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Meso Template Flora Mesoamericana, Volumen 2 (2), Lepidobotryaceae, página 1 de 3 Última revisión del autor 12 oct. 2007. Initialmente publicado en el sitio Internet de la Flora Mesoamericana, 6 mar. 2012. 109. LEPIDOBOTRYACEAE Descripción de la familia por J.F. Morales. Árboles, dioicos, siempreverdes. Hojas alternas, simples (aparentemente unifoliolado- compuestas), enteras, articuladas con el pecíolo; estípulas en pares, pronto caducas; estipelas solitarias, pronto caducas. Inflorescencias terminales o subterminales, opuestas a las hojas. Flores pequeñas, actinomorfas, funcionalmente unisexuales; perianto en 2 verticilos, los sépalos y pétalos 5, imbricados; estambres 10, basalmente connatos en un tubo estaminal nectarífero, los filamentos antipétalos más largos que los filamentos antisépalos, las anteras basifijas (Mesoamérica) o versátiles; pistilo 1, el ovario súpero, 2(3)-locular, la placentación axial y apical, cada lóculo con 2 óvulos péndulos ovules; estilos ausentes o casi ausentes; estigmas capitados o inconspicuamente 2-lobados. Frutos en cápsulas septicidas; exocarpo y endocarpo separados, el exocarpo coriáaceo a leñoso, con dehiscencia irregular, el endocarpo cartilaginoso a cartáceo; semillas 1(2), negras, ariladas en el ápice; endospermo ausente. 2 gen., 2 spp. América tropical y África Occidental. El género monotípico Lepidobotrys está restringido a África Occidental. Algunos autores han considerado a la familia Lepidobotryaceae como parte de Oxalidaceae, pero en la actualidad se piensa que está más cercanamente relacionada con Celastraceae. 1. Ruptiliocarpon Hammel et N. Zamora Por J.F. Morales. Árboles; madera con médula con vestidura. Inflorescencias en panículas de espigas. Flores crípticamente unisexuales; gineceo 2-mero; filamentos relativamente cortos; anteras basifijas, el conectivo apiculado; estilo no bien diferenciado. Cápsulas con el exocarpo leñoso y el endocarpo cartilaginoso. 1 sp. Mesoamérica, Colombia, Surinam, Ecuador, Perú. Bibliografía: Hammel, B.E. y Zamora, N.A. Novon 3: 408-417 (1993). Morales, J.F. Man. Pl. Costa Rica 6: 198-199 (2007). Flora Mesoamericana, Volumen 2 (2), Lepidobotryaceae, página 2 de 3 1. Ruptiliocarpon caracolito Hammel et N. Zamora, Novon 3: 408 (1993). Holotipo: Costa Rica, Herrera y Martínez 2310 (CR!). Ilustr.: Hammel y Zamora, Novon 3: 410, t. 1 (1993). N.v.: Caracolito, cedro caracolito, CR. Árboles 10-25 m; corteza generalmente grisácea, glabra; ramitas cilíndricas o subcilíndricas; estípulas 1-1.5 mm, ensiformes, prontamente caducas. Hojas unifolioladas, alternas; pecíolo 0.5- 1.6 cm, basalmente pulvinular; folíolo con el peciólulo 2-4 mm, pulvinular sobre toda su longitud, articulado donde se junta con el pecíolo, la estipela 4-5 mm, linear; lámina 6-16 × 3-7 cm, elíptica, obtusa a redondeada en la base, cuspidada a cuspidado-acuminada en el ápice, entera, glabrescente, pinnatinervia, los nervios terciarios conspicuamente reticulados en el envés. Inflorescencias en panículas laxas de 1-3(4) espigas 2.3-8(-10) cm; pedúnculo 1-2.2 cm. Flores crípticamente unisexuales, globosas a ovoides, cada una rodeada por 3 brácteas; sépalos 5, 1.5-2 mm, separados, imbricados, ciliados; pétalos 5, 3-4 mm, separados, imbricados distalmente, ciliados; estambres c. 3 mm (más cortos en las flores pistiladas), en 2 series alternas de longitud diferente, los filamentos unidos en un tubo nectarífero c. 0.5 mm; anteras sin polen en las flores pistiladas, el conectivo proyectado en un pequeño apéndice pubescente; ovario más pequeño en las flores estaminadas; óvulos 2 por lóculo (rudimentarios en las flores estaminadas); estigma indistintamente 2-lobulado. Capsulas septicidas, el exocarpo quebrándose irregularmente hasta exponer las 2 válvulas óseas del endocarpo, una que envuelve totalmente la semilla y la otra (generalmente) vacía, ambas que luego se desprenden; semilla generalmente 1, 2.5-3.3 cm, negra, ovoide a elipsoide, lustrosa, el 1/3 apical envuelto por un arilo rojo anaranjado, que se vuelve péndulo (por el arilo) al final de un tiro coriáceo (partición del ovario) conectado al receptáculo. Bosques muy húmedos, márgenes de ríos, quebradas y ocasional en pastizales. N (Rueda et al. 17950, MO); CR (Morales 7631, MO); P (McPherson 9513, MO). 0-1100 m. (Mesoamérica, Colombia, Ecuador, Perú.) Ruptiliocarpon caracolito se reconoce por las hojas unifolioladas, con el pecíolo y peciólulo pulvinulares, las inflorescencias paniculadas conformadas por espigas de flores muy pequeñas y los frutos capsulares con el exocarpo coriáceo a leñoso, que se abre en forma irregular al madurar hasta exponer las dos válvulas óseas del endocarpo (la más grande que parece la concha de un caracol y de ahí el nombre vernacular). Por las hojas unifolioladas, se puede confundir en estado estéril con varios miembros de la familia Fabaceae, p.ej., Lecointea amazonica Ducke, Lonchocarpus unifoliolatus Benth. o alguna especies de Swartzia. Flora Mesoamericana, Volumen 2 (2), Lepidobotryaceae, página 3 de 3 Es factible que el material de Perú y Surinam, casi todo en estado estéril, abarque una o dos spp. distintas, sin describir. .
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