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Phuket Mar. Biol. Cent. Res. Bull. 75: 27–44 (2018) 3

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Phuket Mar. Biol. Cent. Res. Bull. 75: 27–44 (2018) 3 Phuket mar. biol. Cent. Res. Bull. 75: 27–44 (2018) 3 NEW RECORDS OF AMMONOIDS (CEPHALOPODA: AMMONOIDEA) FROM PENINSULAR THAILAND Kittichai Tongtherm1, Jaruwat Nabhitabhata2*, Paladej Srisuk3 and Thammarat Nutadhira4 1Department of Biology, Faculty of Science, Prince of Songkla University, Hatyai, Songkhla 90112, Thailand 2Excellence Centre for Biodiversity of Peninsular Thailand (CBIPT), Faculty of Science, Prince of Songkla University, Hatyai, Songkhla 90112, Thailand 3Palaeontological Research and Education Center, Mahasarakham University, Kantarawichai, Mahasarakham 44150, Thailand 4Natural History Museum, Kampangwittaya School, La-gnu, Satun 91110, Thailand *Corresponding Author: [email protected], [email protected] ABSTRACT: Four taxa of ammonoid cephalopods have been discovered in the Devonian, Carboniferous, Permian and Triassic strata in Peninsular Thailand. All taxa are new records of ammonoids in Thailand. They comprise the Devonian ammonnoid ?Parawocklumeriidae gen. et sp. indet., the Carboniferous form Goniatites cf. eganensis Korn and Titus, 2011, the Permian ammonoid Cyclolobus sp. indet. from Satun province, on the western part of the peninsula, and the Triassic ammonoid Ceratitina gen. et sp. indet. from Phatthalung province, on the eastern part of the peninsula. Key words: new records, ammonoids, Cephalopoda, peninsular Thailand. INTRODUCTION IURP .RK 0DSUDR R൶FLDOO\ .R 3UDR Phuket province. However, this species name Several authors have documented ammonoids might currently be a synonym of a marine caeno- IURP 7KDLODQG 7KH ¿UVW UHFRUG RI DPPRQRLG gastropod, Hipponix antiquatus (Linnaeus, 1767) cephalopod fossils in Thailand was reported by of the family Hipponicidae (WoRMS, 2017). All Reed (1920), based on specimens from Carbonif- published occurrences are listed in Table 1. Am- erous strata in Srinagarindra district, Phatthalung monoids are overall rather rare in Thailand, thus province, on the eastern part of the peninsula (Ta- making almost every new record relevant for a ble 1). The record comprised three taxa, Goniatites publication. (?) sp. (as Glyphioceras (?) sp.), Prolecanites (?) This study focuses on identification and sp. and PronoritesD൵cyclolobus (Phillips, 1836). description of newly recorded ammonoids, adding Later, Suvatti (1938; 1967) reported an ammonoid information to previous records of cephalopod fauna of unknown age: Amalthea antiquate Linnaeus, in Peninsular Thailand. 28 Phuket mar. biol. Cent. Res. Bull. 2 Table 1. Previous records of ammonoids in Thailand. Locality Age Taxon References (Province, Region) - Amalthea antiquate Linnaeus, 1767 Phuket, Suvatti (1938; (Gastropod?) Southwestern 1967) Carboniferous Glyphioceras (?) sp., Prolecanites (?) sp., Phatthalung, Reed (1920) Pronorites D൵. cyclolobus (Phillips, 1836) Southeastern Goniatites sp. (as Glyphioceras (?) sp.), Phatthalung, Pitakpaivan et al. Prolecanites (?) sp. Southeastern (1969) Pronorites D൵. cyclolobus (Phillips, 1836) Stenopronorites uralensis (Karpinsky 1889) Mae Hong Son, Ingavat-Helmcke (as Pronorites D൵. uralensis Karpinsky 1889) Northern (1994) Protocanites sp. Mae Hong Son, Ishibashi et al. Northern (1997) Branneroceras sp., Bisatoceras sp., Loei, Northeastern Neogastrioceras sp., Syngastrioceras sp. Bisatoceras sp., Neogastrioceras sp., Loei, Northeastern Fujikawa et al. Syngastrioceras sp., (1999) Branneroceras branneri Smith, 1896, Diaboloceras involutum Nassichuk 1975, Cravenoceras (?) sp. Mae Hong Son, Fujikawa and Northern Ishibashi (1999 cited after Fujikawa and Ishibashi, 2000) Branneroceras (?) sp., Cravenoceras (?) sp., Loei, Northeastern Gastrioceras spp., Paralegoceras sp., Pseudoparalegoceras sp. Stenopronorites cf. uralensis (Karpinsky, Satun, House (2004) in 1889), Southwestern Wongwanich et al. Syngastrioceras sp. indet., ?Glaphyritid indet. (2004) Neogluphioceras subcirculare (Miller, 1889) Satun, Tongtherm et al. Southwestern (2016) Permian AgathicerasD൵. suessi Gemmellaro, 1887 Loei and Pitakpaivan et al. Khon Kaen, (1969) Northeastern; Propinacoceras sp. Khon Kaen, Northeastern Agathiceras mediterraneum Tumanskaya, Nakhon Glenister et al. 1949, Ratchasima, (1990) Miklukhoceras cf. pamiricum Pavlov, 1967, Northeastern Perrinites cf. hilli Smith, 1903 Prostacheoceras cf. oshense (Toumanskaya, 1938) 3 30 Phuket mar. biol. Cent. Res. Bull. 2 Locality Age Taxon References (Province, Region) AgathicerasD൵. suessi Gemmellaro, 1887 Loei, Northeastern and Phatthalung, Southeastern Agathiceras mediterraneum Tumanskaya, Nakhon Zhou and 1949, Ratchasima, Liengjarern (1997) Miklukhoceras cf. pamiricum Pavlov, 1967, Northeastern Perrinites cf. hilli Smith, 1903, Prostacheoceras cf. oshense (Tumanskaya, 1938) Huananoceras sp., Protoceras sp., Lampang, Fujikawa and Pseudotirolotes sp., Northern Ishibashi (1999 ']KX¿WHVD൵. spinosus Shevyrev, 1965 cited after (as Paratirolites D൵. spinosus Shevyrev, 1965), Fujikawa and Pseudogastrioceras D൵. szechuanense Chao, Ishibashi, 2000) 1965, 7DSDVKDQLWHVÀRULIRUPLV Zhao and Liang, 1965 Tapashanites changxingensis , Zhao, Liang, and Zheng, 1978 Tapashanites sp. , Xenodiscus sp. Aganthiceras sp., Prostacheoceras spp., Loei, Northeastern Zhou and Akmilleria electraensis (Plummer and Scott, Liengjarern (2004) 1937), Bamyaniceras loeiense (Ishibashi, Fujikawa and Nakornsri, 1996), Metaperrinites ishibashi Zhou and Liengjarern, 2004, Neopronorites cf. darvasicus Leonova, 1988, Properrinites boesei Plummer and Scott, 1937, Popanoceras cf. sobolewskyanum (de Verneuil, 1845), (as Popanoceras cf. sobolewskyanus (de Verneuil, 1845)), Sicanites cf. notabilis Ruzhencev, 1940 Agathiceras mediterraneum Toumanskaya, Nakhon 1949 Ratchasima, Neocrimites nalivkini (Toumanskaya, 1949), Northeastern Parapronorites rectus Leonova, 1989, Perrinites tardus Miller and Furnish, 1940, Prostacheoceras alter Leonova, 1989 Perrinites gouldi Miller and Furnish, 1940 Saraburi, Central Agathiceras suessi Gemmellaro, 1887, Phatthalung, Fujikawa et al. Agathiceras girtyi Böse, 1919, Southeastern (2005) Aganthiceras? sp., Miklukhoceras sp., Neocrimites sp. Tapashanites yaowalakae Ishibashi, Nakornsri Lampang, DMR (2006) and Nagai, 1994 Northern 31 New records of ammonoids (Cephalopoda: Ammonoidea) from Peninsular Thailand 3 Locality Age Taxon References (Province, Region) Bamyaniceras loeiense (Ishibashi, Loei, Northeastern Fujikawa and Nakornsri, 1996), (as Artinskia loeiensis Ishibashi, Fujikawa, and Nakornsri, 1996), Metaperrinites? sp., Agathiceras sp. Loei, Northeastern Fujikawa and Ishibashi (2006) Triassic Balatonites sp. indet. I, II, Beyrichites sp., Lampang, Kummel (1960) Sturia sp. indet., Northern Cladiscites cf. beyrichi Welter, 1914, Joannites cf. klipsteini (Mojsisovics, 1875), Lobites cf. ellipticus (Hauer, 1860), Ptychites cf. rectangulatus Kraus, 1916, Trachyceras (Paratrachyceras) cf. regoledanum (Mojsisovics, 1869), Tropigymnites cf. chandra (Diener, 1895) Beyrichites sp. indet., Balatonites sp. indet. I, II, Lampang, Pitakpaivan et al. Sturia sp. indet., Northern (1969) Cladiscites cf. beyrichi Welter, 1914, Joannites cf. klipsteini (Mojsisovics, 1875), Lobites cf. ellipticus (Hauer, 1860), Ptychites cf. rectangulatus Kraus, 1916, Trachyceras (Paratrachyceras) cf. regoledanum (Mojsisovics, 1869), Tropigymnites cf. chandra (Diener, 1895) BalatonitesD൵. balatonicum (Mojsisovics, Northern Komalarjun 1873), and Sato (1964 Balatonites sp., Leiophyllites sp., cited after Ophiceras sp., Chonglakmani, ClionitesD൵. barwicki Johnson, 1941, 1983) Joannites maehuatensis Chonglakmani, 1983, Joannies nonghoiensis Chonglakmani, 1983, Leiophyllites cf. pitamaha Diener, 1895, Ophiceras cf. chamunda Diener, 1897, Ophiceras (Discophiceras) sp. nov. cf. subkyokticum Spath, 1930, Paratrachyceras cf. regoledanum (Mojsisovics, 1931), Protrachyceras cf. longobardicum (Mojsisovics, 1882), Sirenites senticosus (Dittmar, 1866), Trachyceras cf. aon (Munster, 1834) Beyrichites cf. srikanta (Diener, 1895), Lampang, Beyrichites sp., Sturia sp. indet. Northern Trachyceras sp. Tak, Western 32 Phuket mar. biol. Cent. Res. Bull. 2 Locality Age Taxon References (Province, Region) Anatomites D൵. rotundus Mojsisovics, 1893, Northern Chonglakmani Anatomites sp., Anatomites sp. indet., (1983) Balatonites D൵. balatonicus Mojsisovics, 1873 (as Balatonites D൵. balatonicum Mojsisovics, 1873) Buchites sp. indet., Beyrichites ? sp., Beyrichites cf. srikanta (Diener) Griesbachites sp. indet., Hollandites cf. ravana (Diener) Hollandites cf. roxburghii (Diener) Hollandites cf. visvakarma (Diener) Joannites maehuatensis Chonglakmani, 1983 Joannies nonghoiensis Chonglakmani, 1983, Juvanites cf. idenburgi Welter Leiophyllites cf. pitamaha Diener, 1895 Lobites cf. ellipticus Hauer, 1860 Ophiceras (Discophiceras) cf. subkyoticum Spath, 1930 Ophiceras (Lytophiceras) cf. chamunda Diener Protachyceras cf. longobardicum Mojsisovics, Paratrachyceras cf. regoledanum Mojsisovics, Paratrachyceras sp., Sturia sp. indet., Ptychites cf. rectangulatus Kraus Sirenites (Sirenites) senticosus Dittmar, 1866 Trachyceras cf. aon Munster, 1834 Tropigymnites cf. chandra (Diener) Zestoceras barwicki Johnson, 1941 (as Clionites D൵. barwicki Johnson, 1941) Choristoceras ammonitiforme Guambel, Tak, Western 1861, Tmetoceras dhanarajatai Sato, 1964, Trachyceras sp. Aplococeras parrus (Smith, 1914), Phatthalung, Tongtherm et al. Arianites musacchi Arthaber, 1911, Southeastern (2016) Audaxlytoceras audax Meneghini, 1881, Bosnites clathratus Hyatt& Smith, 1897, (SLFHOWLWHVJHQ¿L Arthaber, 1911, Eutomoceras D൵. laubei Meek, 1877, Eutomoceras dunni Smith,
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