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Generic Composition of the New World Archipini (Lepidoptera, Tortricidae) with Description of Two New Genera and Two New Species J

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Generic Composition of the New World Archipini (Lepidoptera, Tortricidae) with Description of Two New Genera and Two New Species J Miscel.lania Zoologica 20.1 (1997) 125 Generic composition of the New World Archipini (Lepidoptera, Tortricidae) with description of two new genera and two new species J. Razowski Razowski, J , 1997 Generic composition of the New World Archipini (Lepidoptera, Tortricidae) with description of two new genera and two new species MISCZool, 20 1 125-130 Gener~ccornpos~t~on of the New World Archlplni (Lepldoptera, Tortrlcldae) wlth descrlptlon of two new genera and two new spec~es- The geographic distribution of the genera of Archipini from the New World is discussed and compared with that of other regions The 11 genera found in the Nearctic subregion are al1 known frorn the Palaearctic subregion In the Neotropical region there are six genera only, three being endernic Two genera, viz, Tacertaenia with T polonorurn and Sychnovalva with S syrrhapta are described as new Key words Lepidoptera, Tortricidae, Archipini, New World (Rebut: 24 V 96; Acceptació cond~cional: 16 XI 96; Acc. definitiva: 28 1 97) J Razowskl, lnst of Systernatlcs and Evolut~onof An~rnals, Pol~shAcaderny of Sc~ences,31-016 Kraków, Slawkowska 17, Poland lntroduction subregion which is the best studied area, 25 genera occur of which 12 are shared The generic composition of the tortricid with the Oriental region. tribes of the New World has not been dis- The aim of this work is to present a cussed previously. Moreover, data on the preliminary summary of the generic com- Neotropical faunas are still very incomplete. position of the tribe Archipini in the New The tribe Archipini has a world-wide dis- World. Although a comparison with that of tribution and the majority of genera and other tribes is at present impossible, it is species occur in the Oriental region and the clear that the Archipini are poorly repre- Palaearctic subregion. In the Palaearctic sented in the Neotropical region. ISSN 021 1-6529 O 1997 Museu de Zoologia l - -- - 126 Razowski Material and methods gion, not expanding far south (except for one widely distributed species recorded in The present data are based on the material the south from California and Florida, and deposited in the largest entomological in- central Mexico). stitutions, which are mainly American, and The genus Pandemis is abundant in spe- in some private collections. cies in temperate and tropical Asia as well as in the Afrotropical region where it oc- curs in Madagascar and South Africa. The Distribution of the genera of Archipini Nearctic species reaches as far south as Cali- in the New World fornia. Two genera, Argyrotaenia, Stepyhens, The Archipini are represented by only 15 1852 and Clepsis, occur over a wide area genera in the New World. Only nine genera of the New World, the former from north- are recorded from the Nearctic subregion. ern Canada to Argentina and the latter to Cudonigera Obraztsov & Powell, 1977 is Peru. Argyrotaenia is represented in the endemic in this area. This monotypical genus Holarctic region by a single very widely is very close to Choristioneura Hübner, 1825 distributed Palaearctic species and by over differing only in one character of the fe- 35 Nearctic species. A similar number of male genitalia. species occurs in the Neotropical region. Two genera, viz., Batodes Guenée,1845 There are only slight differences between and probably Cacoecimorpha Obraztsov, the species and only a few South Ameri- 1954 were artificially introduced into North can species are better differentiated, par- America. ticularly concerning the genitalia. The Four genera (Archips Hübner, 1825, number of species of Clepsis increases Pandemis Hübner, 1825, Adoxophyes Mey- slightly from Canada to Mexico, and in rick, 1881 and Clepsis Guenée, 1845) are South America the genus is represented widely distributed in the Palaearctic sub- by five species in Peru, Venezuela and region and the Oriental region. The distri- Guatemala and five other species in Co- bution of Adoxophyes is even wider as there lombia. On basis of unexamined material are several species in the Australian region the author assumes that this genus is more and in North America there are two species abundant in species, specially in the moun- distributed in its northern parts. tains of Peru and Colombia. Three of the The genera Syndemis Hübner, 1825 and species closely related with C. smicrotes Aphelia Hübner, 1825 are Holarctic in dis- (Walsingham, 1914) occur in Canada and tribution; the former is represented in 14 in Mexico. All Central American and the Palaearctic subregion by a single spe- South American species belong to this cies, and the latter by numerous species group. Several species described or placed and al1 subgenera. The Nearctic species of in Smicrotes Clemens, 1860 have been Aphelia Hübner belong to two subgenera; incorrectly included in P~cholomaSte- one species is included in Zolotherses phens, 1829 which is exclusively a Palaearctic Lederer, 1859 four in Aphelia s. str. They genus. Of the remaining species of Clepsis are widely distributed throughout North one is of Holarctic distribution, and six America from Alaska and Quebec to Texas. are Nearctic, al1 occurring in the northern Archips Hübner is known from al1 of and central areas of North America and Asia and in North America is represented more southwards in the mountains (e.g. by 21 species distributed mainly in the north- in California). ern and central parts of the subregion but Another Nearctic genus, Durangarchips reaching southwards as far as California, Powell, 1991 is monotypical and known Texas and Florida. only from Durango (Mexico). Choristoneura has a similar distribution In the Neotropical region, apart from but does not penetrate into the Oriental the last two genera, occurs ldolatteria Wal- region. Seventeen species occur in North singham, 1913 with eight species distrib- America and are known mainly from the uted between Guatemala and Bolivia and northern and central parts of the subre- two genera described below, al1 most prob- Miscel.lania Zoologica 20.1 (1997) ably endemic in this region. They are de- der, apophyses moderate; sterigma rather scribed from Southern Brazil, but their range broad in median part, with slender lateral may be much larger. arms well sclerotized along anterior edges, As it has not been possible to examine ostium bursae very small; colliculum small, any specimens of Nesochoris Clarke, 1965 membranous; ductus bursae rather short the systematic position of this genus can- with ductus seminalis extending post-medi- not yet be discussed. ally; signum absent. The tribe Archipini, seems to be repre- sented in the New World by a rather small Remarks number of the genera and species. Some Supposedly closest to Argyrotaenia as the endemic small genera may yet be discov- shape of forewings, coloration and sexual ered in very little studied territories, spe- dimorphism show. The genitalia are similar cially in the mountains of the western part only in the shape of transtilla, aedeagus of the Neotropical region. Many new spe- and bursa copulatrix which, however, is cies may be added to Argyrotaenia and variable in Argyrotaenia (the sclerite of the Clepsis. More comprehensive collecting may basal part of ductus bursae is often miss- add many taxa as it did, for instance, in ing). The supposed autapomorphies are Mexico. the broad, rounded apically dista1 parts of the uncus, the arms of gnathos connected by means of a membrane, the atrophy of Descriptions of new taxa median, terminal part of the gnathos, the enlarged, somewhat stronger sclerotized Tacertaenia n. gen. dorsal termination of the valva. The ab- sence of the cornuti and the signum are Type-species: Tacertaenia polonorum n. sp. convergent reductions often observed in this subfamily. The new genus is also close Externally similar to the representatives to the new genus described below although of Argyrotaenia Stephens. Venation and it differs in the absence of median plate of sexual dimorphism as in the mentioned the gnathos and its lateral processes, and genus. the absence of the signum. Male genitalia (figs 1-3) Tegumen very broad, with small, narrow Tacertaenia polonorum n. sp. ends of pedunculi; uncus very large, bifid postmedially, with rounded apical parts; Male socius small, membranous, hairy; gnathos Alar expanse 13-15 mm; labial palpus as arms broad, with terminal parts plate- long as diameter of eye, ferruginous to rust shaped, apically rounded, connected with yellowish, head, thorax and forewing pat- one another by means of broad membrane, tern concolorous; the latter consisting of without medial plate; vinculum small, as in dorso-basal suffusion, median fascia inter- Argyrotaenia; valva elongate, with dorso- rupted subcostally and subtriangular, large terminal part expanding slightly dorsally, subapical blotch; ground-colour much paler more strongly sclerotized than the remain- and creamer, sprinkled rust; fringes in torna1 ing costal area, sacculus as long as valva, half concolorous with ground-colour, other- angulate at its end, extending dorsally wise with pattern. Hindwing dirty cream, along its caudal edge, provided with minute tinged grey in anal area, with yellow in free termination, fold distinct, pulvinus ill- remaining periphery; fringes concolourous defined; transtilla band-shaped, broaden- with adjacent parts of wing. Variation: pat- ing slightly basally; juxta ovate; aedeagus tern occasionally brown-grey, more or less slender, curved, with short coecum penis, distinct, ground-colour at dorsum pale. Fe- cornuti missing. male (17-19 mm) almost unicolorous rust, or with weak darker pattern before apex and Female genitalia (fig. 4) tornus; hindwing orange-ferruginous, paler Eighth tergite and papilla analis large, slen- basally, brown-grey in anal area. l 128 Razowski Figs. 1-7. Male and female genitalia. 1-4. Tacertaenia polonorum n. sp.: 1-3. cf paratype.; 4. 9 paratype. 5-7. Sychnovalva syrrhapta n. sp.: 5,6. cf holotype; 7. 9 paratype. Genitalia del macho y de la hembra. 1-4. Tacertaenia polonorum sp.
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