportion of the total abundance of Carabidae in the collection period. Pinon Canyon Maneuver Site, 2008.

Fig. 5 species abundance calculated as a proportion of the total abundace of Asilidae in the collection period. Pinon Canyon Maneuver Site, 2008.

30.0%

25.0%

20.0%

15.0%

10.0%

5.0%

0.0%

Fig. 6 species abundance calculated as a proportion of the total abundance of in the collection period. Pinon Canyon Maneuver Site, 2008.

Fig. 7 Most abundant species calculated as a proportion of the total abundance of Orthoptera in the collection period. Pinon Canyon Maneuver Site, 2008.

Fig. 8 Moderately abundant Orthoptera species calculated as a proportion of the total abundance of Orthoptera in the collection period. Pinon Canyon Maneuver Site, 2008.

Fig. 9 Least abundant Orthoptera species calculated as a proportion of the total abundance of Orthoptera in the collection period. Pinon Canyon Maneuver Site, 2008.

Fig. 10 Apiformes dominant species comparison between 2007 and 2008. Each species as a percentage of the entire taxa. Pinon Canyon Maneuver Site, 2008.

Fig. 11 Carabidae dominant species comparison between 2007 and 2008. Each species as a percentage of the entire taxa. Pinon Canyon Maneuver Site, 2008.

Fig. 12 Asilidae dominant species comparison between 2007 and 2008. Each species as a percentage of the entire taxa. Pinon Canyon Maneuver Site, 2008.

2009 Butterflies

Other

Callophrys gryneus communis Hesperia pahaska Hesperia uncas picta

Nathalis iole Euptoieta claudia Pontia protodice

Cercyonis pegala

Speyeria coronis Colias philodice Brephidium exile

Fig. 13 Butterfly dominant species comparison between 2007, 2008 and 2009. Each species as a percentage of the entire taxa. Pinon Canyon Maneuver Site, 2009

Fig. 14 Orthoptera dominant species comparison between 2007 and 2008. Each species as a percentage of the entire taxa. Pinon Canyon Maneuver Site, 2008.

Fig. 15 dominant species comparison between 2007 and 2008. Each species as a percentage of the entire taxa. Pinon Canyon Maneuver Site, 2008.

The only species of concern observed in 2007, rita (Mattoni, [1966]) was not observed in 2008. In an effort to document E. rita in

2008, we searched for host plants, larvae and adults of this species while carrying out other collection work, in additon to the butterfly survey technique that was used to document the species in 2007. We collected two individuals of a , Alypia octomaculata (Fabricius), in the

Cottonwood habitat. The occurrence of the moth at Pinon Canyon will be a range extension if a persistent population can be demonstrated. We

will look for A. octomaculata larva and its plant hosts in 2009 to verify a persistant population. We collected 86 new species this year. The biology and behavor of the individual species will determine a species presence in a habitat based on yearly environmental variations. An updated species list can be found in Appendix 1. Species added to the list in 2008 are in bold.

Habitat Results

Fig. 16 Sand sagebrush habitat in June of 2007 on the left and June of 2008 on the right to demonstrate differences in habitat quality. Pinon Canyon Maneuver Site, 2008.

Habitat quality was relatively poor in 2008 when compared with 2007, based on visual assessment of the habitat by the presence of green,

flowering plants or moisture and the overall lack of rain in 2008 (Fig. 16). The habitat resources were scarce in most plant communities we studied

throughout the collection period with the exception of cottonwood habitat that is associated with a continuous water source. There was a general

trend for species to be found in a more restricted set of habitats in 2008 than in 2007 and more individuals were collected near permanent water

sources. Several species, like Amara quenseli, Nathalis iole, and discolor that were present in multiple habitats in 2007 were only

observed in a single or few habitats in 2008 (Appendix 2). In contrast, some species, like Anthophora sp., Efferia sp., Psilocurus camposi, and Amara carinata that were found in few habitats in 2007 were found in additional habitats in 2008 (Appendix 2). The presence of a species like Arphia conspersa, Melanoplus flavidus, or Philonicus limpidipennis in different habitats from 2007 demonstraites either an increase the amount of time actively searching for food and mates or that species’ ability to outcompete other species that are less drought tolerant.

Table 2: Total specimens, number of species, and diversity indices for collections from nine sites at Pinon Canyon Maneuver Site, 2008.

Total Individuals Habitat Shannon's Index Shannon's Evenness Modified Simpson's Index Species collected

JUMO/BOER 3.32 0.05 0.93 372 64 BOGR/HIJA 2.34 0.04 0.80 980 57 ARFI/YUGL2 2.68 0.04 0.86 766 60 ATCA/SPAI 2.78 0.05 0.88 490 56 ARFI/YUGL1 2.80 0.05 0.88 588 60 PODE/BRJA 2.50 0.03 0.81 1,262 80 OPIM/BOGR 2.03 0.04 0.78 746 46 GLME/FRJA 1.97 0.05 0.68 453 40 TA spp. 1.46 0.10 0.69 513 14 While diversity was relativley high in 2008, evenness was very low. High diversity with a low evenness indicated that there was still a high

number of species in the habitat and that they were relativley abundant, but the likelyhood of encountering any particular species at a single collection event or time period was low. The low evenness indicates the population was unpredicable and unstable. The juniper habitat had the highest diversity and had a more stable community of than any other habitat. Most of the habitats were comparable in diversity and evenness, except the Tamarix sp. site. It had a low diversity and a very low evenness. This indicates that most species are not consistantly using this habitat and are most likely vagrant individuals who are just passing through. The highest number of individuals were collected at the cottonwood habitat, where water was available throughout the year. Species present in the habitats were much lower when compared to 2007.

Sampling Results

Pitfall sampling was the most productive collection technique for 2008, yielding the most individuals and species. Although the diversity was slightly less than the other techniques, Shannon’s diversity (2.96-2.87) and evenness of Shannon’s diverstiy (0.078-0.026) were very similar for all collection techniques (Table 3). Evenness for all categories was less than 0.10 (low), with samples being very variable (Table 3). Surveys were still important techniques although the Asilid and Apiformes surveys were relatively unsucessful (Table 3). The low collection success was likely caused by the climatic conditions and little persistance of suitable food for these groups in any one location. The current collection techniques appear to be satisfactory and we will conduct the 2009 surveys in the same manner.

Table 3: Total specimens, number of species, and diversity indices for three collection methods at Pinon Canyon Maneuver Site, 2008.

Total Collection Shannon's Shannon's Modified individuals Species method Index Evenness Simpson's Index collected

Pitfalls Trap 5,515 112 2.87 0.026 0.89

Surveys 348 63 2.96 0.047 0.88 Malaise Trap 323 39 2.93 0.075 0.91 Conclusions

The biodiversity project at Pinon Canyon Manuever Site is continuing to add species to the species list as well as gaining valuable insight into habitat presence of a species as a preliminary indication of habitat use. Diversity remained similar between years independent of climate differences, but the ability to document all species in an area was reduced as indicated by the low evenness of Shannon’s diversity and the low abundance of species that were collected in 2008. Habitat presence of species will shift with changing climatic events. The difference in climate from year to year strongly influences the collection and analysis of results. Climatic differences confound habitat usage. Without multiple years of data collected in similar climatic years, general habitat use is difficult to determine. The third summer of collection will allow for better comparisons to be made with more accurate conclusions about a species use of a habitat.

Appendix 1: Species list with 2008 additions in bold. Pinon Canyon Maneuver Site 2008.

Orthoptera (grasshoppers, crickets, katydids)

Family Scientific Name Common Name

Acrididae Acrolophitus hirtipes Green Fool Grasshopper Acrididae Aeoloplides turnbulli Russian Thistle Grasshopper Acrididae Ageneotettix deorum White-whiskered Grasshopper Acrididae Amphitornus coloradus Striped Slant-faced Grasshopper Acrididae Arphia pseudonietana Northwestern Red-Winged Grasshopper Acrididae Arphia conspersa Speckle-winged Rangeland Grasshopper Acrididae Arphia simplex Plains Yellow-winged Grasshopper Acrididae Aulocara elliotti Big Headed Grasshopper Acrididae Aulocara femoratum White Cross Grasshopper Acrididae Boopedon gracile Graceful Range Grasshopper Acrididae Boopedon nubilum Ebony Grasshopper Acrididae Camnula pellucida Clear-winged Grasshopper Acrididae Chloealtis abdominalis Thomas's Broad-Winged Grasshopper Acrididae Chloealtis conspersa Sprinkled Broad-Winged Grasshopper Acrididae Chorthippus curtipennis Marsh Meadow Grasshopper Acrididae Chortophaga viridifasciata Northern Green-striped Grasshopper Acrididae Circotettix rabula Wrangler Grasshopper Acrididae Crenulated Grasshopper Acrididae Spotted Winged Grasshopper Acrididae Dactylotum bicolor Pictured Grasshopper Acrididae Derotmema haydeni Hayden’s Grasshopper Acrididae Eritettix simplex Velvet Striped Grasshopper Acrididae Hadrotettix magnificus Magnificent Grasshopper Acrididae Hesperotettix speciosus Western Grass-Green Grasshopper Acrididae Hadrotettix trifasciatus Three Banded Range Grasshopper Acrididae Hesperotettix viridis Snakeweed Grasshopper Acrididae Hippiscus ocelote Wrinkled Grasshopper Acrididae Hippopedon capito Apache Grasshopper Acrididae Hypochlora alba Cudweed Grasshopper Acrididae Leprus cyaneus Blue-Winged Grasshopper Acrididae Leprus robustus Acrididae Melanoplus spp Spurthroated Grasshoppers Acrididae Melanoplus angustipennis Narrow-winged Sand Grasshopper Acrididae Melanoplus arizonae Arizona Spur-throated Grasshopper Acrididae Melanoplus bivittatus Two-Striped Grasshopper Acrididae Melanoplus borealis Northern Grasshopper

Acrididae Melanoplus bowditchi Sagebrush Grasshopper Acrididae Melanoplus confusus Little Pasture Spurthroated Grasshopper Acrididae Melanoplus differentialis Differential Grasshopper Acrididae Melanoplus discolor Contrasting Spurthroated Grasshopper Acrididae Melanoplus dodgei Dodge's Spurthroated Grasshopper Acrididae Melanoplus fasciatus Huckleberry Spur-Throat Grasshopper

Orthoptera (grasshoppers, crickets, katydids) Acrididae Melanoplus flavidus Yellowish Spurthroated Grasshopper Acrididae Melanoplus femurrubrum Red-legged Grasshopper Acrididae Melanoplus foedus Striped Sand Grasshopper Acrididae Melanoplus gladstoni Gladston’s Spurthroated Grasshopper Acrididae Melanoplus glaucipes Glaucous-Legged Spurthroated Grasshopper Acrididae Melanoplus lakinus Lakin Grasshopper Acrididae Melanoplus occidentalis Flabellate Grasshopper Acrididae Melanoplus packardii Packard Grasshopper Acrididae Melanoplus ponderosus Ponderous Spurthroated Grasshopper Acrididae Melanoplus regalis Regal Spurthroated Grasshopper Acrididae Melanoplus sanguinipes Migratory Grasshopper Acrididae Melanoplus splendidus Splendid Grasshopper Acrididae Melanoplus tristis Acrididae Melanoplus yarrowii Yarrow's Spurthroated Grasshopper Acrididae Mermiria bivittata Two-striped Mermiria Grasshopper Acrididae Mermiria picta Painted Slantfaced Grasshopper Acrididae Mestobregma plattei Platt Range Grasshopper Acrididae Metator pardalinus Pard Grasshopper Acrididae Opeia obscura Obscure Grasshopper Acrididae Orphulella speciosa Pasture Grasshopper Acrididae Pardalophora apiculata Coral-winged Grasshopper Acrididae Paropomala wyomingensis Wyoming Toothpick Grasshopper Acrididae Phlibostroma quadrimaculatum Four-spotted Grasshopper Acrididae Phoetaliotes nebrascensis Large-headed Grasshopper Acrididae Psoloessa delicatula Brown-Spotted Range Grasshopper Acrididae Psoloessa texana Texas Spotted Range Grasshopper Acrididae Pseudopomala brachyptera Big Slender Grasshopper Acrididae Schistocerca obscura Obscure Bird Grasshopper Acrididae Spharagemon collare Mottled Sand Grasshopper Acrididae Spharagemon equale Orange-legged Grasshopper Acrididae Stethophyma gracile Graceful Sedge Grasshopper Acrididae Syrbula admirabilis Admirable Grasshopper Acrididae Syrbula montezuma Slantfaced Grasshopper

Acrididae Trachyrhachys coronata Crowned Grasshopper Acrididae Trachyrhachys kiowa Kiowa range Grasshopper Acrididae Trimerotropis sp Slender Grasshopper Acrididae Trimerotropis gracilis Thomas' Slender Grasshopper Acrididae Trimerotropis latifasciata Broad Banded Grasshopper Acrididae Trimerotropis melanoptera Black-winged Grasshopper Acrididae Trimerotropis pallidipennis Pallid-winged Grasshopper Acrididae Tropidolophus formosus Great Crested Grasshopper Acrididae Xanthippus corallipes Red Shanked Grasshopper Gryllidae Acheta domesticus House Cricket Gryllidae Allonemobius fasciatus Striped Ground Cricket Gryllidae Cycloptilum sp Gryllidae Eunemobius sp Carolina Ground Cricket Gryllidae Gryllus pennsylvanicus Fall Field Cricket (Black field cricket) Oedipodinae Encoptolophus costalis Dusky Grasshopper Rhaphidophoridae Ceuthophilus spp Camel cricket Orthoptera (grasshoppers, crickets, katydids)

Romaleidae Brachystola magna Plains Lubber Grasshopper Tetrigidae Tetrix ornata Ornate Pygmy Grasshopper Tettigoniidae Microcentrum rhomhbifolium Angle-Winged Katydid Tettigoniidae Pseudophyllinae sp Katydid Tettigoniidae Scudderia furcata Fork-tailed Bush Katydid

Odonata (dragonflies and damselflies)

Family Scientific Name Common Name Calopterygidae Broad-Winged Damselflies Coenagrionidae Pond Damselflies Cordulidae Emeralds Gomphidae Clubtail Dragonflies Libellulidae Libellula pulchella Twelve Spotted Skimmer Libellulidae Tramea lacerata Black Saddlebags

Neuroptera

Family Scientific Name Common Name Chrysopidae Lacewing Mantispidae Mantis Myrmeleontidae Ant lion

Coleoptera ()

Family Scientific Name Common Name Anthicidae Notoxus sp Ant-like Flower Buprestidae Metallic Wood Boring Beetles Cantharidae Cantharis sp Soldier Beetles Carabidae Agonum cyclifer Carabidae Agonum decorum Ground Beetle Carabidae Agonum extensicolle Ground Beetle Carabidae Agonum placidum Ground Beetle Carabidae Agonum punctiforme Ground Beetle Carabidae Amara sp Ground Beetle Carabidae Amara carinata Ground Beetle Carabidae Amara deparca Ground Beetle Carabidae Amara quenseli Ground Beetle Carabidae Amara thoracica Ground Beetle Carabidae Amblycheila cylindriformis Great Plains Giant Tiger Beetle Carabidae Badister obtusus Ground Beetle Carabidae Brachinus sp Ground Beetle Carabidae Brachinus phaeocerus Ground Beetle Carabidae Calathus sp Ground Beetle Carabidae Calathus opaculus Ground Beetle Carabidae Calosoma sp Ground Beetle Carabidae Calosoma affine Ground Beetle Carabidae Calosoma marginake Ground Beetle Coleoptera (beetles)

Carabidae Calosoma peregrinator Ground Beetle Carabidae Calosoma scrutator Ground Beetle Carabidae Chlaenius tricolor Vivid metallic ground beetles Carabidae Chlaenius s sericeus Ground Beetle Carabidae Chlaenius t tomentosus Ground Beetle Carabidae Cicindela sp Ground Beetle Carabidae Cicindela belfragei Loamy-ground Tiger Beetle Carabidae Cicindela formosa Big Sand Tiger Beetle Carabidae Cicindela n nigrocoerulea Ground Beetle Carabidae Cicindela o obsoleta Ground Beetle Carabidae Cicindela p punctulata Ground Beetle Carabidae Cicindela s scutellaris Festive Tiger Beetle Carabidae Cratacanthus dubius Ground Beetle Carabidae Cyclotrachelus constrictus Ground Beetle Carabidae Cyclotrachelus substriatus Ground Beetle Carabidae Cymindis interior Ground Beetle

Carabidae Cymindis planipennis Ground Beetle Carabidae Cymindis punctiger Ground Beetle Carabidae Dicaelus l laevipennis Ground Beetle Carabidae Diplocheila obtusa Ground Beetle Carabidae Euryderus sp Ground Beetle Carabidae Euryderus grossus Ground Beetle Carabidae Galerita sp Ground Beetle Carabidae Galerita janus Ground Beetle Carabidae Harpalus sp Ground Beetle Carabidae Harpalus caliginosus Ground Beetle Carabidae Harpalus fraternus Ground Beetle Carabidae Harpalus paratus Ground Beetle Carabidae Harpalus pensylvanicus Ground Beetle Carabidae Hartonymus alternatus Ground Beetle Carabidae Lebia viridis Ground Beetle Carabidae Micrixys distincta Ground Beetle Carabidae Pasimachus californicus Ground Beetle Carabidae Pasimachus elongatus Ground Beetle Carabidae Piosoma setosum Ground Beetle Carabidae sp Ground Beetle Carabidae Poecilus chalcites Ground Beetle Carabidae Poecilus cyanicolor Ground Beetle Carabidae Poecilus l lucublandus Ground Beetle Carabidae Poecilus scitulus Ground Beetle Carabidae Pterostichus commutabilis Ground Beetle Carabidae Rhadine dissecta sp Ground Beetle Carabidae Scarites subterraneus Big Headed Ground Beetle Carabidae Scarites quadriceps Ground Beetle Cerambycidae Oberea sp Long-horned Wood-boring Beetles Cerambycidae Prionus sp Long-horned Wood-boring Beetles Chrysomelidae Calligrapha sp Leaf Beetles Chrysomelidae Diabrotica sp Leaf Beetles Coleoptera (beetles)

Cleridae Checkered Flower Beetles Anatis lecontei Large Ladybeetle Coccinellidae Coccinella septempunctata Seven-spotted Ladybeetle Coccinellidae Hippodamia parenthesis Coccinellidae Hippodamia convergens Convergent Ladybeetle Coccinellidae Myzia interrupta Curculiondiae Weevils Dytiscidae Predacious Diving Beetles

Elateridae Click Beetles Histeridae Clown Beetles Lampyridae Fireflies Meloidae Epicauta sp Blister Beetles Nitidulidae Capophilus sp Sap Beetles Scarabidae Aphodices sp Scarabidae Canthon sp Dung Beetle Scarabidae Eurphoria inda Scarabidae Melanocanthon sp Dung Beetle Sciritidae Marsh Beetles Silphidae Nicrophorus sp Burying Beetles Silphidae Silpha sp Carrion Beetles Staphylinidae Aleochara sp Rove Beetles Tenebrionidae Diaperis sp Darkling Beetles Tenebrionidae Eleodes sp Darkling Beetles Trogidae Hide Beetles

Lepidoptera (butterflies and )

Family Scientific Name Common Name Arctiidae Grammia sp Tiger Moths Arctiidae Hyproprea sp Geometridae Geometrid Moths Hesperiidae Amblyscirtes vialis Common Roadside Hesperiidae Hesperia pahaska Pahaska Skipper Hesperiidae Hesperia uncas Uncas Skipper Hesperiidae Hesperopsis alpheus Saltbush Sootywing Hesperiidae Megathymus streckeri Strecker's Giant Skipper Hesperiidae Pholisora catullus Common Sootywing Hesperiidae Pyrgus communis Common Checkered Skipper Hesperiidae Pyrgus scriptura Small Checkered Skipper Hesperiidae Thorybes pylades Northern Cloudywing Brephidium exile Western Pygmy Blue Lycaenidae Callophrys gryneas Juniper Hairstreak Lycaenidae Euphilotes rita Rita Blue Lycaenidae Hemiargus isola Reakirt’s Blue Lycaenidae Leptotes marina Marine Blue Lycaenidae Plebejus acmon Acmon Blue Lycaenidae Plebejus saepiolus Greenish Blue (butterflies and moths)

Lycaenidae Strymon melinus Grey Hairstreak Noctuidae Millers or Owlet Moths Noctuidae Alypia octomaculata Eight-spotted Forester Anaea andria Goatweed Leafwing Nymphalidae Cercyonis oetus Small Wood Nymph Nymphalidae Cercyonis pegala Common Wood Nymph Nymphalidae Danaus plexippus Monarch Nymphalidae Euptoieta claudia Variegated Fritillary Nymphalidae Junonia coenia Common Buckeye Nymphalidae Limenitis archippus Viceroy Nymphalidae Nymphalis antiopa Mourning Cloak Nymphalidae Phyciodes campestris Field Crescent Nymphalidae Mylitta Crescent Nymphalidae Phyciodes pallida Pale Crescent Nymphalidae Phyciodes picta Painted Crescent Nymphalidae Phyciodes tharos Pearl Crescent Nymphalidae Polygonia interrogationis Question Mark Nymphalidae Speyeria coronis Coronis Fritillary Nymphalidae Speyeria edwardsii Edward's Fritillary Nymphalidae Thessalia fulvia Fulvia Checkerspot Nymphalidae Vanessa atalanta Red Admiral Nymphalidae Vanessa cardui Painted Lady Nymphalidae Vanessa virginiensis American Lady Papilionidae Papilio mulitcaudata Two-tailed Swallowtail Papilionidae Papilio polyxenes Black Swallowtail Papilionidae Papilio rutulus Western Tiger Swallowtail Pieridae Colias cesonia Southern Dogface Pieridae Colias eurytheme Orange Sulphur Pieridae Colias philodice Clouded Sulphur Pieridae Eurema nicippe Sleepy Orange Pieridae Nathalis iole Dainty Sulphur Pieridae Pieris rapae Cabbage White Pieridae Pontia occidentalis Western White Pieridae Pontia protodice Checkered White Pyralidae Pyralid Moth Hyles lineata White Lined Sphinx Moth Sphingidae juanita Juanita Sphinx Moth

Diptera ()

Family Scientific Name Common Name Asilidae Albibarbefferia sp Robber Flies

Asilidae Albibarbefferia albibarbis Robber Flies Asilidae Albibarbefferia bicolor Robber Flies Asilidae Albibarbefferia leucocoma Robber Flies Asilidae Aridefferia prattii Robber Flies Asilidae Aridefferia subarida Robber Flies Asilidae Asilus auriannulatus Robber Flies Diptera (flies)

Asilidae Asilus sp Robber Flies Asilidae Atomosia sp Robber Flies Asilidae Atomosia melanopogon Robber Flies Asilidae Atomosia nuda Robber Flies Asilidae Atomosia puella Robber Flies Asilidae Carinefferia costalis Robber Flies Asilidae Diogmites sp Robber Flies Asilidae Diogmites neoternatus Robber Flies Asilidae "Efferia" spp Robber Flies Asilidae Efferia anomala Robber Flies Asilidae Efferia apache Robber Flies Asilidae Efferia bryanti Robber Flies Asilidae Efferia duncani Robber Flies Asilidae Laphystia varipes Robber Flies Asilidae Lasiopogon sp Robber Flies Asilidae Lasiopogon quadrivittatus Robber Flies Asilidae Leptogaster sp Robber Flies Asilidae Leptogaster brevicornis Robber Flies Asilidae Leptogaster weslacensis Robber Flies Asilidae Nerax beameri Robber Flies Asilidae Ospriocerus sp Robber Flies Asilidae Ospriocerus abdominalis Robber Flies Asilidae Philonicus sp Robber Flies Asilidae Philonicus arizonensis Robber Flies Asilidae Philonicus limpidipennis Robber Flies Asilidae Pogoniefferia sp Robber Flies Asilidae Pogoniefferia bicaudata Robber Flies Asilidae Pogoniefferia helenae Robber Flies Asilidae Pogoniefferia texana Robber Flies Asilidae Pogonosoma sp Robber Flies Asilidae Proctacanthella exquisita Robber Flies Asilidae Proctacanthus sp Robber Flies Asilidae Promachus sp Robber Flies Asilidae Psilocurus sp Robber Flies

Asilidae Psilocurus arizonensis Robber Flies Asilidae Psilocurus birdi Robber Flies Asilidae Psilocurus camposi Robber Flies Asilidae Stenopogon sp Robber Flies Asilidae Stichopogon sp Robber Flies Culicidae Mosquitoes Muscidae Musca domestica House Fly Mydidae sp Mydas Fly

Scorpiones

Family Scientific Name Common Name Buthidae Centruroides vittatus Scorpion

Hymenoptera (bees) Family Scientific Name Common Name confederata Andrenidae Andrena helianthi Andrenidae Andrena merriami Andrenidae Perdita spp Andrenidae Protandrena spp Anthophora spp Apidae Bombus spp Bumblebee Apidae diminuta Apidae Diadasia enavata Apidae Diadasia olivacea Apidae Diadasia rinconis rinconis Apidae Eucera spp Apidae Holcopasites tegularis Apidae Melecta sp Apidae Melissodes intorta Apidae Melissodes spp Apidae Neolarra verbesinae Apidae Nomada sp Apidae Svastra obliqua Apidae Tetralphiella sp Apidae Triepeolus sp Apidae Xenoglossa sp Colletidae Colletes spp Digger Bee Halictidae Agapostemon angelicus Halictidae Agapostemon angelicus or texanus

Halictidae Agapostemon sp Halictidae Augochlora sp Halictidae Augochloropsis sp Halictidae Dieunomia sp Halictidae Halictus ligatus Halictidae Halictus virgatellus or provancheri Halictidae Lasioglossum spp Halictidae Nomia sp Alkali Bee Halictidae Sphecodes sp Megachilidae Athidium sp Megachilidae Dianthidium spp Megachilidae Megachile amica Megachilidae Megachile brevis Megachilidae Megachile brevis onobrychidis Megachilidae Osmia spp Braconidae Formicidae Ants Ichneumonidae Ichneumonid Mutilidae Dasymatina sp Velvet Ants Pompilidae Pepsis sp Wasps Sphecidae Thread-wasted Vespidae Polistes sp Paper Wasp Vespidae Vespula sp Appendix 2: Presence or absence of a species for each year. Black indicates the 2007 occurrence and 2008 is indicated with grey. Pinon Canyon Maneuver Site 2008. Carabidae ARFI/ ATCA/ BOGR/ GLME/ JUMO/ OPIM/ PIPO/ PODE/ Species YUGL SPAI HIJA FRJA BOER BOGR SONU BRJA TA sp. Agonum cyclifer Agonum decorum Agonum extensicolle Agonum placidum Amara carinata Amara deparca Amara quenseli Amara thoracica Amblycheila cylindriformis Badister obtusus Brachinus phaeocerus

Carabidae ARFI/ ATCA/ BOGR/ GLME/ JUMO/ OPIM/ PIPO/ PODE/ Species YUGL SPAI HIJA FRJA BOER BOGR SONU BRJA TA sp. Calathus opaculus Calosoma affine Calosoma scrutator Chlaenius s. sericeus Chlaenius t. tomentosus Chlaenius tricolor Cicindela n. nigrocoerulea Cicindela o. obsoleta Cicindela obsoleta Cicindela p. punctulata Cicindela s. scutellaris Cicindela scutellaris Cratacanthus dubius Cyclotrachelus constrictus Cyclotrachelus substriatus Cymindis interior Cymindis planipennis Cymindis punctiger Dicaelus l. laevipennis Diplocheila obtusa Euryderus grossus Galerita janus Harpalus caliginosus Harpalus fraternus Harpalus paratus Harpalus pensylvanicus Lebia viridis Micrixys distincta Pasimachus californicus Pasimachus elongatus Piosoma setosum Poecilus chalcites

Carabidae ARFI/ ATCA/ BOGR/ GLME/ JUMO/ OPIM/ PIPO/ PODE/ Species YUGL SPAI HIJA FRJA BOER BOGR SONU BRJA TA sp. Poecilus cyanicolor Poecilus l. lucublandus Poecilus lucublandus Poecilus scitulus Pterostichus commutabilis Rhadine dissecta sp. Scarites subterraneus

Lepidoptera ARFI/ ATCA/ BOGR/ GLME/ JUMO/ OPIM/ PIPO/ PODE/ Species YUGL SPAI HIJA FRJA BOER BOGR SONU BRJA TA sp. Amblyscirtes vialis Anaea andria Brephidium exile Callophrys gryneas Cercyonis oetus Cercyonis pegala Colias cesonia Colias eurytheme Colias philodice Danaus plexippus Euphilotes rita Euptoieta claudia Eurema nicippe Hemiargus isola Hesperia pahaska Hesperia uncas Hesperopsis alpheus Leptotes marina Limenitis archippus Lycaeides melissa

Lepidoptera ARFI/ ATCA/ BOGR/ GLME/ JUMO/ OPIM/ PIPO/ PODE/ Species YUGL SPAI HIJA FRJA BOER BOGR SONU BRJA TA sp. Megathymus streckeri Nathalis iole Nymphalis antiopa Papilio multicaudata Papilio polyxenes Papilio rutulus Pholisora catullus Phyciodes campestris Phyciodes mylitta Phyciodes pallida Phyciodes picta Phyciodes tharos Pieris rapae Plebejus acmon Plebejus saepiolus Polygonia interrogationis Pontia occidentalis Pontia protodice Pyrgus communis Pyrgus scriptura Speyeria coronis Speyeria edwardsii Strymon melinus Thessalia fulvia Thorybes pylades Vanessa cardui Vanessa virginiensis

Orthoptera ARFI/ ATCA/ BOGR/ GLME/ JUMO/ OPIM/ PIPO/ PODE/ Scientific Names YUGL SPAI HIJA FRJA BOER BOGR SONU BRJA TA sp. Orthoptera ARFI/ ATCA/ BOGR/ GLME/ JUMO/ OPIM/ PIPO/ PODE/ Scientific Names YUGL SPAI HIJA FRJA BOER BOGR SONU BRJA TA sp. Acheta domesticus Acrolophitus hirtipes Aeoloplides turnbulli Ageneotettix deorum Amphitornus coloradus Arphia conspersa Arphia pseudonietana Arphia simplex Aulocara elliotti Aulocara femoratum Boopedon gracile Boopedon nubilum Brachystola magna Cactylotum bicolor Camnula pellucida Ceuthophilus sp. Chloeatis abdominalis Choealtis conspersa Chorthippus curtipennis Chortophaga viridifasciata Circotettix rabula Cordillacris crenulata Cordillacris occipitalis Cycloptilum sp. Dactylotum bicolor Derotmema haydeni Encoptolophus costalis Eritettix simplex Eunemobius sp. Gryllus pennsylvanicus Hadrotettix magnificus Hadrotettix trifasciatus

Orthoptera ARFI/ ATCA/ BOGR/ GLME/ JUMO/ OPIM/ PIPO/ PODE/ Scientific Names YUGL SPAI HIJA FRJA BOER BOGR SONU BRJA TA sp. Hesperotettix speciosus Hesperotettix viridis Hesperotettix wyomingensis Hippiscus ocelote Hippopedon capito Hypochlora alba Melanoplus angustipennis Melanoplus arizonae Melanoplus bivittatus Melanoplus borealis Melanoplus bowditchi Melanoplus confusus Melanoplus differentialis Melanoplus discolor Melanoplus dodgei Melanoplus fasciatus Melanoplus femurrubrum Melanoplus flavidus Melanoplus foedus Melanoplus gladstoni Melanoplus glaucipes Melanoplus lakinus Melanoplus occidentalis Melanoplus packardii Melanoplus ponderosus Melanoplus regalis Melanoplus sanguinipes Melanoplus sp. Melanoplus splendidus Melanoplus tristis Melanoplus yarrowii Mermiria bivittata

Orthoptera ARFI/ ATCA/ BOGR/ GLME/ JUMO/ OPIM/ PIPO/ PODE/ Scientific Names YUGL SPAI HIJA FRJA BOER BOGR SONU BRJA TA sp. Mermiria picta Mestobregma plattei Metator pardalinus Microentrum sp. Opeia obscura Orphulella speciosa Pardalophora apiculata Paropomala wyomingensis Phlibostroma quadrimaculatum Phoetaliotes nebrascensis Pseudophyllinae sp. Pseudopomala brachyptera Psoloessa delicatula Psoloessa texana Rhaphidophoridae sp. Schistocerca obscura Scudderia furcata Spharagemon collare Spharagemon equale Stethophyma gracile Syrbula admirabilis Syrbula montezuma Tetrix ornata Trachyrhachys coronata Trachyrhachys kiowa Trimerotropis gracilis Trimerotropis latifasciata Trimerotropis melanoptera Trimerotropis pallidipennis Trimerotropis sp. Tropidolophus formosus

Orthoptera ARFI/ ATCA/ BOGR/ GLME/ JUMO/ OPIM/ PIPO/ PODE/ Scientific Names YUGL SPAI HIJA FRJA BOER BOGR SONU BRJA TA sp. Xanthippus corallipes

Hymenoptera ARFI/ ATCA/ BOGR/ GLME/ JUMO/ OPIM/ PIPO/ PODE/ Species YUGL SPAI HIJA FRJA BOER BOGR SONU BRJA TA sp. Agapostemon angelicus Agapostemon angelicus or texanus Agapostemon sp. Andrena confederata Andrena helianthi Andrena merriami Anthophora sp. 1 Anthophora sp. 2 Anthophora sp. 3 Athidium sp. Augochlora sp. Augochloropsis sp. Bombus sp. 1 Bombus spp. Colletes spp. Diadasia diminuta Diadasia enavata Diadasia olivacea Diadasia rinconis rinconis Dianthidium sp. 1 Dianthidium sp. 2 Dieunomia sp. Eucera sp. Eucera sp. 1 Eucera sp. 2

Hymenoptera ARFI/ ATCA/ BOGR/ GLME/ JUMO/ OPIM/ PIPO/ PODE/ Species YUGL SPAI HIJA FRJA BOER BOGR SONU BRJA TA sp. Halictus ligatus Halictus virgatellus or provancheri Holcopasites tegularis Lasioglossum spp. 1 Lasioglossum spp. 2 Megachile amica Megachile brevis Megachile brevis onobrychidis Melecta sp. Melissodes intorta Melissodes sp. 1 Melissodes sp. 2 Neolarra verbesinae Nomia sp. Osmia spp. Perdita spp. Protandrena sp. 1 Protandrena sp. 2 Sphecodes sp. Svastra obliqua Tetralphiella sp. Triepeolus sp. Xenoglossa sp.

Asilidae

ARFI/ PIPO/SON Species YUGL ATCA/SPAI BOGR/HIJA GLME/FRJA JUMO/BOER OPIM/BOGR U PODE/BRJA

"Efferia" spp.

Albibarbefferia albibarbis

Albibarbefferia bicolor

Albibarbefferia leucocoma

Albibarbefferia spp.

Aridefferia prattii

Aridefferia subarida 1

Asilus auriannulatus

Atomosia melanopogon

Atomosia nuda 4

Atomosia puella

Carinefferia costalis

Diogmites neoternatus

Efferia candida

Efferia spp.

Haplopogon sp.

Laphystia varipes

Lasiopogon quadrivittatus

Leptogaster brevicornis

Leptogaster weslacensis

Nerax beameri

Ospriocerus abdominalis

Philonicus arizonensis 1

Philonicus limpidipennis 6

Pogoniefferia bicaudata

Pogoniefferia helenae Asilidae ARFI/ JUMO/BOE PIPO/SON PODE/ Species YUGL ATCA/SPAI BOGR/HIJA GLME/FRJA R OPIM/BOGR U BRJA

Pogoniefferia texana

Proctacanthella exquisita

Psilocurus birdi

Psilocurus camposi

Scleropogon arizonensis

Stenopogon albibasis

Stichopogon sp. 1