The Pennsylvania State University the Graduate School College of the Liberal Arts

Total Page:16

File Type:pdf, Size:1020Kb

The Pennsylvania State University the Graduate School College of the Liberal Arts The Pennsylvania State University The Graduate School College of the Liberal Arts THE EL GIGANTE ROCK SHELTER, HONDURAS A Thesis in Anthropology by Timothy E. Scheffler © 2008 Timothy E. Scheffler Submitted in Partial Fulfillment of the Requirements for the Degree of Doctor of Philosophy May 2008 The thesis of Timothy E. Scheffler has been reviewed and approved* by the following: Kenneth G. Hirth Professor of Archaeological Anthropology Thesis Advisor Chair of Committee Lee Ann Newsom Associate Professor of Archaeological Anthropology William Sanders Professor Emeritus of Archaeological Anthropology Alfred Traverse Professor of Emeritus of Palynology David Webster Professor of Archaeological Anthropology Nina Jablonski Professor of Anthropology Head of the Department of Anthropology *Signatures on file in the Graduate School ii Abstract: The site of El Gigante, La Paz, Honduras is unique in Central America for its very well preserved organic remains and lengthy archaeological sequence. Preliminary analyses of the ceramics, lithics, botanical and faunal material are presented in a cultural sequence beginning at 9,480 B.C. The botanical remains recovered from the site are inventoried and several species are subjected to preliminary morphometric analyses in order to characterize the assemblage, and assess changes in plant phenotypes through time. The operation of "directional" (artificial) selection is evident at the site for several species, including avocado (Persea americana), squash (Cucurbita sp., cf. C. pepo), and bottle gourd (Lagenaria siceraria). While varieties of squashes may have been cultivated in the Archaic, clearly domesticated maize and beans appear to have been introduced in the Formative. However, stratigraphic disconformities in the period of c. 5,000 – 2,000 B.C., may bias this conclusion. From early in the sequence, botanical diversity is high, despite changing subsistence specialization and generalization evident in other artifact assemblages. Many undomesticated species are utilized prehistorically and are not lost from the diet despite the intensification of field-based food production of domesticated species. An emphasis is placed on suites of plants, both wild and domesticated, annual and perennial, whose use is integrated through time El Gigante. Together these subsistence systems afforded flexible, energetically efficient and risk minimizing choices to prehistoric foragers. Low level food production is evident as early as the Archaic period. Tree crops were one focus of subsistence practices. The augmentation and management of perennials is inferred from their consistency and prominence in the archaeological record. However, there is also evidence for the use of annual grasses during this time. The data invites further investigation into the landscape level paleoecological past around El Gigante. iii Table of Contents List of Figures................................................................................................................... vii List of Tables ..................................................................................................................... ix List of Photographs.............................................................................................................x Acknowledgements........................................................................................................... xii CHAPTER 1: THE SITE OF EL GIGANTE .................................................................... 1 Introduction ..................................................................................................................... 1 The Discovery of El Gigante........................................................................................... 3 The Current Project ......................................................................................................... 5 Dissertation Goals............................................................................................................ 6 CHAPTER 2: THE ENVIRONMENTAL CONTEXT OF THE EL GIGANTE SITE, PAST AND PRESENT..................................................................................................... 21 Environment and subsistence ........................................................................................ 21 The climate and soils of highland Honduras ................................................................. 23 Regional Geology.......................................................................................................... 24 Relevant Climate Studies............................................................................................... 27 Vegetation history and anthropogenic disturbance ....................................................... 31 The Modern Vegetation Community............................................................................. 34 Conclusions ................................................................................................................... 37 CHAPTER 3: EXCAVATION AND STRATIGRAPHY............................................... 49 Introduction and Methods.............................................................................................. 49 Stratigraphy ................................................................................................................... 56 Site Concordance........................................................................................................ 63 CHAPTER 4: THE EXCAVATION RESULTS............................................................. 85 Radiocarbon Dating and the Cultural Sequence............................................................ 85 Disturbances and Gaps in the Archaeological Record .................................................. 89 Feature Descriptions...................................................................................................... 92 Feature Summary ..................................................................................................... 102 Sedimentary Analyses ................................................................................................. 104 Ash Comparison....................................................................................................... 104 Soil Carbon Content................................................................................................. 105 Fire Cracked Rock.................................................................................................... 106 Conclusions ................................................................................................................. 107 CHAPTER 5: MATERIAL CULTURE AND FAUNAL REMAINS .......................... 121 Ceramics ...................................................................................................................... 121 Flaked lithic artifacts ................................................................................................... 125 Projectile Points........................................................................................................ 131 Ground Stone Artifacts............................................................................................. 134 Faunal Remains ........................................................................................................... 137 Declining Large Game through the Archaic ............................................................ 139 Pleistocene Fauna..................................................................................................... 143 Human Skeletal Remains............................................................................................. 145 Worked bone and shell ................................................................................................ 146 Textiles ........................................................................................................................ 147 Pictographs .................................................................................................................. 150 iv CHAPTER 6: THE PLANT REMAINS........................................................................ 168 Introduction to the Assemblage................................................................................... 168 Macrobotanical methods.............................................................................................. 174 Macrobotanical Results ............................................................................................... 178 Esperanza Phase Remains ........................................................................................ 178 Agave/Maguey ...................................................................................................... 178 Hog Plum/Ciruela................................................................................................. 181 Sapotaceae............................................................................................................. 187 Avocado ................................................................................................................ 191 Palm ...................................................................................................................... 194 Wild beans ............................................................................................................ 196 Acorns................................................................................................................... 197 Other
Recommended publications
  • New Lives for Ancient and Extinct Crops
    Book Review New Lives for Ancient and Extinct Crops Edited by Paul E. Minnis. 2014. The University of Arizona Press, Tucson. 288 pp. $65.00 (hardcover). ISBN: 978-0-8165-3062- 5. Reviewed by Thomas C. Hart Reviewer address: Department of Anthropology, University of Texas, 2201 Speedway Stop C3200, SAC 4.102, Austin, TX 78712, USA. Email: [email protected] Received: February 8, 2015 Volume: 6(1):116-118 Published: August 19, 2015 © 2015 Society of Ethnobiology In this edited volume, Paul E. Minnis and the chapter In the first chapter, Gayle J. Fritz describes how authors successfully illustrate how archaeological, maygrass (Phalaris caroliniana Walter Poaceae.) was an ethnohistorical, and ethnobotanical data can be important North American grass that was a part of effectively synthesized to provide a detailed account the Eastern Agricultural Complex for at least 3,000 of how ancient and extinct crops were used, as well as years. The earliest evidence for maygrass is found the potential they hold for diversifying global food during the Late Archaic in Illinois, Tennessee, and stocks. The authors explore the ancient uses and Kentucky. Eventually this crop spread out of its contemporary large scale agricultural potentials of native range and encompassed an area from Wiscon- maygrass, goosefoot, sumpweed or marshelder, agave, sin and Pennsylvania, in the north, to Texas and little barley grass, chia, arrowroot, leren, and sama (or Georgia, in the south. This very small seeded grass bitter vetch). Minnis suggests that by looking to the was an important component of the ritual feasts that past, researchers can “pre-screen” species in terms of took place at Cahokia, as evidenced by the abundant looking for genetic material from ancient taxa that remains recovered from sub-Mound 51.
    [Show full text]
  • Principles and Practice of Forest Landscape Restoration Case Studies from the Drylands of Latin America Edited by A.C
    Principles and Practice of Forest Landscape Restoration Case studies from the drylands of Latin America Edited by A.C. Newton and N. Tejedor About IUCN IUCN, International Union for Conservation of Nature, helps the world find pragmatic solutions to our most pressing environment and development challenges. IUCN works on biodiversity, climate change, energy, human livelihoods and greening the world economy by supporting scientific research, managing field projects all over the world, and bringing governments, NGOs, the UN and companies together to develop policy, laws and best practice. IUCN is the world’s oldest and largest global environmental organization, with more than 1,000 government and NGO members and almost 11,000 volunteer experts in some 160 countries. IUCN’s work is supported by over 1,000 staff in 60 offices and hundreds of partners in public, NGO and private sectors around the world. www.iucn.org Principles and Practice of Forest Landscape Restoration Case studies from the drylands of Latin America Principles and Practice of Forest Landscape Restoration Case studies from the drylands of Latin America Edited by A.C. Newton and N. Tejedor This book is dedicated to the memory of Margarito Sánchez Carrada, a student who worked on the research project described in these pages. The designation of geographical entities in this book, and the presentation of the material, do not imply the expression of any opinion whatsoever on the part of IUCN or the European Commission concerning the legal status of any country, territory, or area, or of its authorities, or concerning the delimitation of its frontiers or boundaries.
    [Show full text]
  • A New Record of Domesticated Little Barley (Hordeum Pusillum Nutt.) in Colorado: Travel, Trade, Or Independent Domestication
    UC Davis UC Davis Previously Published Works Title A New Record of Domesticated Little Barley (Hordeum pusillum Nutt.) in Colorado: Travel, Trade, or Independent Domestication Permalink https://escholarship.org/uc/item/1v84t8z1 Journal KIVA, 83(4) ISSN 0023-1940 Authors Graham, AF Adams, KR Smith, SJ et al. Publication Date 2017-10-02 DOI 10.1080/00231940.2017.1376261 Peer reviewed eScholarship.org Powered by the California Digital Library University of California KIVA Journal of Southwestern Anthropology and History ISSN: 0023-1940 (Print) 2051-6177 (Online) Journal homepage: http://www.tandfonline.com/loi/ykiv20 A New Record of Domesticated Little Barley (Hordeum pusillum Nutt.) in Colorado: Travel, Trade, or Independent Domestication Anna F. Graham, Karen R. Adams, Susan J. Smith & Terence M. Murphy To cite this article: Anna F. Graham, Karen R. Adams, Susan J. Smith & Terence M. Murphy (2017): A New Record of Domesticated Little Barley (Hordeum pusillum Nutt.) in Colorado: Travel, Trade, or Independent Domestication, KIVA, DOI: 10.1080/00231940.2017.1376261 To link to this article: http://dx.doi.org/10.1080/00231940.2017.1376261 View supplementary material Published online: 12 Oct 2017. Submit your article to this journal View related articles View Crossmark data Full Terms & Conditions of access and use can be found at http://www.tandfonline.com/action/journalInformation?journalCode=ykiv20 Download by: [184.99.134.102] Date: 12 October 2017, At: 06:14 kiva, 2017, 1–29 A New Record of Domesticated Little Barley (Hordeum pusillum Nutt.) in Colorado: Travel, Trade, or Independent Domestication Anna F. Graham1, Karen R. Adams2, Susan J. Smith3, and Terence M.
    [Show full text]
  • Noxious and Rangeland Weed Management: Getting the Most out of Spraying Mark Pederson Dow Agrosciences [email protected]
    Noxious and Rangeland Weed Management: Getting the Most Out of Spraying Mark Pederson Dow AgroSciences [email protected] DOW RESTRICTED - For internal use only Thanks to Pend Oreille County. DOW RESTRICTED - For internal use only Successful Herbicide Weed Control • Why does the herbicide work better one year and not the next? • What changes from year to year? • Is your equipment in good working order? • Just keeping on doing what you always have done… • And get what you always got! DOW RESTRICTED - For internal use only The Variables • Weather • Growing conditions • Temperatures • Wind • Water volume per acre applied • Equipment wear and calibration • You, you, you DOW RESTRICTED - For internal use only Start with the End in Mind • Visualize what you want to accomplish with every spray job – set expectations • Review records and see what has changed • Are the growing conditions and weeds conducive to weed control? • Recalibrate every year and even do a check up during the spray season • Calibrate every applicator that applies products DOW RESTRICTED - For internal use only Rates, Carrier & Equipment • Select herbicide product that: – controls the majority of target weeds – use the correct rates (don’t scrimp) • Water carrier per acre – use enough to get the product to the desired site of activity (if herbicide is soil active, it must penetrate the vegetative canopy to get benefit) – Use Syltac or R-11 • Tune-up sprayers and do timely repairs DOW RESTRICTED - For internal use only Adjuvants/Surfactants • Always use Syltac or R-11 • Use
    [Show full text]
  • Crops Before Corn an INVESTIGATION of EASTERN AGRICULTURAL COMPLEX PLANTS at the TOM JONES SITE, ARKANSAS
    Crops Before Corn AN INVESTIGATION OF EASTERN AGRICULTURAL COMPLEX PLANTS AT THE TOM JONES SITE, ARKANSAS ROSIE BLEWITT-GOLSCH Corn, Beans, and Squash Eastern Agricultural Complex • Initial domestication ca. 4000 – 3000 BP • Earliest occurrences found in upper South and lower Midwest • Initial evidence of EAC from rock shelters, but difficult to prove that they were actually grown as crops in prehistory Wild-Cultivated-Domesticated Spectrum Activities Outcomes • No intervention • Cultivation - preparing a seed bed, garden, field, • Encourage/tend planting the seeds Clearing out competing however, identical trees to wild plants Bringing water to plants • Domestication - putting during drought different selective Pruning plants pressures on plants and animals; generally genetic Burning and/or phenotypic • Control reproduction plants dependent Transplanting on people Collecting and planting seeds Eastern Agricultural Complex Plant Species • Sumpweed (Iva annua) • Sunflower (Helianthus annus) • Maygrass (Phalaris caroliniana) • Goosefoot (Chenopodium berlandieri) • Knotweed (Polygonum erectum) • Little Barley (Hordeum pusillum) • Squash (Cucurbita pepo) Squash and Gourds • Bottle gourd (date?) • Squash (4400 BP) • Produce starchy seeds but not much flesh • Rinds can be used as floats or containers • Domestication marked by seed enlargement and increase in rind thickness Squash Bottle Gourd Sunflower and Sumpweed • Sunflower (4300 BP) • Sumpweed (3900 BP) • Oily seeds • Domestication marked by increase in seed size Sunflower Sumpweed Goosefoot and Erect Knotweed • Goosefoot (3500 BP) • Erect knotweed (2500 BP) • Starchy seeds, Fall food source • Seed coat thickness, larger perisperm Knotweed Goosefoot Goosefoot Wild goosefoot Domesticated goosefoot Domesticated? variant Wild? variant Erect knotweed – two phenotypes Longer/shorter Thick/thin seed coat Maygrass and Little Barley • Maygrass (date?) • Little barley (date?) • Starchy seeds, Spring resources • Geographic range, quantity little barley Maygrass WHEAT Middle East Domesticated ~7,000 B.C.
    [Show full text]
  • Mistaken Identity? Invasive Plants and Their Native Look-Alikes: an Identification Guide for the Mid-Atlantic
    Mistaken Identity ? Invasive Plants and their Native Look-alikes an Identification Guide for the Mid-Atlantic Matthew Sarver Amanda Treher Lenny Wilson Robert Naczi Faith B. Kuehn www.nrcs.usda.gov http://dda.delaware.gov www.dsu.edu www.dehort.org www.delawareinvasives.net Published by: Delaware Department Agriculture • November 2008 In collaboration with: Claude E. Phillips Herbarium at Delaware State University • Delaware Center for Horticulture Funded by: U.S. Department of Agriculture Natural Resources Conservation Service Cover Photos: Front: Aralia elata leaf (Inset, l-r: Aralia elata habit; Aralia spinosa infloresence, Aralia elata stem) Back: Aralia spinosa habit TABLE OF CONTENTS About this Guide ............................1 Introduction What Exactly is an Invasive Plant? ..................................................................................................................2 What Impacts do Invasives Have? ..................................................................................................................2 The Mid-Atlantic Invasive Flora......................................................................................................................3 Identification of Invasives ..............................................................................................................................4 You Can Make a Difference..............................................................................................................................5 Plant Profiles Trees Norway Maple vs. Sugar
    [Show full text]
  • Grasses of the Texas Hill Country: Vegetative Key and Descriptions
    Hagenbuch, K.W. and D.E. Lemke. 2015. Grasses of the Texas Hill Country: Vegetative key and descriptions. Phytoneuron 2015-4: 1–93. Published 7 January 2015. ISSN 2153 733X GRASSES OF THE TEXAS HILL COUNTRY: VEGETATIVE KEY AND DESCRIPTIONS KARL W. HAGENBUCH Department of Biological Sciences San Antonio College 1300 San Pedro Avenue San Antonio, Texas 78212-4299 [email protected] DAVID E. LEMKE Department of Biology Texas State University 601 University Drive San Marcos, Texas 78666-4684 [email protected] ABSTRACT A key and a set of descriptions, based solely on vegetative characteristics, is provided for the identification of 66 genera and 160 grass species, both native and naturalized, of the Texas Hill Country. The principal characters used (features of longevity, growth form, roots, rhizomes and stolons, culms, leaf sheaths, collars, auricles, ligules, leaf blades, vernation, vestiture, and habitat) are discussed and illustrated. This treatment should prove useful at times when reproductive material is not available. Because of its size and variation in environmental conditions, Texas provides habitat for well over 700 species of grasses (Shaw 2012). For identification purposes, the works of Correll and Johnston (1970); Gould (1975) and, more recently, Shaw (2012) treat Texas grasses in their entirety. In addition to these comprehensive works, regional taxonomic treatments have been done for the grasses of the Cross Timbers and Prairies (Hignight et al. 1988), the South Texas Brush Country (Lonard 1993; Everitt et al. 2011), the Gulf Prairies and Marshes (Hatch et al. 1999), and the Trans-Pecos (Powell 1994) natural regions. In these, as well as in numerous other manuals and keys, accurate identification of grass species depends on the availability of reproductive material.
    [Show full text]
  • Redalyc.Tree and Tree-Like Species of Mexico: Apocynaceae, Cactaceae
    Revista Mexicana de Biodiversidad ISSN: 1870-3453 [email protected] Universidad Nacional Autónoma de México México Ricker, Martin; Valencia-Avalos, Susana; Hernández, Héctor M.; Gómez-Hinostrosa, Carlos; Martínez-Salas, Esteban M.; Alvarado-Cárdenas, Leonardo O.; Wallnöfer, Bruno; Ramos, Clara H.; Mendoza, Pilar E. Tree and tree-like species of Mexico: Apocynaceae, Cactaceae, Ebenaceae, Fagaceae, and Sapotaceae Revista Mexicana de Biodiversidad, vol. 87, núm. 4, diciembre, 2016, pp. 1189-1202 Universidad Nacional Autónoma de México Distrito Federal, México Available in: http://www.redalyc.org/articulo.oa?id=42548632003 How to cite Complete issue Scientific Information System More information about this article Network of Scientific Journals from Latin America, the Caribbean, Spain and Portugal Journal's homepage in redalyc.org Non-profit academic project, developed under the open access initiative Available online at www.sciencedirect.com Revista Mexicana de Biodiversidad Revista Mexicana de Biodiversidad 87 (2016) 1189–1202 www.ib.unam.mx/revista/ Taxonomy and systematics Tree and tree-like species of Mexico: Apocynaceae, Cactaceae, Ebenaceae, Fagaceae, and Sapotaceae Especies arbóreas y arborescentes de México: Apocynaceae, Cactaceae, Ebenaceae, Fagaceae y Sapotaceae a,∗ b a a Martin Ricker , Susana Valencia-Avalos , Héctor M. Hernández , Carlos Gómez-Hinostrosa , a b c Esteban M. Martínez-Salas , Leonardo O. Alvarado-Cárdenas , Bruno Wallnöfer , a a Clara H. Ramos , Pilar E. Mendoza a Herbario Nacional de México (MEXU), Departamento
    [Show full text]
  • Las Palmeras En El Marco De La Investigacion Para El
    REVISTA PERUANA DE BIOLOGÍA Rev. peru: biol. ISSN 1561-0837 Volumen 15 Noviembre, 2008 Suplemento 1 Las palmeras en el marco de la investigación para el desarrollo en América del Sur Contenido Editorial 3 Las comunidades y sus revistas científicas 1he scienrific cornmuniries and their journals Leonardo Romero Presentación 5 Laspalmeras en el marco de la investigación para el desarrollo en América del Sur 1he palrns within the framework ofresearch for development in South America Francis Kahny CésarArana Trabajos originales 7 Laspalmeras de América del Sur: diversidad, distribución e historia evolutiva 1he palms ofSouth America: diversiry, disrriburíon and evolutionary history Jean-Christopbe Pintaud, Gloria Galeano, Henrik Balslev, Rodrigo Bemal, Fmn Borchseníus, Evandro Ferreira, Jean-Jacques de Gran~e, Kember Mejía, BettyMillán, Mónica Moraes, Larry Noblick, FredW; Staufl'er y Francis Kahn . 31 1he genus Astrocaryum (Arecaceae) El género Astrocaryum (Arecaceae) . Francis Kahn 49 1he genus Hexopetion Burret (Arecaceae) El género Hexopetion Burret (Arecaceae) Jean-Cbristopbe Pintand, Betty MiJJány Francls Kahn 55 An overview ofthe raxonomy ofAttalea (Arecaceae) Una visión general de la taxonomía de Attalea (Arecaceae) Jean-Christopbe Pintaud 65 Novelties in the genus Ceroxylon (Arecaceae) from Peru, with description ofa new species Novedades en el género Ceroxylon (Arecaceae) del Perú, con la descripción de una nueva especie Gloria Galeano, MariaJosé Sanín, Kember Mejía, Jean-Cbristopbe Pintaud and Betty MiJJán '73 Estatus taxonómico
    [Show full text]
  • Discover Illinois Archaeology
    Discover Illinois Archaeology ILLINOIS ASSOCIATION FOR ADVANCEMENT OF ARCHAEOLOGY ILLINOIS ARCHAEOLOGICAL SURVEY Discover Illinois Archaeology Illinois’ rich cultural heritage began more collaborative effort by 18 archaeologists from than 12,000 years ago with the arrival of the across the state, with a major contribution by ancestors of today’s Native Americans. We learn Design Editor Kelvin Sampson. Along with sum- about them through investigations of the remains maries of each cultural period and highlights of they left behind, which range from monumental regional archaeological research, we include a earthworks with large river-valley settlements to short list of internet and print resources. A more a fragment of an ancient stone tool. After the extensive reading list can be found at the Illinois arrival of European explorers in the late 1600s, a Association for Advancement of Archaeology succession of diverse settlers added to our cul- web site www.museum.state.il.us/iaaa/DIA.pdf. tural heritage, leading to our modern urban com- We hope that by reading this summary of munities and the landscape we see today. Ar- Illinois archaeology, visiting a nearby archaeo- chaeological studies allow us to reconstruct past logical site or museum exhibit, and participating environments and ways of life, study the rela- in Illinois Archaeology Awareness Month pro- tionship between people of various cultures, and grams each September, you will become actively investigate how and why cultures rise and fall. engaged in Illinois’ diverse past and DISCOVER DISCOVER ILLINOIS ARCHAEOLOGY, ILLINOIS ARCHAEOLOGY. summarizing Illinois culture history, is truly a Alice Berkson Michael D. Wiant IIILLINOIS AAASSOCIATION FOR CONTENTS AAADVANCEMENT OF INTRODUCTION.
    [Show full text]
  • An Updated Infrageneric Classification of the North American Oaks
    Article An Updated Infrageneric Classification of the North American Oaks (Quercus Subgenus Quercus): Review of the Contribution of Phylogenomic Data to Biogeography and Species Diversity Paul S. Manos 1,* and Andrew L. Hipp 2 1 Department of Biology, Duke University, 330 Bio Sci Bldg, Durham, NC 27708, USA 2 The Morton Arboretum, Center for Tree Science, 4100 Illinois 53, Lisle, IL 60532, USA; [email protected] * Correspondence: [email protected] Abstract: The oak flora of North America north of Mexico is both phylogenetically diverse and species-rich, including 92 species placed in five sections of subgenus Quercus, the oak clade centered on the Americas. Despite phylogenetic and taxonomic progress on the genus over the past 45 years, classification of species at the subsectional level remains unchanged since the early treatments by WL Trelease, AA Camus, and CH Muller. In recent work, we used a RAD-seq based phylogeny including 250 species sampled from throughout the Americas and Eurasia to reconstruct the timing and biogeography of the North American oak radiation. This work demonstrates that the North American oak flora comprises mostly regional species radiations with limited phylogenetic affinities to Mexican clades, and two sister group connections to Eurasia. Using this framework, we describe the regional patterns of oak diversity within North America and formally classify 62 species into nine major North American subsections within sections Lobatae (the red oaks) and Quercus (the Citation: Manos, P.S.; Hipp, A.L. An Quercus Updated Infrageneric Classification white oaks), the two largest sections of subgenus . We also distill emerging evolutionary and of the North American Oaks (Quercus biogeographic patterns based on the impact of phylogenomic data on the systematics of multiple Subgenus Quercus): Review of the species complexes and instances of hybridization.
    [Show full text]
  • An Overview of the Taxonomy of Attalea (Arecaceae)
    Rev. peru. biol. 15(supl. 1): 055- 063 (Noviembre 2008) Las palmeras en América del Sur An overviewVersión of the Online taxonomy ISSN of1727-9933 ATTALEA © Facultad de Ciencias Biológicas UNMSM An overview of the taxonomy of Attalea (Arecaceae) Una visión general de la taxonomía de Attalea (Arecaceae) Jean-Christophe Pintaud IRD, UMR DIA-PC/DYNADIV, La- boratoire Genetrop, Centre IRD de Abstract Montpellier, 911 Av. Agropolis, BP The genus Attalea (Arecaceae) is distributed in continental habitats of the Neotropical region and in some 64501, 34394 Montpellier Cedex 5, France. Caribbean islands. Life forms of Attalea species vary from small acaulescent palms to tall and massive palms, always solitary. The ecological range of the genus encompasses most of the Neotropical ecosystems, from coastal sand dunes to sub-Andean forests up to 1600 m in elevation, lowland wet to dry forests, savannas, swamps, etc. The taxonomy of the genus has been poorly understood due to conflicting genus and species concepts that exist since the last decades. Taxonomical problems have been caused by the lack of adequate material, especially species of large size, loss of many types and difficulties in interpreting hybrids. In this article, I review the most recent taxonomic literature on Attalea. The number of species in Attalea varies from 29 to 67 depending on different authors, with a maximum estimate of 73 species when combining the revised publications. There is a consensus for the validity of 20 species among modern palm taxonomists. The most conflicting species or group of species are discussed in detail as well as the taxonomic significance of some characters such as the pattern of insertion of staminate flowers on rachillae, insertion of pinnae on rachis, and arrangement of fibrous strands in the endocarp.
    [Show full text]