Chapter III —Order Ephemeroptera
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Introduction
INTRODUCTION A BRIEF HISTORY OF MAYFLY of larvae could be made by the investigator, who did CLASSIFICATION not know the taxonomic significance of many characters shown on them. Many supraspecies taxa established by Eaton were natural, although they did In the early classifications (Linnaeus 1758, et al.) all not have sufficient diagnoses. mayflies, constituting a single holophyletic genus Later (Lestage 1917, et al.) ephemeropterologists Ephemera Linnaeus 1758 (placed to artificial order paid more and more attention to larval characters Neuroptera), were divided into two groups according rather than to imaginal ones, and established classifi- to the number of imaginal caudalii – 3 or 2. Each of cations based mainly or solely on larval characters. these groups was actually polyphyletic. The imagi- Since the artificial Linnaean order Neuroptera nal paracercus is developed in the majority of Euro- was completely divided into smaller natural orders pean Furcatergaliae and vestigial in the majority of (the process started by Burmeister 1829, and European Tridentiseta and Branchitergaliae; thus if finished by Packard 1886 and Handlirsch 1903), one studies superficially the European species only, mayflies got ordinal rank and were divided into a an impression could appear that this character allows number of families and superfamilies, which in large one to divide mayflies into natural groups. However, degree corresponded to sections, series and groups more detailed examination of mayflies reveals that proposed by Eaton (1883–1888) to the former family representatives with 3 and 2 caudalii occur in many Ephemeridae. Basing mainly on larval characters, evidently holophyletic taxa (see Index of characters authors of new classifications changed many of [2.3.20]). -
The Mayfly Newsletter: Vol
Volume 20 | Issue 2 Article 1 1-9-2018 The aM yfly Newsletter Donna J. Giberson The Permanent Committee of the International Conferences on Ephemeroptera, [email protected] Follow this and additional works at: https://dc.swosu.edu/mayfly Part of the Biology Commons, Entomology Commons, Systems Biology Commons, and the Zoology Commons Recommended Citation Giberson, Donna J. (2018) "The aM yfly eN wsletter," The Mayfly Newsletter: Vol. 20 : Iss. 2 , Article 1. Available at: https://dc.swosu.edu/mayfly/vol20/iss2/1 This Article is brought to you for free and open access by the Newsletters at SWOSU Digital Commons. It has been accepted for inclusion in The Mayfly eN wsletter by an authorized editor of SWOSU Digital Commons. An ADA compliant document is available upon request. For more information, please contact [email protected]. The Mayfly Newsletter Vol. 20(2) Winter 2017 The Mayfly Newsletter is the official newsletter of the Permanent Committee of the International Conferences on Ephemeroptera In this issue Project Updates: Development of new phylo- Project Updates genetic markers..................1 A new study of Ephemeroptera Development of new phylogenetic markers to uncover island in North West Algeria...........3 colonization histories by mayflies Sereina Rutschmann1, Harald Detering1 & Michael T. Monaghan2,3 Quest for a western mayfly to culture...............................4 1Department of Biochemistry, Genetics and Immunology, University of Vigo, Spain 2Leibniz-Institute of Freshwater Ecology and Inland Fisheries, Berlin, Germany 3 Joint International Conf. Berlin Center for Genomics in Biodiversity Research, Berlin, Germany Items for the silent auction at Email: [email protected]; [email protected]; [email protected] the Aracruz meeting (to sup- port the scholarship fund).....6 The diversification of evolutionary young species (<20 million years) is often poorly under- stood because standard molecular markers may not accurately reconstruct their evolutionary How to donate to the histories. -
Classification Scheme of Freshwater Aquatic Organisms Freshwater Keys: Classification
Compendium of Recommended Keys for British Columbia Freshwater Organisms: Part 3 Classification Scheme of Freshwater Aquatic Organisms Freshwater Keys: Classification Table of Contents TABLE OF CONTENTS.............................................................................................................................. 2 INTRODUCTION......................................................................................................................................... 4 KINGDOM MONERA................................................................................................................................. 5 KINGDOM PROTISTA............................................................................................................................... 5 KINGDOM FUNGI ...................................................................................................................................... 5 KINGDOM PLANTAE ................................................................................................................................ 6 KINGDOM ANIMALIA .............................................................................................................................. 8 SUBKINGDOM PARAZOA ........................................................................................................................ 8 SUBKINGDOM EUMETAZOA.................................................................................................................. 8 2 Freshwater Keys: Classification 3 Freshwater Keys: Classification -
Checklist of Mayflies (Ephemeroptera, Insecta) from Iraq
M.CHECKLIST S. Abdul-Rassoul Iraq Nat. Hist. Mus. Publ. (May, 2020) no. 37, 16 pp. Checklist of Mayflies (Ephemeroptera, Insecta) from Iraq M. S. Abdul-Rassoul Iraq Natural History Research Center and Museum (Emeritus professor), University of Baghdad, Baghdad, Iraq. Corresponding author: [email protected] Received Date: 05January 2020, Accepted Date: 11 May 2020, Published Date: 31 May 2020 Abstract This work provides the updated checklist of the mayfly (Ephemeroptera) fauna of Iraq; it consists of 30 species belonging to 18 genera and 10 families. The highest number of species belongs to the families of Baetidae and Heptageniidae, whereas the lowest in Ephemeridae, Leptophlebiidae and Palingeniidae that appeared with only one species for each of them. The current investigation is a contribution to the knowledge regarding the biodiversity of mayflies, with mentioning the synonyms of the species and correcting the scientific names that found in previous publications in Iraq. Keywords: Ephemeroptera, Insecta, Iraq, Mayflies, Museum. Introduction Ephemeroptera with two pairs of wings; the fore much wider than the hind wings, which in some genera are reduced in size or even wholly absent; venation system is of considerable importance in classification; further small veins locate between the main veins are termed intercalary veins. Mouth parts entirely vestigial; compound eyes of male larger than at female, divided in some species, and the 1 Checklist of Mayflies upper part prominent as turret shaped; forelegs in males often extremely long; abdominal terminates with two or three long and many jointed filaments, the median one in certain genera being reduced; the male carries a pair of jointed claspers arising from a plate on the 9thsternite (Kimmins, 1950). -
Toward a Phylogenetic Classification of the Ephemeroptera (Lnsecta): a Commentary on Systematics
FORUM Toward a Phylogenetic Classification of the Ephemeroptera (lnsecta): A Commentary on Systematics W. P. MCCAFFERTY Department of Entomology, Purdue University, West Lafayette, Indiana 47907 Ann. Entomol. Soc. Am. 84(4): 343-360 (1991) ABSTRACT Higher classifications of Ephemeroptera formulated in light of phylogenetic hypotheses have been essentially evolutionary in that they have incorporated paraphyletic taxa. The term "paraphyletic" has had a confused history because systematists have applied it to different concepts; for Ephemeroptera, it has denoted one type of nonpolyphyletic grouping. Such paraphyletic taxa were used so that large degrees of character (presumably adaptive) differentiation could be expressed, but resulting classifications lack any means of consistently expressing such gaps, and their ability to express genealogy is compromised. Phylogenetic classifications are now strongly advocated. Not only can the increasing number of phylogenetic hypotheses based on cladistic analysis be objectively expressed, but many traditional taxa and categories can be conserved by employing sequencing conventions. The philosophy and history of competing classificatory systems are discussed. A phylogenetic methodology is employed when possible in proposed revisions of the higher taxa of Ephem eroptera. New cladistic data are presented and previous data reviewed. Major lineages formerly part of the paraphyletic family Siphlonuridae are classified as families within several higher taxa to express phylogeny and at the same time maintain -
Landasoldan1985p1.Pdf
CESKOSLOVENSKA STUDIE CSAV AKADEMIE VED Phylogeny and higher classification of the order Ephemeroptera: a discussion from the comparative anatomical point of view CESKOSLOVENSKA AKADEMIE VED Scietific Editor Prof. Ing. Dr. Antonin Pfeffer, DrSc. Scientific Adviser Doc. RNDr. Jaroslav Hrbacek, CSc. Vladimir Landa and Tomas Soldan Phylogeny and higher classification of the order Ephemeroptera: a discussion from the comparative anatomical point of view ACADEMIA nakladatelstvi Ceskoslovenske akademie ved Praha 198.5 ©Vladimir Landa, Tomas Soldan 1985 Contents Introduction ••••••••••••••••••••••••••••••••••••••••••••••••••• • • • • 7 Review of higher classification of the Ephemeroptera................ 9 Material and methods • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • 23 Arrangemettt and anagenesis of organ systems studied •••••••••••••••• 26 Ventral nerve cord •• •......................................... 26 Tracheal system • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • 34 Alimentary canal ••••.••••••••••••••••••••••••••••••••••••• • • • • 43 Malpighian tubules • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • 49 Internal reproductive organs •••••••••••••••••••••••••••••••••• 58 Discussion •••••••••••••••••••••••••••••••••••••••••••••••••• • • • • • • • 69 Suborders ••••••••••••••••••••••••••••••••••••••••••••••••• • • · • 70 Superfamilies • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • -
The Higher Classification of the Ephemeroptera and Its Evolutionary Basis1''
The Higher Classification of the Ephemeroptera and Its Evolutionary Basis1'' W. P. McCAFFERTY3 AND GEORGE F. EDMUNDS, JR.4 ABSTRACT Ann. Entomol. Soc. Am. 72: 5-12 (1979) The higher classification of the extant Ephemeroptera of the world is reviewed. The su- borders Schistonota (to include the Baetoidea, Leptophlebioidea, and Ephemeroidea) and Pannota (to include the Ephemerelloidea, Caenoidea, and Prosopistomatoidea) are introduced and defined on the basis of thoracic structure and other characters. The families Ephemerel- lidae and Tricorythidae are removed from the Leptophlebioidea and placed as the Ephemer- elloidea. The phyletic basis for this classification and the evolutionary history of the super- families is presented. The Siphlonuridae and Leptophlebiidae are paraphyletic groups as constituted. All extant mayfly lineages take their origin within the Siphlonuridae and their derivations are discussed. The Leptophlebiidae gave rise to the Ephemeroidea. Classificatory modifications in the makeup of the Siphlonuridae, Siphlaenigmatidae, Baetidae, Ametropod- idae, Oligoneuriidae, Ephemerellidae, and Tricorythidae are discussed. On the basis of phy- Downloaded from https://academic.oup.com/aesa/article/72/1/5/136951 by guest on 02 October 2021 letic intermediacy, the Siphlaenigmatinae is recognized as a subfamily of Baetidae, and the Isonychiinae and Coloburiscinae are recognized as subfamilies of Oligoneuriidae. Evolutionary relationships are becoming relatively for these modifications, along with evidences for the well understood among higher groups of Ephemerop- paraphyletic nature of major stem-groups are treated tera. Thus, this group of insects is a model for studies herein. integrating phylogeny, classification, and biogeography. Suborders Reasons for this level of understanding are many. The order's size has helped make systematic studies ap- New evidence from the thorax along with other data proachable from a world perspective. -
Irish Biodiversity: a Taxonomic Inventory of Fauna
Irish Biodiversity: a taxonomic inventory of fauna Irish Wildlife Manual No. 38 Irish Biodiversity: a taxonomic inventory of fauna S. E. Ferriss, K. G. Smith, and T. P. Inskipp (editors) Citations: Ferriss, S. E., Smith K. G., & Inskipp T. P. (eds.) Irish Biodiversity: a taxonomic inventory of fauna. Irish Wildlife Manuals, No. 38. National Parks and Wildlife Service, Department of Environment, Heritage and Local Government, Dublin, Ireland. Section author (2009) Section title . In: Ferriss, S. E., Smith K. G., & Inskipp T. P. (eds.) Irish Biodiversity: a taxonomic inventory of fauna. Irish Wildlife Manuals, No. 38. National Parks and Wildlife Service, Department of Environment, Heritage and Local Government, Dublin, Ireland. Cover photos: © Kevin G. Smith and Sarah E. Ferriss Irish Wildlife Manuals Series Editors: N. Kingston and F. Marnell © National Parks and Wildlife Service 2009 ISSN 1393 - 6670 Inventory of Irish fauna ____________________ TABLE OF CONTENTS Executive Summary.............................................................................................................................................1 Acknowledgements.............................................................................................................................................2 Introduction ..........................................................................................................................................................3 Methodology........................................................................................................................................................................3 -
ORDEN EPHEMEROPTERA.Pdf
UNIVERSIDAD NACIONAL DE CAJAMARCA FACULTAD DE CIENCIAS AGRARIAS ESCUELA ACADÉMICO PROFESIONAL DE AGRONOMÍA EL ORDEN EPHEMEROPTERA EN LA CUENCA EL RONQUILLO - CAJAMARCA Para Optar el Título Profesional de: INGENIERO AGRÓNOMO Presentado por el Bachiller: ERICK MAURICIO MEZA PAREDES Asesores: Ing. Agr. Alonso Vela Ahumada Ing. Agr. Mg. Sc. Jhon Anthony Vergara Copacondori CAJAMARCA - PERÚ 2019 1 2 DEDICATORIA A mi madre por su apoyo constante. A mi padre, que desde el cielo guía mis pasos para poder seguir adelante. A mi pareja por su amor, paciencia y motivación en los momentos más difícil y finalmente a mis hermanos por inspirarme a cumplir mis metas. 3 AGRADECIMIENTO A mis asesores, por su tiempo, sabiduría, amistad y por su apoyo a culminar este proyecto de investigación. A mi madre y a mi pareja por su apoyo incondicional. 4 ÍNDICE GENERAL CONTENIDO Página DEDICATORIA AGRADECIMIENTO ÍNDICE GENERAL ÍNDICE DE TABLAS ÍNDICE DE FIGURAS ÍNDICE DE ANEXOS RESUMEN ABSTRACT CAPÍTULO I. INTRODUCCIÓN 1 1.1 Formulación del problema 1 1.2 Objetivo de la investigación 2 1.3 Hipótesis de la investigación 2 CAPÍTULO II. REVISIÓN DE LITERATURA 3 2.1 Antecedentes de la investigación 3 2.2 Bases teóricas 5 2.2.1 Orden Ephemeroptera 5 a. Morfología 6 a.1 Náyade 7 a.1.1 Cabeza 7 a.1.2 Tórax 7 a.1.3 Abdomen 8 a.2 Subimago 8 a.3 Imago o adulto 8 a.3.1 Cabeza 9 a.3.2 Tórax 9 a.3.3 Abdomen 9 b. Biología y ecología 10 b.1 Náyade 10 b.2 Subimago 11 b.3 Imago o adulto 11 c. -
Author's Personal Copy
Author's personal copy Provided for non-commercial research and educational use only. Not for reproduction, distribution or commercial use. This chapter was originally published in the book Thorp and Covich's Freshwater Invertebrates Volume 1. The copy attached is provided by Elsevier for the author's benefit and for the benefit of the author's institution, for non-commercial research, and educational use. This includes without limitation use in instruction at your institution, distribution to specific colleagues, and providing a copy to your institution's administrator. All other uses, reproduction and distribution, including without limitation commercial reprints, selling or licensing copies or access, or posting on open internet sites, your personal or institution’s website or repository, are prohibited. For exceptions, permission may be sought for such use through Elsevier’s permissions site at: http://www.elsevier.com/locate/permissionusematerial From Sartori, M., Brittain, J.E., 2015. Order Ephemeroptera. In: Thorp, J., Rogers, D.C. (Eds.), Ecology and General Biology: Thorp and Covich's Freshwater Invertebrates, Academic Press, 873–891. ISBN: 9780123850263 Copyright © 2015 Elsevier Inc. All rights reserved. Academic Press Author's personal copy Chapter 34 Order Ephemeroptera Michel Sartori Museum of Zoology, Palais de Rumine, Lausanne, Switzerland John E. Brittain Natural History Museum, University of Oslo, Oslo, Norway Chapter Outline Introduction To Mayflies (Ephemeroptera) 873 Flight Activity 883 Brief History and Paleontology -
Distribution and Biology of the Mayflies (Ephemeroptera) of Western Ghats
Distribution and Biology of the Mayflies (Ephemeroptera) of Western Ghats C. Balachandran, M. D. Subhash Chandran and T. V. Ramachandra Energy and Wetlands Research Group, Centre for Ecological Sciences, Indian Institute of Science, Bangalore – 560 012 Email – [email protected], [email protected], [email protected] Introduction Insects are perhaps the most successful creatures in the whole of the animal kingdom, considering their prolific numbers of species (of estimated eight million species, over one million species described) and individuals, their extraordinarily adaptations to live in diverse habitats of land, water and air. They are invertebrates (having no backbone), belonging to the Phylum Arthropods which are characterised by hard, protective outer shield and three pairs of jointed legs. Insects, which belong to a Class Insecta, are different from other arthropods, like spiders and mites, centipedes and millipedes, crabs and shrimps etc. which have more number of legs. Most insects also have wings, which are lacking in other arthropods. Winged insects have wider distributional coverage than their wingless counterparts. Insects play crucial role in ecosystem functions such as decomposition, nutrient cycling, pollination, predation, parasitism and maintenance of community structure wherever they live. Ephemeroptera is an order of insects, comprising of over 3000 species of insects, popularly referred to as mayflies, which are confined to fresh water, at least for part of their life cycles. The name Ephemeroptera (ephemeros means "short-lived", pteron is "wing") refers to the winged adults, which have lifespans from just 1-2 hours to few days, a maximum of 14 days. Adult mayfly species are terrestrial, the immature called naiads (nymph), are aquatic. -
Ephemeroptera)
Brigham Young University BYU ScholarsArchive Theses and Dissertations 2004-11-29 Evolution and phylogeny of basal winged insects with emphasis on mayflies (Ephemeroptera) Thomas H. Ogden Brigham Young University - Provo Follow this and additional works at: https://scholarsarchive.byu.edu/etd Part of the Biology Commons BYU ScholarsArchive Citation Ogden, Thomas H., "Evolution and phylogeny of basal winged insects with emphasis on mayflies (Ephemeroptera)" (2004). Theses and Dissertations. 207. https://scholarsarchive.byu.edu/etd/207 This Dissertation is brought to you for free and open access by BYU ScholarsArchive. It has been accepted for inclusion in Theses and Dissertations by an authorized administrator of BYU ScholarsArchive. For more information, please contact [email protected], [email protected]. EVOLUTION AND PHYLOGENY OF BASAL WINGED INSECTS WITH EMPHASIS ON MAYFLIES (EPHEMEROPTERA) By T. Heath Ogden A dissertation submitted to the faculty of Brigham Young University In partial fulfillment of the requirement for the degree of Doctor of Philosophy Department of Integrative Biology Brigham Young University December 2004 BRIGHAM YOUNG UNIVERSITY GRADUATE COMMITTEE APPROVAL of dissertation submitted by T. Heath Ogden This dissertation has been read by each member of the following graduate committee and by majority vote has been found to be satisfactory. __________________________ ________________________________________ Date Michael F. Whiting, Chair __________________________ ________________________________________